182 lines
20 KiB
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182 lines
20 KiB
XML
<document id="2E131BEBE91D47C617F649F41C8F4594" ID-CLB-Dataset="306474" ID-DOI="10.11646/zootaxa.5542.1.1" ID-GBIF-Dataset="b349e73d-c49e-418c-81f7-c5d32e136362" ID-ISSN="1175-5334" ID-Zenodo-Dep="14385386" ID-ZooBank="2DB3A5B7-4292-4CD9-B6D8-FA97EB48DD16" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1733917054948" checkinUser="plazi" docAuthor="Moghaddam, Masumeh & Watson, Gillian W." docDate="2024" docId="981B8798FFAC397C92DA905D30ACDE14" docLanguage="en" docName="zootaxa.5542.1.1.pdf" docOrigin="Zootaxa 5542 (1)" docSource="https://doi.org/10.11646/zootaxa.5542.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Coccus Linnaeus" docType="treatment" docVersion="5" lastPageNumber="52" masterDocId="6422FFE0FF9D394F924D97543531DD43" masterDocTitle="The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 3 The Soft Scales (Coccidae) And Other Families" masterLastPageNumber="202" masterPageNumber="1" pageNumber="50" updateTime="1734101643649" updateUser="ExternalLinkService">
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<mods:titleInfo id="7A71CD07E0ECDD110CAFB6ADB0B3EB6F">
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<mods:title id="49810D4C6B95CC3A49581EB4804F48EA">The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 3 The Soft Scales (Coccidae) And Other Families</mods:title>
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<mods:name id="D449EBC183FE5A45A0F033912684178C" type="personal">
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<mods:namePart id="4F0E5456D77A6AF7E3A5400D019D724F">Moghaddam, Masumeh</mods:namePart>
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<mods:affiliation id="F83BE3F45E20AFAE46EE0EA98DD05AA7">Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran.</mods:affiliation>
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<mods:namePart id="A387E4B22672FF08B5ACBAA46165C616">Watson, Gillian W.</mods:namePart>
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<mods:nameIdentifier id="0129CC2C9172A5C1A9D361BFD27492A7" type="ORCID">0000-0001-9914-0094</mods:nameIdentifier>
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<mods:affiliation id="9AAC3C0EF64C57B80C5F4C1D68A9739C">Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. & Science: Research, Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K. gillian. watson @ nhm. ac. uk; https: // orcid. org / 0000 - 0001 - 9914 - 0094 * Corresponding author. moghadam @ iripp. ir; https: // orcid. org / 0000 - 0003 - 0908 - 838 X & Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran.</mods:affiliation>
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<mods:nameIdentifier id="93188ADF063524250475B8A4891AAA28" type="email">gillian.watson@nhm.ac.uk</mods:nameIdentifier>
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<mods:title id="ECC85D22E3D7AE2F74EAC67F53F45969">Zootaxa</mods:title>
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<mods:part id="982A264CB0CED372BAC2F423C18F5EC7">
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<mods:date id="7BA39F3CEBA64D9D899BB4C6C04CA7BF">2024</mods:date>
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<mods:number id="5DEDA08141E0CD6743DE0D11F4DF5C18">2024-11-29</mods:number>
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<mods:number id="5566C1D7E5B8B339E0D0F238FD8613A7">5542</mods:number>
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<treatment id="981B8798FFAC397C92DA905D30ACDE14" LSID="urn:lsid:plazi:treatment:981B8798FFAC397C92DA905D30ACDE14" httpUri="http://treatment.plazi.org/id/981B8798FFAC397C92DA905D30ACDE14" lastPageId="51" lastPageNumber="52" pageId="49" pageNumber="50">
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<subSubSection id="58A86505FFAC397E92DA905D34EEDA60" box="[151,479,1801,1827]" pageId="49" pageNumber="50" type="nomenclature">
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<paragraph id="100D368EFFAC397E92DA905D34EEDA60" blockId="49.[151,479,1801,1827]" box="[151,479,1801,1827]" pageId="49" pageNumber="50">
