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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.224.102752" ID-Pensoft-Pub="1314-2003-224-183" ID-Pensoft-UUID="AB2BB31BE1A25528A6D78E191D0AEF91" ModsDocID="1314-2003-224-183" checkinTime="1680938996089" checkinUser="pensoft" docAuthor="Zou, Dian-yang, Cao, Guan-long, Zhang, Jin-guo, Li, Lang &amp; Li, Jie" docDate="2023" docId="26B9495DCE415454894380B415741874" docLanguage="en" docName="PhytoKeys 224: 183-192" docOrigin="PhytoKeys 224" docPubDate="2023-04-07" docSource="http://dx.doi.org/10.3897/phytokeys.224.102752" docTitle="Endiandra macrocarpa D. Y. Zou, Lang Li &amp; J. Li 2023, sp. nov." docType="treatment" docVersion="1" id="AB2BB31BE1A25528A6D78E191D0AEF91" lastPageNumber="183" masterDocId="AB2BB31BE1A25528A6D78E191D0AEF91" masterDocTitle="Endiandra macrocarpa (Lauraceae), a new tree species from south-western China" masterLastPageNumber="192" masterPageNumber="183" pageNumber="183" updateTime="1680938996089" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Endiandra macrocarpa (Lauraceae), a new tree species from south-western China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zou, Dian-yang</mods:namePart>
<mods:affiliation>Plant Phylogenetics and Conservation Group, Center for Integrative Conservation &amp; Yunnan Key Laboratory for Conservation of Tropical Rainforests and Asian Elephants, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Mengla, Yunnan 666303, China &amp; University of Chinese Academy of Sciences, Beijing 100049, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cao, Guan-long</mods:namePart>
<mods:affiliation>University of Chinese Academy of Sciences, Beijing 100049, China &amp; State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, Beijing 100093, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhang, Jin-guo</mods:namePart>
<mods:affiliation>Administration Bureau of Maguan Gulinqing Provincial Nature Reserve, Wenshan, Yunnan 663000, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Lang</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7439-2030</mods:nameIdentifier>
<mods:affiliation>Plant Phylogenetics and Conservation Group, Center for Integrative Conservation &amp; Yunnan Key Laboratory for Conservation of Tropical Rainforests and Asian Elephants, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Mengla, Yunnan 666303, China</mods:affiliation>
<mods:nameIdentifier type="email">lilang@xtbg.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Jie</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8067-749X</mods:nameIdentifier>
<mods:affiliation>Plant Phylogenetics and Conservation Group, Center for Integrative Conservation &amp; Yunnan Key Laboratory for Conservation of Tropical Rainforests and Asian Elephants, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Mengla, Yunnan 666303, China</mods:affiliation>
<mods:nameIdentifier type="email">jieli@xtbg.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-04-07</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>224</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>183</mods:start>
<mods:end>192</mods:end>
</mods:extent>
</mods:part>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.224.102752</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.224.102752</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-224-183</mods:identifier>
<mods:identifier type="Pensoft-UUID">AB2BB31BE1A25528A6D78E191D0AEF91</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:26B9495DCE415454894380B415741874" httpUri="http://treatment.plazi.org/id/26B9495DCE415454894380B415741874" lastPageNumber="183" pageId="0" pageNumber="183">
<subSubSection pageId="0" pageNumber="183" type="nomenclature">
<paragraph pageId="0" pageNumber="183">
