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<document id="773581979CA5707DA98890B309CFFC22" ID-CLB-Dataset="55793" ID-DOI="10.1080/00222933.2013.791936" ID-GBIF-Dataset="6591e69b-631a-4c4d-bf83-db8c70d1ad05" ID-ISSN="1464-5262" ID-Zenodo-Dep="5198102" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" checkinTime="1628884845165" checkinUser="felipe" docAuthor="Zhou, Yu-Lingzi &amp; Zhou, Hong-Zhang" docDate="2013" docId="A10D4838C345374725FAFDE719A3F91F" docLanguage="en" docName="JNatHist.47.45-46.2869-2904.pdf" docOrigin="Journal of Natural History 47 (45 - 46)" docSource="http://dx.doi.org/10.1080/00222933.2013.791936" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.6:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="6" docTitle="Liotesba subsimilis Cameron 1945" docType="treatment" docVersion="12" lastPageNumber="2898" masterDocId="5D343040C35B37582413FFC11B0DFF84" masterDocTitle="Taxonomy and phylogeny of the genus Liotesba Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae: Xantholinini) with descriptions of four new species" masterLastPageNumber="2904" masterPageNumber="2869" pageNumber="2897" updateTime="1708086164713" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="872222AD3E5CFB51A84BB50B1765E06D">Taxonomy and phylogeny of the genus Liotesba Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae: Xantholinini) with descriptions of four new species</mods:title>
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<mods:name id="749F1921B9FCAE878131881E7C1AC74A" type="personal">
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<mods:roleTerm id="4783BEBBF534A9148BC875C156D15602">Author</mods:roleTerm>
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<mods:namePart id="EE20C9EECCB55CCE1A3E5646AF0BFC1D">Zhou, Yu-Lingzi</mods:namePart>
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</mods:role>
<mods:namePart id="AC8F9F6378351A717E858BA56C15A9B1">Zhou, Hong-Zhang</mods:namePart>
</mods:name>
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<mods:title id="092AEB7059DD415E6A51B58301FB729B">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="FA5CF54501F310F0DA800F0D160033CF">
<mods:date id="DF5C6F4C26BC68E03B23EA1D373B854F">2013</mods:date>
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<mods:number id="C1075D9B1EF801CA95D8AECFF25B547F">2013-08-08</mods:number>
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<mods:number id="BBE1B47AEB6D6775396E87DA5ECC432F">47</mods:number>
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<mods:detail id="7E7E2C6D26673F59928315E83DC81762" type="issue">
<mods:number id="7B266DB723F92D34B6FAD6D533FDA90A">45 - 46</mods:number>
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<mods:start id="F1E5BDEB11AB3B532C076A750BD77503">2869</mods:start>
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<mods:url id="152B475EC30058A115433E829499E065">http://dx.doi.org/10.1080/00222933.2013.791936</mods:url>
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<mods:classification id="8631088258C0B481F898038AD2B48032">journal article</mods:classification>
<mods:identifier id="9570B2B0A3783CB608BFBBD6327659DC" type="CLB-Dataset">55793</mods:identifier>
<mods:identifier id="B04102899B9C29C30D423BB8D8F67F20" type="DOI">10.1080/00222933.2013.791936</mods:identifier>
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<mods:identifier id="56C3EB3957B669C3F99A0E6EB9C791AF" type="ISSN">1464-5262</mods:identifier>
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</mods:mods>
<treatment id="A10D4838C345374725FAFDE719A3F91F" ID-GBIF-Taxon="194873660" LSID="urn:lsid:plazi:treatment:A10D4838C345374725FAFDE719A3F91F" httpUri="http://treatment.plazi.org/id/A10D4838C345374725FAFDE719A3F91F" lastPageId="31" lastPageNumber="2898" pageId="30" pageNumber="2897">