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<heading id="4B4581E2FFAC397E92DA905D34EEDA60" bold="true" box="[151,479,1801,1827]" fontSize="11" level="2" pageId="49" pageNumber="50" reason="3">
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<emphasis id="22C6EA9CFFAC397E92DA905D34EEDA60" bold="true" box="[151,479,1801,1827]" pageId="49" pageNumber="50">
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Genus
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<taxonomicName id="D7B24D0DFFAC397E92A3905D34EEDA60" ID-CoL="3RN8" authority="Linnaeus" authorityName="Linnaeus" box="[238,479,1801,1827]" class="Insecta" family="Coccidae" genus="Coccus" kingdom="Animalia" order="Hemiptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">
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<emphasis id="22C6EA9CFFAC397E92A3905D3452DA60" bold="true" box="[238,355,1801,1827]" italics="true" pageId="49" pageNumber="50">COCCUS</emphasis>
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Linnaeus
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="58A86505FFAC397C92DA901A3767DC92" lastPageId="51" lastPageNumber="52" pageId="49" pageNumber="50" type="reference_group">
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<paragraph id="100D368EFFAC397E92DA901A36E6DAC0" blockId="49.[151,1437,1869,1924]" pageId="49" pageNumber="50">
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<treatmentCitationGroup id="30A211A0FFAC397E92DA901A36E6DAC0" pageId="49" pageNumber="50">
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<treatmentCitation id="9113109FFFAC397E92DA901A348DDA26" ID-DOI="http://doi.org/10.5281/zenodo.3914541" ID-Zenodo-Dep="3914541" LSID="urn:lsid:zoobank.org:act:D7AFF012-1B2A-4F9B-BA23-795DCC253C10" author="Linnaeus, C." box="[151,444,1869,1893]" httpUri="http://treatment.plazi.org/id/65EAD2D21B241EC2A6551017B542B7D6" page="455" pageId="49" pageNumber="50" year="1758">
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<taxonomicName id="D7B24D0DFFAC397E92DA901A348DDA26" ID-CoL="3RN8" authority="Linnaeus, 1758: 455" authorityName="Linnaeus" authorityPageNumber="455" authorityYear="1758" box="[151,444,1869,1893]" class="Insecta" family="Coccidae" genus="Coccus" kingdom="Animalia" order="Hemiptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">
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<emphasis id="22C6EA9CFFAC397E92DA901A35D3DA26" box="[151,226,1870,1893]" italics="true" pageId="49" pageNumber="50">Coccus</emphasis>
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<bibRefCitation id="74234B7FFFAC397E92A59019348DDA26" author="Linnaeus, C." box="[232,444,1869,1893]" pageId="49" pageNumber="50" refId="ref104704" refString="Linnaeus, C. (1758) Systema Naturae, per regna tria naturae, secundum classes, ordines, genera, species cum characteribus, differentiis, synonymis, locis. Laurentii Salvii, Holmiae, (4) + 824 pp. https: // doi. org / 10.5962 / bhl. title. 542" type="book" year="1758">Linnaeus, 1758: 455</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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<typeStatus id="CF09882CFFAC397E9388901A34C6DA25" box="[453,503,1870,1894]" pageId="49" pageNumber="50">Type</typeStatus>
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species:
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<taxonomicName id="D7B24D0DFFAC397E9018901A36B1DA26" authority="Linnaeus" authorityName="Linnaeus" box="[597,896,1869,1893]" class="Insecta" family="Coccidae" genus="Coccus" kingdom="Animalia" order="Hemiptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="hesperidum">
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<emphasis id="22C6EA9CFFAC397E9018901A362CDA26" box="[597,797,1869,1893]" italics="true" pageId="49" pageNumber="50">Coccus hesperidum</emphasis>
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Linnaeus
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</taxonomicName>
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by subsequent designation.