<taxonomicName LSID="26B9495D-CE41-5454-8943-80B415741874" authority="D. Y. Zou, Lang Li &amp; J. Li" authorityName="D. Y. Zou, Lang Li &amp; J. Li" authorityYear="2023" class="Magnoliopsida" family="Lauraceae" genus="Endiandra" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endiandra macrocarpa" order="Laurales" pageId="0" pageNumber="183" phylum="Tracheophyta" rank="species" species="macrocarpa" status="sp. nov.">Endiandra macrocarpa D.Y.Zou, Lang Li &amp; J.Li</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="183">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="183" type="description">
<paragraph pageId="0" pageNumber="183">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Morphology of Endiandra macrocarpa A tree habit B flowering branchlet C fruiting branchlet displaying immature fruit D mature fruits. Photographed by Lang Li and Guan-long Cao." figureDoi="10.3897/phytokeys.224.102752.figure1" httpUri="https://binary.pensoft.net/fig/834766" pageId="0" pageNumber="183">Figs 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Morphology of a flower of Endiandra macrocarpa. A outer tepals, abaxial and adaxial side B inner tepals, abaxial and adaxial side C stamen, abaxial side D pistil E flower, top view F flower, longitudinal section. Photographed by Dian-yang Zou." figureDoi="10.3897/phytokeys.224.102752.figure2" httpUri="https://binary.pensoft.net/fig/834767" pageId="0" pageNumber="183">, 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="183" type="diagnosis">
<paragraph pageId="0" pageNumber="183">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="183">
Compared to other
<taxonomicName class="Magnoliopsida" family="Lauraceae" genus="Endiandra" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endiandra" order="Laurales" pageId="0" pageNumber="183" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="183">Endiandra</emphasis>
</taxonomicName>
species occurring in south China and the adjacent regions in Indochina, this species is mainly characterised by its much larger ellipsoidal fruits (up to 11
<normalizedToken originalValue="×">x</normalizedToken>
6 cm), as well as glabrous branchlets and puberulent inflorescences.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="183" type="materials_examined">
<paragraph pageId="0" pageNumber="183">
<typeStatus>Type</typeStatus>
.
</paragraph>
<paragraph pageId="0" pageNumber="183">
<materialsCitation collectingDate="2022-05-12" country="China" county="Maguan County" elevation="800" location="Houcao Village" municipality="Gulinqing Town" specimenCount="1" stateProvince="Yunnan Province" typeStatus="Holotype">
<collectingCountry name="China">China</collectingCountry>
.
<collectingRegion country="China" name="Yunnan">Yunnan Province</collectingRegion>
:
<collectingCounty>Maguan County</collectingCounty>
,
<collectingMunicipality>Gulinqing Town</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:26B9495DCE415454894380B415741874:21D76692448478D54E6C38618B582D21" country="China" county="Maguan County" municipality="Gulinqing Town" name="Houcao Village" stateProvince="Yunnan Province">Houcao Village</location>
, in tropical montane forest near the village and is strongly disturbed by human activities, ca.
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">800 m</quantity>
a.s.l.
</elevation>
,
<collectingDate value="2022-05-12">12 May 2022</collectingDate>
, flowering,
<emphasis italics="true" pageId="0" pageNumber="183">Lang Li and Dian-yang Zou, 2022028</emphasis>
(
<typeStatus>Holotype</typeStatus>
: HITBC!; Isotypes: HITBC!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="183" type="description">
<paragraph pageId="0" pageNumber="183">Description.</paragraph>
<paragraph pageId="0" pageNumber="183">
Trees evergreen, up to 15 m tall (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Morphology of Endiandra macrocarpa A tree habit B flowering branchlet C fruiting branchlet displaying immature fruit D mature fruits. Photographed by Lang Li and Guan-long Cao." figureDoi="10.3897/phytokeys.224.102752.figure1" httpUri="https://binary.pensoft.net/fig/834766" pageId="0" pageNumber="183">1A</figureCitation>