<subSubSection id="61BEAAA5C345374625FAFDE71873FDBA" box="[489,894,550,574]" pageId="30" pageNumber="2897" type="nomenclature">
<paragraph id="291BF92EC345374625FAFDE71873FDBA" blockId="30.[489,894,550,574]" box="[489,894,550,574]" pageId="30" pageNumber="2897">
<heading id="72534E42C345374625FAFDE71873FDBA" box="[489,894,550,574]" centered="true" fontSize="10" level="2" pageId="30" pageNumber="2897" reason="2">
<taxonomicName id="EEA482ADC345374625FAFDE71873FDBA" ID-CoL="3V87W" authority="(Cameron, 1945)" authorityName="Cameron" authorityYear="1945" box="[489,894,550,574]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="2897" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="1BD0253CC345374625FAFDE719B4FDBA" bold="true" box="[489,697,550,574]" italics="true" pageId="30" pageNumber="2897">Liotesba subsimilis</emphasis>
(
<bibRefCitation id="4D3584DFC345374626DBFDE71878FDBA" author="Cameron M" box="[712,885,550,574]" pageId="30" pageNumber="2897" pagination="63 - 69" refId="ref17526" refString="Cameron M. 1945. Descriptions of new Staphylinidae (Coleoptera). Proc R Entomol Soc Lond. 14: 63 - 69." type="journal article" year="1945">Cameron, 1945</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61BEAAA5C345374624C0FDA71A55FC99" pageId="30" pageNumber="2897" type="reference_group">
<paragraph id="291BF92EC345374624C0FDA71A55FC99" blockId="30.[211,1172,614,894]" pageId="30" pageNumber="2897">
<treatmentCitationGroup id="09B4DE00C345374624C0FDA71A55FC99" pageId="30" pageNumber="2897">
<treatmentCitation id="A805DF3FC345374624C0FDA71AB9FDFA" author="Cameron" box="[211,436,614,638]" page="68" pageId="30" pageNumber="2897" year="1945">
<bibRefCitation id="4D3584DFC345374624C0FDA71AB9FDFA" author="Cameron M" box="[211,436,614,638]" pageId="30" pageNumber="2897" pagination="63 - 69" refId="ref17526" refString="Cameron M. 1945. Descriptions of new Staphylinidae (Coleoptera). Proc R Entomol Soc Lond. 14: 63 - 69." type="journal article" year="1945">Cameron, 1945: 68</bibRefCitation>
</treatmentCitation>
(orig.
<taxonomicName id="EEA482ADC34537462618FDA71807FDFA" ID-CoL="6JYSV" authorityName="Cameron" authorityYear="1945" box="[523,778,614,638]" class="Insecta" family="Staphylinidae" genus="Gauropterus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="2897" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="1BD0253CC34537462618FDA71807FDFA" box="[523,778,614,638]" italics="true" pageId="30" pageNumber="2897">Gauropterus subsimilis</emphasis>
</taxonomicName>
;
<typeStatus id="F61F478CC3453746270CFDA71842FDFA" box="[799,847,614,638]" pageId="30" pageNumber="2897">type</typeStatus>
locality: United Provinces: Chakrata district, Mundali);
<treatmentCitation id="A805DF3FC34537462630FD44180DFD19" author="Bordoni A" box="[547,768,645,669]" page="317" pageId="30" pageNumber="2897" year="2002">
<bibRefCitation id="4D3584DFC34537462630FD44180DFD19" author="Bordoni A" box="[547,768,645,669]" pageId="30" pageNumber="2897" pagination="1 - 998" refId="ref17034" refString="Bordoni A. 2002. Xantholinini of the Oriental Region (Coleoptera: Staphylinidae). Classification, phylogeny and taxonomic revision. Museo Regionale di Scienze Naturali Monografie (Turin). 33: 1 - 998." type="journal article" year="2002">Bordoni, 2002: 317</bibRefCitation>
</treatmentCitation>
(
<taxonomicName id="EEA482ADC34537462701FD47187CFD1A" ID-CoL="8VYYF" box="[786,881,646,670]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="2897" phylum="Arthropoda" rank="genus">
<emphasis id="1BD0253CC34537462701FD47187CFD1A" box="[786,881,646,670]" italics="true" pageId="30" pageNumber="2897">Liotesba</emphasis>
</taxonomicName>
; characters;
<collectingCountry id="51B3B9BEC34537462014FD441F4BFD19" box="[1031,1094,645,669]" name="India" pageId="30" pageNumber="2897">India</collectingCountry>
,
<collectingRegion id="EB6037CCC34537462047FD471A3CFD39" country="India" name="Uttar Pradesh" pageId="30" pageNumber="2897">Uttar Pradesh</collectingRegion>
, W-Almora Div., Sunderdhunga, Wantisan, Mundali Chakrata;
<collectingCountry id="51B3B9BEC34537462054FD641F82FD39" box="[1095,1167,677,701]" name="Nepal" pageId="30" pageNumber="2897">Nepal</collectingCountry>
,
<collectingRegion id="EB6037CCC345374624C0FD041A6DFD59" box="[211,352,709,733]" country="Nepal" name="Dhawalagiri" pageId="30" pageNumber="2897">Dhawalagiri</collectingRegion>
, Myagdi Distr., Ropchani, north Sanchur, Kathmandu Distr., Kakani, Kathmandu Distr., Godavari);
<treatmentCitation id="A805DF3FC34537462628FD241810FD79" author="Smetana A" box="[571,797,741,765]" page="691" pageId="30" pageNumber="2897" year="2004">
<bibRefCitation id="4D3584DFC34537462628FD241810FD79" author="Smetana A" box="[571,797,741,765]" pageId="30" pageNumber="2897" pagination="687 - 698" refId="ref18057" refString="Smetana A. 2004. Staphylinidae, tribe Xantholinini. In: Lobl I, Smetana A, editors. Catalogue of Palaearctic Coleoptera, Vol. 2. Stenstrup: Apollo Books; p. 687 - 698." type="book chapter" year="2004">Smetana, 2004: 691</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="A805DF3FC345374627A6FD241F99FD79" author="Bordoni A" box="[949,1172,741,765]" page="71" pageId="30" pageNumber="2897" year="2007">
<bibRefCitation id="4D3584DFC345374627A6FD241F99FD79" author="Bordoni A" box="[949,1172,741,765]" pageId="30" pageNumber="2897" pagination="67 - 78" refId="ref17403" refString="Bordoni A. 2007 b. Xantholinini (Coleoptera staphylinidae). Supplement to the catalogue Palaearctic coleoptera, volume 2. Bollettino della Societa Entomologica Italiana. 139: 67 - 78." type="journal article" year="2007">Bordoni, 2007b: 71</bibRefCitation>
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="61BEAAA5C345374624EDFCE418AFFCF9" pageId="30" pageNumber="2897" type="synonymic_list">
<paragraph id="291BF92EC345374624EDFCE418AFFCF9" blockId="30.[211,1172,614,894]" pageId="30" pageNumber="2897">
<emphasis id="1BD0253CC345374624EDFCE41A7BFCB9" bold="true" box="[254,374,805,829]" pageId="30" pageNumber="2897">Synonyms.</emphasis>
<taxonomicName id="EEA482ADC34537462594FCE41873FCB9" authority="Coiffait, 1982: 248" authorityName="Coiffait" authorityPageNumber="248" authorityYear="1982" box="[391,894,805,829]" class="Insecta" family="Staphylinidae" genus="Gauropterus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="2897" phylum="Arthropoda" rank="species" species="nepalicus">
<emphasis id="1BD0253CC34537462594FCE4198BFCB9" box="[391,646,805,829]" italics="true" pageId="30" pageNumber="2897">Gauropterus nepalicus</emphasis>
<bibRefCitation id="4D3584DFC3453746268BFCE41873FCB9" author="Coiffait H" box="[664,894,805,829]" pageId="30" pageNumber="2897" pagination="231 - 302" refId="ref17625" refString="Coiffait H. 1982. Staphylinides (Col.) de la region himalayenne et de l'Inde (I. Xantholininae, Staphylininae et Paederinae). Entomol Basiliensia. 7: 231 - 302." type="journal article" year="1982">Coiffait, 1982: 248</bibRefCitation>
</taxonomicName>
(
<typeStatus id="F61F478CC3453746278BFCE418C7FCB9" box="[920,970,805,829]" pageId="30" pageNumber="2897">type</typeStatus>
locality:
<collectingCountry id="51B3B9BEC34537462054FCE41F82FCB9" box="[1095,1167,805,829]" name="Nepal" pageId="30" pageNumber="2897">Nepal</collectingCountry>
: Godavari,
<quantity id="EE5C54CBC34537462547FC841AFEFCD9" box="[340,499,837,861]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="30" pageNumber="2897" unit="m" value="1600.0" valueMax="1700.0" valueMin="1500.0">1500–1700 m</quantity>
);
<bibRefCitation id="4D3584DFC3453746261CFC8419FAFCD9" author="Bordoni A" box="[527,759,837,861]" pageId="30" pageNumber="2897" pagination="1 - 998" refId="ref17034" refString="Bordoni A. 2002. Xantholinini of the Oriental Region (Coleoptera: Staphylinidae). Classification, phylogeny and taxonomic revision. Museo Regionale di Scienze Naturali Monografie (Turin). 33: 1 - 998." type="journal article" year="2002">Bordoni, 2002: 317</bibRefCitation>
(synonym of
<taxonomicName id="EEA482ADC345374627B9FC841F8EFCD9" baseAuthorityName="Cameron" baseAuthorityYear="1945" box="[938,1155,837,861]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="2897" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="1BD0253CC345374627B9FC841F8EFCD9" box="[938,1155,837,861]" italics="true" pageId="30" pageNumber="2897">Liotesba subsimilis</emphasis>
</taxonomicName>
);
<bibRefCitation id="4D3584DFC345374624C0FCA41AA3FCF9" author="Smetana A" box="[211,430,869,893]" pageId="30" pageNumber="2897" pagination="687 - 698" refId="ref18057" refString="Smetana A. 2004. Staphylinidae, tribe Xantholinini. In: Lobl I, Smetana A, editors. Catalogue of Palaearctic Coleoptera, Vol. 2. Stenstrup: Apollo Books; p. 687 - 698." type="book chapter" year="2004">Smetana, 2004: 691</bibRefCitation>
(catalogue; synonym of
<taxonomicName id="EEA482ADC345374626D2FCA4189CFCF9" baseAuthorityName="Cameron" baseAuthorityYear="1945" box="[705,913,869,893]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="2897" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="1BD0253CC345374626D2FCA4189CFCF9" box="[705,913,869,893]" italics="true" pageId="30" pageNumber="2897">Liotesba subsimilis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="61BEAAA5C345374624C0FC6518B9FC78" pageId="30" pageNumber="2897" type="materials_examined">
<paragraph id="291BF92EC345374624C0FC6518B9FC78" blockId="30.[211,1171,932,1021]" pageId="30" pageNumber="2897">
<emphasis id="1BD0253CC345374624C0FC651ABEFC38" bold="true" box="[211,435,932,956]" pageId="30" pageNumber="2897">Material examined.</emphasis>
<materialsCitation id="99CCF373C345374625D1FC6518BDFC78" ID-GBIF-Occurrence="3757895311" collectingDate="1933-06-19" collectionCode="BMNH" collectorName="M. Cameron. Bequest." determinerName="Bordoni" pageId="30" pageNumber="2897" specimenCount="1">
female, with labels as follows: “
<typeStatus id="F61F478CC34537462754FC651889FC38" box="[839,900,932,956]" pageId="30" pageNumber="2897">Type</typeStatus>
”/“on wing”/“Mundali Chakrata, U.P., B.M. Bhatis,
<date id="5D1ADFEEC3453746260EFC051996FC58" box="[541,667,964,988]" pageId="30" pageNumber="2897" value="1933-06-19">
<collectingDate id="4D5E2606C3453746260EFC051996FC58" box="[541,667,964,988]" pageId="30" pageNumber="2897" value="1933-06-19">19.VI.1933</collectingDate>
</date>
”/“762”/“
<collectorName id="84519CF8C3453746271EFC051F02FC58" box="[781,1039,964,988]" pageId="30" pageNumber="2897">M. Cameron. Bequest.</collectorName>
B.M. 1955- 147”/“
<taxonomicName id="EEA482ADC34537462531FC241941FC78" authority="(Cam.)" authorityName="Tempora" baseAuthorityName="Cameron" baseAuthorityYear="1945" box="[290,588,996,1021]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="2897" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="1BD0253CC34537462531FC241AFFFC79" box="[290,498,997,1021]" italics="true" pageId="30" pageNumber="2897">Liotesba subsimilis</emphasis>
(Cam.)
</taxonomicName>
,
<determinerName id="DCC715EAC34537462645FC2519BEFC78" box="[598,691,996,1020]" pageId="30" pageNumber="2897">Bordoni</determinerName>
det., 1998”. (
<collectionCode id="4FB561EBC34537462758FC2518A5FC78" box="[843,936,996,1020]" pageId="30" pageNumber="2897">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="61BEAAA5C345374724C0FBF31877FA5F" lastPageId="31" lastPageNumber="2898" pageId="30" pageNumber="2897" type="description">
<paragraph id="291BF92EC345374624C0FBF31A5FFBCD" blockId="30.[211,1171,1074,1264]" box="[211,338,1074,1097]" pageId="30" pageNumber="2897">
<heading id="72534E42C345374624C0FBF31A5FFBCD" box="[211,338,1074,1097]" fontSize="10" level="3" pageId="30" pageNumber="2897" reason="5">
<emphasis id="1BD0253CC345374624C0FBF31A5FFBCD" box="[211,338,1074,1097]" italics="true" pageId="30" pageNumber="2897">Description</emphasis>
</heading>
</paragraph>
<paragraph id="291BF92EC345374624C0FB9818A3FB14" blockId="30.[211,1171,1074,1264]" pageId="30" pageNumber="2897">
<emphasis id="1BD0253CC345374624C0FB981A7DFBF5" bold="true" box="[211,368,1113,1137]" pageId="30" pageNumber="2897">Measurement.</emphasis>
BL =
<quantity id="EE5C54CBC345374625DAFB98193CFBF4" box="[457,561,1113,1137]" metricMagnitude="-3" metricUnit="m" metricValue="6.23" pageId="30" pageNumber="2897" unit="mm" value="6.23">6.23 mm</quantity>
, FL =
<quantity id="EE5C54CBC3453746269EFB9819FBFBF4" box="[653,758,1113,1137]" metricMagnitude="-3" metricUnit="m" metricValue="3.53" pageId="30" pageNumber="2897" unit="mm" value="3.53">3.53 mm</quantity>
, HL =
<quantity id="EE5C54CBC34537462745FB9818B3FBF4" box="[854,958,1113,1137]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="30" pageNumber="2897" unit="mm" value="1.03">1.03 mm</quantity>
, HW =
<quantity id="EE5C54CBC34537462034FB981F82FBF4" box="[1063,1167,1113,1137]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="30" pageNumber="2897" unit="mm" value="0.88">0.88 mm</quantity>
, PL =
<quantity id="EE5C54CBC34537462504FBB81A71FB14" box="[279,380,1145,1169]" metricMagnitude="-3" metricUnit="m" metricValue="1.12" pageId="30" pageNumber="2897" unit="mm" value="1.12">1.12 mm</quantity>
, PW =
<quantity id="EE5C54CBC345374625C0FBB8193AFB14" box="[467,567,1145,1169]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" pageId="30" pageNumber="2897" unit="mm" value="0.81">0.81 mm</quantity>
, EL =
<quantity id="EE5C54CBC34537462694FBB819E1FB14" box="[647,748,1145,1169]" metricMagnitude="-3" metricUnit="m" metricValue="1.26" pageId="30" pageNumber="2897" unit="mm" value="1.26">1.26 mm</quantity>
, EW =
<quantity id="EE5C54CBC34537462756FBB818A4FB14" box="[837,937,1145,1169]" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" pageId="30" pageNumber="2897" unit="mm" value="0.94">0.94 mm</quantity>
.
</paragraph>
<paragraph id="291BF92EC345374624EDFB591A2EFB74" blockId="30.[211,1171,1074,1264]" pageId="30" pageNumber="2897">
Body nearly cylindrical, small-sized. Body dark brown, except elytra
<emphasis id="1BD0253CC345374627EDFB591F58FB34" box="[1022,1109,1176,1200]" pageId="30" pageNumber="2897">reddish</emphasis>
. Legs and antennae reddish brown, except scape dark brown. Maxillary and labial palpi brown.
</paragraph>
<paragraph id="291BF92EC345374724C0FAD91A07FF5B" blockId="30.[211,1172,1304,1648]" lastBlockId="31.[143,1103,167,223]" lastPageId="31" lastPageNumber="2898" pageId="30" pageNumber="2897">
<emphasis id="1BD0253CC345374624C0FAD91A1AFAB4" bold="true" box="[211,279,1304,1328]" pageId="30" pageNumber="2897">Head.</emphasis>
Rectangular (HL to HW ratio 1.2), tempora straight, posterior angles projecting outwards. Dorsal integument shiny, rather smooth, extensively bearing micropunctures, with five or six sparsely scattered large punctures on posterior half of head, also with a row of large punctures on deflexed portion of tempora. Each side of cranium with anterolateral puncture near antennal insertion, midlateral puncture about a distance of three punctures’ diameter from dorsal margin of eye, temporal puncture at the posterior one-fifth, and occipital puncture at the lateral third. Frontal furrows relatively short, anteocular furrows absent. Eye small, shorter than one-third the length of tempora (eye: tempora = 0.19:
<quantity id="EE5C54CBC3453746267FF9D619D9F9AB" box="[620,724,1559,1583]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="30" pageNumber="2897" unit="mm" value="0.59">0.59 mm</quantity>
), not protruding laterad; one large and one small puncture next to dorsal margin of eye; five or six larger punctures confluent to a deep furrow behind eye. Epistoma narrow, narrowed forwards. Distance between antennal insertions
<quantity id="EE5C54CBC344374725DDFF66193EFF3B" box="[462,563,167,191]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="31" pageNumber="2898" unit="mm" value="0.27">0.27 mm</quantity>
, being subequal to distance from antenna to eye (
<quantity id="EE5C54CBC34437472484FF061BF3FF5B" box="[151,254,199,223]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="31" pageNumber="2898" unit="mm" value="0.29">0.29 mm</quantity>
).
</paragraph>
<paragraph id="291BF92EC3443747249CFEC61A0BFE1B" blockId="31.[143,1104,263,415]" pageId="31" pageNumber="2898">
<emphasis id="1BD0253CC3443747249CFEC61BF3FE9B" bold="true" box="[143,254,263,287]" pageId="31" pageNumber="2898">Antennae.</emphasis>
Scape stout, thickened apically, slightly longer than three subsequent antennomeres combined,
<quantity id="EE5C54CBC344374725A6FEE61916FEBB" box="[437,539,295,319]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="31" pageNumber="2898" unit="mm" value="0.43">0.43 mm</quantity>
; antennomere II globular,
<quantity id="EE5C54CBC3443747275EFEE618B9FEBB" box="[845,948,295,319]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="31" pageNumber="2898" unit="mm" value="0.11">0.11 mm</quantity>
; III distinctly elongate, observably longer than II,
<quantity id="EE5C54CBC34437472634FE861981FEDB" box="[551,652,327,351]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="31" pageNumber="2898" unit="mm" value="0.14">0.14 mm</quantity>
; IV and V subequal in length,
<quantity id="EE5C54CBC344374727F5FE861F46FEDB" box="[998,1099,327,351]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="31" pageNumber="2898" unit="mm" value="0.08">0.08 mm</quantity>
; last antennomere short,
<quantity id="EE5C54CBC344374725B3FEA61909FEFB" box="[416,516,359,383]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="31" pageNumber="2898" unit="mm" value="0.18">0.18 mm</quantity>
, distinctly shorter than two preceding antennomeres combined.
</paragraph>
<paragraph id="291BF92EC3443747249CFE071AB3FDFA" blockId="31.[143,1104,454,638]" pageId="31" pageNumber="2898">
<emphasis id="1BD0253CC3443747249CFE071A14FE5A" bold="true" box="[143,281,454,478]" pageId="31" pageNumber="2898">Mouthparts.</emphasis>
Labrum transverse, with a pair of median teeth, and a lateral tooth on each side; median teeth protruding forwards, and lateral teeth slight protruding. Mandibles falciform, right one with a sharp tooth on inner edge. Maxillary palpus elongate, second segment longest, last segment fusiform, as long as the penultimate, apex subtruncated. Labial palpus slender, each segment in same length, last segment slightly dilated subapically.
</paragraph>
<paragraph id="291BF92EC3443747249CFD6418BAFD39" blockId="31.[143,951,677,701]" box="[143,951,677,701]" pageId="31" pageNumber="2898">
<emphasis id="1BD0253CC3443747249CFD641BDCFD39" bold="true" box="[143,209,677,701]" pageId="31" pageNumber="2898">Neck.</emphasis>
Average in wide (
<quantity id="EE5C54CBC34437472584FD641AF3FD39" box="[407,510,677,701]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="31" pageNumber="2898" unit="mm" value="0.36">0.36 mm</quantity>
), narrower than half the width of head.
</paragraph>
<paragraph id="291BF92EC3443747249CFD241A8AFC19" blockId="31.[143,1104,741,925]" pageId="31" pageNumber="2898">
<emphasis id="1BD0253CC3443747249CFD241A09FD79" bold="true" box="[143,260,741,765]" pageId="31" pageNumber="2898">Pronotum.</emphasis>
Subrectangular (PL to PW ratio 1.4), slightly longer, narrower than head. Widened anteriad, lateral margins slightly sinuate, anterior angles projecting, posterior angles broadly rounded. Dorsal integument extensively covered with micropunctures, with two medium-sized punctures on each anterolateral corner, one puncture on each posterolateral corner, and also with other irregular punctures near anterior and lateral margins of pronotum.
</paragraph>
<paragraph id="291BF92EC3443747249CFC051F18FC58" blockId="31.[143,1045,964,989]" box="[143,1045,964,989]" pageId="31" pageNumber="2898">
<emphasis id="1BD0253CC3443747249CFC051A36FC58" bold="true" box="[143,315,964,988]" pageId="31" pageNumber="2898">Mesoscutellum.</emphasis>
Shiny, rugose, with several irregularly scattered, small punctures.
</paragraph>
<paragraph id="291BF92EC3443747249CFBC519B2FB18" blockId="31.[143,1104,1028,1180]" pageId="31" pageNumber="2898">
<emphasis id="1BD0253CC3443747249CFBC51BD0FB98" bold="true" box="[143,221,1028,1052]" pageId="31" pageNumber="2898">Elytra.</emphasis>
Subrectangular, elongate (EL to EW ratio 1.3), longer, wider than pronotum. Humeri well developed, lateral margins divergent posteriorly, hind margin rounded. Dorsal integument shiny, flattened, without microsculpture; each elytron with a row of smaller punctures along median suture, a row along midwidth, also with three or four rows of larger punctures on deflexed portion.
</paragraph>
<paragraph id="291BF92EC3443747249CFB021892FA9F" blockId="31.[143,1103,1219,1308]" pageId="31" pageNumber="2898">
<emphasis id="1BD0253CC3443747249CFB021BC1FB5F" bold="true" box="[143,204,1219,1243]" pageId="31" pageNumber="2898">Legs.</emphasis>
First four segments of protarsi heart-shaped, dilated, and those of mesotarsi and metatarsi slender. Last segment of protarsi and mesotarsi shorter than the II–IV, respectively; last segment of metatarsi subequal to the length of II–IV.
</paragraph>
<paragraph id="291BF92EC3443747249CFA821877FA5F" blockId="31.[143,1104,1347,1499]" pageId="31" pageNumber="2898">
<emphasis id="1BD0253CC3443747249CFA821BF3FADF" bold="true" box="[143,254,1347,1371]" pageId="31" pageNumber="2898">Abdomen.</emphasis>
Cylindrical, broadest at segment VI. Tergites III–VII shiny, each segment covered with shallow transverse microstriae, with densely distributed punctures, distance between punctures about a puncture’s diameter. Each tergite with a basal impression near anterior margin, rugose, with dense, large punctures. All abdominal sternites shiny, with microstriae and punctures as those on tergites.
</paragraph>
</subSubSection>
<subSubSection id="61BEAAA5C3443747249CF9C3196BF99E" box="[143,614,1538,1562]" pageId="31" pageNumber="2898" type="distribution">
<paragraph id="291BF92EC3443747249CF9C3196BF99E" blockId="31.[143,614,1538,1562]" box="[143,614,1538,1562]" pageId="31" pageNumber="2898">
<emphasis id="1BD0253CC3443747249CF9C31A11F99E" bold="true" box="[143,284,1538,1562]" pageId="31" pageNumber="2898">Distribution.</emphasis>
<collectingCountry id="51B3B9BEC34437472530F9C31A52F99E" box="[291,351,1538,1562]" name="India" pageId="31" pageNumber="2898">India</collectingCountry>
(North region);
<collectingCountry id="51B3B9BEC3443747260AF9C3196CF99E" box="[537,609,1538,1562]" name="Nepal" pageId="31" pageNumber="2898">Nepal</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="61BEAAA5C3443747249CF98319A3F91F" pageId="31" pageNumber="2898" type="discussion">
<paragraph id="291BF92EC3443747249CF98319A3F91F" blockId="31.[143,1103,1602,1691]" pageId="31" pageNumber="2898">
<emphasis id="1BD0253CC3443747249CF9831BF5F9DE" bold="true" box="[143,248,1602,1626]" pageId="31" pageNumber="2898">Remarks.</emphasis>
This species can be distinguished from the other congeners by projecting laterad hind angles of head, protruding anterior angles of pronotum and a deep furrow formed by five or six large punctures behind eye.
</paragraph>
</subSubSection>
</treatment>
</document>
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