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<treatmentCitation id="9113109FFFAC397E96ED901935E7DAC7" author="Costa, O. G." page="6" pageId="49" pageNumber="50" year="1828">
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<taxonomicName id="D7B24D0DFFAC397E96ED901935E7DAC7" authority="Costa, 1828: 6" authorityName="Costa" authorityPageNumber="6" authorityYear="1828" class="Insecta" family="Coccidae" genus="Calymmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">
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<emphasis id="22C6EA9CFFAC397E96ED90193024DA26" box="[1184,1301,1869,1893]" italics="true" pageId="49" pageNumber="50">Calymmata</emphasis>
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<bibRefCitation id="74234B7FFFAC397E9751901935E7DAC7" author="Costa, O. G." pageId="49" pageNumber="50" refId="ref100048" refString="Costa, O. G. (1828) Prospetto di una nuova divisione metodica del genere Coccus, Lin. Lam. Ec. Dalla Tipografia Trani., Napoli, 8 pp." type="book" year="1828">Costa, 1828: 6</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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<treatmentCitation id="9113109FFFAC397E92AC903834D8DAC7" author="Costa, O. G." box="[225,489,1900,1924]" page="8" pageId="49" pageNumber="50" year="1829">
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<taxonomicName id="D7B24D0DFFAC397E92AC903834D8DAC7" authority="Costa, 1829: 8" authorityName="Costa" authorityPageNumber="8" authorityYear="1829" box="[225,489,1900,1924]" class="Insecta" family="Coccidae" genus="Calypticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">
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<emphasis id="22C6EA9CFFAC397E92AC9038347CDAC7" box="[225,333,1900,1924]" italics="true" pageId="49" pageNumber="50">Calypticus</emphasis>
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<bibRefCitation id="74234B7FFFAC397E931E903834D8DAC7" author="Costa, O. G." box="[339,489,1900,1924]" pageId="49" pageNumber="50" refId="ref100083" refString="Costa, O. G. (1829) Fauna del Regno di Napoli, famiglia de coccinigliferi, o de' gallinsetti. Emitteri, Napoli, 23 pp." type="book" year="1829">Costa, 1829: 8</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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<treatmentCitation id="9113109FFFAC397E93BE903936E2DAC0" author="Tao, C. C. C. & Wong, C. Y. & Chang, Y. C." box="[499,979,1900,1924]" page="77" pageId="49" pageNumber="50" year="1983">
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<taxonomicName id="D7B24D0DFFAC397E93BE903936E2DAC0" authority="Tao, Wong & Chang, 1983: 77" authorityName="Tao, Wong & Chang" authorityPageNumber="77" authorityYear="1983" box="[499,979,1900,1924]" class="Insecta" family="Coccidae" genus="Taiwansaissetia" kingdom="Animalia" order="Hemiptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">
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<emphasis id="22C6EA9CFFAC397E93BE903937A5DAC7" box="[499,660,1901,1924]" italics="true" pageId="49" pageNumber="50">Taiwansaissetia</emphasis>
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<bibRefCitation id="74234B7FFFAC397E90D7903836E2DAC0" author="Tao, C. C. C. & Wong, C. Y. & Chang, Y. C." box="[666,979,1900,1924]" pageId="49" pageNumber="50" pagination="57 - 107" refId="ref109521" refString="Tao, C. C. C., Wong, C. Y. & Chang, Y. C. (1983) Monograph of Coccidae of Taiwan, Republic of China (Homoptera: Coccoidea). Journal of the Taiwan Museum, 36, 57 - 107." type="journal article" year="1983">Tao, Wong & Chang, 1983: 77</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="100D368EFFAC397C92DA90E63767DC92" blockId="49.[151,1436,1970,2033]" lastBlockId="51.[151,1437,151,537]" lastPageId="51" lastPageNumber="52" pageId="49" pageNumber="50">
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<emphasis id="22C6EA9CFFAC397E92DA90E637B3DA8F" bold="true" box="[151,642,1970,1996]" pageId="49" pageNumber="50">Diagnostic characters of the adult female:</emphasis>
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Live adult female oval / elongate / pyriform, usually flat to only slightly convex. Slide-mounted, body oval; dorsal surface membranous, gradually increasing in thickness at maturity, with small areolations present. Dorsum with short setae setose / spinose / clavate / cylindrical. Ventral surface with short setae. Pregenital disc-pores mainly each with 10 loculi, present around anogenital fold or extending anteriorly, even as far forward as head. Spiracular disc-pores each with 5 loculi. Preopercular pores present or absent. Dorsal tubular ducts and dorsal submarginal tubercles sometimes present (usually on submargin). Ventral tubular ducts present or absent; when present, located medially or occasionally forming a sparse to moderately dense submarginal band. Marginal setae simple or branched distally; stigmatic clefts deep or shallow, each containing 3 stigmatic setae. Antennae usually with 7 or 8 segments. Legs usually well developed, with or without a tibio-tarsal articulatory sclerosis. Anal cleft of moderate depth. Anal plates together quadrate, sometimes with subdiscal setae but never with discal setae. Anal ring with 6 or 8 setae.