). Bark brownish-grey. Branchlets brown, terete, with blunt ridges and striate when dry, glabrous, slightly warty. Leaves alternate; petiole 1-1.5 cm, concave-convex, glabrous; leaf blade greenish and opaque abaxially, green and shiny adaxially, elliptic or oblong-elliptic, 5-16
<normalizedToken originalValue="×">x</normalizedToken>
3-7 cm, thinly leathery. Mid-rib elevated on both surfaces, but rather conspicuous abaxially, lateral veins 5-8 pairs, slightly elevated abaxially, conspicuous adaxially, veins and veinlets reticulate, base cuneate to obtuse, mostly asymmetric, apex acuminate with obtuse acumen or obtuse with acute acumen. Panicle axillary, 4-8(10) cm, puberulent (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Morphology of Endiandra macrocarpa A tree habit B flowering branchlet C fruiting branchlet displaying immature fruit D mature fruits. Photographed by Lang Li and Guan-long Cao." figureDoi="10.3897/phytokeys.224.102752.figure1" httpUri="https://binary.pensoft.net/fig/834766" pageId="0" pageNumber="183">1B</figureCitation>
). Pedicels slender, 1-3 mm, thickened after anthesis. Flowers yellow, scented, ca. 3 mm. Perianth fleshy, unequal, outer ones slightly larger, broadly ovate, 3
<normalizedToken originalValue="×">x</normalizedToken>
2 mm (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Morphology of a flower of Endiandra macrocarpa. A outer tepals, abaxial and adaxial side B inner tepals, abaxial and adaxial side C stamen, abaxial side D pistil E flower, top view F flower, longitudinal section. Photographed by Dian-yang Zou." figureDoi="10.3897/phytokeys.224.102752.figure2" httpUri="https://binary.pensoft.net/fig/834767" pageId="0" pageNumber="183">2A</figureCitation>
), adaxially pilose; inner ones smaller, ovate, 2.5
<normalizedToken originalValue="×">x</normalizedToken>
1.8 mm, adaxially densely villous (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Morphology of a flower of Endiandra macrocarpa. A outer tepals, abaxial and adaxial side B inner tepals, abaxial and adaxial side C stamen, abaxial side D pistil E flower, top view F flower, longitudinal section. Photographed by Dian-yang Zou." figureDoi="10.3897/phytokeys.224.102752.figure2" httpUri="https://binary.pensoft.net/fig/834767" pageId="0" pageNumber="183">2B</figureCitation>
). Fertile stamens 3, triangular, ca. 2 mm, eglandular, puberulent; anthers thick, stalkless, 2-celled, cells extrorse, tightly adnate to each other; staminodes absent (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Morphology of a flower of Endiandra macrocarpa. A outer tepals, abaxial and adaxial side B inner tepals, abaxial and adaxial side C stamen, abaxial side D pistil E flower, top view F flower, longitudinal section. Photographed by Dian-yang Zou." figureDoi="10.3897/phytokeys.224.102752.figure2" httpUri="https://binary.pensoft.net/fig/834767" pageId="0" pageNumber="183">2C, E</figureCitation>
). Ovary ovoid, ca. 1.2 mm; style short; stigma punctate (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Morphology of a flower of Endiandra macrocarpa. A outer tepals, abaxial and adaxial side B inner tepals, abaxial and adaxial side C stamen, abaxial side D pistil E flower, top view F flower, longitudinal section. Photographed by Dian-yang Zou." figureDoi="10.3897/phytokeys.224.102752.figure2" httpUri="https://binary.pensoft.net/fig/834767" pageId="0" pageNumber="183">2D</figureCitation>
). Fruit ellipsoid or long ellipsoid, up to 11
<normalizedToken originalValue="×">x</normalizedToken>
6 cm, immature fruit green, yellow when mature, smooth, glabrous, apex bluntly apiculate (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Morphology of Endiandra macrocarpa A tree habit B flowering branchlet C fruiting branchlet displaying immature fruit D mature fruits. Photographed by Lang Li and Guan-long Cao." figureDoi="10.3897/phytokeys.224.102752.figure1" httpUri="https://binary.pensoft.net/fig/834766" pageId="0" pageNumber="183">1C, D</figureCitation>
). Seed endocarp light brown with a darker brown network of both broad and fine, slightly raised veins. Fruit stalk brown, up to 5 mm in diam. at apex, glabrous.