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</paragraph>
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</subSubSection>
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<caption id="44CD6606FFAF397D92DA90093615DA36" ID-DOI="http://doi.org/10.5281/zenodo.14385458" ID-Zenodo-Dep="14385458" box="[151,804,1885,1909]" httpUri="https://zenodo.org/record/14385458/files/figure.png" pageId="50" pageNumber="51" startId="50.[151,250,1885,1909]" targetBox="[161,1431,191,1861]" targetPageId="50" targetType="figure">
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<paragraph id="100D368EFFAF397D92DA90093615DA36" blockId="50.[151,804,1885,1909]" box="[151,804,1885,1909]" pageId="50" pageNumber="51">
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<emphasis id="22C6EA9CFFAF397D92DA90093411DA36" bold="true" box="[151,288,1885,1909]" pageId="50" pageNumber="51">FIGURE 17.</emphasis>
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Adult female of
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<taxonomicName id="D7B24D0DFFAF397D938690093615DA36" authority="Del Guercio." authorityName="Del Guercio." box="[459,804,1885,1909]" class="Insecta" family="Coccidae" genus="Ceroplastes" kingdom="Animalia" order="Hemiptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="sinensis">
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<emphasis id="22C6EA9CFFAF397D9386900937ABDA36" box="[459,666,1885,1909]" italics="true" pageId="50" pageNumber="51">Ceroplastes sinensis</emphasis>
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Del Guercio.
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</taxonomicName>
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</paragraph>
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</caption>
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<subSubSection id="58A86505FFAE397C928A968E3772DF5A" pageId="51" pageNumber="52" type="discussion">
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<paragraph id="100D368EFFAE397C928A968E3772DF5A" blockId="51.[151,1437,151,537]" pageId="51" pageNumber="52">
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<emphasis id="22C6EA9CFFAE397C928A968E3463DCB7" bold="true" box="[199,338,474,500]" pageId="51" pageNumber="52">Comments:</emphasis>
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Worldwide, 114 species are included in
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<taxonomicName id="D7B24D0DFFAE397C9162968F31EADCB6" authority="(Garcia Morales et al. 2016)" baseAuthorityName="Garcia Morales" baseAuthorityYear="2016" box="[815,1243,475,501]" class="Insecta" family="Coccidae" genus="Coccus" kingdom="Animalia" order="Hemiptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
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<emphasis id="22C6EA9CFFAE397C9162968F36B0DCB6" box="[815,897,475,501]" italics="true" pageId="51" pageNumber="52">Coccus</emphasis>
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(
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<bibRefCitation id="74234B7FFFAE397C91D9968F31E5DCB6" author="Garcia Morales M. & Denno B. D. & Miller D. R. & Miller G. L. & Ben-Dov Y. & Hardy N. B." box="[916,1236,475,501]" pageId="51" pageNumber="52" refId="ref101399" refString="Garcia Morales M., Denno B. D., Miller D. R., Miller G. L., Ben-Dov Y. & Hardy N. B. (2016) ScaleNet: a literature-based model of scale insect biology and systematics. Database. Available from: http: // scalenet. info (accessed 31 January 2024) https: // doi. org / 10.1093 / database / bav 118" type="url" year="2016">
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García Morales
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<emphasis id="22C6EA9CFFAE397C961C968F31B4DCB6" box="[1105,1157,475,501]" italics="true" pageId="51" pageNumber="52">et al</emphasis>
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. 2016
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</bibRefCitation>
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)
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</taxonomicName>
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; two species are known from