</paragraph>
<caption doi="10.3897/phytokeys.224.102752.figure1" httpUri="https://binary.pensoft.net/fig/834766" pageId="0" pageNumber="183" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="183">
<emphasis bold="true" pageId="0" pageNumber="183">Figure 1.</emphasis>
Morphology of
<taxonomicName authorityName="D. Y. Zou, Lang Li &amp; J. Li" authorityYear="2023" class="Magnoliopsida" family="Lauraceae" genus="Endiandra" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endiandra macrocarpa" order="Laurales" pageId="0" pageNumber="183" phylum="Tracheophyta" rank="species" species="macrocarpa">
<emphasis italics="true" pageId="0" pageNumber="183">Endiandra macrocarpa</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="183">A</emphasis>
tree habit
<emphasis bold="true" pageId="0" pageNumber="183">B</emphasis>
flowering branchlet
<emphasis bold="true" pageId="0" pageNumber="183">C</emphasis>
fruiting branchlet displaying immature fruit
<emphasis bold="true" pageId="0" pageNumber="183">D</emphasis>
mature fruits. Photographed by Lang Li and Guan-long Cao.
</paragraph>
</caption>
<caption doi="10.3897/phytokeys.224.102752.figure2" httpUri="https://binary.pensoft.net/fig/834767" pageId="0" pageNumber="183" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="183">
<emphasis bold="true" pageId="0" pageNumber="183">Figure 2.</emphasis>
Morphology of a flower of
<taxonomicName authorityName="D. Y. Zou, Lang Li &amp; J. Li" authorityYear="2023" class="Magnoliopsida" family="Lauraceae" genus="Endiandra" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endiandra macrocarpa" order="Laurales" pageId="0" pageNumber="183" phylum="Tracheophyta" rank="species" species="macrocarpa">
<emphasis italics="true" pageId="0" pageNumber="183">Endiandra macrocarpa</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="183">A</emphasis>
outer tepals, abaxial and adaxial side
<emphasis bold="true" pageId="0" pageNumber="183">B</emphasis>
inner tepals, abaxial and adaxial side
<emphasis bold="true" pageId="0" pageNumber="183">C</emphasis>
stamen, abaxial side
<emphasis bold="true" pageId="0" pageNumber="183">D</emphasis>
pistil
<emphasis bold="true" pageId="0" pageNumber="183">E</emphasis>
flower, top view
<emphasis bold="true" pageId="0" pageNumber="183">F</emphasis>
flower, longitudinal section. Photographed by Dian-yang Zou.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="183" type="phenology">
<paragraph pageId="0" pageNumber="183">Phenology.</paragraph>
<paragraph pageId="0" pageNumber="183">Flowering from April to May and fruiting from July to October.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="183" type="distribution">
<paragraph pageId="0" pageNumber="183">Distribution and habitat.</paragraph>
<paragraph pageId="0" pageNumber="183">
Currently known only from the type locality in Maguan County, Yunnan Province, south-western China (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distribution of E. macrocarpa (black triangle), E. hainanensis (red dot), E. dolichocarpa (green dot) and E. coriacea (yellow dot) in China." figureDoi="10.3897/phytokeys.224.102752.figure3" httpUri="https://binary.pensoft.net/fig/834768" pageId="0" pageNumber="183">3</figureCitation>
). Tropical montane forest in valley, on clay loam soil mixed limestone; ca. 800 m a.s.l.
</paragraph>
<caption doi="10.3897/phytokeys.224.102752.figure3" httpUri="https://binary.pensoft.net/fig/834768" pageId="0" pageNumber="183" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="183">
<emphasis bold="true" pageId="0" pageNumber="183">Figure 3.</emphasis>
Distribution of
<taxonomicName class="Magnoliopsida" family="Lauraceae" kingdom="Plantae" lsidName="E. macrocarpa" order="Laurales" pageId="0" pageNumber="183" phylum="Tracheophyta" rank="species" species="macrocarpa">
<emphasis italics="true" pageId="0" pageNumber="183">E. macrocarpa</emphasis>
</taxonomicName>
(black triangle),
<taxonomicName class="Magnoliopsida" family="Lauraceae" kingdom="Plantae" lsidName="E. hainanensis" order="Laurales" pageId="0" pageNumber="183" phylum="Tracheophyta" rank="species" species="hainanensis">
<emphasis italics="true" pageId="0" pageNumber="183">E. hainanensis</emphasis>
</taxonomicName>
(red dot),
<taxonomicName class="Magnoliopsida" family="Lauraceae" kingdom="Plantae" lsidName="E. dolichocarpa" order="Laurales" pageId="0" pageNumber="183" phylum="Tracheophyta" rank="species" species="dolichocarpa">
<emphasis italics="true" pageId="0" pageNumber="183">E. dolichocarpa</emphasis>
</taxonomicName>
(green dot) and
<taxonomicName class="Magnoliopsida" family="Lauraceae" kingdom="Plantae" lsidName="E. coriacea" order="Laurales" pageId="0" pageNumber="183" phylum="Tracheophyta" rank="species" species="coriacea">
<emphasis italics="true" pageId="0" pageNumber="183">E. coriacea</emphasis>
</taxonomicName>
(yellow dot) in China.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="183" type="etymology">
<paragraph pageId="0" pageNumber="183">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="183">
The specific epithet &quot;
<taxonomicName authorityName="D. Y. Zou, Lang Li &amp; J. Li" authorityYear="2023" class="Magnoliopsida" family="Lauraceae" genus="Endiandra" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endiandra macrocarpa" order="Laurales" pageId="0" pageNumber="183" phylum="Tracheophyta" rank="species" species="macrocarpa">
<emphasis italics="true" pageId="0" pageNumber="183">Endiandra macrocarpa</emphasis>
</taxonomicName>
&quot; of the new species refers to its very large fruits compared to the other species within the genus
<taxonomicName class="Magnoliopsida" family="Lauraceae" genus="Endiandra" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endiandra" order="Laurales" pageId="0" pageNumber="183" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="183">Endiandra</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="183" type="preliminary conservation status">
<paragraph pageId="0" pageNumber="183">Preliminary conservation status.</paragraph>
<paragraph pageId="0" pageNumber="183">
Currently,
<taxonomicName class="Magnoliopsida" family="Lauraceae" kingdom="Plantae" lsidName="E. macrocarpa" order="Laurales" pageId="0" pageNumber="183" phylum="Tracheophyta" rank="species" species="macrocarpa">
<emphasis italics="true" pageId="0" pageNumber="183">E. macrocarpa</emphasis>
</taxonomicName>
is only known from the type locality in Maguan County, Yunnan Province, south-western China with only one mature individual. It is located in tropical montane forest near the village, which is strongly disturbed by human activities. So far, no other occurrence of this species in south-eastern Yunnan and the adjacent regions has been found or reported. Further in-depth field surveys are suggested in order to find more individuals and locations of the species. Thus, the preliminary conservation status of
<taxonomicName class="Magnoliopsida" family="Lauraceae" kingdom="Plantae" lsidName="E. macrocarpa" order="Laurales" pageId="0" pageNumber="183" phylum="Tracheophyta" rank="species" species="macrocarpa">
<emphasis italics="true" pageId="0" pageNumber="183">E. macrocarpa</emphasis>
</taxonomicName>
is not assessed in the present study.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="183" type="additional specimen examined">
<paragraph pageId="0" pageNumber="183">Additional specimen examined.</paragraph>
<paragraph pageId="0" pageNumber="183">
China. Yunnan Province: Maguan County, Gulinqing Town, Houcao Village, in tropical montane forest near the village, strongly disturbed by human activities, ca. 800 m a.s.l., 26 August 2020, fruiting,
<emphasis italics="true" pageId="0" pageNumber="183">Lang Li and Guan-long Cao, 2020158</emphasis>
(HITBC!); 25 April 2021, flowering,
<emphasis italics="true" pageId="0" pageNumber="183">Lang Li and Guan-long Cao, 2021029</emphasis>
(HITBC!); 28 October 2021, mature fruits,
<emphasis italics="true" pageId="0" pageNumber="183">Lang Li and Dian-yang Zou, 2021081</emphasis>
(HITBC!).
</paragraph>
</subSubSection>
</treatment>
</document>