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<collectingCountry id="68A5761EFFAE397C936596AB3464DF5A" box="[296,341,511,537]" name="Iran" pageId="51" pageNumber="52">Iran</collectingCountry>
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(
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<bibRefCitation id="74234B7FFFAE397C932E96AB3706DF5A" author="Moghaddam, M." box="[355,567,511,537]" pageId="51" pageNumber="52" pagination="1 - 92" refId="ref105801" refString="Moghaddam, M. (2013) An annotated checklist of the scale insects of Iran (Hemiptera, Sternorrhyncha, Coccoidea) with new records and distribution data. ZooKeys, 334, 1 - 92. https: // doi. org / 10.3897 / zookeys. 334.5818" type="journal article" year="2013">Moghaddam 2013</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="58A86505FFAE397C92DA953F30ACDE14" pageId="51" pageNumber="52" type="key">
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<paragraph id="100D368EFFAE397C92DA953F36C6DFC6" blockId="51.[151,1015,618,645]" box="[151,1015,618,645]" pageId="51" pageNumber="52">
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<emphasis id="22C6EA9CFFAE397C92DA953F371DDFC6" bold="true" box="[151,556,618,645]" pageId="51" pageNumber="52">
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Key to species of
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<taxonomicName id="D7B24D0DFFAE397C9323953F34F7DFC6" ID-CoL="3RN8" box="[366,454,619,645]" class="Insecta" family="Coccidae" genus="Coccus" kingdom="Animalia" order="Hemiptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="22C6EA9CFFAE397C9323953F34F7DFC6" bold="true" box="[366,454,619,645]" italics="true" pageId="51" pageNumber="52">Coccus</emphasis>
|
|
</taxonomicName>
|
|
in
|
|
<collectingCountry id="68A5761EFFAE397C93A0953F3717DFC6" box="[493,550,619,645]" name="Iran" pageId="51" pageNumber="52">Iran</collectingCountry>
|
|
,
|
|
</emphasis>
|
|
based on slide-mounted adult females.
|
|
</paragraph>
|
|
<keyStep id="AB462E2BFFAE397C92DA95E430ACDE14" pageId="51" pageNumber="52">
|
|
<key id="642361BFFFAE397C92DA95E430ACDE14" pageId="51" pageNumber="52">
|
|
<paragraph id="100D368EFFAE397C92DA95E430ADDE43" blockId="51.[151,1437,688,856]" pageId="51" pageNumber="52">
|
|
<keyLead id="AB4395BBFFAE397C92DA95E430ADDE43" pageId="51" pageNumber="52">
|
|
1(0) Submarginal duct tubercles present. Antennae each with 7 segments. Ventral tubular ducts present medially near attachments of legs (especially on mesothorax), absent from abdominal submargin. Each claw without a denticle. Dorsal setae bluntly pointed............................................................................
|
|
<taxonomicName id="D7B24D0DFFAE397C968795BE30ADDE43" authority="Linnaeus" authorityName="Linnaeus" box="[1226,1436,746,768]" class="Insecta" family="Coccidae" genus="Coccus" kingdom="Animalia" order="Hemiptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="hesperidum">
|
|
<emphasis id="22C6EA9CFFAE397C968795BE300FDE43" bold="true" box="[1226,1342,746,768]" italics="true" pageId="51" pageNumber="52">hesperidum</emphasis>
|
|
Linnaeus
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="100D368EFFAE397C92DA945C30ACDE14" blockId="51.[151,1437,688,856]" pageId="51" pageNumber="52">
|
|
<keyLead id="AB4395BBFFAE397C92DA945C30ACDE14" pageId="51" pageNumber="52">
|
|
- Submarginal duct tubercles absent. Antennae each with 8 segments. Ventral tubular ducts absent medially near attachments of mesothoracic legs but present on abdominal submargin. Each claw with a denticle. Dorsal setae sharply pointed.................................................................................... ..
|
|
<taxonomicName id="D7B24D0DFFAE397C9627941630ACDE14" ID-CoL="WMP5" authority="(Kuwana)" baseAuthorityName="Kuwana" baseAuthorityYear="1914" box="[1130,1437,834,856]" class="Insecta" family="Coccidae" genus="Coccus" kingdom="Animalia" order="Hemiptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="pseudomagnoliarum">
|
|
<emphasis id="22C6EA9CFFAE397C962794163007DE1B" bold="true" box="[1130,1334,834,856]" italics="true" pageId="51" pageNumber="52">pseudomagnoliarum</emphasis>
|
|
(Kuwana)
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
</key>
|
|
</keyStep>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |