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<document ID-DOI="10.1093/zoolinnean/zlab015" ID-GBIF-Dataset="cc82accd-7fe1-4f9f-a60d-c7c121456114" ID-ISSN="0024-4082" ID-Zenodo-Dep="7551985" ID-ZooBank="42E7A370-FCD8-492A-8604-C42863BDE6E2" checkinTime="1647256475265" checkinUser="plazi" docAuthor="Cupello, Mario, Ribeiro-Costa, Cibele S. &amp; Vaz-De-Mello, Fernando Z." docDate="2022" docId="27085036DC08FFE53ECE08B8DE29F8A1" docLanguage="en" docName="zlab015.pdf" docOrigin="Zoological Journal of the Linnean Society 194" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-.journal_article" docStyleVersion="7" docTitle="Bolbites onitoides Harold des Cupello 1868" docType="treatment" docVersion="6" lastPageNumber="997" masterDocId="DB31284EDC00FFFC3E380D0BDB22FFF3" masterDocTitle="The evolution of Bolbites onitoides (Coleoptera: Scarabaeidae: Phanaeini): its phylogenetic significance, geographical polychromatism and the subspecies problem" masterLastPageNumber="1034" masterPageNumber="973" pageNumber="981" updateTime="1674153317425" updateUser="ExternalLinkService">
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<mods:title>The evolution of Bolbites onitoides (Coleoptera: Scarabaeidae: Phanaeini): its phylogenetic significance, geographical polychromatism and the subspecies problem</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Cupello, Mario</mods:namePart>
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<mods:namePart>Ribeiro-Costa, Cibele S.</mods:namePart>
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<mods:namePart>Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:date>2022</mods:date>
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<mods:number>2021-07-12</mods:number>
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<taxonomicName ID-CoL="5WMQM" ID-ENA="377182" authority="HAROLD, 1868" authorityName="Harold des Cupello" authorityYear="1868" box="[246,659,1458,1483]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1013" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[246,488,1459,1482]" italics="true" pageId="8" pageNumber="981">BOLBITES ONITOIDES</emphasis>
HAROLD, 1868
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(
<figureCitation box="[394,511,1497,1522]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-10="Figure 11" captionStart-11="Figure 12" captionStart-12="Figure 13" captionStart-13="Figure 14" captionStart-14="Figure 15" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStart-5="Figure 6" captionStart-6="Figure 7" captionStart-7="Figure 8" captionStart-8="Figure 9" captionStart-9="Figure 10" captionStartId-0="2.[145,223,1852,1874]" captionStartId-1="4.[145,223,1515,1537]" captionStartId-10="32.[145,223,1631,1653]" captionStartId-11="33.[163,241,1286,1308]" captionStartId-12="37.[163,242,1148,1170]" captionStartId-13="38.[145,225,1140,1162]" captionStartId-14="39.[164,242,1171,1193]" captionStartId-2="10.[145,223,1255,1277]" captionStartId-3="11.[163,245,1393,1415]" captionStartId-4="12.[145,223,1538,1560]" captionStartId-5="13.[164,244,1206,1228]" captionStartId-6="14.[145,222,730,752]" captionStartId-7="15.[163,241,1105,1127]" captionStartId-8="16.[145,225,1732,1754]" captionStartId-9="30.[145,223,1487,1509]" captionTargetBox-0="[217,1343,203,1809]" captionTargetBox-1="[274,1291,204,1473]" captionTargetBox-10="[147,1428,197,1590]" captionTargetBox-11="[166,1446,198,1244]" captionTargetBox-12="[166,1446,198,1105]" captionTargetBox-13="[147,1428,197,1100]" captionTargetBox-14="[166,1446,198,1117]" captionTargetBox-2="[209,1358,197,1213]" captionTargetBox-3="[166,1447,198,1350]" captionTargetBox-4="[147,1408,197,1497]" captionTargetBox-5="[165,1446,197,1164]" captionTargetBox-6="[147,1429,197,689]" captionTargetBox-7="[181,784,202,1062]" captionTargetBox-8="[305,1265,198,1689]" captionTargetBox-9="[146,1423,197,1438]" captionTargetId-0="figure-25@2.[207,1362,195,1812]" captionTargetId-1="figure-160@4.[271,1298,195,1475]" captionTargetId-10="figure-25@32.[145,1428,195,1591]" captionTargetId-11="figure-283@33.[163,1449,195,1247]" captionTargetId-12="figure-332@37.[163,1449,195,1108]" captionTargetId-13="figure-334@38.[145,1428,195,1101]" captionTargetId-14="figure-371@39.[163,1449,195,1131]" captionTargetId-2="figure-230@10.[207,1362,195,1215]" captionTargetId-3="figure-147@11.[163,1450,195,1353]" captionTargetId-4="figure-25@12.[145,1428,195,1498]" captionTargetId-5="figure-317@13.[163,1446,195,1166]" captionTargetId-6="figure-598@14.[145,1429,195,690]" captionTargetId-7="figure-551@15.[163,784,195,1065]" captionTargetId-8="figure-25@16.[302,1268,195,1692]" captionTargetId-9="figure-25@30.[145,1425,195,1447]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-10="32" captionTargetPageId-11="33" captionTargetPageId-12="37" captionTargetPageId-13="38" captionTargetPageId-14="39" captionTargetPageId-2="10" captionTargetPageId-3="11" captionTargetPageId-4="12" captionTargetPageId-5="13" captionTargetPageId-6="14" captionTargetPageId-7="15" captionTargetPageId-8="16" captionTargetPageId-9="30" captionText-0="Figure 1. The wide colour variation shown by Bolbites onitoides. A, copper (male). B, red (male). C, Copper individual with greenish pronotum and elytra (female). D, atypical bicoloured individual with unusually green elytra (female). E, F, typical bicoloured individuals (males). GI, blue individuals (males). Although Martínez (1959, 1987) thought that two subspecies could be recognized one for red individuals and the other for blue ones two facts observed by us do not support this idea.First, two kinds of intermediates exist connecting the two main colour patterns of red (B) and blue individuals (GI): specimens with a red pronotum and blue or green elytra (DF), which we call the discrete bicoloured intermediates, and the second being the gradual colour-blended intermediates, or specimens that lie at the centre of the variation spectrum between the two colour ends, which includes specimens showing dorsal colouration entirely coppery (A), coppery with greenish reflections (C), and coppery with reddish reflections. Secondly, no other features were found to vary geographically along with colour. Therefore, we interpret the plethora of colour variants as intraspecific variation of a single monotypic species." captionText-1="Figure 2. Ventral sexual dimorphism in Bolbites onitoides. A, lateral view of a male. B, C, ventral view: B, male: C, female. Note the conspicuous ventral sexual dimorphism in this species, including the presence of several teeth and spines all over the body of the male, the differences in the overall shape of the protibiae, the pilosity of the metacoxae, and colour of the metaventrite." captionText-10="Figure 11. Geographical range of B. onitoides showing the differential distribution of the colour variants. It is indisputable that there is a strong correlation between colour and geographical provenance in the species: blue individuals are abundant at the western-half of the distribution, while red specimens are dominant at the eastern-half. Nevertheless, two kinds of intermediates the discrete bicoloured and the gradual colour-blended are found throughout the range of B. onitoides, but nowhere are they the dominant variant. Note that there is a zone of overlap between the distribution of red and blue variants in Paraguay and Argentina, and that three localities recorded red, blue and bicoloured individuals and a fourth recorded all four variants." captionText-11="Figure 12. Distribution of each colour variant showed in separate maps for an easier discrimination. See legend for Figure 10 for more details." captionText-12="Figure 13. Predicted areas of environmental suitability (temperature and precipitation) for the red (upper row) and blue (lower row) variants of Bolbites onitoides within the range of the species. Note that the area of suitable environmental conditions for each colour variant within the range of B. onitoides, particularly when precipitation is taken into account, corresponds largely to the actual range of each variant. Furthermore, these areas largely do not overlap with each other. Confirming our initial hypothesis, this result indicates that the differential distribution of the red and blue variants across the populations of B. onitoides is likely a response to varying environmental conditions." captionText-13="Figure 14. Predicted areas of environmental suitability (temperature and precipitation) for the bicoloured (upper row) and coppery (lower row) variants of Bolbites onitoides within the range of the species. Note that the area of environmental suitability for each of these variants matches their observed geographical ranges and is largely corresponded to the range of the species as a whole (particularly the bicoloured one). They broadly overlap with each other and with the suitable areas for the red and blue variants (Fig. 13)." captionText-14="Figure 15. Predicted area of environmental suitability for Bolbites onitoides in South America (to the left) and the realized geographical range of the species (to the right)." captionText-2="Figure 3. Dorsal sexual monomorphism in Bolbites onitoides. A, frontal view of a male. B, C, head and pronotum: B, male; C, female. Note how little the sexes differ dorsally. Unlike most groups of Phanaeina, males of B. onitoides lack any cephalic or pronotal horns and have a pronotal ridge identical to that of the females. This dorsal monomorphism is in striking contrast with the sexual dimorphism found on the venter. Note, however, the sexually dimorphic extension of iridescence over the clypeus, males having at least its basal half metallic, while females are entirely (or almost entirely) dull black." captionText-3="Figure 4. Meso- and metaventrite in Bolbites and Gromphadina. A, Bolbites onitoides. B, Gromphas inermis. C, D, Oruscatus davus: C, male; D, female. The carina of the mesofurcal brace constitutes the posterior branch of the mesoventral ridge and gives it the shape of a cross (orange arrow). This cross-shaped ridge is typical of the Gromphadina and is also present in at least some Phanaeina. Note that, while in the Gromphadina the posterior branch connects the mesoventral ridge to the anterior edge of the metaventrite, no such connection exists in most Bolbites and and the other Phanaeina whose mesoventral ridge is T-shaped. In Oruscatus, although present in both sexes, the posterior branch of the mesoventral ridge is much more clearly delimited in males than in females; in females, the posterior branch is usually incomplete and interrupted, while in males it is always sharp and continuous. Note also other differences between Bolbites and the Gromphadina relating to the shape of the metaventral anterior prominence and its integument and lateral pilosity." captionText-4="Figure 5. Legs. AE, sexual dimorphism on the front legs of B. onitoides. A, B, profemora: A, male; B, female. Note that the male profemur is claviform and has a strong emargination on its posterior edge (indicated by the red arrow), while that of the female is shorter and has a simply rounded aspect. CE, protibiae: C, D, dorsal view of male (C) and female (D). Observe that the male protibia is much longer and slenderer than that of the female and has a well-delimited tuft of long setae at its internal apical angle. E, lateral view of the male protibia. In contrast to females, males have a series of sharp, uneven teeth along the ventral protibial carina. Red arrow indicates the central tooth, which is longer than the others. F, G, meso- and metatarsi of B. onitoides. Unlike all other Phanaeini, which are clawless, B. onitoides has a pair of well-developed claws on the apical meso- and metatarsomeres. A small prolongation raises from underneath the claws and project itself anteriorly (indicated by the red arrow in F); this prolongation seems to be homologous to the tapered apex of the apical meso- and metatarsomeres of the other Phanaeini and is probably a synapomorphy of the clade. H, metanepisternum and its long tab typical of the Phanaeina." captionText-5="Figure 6. Sexual dimorphism on the hind legs of B. onitoides. A, B, metacoxae: A, male; B, female. Note the plate-like projections and the long spines directed posteriorly in the male and which are lacking in the female. C, D, metafemora and metatibiae: C, male; D, female. Observe the ridge on the posterior edge of the metafemora and the sharp ridge on the external and internal edges of the metatibiae of the males, all of which are absent in females." captionText-6="Figure 7. Sexual dimorphism on the pygidium and at the apex of the sutural margin of the elytra of B. onitoides: A, male; B, female. Orange arrow shows the short spines of the apex of the sutural margin of the elytra, whereas red arrow indicates the long spines raising from the basolateral angles of the pygidium of males. Both structures are absent in females." captionText-7="Figure 8. Genitalia of both sexes of B. onitoides. A, female: spermatheca. BD, male: B, genital segment (i.e. nineth abdominal segment). C, D, tegmen. C, dorsal view. D, lateral view. LSP = lateral sclerotized plate; MSP = medial sclerotized plate." captionText-8="Figure 9. Male genitalia: Endophallus of B. onitoides. A, B, general view of the extracted endophallus. A, dorsal view. B, ventral view. CG, endophallites: C, axial + subaxial endophallite complex (dissected). D, fronto-lateral peripheral endophallite (ventral view). E, medial endophallite (dorsal view). FH, superior right peripheral endophallite from different views: F, G, ventral view: F, seahorse-shaped; G, F-shaped. H, lateral view.A = axial endophallite; SA = subaxial endophallite; MS = medial endophallite; FLP = fronto-lateral peripheral endophallite; SRP = superior right peripheral endophallite." captionText-9="Figure 10. Phylogenetic placement of Bolbites onitoides among the Phanaeini and the evolution of iridescent colouration in this clade. A growing body of evidence has accumulated over the past two decades confirming that the horneless Bolbites is a phanaeine lineage that is more closely associated to horned Phanaeina than to the primitively hornless Gromphadina (see references in the text). If this is correct, the evolution of iridescence from the plesiomorphic opaque condition preceded that of the typically male cephalic and pronotal ornamentation found widely among the Phanaeina.As a consequence, the selective pressure that led to the evolution of the two features cannot have been the same. A more likely scenario is that iridescence first evolved serving another function in the common ancestors of the Phanaeini e.g. thermoregulation or aposematism and was inherited serving this function by the the Gromphadina and Phanaeina alike. Then, iridescence was putatively co-opted independently during the parallel evolution of cephalic horns by the Phanaeina less Bolbites and the gromphadine clade Gromphas aeruginosa + G. lemoinei to serve a new role in the evaluation of male horns during sexual interactions. Sexual selection thus co-opted for a new use a feature that had originally evolved under natural selection serving another function. This characterizes iridescence as an exaptation for horn evaluation, not an adaptation for this function. Species illustrated (from top to bottom): Eucranium cyclosoma Burmeister, 1861, Oruscatus davus (Erichson, 1847), Gromphas lemoinei Waterhouse, 1891, G. dichroa Blanchard, 1846, Bolbites onitoides Harold, 1868, a red Coprophanaeus saphirinus (Sturm, 1826), a blue C. saphirinus, and Sulcophanaeus menelas (Castelnau, 1840)." figureDoi-0="http://doi.org/10.5281/zenodo.7551987" figureDoi-1="http://doi.org/10.5281/zenodo.7551989" figureDoi-10="http://doi.org/10.5281/zenodo.7552016" figureDoi-11="http://doi.org/10.5281/zenodo.7552018" figureDoi-12="http://doi.org/10.5281/zenodo.7552020" figureDoi-13="http://doi.org/10.5281/zenodo.7552022" figureDoi-14="http://doi.org/10.5281/zenodo.7552024" figureDoi-2="http://doi.org/10.5281/zenodo.7551991" figureDoi-3="http://doi.org/10.5281/zenodo.7551994" figureDoi-4="http://doi.org/10.5281/zenodo.7551998" figureDoi-5="http://doi.org/10.5281/zenodo.7552000" figureDoi-6="http://doi.org/10.5281/zenodo.7552002" figureDoi-7="http://doi.org/10.5281/zenodo.7552004" figureDoi-8="http://doi.org/10.5281/zenodo.7552012" figureDoi-9="http://doi.org/10.5281/zenodo.7552014" httpUri-0="https://zenodo.org/record/7551987/files/figure.png" httpUri-1="https://zenodo.org/record/7551989/files/figure.png" httpUri-10="https://zenodo.org/record/7552016/files/figure.png" httpUri-11="https://zenodo.org/record/7552018/files/figure.png" httpUri-12="https://zenodo.org/record/7552020/files/figure.png" httpUri-13="https://zenodo.org/record/7552022/files/figure.png" httpUri-14="https://zenodo.org/record/7552024/files/figure.png" httpUri-2="https://zenodo.org/record/7551991/files/figure.png" httpUri-3="https://zenodo.org/record/7551994/files/figure.png" httpUri-4="https://zenodo.org/record/7551998/files/figure.png" httpUri-5="https://zenodo.org/record/7552000/files/figure.png" httpUri-6="https://zenodo.org/record/7552002/files/figure.png" httpUri-7="https://zenodo.org/record/7552004/files/figure.png" httpUri-8="https://zenodo.org/record/7552012/files/figure.png" httpUri-9="https://zenodo.org/record/7552014/files/figure.png" pageId="8" pageNumber="981">FIGS 115</figureCitation>
)
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<treatmentCitation author="Fabre J-H" box="[566,756,1598,1620]" page="73" pageId="8" pageNumber="981" year="1899">
<bibRefCitation author="Fabre J-H" box="[566,756,1598,1620]" pageId="8" pageNumber="981" refId="ref48519" refString="Fabre J-H. 1899. Souvenirs entomologiques (sixieme serie). Etudes sur l'instict et les moeurs des insectes. Paris: Librairie Ch. Delagrave, 418." type="book" year="1899">Fabre, 1899: 73</bibRefCitation>
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<bibRefCitation author="Judulien F" box="[228,454,1629,1651]" pageId="8" pageNumber="981" pagination="371 - 380" refId="ref50626" refString="Judulien F. 1899. Quelques notes sur plusieurs coprophages de Buenos Aires. Revista del Museo de La Plata 9: 371 - 380. [Frere Judulien was the religious name of the French- Argentinian entomologist Jean (Juan) Brethes (1871 - 1928) and it was with this name he signed his early works (Doello- Jurado, 1928)]." type="journal article" year="1899">Judulien, 1899: 376</bibRefCitation>
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, figs 5, 7;
<treatmentCitation author="Laloy L" box="[570,756,1629,1651]" page="171" pageId="8" pageNumber="981" year="1902">
<bibRefCitation author="Laloy L" box="[570,756,1629,1651]" pageId="8" pageNumber="981" pagination="169 - 171" refId="ref51055" refString="Laloy L. 1902. Moeurs des coleopteres coprophages, specialement de La Plata. Le Naturaliste, 2 e Serie 16: 169 - 171." type="journal article" year="1902">Laloy, 1902: 171</bibRefCitation>
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;
<treatmentCitation author="Kolbe HJ" box="[145,343,1660,1682]" page="487" pageId="8" pageNumber="981" year="1905">
<bibRefCitation author="Kolbe HJ" box="[145,343,1660,1682]" pageId="8" pageNumber="981" pagination="475 - 594" refId="ref50884" refString="Kolbe HJ. 1905. Uber die Lebensweise und die geographische Verbreitung der coprophagen Lamellicornier. Zoologischen Jahrbucher, Supplement 8: 475 - 594." type="journal article" year="1905">Kolbe, 1905: 487</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Bruch C" box="[356,559,1660,1682]" page="189" pageId="8" pageNumber="981" year="1911">
<bibRefCitation author="Bruch C" box="[356,559,1660,1682]" pageId="8" pageNumber="981" pagination="143 - 225" refId="ref46594" refString="Bruch C. 1911. Catalogo sistematico de los coleopteros de la Republica Argentina. Revista del Museo de la Plata 17: 143 - 225." type="journal article" year="1911">Bruch, 1911: 189</bibRefCitation>
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;
<treatmentCitation author="Gillet JJE" box="[572,755,1660,1682]" page="80" pageId="8" pageNumber="981" year="1911">
<bibRefCitation author="Gillet JJE" box="[572,755,1660,1682]" pageId="8" pageNumber="981" refId="ref49242" refString="Gillet JJE. 1911. Coleopterorum catalogus. Pars 38: Scarabaeidae: Coprinae I. Berlin: W. Junk, 100." type="book" year="1911">Gillet, 1911: 80</bibRefCitation>
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;
<treatmentCitation author="Fabre J-H" box="[145,398,1690,1712]" page="242" pageId="8" pageNumber="981" year="1919">
<bibRefCitation author="Fabre J-H" box="[145,398,1690,1712]" pageId="8" pageNumber="981" refId="ref48549" refString="Fabre J-H. 1919. The glow-worm and other beetles. Toronto: McClelland &amp; Stewart, 488." type="book" year="1919">Fabre, 1919: 242244</bibRefCitation>
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;
<treatmentCitation author="Lucas R" box="[410,609,1691,1712]" page="141" pageId="8" pageNumber="981" year="1920">
<bibRefCitation author="Lucas R" box="[410,609,1691,1712]" pageId="8" pageNumber="981" refId="ref51288" refString="Lucas R. 1920. Catalogus alphabeticus generum et subgenerum Coleopterorum orbis terrarum totius (famil., trib., subtr., sect. incl.). Pars I. Berlin: Nicolaische." type="book" year="1920">Lucas, 1920: 141</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="d'Olsoufieff G" page="20" pageId="8" pageNumber="981" year="1924">
<bibRefCitation author="d'Olsoufieff G" pageId="8" pageNumber="981" pagination="4 - 201" refId="ref52767" refString="d'Olsoufieff G. 1924. Les Phanaeides (Coleoptera - Lamellicornia), famille Scarabaeidae - tr. Coprini. Insecta 13: 4 - 201." type="journal article" year="1924">dOlsoufieff, 1924: 20</bibRefCitation>
</treatmentCitation>
, 58, 136, 138, pl. 3, figs 1, 10;
<treatmentCitation author="Janssens A" page="1" pageId="8" pageNumber="981" year="1940">
<bibRefCitation author="Janssens A" pageId="8" pageNumber="981" pagination="1 - 7" refId="ref50591" refString="Janssens A. 1940. Contribution a l'Etude des Coleopteres lamellicornes coprophages. II. - Remarques sur quelques Phanaeides. Bulletin du Musee Royal d'Histoire Naturelle de Belgique 16: 1 - 7." type="journal article" year="1940">Janssens, 1940: 1</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Balthasar V" box="[170,404,1752,1774]" page="350" pageId="8" pageNumber="981" year="1941">
<bibRefCitation author="Balthasar V" box="[170,404,1752,1774]" pageId="8" pageNumber="981" pagination="337 - 358" refId="ref46125" refString="Balthasar V. 1941. Scarabaeidae Laparostictae (Col.). Beitrage Fauna Perus 1: 337 - 358." type="journal article" year="1941">Balthasar, 1941: 350</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Pessoa SB &amp; Lane F" box="[414,756,1752,1773]" page="470" pageId="8" pageNumber="981" year="1941">
<bibRefCitation author="Pessoa SB &amp; Lane F" box="[414,756,1752,1773]" pageId="8" pageNumber="981" pagination="389 - 504" refId="ref53199" refString="Pessoa SB, Lane F. 1941. Coleopteros necrofagos de interesse medico-legal. Ensaio monografico sobre a familia Scarabaeidae de S. Paulo e regioes vizinhas. Arquivos de Zoologia do Estado de Sao Paulo 2: 389 - 504." type="journal article" year="1941">Pessôa &amp; Lane, 1941: 470471</bibRefCitation>
</treatmentCitation>
, fig. 32, pl. 10, fig. 75;
<treatmentCitation author="Blackwelder RE" box="[374,634,1782,1804]" page="208" pageId="8" pageNumber="981" year="1944">
<bibRefCitation author="Blackwelder RE" box="[374,634,1782,1804]" pageId="8" pageNumber="981" pagination="189 - 341" refId="ref46316" refString="Blackwelder RE. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America. Part 2. United States National Museum Bulletin 185: 189 - 341." type="journal article" year="1944">Blackwelder, 1944: 208</bibRefCitation>
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;
<treatmentCitation author="Balthasar V" page="335" pageId="8" pageNumber="981" year="1951">
<bibRefCitation author="Balthasar V" pageId="8" pageNumber="981" pagination="322 - 344" refId="ref46146" refString="Balthasar V. 1951. Scarabaeidae Laparostictae (Col.). Beitrage Fauna Perus 2: 322 - 344." type="journal article" year="1951">Balthasar, 1951: 335</bibRefCitation>
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;
<treatmentCitation author="Martinez A" box="[272,496,1813,1835]" page="94" pageId="8" pageNumber="981" year="1959">
<bibRefCitation author="Martinez A" box="[272,496,1813,1835]" pageId="8" pageNumber="981" pagination="1 - 126" refId="ref51548" refString="Martinez A. 1959. Catalogo de los Scarabaeidae Argentinos. Revista del Museo Argentino de Ciencias Naturales ' Bernardino Rivadavia', Ciencias Zoologicas 5: 1 - 126." type="journal article" year="1959">Martínez, 1959: 94</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Halffter G &amp; Matthews EG" page="132" pageId="8" pageNumber="981" year="1966">
<bibRefCitation author="Halffter G &amp; Matthews EG" pageId="8" pageNumber="981" pagination="1 - 312" refId="ref49650" refString="Halffter G, Matthews EG. 1966. The natural history of dung beetles of the subfamily Scarabaeinae (Coleoptera: Scarabaeidae). Folia Entomologica Mexicana 12 - 14: 1 - 312." type="journal article" year="1966">Halffter &amp; Matthews, 1966: 132</bibRefCitation>
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;
<treatmentCitation author="Krantz GW" box="[268,459,1844,1866]" page="12" pageId="8" pageNumber="981" year="1967">
<bibRefCitation author="Krantz GW" box="[268,459,1844,1866]" pageId="8" pageNumber="981" pagination="1 - 89" refId="ref50988" refString="Krantz GW. 1967. A review of the genus Holocelaeno Berlese, 1910 (Acarina: Macrochelidae). Acarologia 9 (Suppl.): 1 - 89." type="journal article" year="1967">Krantz, 1967: 12</bibRefCitation>
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, 48, 53, 55, 8687; Haffter &amp; Matthews, 1971: 159;
<treatmentCitation author="Edmonds WD" box="[447,696,1874,1896]" page="818" pageId="8" pageNumber="981" year="1972">
<bibRefCitation author="Edmonds WD" box="[447,696,1874,1896]" pageId="8" pageNumber="981" pagination="731 - 874" refId="ref48078" refString="Edmonds WD. 1972. Comparative skeletal morphology, systematics and evolution of the Phanaeine dung beetles (Coleoptera: Scarabaeidae). University of Kansas Science Bulletin 49: 731 - 874." type="journal article" year="1972">Edmonds, 1972: 818</bibRefCitation>
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, 854;
<treatmentCitation author="Halffter G &amp; Edmonds WD" box="[809,1182,197,219]" page="86" pageId="8" pageNumber="981" year="1982">
<bibRefCitation author="Halffter G &amp; Edmonds WD" box="[809,1182,197,219]" pageId="8" pageNumber="981" refId="ref49587" refString="Halffter G, Edmonds WD. 1982. The nesting behavior of dung beetles (Scarabaeinae). An ecological and evolutive approach. Mexico: Instituto de Ecologia, 176." type="book" year="1982">Halffter &amp; Edmonds, 1982: 86</bibRefCitation>
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, 89;
<treatmentCitation author="Kirk AA" box="[1242,1420,197,219]" page="53" pageId="8" pageNumber="981" year="1992">
<bibRefCitation author="Kirk AA" box="[1242,1420,197,219]" pageId="8" pageNumber="981" pagination="45 - 54" refId="ref50693" refString="Kirk AA. 1992. Dung beetles (Coleoptera: Scarabaeidae) active in patchy forest and pasture habitats in Santa Cruz province, Bolivia, during spring. Folia Entomologica Mexicana 84: 45 - 54." type="journal article" year="1992">Kirk, 1992: 53</bibRefCitation>
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;
<treatmentCitation author="Luzzatto M &amp; Monteresino E &amp; Zunino M &amp; Barbero E" box="[809,1092,228,250]" page="63" pageId="8" pageNumber="981" year="1994">
<bibRefCitation author="Luzzatto M &amp; Monteresino E &amp; Zunino M &amp; Barbero E" box="[809,1092,228,250]" pageId="8" pageNumber="981" pagination="61 - 65" refId="ref51325" refString="Luzzatto M, Monteresino E, Zunino M, Barbero E. 1994. Material per un'analisi del popolamento di scarabeidi coprofagi delle zone aride del cono sud del Sudamerica. Bollettino della Societa Entomologica Italiana 126: 61 - 65." type="journal article" year="1994">
Luzzatto
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;
<treatmentCitation author="Cabrera Walsh G &amp; Gandolfo D" page="582" pageId="8" pageNumber="981" year="1996">
<bibRefCitation author="Cabrera Walsh G &amp; Gandolfo D" pageId="8" pageNumber="981" pagination="581 - 588" refId="ref46837" refString="Cabrera Walsh G, Gandolfo D. 1996. Nidification of thirteen common Argentine dung beetles (Scarabaeidae: Scarabaeinae). Annals of the Entomological Society of America 89: 581 - 588." type="journal article" year="1996">Cabrera Walsh &amp; Gandolfo, 1996: 582587</bibRefCitation>
</treatmentCitation>
, fig. 1i;
<treatmentCitation author="Monteresino E &amp; Martinez A &amp; Zunino M" box="[1075,1420,258,280]" page="109" pageId="8" pageNumber="981" year="1996">
<bibRefCitation author="Monteresino E &amp; Martinez A &amp; Zunino M" box="[1075,1420,258,280]" pageId="8" pageNumber="981" pagination="101 - 117" refId="ref52229" refString="Monteresino E, Martinez A, Zunino M. 1996. Los Scarabaeinae (Coleoptera, Scarabaeidae) de la provincia de Cordoba, Argentina. In: di Tada IE, Bucher EH, eds. Biodiversidad de la Provincia de Cordoba. Fauna, Vol. 1. Cordoba: Universidad Nacional de Rio Cuarto, 101 - 117." type="book chapter" year="1996">
Monteresino
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;
<treatmentCitation author="Morelli E &amp; Gonzalez-Vainer P" box="[809,1253,289,311]" page="197" pageId="8" pageNumber="981" year="1997">
<bibRefCitation author="Morelli E &amp; Gonzalez-Vainer P" box="[809,1253,289,311]" pageId="8" pageNumber="981" pagination="197" refId="ref52291" refString="Morelli E, Gonzalez-Vainer P. 1997. Dung beetles (Coleoptera: Scarabaeidae) inhabiting bovine and ovine droppings in Uruguayan prairies. The Coleopterists Bulletin 51: 197." type="journal article" year="1997">Morelli &amp; Gonzalez-Vainer, 1997: 197</bibRefCitation>
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;
<treatmentCitation author="Morelli E &amp; Gonzalez-Vainer P &amp; Baz A" page="54" pageId="8" pageNumber="981" year="2002">
<bibRefCitation author="Morelli E &amp; Gonzalez-Vainer P &amp; Baz A" pageId="8" pageNumber="981" pagination="53 - 57" refId="ref52322" refString="Morelli E, Gonzalez-Vainer P, Baz A. 2002. Coprophagous beetles (Coleoptera: Scarabaeoidea) in Uruguayan prairies: abundance, diversity and seasonal occurrence. Studies on Neotropical Fauna and Environment 37: 53 - 57." type="journal article" year="2002">
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<treatmentCitation author="Philips TK &amp; Edmonds WD &amp; Scholtz CH" box="[917,1184,320,342]" page="46" pageId="8" pageNumber="981" year="2004">
<bibRefCitation author="Philips TK &amp; Edmonds WD &amp; Scholtz CH" box="[917,1184,320,342]" pageId="8" pageNumber="981" pagination="43 - 63" refId="ref53431" refString="Philips TK, Edmonds WD, Scholtz CH. 2004 b. A phylogenetic analysis of the New World tribe Phanaeini (Coleoptera: Scarabaeidae: Scarabaeinae): hypotheses on relationship and origins. Insect Systematics and Evolution 35: 43 - 63." type="journal article" year="2004">
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, 56, 5859, 62, figs 2, 18, 30;
<treatmentCitation author="Hamel-Leigue AC &amp; Mann DJ &amp; Vaz-de-Mello FZ &amp; Herzog SK" box="[886,1199,350,372]" page="6" pageId="8" pageNumber="981" year="2006">
<bibRefCitation author="Hamel-Leigue AC &amp; Mann DJ &amp; Vaz-de-Mello FZ &amp; Herzog SK" box="[886,1199,350,372]" pageId="8" pageNumber="981" pagination="1 - 18" refId="ref49745" refString="Hamel-Leigue AC, Mann DJ, Vaz-de-Mello FZ, Herzog SK. 2006. Hacia un inventario de los escarabajos peloteros (Coleoptera: Scarabaeidae) de Bolivia: primera compilacion de los generos y especies registrados para el pais. Revista Boliviana de Ecologia y Conservacion Ambiental 20: 1 - 18." type="journal article" year="2006">
Hamel-Leigue
<emphasis box="[1053,1107,350,372]" italics="true" pageId="8" pageNumber="981">et al.</emphasis>
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</bibRefCitation>
</treatmentCitation>
, figs 4142;
<treatmentCitation author="Krajcik M" page="16" pageId="8" pageNumber="981" year="2006">
<bibRefCitation author="Krajcik M" pageId="8" pageNumber="981" pagination="1 - 189" refId="ref50932" refString="Krajcik M. 2006. Checklist of Scarabaeoidea of the world. 1. Scarabaeinae (Coleoptera: Scarabaeidae: Scarabaeinae). Anima. X, Supplement 3: 1 - 189." type="journal article" year="2006">Krajcik, 2006: 16</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Ocampo FC &amp; Hawks DC" box="[933,1298,381,403]" page="562" pageId="8" pageNumber="981" year="2006">
<bibRefCitation author="Ocampo FC &amp; Hawks DC" box="[933,1298,381,403]" pageId="8" pageNumber="981" pagination="557 - 570" refId="ref52651" refString="Ocampo FC, Hawks DC. 2006. Molecular phylogenetics and evolution of the food relocation behaviour of the dung beetle tribe Eucraniini (Coleoptera: Scarabaeidae: Scarabaeinae). Invertebrate Systematics 20: 557 - 570." type="journal article" year="2006">Ocampo &amp; Hawks, 2006: 562</bibRefCitation>
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, 564567, figs 58;
<treatmentCitation author="Hamel-Leigue AC &amp; Herzog SK &amp; Mann DJ &amp; Larsen TH &amp; Gill BD &amp; Edmonds WD &amp; Spector S" box="[908,1237,412,434]" page="49" pageId="8" pageNumber="981" year="2009">
<bibRefCitation author="Hamel-Leigue AC &amp; Herzog SK &amp; Mann DJ &amp; Larsen TH &amp; Gill BD &amp; Edmonds WD &amp; Spector S" box="[908,1237,412,434]" pageId="8" pageNumber="981" pagination="43 - 95" refId="ref49799" refString="Hamel-Leigue AC, Herzog SK, Mann DJ, Larsen TH, Gill BD, Edmonds WD, Spector S. 2009. Distribucion e historia natural de escarabajos de la tribu Phanaeini (Coleoptera: Scarabaeidae: Scarabaeinae) en Bolivia. Kempffiana 5: 43 - 95." type="journal article" year="2009">
Hamel-Leigue
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, 2009: 49
</bibRefCitation>
</treatmentCitation>
, 56, 69, 85, fig. 8, photos 12;
<treatmentCitation author="Scholtz CH" box="[943,1217,442,464]" page="254" pageId="8" pageNumber="981" year="2009">
<bibRefCitation author="Scholtz CH" box="[943,1217,442,464]" pageId="8" pageNumber="981" pagination="225 - 327" refId="ref54323" refString="Scholtz CH. 2009 b. Section C. Phylogeny of the Scarabaeinae. In: Scholtz CH, Davis ALV, Kryger U, eds. Evolutionary biology and conservation of dung beetles. Sofia: Pensoft Publishers, 225 - 327." type="book chapter" year="2009">Scholtz, 2009b: 254255</bibRefCitation>
</treatmentCitation>
, figs 12.6ab; Vazde-Mello &amp; Grossi, 2010: 220;
<treatmentCitation author="Vaz-de-Mello FZ &amp; Edmonds WD &amp; Ocampo FC &amp; Schoolmeesters P" page="44" pageId="8" pageNumber="981" year="2011">
<bibRefCitation author="Vaz-de-Mello FZ &amp; Edmonds WD &amp; Ocampo FC &amp; Schoolmeesters P" pageId="8" pageNumber="981" pagination="1 - 73" refId="ref55440" refString="Vaz-de-Mello FZ, Edmonds WD, Ocampo FC, Schoolmeesters P. 2011. A multilingual key to the genera and subgenera of the subfamily Scarabaeinae of the New World (Coleoptera: Scarabaeidae). Zootaxa 2854: 1 - 73." type="journal article" year="2011">
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</bibRefCitation>
</treatmentCitation>
, 51, fig. 22;
<treatmentCitation author="Krajcik M" box="[986,1195,504,526]" page="54" pageId="8" pageNumber="981" year="2012">
<bibRefCitation author="Krajcik M" box="[986,1195,504,526]" pageId="8" pageNumber="981" pagination="1 - 278" refId="ref50966" refString="Krajcik M. 2012. Checklist of the world Scarabaeoidea. Anima. X, Supplement 5: 1 - 278." type="journal article" year="2012">Krajcik, 2012: 54</bibRefCitation>
</treatmentCitation>
; Cupello &amp; Vaz-de-Mello, 2013: 441442;
<treatmentCitation author="Cupello M &amp; Vaz-de-Mello FZ" page="16" pageId="8" pageNumber="981" year="2015">
<bibRefCitation author="Cupello M &amp; Vaz-de-Mello FZ" pageId="8" pageNumber="981" pagination="1 - 27" refId="ref47264" refString="Cupello M, Vaz-de-Mello FZ. 2015. A new species and the phylogeny of the South American genus Gromphas Brulle, 1837 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini). Journal of Natural History (online edition): 1 - 27." type="book chapter" year="2015">Cupello &amp; Vaz-de-Mello, 2015: 16</bibRefCitation>
</treatmentCitation>
, 1819, figs 7C, 8C;
<treatmentCitation author="Alonso CBG &amp; Zurita GA &amp; Bellocq MI" box="[1061,1321,565,587]" page="299" pageId="8" pageNumber="981" year="2019">
<bibRefCitation author="Alonso CBG &amp; Zurita GA &amp; Bellocq MI" box="[1061,1321,565,587]" pageId="8" pageNumber="981" pagination="296 - 308" refId="ref45962" refString="Alonso CBG, Zurita GA, Bellocq MI. 2019. Livestock areas with canopy cover sustain dung beetle diversity in the humid subtropical Chaco forest. Insect Conservation and Diversity 12: 296 - 308." type="journal article" year="2019">Alonso et al., 2019: 299</bibRefCitation>
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;
<treatmentCitation author="Cabrera Walsh G" page="761" pageId="8" pageNumber="981" year="2020">
<bibRefCitation author="Cabrera Walsh G" pageId="8" pageNumber="981" pagination="760 - 762" refId="ref46808" refString="Cabrera Walsh G. 2020. Dung beetle (Coleoptera: Scarabaeidae: Scarabaeinae) coloration and foraging time. The Coleopterists Bulletin 74: 760 - 762." type="journal article" year="2020">Cabrera Walsh, 2020: 761</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Gillett CPT &amp; Toussaint EFA" box="[1017,1365,596,618]" page="665" pageId="8" pageNumber="981" year="2020">
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</treatmentCitation>
, 667, 670, 673674, figs 34.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="8.[809,1425,680,734]" pageId="8" pageNumber="981">
<treatmentCitationGroup pageId="8" pageNumber="981">
<taxonomicName authority="HAROLD, 1868" authorityName="Harold des Cupello" authorityYear="1868" box="[809,1024,681,702]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1013" phylum="Arthropoda" rank="species" species="onithoides">
<emphasis box="[809,1024,681,702]" italics="true" pageId="8" pageNumber="981">Bolbites onithoides</emphasis>
</taxonomicName>
[incorrect subsequent spelling]:
<treatmentCitation author="Di Iorio O &amp; Turienzo P" page="21" pageId="8" pageNumber="981" year="2015">
<bibRefCitation author="Di Iorio O &amp; Turienzo P" pageId="8" pageNumber="981" pagination="13 - 42" refId="ref47726" refString="Di Iorio O, Turienzo P. 2015. Campo de Mayo: un area natural historica en los alrededores de la ciudad de Buenos Aires, Argentina, que merece ser declarada reserva natural. Revista del Museo Argentino de Ciencias Naturales, Nueva Serie 17: 13 - 42." type="journal article" year="2015">Di Iorio &amp; Turienzo 2015: 21</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection lastPageId="25" lastPageNumber="998" pageId="8" pageNumber="981" type="materials_examined">
<paragraph blockId="8.[809,1425,766,1279]" pageId="8" pageNumber="981">
<emphasis box="[809,1079,767,788]" italics="true" pageId="8" pageNumber="981">
Type series:
<typeStatus box="[959,1073,767,788]" pageId="8" pageNumber="981">Lectotype</typeStatus>
:
</emphasis>
Male,
<emphasis bold="true" box="[1166,1385,766,788]" pageId="8" pageNumber="981">here designated</emphasis>
(1, Harolds handwriting, bordered in red,
<collectingRegion box="[1285,1424,797,819]" country="Uruguay" name="Montevideo" pageId="8" pageNumber="981">Montevideo</collectingRegion>
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[809,940,828,850]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="981" phylum="Arthropoda" rank="species" species="onitoides">B. onitoides</taxonomicName>
t. Harold; 2, bordered in black, Ex-Musaeo E. Harold; 3, red paper, LECTOTYPE; 4, green paper bordered in black, Muséum Paris ex Coll. R. Oberthür 1952; 5, MCs handwriting, red paper,
<typeStatus box="[1245,1402,920,941]" pageId="8" pageNumber="981">LECTOTYPE</typeStatus>
Ƌ
<taxonomicName authority="Harold des Cupello" authorityName="Harold des Cupello" authorityYear="1868" box="[809,1237,950,972]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="981" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[809,1009,950,971]" italics="true" pageId="8" pageNumber="981">Bolbites onitoides</emphasis>
Harold des Cupello
</taxonomicName>
&amp; Vaz-de-Mello, 2019),
<emphasis bold="true" box="[889,976,981,1002]" pageId="8" pageNumber="981">MNHN</emphasis>
ex Edgar von Harold and R. Oberthür collections.
<emphasis box="[954,1247,1012,1033]" italics="true" pageId="8" pageNumber="981">
Potential
<typeStatus box="[1073,1247,1012,1033]" pageId="8" pageNumber="981">paralectotypes</typeStatus>
</emphasis>
(see Remarks below):
<emphasis bold="true" box="[905,927,1042,1063]" pageId="8" pageNumber="981">1.</emphasis>
male (1, bordered in black, Ex-Musaeo E. Harold; 2, green paper bordered in black, Muséum Paris ex Coll.
<collectionCode box="[971,992,1104,1125]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="981">R</collectionCode>
. Oberthür 1952),
<emphasis bold="true" box="[1203,1291,1104,1125]" pageId="8" pageNumber="981">MNHN</emphasis>
ex Edgar von Harold and R. Oberthür collections.
<emphasis bold="true" box="[1282,1303,1134,1155]" pageId="8" pageNumber="981">2.</emphasis>
female (1, bordered in black, Ex-Musaeo E. Harold; 2, green paper bordered in black, Muséum Paris ex Coll.
<collectionCode box="[809,830,1227,1248]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="981">R</collectionCode>
. Oberthür 1952),
<emphasis bold="true" box="[1031,1118,1226,1247]" pageId="8" pageNumber="981">MNHN</emphasis>
ex Edgar von Harold and R. Oberthür collections.
</paragraph>
<paragraph blockId="8.[809,1218,1320,1342]" box="[809,1218,1320,1342]" pageId="8" pageNumber="981">
<materialsCitation ID-GBIF-Occurrence="3704841550" box="[809,1215,1320,1342]" country="Uruguay" location="Montevideo" pageId="8" pageNumber="981" specimenCount="1" stateProvince="Montevideo" typeStatus="holotype">
<emphasis box="[809,958,1320,1341]" italics="true" pageId="8" pageNumber="981">
<typeStatus box="[809,862,1320,1341]" pageId="8" pageNumber="981">Type</typeStatus>
locality:
</emphasis>
<collectingCountry box="[965,1069,1320,1341]" name="Uruguay" pageId="8" pageNumber="981">Uruguay</collectingCountry>
:
<collectingRegion box="[1079,1215,1320,1342]" country="Uruguay" name="Montevideo" pageId="8" pageNumber="981">Montevideo</collectingRegion>
</materialsCitation>
.
</paragraph>
<paragraph blockId="8.[809,1425,1382,1895]" lastBlockId="9.[163,792,197,711]" lastPageId="9" lastPageNumber="982" pageId="8" pageNumber="981">
<emphasis box="[809,941,1382,1403]" italics="true" pageId="8" pageNumber="981">Etymology:</emphasis>
The specific name
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[1182,1290,1382,1403]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="981" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[1182,1290,1382,1403]" italics="true" pageId="8" pageNumber="981">onitoides</emphasis>
</taxonomicName>
, a noun in the nominative case used in apposition, translates into English as the one like
<taxonomicName authorityName="Fabricius" authorityYear="1798" box="[1195,1273,1444,1465]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="981" phylum="Arthropoda" rank="genus">
<emphasis box="[1195,1273,1444,1465]" italics="true" pageId="8" pageNumber="981">Onitis</emphasis>
</taxonomicName>
and refers to the overall similarity of
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[1120,1253,1474,1496]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="981" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[1120,1253,1474,1496]" italics="true" pageId="8" pageNumber="981">B. onitoides</emphasis>
</taxonomicName>
to members of the genus
<taxonomicName authorityName="Fabricius" authorityYear="1798" box="[927,997,1505,1526]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="981" phylum="Arthropoda" rank="genus">
<emphasis box="[927,997,1505,1526]" italics="true" pageId="8" pageNumber="981">Onitis</emphasis>
</taxonomicName>
. It is formed from the combination of the name
<taxonomicName authorityName="Fabricius" authorityYear="1798" box="[918,987,1536,1557]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="981" phylum="Arthropoda" rank="genus">
<emphasis box="[918,987,1536,1557]" italics="true" pageId="8" pageNumber="981">Onitis</emphasis>
</taxonomicName>
plus the suffix -oides, the latter derived from the combination of the linking vowel -
<emphasis box="[1308,1320,1567,1588]" italics="true" pageId="8" pageNumber="981">o</emphasis>
- and the Ancient Greek noun εἶδος (
<emphasis box="[1115,1172,1597,1618]" italics="true" pageId="8" pageNumber="981">eidos</emphasis>
) for form or likeness (
<bibRefCitation author="Brown RW" box="[817,961,1627,1649]" pageId="8" pageNumber="981" refId="ref46537" refString="Brown RW. 1956. Composition of scientific words. A manual of methods and a lexicon of materials for the practice of logotechnics. Washington, DC: Smithsonian Books, 882." type="book" year="1956">Brown, 1956</bibRefCitation>
;
<bibRefCitation author="Papavero N" box="[975,1149,1628,1649]" pageId="8" pageNumber="981" pagination="151 - 167" refId="ref52964" refString="Papavero N. 1994. Rudimentos de grego. In: Papavero N, ed. Fundamentos praticos de taxonomia zoologica (colecoes, bibliografia, nomenclatura). Sao Paulo: Editora Unesp, 151 - 167." type="book chapter" year="1994">Papavero, 1994</bibRefCitation>
;
<bibRefCitation author="Wiktionary" box="[1162,1357,1628,1650]" pageId="8" pageNumber="981" refId="ref55720" refString="Wiktionary. 2020. Wiktionary. Available at: https: // en. wiktionary. org / wiki / (last accessed on 29 October 2020)." type="url" year="2020">Wiktionary, 2020</bibRefCitation>
). Our interpretation of the name as being a noun instead of as an adjective follows Article 31.2.2 of the Code. According to
<bibRefCitation author="Harold E" box="[836,1004,1719,1741]" pageId="8" pageNumber="981" pagination="979 - 1346" refId="ref49928" refString="Harold E. 1869 a. Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae. Munich: E. H. Gummi, 979 - 1346." type="book chapter" year="1869">Harold (1869a)</bibRefCitation>
, Fabriciuss (1798) genus name
<taxonomicName authorityName="Fabricius" authorityYear="1798" box="[1356,1425,1720,1741]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="981" phylum="Arthropoda" rank="genus">
<emphasis box="[1356,1425,1720,1741]" italics="true" pageId="8" pageNumber="981">Onitis</emphasis>
</taxonomicName>
derives from a Greek word spelled by Harold as ὄνἰς and translated into Latin as
<emphasis box="[1164,1367,1781,1803]" italics="true" pageId="8" pageNumber="981">stercus asininum</emphasis>
(i.e. ass dung).
<bibRefCitation author="Brulle A" box="[934,1086,1811,1834]" pageId="8" pageNumber="981" pagination="225 - 448" refId="ref46659" refString="Brulle A. 1838. [livraison 8: pp. 225 - 448]. In: Brulle A, eds. 1837 - 1838. Histoire naturelle des insectes, traitant de leur organisation et de leurs moeurs en general, par M. V. Audouin, et comprenant leur classification et la description des especes, par M. A. Brulle. Le tout accompagne de planches gravees sur acier, d'apres des peintures executees pour cette edition sur la collection du Museum de Paris. Tome VI. Coleopteres III. Paris: F. D. Pillot, 448 + pls 18, 19, 19 bis, 20, 25 [publication date and authorship based on Bousquet 2016]." type="journal article" year="1838">Brullé (1838)</bibRefCitation>
, in contrast, states the name is derived from the Greek word ὀνῖτις (
<emphasis box="[1268,1333,1843,1864]" italics="true" pageId="8" pageNumber="981">onitis</emphasis>
), which he said refers to an unspecified plant name. Greek dictionaries and lexica indeed list ὀνῖτις as a feminine noun and say it refers to acrid herbs such as marjoram,
<emphasis box="[163,287,260,281]" italics="true" pageId="9" pageNumber="982">Origanum</emphasis>
L. (
<bibRefCitation author="Liddell HG &amp; Scott R" box="[339,601,260,282]" pageId="9" pageNumber="982" refId="ref51155" refString="Liddell HG, Scott R. 1897. A Greek-English lexicon. Revised and augmented throughout with the cooperation of professor Drisler, of Columbia College, New York, 8 th edn. New York: Harper &amp; Brothers." type="book" year="1897">Liddell &amp; Scott, 1897</bibRefCitation>
;
<bibRefCitation author="Brown RW" box="[619,771,259,281]" pageId="9" pageNumber="982" refId="ref46537" refString="Brown RW. 1956. Composition of scientific words. A manual of methods and a lexicon of materials for the practice of logotechnics. Washington, DC: Smithsonian Books, 882." type="book" year="1956">Brown, 1956</bibRefCitation>
;
<bibRefCitation author="Wiktionary" box="[163,359,290,312]" pageId="9" pageNumber="982" refId="ref55720" refString="Wiktionary. 2020. Wiktionary. Available at: https: // en. wiktionary. org / wiki / (last accessed on 29 October 2020)." type="url" year="2020">Wiktionary, 2020</bibRefCitation>
). The word suggested by Harold, on the other hand, is not found with that spelling in Greek dictionaries, but the similar form ὄνος (
<emphasis box="[604,654,352,373]" italics="true" pageId="9" pageNumber="982">ónos</emphasis>
) is present and indeed translates as ass or donkey (
<bibRefCitation author="Liddell HG &amp; Scott R" pageId="9" pageNumber="982" refId="ref51155" refString="Liddell HG, Scott R. 1897. A Greek-English lexicon. Revised and augmented throughout with the cooperation of professor Drisler, of Columbia College, New York, 8 th edn. New York: Harper &amp; Brothers." type="book" year="1897">Liddell &amp; Scott, 1897</bibRefCitation>
;
<bibRefCitation author="Wiktionary" box="[296,486,413,435]" pageId="9" pageNumber="982" refId="ref55720" refString="Wiktionary. 2020. Wiktionary. Available at: https: // en. wiktionary. org / wiki / (last accessed on 29 October 2020)." type="url" year="2020">Wiktionary, 2020</bibRefCitation>
). If Harold is correct, then it is possible that
<bibRefCitation author="Fabricius JC" box="[366,550,444,466]" pageId="9" pageNumber="982" refId="ref48568" refString="Fabricius JC. 1798. Supplementum entomologiae systematicae. Copenhagen: Proft &amp; Storch." type="book" year="1798">Fabricius (1798)</bibRefCitation>
derived
<taxonomicName authorityName="Fabricius" authorityYear="1798" box="[648,718,444,465]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="982" phylum="Arthropoda" rank="genus">
<emphasis box="[648,718,444,465]" italics="true" pageId="9" pageNumber="982">Onitis</emphasis>
</taxonomicName>
from the combination of the stem of ὄνος (viz., ὄν–,
<emphasis box="[681,708,475,496]" italics="true" pageId="9" pageNumber="982">on</emphasis>
-) plus the suffix -ῖτις (-
<emphasis box="[352,389,505,526]" italics="true" pageId="9" pageNumber="982">itis</emphasis>
), the feminine equivalent of -ῑ́της (
<emphasis box="[171,220,536,557]" italics="true" pageId="9" pageNumber="982">-ites</emphasis>
). Under either Harolds or Brullés suggestion,
<taxonomicName authorityName="Fabricius" authorityYear="1798" box="[163,235,567,588]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="982" phylum="Arthropoda" rank="genus">
<emphasis box="[163,235,567,588]" italics="true" pageId="9" pageNumber="982">Onitis</emphasis>
</taxonomicName>
would be feminine in gender, which disagrees with the way
<bibRefCitation author="Fabricius JC" box="[320,498,597,619]" pageId="9" pageNumber="982" refId="ref48568" refString="Fabricius JC. 1798. Supplementum entomologiae systematicae. Copenhagen: Proft &amp; Storch." type="book" year="1798">Fabricius (1798</bibRefCitation>
,
<bibRefCitation author="Fabricius JC" box="[511,566,597,618]" pageId="9" pageNumber="982" refId="ref48583" refString="Fabricius JC. 1801. Systema eleutheratorum, secundum ordines, genera, species, adjectis synonymis, locis, observationibus, descriptionibus. Tomus I. Kiel: Bibliopolii Academici Novi [publication date following Bousquet 2016]." type="journal volume" year="1801">1801</bibRefCitation>
) and (as far as we know) all other authors have ever treated the name, i.e. as masculine, including equivalent names in botanical nomenclature (
<bibRefCitation author="IPNI" box="[331,453,689,710]" pageId="9" pageNumber="982" refId="ref50491" refString="IPNI. 2021. International Plant Name Index. Available at www. ipni. org." type="book" year="2021">IPNI, 2021</bibRefCitation>
).
</paragraph>
<paragraph blockId="9.[163,780,751,1111]" pageId="9" pageNumber="982">
<emphasis box="[163,492,751,773]" italics="true" pageId="9" pageNumber="982">
Diagnosis:
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[290,492,751,772]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="982" phylum="Arthropoda" rank="species" species="onitoides">Bolbites onitoides</taxonomicName>
</emphasis>
is readily identified as a
<taxonomicName box="[163,284,782,804]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1032" phylum="Arthropoda" rank="tribe" tribe="Phanaeini">Phanaeini</taxonomicName>
by the presence of iridescent colouration, eyes completely divided by the canthus into a dorsal and a ventral portion, a long metanepisternal tab (
<figureCitation box="[172,262,874,896]" captionStart="Figure 5" captionStartId="12.[145,223,1538,1560]" captionTargetBox="[147,1408,197,1497]" captionTargetId="figure-25@12.[145,1428,195,1498]" captionTargetPageId="12" captionText="Figure 5. Legs. AE, sexual dimorphism on the front legs of B. onitoides. A, B, profemora: A, male; B, female. Note that the male profemur is claviform and has a strong emargination on its posterior edge (indicated by the red arrow), while that of the female is shorter and has a simply rounded aspect. CE, protibiae: C, D, dorsal view of male (C) and female (D). Observe that the male protibia is much longer and slenderer than that of the female and has a well-delimited tuft of long setae at its internal apical angle. E, lateral view of the male protibia. In contrast to females, males have a series of sharp, uneven teeth along the ventral protibial carina. Red arrow indicates the central tooth, which is longer than the others. F, G, meso- and metatarsi of B. onitoides. Unlike all other Phanaeini, which are clawless, B. onitoides has a pair of well-developed claws on the apical meso- and metatarsomeres. A small prolongation raises from underneath the claws and project itself anteriorly (indicated by the red arrow in F); this prolongation seems to be homologous to the tapered apex of the apical meso- and metatarsomeres of the other Phanaeini and is probably a synapomorphy of the clade. H, metanepisternum and its long tab typical of the Phanaeina." figureDoi="http://doi.org/10.5281/zenodo.7551998" httpUri="https://zenodo.org/record/7551998/files/figure.png" pageId="9" pageNumber="982">Fig. 5H</figureCitation>
), and its metaventrite with an acuminate anteromedian prominence flanked by long setae (
<figureCitation captionStart="Figure 4" captionStartId="11.[163,245,1393,1415]" captionTargetBox="[166,1447,198,1350]" captionTargetId="figure-147@11.[163,1450,195,1353]" captionTargetPageId="11" captionText="Figure 4. Meso- and metaventrite in Bolbites and Gromphadina. A, Bolbites onitoides. B, Gromphas inermis. C, D, Oruscatus davus: C, male; D, female. The carina of the mesofurcal brace constitutes the posterior branch of the mesoventral ridge and gives it the shape of a cross (orange arrow). This cross-shaped ridge is typical of the Gromphadina and is also present in at least some Phanaeina. Note that, while in the Gromphadina the posterior branch connects the mesoventral ridge to the anterior edge of the metaventrite, no such connection exists in most Bolbites and and the other Phanaeina whose mesoventral ridge is T-shaped. In Oruscatus, although present in both sexes, the posterior branch of the mesoventral ridge is much more clearly delimited in males than in females; in females, the posterior branch is usually incomplete and interrupted, while in males it is always sharp and continuous. Note also other differences between Bolbites and the Gromphadina relating to the shape of the metaventral anterior prominence and its integument and lateral pilosity." figureDoi="http://doi.org/10.5281/zenodo.7551994" httpUri="https://zenodo.org/record/7551994/files/figure.png" pageId="9" pageNumber="982">Fig. 4A</figureCitation>
). In turn, it differs from all other phanaeines by having meso- and metatarsi with claws and from other Phanaeina by its antennae with free apical lamellae. Its unusual lack of dorsal sexual dimorphism while having a rich ventral one is also an unique condition among the Phanaeina.
</paragraph>
<paragraph blockId="9.[163,779,1146,1536]" pageId="9" pageNumber="982">
<emphasis box="[163,534,1146,1168]" italics="true" pageId="9" pageNumber="982">Redescription: Colouration:</emphasis>
Clypeus sexually dimorphic (see below). Frons, paraocular areas, pronotum and elytra mainly metallic light to dark blue in western populations (
<figureCitation box="[501,619,1238,1260]" captionStart="Figure 1" captionStartId="2.[145,223,1852,1874]" captionTargetBox="[217,1343,203,1809]" captionTargetId="figure-25@2.[207,1362,195,1812]" captionTargetPageId="2" captionText="Figure 1. The wide colour variation shown by Bolbites onitoides. A, copper (male). B, red (male). C, Copper individual with greenish pronotum and elytra (female). D, atypical bicoloured individual with unusually green elytra (female). E, F, typical bicoloured individuals (males). GI, blue individuals (males). Although Martínez (1959, 1987) thought that two subspecies could be recognized one for red individuals and the other for blue ones two facts observed by us do not support this idea.First, two kinds of intermediates exist connecting the two main colour patterns of red (B) and blue individuals (GI): specimens with a red pronotum and blue or green elytra (DF), which we call the discrete bicoloured intermediates, and the second being the gradual colour-blended intermediates, or specimens that lie at the centre of the variation spectrum between the two colour ends, which includes specimens showing dorsal colouration entirely coppery (A), coppery with greenish reflections (C), and coppery with reddish reflections. Secondly, no other features were found to vary geographically along with colour. Therefore, we interpret the plethora of colour variants as intraspecific variation of a single monotypic species." figureDoi="http://doi.org/10.5281/zenodo.7551987" httpUri="https://zenodo.org/record/7551987/files/figure.png" pageId="9" pageNumber="982">Fig. 1G, H</figureCitation>
) and metallic red in eastern populations (
<figureCitation box="[480,562,1269,1291]" captionStart="Figure 1" captionStartId="2.[145,223,1852,1874]" captionTargetBox="[217,1343,203,1809]" captionTargetId="figure-25@2.[207,1362,195,1812]" captionTargetPageId="2" captionText="Figure 1. The wide colour variation shown by Bolbites onitoides. A, copper (male). B, red (male). C, Copper individual with greenish pronotum and elytra (female). D, atypical bicoloured individual with unusually green elytra (female). E, F, typical bicoloured individuals (males). GI, blue individuals (males). Although Martínez (1959, 1987) thought that two subspecies could be recognized one for red individuals and the other for blue ones two facts observed by us do not support this idea.First, two kinds of intermediates exist connecting the two main colour patterns of red (B) and blue individuals (GI): specimens with a red pronotum and blue or green elytra (DF), which we call the discrete bicoloured intermediates, and the second being the gradual colour-blended intermediates, or specimens that lie at the centre of the variation spectrum between the two colour ends, which includes specimens showing dorsal colouration entirely coppery (A), coppery with greenish reflections (C), and coppery with reddish reflections. Secondly, no other features were found to vary geographically along with colour. Therefore, we interpret the plethora of colour variants as intraspecific variation of a single monotypic species." figureDoi="http://doi.org/10.5281/zenodo.7551987" httpUri="https://zenodo.org/record/7551987/files/figure.png" pageId="9" pageNumber="982">Fig. 1B</figureCitation>
), both phases with different degrees of greenish reflections over dorsum; throughout the range, specimens sometimes with metallic red head and pronotum and metallic blue or green elytra (
<figureCitation box="[323,439,1391,1413]" captionStart="Figure 1" captionStartId="2.[145,223,1852,1874]" captionTargetBox="[217,1343,203,1809]" captionTargetId="figure-25@2.[207,1362,195,1812]" captionTargetPageId="2" captionText="Figure 1. The wide colour variation shown by Bolbites onitoides. A, copper (male). B, red (male). C, Copper individual with greenish pronotum and elytra (female). D, atypical bicoloured individual with unusually green elytra (female). E, F, typical bicoloured individuals (males). GI, blue individuals (males). Although Martínez (1959, 1987) thought that two subspecies could be recognized one for red individuals and the other for blue ones two facts observed by us do not support this idea.First, two kinds of intermediates exist connecting the two main colour patterns of red (B) and blue individuals (GI): specimens with a red pronotum and blue or green elytra (DF), which we call the discrete bicoloured intermediates, and the second being the gradual colour-blended intermediates, or specimens that lie at the centre of the variation spectrum between the two colour ends, which includes specimens showing dorsal colouration entirely coppery (A), coppery with greenish reflections (C), and coppery with reddish reflections. Secondly, no other features were found to vary geographically along with colour. Therefore, we interpret the plethora of colour variants as intraspecific variation of a single monotypic species." figureDoi="http://doi.org/10.5281/zenodo.7551987" httpUri="https://zenodo.org/record/7551987/files/figure.png" pageId="9" pageNumber="982">Fig. 1DF</figureCitation>
) or dorsum entirely metallic coppery (
<figureCitation box="[268,350,1422,1444]" captionStart="Figure 1" captionStartId="2.[145,223,1852,1874]" captionTargetBox="[217,1343,203,1809]" captionTargetId="figure-25@2.[207,1362,195,1812]" captionTargetPageId="2" captionText="Figure 1. The wide colour variation shown by Bolbites onitoides. A, copper (male). B, red (male). C, Copper individual with greenish pronotum and elytra (female). D, atypical bicoloured individual with unusually green elytra (female). E, F, typical bicoloured individuals (males). GI, blue individuals (males). Although Martínez (1959, 1987) thought that two subspecies could be recognized one for red individuals and the other for blue ones two facts observed by us do not support this idea.First, two kinds of intermediates exist connecting the two main colour patterns of red (B) and blue individuals (GI): specimens with a red pronotum and blue or green elytra (DF), which we call the discrete bicoloured intermediates, and the second being the gradual colour-blended intermediates, or specimens that lie at the centre of the variation spectrum between the two colour ends, which includes specimens showing dorsal colouration entirely coppery (A), coppery with greenish reflections (C), and coppery with reddish reflections. Secondly, no other features were found to vary geographically along with colour. Therefore, we interpret the plethora of colour variants as intraspecific variation of a single monotypic species." figureDoi="http://doi.org/10.5281/zenodo.7551987" httpUri="https://zenodo.org/record/7551987/files/figure.png" pageId="9" pageNumber="982">Fig. 1A</figureCitation>
). Venter, including ventral surface of legs, and pygidium always ranging from dark blue to light green, never red or copper (
<figureCitation box="[543,656,1483,1505]" captionStart="Figure 2" captionStartId="4.[145,223,1515,1537]" captionTargetBox="[274,1291,204,1473]" captionTargetId="figure-160@4.[271,1298,195,1475]" captionTargetPageId="4" captionText="Figure 2. Ventral sexual dimorphism in Bolbites onitoides. A, lateral view of a male. B, C, ventral view: B, male: C, female. Note the conspicuous ventral sexual dimorphism in this species, including the presence of several teeth and spines all over the body of the male, the differences in the overall shape of the protibiae, the pilosity of the metacoxae, and colour of the metaventrite." figureDoi="http://doi.org/10.5281/zenodo.7551989" httpUri="https://zenodo.org/record/7551989/files/figure.png" pageId="9" pageNumber="982">Fig. 2B, C</figureCitation>
); centre of metaventrite sexually dimorphic (see below).
</paragraph>
<paragraph blockId="9.[163,779,1566,1895]" lastBlockId="9.[827,1443,197,433]" pageId="9" pageNumber="982">
<emphasis box="[163,229,1566,1587]" italics="true" pageId="9" pageNumber="982">Head:</emphasis>
Clypeus at anterior edge with two small, rounded teeth separated by a broad U-shaped emargination; anterior edge only slightly bent upwards; integument entirely covered by net of transverse ridges; clypeal process a rounded transverse carina. Junction of clypeus and paraocular areas deeply notched on outer edge. Paraocular areas and frons with integument similar to clypeus, but ridges less well-formed and, posteriorly, sometimes becoming individual granules. Cephalic process raised between clypeus and frons into a straight transverse carina (
<figureCitation box="[387,454,1873,1895]" captionStart="Figure 3" captionStartId="10.[145,223,1255,1277]" captionTargetBox="[209,1358,197,1213]" captionTargetId="figure-230@10.[207,1362,195,1215]" captionTargetPageId="10" captionText="Figure 3. Dorsal sexual monomorphism in Bolbites onitoides. A, frontal view of a male. B, C, head and pronotum: B, male; C, female. Note how little the sexes differ dorsally. Unlike most groups of Phanaeina, males of B. onitoides lack any cephalic or pronotal horns and have a pronotal ridge identical to that of the females. This dorsal monomorphism is in striking contrast with the sexual dimorphism found on the venter. Note, however, the sexually dimorphic extension of iridescence over the clypeus, males having at least its basal half metallic, while females are entirely (or almost entirely) dull black." figureDoi="http://doi.org/10.5281/zenodo.7551991" httpUri="https://zenodo.org/record/7551991/files/figure.png" pageId="9" pageNumber="982">Fig. 3</figureCitation>
); carina usually with three (smaller specimens) or five (larger specimens) tenuous elevations, the central one higher than the lateral ones, elevations never acute or with aspect of tubercles or teeth such as those of many
<emphasis box="[1135,1346,289,311]" italics="true" pageId="9" pageNumber="982">
<taxonomicName authorityName="d'Olsoufieff" authorityYear="1924" box="[1135,1309,289,310]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="982" phylum="Arthropoda" rank="genus" sensu="stricto">Coprophanaeus</taxonomicName>
<taxonomicNameLabel box="[1314,1346,290,311]" pageId="9" pageNumber="982" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
(in worn specimens, carina truncate, without elevations). Eyes completely divided into dorsal and ventral portions by prolongation of canthus; dorsal portion slightly smaller than ventral portion.
</paragraph>
<paragraph blockId="9.[827,1444,474,1385]" pageId="9" pageNumber="982">
<emphasis box="[827,914,474,495]" italics="true" pageId="9" pageNumber="982">Thorax:</emphasis>
Pronotum on anterior region with transverse C-shaped ridge flanked by shallow depressions (
<figureCitation box="[834,894,535,557]" captionStart="Figure 3" captionStartId="10.[145,223,1255,1277]" captionTargetBox="[209,1358,197,1213]" captionTargetId="figure-230@10.[207,1362,195,1215]" captionTargetPageId="10" captionText="Figure 3. Dorsal sexual monomorphism in Bolbites onitoides. A, frontal view of a male. B, C, head and pronotum: B, male; C, female. Note how little the sexes differ dorsally. Unlike most groups of Phanaeina, males of B. onitoides lack any cephalic or pronotal horns and have a pronotal ridge identical to that of the females. This dorsal monomorphism is in striking contrast with the sexual dimorphism found on the venter. Note, however, the sexually dimorphic extension of iridescence over the clypeus, males having at least its basal half metallic, while females are entirely (or almost entirely) dull black." figureDoi="http://doi.org/10.5281/zenodo.7551991" httpUri="https://zenodo.org/record/7551991/files/figure.png" pageId="9" pageNumber="982">Fig. 3</figureCitation>
); pronotal ridge at most as wide as cephalic carina (
<figureCitation box="[835,900,566,588]" captionStart="Figure 3" captionStartId="10.[145,223,1255,1277]" captionTargetBox="[209,1358,197,1213]" captionTargetId="figure-230@10.[207,1362,195,1215]" captionTargetPageId="10" captionText="Figure 3. Dorsal sexual monomorphism in Bolbites onitoides. A, frontal view of a male. B, C, head and pronotum: B, male; C, female. Note how little the sexes differ dorsally. Unlike most groups of Phanaeina, males of B. onitoides lack any cephalic or pronotal horns and have a pronotal ridge identical to that of the females. This dorsal monomorphism is in striking contrast with the sexual dimorphism found on the venter. Note, however, the sexually dimorphic extension of iridescence over the clypeus, males having at least its basal half metallic, while females are entirely (or almost entirely) dull black." figureDoi="http://doi.org/10.5281/zenodo.7551991" httpUri="https://zenodo.org/record/7551991/files/figure.png" pageId="9" pageNumber="982">Fig. 3</figureCitation>
); without horns or any other ornamentation in both sexes (
<figureCitation box="[969,1060,597,619]" captionStart="Figure 2" captionStartId="4.[145,223,1515,1537]" captionTargetBox="[274,1291,204,1473]" captionTargetId="figure-160@4.[271,1298,195,1475]" captionTargetPageId="4" captionText="Figure 2. Ventral sexual dimorphism in Bolbites onitoides. A, lateral view of a male. B, C, ventral view: B, male: C, female. Note the conspicuous ventral sexual dimorphism in this species, including the presence of several teeth and spines all over the body of the male, the differences in the overall shape of the protibiae, the pilosity of the metacoxae, and colour of the metaventrite." figureDoi="http://doi.org/10.5281/zenodo.7551989" httpUri="https://zenodo.org/record/7551989/files/figure.png" pageId="9" pageNumber="982">Figs 2A</figureCitation>
,
<figureCitation box="[1077,1091,597,618]" captionStart="Figure 3" captionStartId="10.[145,223,1255,1277]" captionTargetBox="[209,1358,197,1213]" captionTargetId="figure-230@10.[207,1362,195,1215]" captionTargetPageId="10" captionText="Figure 3. Dorsal sexual monomorphism in Bolbites onitoides. A, frontal view of a male. B, C, head and pronotum: B, male; C, female. Note how little the sexes differ dorsally. Unlike most groups of Phanaeina, males of B. onitoides lack any cephalic or pronotal horns and have a pronotal ridge identical to that of the females. This dorsal monomorphism is in striking contrast with the sexual dimorphism found on the venter. Note, however, the sexually dimorphic extension of iridescence over the clypeus, males having at least its basal half metallic, while females are entirely (or almost entirely) dull black." figureDoi="http://doi.org/10.5281/zenodo.7551991" httpUri="https://zenodo.org/record/7551991/files/figure.png" pageId="9" pageNumber="982">3</figureCitation>
); integument almost entirely sculptured, postero-central region usually with small rounded impressions that become progressively more confluent towards anterior region, giving a rugose aspect to integument; anterolateral angles usually covered by shiny scale-like granules; lateral margin irregularly serrate, with sexually dimorphic presence of posterior tab; anterior margin not interrupted behind eyes; posterior margin complete; lateral fossae deep, broad, well developed, smooth and shiny centrally; with two deep, well-developed, elongate posterior fossae. Hypomera, mesepimera, metanepisterna and outer lobes of metaventrite covered by abundant long, black setae. Metanepisterna with long dorsal tab covering part of epipleura, which is evidently depressed at this point (
<figureCitation box="[902,988,1057,1079]" captionStart="Figure 5" captionStartId="12.[145,223,1538,1560]" captionTargetBox="[147,1408,197,1497]" captionTargetId="figure-25@12.[145,1428,195,1498]" captionTargetPageId="12" captionText="Figure 5. Legs. AE, sexual dimorphism on the front legs of B. onitoides. A, B, profemora: A, male; B, female. Note that the male profemur is claviform and has a strong emargination on its posterior edge (indicated by the red arrow), while that of the female is shorter and has a simply rounded aspect. CE, protibiae: C, D, dorsal view of male (C) and female (D). Observe that the male protibia is much longer and slenderer than that of the female and has a well-delimited tuft of long setae at its internal apical angle. E, lateral view of the male protibia. In contrast to females, males have a series of sharp, uneven teeth along the ventral protibial carina. Red arrow indicates the central tooth, which is longer than the others. F, G, meso- and metatarsi of B. onitoides. Unlike all other Phanaeini, which are clawless, B. onitoides has a pair of well-developed claws on the apical meso- and metatarsomeres. A small prolongation raises from underneath the claws and project itself anteriorly (indicated by the red arrow in F); this prolongation seems to be homologous to the tapered apex of the apical meso- and metatarsomeres of the other Phanaeini and is probably a synapomorphy of the clade. H, metanepisternum and its long tab typical of the Phanaeina." figureDoi="http://doi.org/10.5281/zenodo.7551998" httpUri="https://zenodo.org/record/7551998/files/figure.png" pageId="9" pageNumber="982">Fig. 5H</figureCitation>
). Mesoventrite constricted at centre by anterior projection of central lobe of metaventrite (
<figureCitation captionStart="Figure 4" captionStartId="11.[163,245,1393,1415]" captionTargetBox="[166,1447,198,1350]" captionTargetId="figure-147@11.[163,1450,195,1353]" captionTargetPageId="11" captionText="Figure 4. Meso- and metaventrite in Bolbites and Gromphadina. A, Bolbites onitoides. B, Gromphas inermis. C, D, Oruscatus davus: C, male; D, female. The carina of the mesofurcal brace constitutes the posterior branch of the mesoventral ridge and gives it the shape of a cross (orange arrow). This cross-shaped ridge is typical of the Gromphadina and is also present in at least some Phanaeina. Note that, while in the Gromphadina the posterior branch connects the mesoventral ridge to the anterior edge of the metaventrite, no such connection exists in most Bolbites and and the other Phanaeina whose mesoventral ridge is T-shaped. In Oruscatus, although present in both sexes, the posterior branch of the mesoventral ridge is much more clearly delimited in males than in females; in females, the posterior branch is usually incomplete and interrupted, while in males it is always sharp and continuous. Note also other differences between Bolbites and the Gromphadina relating to the shape of the metaventral anterior prominence and its integument and lateral pilosity." figureDoi="http://doi.org/10.5281/zenodo.7551994" httpUri="https://zenodo.org/record/7551994/files/figure.png" pageId="9" pageNumber="982">Fig. 4A</figureCitation>
). Metaventrite with well-developed anteromedian prominence with fairly acute apex and flanked by long and dense black setation, laterals without granules typical of
<taxonomicName authorityName="Brulle" authorityYear="1838" box="[855,974,1210,1231]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="982" phylum="Arthropoda" rank="genus">
<emphasis box="[855,974,1210,1231]" italics="true" pageId="9" pageNumber="982">Gromphas</emphasis>
</taxonomicName>
(
<figureCitation box="[991,1073,1210,1232]" captionStart="Figure 4" captionStartId="11.[163,245,1393,1415]" captionTargetBox="[166,1447,198,1350]" captionTargetId="figure-147@11.[163,1450,195,1353]" captionTargetPageId="11" captionText="Figure 4. Meso- and metaventrite in Bolbites and Gromphadina. A, Bolbites onitoides. B, Gromphas inermis. C, D, Oruscatus davus: C, male; D, female. The carina of the mesofurcal brace constitutes the posterior branch of the mesoventral ridge and gives it the shape of a cross (orange arrow). This cross-shaped ridge is typical of the Gromphadina and is also present in at least some Phanaeina. Note that, while in the Gromphadina the posterior branch connects the mesoventral ridge to the anterior edge of the metaventrite, no such connection exists in most Bolbites and and the other Phanaeina whose mesoventral ridge is T-shaped. In Oruscatus, although present in both sexes, the posterior branch of the mesoventral ridge is much more clearly delimited in males than in females; in females, the posterior branch is usually incomplete and interrupted, while in males it is always sharp and continuous. Note also other differences between Bolbites and the Gromphadina relating to the shape of the metaventral anterior prominence and its integument and lateral pilosity." figureDoi="http://doi.org/10.5281/zenodo.7551994" httpUri="https://zenodo.org/record/7551994/files/figure.png" pageId="9" pageNumber="982">Fig. 4A</figureCitation>
); in lateral view, anteromedian prominence vertical anteriorly, not projected, at most only slighly indicated apically (
<figureCitation box="[1108,1185,1271,1293]" captionStart="Figure 2" captionStartId="4.[145,223,1515,1537]" captionTargetBox="[274,1291,204,1473]" captionTargetId="figure-160@4.[271,1298,195,1475]" captionTargetPageId="4" captionText="Figure 2. Ventral sexual dimorphism in Bolbites onitoides. A, lateral view of a male. B, C, ventral view: B, male: C, female. Note the conspicuous ventral sexual dimorphism in this species, including the presence of several teeth and spines all over the body of the male, the differences in the overall shape of the protibiae, the pilosity of the metacoxae, and colour of the metaventrite." figureDoi="http://doi.org/10.5281/zenodo.7551989" httpUri="https://zenodo.org/record/7551989/files/figure.png" pageId="9" pageNumber="982">Fig. 2A</figureCitation>
); centre of metaventrite glabrous and covered by sexually dimorphic ill-delimited punctation, never smooth, some specimens with tenuous, ill-delimited mid-longitudinal groove.
</paragraph>
<paragraph blockId="9.[827,1444,1421,1903]" lastBlockId="10.[145,760,1456,1540]" lastPageId="10" lastPageNumber="983" pageId="9" pageNumber="982">
<emphasis box="[827,887,1421,1442]" italics="true" pageId="9" pageNumber="982">Legs:</emphasis>
Shape of profemora, protibiae, protibial spur, protibial ventral carina, metacoxae, metafemora and metatibiae sexually dimorphic (see below). All femora with long, black setae (
<figureCitation box="[1172,1288,1512,1535]" captionStart="Figure 5" captionStartId="12.[145,223,1538,1560]" captionTargetBox="[147,1408,197,1497]" captionTargetId="figure-25@12.[145,1428,195,1498]" captionTargetPageId="12" captionText="Figure 5. Legs. AE, sexual dimorphism on the front legs of B. onitoides. A, B, profemora: A, male; B, female. Note that the male profemur is claviform and has a strong emargination on its posterior edge (indicated by the red arrow), while that of the female is shorter and has a simply rounded aspect. CE, protibiae: C, D, dorsal view of male (C) and female (D). Observe that the male protibia is much longer and slenderer than that of the female and has a well-delimited tuft of long setae at its internal apical angle. E, lateral view of the male protibia. In contrast to females, males have a series of sharp, uneven teeth along the ventral protibial carina. Red arrow indicates the central tooth, which is longer than the others. F, G, meso- and metatarsi of B. onitoides. Unlike all other Phanaeini, which are clawless, B. onitoides has a pair of well-developed claws on the apical meso- and metatarsomeres. A small prolongation raises from underneath the claws and project itself anteriorly (indicated by the red arrow in F); this prolongation seems to be homologous to the tapered apex of the apical meso- and metatarsomeres of the other Phanaeini and is probably a synapomorphy of the clade. H, metanepisternum and its long tab typical of the Phanaeina." figureDoi="http://doi.org/10.5281/zenodo.7551998" httpUri="https://zenodo.org/record/7551998/files/figure.png" pageId="9" pageNumber="982">Figs 5A, B</figureCitation>
,
<figureCitation box="[1300,1362,1513,1535]" captionStart="Figure 6" captionStartId="13.[164,244,1206,1228]" captionTargetBox="[165,1446,197,1164]" captionTargetId="figure-317@13.[163,1446,195,1166]" captionTargetPageId="13" captionText="Figure 6. Sexual dimorphism on the hind legs of B. onitoides. A, B, metacoxae: A, male; B, female. Note the plate-like projections and the long spines directed posteriorly in the male and which are lacking in the female. C, D, metafemora and metatibiae: C, male; D, female. Observe the ridge on the posterior edge of the metafemora and the sharp ridge on the external and internal edges of the metatibiae of the males, all of which are absent in females." figureDoi="http://doi.org/10.5281/zenodo.7552000" httpUri="https://zenodo.org/record/7552000/files/figure.png" pageId="9" pageNumber="982">6C, D</figureCitation>
); setae denser on profemora than on meso- and metafemora. Profemora with small acute tubercle on ventral surface (
<figureCitation box="[835,944,1604,1627]" captionStart="Figure 5" captionStartId="12.[145,223,1538,1560]" captionTargetBox="[147,1408,197,1497]" captionTargetId="figure-25@12.[145,1428,195,1498]" captionTargetPageId="12" captionText="Figure 5. Legs. AE, sexual dimorphism on the front legs of B. onitoides. A, B, profemora: A, male; B, female. Note that the male profemur is claviform and has a strong emargination on its posterior edge (indicated by the red arrow), while that of the female is shorter and has a simply rounded aspect. CE, protibiae: C, D, dorsal view of male (C) and female (D). Observe that the male protibia is much longer and slenderer than that of the female and has a well-delimited tuft of long setae at its internal apical angle. E, lateral view of the male protibia. In contrast to females, males have a series of sharp, uneven teeth along the ventral protibial carina. Red arrow indicates the central tooth, which is longer than the others. F, G, meso- and metatarsi of B. onitoides. Unlike all other Phanaeini, which are clawless, B. onitoides has a pair of well-developed claws on the apical meso- and metatarsomeres. A small prolongation raises from underneath the claws and project itself anteriorly (indicated by the red arrow in F); this prolongation seems to be homologous to the tapered apex of the apical meso- and metatarsomeres of the other Phanaeini and is probably a synapomorphy of the clade. H, metanepisternum and its long tab typical of the Phanaeina." figureDoi="http://doi.org/10.5281/zenodo.7551998" httpUri="https://zenodo.org/record/7551998/files/figure.png" pageId="9" pageNumber="982">Fig. 5A, B</figureCitation>
). Protibiae with four external teeth (
<figureCitation captionStart="Figure 5" captionStartId="12.[145,223,1538,1560]" captionTargetBox="[147,1408,197,1497]" captionTargetId="figure-25@12.[145,1428,195,1498]" captionTargetPageId="12" captionText="Figure 5. Legs. AE, sexual dimorphism on the front legs of B. onitoides. A, B, profemora: A, male; B, female. Note that the male profemur is claviform and has a strong emargination on its posterior edge (indicated by the red arrow), while that of the female is shorter and has a simply rounded aspect. CE, protibiae: C, D, dorsal view of male (C) and female (D). Observe that the male protibia is much longer and slenderer than that of the female and has a well-delimited tuft of long setae at its internal apical angle. E, lateral view of the male protibia. In contrast to females, males have a series of sharp, uneven teeth along the ventral protibial carina. Red arrow indicates the central tooth, which is longer than the others. F, G, meso- and metatarsi of B. onitoides. Unlike all other Phanaeini, which are clawless, B. onitoides has a pair of well-developed claws on the apical meso- and metatarsomeres. A small prolongation raises from underneath the claws and project itself anteriorly (indicated by the red arrow in F); this prolongation seems to be homologous to the tapered apex of the apical meso- and metatarsomeres of the other Phanaeini and is probably a synapomorphy of the clade. H, metanepisternum and its long tab typical of the Phanaeina." figureDoi="http://doi.org/10.5281/zenodo.7551998" httpUri="https://zenodo.org/record/7551998/files/figure.png" pageId="9" pageNumber="982">Fig. 5C, D</figureCitation>
); ventral surface covered by series of oblique ridges. Protarsi absent in both sexes. Meso- and metatarsi with five tarsomeres and short, curved tarsal claws (
<figureCitation box="[835,948,1727,1749]" captionStart="Figure 5" captionStartId="12.[145,223,1538,1560]" captionTargetBox="[147,1408,197,1497]" captionTargetId="figure-25@12.[145,1428,195,1498]" captionTargetPageId="12" captionText="Figure 5. Legs. AE, sexual dimorphism on the front legs of B. onitoides. A, B, profemora: A, male; B, female. Note that the male profemur is claviform and has a strong emargination on its posterior edge (indicated by the red arrow), while that of the female is shorter and has a simply rounded aspect. CE, protibiae: C, D, dorsal view of male (C) and female (D). Observe that the male protibia is much longer and slenderer than that of the female and has a well-delimited tuft of long setae at its internal apical angle. E, lateral view of the male protibia. In contrast to females, males have a series of sharp, uneven teeth along the ventral protibial carina. Red arrow indicates the central tooth, which is longer than the others. F, G, meso- and metatarsi of B. onitoides. Unlike all other Phanaeini, which are clawless, B. onitoides has a pair of well-developed claws on the apical meso- and metatarsomeres. A small prolongation raises from underneath the claws and project itself anteriorly (indicated by the red arrow in F); this prolongation seems to be homologous to the tapered apex of the apical meso- and metatarsomeres of the other Phanaeini and is probably a synapomorphy of the clade. H, metanepisternum and its long tab typical of the Phanaeina." figureDoi="http://doi.org/10.5281/zenodo.7551998" httpUri="https://zenodo.org/record/7551998/files/figure.png" pageId="9" pageNumber="982">Fig. 5F, G</figureCitation>
); tarsomere I to IV progressively smaller; tarsomere V (apical) longer than IV and with strong apical projection rising from beneath tarsal claws (
<figureCitation box="[835,917,1819,1841]" captionStart="Figure 5" captionStartId="12.[145,223,1538,1560]" captionTargetBox="[147,1408,197,1497]" captionTargetId="figure-25@12.[145,1428,195,1498]" captionTargetPageId="12" captionText="Figure 5. Legs. AE, sexual dimorphism on the front legs of B. onitoides. A, B, profemora: A, male; B, female. Note that the male profemur is claviform and has a strong emargination on its posterior edge (indicated by the red arrow), while that of the female is shorter and has a simply rounded aspect. CE, protibiae: C, D, dorsal view of male (C) and female (D). Observe that the male protibia is much longer and slenderer than that of the female and has a well-delimited tuft of long setae at its internal apical angle. E, lateral view of the male protibia. In contrast to females, males have a series of sharp, uneven teeth along the ventral protibial carina. Red arrow indicates the central tooth, which is longer than the others. F, G, meso- and metatarsi of B. onitoides. Unlike all other Phanaeini, which are clawless, B. onitoides has a pair of well-developed claws on the apical meso- and metatarsomeres. A small prolongation raises from underneath the claws and project itself anteriorly (indicated by the red arrow in F); this prolongation seems to be homologous to the tapered apex of the apical meso- and metatarsomeres of the other Phanaeini and is probably a synapomorphy of the clade. H, metanepisternum and its long tab typical of the Phanaeina." figureDoi="http://doi.org/10.5281/zenodo.7551998" httpUri="https://zenodo.org/record/7551998/files/figure.png" pageId="9" pageNumber="982">Fig. 5F</figureCitation>
). Metacoxae usually with long setae at apex (
<figureCitation box="[835,951,1850,1872]" captionStart="Figure 6" captionStartId="13.[164,244,1206,1228]" captionTargetBox="[165,1446,197,1164]" captionTargetId="figure-317@13.[163,1446,195,1166]" captionTargetPageId="13" captionText="Figure 6. Sexual dimorphism on the hind legs of B. onitoides. A, B, metacoxae: A, male; B, female. Note the plate-like projections and the long spines directed posteriorly in the male and which are lacking in the female. C, D, metafemora and metatibiae: C, male; D, female. Observe the ridge on the posterior edge of the metafemora and the sharp ridge on the external and internal edges of the metatibiae of the males, all of which are absent in females." figureDoi="http://doi.org/10.5281/zenodo.7552000" httpUri="https://zenodo.org/record/7552000/files/figure.png" pageId="9" pageNumber="982">Fig. 6A, B</figureCitation>
); sometimes setae absent due to abrasion (but setiferous punctures still visible over integument surface) or surface naturally glabrous; density of setation sexually dimorphic (see below). Metatibial spur gently curved apically.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7551991" ID-Zenodo-Dep="7551991" httpUri="https://zenodo.org/record/7551991/files/figure.png" pageId="10" pageNumber="983" startId="10.[145,223,1255,1277]" targetBox="[209,1358,197,1213]" targetPageId="10">
<paragraph blockId="10.[145,1425,1255,1394]" pageId="10" pageNumber="983">
<emphasis bold="true" box="[145,249,1255,1277]" pageId="10" pageNumber="983">Figure 3.</emphasis>
Dorsal sexual monomorphism in
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[606,790,1255,1277]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="983" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[606,790,1255,1277]" italics="true" pageId="10" pageNumber="983">Bolbites onitoides</emphasis>
</taxonomicName>
. A, frontal view of a male. B, C, head and pronotum: B, male; C, female. Note how little the sexes differ dorsally. Unlike most groups of Phanaeina, males of
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[1121,1241,1284,1306]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="983" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[1121,1241,1284,1306]" italics="true" pageId="10" pageNumber="983">B. onitoides</emphasis>
</taxonomicName>
lack any cephalic or pronotal horns and have a pronotal ridge identical to that of the females. This dorsal monomorphism is in striking contrast with the sexual dimorphism found on the venter. Note, however, the sexually dimorphic extension of iridescence over the clypeus, males having at least its basal half metallic, while females are entirely (or almost entirely) dull black.
</paragraph>
</caption>
<paragraph blockId="10.[145,761,1580,1909]" lastBlockId="10.[809,1425,1456,1601]" pageId="10" pageNumber="983">
<emphasis box="[145,223,1580,1601]" italics="true" pageId="10" pageNumber="983">Elytra:</emphasis>
With nine thin, ill-delimited striae, sometimes almost impossible to discern from the rest of integument except on bright humeri and apical calluses; ninth stria adjacent to epipleura. Integument with diffuse shine by being mostly covered by foveolate microsculpture (microfoveae only clearly discernible from one another at 70× magnification or more); bright, smooth areas limited to sutural margin, humeri and apical calluses; no granules or irregularities of any kind. Sutural margin elevated, highly shiny and smooth or with sparse micropunctures; apex sexually dimorphic. Humeri projected, usually entirely punctate and shiny. Apical calluses usually punctate and shiny. Base of elytra deeply depressed between third stria and humerus. Epipleura broad, with integument similar to most of rest of elytra.
</paragraph>
<paragraph blockId="10.[809,1426,1642,1909]" lastBlockId="11.[163,779,1678,1792]" lastPageId="11" lastPageNumber="984" pageId="10" pageNumber="983">
<emphasis box="[809,921,1642,1663]" italics="true" pageId="10" pageNumber="983">Abdomen:</emphasis>
With six ventrites. Ventrite I acute between apices of metacoxae (
<figureCitation box="[1087,1224,1672,1694]" captionStart="Figure 6" captionStartId="13.[164,244,1206,1228]" captionTargetBox="[165,1446,197,1164]" captionTargetId="figure-317@13.[163,1446,195,1166]" captionTargetPageId="13" captionText="Figure 6. Sexual dimorphism on the hind legs of B. onitoides. A, B, metacoxae: A, male; B, female. Note the plate-like projections and the long spines directed posteriorly in the male and which are lacking in the female. C, D, metafemora and metatibiae: C, male; D, female. Observe the ridge on the posterior edge of the metafemora and the sharp ridge on the external and internal edges of the metatibiae of the males, all of which are absent in females." figureDoi="http://doi.org/10.5281/zenodo.7552000" httpUri="https://zenodo.org/record/7552000/files/figure.png" pageId="10" pageNumber="983">Fig. 6A -B</figureCitation>
). Ventrites IV glabrous and with integument covered by thin alveolar microsculpture (alveoli only clearly discernible at 80× magnification); ventrite VI with similar integument on laterals and smooth at centre. Ventrite VI narrow, much shorter than the others, as wide in males as in females. Pygidium with integument entirely coated by thin alveolar microsculpture similar to that of ventrites and by small, rounded impressions, not usual punctures (
<figureCitation box="[170,229,1709,1731]" captionStart="Figure 7" captionStartId="14.[145,222,730,752]" captionTargetBox="[147,1429,197,689]" captionTargetId="figure-598@14.[145,1429,195,690]" captionTargetPageId="14" captionText="Figure 7. Sexual dimorphism on the pygidium and at the apex of the sutural margin of the elytra of B. onitoides: A, male; B, female. Orange arrow shows the short spines of the apex of the sutural margin of the elytra, whereas red arrow indicates the long spines raising from the basolateral angles of the pygidium of males. Both structures are absent in females." figureDoi="http://doi.org/10.5281/zenodo.7552002" httpUri="https://zenodo.org/record/7552002/files/figure.png" pageId="11" pageNumber="984">Fig. 7</figureCitation>
); basal margin complete, never interrupted by midlongitudinal groove of propygidium, sexually dimorphic (see below).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7551994" ID-Zenodo-Dep="7551994" httpUri="https://zenodo.org/record/7551994/files/figure.png" pageId="11" pageNumber="984" startId="11.[163,245,1393,1415]" targetBox="[166,1447,198,1350]" targetPageId="11">
<paragraph blockId="11.[163,1443,1393,1650]" pageId="11" pageNumber="984">
<emphasis bold="true" box="[163,273,1393,1415]" pageId="11" pageNumber="984">Figure 4.</emphasis>
Meso- and metaventrite in
<taxonomicName authorityName="Harold" authorityYear="1868" box="[589,677,1393,1414]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="984" phylum="Arthropoda" rank="genus">
<emphasis box="[589,677,1393,1414]" italics="true" pageId="11" pageNumber="984">Bolbites</emphasis>
</taxonomicName>
and Gromphadina. A,
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[931,1125,1393,1414]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="984" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[931,1125,1393,1414]" italics="true" pageId="11" pageNumber="984">Bolbites onitoides</emphasis>
</taxonomicName>
. B,
<taxonomicName authorityName="Harold, Black" authorityYear="1869" box="[1170,1375,1393,1415]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="984" phylum="Arthropoda" rank="species" species="inermis">
<emphasis box="[1170,1375,1393,1415]" italics="true" pageId="11" pageNumber="984">Gromphas inermis</emphasis>
</taxonomicName>
. C, D,
<taxonomicName baseAuthorityName="Erichson" baseAuthorityYear="1847" box="[163,338,1423,1445]" class="Insecta" family="Scarabaeidae" genus="Oruscatus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="984" phylum="Arthropoda" rank="species" species="davus">
<emphasis box="[163,338,1423,1445]" italics="true" pageId="11" pageNumber="984">Oruscatus davus</emphasis>
</taxonomicName>
: C, male; D, female. The carina of the mesofurcal brace constitutes the posterior branch of the mesoventral ridge and gives it the shape of a cross (orange arrow). This cross-shaped ridge is typical of the Gromphadina and is also present in at least some Phanaeina. Note that, while in the Gromphadina the posterior branch connects the mesoventral ridge to the anterior edge of the metaventrite, no such connection exists in most
<taxonomicName authorityName="Harold" authorityYear="1868" box="[1034,1120,1511,1533]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="984" phylum="Arthropoda" rank="genus">
<emphasis box="[1034,1120,1511,1533]" italics="true" pageId="11" pageNumber="984">Bolbites</emphasis>
</taxonomicName>
and and the other Phanaeina whose mesoventral ridge is T-shaped. In
<taxonomicName authorityName="Bates" authorityYear="1870" box="[584,691,1540,1562]" class="Insecta" family="Scarabaeidae" genus="Oruscatus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="984" phylum="Arthropoda" rank="genus">
<emphasis box="[584,691,1540,1562]" italics="true" pageId="11" pageNumber="984">Oruscatus</emphasis>
</taxonomicName>
, although present in both sexes, the posterior branch of the mesoventral ridge is much more clearly delimited in males than in females; in females, the posterior branch is usually incomplete and interrupted, while in males it is always sharp and continuous. Note also other differences between
<taxonomicName authorityName="Harold" authorityYear="1868" box="[1268,1353,1599,1621]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="984" phylum="Arthropoda" rank="genus">
<emphasis box="[1268,1353,1599,1621]" italics="true" pageId="11" pageNumber="984">Bolbites</emphasis>
</taxonomicName>
and the Gromphadina relating to the shape of the metaventral anterior prominence and its integument and lateral pilosity.
</paragraph>
</caption>
<paragraph blockId="11.[163,780,1830,1914]" lastBlockId="13.[163,780,1360,1689]" lastPageId="13" lastPageNumber="986" pageId="11" pageNumber="984">
<emphasis box="[163,477,1830,1852]" italics="true" pageId="11" pageNumber="984">Genitalia: Genital segment:</emphasis>
Mostly membranous (
<figureCitation captionStart="Figure 8" captionStartId="15.[163,241,1105,1127]" captionTargetBox="[181,784,202,1062]" captionTargetId="figure-551@15.[163,784,195,1065]" captionTargetPageId="15" captionText="Figure 8. Genitalia of both sexes of B. onitoides. A, female: spermatheca. BD, male: B, genital segment (i.e. nineth abdominal segment). C, D, tegmen. C, dorsal view. D, lateral view. LSP = lateral sclerotized plate; MSP = medial sclerotized plate." figureDoi="http://doi.org/10.5281/zenodo.7552004" httpUri="https://zenodo.org/record/7552004/files/figure.png" pageId="11" pageNumber="984">Fig. 8B</figureCitation>
). Medial sclerite paired and elongate. Lateral sclerotized plates strongly sclerotized.
<emphasis box="[655,778,1892,1913]" italics="true" pageId="11" pageNumber="984">Aedeagus:</emphasis>
Parameres tapered towards apex in both lateral and dorsal views (
<figureCitation box="[988,1104,1709,1731]" captionStart="Figure 8" captionStartId="15.[163,241,1105,1127]" captionTargetBox="[181,784,202,1062]" captionTargetId="figure-551@15.[163,784,195,1065]" captionTargetPageId="15" captionText="Figure 8. Genitalia of both sexes of B. onitoides. A, female: spermatheca. BD, male: B, genital segment (i.e. nineth abdominal segment). C, D, tegmen. C, dorsal view. D, lateral view. LSP = lateral sclerotized plate; MSP = medial sclerotized plate." figureDoi="http://doi.org/10.5281/zenodo.7552004" httpUri="https://zenodo.org/record/7552004/files/figure.png" pageId="11" pageNumber="984">Fig. 8C, D</figureCitation>
); in dorsal view, with conical aspect (
<figureCitation box="[923,1011,1740,1762]" captionStart="Figure 8" captionStartId="15.[163,241,1105,1127]" captionTargetBox="[181,784,202,1062]" captionTargetId="figure-551@15.[163,784,195,1065]" captionTargetPageId="15" captionText="Figure 8. Genitalia of both sexes of B. onitoides. A, female: spermatheca. BD, male: B, genital segment (i.e. nineth abdominal segment). C, D, tegmen. C, dorsal view. D, lateral view. LSP = lateral sclerotized plate; MSP = medial sclerotized plate." figureDoi="http://doi.org/10.5281/zenodo.7552004" httpUri="https://zenodo.org/record/7552004/files/figure.png" pageId="11" pageNumber="984">Fig. 8C</figureCitation>
); without hooks or notches; coated dorsally by wide area of whitish, glabrous integument. Phallobase, in ventral view, without median incision on basal edge or mid-longitudinal sulcus. Endophallus with five endophallites as follows (
<figureCitation box="[1228,1342,1862,1884]" captionStart="Figure 9" captionStartId="16.[145,225,1732,1754]" captionTargetBox="[305,1265,198,1689]" captionTargetId="figure-25@16.[302,1268,195,1692]" captionTargetPageId="16" captionText="Figure 9. Male genitalia: Endophallus of B. onitoides. A, B, general view of the extracted endophallus. A, dorsal view. B, ventral view. CG, endophallites: C, axial + subaxial endophallite complex (dissected). D, fronto-lateral peripheral endophallite (ventral view). E, medial endophallite (dorsal view). FH, superior right peripheral endophallite from different views: F, G, ventral view: F, seahorse-shaped; G, F-shaped. H, lateral view.A = axial endophallite; SA = subaxial endophallite; MS = medial endophallite; FLP = fronto-lateral peripheral endophallite; SRP = superior right peripheral endophallite." figureDoi="http://doi.org/10.5281/zenodo.7552012" httpUri="https://zenodo.org/record/7552012/files/figure.png" pageId="11" pageNumber="984">Fig. 9A, B</figureCitation>
): medial endophallite elongate and bent basally (
<figureCitation box="[1337,1426,1893,1915]" captionStart="Figure 9" captionStartId="16.[145,225,1732,1754]" captionTargetBox="[305,1265,198,1689]" captionTargetId="figure-25@16.[302,1268,195,1692]" captionTargetPageId="16" captionText="Figure 9. Male genitalia: Endophallus of B. onitoides. A, B, general view of the extracted endophallus. A, dorsal view. B, ventral view. CG, endophallites: C, axial + subaxial endophallite complex (dissected). D, fronto-lateral peripheral endophallite (ventral view). E, medial endophallite (dorsal view). FH, superior right peripheral endophallite from different views: F, G, ventral view: F, seahorse-shaped; G, F-shaped. H, lateral view.A = axial endophallite; SA = subaxial endophallite; MS = medial endophallite; FLP = fronto-lateral peripheral endophallite; SRP = superior right peripheral endophallite." figureDoi="http://doi.org/10.5281/zenodo.7552012" httpUri="https://zenodo.org/record/7552012/files/figure.png" pageId="11" pageNumber="984">Fig. 9E</figureCitation>
); superior right peripheral (
<collectionCode box="[461,513,1361,1383]" country="USA" lsid="urn:lsid:biocol.org:col:15742" name="Boise State University" pageId="13" pageNumber="986" type="Herbarium">SRP</collectionCode>
) endophallite elongate, bent apically and slightly variable in overall shape, depending on specimen (
<figureCitation box="[469,591,1422,1444]" captionStart="Figure 9" captionStartId="16.[145,225,1732,1754]" captionTargetBox="[305,1265,198,1689]" captionTargetId="figure-25@16.[302,1268,195,1692]" captionTargetPageId="16" captionText="Figure 9. Male genitalia: Endophallus of B. onitoides. A, B, general view of the extracted endophallus. A, dorsal view. B, ventral view. CG, endophallites: C, axial + subaxial endophallite complex (dissected). D, fronto-lateral peripheral endophallite (ventral view). E, medial endophallite (dorsal view). FH, superior right peripheral endophallite from different views: F, G, ventral view: F, seahorse-shaped; G, F-shaped. H, lateral view.A = axial endophallite; SA = subaxial endophallite; MS = medial endophallite; FLP = fronto-lateral peripheral endophallite; SRP = superior right peripheral endophallite." figureDoi="http://doi.org/10.5281/zenodo.7552012" httpUri="https://zenodo.org/record/7552012/files/figure.png" pageId="13" pageNumber="986">Fig. 9FH</figureCitation>
); axial (A) and subaxial (
<collectionCode box="[280,316,1453,1475]" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="13" pageNumber="986">SA</collectionCode>
) endophallites placed tightly together in resting position (
<figureCitation box="[391,475,1483,1505]" captionStart="Figure 9" captionStartId="16.[145,225,1732,1754]" captionTargetBox="[305,1265,198,1689]" captionTargetId="figure-25@16.[302,1268,195,1692]" captionTargetPageId="16" captionText="Figure 9. Male genitalia: Endophallus of B. onitoides. A, B, general view of the extracted endophallus. A, dorsal view. B, ventral view. CG, endophallites: C, axial + subaxial endophallite complex (dissected). D, fronto-lateral peripheral endophallite (ventral view). E, medial endophallite (dorsal view). FH, superior right peripheral endophallite from different views: F, G, ventral view: F, seahorse-shaped; G, F-shaped. H, lateral view.A = axial endophallite; SA = subaxial endophallite; MS = medial endophallite; FLP = fronto-lateral peripheral endophallite; SRP = superior right peripheral endophallite." figureDoi="http://doi.org/10.5281/zenodo.7552012" httpUri="https://zenodo.org/record/7552012/files/figure.png" pageId="13" pageNumber="986">Fig. 9C</figureCitation>
), A endophallite forked at one end and acuminate at the other, SA endophallite broad and curved; fronto-lateral peripheral (
<collectionCode box="[717,771,1545,1567]" pageId="13" pageNumber="986">FLP</collectionCode>
) endophallite fairly reduced, triangular, scale-like
<figureCitation captionStart="Figure 9" captionStartId="16.[145,225,1732,1754]" captionTargetBox="[305,1265,198,1689]" captionTargetId="figure-25@16.[302,1268,195,1692]" captionTargetPageId="16" captionText="Figure 9. Male genitalia: Endophallus of B. onitoides. A, B, general view of the extracted endophallus. A, dorsal view. B, ventral view. CG, endophallites: C, axial + subaxial endophallite complex (dissected). D, fronto-lateral peripheral endophallite (ventral view). E, medial endophallite (dorsal view). FH, superior right peripheral endophallite from different views: F, G, ventral view: F, seahorse-shaped; G, F-shaped. H, lateral view.A = axial endophallite; SA = subaxial endophallite; MS = medial endophallite; FLP = fronto-lateral peripheral endophallite; SRP = superior right peripheral endophallite." figureDoi="http://doi.org/10.5281/zenodo.7552012" httpUri="https://zenodo.org/record/7552012/files/figure.png" pageId="13" pageNumber="986">Fig. 9D</figureCitation>
.
<emphasis box="[211,376,1606,1627]" italics="true" pageId="13" pageNumber="986">Spermatheca:</emphasis>
C-shaped, apical branch slender, tapered and with rounded apex, basal branch broader (
<figureCitation box="[171,253,1667,1689]" captionStart="Figure 8" captionStartId="15.[163,241,1105,1127]" captionTargetBox="[181,784,202,1062]" captionTargetId="figure-551@15.[163,784,195,1065]" captionTargetPageId="15" captionText="Figure 8. Genitalia of both sexes of B. onitoides. A, female: spermatheca. BD, male: B, genital segment (i.e. nineth abdominal segment). C, D, tegmen. C, dorsal view. D, lateral view. LSP = lateral sclerotized plate; MSP = medial sclerotized plate." figureDoi="http://doi.org/10.5281/zenodo.7552004" httpUri="https://zenodo.org/record/7552004/files/figure.png" pageId="13" pageNumber="986">Fig. 8A</figureCitation>
). Spermathecal gland not observed.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7551998" ID-Zenodo-Dep="7551998" httpUri="https://zenodo.org/record/7551998/files/figure.png" pageId="12" pageNumber="985" startId="12.[145,223,1538,1560]" targetBox="[147,1408,197,1497]" targetPageId="12">
<paragraph blockId="12.[145,1425,1538,1853]" pageId="12" pageNumber="985">
<emphasis bold="true" box="[145,248,1538,1560]" pageId="12" pageNumber="985">Figure 5.</emphasis>
Legs. AE, sexual dimorphism on the front legs of
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[782,903,1538,1560]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="985" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[782,903,1538,1560]" italics="true" pageId="12" pageNumber="985">B. onitoides</emphasis>
</taxonomicName>
. A, B, profemora: A, male; B, female. Note that the male profemur is claviform and has a strong emargination on its posterior edge (indicated by the red arrow), while that of the female is shorter and has a simply rounded aspect. CE, protibiae: C, D, dorsal view of male (C) and female (D). Observe that the male protibia is much longer and slenderer than that of the female and has a well-delimited tuft of long setae at its internal apical angle. E, lateral view of the male protibia. In contrast to females, males have a series of sharp, uneven teeth along the ventral protibial carina. Red arrow indicates the central tooth, which is longer than the others. F, G, meso
<emphasis bold="true" box="[1370,1378,1685,1706]" pageId="12" pageNumber="985">-</emphasis>
and metatarsi of
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[279,400,1714,1736]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="985" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[279,400,1714,1736]" italics="true" pageId="12" pageNumber="985">B. onitoides</emphasis>
</taxonomicName>
. Unlike all other
<taxonomicName box="[584,693,1714,1736]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1032" phylum="Arthropoda" rank="tribe" tribe="Phanaeini">Phanaeini</taxonomicName>
, which are clawless,
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[911,1032,1714,1736]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="985" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[911,1032,1714,1736]" italics="true" pageId="12" pageNumber="985">B. onitoides</emphasis>
</taxonomicName>
has a pair of well-developed claws on the apical meso- and metatarsomeres. A small prolongation raises from underneath the claws and project itself anteriorly (indicated by the red arrow in F); this prolongation seems to be homologous to the tapered apex of the apical meso- and metatarsomeres of the other
<taxonomicName box="[447,554,1802,1824]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1032" phylum="Arthropoda" rank="tribe" tribe="Phanaeini">Phanaeini</taxonomicName>
and is probably a synapomorphy of the clade. H, metanepisternum and its long tab typical of the Phanaeina.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7552000" ID-Zenodo-Dep="7552000" httpUri="https://zenodo.org/record/7552000/files/figure.png" pageId="13" pageNumber="986" startId="13.[164,244,1206,1228]" targetBox="[165,1446,197,1164]" targetPageId="13">
<paragraph blockId="13.[163,1442,1206,1316]" pageId="13" pageNumber="986">
<emphasis bold="true" box="[164,271,1206,1228]" pageId="13" pageNumber="986">Figure 6.</emphasis>
Sexual dimorphism on the hind legs of
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[708,833,1206,1228]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="986" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[708,833,1206,1228]" italics="true" pageId="13" pageNumber="986">B. onitoides</emphasis>
</taxonomicName>
. A, B, metacoxae: A, male; B, female. Note the plate-like projections and the long spines directed posteriorly in the male and which are lacking in the female. C, D, metafemora and metatibiae: C, male; D, female. Observe the ridge on the posterior edge of the metafemora and the sharp ridge on the external and internal edges of the metatibiae of the males, all of which are absent in females.
</paragraph>
</caption>
<paragraph blockId="13.[163,779,1730,1905]" lastBlockId="14.[809,1424,855,907]" lastPageId="14" lastPageNumber="987" pageId="13" pageNumber="986">
<emphasis box="[163,479,1730,1751]" italics="true" pageId="13" pageNumber="986">Sexual dimorphism: Male:</emphasis>
Clypeus anterior-half or one-third dull black and posterior-half or two-thirds iridescent, with same colour as rest of head (
<figureCitation box="[656,764,1791,1813]" captionStart="Figure 3" captionStartId="10.[145,223,1255,1277]" captionTargetBox="[209,1358,197,1213]" captionTargetId="figure-230@10.[207,1362,195,1215]" captionTargetPageId="10" captionText="Figure 3. Dorsal sexual monomorphism in Bolbites onitoides. A, frontal view of a male. B, C, head and pronotum: B, male; C, female. Note how little the sexes differ dorsally. Unlike most groups of Phanaeina, males of B. onitoides lack any cephalic or pronotal horns and have a pronotal ridge identical to that of the females. This dorsal monomorphism is in striking contrast with the sexual dimorphism found on the venter. Note, however, the sexually dimorphic extension of iridescence over the clypeus, males having at least its basal half metallic, while females are entirely (or almost entirely) dull black." figureDoi="http://doi.org/10.5281/zenodo.7551991" httpUri="https://zenodo.org/record/7551991/files/figure.png" pageId="13" pageNumber="986">Fig. 3A, B</figureCitation>
); paraocular areas and frons usually almost completely iridescent. Pronotum raised into acute posterior tab on lateral margin (
<figureCitation box="[402,526,1883,1905]" captionStart="Figure 3" captionStartId="10.[145,223,1255,1277]" captionTargetBox="[209,1358,197,1213]" captionTargetId="figure-230@10.[207,1362,195,1215]" captionTargetPageId="10" captionText="Figure 3. Dorsal sexual monomorphism in Bolbites onitoides. A, frontal view of a male. B, C, head and pronotum: B, male; C, female. Note how little the sexes differ dorsally. Unlike most groups of Phanaeina, males of B. onitoides lack any cephalic or pronotal horns and have a pronotal ridge identical to that of the females. This dorsal monomorphism is in striking contrast with the sexual dimorphism found on the venter. Note, however, the sexually dimorphic extension of iridescence over the clypeus, males having at least its basal half metallic, while females are entirely (or almost entirely) dull black." figureDoi="http://doi.org/10.5281/zenodo.7551991" httpUri="https://zenodo.org/record/7551991/files/figure.png" pageId="13" pageNumber="986">Fig. 3A, B</figureCitation>
). Metaventrite with punctation at centre more smoothly marked than in females. Anterior leg longer than in females (
<figureCitation box="[1346,1427,1391,1413]" captionStart="Figure 3" captionStartId="10.[145,223,1255,1277]" captionTargetBox="[209,1358,197,1213]" captionTargetId="figure-230@10.[207,1362,195,1215]" captionTargetPageId="10" captionText="Figure 3. Dorsal sexual monomorphism in Bolbites onitoides. A, frontal view of a male. B, C, head and pronotum: B, male; C, female. Note how little the sexes differ dorsally. Unlike most groups of Phanaeina, males of B. onitoides lack any cephalic or pronotal horns and have a pronotal ridge identical to that of the females. This dorsal monomorphism is in striking contrast with the sexual dimorphism found on the venter. Note, however, the sexually dimorphic extension of iridescence over the clypeus, males having at least its basal half metallic, while females are entirely (or almost entirely) dull black." figureDoi="http://doi.org/10.5281/zenodo.7551991" httpUri="https://zenodo.org/record/7551991/files/figure.png" pageId="13" pageNumber="986">Fig. 3A</figureCitation>
), articulation between profemora and protibiae visible from above in pinned specimens mounted in traditional position. Profemora claviform, posterior edge with deep basal emargination followed by acute spine (
<figureCitation box="[914,1004,1544,1566]" captionStart="Figure 5" captionStartId="12.[145,223,1538,1560]" captionTargetBox="[147,1408,197,1497]" captionTargetId="figure-25@12.[145,1428,195,1498]" captionTargetPageId="12" captionText="Figure 5. Legs. AE, sexual dimorphism on the front legs of B. onitoides. A, B, profemora: A, male; B, female. Note that the male profemur is claviform and has a strong emargination on its posterior edge (indicated by the red arrow), while that of the female is shorter and has a simply rounded aspect. CE, protibiae: C, D, dorsal view of male (C) and female (D). Observe that the male protibia is much longer and slenderer than that of the female and has a well-delimited tuft of long setae at its internal apical angle. E, lateral view of the male protibia. In contrast to females, males have a series of sharp, uneven teeth along the ventral protibial carina. Red arrow indicates the central tooth, which is longer than the others. F, G, meso- and metatarsi of B. onitoides. Unlike all other Phanaeini, which are clawless, B. onitoides has a pair of well-developed claws on the apical meso- and metatarsomeres. A small prolongation raises from underneath the claws and project itself anteriorly (indicated by the red arrow in F); this prolongation seems to be homologous to the tapered apex of the apical meso- and metatarsomeres of the other Phanaeini and is probably a synapomorphy of the clade. H, metanepisternum and its long tab typical of the Phanaeina." figureDoi="http://doi.org/10.5281/zenodo.7551998" httpUri="https://zenodo.org/record/7551998/files/figure.png" pageId="13" pageNumber="986">Fig. 5A</figureCitation>
). Protibiae long and slender, with tuft of long and dense setae apically (
<figureCitation box="[1291,1378,1575,1597]" captionStart="Figure 3" captionStartId="10.[145,223,1255,1277]" captionTargetBox="[209,1358,197,1213]" captionTargetId="figure-230@10.[207,1362,195,1215]" captionTargetPageId="10" captionText="Figure 3. Dorsal sexual monomorphism in Bolbites onitoides. A, frontal view of a male. B, C, head and pronotum: B, male; C, female. Note how little the sexes differ dorsally. Unlike most groups of Phanaeina, males of B. onitoides lack any cephalic or pronotal horns and have a pronotal ridge identical to that of the females. This dorsal monomorphism is in striking contrast with the sexual dimorphism found on the venter. Note, however, the sexually dimorphic extension of iridescence over the clypeus, males having at least its basal half metallic, while females are entirely (or almost entirely) dull black." figureDoi="http://doi.org/10.5281/zenodo.7551991" httpUri="https://zenodo.org/record/7551991/files/figure.png" pageId="13" pageNumber="986">Fig. 3A</figureCitation>
,
<figureCitation box="[1394,1426,1575,1597]" captionStart="Figure 5" captionStartId="12.[145,223,1538,1560]" captionTargetBox="[147,1408,197,1497]" captionTargetId="figure-25@12.[145,1428,195,1498]" captionTargetPageId="12" captionText="Figure 5. Legs. AE, sexual dimorphism on the front legs of B. onitoides. A, B, profemora: A, male; B, female. Note that the male profemur is claviform and has a strong emargination on its posterior edge (indicated by the red arrow), while that of the female is shorter and has a simply rounded aspect. CE, protibiae: C, D, dorsal view of male (C) and female (D). Observe that the male protibia is much longer and slenderer than that of the female and has a well-delimited tuft of long setae at its internal apical angle. E, lateral view of the male protibia. In contrast to females, males have a series of sharp, uneven teeth along the ventral protibial carina. Red arrow indicates the central tooth, which is longer than the others. F, G, meso- and metatarsi of B. onitoides. Unlike all other Phanaeini, which are clawless, B. onitoides has a pair of well-developed claws on the apical meso- and metatarsomeres. A small prolongation raises from underneath the claws and project itself anteriorly (indicated by the red arrow in F); this prolongation seems to be homologous to the tapered apex of the apical meso- and metatarsomeres of the other Phanaeini and is probably a synapomorphy of the clade. H, metanepisternum and its long tab typical of the Phanaeina." figureDoi="http://doi.org/10.5281/zenodo.7551998" httpUri="https://zenodo.org/record/7551998/files/figure.png" pageId="13" pageNumber="986">5C</figureCitation>
); protibial spur shorter and broader than in females, clearly bent downwards (
<figureCitation box="[1123,1206,1636,1658]" captionStart="Figure 3" captionStartId="10.[145,223,1255,1277]" captionTargetBox="[209,1358,197,1213]" captionTargetId="figure-230@10.[207,1362,195,1215]" captionTargetPageId="10" captionText="Figure 3. Dorsal sexual monomorphism in Bolbites onitoides. A, frontal view of a male. B, C, head and pronotum: B, male; C, female. Note how little the sexes differ dorsally. Unlike most groups of Phanaeina, males of B. onitoides lack any cephalic or pronotal horns and have a pronotal ridge identical to that of the females. This dorsal monomorphism is in striking contrast with the sexual dimorphism found on the venter. Note, however, the sexually dimorphic extension of iridescence over the clypeus, males having at least its basal half metallic, while females are entirely (or almost entirely) dull black." figureDoi="http://doi.org/10.5281/zenodo.7551991" httpUri="https://zenodo.org/record/7551991/files/figure.png" pageId="13" pageNumber="986">Fig. 3A</figureCitation>
,
<figureCitation box="[1221,1252,1636,1658]" captionStart="Figure 5" captionStartId="12.[145,223,1538,1560]" captionTargetBox="[147,1408,197,1497]" captionTargetId="figure-25@12.[145,1428,195,1498]" captionTargetPageId="12" captionText="Figure 5. Legs. AE, sexual dimorphism on the front legs of B. onitoides. A, B, profemora: A, male; B, female. Note that the male profemur is claviform and has a strong emargination on its posterior edge (indicated by the red arrow), while that of the female is shorter and has a simply rounded aspect. CE, protibiae: C, D, dorsal view of male (C) and female (D). Observe that the male protibia is much longer and slenderer than that of the female and has a well-delimited tuft of long setae at its internal apical angle. E, lateral view of the male protibia. In contrast to females, males have a series of sharp, uneven teeth along the ventral protibial carina. Red arrow indicates the central tooth, which is longer than the others. F, G, meso- and metatarsi of B. onitoides. Unlike all other Phanaeini, which are clawless, B. onitoides has a pair of well-developed claws on the apical meso- and metatarsomeres. A small prolongation raises from underneath the claws and project itself anteriorly (indicated by the red arrow in F); this prolongation seems to be homologous to the tapered apex of the apical meso- and metatarsomeres of the other Phanaeini and is probably a synapomorphy of the clade. H, metanepisternum and its long tab typical of the Phanaeina." figureDoi="http://doi.org/10.5281/zenodo.7551998" httpUri="https://zenodo.org/record/7551998/files/figure.png" pageId="13" pageNumber="986">5C</figureCitation>
); ventral carina interrupted by series of acute teeth irregular in size, central tooth larger than others (
<figureCitation box="[1213,1296,1697,1720]" captionStart="Figure 5" captionStartId="12.[145,223,1538,1560]" captionTargetBox="[147,1408,197,1497]" captionTargetId="figure-25@12.[145,1428,195,1498]" captionTargetPageId="12" captionText="Figure 5. Legs. AE, sexual dimorphism on the front legs of B. onitoides. A, B, profemora: A, male; B, female. Note that the male profemur is claviform and has a strong emargination on its posterior edge (indicated by the red arrow), while that of the female is shorter and has a simply rounded aspect. CE, protibiae: C, D, dorsal view of male (C) and female (D). Observe that the male protibia is much longer and slenderer than that of the female and has a well-delimited tuft of long setae at its internal apical angle. E, lateral view of the male protibia. In contrast to females, males have a series of sharp, uneven teeth along the ventral protibial carina. Red arrow indicates the central tooth, which is longer than the others. F, G, meso- and metatarsi of B. onitoides. Unlike all other Phanaeini, which are clawless, B. onitoides has a pair of well-developed claws on the apical meso- and metatarsomeres. A small prolongation raises from underneath the claws and project itself anteriorly (indicated by the red arrow in F); this prolongation seems to be homologous to the tapered apex of the apical meso- and metatarsomeres of the other Phanaeini and is probably a synapomorphy of the clade. H, metanepisternum and its long tab typical of the Phanaeina." figureDoi="http://doi.org/10.5281/zenodo.7551998" httpUri="https://zenodo.org/record/7551998/files/figure.png" pageId="13" pageNumber="986">Fig. 5E</figureCitation>
). Metacoxae with plate projections bearing long, acute posterior spine near insertion of trochanters (
<figureCitation box="[1245,1327,1759,1781]" captionStart="Figure 6" captionStartId="13.[164,244,1206,1228]" captionTargetBox="[165,1446,197,1164]" captionTargetId="figure-317@13.[163,1446,195,1166]" captionTargetPageId="13" captionText="Figure 6. Sexual dimorphism on the hind legs of B. onitoides. A, B, metacoxae: A, male; B, female. Note the plate-like projections and the long spines directed posteriorly in the male and which are lacking in the female. C, D, metafemora and metatibiae: C, male; D, female. Observe the ridge on the posterior edge of the metafemora and the sharp ridge on the external and internal edges of the metatibiae of the males, all of which are absent in females." figureDoi="http://doi.org/10.5281/zenodo.7552000" httpUri="https://zenodo.org/record/7552000/files/figure.png" pageId="13" pageNumber="986">Fig. 6A</figureCitation>
); setation of apical surface usually shorter and sparser than in females, not rarely even entirely glabrous. Metafemora straight on anterior edge, posterior edge with large rounded ridge (
<figureCitation box="[1204,1290,1882,1904]" captionStart="Figure 6" captionStartId="13.[164,244,1206,1228]" captionTargetBox="[165,1446,197,1164]" captionTargetId="figure-317@13.[163,1446,195,1166]" captionTargetPageId="13" captionText="Figure 6. Sexual dimorphism on the hind legs of B. onitoides. A, B, metacoxae: A, male; B, female. Note the plate-like projections and the long spines directed posteriorly in the male and which are lacking in the female. C, D, metafemora and metatibiae: C, male; D, female. Observe the ridge on the posterior edge of the metafemora and the sharp ridge on the external and internal edges of the metatibiae of the males, all of which are absent in females." figureDoi="http://doi.org/10.5281/zenodo.7552000" httpUri="https://zenodo.org/record/7552000/files/figure.png" pageId="13" pageNumber="986">Fig. 6C</figureCitation>
). Metatibiae with strong ridges on external and internal edges, internal edges with few small spines (
<figureCitation box="[651,743,886,908]" captionStart="Figure 6" captionStartId="13.[164,244,1206,1228]" captionTargetBox="[165,1446,197,1164]" captionTargetId="figure-317@13.[163,1446,195,1166]" captionTargetPageId="13" captionText="Figure 6. Sexual dimorphism on the hind legs of B. onitoides. A, B, metacoxae: A, male; B, female. Note the plate-like projections and the long spines directed posteriorly in the male and which are lacking in the female. C, D, metafemora and metatibiae: C, male; D, female. Observe the ridge on the posterior edge of the metafemora and the sharp ridge on the external and internal edges of the metatibiae of the males, all of which are absent in females." figureDoi="http://doi.org/10.5281/zenodo.7552000" httpUri="https://zenodo.org/record/7552000/files/figure.png" pageId="14" pageNumber="987">Fig. 6C</figureCitation>
). Sutural margins of elytra with short spines at apex (
<figureCitation box="[219,306,947,969]" captionStart="Figure 7" captionStartId="14.[145,222,730,752]" captionTargetBox="[147,1429,197,689]" captionTargetId="figure-598@14.[145,1429,195,690]" captionTargetPageId="14" captionText="Figure 7. Sexual dimorphism on the pygidium and at the apex of the sutural margin of the elytra of B. onitoides: A, male; B, female. Orange arrow shows the short spines of the apex of the sutural margin of the elytra, whereas red arrow indicates the long spines raising from the basolateral angles of the pygidium of males. Both structures are absent in females." figureDoi="http://doi.org/10.5281/zenodo.7552002" httpUri="https://zenodo.org/record/7552002/files/figure.png" pageId="14" pageNumber="987">Fig. 7A</figureCitation>
, yellow arrow). Pygidium raised into pair of long spines at basolateral angles of pygidial margin, foveate on surface near spines (
<figureCitation box="[617,700,1008,1030]" captionStart="Figure 6" captionStartId="13.[164,244,1206,1228]" captionTargetBox="[165,1446,197,1164]" captionTargetId="figure-317@13.[163,1446,195,1166]" captionTargetPageId="13" captionText="Figure 6. Sexual dimorphism on the hind legs of B. onitoides. A, B, metacoxae: A, male; B, female. Note the plate-like projections and the long spines directed posteriorly in the male and which are lacking in the female. C, D, metafemora and metatibiae: C, male; D, female. Observe the ridge on the posterior edge of the metafemora and the sharp ridge on the external and internal edges of the metatibiae of the males, all of which are absent in females." figureDoi="http://doi.org/10.5281/zenodo.7552000" httpUri="https://zenodo.org/record/7552000/files/figure.png" pageId="14" pageNumber="987">Fig. 6C</figureCitation>
,
<figureCitation box="[714,746,1009,1030]" captionStart="Figure 7" captionStartId="14.[145,222,730,752]" captionTargetBox="[147,1429,197,689]" captionTargetId="figure-598@14.[145,1429,195,690]" captionTargetPageId="14" captionText="Figure 7. Sexual dimorphism on the pygidium and at the apex of the sutural margin of the elytra of B. onitoides: A, male; B, female. Orange arrow shows the short spines of the apex of the sutural margin of the elytra, whereas red arrow indicates the long spines raising from the basolateral angles of the pygidium of males. Both structures are absent in females." figureDoi="http://doi.org/10.5281/zenodo.7552002" httpUri="https://zenodo.org/record/7552002/files/figure.png" pageId="14" pageNumber="987">7A</figureCitation>
); lateral margin partly interrupted near spines in larger specimens.
<emphasis box="[380,477,1070,1091]" italics="true" pageId="14" pageNumber="987">Female:</emphasis>
Clypeus entirely dull black, traces of iridescence at most near small areas adjacent to frons and paraocular areas (
<figureCitation captionStart="Figure 3" captionStartId="10.[145,223,1255,1277]" captionTargetBox="[209,1358,197,1213]" captionTargetId="figure-230@10.[207,1362,195,1215]" captionTargetPageId="10" captionText="Figure 3. Dorsal sexual monomorphism in Bolbites onitoides. A, frontal view of a male. B, C, head and pronotum: B, male; C, female. Note how little the sexes differ dorsally. Unlike most groups of Phanaeina, males of B. onitoides lack any cephalic or pronotal horns and have a pronotal ridge identical to that of the females. This dorsal monomorphism is in striking contrast with the sexual dimorphism found on the venter. Note, however, the sexually dimorphic extension of iridescence over the clypeus, males having at least its basal half metallic, while females are entirely (or almost entirely) dull black." figureDoi="http://doi.org/10.5281/zenodo.7551991" httpUri="https://zenodo.org/record/7551991/files/figure.png" pageId="14" pageNumber="987">Fig. 3C</figureCitation>
); anterior region of paraocular areas and of frons sometimes also dull black. Pronotum not raised into acute tab on lateral margin (
<figureCitation box="[655,744,1223,1245]" captionStart="Figure 3" captionStartId="10.[145,223,1255,1277]" captionTargetBox="[209,1358,197,1213]" captionTargetId="figure-230@10.[207,1362,195,1215]" captionTargetPageId="10" captionText="Figure 3. Dorsal sexual monomorphism in Bolbites onitoides. A, frontal view of a male. B, C, head and pronotum: B, male; C, female. Note how little the sexes differ dorsally. Unlike most groups of Phanaeina, males of B. onitoides lack any cephalic or pronotal horns and have a pronotal ridge identical to that of the females. This dorsal monomorphism is in striking contrast with the sexual dimorphism found on the venter. Note, however, the sexually dimorphic extension of iridescence over the clypeus, males having at least its basal half metallic, while females are entirely (or almost entirely) dull black." figureDoi="http://doi.org/10.5281/zenodo.7551991" httpUri="https://zenodo.org/record/7551991/files/figure.png" pageId="14" pageNumber="987">Fig. 3C</figureCitation>
). Metaventrite with punctation at centre coarser than in males. Anterior leg shorter than in males (
<figureCitation captionStart="Figure 3" captionStartId="10.[145,223,1255,1277]" captionTargetBox="[209,1358,197,1213]" captionTargetId="figure-230@10.[207,1362,195,1215]" captionTargetPageId="10" captionText="Figure 3. Dorsal sexual monomorphism in Bolbites onitoides. A, frontal view of a male. B, C, head and pronotum: B, male; C, female. Note how little the sexes differ dorsally. Unlike most groups of Phanaeina, males of B. onitoides lack any cephalic or pronotal horns and have a pronotal ridge identical to that of the females. This dorsal monomorphism is in striking contrast with the sexual dimorphism found on the venter. Note, however, the sexually dimorphic extension of iridescence over the clypeus, males having at least its basal half metallic, while females are entirely (or almost entirely) dull black." figureDoi="http://doi.org/10.5281/zenodo.7551991" httpUri="https://zenodo.org/record/7551991/files/figure.png" pageId="14" pageNumber="987">Fig. 3C</figureCitation>
), articulation between profemora and protibiae not visible from above. Profemora entirely broad in females, without posterior emargination or spine (
<figureCitation box="[154,245,1407,1429]" captionStart="Figure 5" captionStartId="12.[145,223,1538,1560]" captionTargetBox="[147,1408,197,1497]" captionTargetId="figure-25@12.[145,1428,195,1498]" captionTargetPageId="12" captionText="Figure 5. Legs. AE, sexual dimorphism on the front legs of B. onitoides. A, B, profemora: A, male; B, female. Note that the male profemur is claviform and has a strong emargination on its posterior edge (indicated by the red arrow), while that of the female is shorter and has a simply rounded aspect. CE, protibiae: C, D, dorsal view of male (C) and female (D). Observe that the male protibia is much longer and slenderer than that of the female and has a well-delimited tuft of long setae at its internal apical angle. E, lateral view of the male protibia. In contrast to females, males have a series of sharp, uneven teeth along the ventral protibial carina. Red arrow indicates the central tooth, which is longer than the others. F, G, meso- and metatarsi of B. onitoides. Unlike all other Phanaeini, which are clawless, B. onitoides has a pair of well-developed claws on the apical meso- and metatarsomeres. A small prolongation raises from underneath the claws and project itself anteriorly (indicated by the red arrow in F); this prolongation seems to be homologous to the tapered apex of the apical meso- and metatarsomeres of the other Phanaeini and is probably a synapomorphy of the clade. H, metanepisternum and its long tab typical of the Phanaeina." figureDoi="http://doi.org/10.5281/zenodo.7551998" httpUri="https://zenodo.org/record/7551998/files/figure.png" pageId="14" pageNumber="987">Fig. 5B</figureCitation>
). Protibiae broader and shorter than in males, without well-defined apical tuft of long setae (at most, with few shorter setae on internal apical angle) (
<figureCitation box="[245,338,1499,1521]" captionStart="Figure 5" captionStartId="12.[145,223,1538,1560]" captionTargetBox="[147,1408,197,1497]" captionTargetId="figure-25@12.[145,1428,195,1498]" captionTargetPageId="12" captionText="Figure 5. Legs. AE, sexual dimorphism on the front legs of B. onitoides. A, B, profemora: A, male; B, female. Note that the male profemur is claviform and has a strong emargination on its posterior edge (indicated by the red arrow), while that of the female is shorter and has a simply rounded aspect. CE, protibiae: C, D, dorsal view of male (C) and female (D). Observe that the male protibia is much longer and slenderer than that of the female and has a well-delimited tuft of long setae at its internal apical angle. E, lateral view of the male protibia. In contrast to females, males have a series of sharp, uneven teeth along the ventral protibial carina. Red arrow indicates the central tooth, which is longer than the others. F, G, meso- and metatarsi of B. onitoides. Unlike all other Phanaeini, which are clawless, B. onitoides has a pair of well-developed claws on the apical meso- and metatarsomeres. A small prolongation raises from underneath the claws and project itself anteriorly (indicated by the red arrow in F); this prolongation seems to be homologous to the tapered apex of the apical meso- and metatarsomeres of the other Phanaeini and is probably a synapomorphy of the clade. H, metanepisternum and its long tab typical of the Phanaeina." figureDoi="http://doi.org/10.5281/zenodo.7551998" httpUri="https://zenodo.org/record/7551998/files/figure.png" pageId="14" pageNumber="987">Fig. 5D</figureCitation>
); protibial spur long, spiniform, slightly bent downwards; ventral carina simple, without teeth. Metacoxae unarmed, without plate projection or spines (
<figureCitation box="[419,510,1591,1613]" captionStart="Figure 6" captionStartId="13.[164,244,1206,1228]" captionTargetBox="[165,1446,197,1164]" captionTargetId="figure-317@13.[163,1446,195,1166]" captionTargetPageId="13" captionText="Figure 6. Sexual dimorphism on the hind legs of B. onitoides. A, B, metacoxae: A, male; B, female. Note the plate-like projections and the long spines directed posteriorly in the male and which are lacking in the female. C, D, metafemora and metatibiae: C, male; D, female. Observe the ridge on the posterior edge of the metafemora and the sharp ridge on the external and internal edges of the metatibiae of the males, all of which are absent in females." figureDoi="http://doi.org/10.5281/zenodo.7552000" httpUri="https://zenodo.org/record/7552000/files/figure.png" pageId="14" pageNumber="987">Fig. 6B</figureCitation>
); setation of apical surface usually much longer and denser than in males, only rarely glabrous. Metafemora usually (but not always) slightly curved at basal half of anterior edge (
<figureCitation box="[217,303,1713,1736]" captionStart="Figure 6" captionStartId="13.[164,244,1206,1228]" captionTargetBox="[165,1446,197,1164]" captionTargetId="figure-317@13.[163,1446,195,1166]" captionTargetPageId="13" captionText="Figure 6. Sexual dimorphism on the hind legs of B. onitoides. A, B, metacoxae: A, male; B, female. Note the plate-like projections and the long spines directed posteriorly in the male and which are lacking in the female. C, D, metafemora and metatibiae: C, male; D, female. Observe the ridge on the posterior edge of the metafemora and the sharp ridge on the external and internal edges of the metatibiae of the males, all of which are absent in females." figureDoi="http://doi.org/10.5281/zenodo.7552000" httpUri="https://zenodo.org/record/7552000/files/figure.png" pageId="14" pageNumber="987">Fig. 6D</figureCitation>
); posterior edge unarmed. Metatibiae with sharp teeth along external edge, but without strong ridge on external and internal edges (
<figureCitation captionStart="Figure 6" captionStartId="13.[164,244,1206,1228]" captionTargetBox="[165,1446,197,1164]" captionTargetId="figure-317@13.[163,1446,195,1166]" captionTargetPageId="13" captionText="Figure 6. Sexual dimorphism on the hind legs of B. onitoides. A, B, metacoxae: A, male; B, female. Note the plate-like projections and the long spines directed posteriorly in the male and which are lacking in the female. C, D, metafemora and metatibiae: C, male; D, female. Observe the ridge on the posterior edge of the metafemora and the sharp ridge on the external and internal edges of the metatibiae of the males, all of which are absent in females." figureDoi="http://doi.org/10.5281/zenodo.7552000" httpUri="https://zenodo.org/record/7552000/files/figure.png" pageId="14" pageNumber="987">Fig. 6D</figureCitation>
). Sutural margin of elytra unarmed apically (
<figureCitation captionStart="Figure 7" captionStartId="14.[145,222,730,752]" captionTargetBox="[147,1429,197,689]" captionTargetId="figure-598@14.[145,1429,195,690]" captionTargetPageId="14" captionText="Figure 7. Sexual dimorphism on the pygidium and at the apex of the sutural margin of the elytra of B. onitoides: A, male; B, female. Orange arrow shows the short spines of the apex of the sutural margin of the elytra, whereas red arrow indicates the long spines raising from the basolateral angles of the pygidium of males. Both structures are absent in females." figureDoi="http://doi.org/10.5281/zenodo.7552002" httpUri="https://zenodo.org/record/7552002/files/figure.png" pageId="14" pageNumber="987">Fig. 7B</figureCitation>
). Pygidium gently depressed near basolateral angles; basolateral angles not raised into spines, at most slighly elevated; pygidial lateral margin with emargination less evident than in males (
<figureCitation box="[1300,1387,885,907]" captionStart="Figure 7" captionStartId="14.[145,222,730,752]" captionTargetBox="[147,1429,197,689]" captionTargetId="figure-598@14.[145,1429,195,690]" captionTargetPageId="14" captionText="Figure 7. Sexual dimorphism on the pygidium and at the apex of the sutural margin of the elytra of B. onitoides: A, male; B, female. Orange arrow shows the short spines of the apex of the sutural margin of the elytra, whereas red arrow indicates the long spines raising from the basolateral angles of the pygidium of males. Both structures are absent in females." figureDoi="http://doi.org/10.5281/zenodo.7552002" httpUri="https://zenodo.org/record/7552002/files/figure.png" pageId="14" pageNumber="987">Fig. 7B</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7552002" ID-Zenodo-Dep="7552002" httpUri="https://zenodo.org/record/7552002/files/figure.png" pageId="14" pageNumber="987" startId="14.[145,222,730,752]" targetBox="[147,1429,197,689]" targetPageId="14">
<paragraph blockId="14.[145,1425,730,811]" pageId="14" pageNumber="987">
<emphasis bold="true" box="[145,247,730,752]" pageId="14" pageNumber="987">Figure 7.</emphasis>
Sexual dimorphism on the pygidium and at the apex of the sutural margin of the elytra of
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[1186,1304,730,752]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="987" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[1186,1304,730,752]" italics="true" pageId="14" pageNumber="987">B. onitoides</emphasis>
</taxonomicName>
: A, male; B, female. Orange arrow shows the short spines of the apex of the sutural margin of the elytra, whereas red arrow indicates the long spines raising from the basolateral angles of the pygidium of males. Both structures are absent in females.
</paragraph>
</caption>
<paragraph blockId="14.[809,1425,948,1216]" pageId="14" pageNumber="987">
<emphasis box="[809,1234,948,970]" italics="true" pageId="14" pageNumber="987">Measurements (in millimetres):</emphasis>
<emphasis bold="true" box="[1249,1342,948,969]" pageId="14" pageNumber="987">Males:</emphasis>
<emphasis italics="true" pageId="14" pageNumber="987">Total length (TL):</emphasis>
14.821.5 [average (AV) = 18.8 ± 1.85].
<emphasis box="[809,1282,1009,1031]" italics="true" pageId="14" pageNumber="987">
Partial length (without head;
<collectionCode box="[1227,1262,1010,1031]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="14" pageNumber="987" type="Museum">PL</collectionCode>
):
</emphasis>
11.117.5 (AV = 15.5 ± 1.48).
<emphasis italics="true" pageId="14" pageNumber="987">
Maximum width of pronotum (
<collectionCode box="[817,859,1071,1092]" pageId="14" pageNumber="987">WP</collectionCode>
):
</emphasis>
7.912.7 (AV = 11 ± 1.15).
<emphasis italics="true" pageId="14" pageNumber="987">
Maximum width of elytra (
<collectionCode box="[895,938,1102,1123]" pageId="14" pageNumber="987">WE</collectionCode>
):
</emphasis>
8.212.9 (AV = 11.4 ± 1.11).
<emphasis bold="true" box="[1309,1425,1102,1123]" pageId="14" pageNumber="987">Females:</emphasis>
<emphasis box="[809,850,1133,1154]" italics="true" pageId="14" pageNumber="987">TL:</emphasis>
14.020.7 (AV = 17.9 ± 1.93).
<emphasis box="[1207,1247,1133,1154]" italics="true" pageId="14" pageNumber="987">PL:</emphasis>
11.916.7 (AV: 14.4 ± 1.69).
<emphasis box="[953,999,1164,1185]" italics="true" pageId="14" pageNumber="987">WP:</emphasis>
8.612.5 (AV = 10.7 ± 1.32).
<emphasis box="[1323,1370,1164,1185]" italics="true" pageId="14" pageNumber="987">WE:</emphasis>
8.8 12.6 (AV = 10.9 ± 1.29).
</paragraph>
<paragraph blockId="14.[809,1426,1256,1554]" pageId="14" pageNumber="987">
<emphasis box="[809,1039,1256,1278]" italics="true" pageId="14" pageNumber="987">Geographical range:</emphasis>
Present mostly across xerophytic deciduous forests, savannas and grass- and shrublands from the dry Chaco in southern
<collectingCountry box="[1202,1287,1317,1339]" name="Bolivia" pageId="14" pageNumber="987">Bolivia</collectingCountry>
and northwestern
<collectingCountry box="[912,1025,1349,1370]" name="Paraguay" pageId="14" pageNumber="987">Paraguay</collectingCountry>
and the humid Chaco in eastern
<collectingCountry box="[809,918,1379,1400]" name="Paraguay" pageId="14" pageNumber="987">Paraguay</collectingCountry>
and
<collectingRegion box="[972,1252,1379,1401]" country="Brazil" name="Mato Grosso do Sul" pageId="14" pageNumber="987">Mato Grosso do Sul state</collectingRegion>
in
<collectingCountry box="[1286,1356,1379,1401]" name="Brazil" pageId="14" pageNumber="987">Brazil</collectingCountry>
south to the Uruguayan and southern Brazilian Savanna and the Espinal region in central
<collectingCountry box="[1223,1347,1440,1462]" name="Argentina" pageId="14" pageNumber="987">Argentina</collectingCountry>
. Some populations also present in higher and drier lands along the Argentinian Low Monte region (
<figureCitation box="[1294,1378,1501,1523]" captionStart="Figure 11" captionStartId="32.[145,223,1631,1653]" captionTargetBox="[147,1428,197,1590]" captionTargetId="figure-25@32.[145,1428,195,1591]" captionTargetPageId="32" captionText="Figure 11. Geographical range of B. onitoides showing the differential distribution of the colour variants. It is indisputable that there is a strong correlation between colour and geographical provenance in the species: blue individuals are abundant at the western-half of the distribution, while red specimens are dominant at the eastern-half. Nevertheless, two kinds of intermediates the discrete bicoloured and the gradual colour-blended are found throughout the range of B. onitoides, but nowhere are they the dominant variant. Note that there is a zone of overlap between the distribution of red and blue variants in Paraguay and Argentina, and that three localities recorded red, blue and bicoloured individuals and a fourth recorded all four variants." figureDoi="http://doi.org/10.5281/zenodo.7552016" httpUri="https://zenodo.org/record/7552016/files/figure.png" pageId="14" pageNumber="987">Figs 11</figureCitation>
,
<figureCitation box="[1391,1419,1502,1523]" captionStart="Figure 12" captionStartId="33.[163,241,1286,1308]" captionTargetBox="[166,1446,198,1244]" captionTargetId="figure-283@33.[163,1449,195,1247]" captionTargetPageId="33" captionText="Figure 12. Distribution of each colour variant showed in separate maps for an easier discrimination. See legend for Figure 10 for more details." figureDoi="http://doi.org/10.5281/zenodo.7552018" httpUri="https://zenodo.org/record/7552018/files/figure.png" pageId="14" pageNumber="987">12</figureCitation>
; Supporting Information, Appendix S1).
</paragraph>
<paragraph blockId="14.[809,1425,1595,1739]" pageId="14" pageNumber="987">
<emphasis box="[809,1102,1595,1616]" italics="true" pageId="14" pageNumber="987">Biomes and ecoregions:</emphasis>
Tropical and subtropical grasslands, savannas, and shrublands: Dry
<collectingRegion box="[1345,1419,1625,1647]" country="Argentina" name="Chaco" pageId="14" pageNumber="987">Chaco</collectingRegion>
, Humid
<collectingRegion box="[908,987,1656,1678]" country="Argentina" name="Chaco" pageId="14" pageNumber="987">Chaco</collectingRegion>
, Uruguayan Savanna. Temperate grasslands, savannas, and shrublands: Humid Pampas, Espinal, Low Monte.
</paragraph>
<paragraph blockId="14.[809,1425,1775,1889]" lastBlockId="15.[827,1443,197,1476]" lastPageId="15" lastPageNumber="988" pageId="14" pageNumber="987">
<emphasis box="[809,1066,1775,1796]" italics="true" pageId="14" pageNumber="987">Collecting localities:</emphasis>
<emphasis bold="true" box="[1078,1296,1775,1797]" pageId="14" pageNumber="987">Red specimens:</emphasis>
<materialsCitation ID-GBIF-Occurrence="4016047311" lastPageId="14" lastPageNumber="988" pageId="14" pageNumber="987">
<collectingCountry box="[1307,1423,1775,1796]" name="Bolivia" pageId="14" pageNumber="987">BOLIVIA</collectingCountry>
[?].
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047364" box="[855,1424,1805,1827]" lastPageId="14" lastPageNumber="988" pageId="14" pageNumber="987">
<collectingCountry box="[855,958,1806,1827]" name="Brazil" pageId="14" pageNumber="987">BRAZIL</collectingCountry>
:
<collectingRegion box="[971,1215,1805,1827]" country="Brazil" name="Mato Grosso do Sul" pageId="14" pageNumber="987">Mato Grosso do Sul</collectingRegion>
:
<collectorName box="[1228,1420,1805,1827]" pageId="14" pageNumber="987">Porto Murtinho</collectorName>
;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047330" box="[809,1260,1836,1858]" lastPageId="14" lastPageNumber="988" pageId="14" pageNumber="987">
<collectingRegion box="[809,1045,1836,1858]" country="Brazil" name="Rio Grande do Sul" pageId="14" pageNumber="987">Rio Grande do Sul</collectingRegion>
:
<collectorName box="[1059,1151,1836,1858]" pageId="14" pageNumber="987">Aceguá</collectorName>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC0EFFF23AB60A27DFCAF8B1" box="[1166,1256,1836,1858]" country="Bolivia" name="Guarai" pageId="14" pageNumber="987" stateProvince="Rio Grande do Sul">Guaraí</location>
.
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047374" lastPageId="14" lastPageNumber="988" pageId="14" pageNumber="987">
<collectingCountry box="[1271,1421,1836,1857]" name="Paraguay" pageId="14" pageNumber="987">PARAGUAY</collectingCountry>
:
<collectingRegion box="[809,988,1867,1889]" country="Paraguay" name="Alto Paraguay" pageId="14" pageNumber="987">Alto Paraguay</collectingRegion>
:
<collectorName box="[1002,1183,1867,1889]" pageId="14" pageNumber="987">Puerto Casado</collectorName>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC0EFFF23A960A40DEA9F893" box="[1198,1419,1867,1888]" country="Paraguay" name="Puerto Esperanza" pageId="14" pageNumber="987" stateProvince="Concepcion">Puerto Esperanza</location>
;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047348" box="[163,466,1302,1324]" lastPageId="15" lastPageNumber="988" pageId="15" pageNumber="987">
<collectingRegion box="[163,303,1302,1324]" country="Paraguay" name="Concepcion" pageId="15" pageNumber="988">Concepción</collectingRegion>
:
<collectorName box="[316,462,1303,1324]" pageId="15" pageNumber="988">Puerto Max</collectorName>
;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047377" lastPageId="15" lastPageNumber="988" pageId="15" pageNumber="987">
<collectingRegion box="[476,695,1302,1324]" country="Paraguay" name="Presidente Hayes" pageId="15" pageNumber="988">Presidente Hayes</collectingRegion>
:
<collectorName box="[708,778,1302,1323]" pageId="15" pageNumber="988">Cruce</collectorName>
de los Pioneros;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047315" lastPageId="15" lastPageNumber="988" pageId="15" pageNumber="987">
<collectingRegion box="[364,488,1333,1355]" country="Paraguay" name="Paraguari" pageId="15" pageNumber="988">Paraguarí</collectingRegion>
.
<collectingRegion box="[502,590,1333,1355]" country="Paraguay" name="Guaira" pageId="15" pageNumber="988">Guairá</collectingRegion>
:
<collectorName box="[603,733,1333,1355]" pageId="15" pageNumber="988">Mbocayaty</collectorName>
del
<collectingRegion box="[163,251,1364,1386]" country="Paraguay" name="Guaira" pageId="15" pageNumber="988">Guairá</collectingRegion>
,
<collectorName box="[266,387,1364,1386]" pageId="15" pageNumber="988">Villarrica</collectorName>
;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047368" box="[402,773,1364,1386]" lastPageId="15" lastPageNumber="988" pageId="15" pageNumber="987">
<collectingRegion box="[402,487,1364,1386]" country="Paraguay" name="Itapua" pageId="15" pageNumber="988">Itapuá</collectingRegion>
: San Cosme y Damián
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="4016047397" pageId="15" pageNumber="988">
<collectingCountry box="[163,326,1394,1415]" name="Argentina" pageId="15" pageNumber="988">ARGENTINA</collectingCountry>
:
<collectingRegion box="[337,440,1395,1416]" country="Argentina" name="Formosa" pageId="15" pageNumber="988">Formosa</collectingRegion>
: Formosa. Chaco:
<collectorName box="[652,778,1394,1416]" pageId="15" pageNumber="988">Chacabuco</collectorName>
(
<collectorName box="[171,269,1425,1447]" pageId="15" pageNumber="988">Charata</collectorName>
),
<collectorName box="[289,394,1425,1447]" pageId="15" pageNumber="988">Liberdad</collectorName>
(
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC0FFFF33FA2089AD908FA54" box="[410,554,1425,1447]" country="Argentina" name="Puerto Tirol" pageId="15" pageNumber="988" stateProvince="Corrientes">Puerto Tirol</location>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047382" box="[574,699,1425,1447]" pageId="15" pageNumber="988">
<collectingRegion box="[574,699,1425,1447]" country="Argentina" name="Corrientes" pageId="15" pageNumber="988">Corrientes</collectingRegion>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047338" pageId="15" pageNumber="988">
<collectingRegion country="Argentina" name="Santa Fe" pageId="15" pageNumber="988">Santa Fe</collectingRegion>
:
<collectorName box="[203,405,1456,1478]" pageId="15" pageNumber="988">General Obligado</collectorName>
(
<collectorName box="[420,530,1456,1478]" pageId="15" pageNumber="988">Villa Ana</collectorName>
),
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC0FFFF33C1E08BBD95DFA35" box="[550,639,1456,1478]" country="Argentina" name="Rosario" pageId="15" pageNumber="988" stateProvince="Entre Rios">Rosario</location>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047304" pageId="15" pageNumber="988">
<collectingRegion box="[650,775,1456,1478]" country="Argentina" name="Entre Rios" pageId="15" pageNumber="988">Entre Ríos</collectingRegion>
:
<collectingCountry box="[163,271,1487,1508]" name="Uruguay" pageId="15" pageNumber="988">Uruguay</collectingCountry>
(
<collectorName box="[290,508,1486,1508]" pageId="15" pageNumber="988">Pronunciamiento</collectorName>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047318" box="[532,773,1486,1508]" pageId="15" pageNumber="988">
<collectorName box="[532,645,1486,1508]" pageId="15" pageNumber="988">Misiones</collectorName>
:
<collectorName box="[660,773,1486,1508]" pageId="15" pageNumber="988">Eldorado</collectorName>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047345" box="[163,302,1516,1539]" pageId="15" pageNumber="988">Córdoba [?]</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047353" box="[318,524,1516,1539]" pageId="15" pageNumber="988">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC0FFFF33F0608E6DAFDF9F0" box="[318,479,1517,1539]" country="Uruguay" name="Buenos Aires" pageId="15" pageNumber="988">Buenos Aires</location>
[?].
</materialsCitation>
<materialsCitation box="[534,773,1517,1539]" pageId="15" pageNumber="988">
<collectingCountry box="[534,671,1517,1538]" name="Uruguay" pageId="15" pageNumber="988">URUGUAY</collectingCountry>
:
<collectingRegion box="[682,773,1517,1539]" country="Uruguay" name="Artigas" pageId="15" pageNumber="988">Artigas</collectingRegion>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047373" box="[163,301,1548,1570]" pageId="15" pageNumber="988">
<collectingRegion box="[163,225,1548,1570]" country="Uruguay" name="Salto" pageId="15" pageNumber="988">Salto</collectingRegion>
:
<collectorName box="[234,301,1548,1570]" pageId="15" pageNumber="988">Belén</collectorName>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047351" box="[311,515,1548,1570]" pageId="15" pageNumber="988">
<collectingRegion box="[311,390,1548,1570]" country="Uruguay" name="Rivera" pageId="15" pageNumber="988">Rivera</collectingRegion>
:
<collectorName box="[398,515,1548,1570]" pageId="15" pageNumber="988">Vichadero</collectorName>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047347" box="[524,775,1548,1570]" pageId="15" pageNumber="988">
<collectingRegion box="[524,667,1548,1570]" country="Uruguay" name="Tacuarembo" pageId="15" pageNumber="988">Tacuarembó</collectingRegion>
:
<collectorName box="[677,775,1548,1569]" pageId="15" pageNumber="988">Zapucay</collectorName>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047329" box="[163,400,1578,1600]" pageId="15" pageNumber="988">
<collectingRegion box="[163,278,1578,1600]" country="Uruguay" name="Paysandu" pageId="15" pageNumber="988">Paysandú</collectingRegion>
: Paysandú
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047322" box="[409,547,1578,1600]" pageId="15" pageNumber="988">
<collectingRegion box="[409,547,1578,1600]" country="Uruguay" name="Cerro Largo" pageId="15" pageNumber="988">Cerro Largo</collectingRegion>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047309" box="[556,645,1578,1600]" pageId="15" pageNumber="988">
<collectingRegion box="[556,645,1578,1600]" country="Uruguay" name="Colonia" pageId="15" pageNumber="988">Colonia</collectingRegion>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047370" pageId="15" pageNumber="988">
<collectingRegion box="[654,775,1578,1600]" country="Uruguay" name="Canelones" pageId="15" pageNumber="988">Canelones</collectingRegion>
:
<collectorName box="[163,327,1609,1631]" pageId="15" pageNumber="988">Canelón Chico</collectorName>
,
<collectingRegion box="[334,452,1609,1631]" country="Uruguay" name="Canelones" pageId="15" pageNumber="988">Canelones</collectingRegion>
(
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC0FFFF33FE80B42D909F9AC" box="[464,555,1609,1631]" country="Uruguay" name="Juanico" pageId="15" pageNumber="988" stateProvince="Canelones">Juanicó</location>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047371" pageId="15" pageNumber="988">
<collectingRegion box="[571,707,1609,1631]" country="Uruguay" name="Montevideo" pageId="15" pageNumber="988">Montevideo</collectingRegion>
:
<collectorName box="[714,779,1609,1630]" pageId="15" pageNumber="988">Cerro</collectorName>
de
<collectingRegion box="[196,331,1640,1662]" country="Uruguay" name="Montevideo" pageId="15" pageNumber="988">Montevideo</collectingRegion>
, Playa Punta Yeguas.
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047346" pageId="15" pageNumber="988">
<emphasis bold="true" pageId="15" pageNumber="988">Blue and green specimens:</emphasis>
<collectingCountry box="[318,434,1670,1691]" name="Bolivia" pageId="15" pageNumber="988">BOLIVIA</collectingCountry>
:
<collectingRegion box="[446,587,1670,1691]" country="Bolivia" name="Santa Cruz" pageId="15" pageNumber="988">Santa Cruz</collectingRegion>
:
<collectorName box="[598,773,1670,1692]" pageId="15" pageNumber="988">Andrés Ibáñez</collectorName>
,
<collectorName box="[163,279,1701,1723]" pageId="15" pageNumber="988">Cordillera</collectorName>
(
<collectorName box="[293,376,1701,1723]" pageId="15" pageNumber="988">Camiri</collectorName>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC0FFFF33FB80BAEDADBF948" box="[384,505,1701,1723]" country="Bolivia" name="Tatarenda" pageId="15" pageNumber="988" stateProvince="Tarija">Tatarenda</location>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047355" pageId="15" pageNumber="988">
<collectingRegion box="[523,595,1701,1723]" country="Bolivia" name="Tarija" pageId="15" pageNumber="988">Tarija</collectingRegion>
: Gran
<collectingCounty box="[669,739,1701,1723]" pageId="15" pageNumber="988">Chaco</collectingCounty>
(
<collectorName pageId="15" pageNumber="988">El Bagual</collectorName>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047380" box="[255,468,1731,1753]" pageId="15" pageNumber="988">
Yacuiba:
<collectingMunicipality box="[355,468,1732,1753]" pageId="15" pageNumber="988">Yaguacua</collectingMunicipality>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047321" box="[476,629,1731,1753]" pageId="15" pageNumber="988">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC0FFFF33FE40BC8D94BF92A" box="[476,617,1731,1753]" country="Bolivia" county="Chaco" municipality="Yaguacua" name="Villamontes" pageId="15" pageNumber="988" stateProvince="Tarija">Villamontes</location>
).
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047394" pageId="15" pageNumber="988">
<collectingCountry box="[634,776,1732,1753]" name="Paraguay" pageId="15" pageNumber="988">PARAGUAY</collectingCountry>
: [unspecified department]:
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC0FFFF33FDB0BE9D9A6F90B" box="[483,644,1762,1784]" country="Paraguay" name="Trans-Chaco'" pageId="15" pageNumber="988" stateProvince="Boqueron">Trans-Chaco</location>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047327" pageId="15" pageNumber="988">
<collectingRegion box="[658,775,1762,1784]" country="Paraguay" name="Boqueron" pageId="15" pageNumber="988">Boquerón</collectingRegion>
:
<collectorName box="[163,289,1793,1815]" pageId="15" pageNumber="988">Filadelfia</collectorName>
,
<collectorName box="[306,529,1793,1815]" pageId="15" pageNumber="988">Fortín Guachalla</collectorName>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047376" pageId="15" pageNumber="988">
<collectingRegion box="[545,773,1793,1815]" country="Paraguay" name="Presidente Hayes" pageId="15" pageNumber="988">Presidente Hayes</collectingRegion>
:
<collectorName box="[163,232,1824,1845]" pageId="15" pageNumber="988">Cruce</collectorName>
de los
<collectorName box="[316,419,1823,1845]" pageId="15" pageNumber="988">Pioneros</collectorName>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC0FFFF33F880A2BD97BF8C6" box="[432,601,1823,1845]" country="Paraguay" name="Pozo Colorado" pageId="15" pageNumber="988" stateProvince="Presidente Hayes">Pozo Colorado</location>
.
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047326" pageId="15" pageNumber="988">
<collectingCountry box="[612,775,1824,1845]" name="Argentina" pageId="15" pageNumber="988">ARGENTINA</collectingCountry>
:
<collectingRegion box="[163,229,1854,1876]" country="Argentina" name="Jujuy" pageId="15" pageNumber="988">Jujuy</collectingRegion>
.
<collectingRegion box="[242,308,1854,1876]" country="Argentina" name="Salta" pageId="15" pageNumber="988">Salta</collectingRegion>
:
<collectorName box="[320,408,1854,1876]" pageId="15" pageNumber="988">Capital</collectorName>
(
<collectingRegion box="[426,491,1854,1876]" country="Argentina" name="Salta" pageId="15" pageNumber="988">Salta</collectingRegion>
),
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047312" box="[514,778,1854,1876]" pageId="15" pageNumber="988">
<collectorName box="[514,618,1854,1876]" pageId="15" pageNumber="988">Cerrillos</collectorName>
(
<collectorName box="[635,766,1854,1876]" pageId="15" pageNumber="988">La Merced</collectorName>
),
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047393" box="[163,276,1885,1907]" pageId="15" pageNumber="988">
<collectingMunicipality box="[163,272,1885,1907]" pageId="15" pageNumber="988">Chicoana</collectingMunicipality>
,
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047367" box="[283,1059,197,1907]" pageId="15" pageNumber="988">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC0FFFF33F230A56DA93F880" box="[283,433,1885,1907]" country="Argentina" municipality="Chicoana" name="General Jose" pageId="15" pageNumber="988" stateProvince="Salta">General José</location>
<collectorName box="[440,607,1885,1907]" pageId="15" pageNumber="988">de San Martín</collectorName>
(
<collectorName box="[622,775,1885,1907]" pageId="15" pageNumber="988">Embarcación</collectorName>
, Salvador Mazza),
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047366" box="[1073,1179,197,219]" pageId="15" pageNumber="988">
<collectorName box="[1073,1179,197,219]" pageId="15" pageNumber="988">La Viña</collectorName>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047388" pageId="15" pageNumber="988">
<collectingRegion box="[1202,1280,197,219]" country="Argentina" name="Chaco" pageId="15" pageNumber="988">Chaco</collectingRegion>
: Chacabuco (Charata)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047325" pageId="15" pageNumber="988">
Misiones: Capital (Posadas), Eldorado (9
<collectorName pageId="15" pageNumber="988">de Julio</collectorName>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047396" box="[906,1437,258,280]" pageId="15" pageNumber="988">
<collectingRegion box="[906,1032,258,280]" country="Argentina" name="Corrientes" pageId="15" pageNumber="988">Corrientes</collectingRegion>
:
<collectorName box="[1043,1185,258,280]" pageId="15" pageNumber="988">General Paz</collectorName>
(
<collectorName box="[1201,1305,258,280]" pageId="15" pageNumber="988">Itá Ibaté</collectorName>
),
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC0FFFF33B150C09DEBFFEEB" box="[1325,1437,258,280]" country="Argentina" name="Ituzaingo" pageId="15" pageNumber="988" stateProvince="Entre Rios">Ituzaingó</location>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047363" box="[827,1288,289,311]" pageId="15" pageNumber="988">
<collectingRegion box="[827,952,289,311]" country="Argentina" name="Entre Rios" pageId="15" pageNumber="988">Entre Ríos</collectingRegion>
:
<collectingCountry box="[963,1065,290,311]" name="Uruguay" pageId="15" pageNumber="988">Uruguay</collectingCountry>
(Pronunciamiento)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047369" pageId="15" pageNumber="988">
<collectingCounty pageId="15" pageNumber="988">Santiago del Estero</collectingCounty>
: Alberdi (Campo Gallo),
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047357" pageId="15" pageNumber="988">
Avellaneda (
<collectingRegion box="[1351,1442,320,342]" country="Uruguay" name="Colonia" pageId="15" pageNumber="988">Colonia</collectingRegion>
Dora, Icaño),
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047328" box="[991,1219,350,372]" pageId="15" pageNumber="988">
Banda (
<collectorName box="[1085,1206,350,372]" pageId="15" pageNumber="988">La Banda</collectorName>
),
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047308" box="[1228,1358,351,372]" pageId="15" pageNumber="988">Barrancas,</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047342" pageId="15" pageNumber="988">
Choya (Frías,
<collectingMunicipality box="[911,1081,381,403]" pageId="15" pageNumber="988">Villa la Punta</collectingMunicipality>
),
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047385" box="[1103,1351,381,403]" pageId="15" pageNumber="988">
Jiménez (
<collectorName box="[1217,1340,381,403]" pageId="15" pageNumber="988">El Charco</collectorName>
),
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047389" pageId="15" pageNumber="988">Loreto, Mitre (Villa Unión),</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047390" box="[1084,1437,410,434]" pageId="15" pageNumber="988">
Moreno (Puna),
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC0FFFF33B3C0C97DEBFFE41" box="[1284,1437,410,434]" country="Uruguay" county="Santiago del Estero" municipality="Villa la Punta" name="Ojo de Agua" pageId="15" pageNumber="988" stateProvince="Colonia">
Ojo
<collectorName box="[1335,1437,410,434]" pageId="15" pageNumber="988">de Água</collectorName>
</location>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047386" pageId="15" pageNumber="988">Tucumán: Burruyacú (Burruyacú), Cerro San Javier, Trancas (Tapia)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047356" box="[1025,1160,473,495]" pageId="15" pageNumber="988">Catamarca</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047302" pageId="15" pageNumber="988">
<collectorName box="[1174,1279,473,495]" pageId="15" pageNumber="988">La Rioja</collectorName>
: General San Martín (Ulapes),
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047365" box="[1042,1437,504,526]" pageId="15" pageNumber="988">Rosario Vera Peñaloza (Chepes)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047314" box="[827,1195,534,556]" pageId="15" pageNumber="988">San Juan: Iglesia (Angualasto)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047372" box="[1208,1437,534,556]" pageId="15" pageNumber="988">San Luis: Belgrano</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047391" box="[827,1442,565,587]" pageId="15" pageNumber="988">
Córdoba: Calamuchita (
<collectingCounty box="[1093,1431,565,587]" pageId="15" pageNumber="988">Villa Yacanto de Calamuchita</collectingCounty>
),
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047313" box="[827,1198,596,618]" pageId="15" pageNumber="988">Ischilín (Copacabana, Quilino),</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047361" pageId="15" pageNumber="988">Jaime Pérez (Jaime Peter),</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047349" box="[915,1442,626,648]" pageId="15" pageNumber="988">
Punilla (
<collectingMunicipality box="[1020,1239,626,648]" pageId="15" pageNumber="988">Capilla del Monte</collectingMunicipality>
, Valle Hermoso),
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047340" box="[827,1441,657,679]" pageId="15" pageNumber="988">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC0FFFF33D030F9AD8E6FD55" box="[827,964,657,679]" country="Uruguay" county="Villa Yacanto de Calamuchita" municipality="Capilla del Monte" name="Rio Cuarto" pageId="15" pageNumber="988" stateProvince="Santa Fe">Río Cuarto</location>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC0FFFF33DEB0F9ADF1DFD55" box="[979,1087,657,679]" country="Uruguay" county="Villa Yacanto de Calamuchita" municipality="Capilla del Monte" name="Rio Seco" pageId="15" pageNumber="988" stateProvince="Santa Fe">Río Seco</location>
, San Alberto (Las Rabonas),
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047320" box="[827,1343,688,710]" pageId="15" pageNumber="988">
Sobremonte (
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC0FFFF33DDA0FBBDE13FD35" box="[994,1329,688,710]" country="Uruguay" county="Villa Yacanto de Calamuchita" municipality="Capilla del Monte" name="San Francisco del Chanar" pageId="15" pageNumber="988" stateProvince="Santa Fe">San Francisco del Chañar</location>
),
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047379" pageId="15" pageNumber="988">
Totoral (
<collectorName box="[834,957,718,740]" pageId="15" pageNumber="988">La Pampa</collectorName>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047336" box="[978,1438,718,740]" pageId="15" pageNumber="988">
<collectingRegion box="[978,1086,718,740]" inferredBySuffix="true" pageId="15" pageNumber="988">Santa Fe</collectingRegion>
:
<collectorName box="[1098,1300,718,740]" pageId="15" pageNumber="988">General Obligato</collectorName>
(
<collectorName box="[1316,1430,718,740]" pageId="15" pageNumber="988">Villa Ana</collectorName>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047339" pageId="15" pageNumber="988">
<collectingCounty box="[827,1070,749,771]" pageId="15" pageNumber="988">Mendoza. La Pampa</collectingCounty>
:
<collectingMunicipality box="[1082,1228,749,771]" pageId="15" pageNumber="988">Lihuel Calel</collectingMunicipality>
(
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC0FFFF33AE50FE6D8EBFCD1" country="Uruguay" county="Mendoza. La Pampa" municipality="Lihuel Calel" name="Parque Nacional Lihue Calel" pageId="15" pageNumber="988" stateProvince="Santa Fe">Parque Nacional Lihué Calel</location>
).
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047395" pageId="15" pageNumber="988">
<emphasis bold="true" box="[988,1281,779,801]" pageId="15" pageNumber="988">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC0FFFF33DE40E00DF4FFCD2" box="[988,1133,779,801]" country="Uruguay" county="Mendoza. La Pampa" municipality="Lihuel Calel" name="Bicoloured" pageId="15" pageNumber="988" stateProvince="Santa Fe">Bicoloured</location>
specimens
</emphasis>
<emphasis bold="true" box="[1281,1286,779,801]" pageId="15" pageNumber="988">:</emphasis>
<collectingCountry box="[1295,1440,780,801]" name="Paraguay" pageId="15" pageNumber="988">PARAGUAY</collectingCountry>
: [unspecified department]:
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC0FFFF33A4F0E21DE35FCB3" box="[1143,1303,810,832]" country="Paraguay" name="Trans-Chaco'" pageId="15" pageNumber="988" stateProvince="Presidente Hayes">Trans-Chaco</location>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047319" pageId="15" pageNumber="988">
<collectingRegion country="Paraguay" name="Presidente Hayes" pageId="15" pageNumber="988">Presidente Hayes</collectingRegion>
:
<collectorName box="[917,987,841,862]" pageId="15" pageNumber="988">Cruce</collectorName>
de los
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC0FFFF33A0B0E42DFBFFCAC" box="[1075,1181,841,863]" country="Paraguay" name="Pioneros" pageId="15" pageNumber="988" stateProvince="Presidente Hayes">Pioneros</location>
.
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047332" pageId="15" pageNumber="988">
<collectingCountry box="[1194,1361,841,862]" name="Argentina" pageId="15" pageNumber="988">ARGENTINA</collectingCountry>
:
<collectingRegion box="[1373,1438,841,863]" country="Argentina" name="Salta" pageId="15" pageNumber="988">Salta</collectingRegion>
:
<collectorName box="[827,935,872,894]" pageId="15" pageNumber="988">La Viña.</collectorName>
<collectingRegion box="[945,1064,872,894]" country="Argentina" name="Tucuman" pageId="15" pageNumber="988">Tucumán</collectingRegion>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047317" pageId="15" pageNumber="988">
<collectingRegion box="[1080,1336,872,894]" country="Argentina" name="Santiago del Estero" pageId="15" pageNumber="988">Santiago del Estero</collectingRegion>
:
<collectorName box="[1350,1441,872,894]" pageId="15" pageNumber="988">Alberdi</collectorName>
(
<collectorName box="[835,1001,902,924]" pageId="15" pageNumber="988">Campo Gallo</collectorName>
), Choya (
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC0FFFF33A5B0E8DDF85FC6F" box="[1123,1191,902,924]" country="Argentina" name="Frias" pageId="15" pageNumber="988" stateProvince="Chaco">Frías</location>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047333" pageId="15" pageNumber="988">
<collectingRegion box="[1216,1294,902,924]" country="Argentina" name="Chaco" pageId="15" pageNumber="988">Chaco</collectingRegion>
: Chacabuco (Charata)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047306" pageId="15" pageNumber="988">
Misiones: Cainguás (
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC0FFFF33ADD0EAEDEB6FC49" box="[1253,1428,933,955]" country="Argentina" name="Dos de Mayo" pageId="15" pageNumber="988" stateProvince="Corrientes">
Dos
<collectorName box="[1315,1428,933,955]" pageId="15" pageNumber="988">de Mayo</collectorName>
</location>
), Eldorado
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047384" box="[951,1209,964,986]" pageId="15" pageNumber="988">
<collectingRegion box="[951,1081,964,986]" country="Argentina" name="Corrientes" pageId="15" pageNumber="988">Corrientes</collectingRegion>
:
<collectorName box="[1093,1209,964,986]" pageId="15" pageNumber="988">Ituzaingó</collectorName>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047387" pageId="15" pageNumber="988">
<collectingRegion box="[1223,1334,964,985]" country="Argentina" name="Santa Fe" pageId="15" pageNumber="988">Santa Fe</collectingRegion>
:
<collectorName pageId="15" pageNumber="988">General Obligado</collectorName>
(
<collectorName box="[948,1063,994,1016]" pageId="15" pageNumber="988">Villa Ana</collectorName>
),
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047359" box="[1084,1296,994,1016]" pageId="15" pageNumber="988">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC0FFFF33A040EE9DFB5FC0B" box="[1084,1175,994,1016]" country="Argentina" name="Rosario" pageId="15" pageNumber="988" stateProvince="Santa Fe">Rosario</location>
,
<collectingRegion box="[1188,1296,994,1016]" country="Argentina" name="Santa Fe" pageId="15" pageNumber="988">Santa Fe</collectingRegion>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047392" pageId="15" pageNumber="988">
<collectingRegion box="[1309,1439,994,1016]" country="Argentina" name="Entre Rios" pageId="15" pageNumber="988">Entre Ríos</collectingRegion>
:
<collectingCountry box="[827,932,1025,1046]" name="Uruguay" pageId="15" pageNumber="988">Uruguay</collectingCountry>
(Pronunciamiento)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047344" pageId="15" pageNumber="988">Córdoba: Jaime Pérez (Jaime Peter),</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047323" box="[1000,1267,1056,1078]" pageId="15" pageNumber="988">Punilla (San Esteban),</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047352" pageId="15" pageNumber="988">
Tulumba (
<collectingRegion country="Uruguay" name="San Jose" pageId="15" pageNumber="988">San José</collectingRegion>
<collectorName box="[891,1074,1086,1108]" pageId="15" pageNumber="988">de la Dormida</collectorName>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047350" pageId="15" pageNumber="988">Buenos Aires: Puan (Felipe Solá),</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047360" box="[914,1332,1117,1139]" pageId="15" pageNumber="988">
San Isidro (
<collectingMunicipality box="[1074,1323,1117,1139]" pageId="15" pageNumber="988">Boulogne Sur Mer</collectingMunicipality>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047303" pageId="15" pageNumber="988">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC0FFFF33B7E0956DFBFFB61" country="Uruguay" municipality="Boulogne Sur Mer" name="Ciudad Autonoma de Buenos Aires" pageId="15" pageNumber="988" stateProvince="San Jose">Ciudad Autónoma de Buenos Aires</location>
: Núñez.
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047331" pageId="15" pageNumber="988">
<collectingCountry box="[1298,1438,1148,1169]" name="Uruguay" pageId="15" pageNumber="988">URUGUAY</collectingCountry>
:
<collectingRegion box="[827,917,1178,1200]" country="Uruguay" name="Artigas" pageId="15" pageNumber="988">Artigas</collectingRegion>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047337" box="[929,1073,1178,1200]" pageId="15" pageNumber="988">
<collectingRegion box="[929,1069,1178,1200]" country="Uruguay" name="Montevideo" pageId="15" pageNumber="988">Montevideo</collectingRegion>
.
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047301" pageId="15" pageNumber="988">
<emphasis bold="true" box="[1081,1325,1178,1200]" pageId="15" pageNumber="988">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC0FFFF33A010991DFB8FB43" box="[1081,1178,1178,1200]" country="Uruguay" name="Copper" pageId="15" pageNumber="988" stateProvince="Artigas">Copper</location>
specimens
</emphasis>
<emphasis bold="true" box="[1325,1331,1178,1200]" pageId="15" pageNumber="988">:</emphasis>
<collectingCountry box="[1339,1439,1178,1199]" name="Brazil" pageId="15" pageNumber="988">BRAZIL</collectingCountry>
:
<collectingRegion box="[827,1047,1209,1231]" country="Brazil" name="Rio Grande do Sul" pageId="15" pageNumber="988">Rio Grande do Sul</collectingRegion>
:
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC0FFFF33A1B09B2DF5AFB3C" box="[1059,1144,1209,1231]" country="Brazil" name="Guarai" pageId="15" pageNumber="988" stateProvince="Rio Grande do Sul">Guaraí</location>
.
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047362" pageId="15" pageNumber="988">
<collectingCountry box="[1156,1320,1209,1230]" name="Argentina" pageId="15" pageNumber="988">ARGENTINA</collectingCountry>
:
<collectingRegion box="[1331,1439,1209,1231]" country="Argentina" name="Mendoza" pageId="15" pageNumber="988">Mendoza</collectingRegion>
.
<collectingRegion box="[827,944,1239,1261]" country="Argentina" name="San Luis" pageId="15" pageNumber="988">San Luis</collectingRegion>
: Belgrano
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047335" box="[1092,1441,1239,1261]" pageId="15" pageNumber="988">Córdoba: Ischilín (Quilino),</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047307" box="[827,1083,1270,1292]" pageId="15" pageNumber="988">Punilla (San Esteban)</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047354" box="[1095,1322,1270,1292]" pageId="15" pageNumber="988">
<collectingRegion box="[1095,1322,1270,1292]" country="Argentina" name="Santiago del Estero" pageId="15" pageNumber="988">Santiago del Estero</collectingRegion>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047305" pageId="15" pageNumber="988">
<collectingRegion box="[1334,1439,1270,1292]" country="Argentina" name="Santa Fe" pageId="15" pageNumber="988">Santa Fe</collectingRegion>
:
<collectorName box="[827,916,1301,1323]" pageId="15" pageNumber="988">Rosario</collectorName>
,
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047378" box="[927,1209,1301,1323]" pageId="15" pageNumber="988">
<collectorName box="[927,1085,1301,1323]" pageId="15" pageNumber="988">San Cristóbal</collectorName>
(Hersilia),
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047341" box="[1217,1322,1301,1322]" pageId="15" pageNumber="988">
<collectingRegion box="[1217,1322,1301,1322]" country="Argentina" name="Santa Fe" pageId="15" pageNumber="988">Santa Fe</collectingRegion>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047310" pageId="15" pageNumber="988">
<collectorName box="[1334,1439,1301,1323]" pageId="15" pageNumber="988">Misiones</collectorName>
:
<collectorName box="[827,938,1331,1353]" pageId="15" pageNumber="988">Eldorado</collectorName>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047334" box="[951,1437,1331,1354]" pageId="15" pageNumber="988">
<collectingRegion box="[951,1084,1332,1354]" country="Argentina" name="Entre Rios" pageId="15" pageNumber="988">Entre Ríos</collectingRegion>
:
<collectingCountry box="[1096,1201,1332,1353]" name="Uruguay" pageId="15" pageNumber="988">Uruguay</collectingCountry>
(Pronunciamiento)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047383" box="[827,1259,1362,1384]" pageId="15" pageNumber="988">
<collectingMunicipality box="[827,1259,1362,1384]" pageId="15" pageNumber="988">Ciudad Autónoma de Buenos Aires</collectingMunicipality>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047381" pageId="15" pageNumber="988">
Buenos Aires:
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC0FFFF33D03087AD8F5FA74" box="[827,983,1393,1415]" country="Uruguay" municipality="Ciudad Autonoma de Buenos Aires" name="Bahia Blanca" pageId="15" pageNumber="988">Bahía Blanca</location>
,
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047358" box="[994,1220,1393,1415]" pageId="15" pageNumber="988">Merlo (Pontevedra),</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4016047316" pageId="15" pageNumber="988">Puan (Felipe Solá), San Nicolás.</materialsCitation>
<materialsCitation box="[985,1223,1423,1445]" pageId="15" pageNumber="988">
<collectingCountry box="[985,1121,1424,1445]" name="Uruguay" pageId="15" pageNumber="988">URUGUAY</collectingCountry>
:
<collectingRegion box="[1132,1223,1423,1445]" country="Uruguay" name="Artigas" pageId="15" pageNumber="988">Artigas</collectingRegion>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047375" box="[1236,1383,1423,1445]" pageId="15" pageNumber="988">
<collectingRegion box="[1236,1301,1423,1445]" country="Uruguay" name="Salto" pageId="15" pageNumber="988">Salto</collectingRegion>
:
<collectorName box="[1313,1383,1423,1445]" pageId="15" pageNumber="988">Belén</collectorName>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047324" pageId="15" pageNumber="988">
<collectingRegion country="Uruguay" name="San Jose" pageId="15" pageNumber="988">San José</collectingRegion>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="4016047343" box="[889,1277,1454,1476]" pageId="15" pageNumber="988">
<collectingRegion box="[889,1025,1454,1476]" country="Uruguay" name="Montevideo" pageId="15" pageNumber="988">Montevideo</collectingRegion>
:
<collectorName box="[1035,1100,1454,1475]" pageId="15" pageNumber="988">Cerro</collectorName>
de
<collectingRegion box="[1141,1277,1454,1476]" country="Uruguay" name="Montevideo" pageId="15" pageNumber="988">Montevideo</collectingRegion>
</materialsCitation>
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7552004" ID-Zenodo-Dep="7552004" httpUri="https://zenodo.org/record/7552004/files/figure.png" pageId="15" pageNumber="988" startId="15.[163,241,1105,1127]" targetBox="[181,784,202,1062]" targetPageId="15">
<paragraph blockId="15.[163,780,1105,1245]" pageId="15" pageNumber="988">
<emphasis bold="true" box="[163,266,1105,1127]" pageId="15" pageNumber="988">Figure 8.</emphasis>
Genitalia of both sexes of
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[545,665,1105,1127]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="988" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[545,665,1105,1127]" italics="true" pageId="15" pageNumber="988">B. onitoides</emphasis>
</taxonomicName>
. A, female: spermatheca. BD, male: B, genital segment (i.e. nineth abdominal segment). C, D, tegmen. C, dorsal view. D, lateral view. LSP = lateral sclerotized plate; MSP = medial sclerotized plate.
</paragraph>
</caption>
<paragraph blockId="15.[827,1443,197,1476]" box="[1277,1281,1454,1476]" pageId="15" pageNumber="988">.</paragraph>
<paragraph blockId="15.[827,1443,1517,1907]" lastBlockId="17.[163,779,197,1875]" lastPageId="17" lastPageNumber="990" pageId="15" pageNumber="988">
<emphasis box="[827,1111,1517,1538]" italics="true" pageId="15" pageNumber="988">Intraspecific variation:</emphasis>
Besides colour, which will be discussed below, the most noticeable variation observed among individuals of
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[1196,1332,1578,1600]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="988" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[1196,1332,1578,1600]" italics="true" pageId="15" pageNumber="988">B. onitoides</emphasis>
</taxonomicName>
concerns the pronotal punctation. This is most evident over the posterior region of the pronotum near the posterior fossae, where punctures range from being dense and almost contiguous with one another to effaced and sparse; sometimes, the region between the posterior fossae lacks any punctures altogether. Although the latter condition seems to be more common in the eastern part of the distribution, populations from the entire geographical range of the species show individuals varying across the full variation spectrum. Other external characters that show some relevant degree of variation are the apical angle of the anteromedian prominence of the metaventrite, which may be more or less projected forwards and pointed depending on the specimen, and the setation at the apex of the metacoxae, which varies from dense to absent in both sexes (though it tends to be denser and longer in females than in males). Even though a strongly acuminate metaventral angle and a denser metacoxal setation are more commonly observed in western individuals, neither of these characters varies consistently with either distribution or dorsal colour variant, and specimens from all populations may show either extreme of variation.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7552012" ID-Zenodo-Dep="7552012" httpUri="https://zenodo.org/record/7552012/files/figure.png" pageId="16" pageNumber="989" startId="16.[145,225,1732,1754]" targetBox="[305,1265,198,1689]" targetPageId="16">
<paragraph blockId="16.[145,1426,1732,1872]" pageId="16" pageNumber="989">
<emphasis bold="true" box="[145,252,1732,1754]" pageId="16" pageNumber="989">Figure 9.</emphasis>
Male genitalia: Endophallus of
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[605,730,1732,1754]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="989" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[605,730,1732,1754]" italics="true" pageId="16" pageNumber="989">B. onitoides</emphasis>
</taxonomicName>
. A, B, general view of the extracted endophallus. A, dorsal view. B, ventral view. CG, endophallites: C, axial + subaxial endophallite complex (dissected). D, fronto-lateral peripheral endophallite (ventral view). E, medial endophallite (dorsal view). FH, superior right peripheral endophallite from different views: F, G, ventral view: F, seahorse-shaped; G, F-shaped. H,
<emphasis bold="true" box="[773,781,1820,1841]" pageId="16" pageNumber="989">l</emphasis>
ateral view. A = axial endophallite; SA = subaxial endophallite; MS = medial endophallite; FLP = fronto-lateral peripheral endophallite; SRP = superior right peripheral endophallite.
</paragraph>
</caption>
<paragraph blockId="17.[163,779,197,1875]" pageId="17" pageNumber="990">
In regard to genital structures, the endophallus of
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[193,332,657,679]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="990" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[193,332,657,679]" italics="true" pageId="17" pageNumber="990">B. onitoides</emphasis>
</taxonomicName>
has one of the most simplified set of endophallites found among the phanaeines, lacking the accessory endophallite that is present in some species of the clade [e.g.
<taxonomicName authority="(Erichson, 1847)" baseAuthorityName="Erichson" baseAuthorityYear="1847" box="[367,773,749,771]" class="Insecta" family="Scarabaeidae" genus="Oruscatus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="990" phylum="Arthropoda" rank="species" species="davus">
<emphasis box="[367,567,749,770]" italics="true" pageId="17" pageNumber="990">Oruscatus davus</emphasis>
(Erichson, 1847)
</taxonomicName>
,
<taxonomicName baseAuthorityName="Perty" baseAuthorityYear="1830" box="[163,430,780,801]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="990" phylum="Arthropoda" rank="species" species="aeruginosa">
<emphasis box="[163,430,780,801]" italics="true" pageId="17" pageNumber="990">Gromphas aeruginosa</emphasis>
</taxonomicName>
and
<taxonomicName authority="(Fabricius, 1781)" authorityName="Pronotum" baseAuthorityName="Fabricius" baseAuthorityYear="1781" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="990" phylum="Arthropoda" rank="species" species="splendidulus">
<emphasis box="[495,778,780,801]" italics="true" pageId="17" pageNumber="990">Phanaeus splendidulus</emphasis>
(Fabricius, 1781)
</taxonomicName>
;
<bibRefCitation author="Tarasov S &amp; Genier F" box="[362,626,810,832]" pageId="17" pageNumber="990" pagination="1 - 86" refId="ref54817" refString="Tarasov S, Genier F. 2015. Innovative bayesian and parsimony phylogeny of dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) enhanced by ontology-based partitioning of morphological characters. PLoS One 10: 1 - 86." type="journal article" year="2015">Tarasov &amp; Génier, 2015</bibRefCitation>
]. But in spite of this simplification, the information stated in one of our previous works (
<bibRefCitation author="Cupello M &amp; Vaz-de-Mello FZ" box="[429,770,872,894]" pageId="17" pageNumber="990" pagination="1 - 27" refId="ref47264" refString="Cupello M, Vaz-de-Mello FZ. 2015. A new species and the phylogeny of the South American genus Gromphas Brulle, 1837 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini). Journal of Natural History (online edition): 1 - 27." type="book chapter" year="2015">Cupello &amp; Vaz-de-Mello, 2015</bibRefCitation>
) that
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[219,353,902,924]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="990" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[219,353,902,924]" italics="true" pageId="17" pageNumber="990">B. onitoides</emphasis>
</taxonomicName>
does not have a medial endophallite (then called medial sclerite) is certainly incorrect, as can be clearly seen in
<figureCitation box="[429,529,964,986]" captionStart="Figure 9" captionStartId="16.[145,225,1732,1754]" captionTargetBox="[305,1265,198,1689]" captionTargetId="figure-25@16.[302,1268,195,1692]" captionTargetPageId="16" captionText="Figure 9. Male genitalia: Endophallus of B. onitoides. A, B, general view of the extracted endophallus. A, dorsal view. B, ventral view. CG, endophallites: C, axial + subaxial endophallite complex (dissected). D, fronto-lateral peripheral endophallite (ventral view). E, medial endophallite (dorsal view). FH, superior right peripheral endophallite from different views: F, G, ventral view: F, seahorse-shaped; G, F-shaped. H, lateral view.A = axial endophallite; SA = subaxial endophallite; MS = medial endophallite; FLP = fronto-lateral peripheral endophallite; SRP = superior right peripheral endophallite." figureDoi="http://doi.org/10.5281/zenodo.7552012" httpUri="https://zenodo.org/record/7552012/files/figure.png" pageId="17" pageNumber="990">Figure 9</figureCitation>
. We do not know the cause of our previous mistake; it is possible that we made wrong notes while dissecting the specimens and the error was repeated in the manuscript. It is worth noting that our observation of the medial endophallite being absent from the two species of
<taxonomicName authority="Bates, 1870" authorityName="Bates" authorityYear="1870" class="Insecta" family="Scarabaeidae" genus="Oruscatus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="990" phylum="Arthropoda" rank="genus">
<emphasis box="[584,700,1117,1138]" italics="true" pageId="17" pageNumber="990">Oruscatus</emphasis>
<bibRefCitation author="Bates HW" pageId="17" pageNumber="990" pagination="173 - 180" refId="ref46167" refString="Bates HW. 1870. On a new genus and some new species of Copridae (Coleoptera-Lamellicornia). Transactions of the Royal Entomological Society of London 18: 173 - 180." type="journal article" year="1870">Bates, 1870</bibRefCitation>
</taxonomicName>
is indeed correct, as can be confirmed in
<bibRefCitation author="Tarasov S &amp; Genier F" pageId="17" pageNumber="990" pagination="1 - 86" refId="ref54817" refString="Tarasov S, Genier F. 2015. Innovative bayesian and parsimony phylogeny of dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) enhanced by ontology-based partitioning of morphological characters. PLoS One 10: 1 - 86." type="journal article" year="2015">Tarasov &amp; Géniers (2015)</bibRefCitation>
illustration of the endophallites of
<taxonomicName baseAuthorityName="Erichson" baseAuthorityYear="1847" box="[163,261,1209,1230]" class="Insecta" family="Scarabaeidae" genus="Oruscatus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="990" phylum="Arthropoda" rank="species" species="davus">
<emphasis box="[163,261,1209,1230]" italics="true" pageId="17" pageNumber="990">O. davus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph blockId="17.[163,779,197,1875]" lastBlockId="17.[827,1443,197,832]" pageId="17" pageNumber="990">
The medial endophallite of
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[509,643,1240,1261]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="990" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[509,643,1240,1261]" italics="true" pageId="17" pageNumber="990">B. onitoides</emphasis>
</taxonomicName>
, despite its overall shape being somewhat conserved among the specimens dissected, shows some minor variations in the development of the lateral projections at its folding area (
<figureCitation box="[318,400,1362,1384]" captionStart="Figure 9" captionStartId="16.[145,225,1732,1754]" captionTargetBox="[305,1265,198,1689]" captionTargetId="figure-25@16.[302,1268,195,1692]" captionTargetPageId="16" captionText="Figure 9. Male genitalia: Endophallus of B. onitoides. A, B, general view of the extracted endophallus. A, dorsal view. B, ventral view. CG, endophallites: C, axial + subaxial endophallite complex (dissected). D, fronto-lateral peripheral endophallite (ventral view). E, medial endophallite (dorsal view). FH, superior right peripheral endophallite from different views: F, G, ventral view: F, seahorse-shaped; G, F-shaped. H, lateral view.A = axial endophallite; SA = subaxial endophallite; MS = medial endophallite; FLP = fronto-lateral peripheral endophallite; SRP = superior right peripheral endophallite." figureDoi="http://doi.org/10.5281/zenodo.7552012" httpUri="https://zenodo.org/record/7552012/files/figure.png" pageId="17" pageNumber="990">Fig. 9E</figureCitation>
). The SRP endophallite, like the medial one, shows some degree of variation, some specimens having one of its extremities bent, giving it a seahorse-like shape in lateral view (
<figureCitation box="[616,696,1454,1476]" captionStart="Figure 9" captionStartId="16.[145,225,1732,1754]" captionTargetBox="[305,1265,198,1689]" captionTargetId="figure-25@16.[302,1268,195,1692]" captionTargetPageId="16" captionText="Figure 9. Male genitalia: Endophallus of B. onitoides. A, B, general view of the extracted endophallus. A, dorsal view. B, ventral view. CG, endophallites: C, axial + subaxial endophallite complex (dissected). D, fronto-lateral peripheral endophallite (ventral view). E, medial endophallite (dorsal view). FH, superior right peripheral endophallite from different views: F, G, ventral view: F, seahorse-shaped; G, F-shaped. H, lateral view.A = axial endophallite; SA = subaxial endophallite; MS = medial endophallite; FLP = fronto-lateral peripheral endophallite; SRP = superior right peripheral endophallite." figureDoi="http://doi.org/10.5281/zenodo.7552012" httpUri="https://zenodo.org/record/7552012/files/figure.png" pageId="17" pageNumber="990">Fig. 9F</figureCitation>
), while others lack such a bend and have a general F-shape in lateral view (
<figureCitation box="[312,393,1516,1538]" captionStart="Figure 9" captionStartId="16.[145,225,1732,1754]" captionTargetBox="[305,1265,198,1689]" captionTargetId="figure-25@16.[302,1268,195,1692]" captionTargetPageId="16" captionText="Figure 9. Male genitalia: Endophallus of B. onitoides. A, B, general view of the extracted endophallus. A, dorsal view. B, ventral view. CG, endophallites: C, axial + subaxial endophallite complex (dissected). D, fronto-lateral peripheral endophallite (ventral view). E, medial endophallite (dorsal view). FH, superior right peripheral endophallite from different views: F, G, ventral view: F, seahorse-shaped; G, F-shaped. H, lateral view.A = axial endophallite; SA = subaxial endophallite; MS = medial endophallite; FLP = fronto-lateral peripheral endophallite; SRP = superior right peripheral endophallite." figureDoi="http://doi.org/10.5281/zenodo.7552012" httpUri="https://zenodo.org/record/7552012/files/figure.png" pageId="17" pageNumber="990">Fig. 9G</figureCitation>
). This variation is correlated with neither population of origin nor dorsal colour. Finally, the A+SA complex (
<figureCitation box="[404,491,1577,1599]" captionStart="Figure 9" captionStartId="16.[145,225,1732,1754]" captionTargetBox="[305,1265,198,1689]" captionTargetId="figure-25@16.[302,1268,195,1692]" captionTargetPageId="16" captionText="Figure 9. Male genitalia: Endophallus of B. onitoides. A, B, general view of the extracted endophallus. A, dorsal view. B, ventral view. CG, endophallites: C, axial + subaxial endophallite complex (dissected). D, fronto-lateral peripheral endophallite (ventral view). E, medial endophallite (dorsal view). FH, superior right peripheral endophallite from different views: F, G, ventral view: F, seahorse-shaped; G, F-shaped. H, lateral view.A = axial endophallite; SA = subaxial endophallite; MS = medial endophallite; FLP = fronto-lateral peripheral endophallite; SRP = superior right peripheral endophallite." figureDoi="http://doi.org/10.5281/zenodo.7552012" httpUri="https://zenodo.org/record/7552012/files/figure.png" pageId="17" pageNumber="990">Fig. 9C</figureCitation>
) and the FLP (
<figureCitation box="[681,770,1577,1599]" captionStart="Figure 9" captionStartId="16.[145,225,1732,1754]" captionTargetBox="[305,1265,198,1689]" captionTargetId="figure-25@16.[302,1268,195,1692]" captionTargetPageId="16" captionText="Figure 9. Male genitalia: Endophallus of B. onitoides. A, B, general view of the extracted endophallus. A, dorsal view. B, ventral view. CG, endophallites: C, axial + subaxial endophallite complex (dissected). D, fronto-lateral peripheral endophallite (ventral view). E, medial endophallite (dorsal view). FH, superior right peripheral endophallite from different views: F, G, ventral view: F, seahorse-shaped; G, F-shaped. H, lateral view.A = axial endophallite; SA = subaxial endophallite; MS = medial endophallite; FLP = fronto-lateral peripheral endophallite; SRP = superior right peripheral endophallite." figureDoi="http://doi.org/10.5281/zenodo.7552012" httpUri="https://zenodo.org/record/7552012/files/figure.png" pageId="17" pageNumber="990">Fig. 9D</figureCitation>
) appear to be the least plastic among the endophallites of
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[191,326,1638,1660]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="990" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[191,326,1638,1660]" italics="true" pageId="17" pageNumber="990">B. onitoides</emphasis>
</taxonomicName>
, the latter being fairly vestigial is this species. The axial endophallite (
<figureCitation box="[527,608,1669,1691]" captionStart="Figure 9" captionStartId="16.[145,225,1732,1754]" captionTargetBox="[305,1265,198,1689]" captionTargetId="figure-25@16.[302,1268,195,1692]" captionTargetPageId="16" captionText="Figure 9. Male genitalia: Endophallus of B. onitoides. A, B, general view of the extracted endophallus. A, dorsal view. B, ventral view. CG, endophallites: C, axial + subaxial endophallite complex (dissected). D, fronto-lateral peripheral endophallite (ventral view). E, medial endophallite (dorsal view). FH, superior right peripheral endophallite from different views: F, G, ventral view: F, seahorse-shaped; G, F-shaped. H, lateral view.A = axial endophallite; SA = subaxial endophallite; MS = medial endophallite; FLP = fronto-lateral peripheral endophallite; SRP = superior right peripheral endophallite." figureDoi="http://doi.org/10.5281/zenodo.7552012" httpUri="https://zenodo.org/record/7552012/files/figure.png" pageId="17" pageNumber="990">Fig. 9C</figureCitation>
), in particular, with one of its ends forked and the other bearing an acuminate prolongation, has a shape reminiscent of that of other phanaeines illustrated in the literature, being particularly similar to that of
<taxonomicName authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Scarabaeidae" genus="Diabroctis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="990" phylum="Arthropoda" rank="species" species="mimas">
<emphasis box="[576,778,1791,1813]" italics="true" pageId="17" pageNumber="990">Diabroctis mimas</emphasis>
(Linnaeus, 1758)
</taxonomicName>
,
<taxonomicName baseAuthorityName="Perty" baseAuthorityYear="1830" box="[362,610,1822,1844]" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="990" phylum="Arthropoda" rank="species" species="aeruginosa">
<emphasis box="[362,610,1822,1844]" italics="true" pageId="17" pageNumber="990">Gromphas aeruginosa</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Erichson" baseAuthorityYear="1847" class="Insecta" family="Scarabaeidae" genus="Oruscatus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="990" phylum="Arthropoda" rank="species" species="davus">
<emphasis italics="true" pageId="17" pageNumber="990">Oruscatus davus</emphasis>
</taxonomicName>
(
<bibRefCitation author="Tarasov S &amp; Genier F" box="[244,506,1853,1875]" pageId="17" pageNumber="990" pagination="1 - 86" refId="ref54817" refString="Tarasov S, Genier F. 2015. Innovative bayesian and parsimony phylogeny of dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) enhanced by ontology-based partitioning of morphological characters. PLoS One 10: 1 - 86." type="journal article" year="2015">Tarasov &amp; Génier, 2015</bibRefCitation>
;
<bibRefCitation author="Valois M &amp; Harada L &amp; Vaz-de-Mello FZ &amp; Silva F" box="[516,710,1853,1875]" pageId="17" pageNumber="990" pagination="347 - 374" refId="ref55098" refString="Valois M, Harada L, Vaz-de-Mello FZ, Silva F. 2018. Synopsis of the genus Diabroctis Gistel, 1857 (Coleoptera: Scarabaeidae: Scarabaeinae) with a new species description. Insect Systematics &amp; Evolution 51: 347 - 374." type="journal article" year="2018">
Valois
<emphasis box="[590,644,1853,1875]" italics="true" pageId="17" pageNumber="990">et al.</emphasis>
, 2018
</bibRefCitation>
). This similarity is likely homologous not only among these taxa, but also to the equally forked shape seen in the
<taxonomicName authorityName="Burmeister" authorityYear="1873" box="[827,951,258,280]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1031" phylum="Arthropoda" rank="tribe" tribe="Eucraniini">Eucraniini</taxonomicName>
(
<bibRefCitation author="Tarasov S &amp; Genier F" box="[967,1242,258,280]" pageId="17" pageNumber="990" pagination="1 - 86" refId="ref54817" refString="Tarasov S, Genier F. 2015. Innovative bayesian and parsimony phylogeny of dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) enhanced by ontology-based partitioning of morphological characters. PLoS One 10: 1 - 86." type="journal article" year="2015">Tarasov &amp; Génier, 2015</bibRefCitation>
), the sister-clade to the phanaeines (see more details below). Therefore, the ancestor of the
<taxonomicName authorityName="Burmeister" authorityYear="1873" box="[1042,1164,319,341]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1031" phylum="Arthropoda" rank="tribe" tribe="Eucraniini">Eucraniini</taxonomicName>
+
<taxonomicName box="[1191,1308,319,341]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1032" phylum="Arthropoda" rank="tribe" tribe="Phanaeini">Phanaeini</taxonomicName>
clade likely also possessed a similar forked axial endophallite, a shape that was directly inherited by
<taxonomicName authorityName="Harold" authorityYear="1868" box="[1290,1388,381,402]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="990" phylum="Arthropoda" rank="genus">
<emphasis box="[1290,1388,381,402]" italics="true" pageId="17" pageNumber="990">Bolbites</emphasis>
</taxonomicName>
and the other mentioned taxa, and from which the more differentiated state found in groups like
<taxonomicName authorityName="MacLeay" authorityYear="1819" box="[1282,1394,442,463]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="990" phylum="Arthropoda" rank="genus">
<emphasis box="[1282,1394,442,463]" italics="true" pageId="17" pageNumber="990">Phanaeus</emphasis>
</taxonomicName>
and
<taxonomicName authority="d'Olsoufieff, 1924" authorityName="d'Olsoufieff" authorityYear="1924" box="[827,1206,473,495]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="990" phylum="Arthropoda" rank="genus">
<emphasis box="[827,1004,473,494]" italics="true" pageId="17" pageNumber="990">Coprophanaeus</emphasis>
<bibRefCitation author="d'Olsoufieff G" box="[1010,1206,473,495]" pageId="17" pageNumber="990" pagination="4 - 201" refId="ref52767" refString="d'Olsoufieff G. 1924. Les Phanaeides (Coleoptera - Lamellicornia), famille Scarabaeidae - tr. Coprini. Insecta 13: 4 - 201." type="journal article" year="1924">dOlsoufieff, 1924</bibRefCitation>
</taxonomicName>
derived. But readers should be cautious as to how these endophallites have been identified in the literature. As confirmed by Marcely Valois (personal communication to
<collectionCode box="[1361,1403,565,586]" country="0" name="Museo de Cipolleti" pageId="17" pageNumber="990">MC</collectionCode>
,
<date pageId="17" pageNumber="990" value="2020-02-13">13 February 2020</date>
), the labelling of the axial and subaxial endophallites of
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1013,1213,626,648]" class="Insecta" family="Scarabaeidae" genus="Diabroctis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="990" phylum="Arthropoda" rank="species" species="mimas">
<emphasis box="[1013,1213,626,648]" italics="true" pageId="17" pageNumber="990">Diabroctis mimas</emphasis>
</taxonomicName>
is inverted in Valois
<emphasis box="[827,884,657,678]" italics="true" pageId="17" pageNumber="990">et al.</emphasis>
s (2018) figure 7. The endophallite identified as the subaxial in that figure is actually the axial, whereas the one labelled as being the axial is the subaxial. The same mistake is present in the illustration of the endophallites of
<taxonomicName authority="Brulle, 1838" authorityName="Brulle" authorityYear="1838" box="[1017,1442,779,801]" class="Insecta" family="Scarabaeidae" genus="Eucranium" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="990" phylum="Arthropoda" rank="species" species="arachnoides">
<emphasis box="[1017,1294,779,801]" italics="true" pageId="17" pageNumber="990">Eucranium arachnoides</emphasis>
<bibRefCitation author="Brulle A" box="[1301,1442,779,801]" pageId="17" pageNumber="990" pagination="225 - 448" refId="ref46659" refString="Brulle A. 1838. [livraison 8: pp. 225 - 448]. In: Brulle A, eds. 1837 - 1838. Histoire naturelle des insectes, traitant de leur organisation et de leurs moeurs en general, par M. V. Audouin, et comprenant leur classification et la description des especes, par M. A. Brulle. Le tout accompagne de planches gravees sur acier, d'apres des peintures executees pour cette edition sur la collection du Museum de Paris. Tome VI. Coleopteres III. Paris: F. D. Pillot, 448 + pls 18, 19, 19 bis, 20, 25 [publication date and authorship based on Bousquet 2016]." type="journal article" year="1838">
Brullé,
<quantity box="[1387,1442,780,801]" metricMagnitude="1" metricUnit="m" metricValue="4.66852" pageId="17" pageNumber="990" unit="in" value="1838.0">1838</quantity>
</bibRefCitation>
</taxonomicName>
in
<bibRefCitation author="Tarasov S &amp; Genier F" box="[855,1152,810,832]" pageId="17" pageNumber="990" pagination="1 - 86" refId="ref54817" refString="Tarasov S, Genier F. 2015. Innovative bayesian and parsimony phylogeny of dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) enhanced by ontology-based partitioning of morphological characters. PLoS One 10: 1 - 86." type="journal article" year="2015">Tarasov &amp; Géniers (2015)</bibRefCitation>
figure 21.
</paragraph>
<paragraph blockId="17.[827,1444,873,1876]" lastBlockId="18.[145,762,197,1875]" lastPageId="18" lastPageNumber="991" pageId="17" pageNumber="990">
<emphasis box="[827,1038,873,894]" italics="true" pageId="17" pageNumber="990">General remarks:</emphasis>
Although a precise number was not mentioned,
<bibRefCitation author="Harold E" box="[1011,1204,903,925]" pageId="17" pageNumber="990" pagination="80 - 86" refId="ref49890" refString="Harold E. 1868 a. Diagnosen neuer Coprophagen. Coleopterologische Hefte 3: 80 - 86." type="journal article" year="1868">Harolds (1868a)</bibRefCitation>
original description of
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[855,990,934,956]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="990" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[855,990,934,956]" italics="true" pageId="17" pageNumber="990">B. onitoides</emphasis>
</taxonomicName>
was certainly based on more than
<specimenCount pageId="17" pageNumber="990" type="generic">one specimen</specimenCount>
, as he characterized both sexes separately. Therefore, the type series was originally composed of an unknown number of male and female
<typeStatus box="[1334,1438,1027,1048]" pageId="17" pageNumber="990">syntypes</typeStatus>
. The male designated above as the
<typeStatus box="[1291,1407,1057,1079]" pageId="17" pageNumber="990">lectotype</typeStatus>
of
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[827,960,1087,1109]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="990" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[827,960,1087,1109]" italics="true" pageId="17" pageNumber="990">B. onitoides</emphasis>
</taxonomicName>
was the only specimen we found that can be confidently regarded as a
<typeStatus box="[1154,1246,1118,1139]" pageId="17" pageNumber="990">syntype</typeStatus>
. It is one of three individuals
<specimenCount box="[991,1111,1149,1171]" pageId="17" pageNumber="990" type="male">two males</specimenCount>
and
<specimenCount box="[1173,1301,1149,1171]" pageId="17" pageNumber="990" type="female">one female</specimenCount>
that once belonged to the Edgar von Harold collection and are now at the MNHN via the René Oberthür collection (cf.
<bibRefCitation author="Horn W &amp; Kahle I &amp; Friese G &amp; Gaedike R" box="[865,1064,1241,1262]" pageId="17" pageNumber="990" pagination="1 - 220" refId="ref50241" refString="Horn W, Kahle I, Friese G, Gaedike R. 1990 a. Collectiones entomologicae. Ein Kompendium uber den Verbleib entomologischer Sammlungen der Welt bis 1960. Teil I: A bis K. Berlin: Akademie der Landwirtschaftswissenschaften der Deutschen Demokratischen Republik, 1 - 220." type="book chapter" year="1990">
Horn
<emphasis box="[929,984,1241,1262]" italics="true" pageId="17" pageNumber="990">et al.</emphasis>
, 1990a
</bibRefCitation>
;
<bibRefCitation author="Cambefort Y" box="[1075,1263,1241,1263]" pageId="17" pageNumber="990" refId="ref46872" refString="Cambefort Y. 2006. Des coleopteres, des collections &amp; des hommes. Paris: Publications Scientifiques du Museum national d'Histoire naturelle, 375." type="book" year="2006">Cambefort, 2006</bibRefCitation>
; Cupello, 2020). Even though all
<specimenCount box="[1019,1207,1271,1293]" pageId="17" pageNumber="990" type="generic">three specimens</specimenCount>
agree with
<bibRefCitation author="Harold E" pageId="17" pageNumber="990" pagination="80 - 86" refId="ref49890" refString="Harold E. 1868 a. Diagnosen neuer Coprophagen. Coleopterologische Hefte 3: 80 - 86." type="journal article" year="1868">Harolds (1868a)</bibRefCitation>
description, including the described colour (Harolds text reads coppery, and the three individuals are indeed copper with greenish reflections), only the now-lectotype bears a label with Harolds handwriting identifying it as
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[1042,1186,1425,1446]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="990" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[1042,1186,1425,1446]" italics="true" pageId="17" pageNumber="990">B. onitoides</emphasis>
</taxonomicName>
. As that same label also shows in Harolds handwriting his typical t. abbreviation for type, there is no doubt that specimen is indeed a
<typeStatus box="[959,1051,1517,1538]" pageId="17" pageNumber="990">syntype</typeStatus>
. In turn, the other
<specimenCount box="[1274,1443,1517,1539]" pageId="17" pageNumber="990" type="generic">two specimens</specimenCount>
lack any labels contemporary to Harold, the oldest one borne by each of them being the typical Ex-Musaeo E. Harold label prepared after Oberthürs purchase of the Harold collection. While for coming from Harolds personal collection there is a potential that these
<specimenCount pageId="17" pageNumber="990" type="generic">two specimens</specimenCount>
are also
<typeStatus box="[1043,1144,1701,1722]" pageId="17" pageNumber="990">syntypes</typeStatus>
(now,
<typeStatus box="[1213,1383,1701,1723]" pageId="17" pageNumber="990">paralectotypes</typeStatus>
), and their lack of a Harold identification label by itself does not necessary rule out that possibility (19
<superScript attach="right" box="[1328,1351,1762,1784]" fontSize="5" pageId="17" pageNumber="990">th-</superScript>
century entomologists often labelled only
<specimenCount box="[1233,1395,1793,1815]" pageId="17" pageNumber="990" type="generic">one specimen</specimenCount>
per series), no conclusive evidence exists confirming that they were indeed studied for the original description. For that reason, we decided to list them in the type series section above as potential paralectotypes. No other candidate
<typeStatus box="[347,454,259,280]" pageId="18" pageNumber="991">syntypes</typeStatus>
were found by us in any other museum, neither did Harold state the collection where the specimens he examined came from.
</paragraph>
<paragraph blockId="18.[145,762,197,1875]" pageId="18" pageNumber="991">
As explained by
<bibRefCitation author="Vaz-de-Mello FZ &amp; Grossi PC" box="[382,754,350,373]" pageId="18" pageNumber="991" pagination="220" refId="ref55410" refString="Vaz-de-Mello FZ, Grossi PC. 2010. First report of Bolbites Harold (Scarabaeidae: Scarabaeinae: Phanaeini) in Brazil. The Coleopterists Bulletin 64: 220." type="journal article" year="2010">Vaz-de-Mello &amp; Grossi (2010)</bibRefCitation>
,
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[145,284,381,403]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="991" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[145,284,381,403]" italics="true" pageId="18" pageNumber="991">B. onitoides</emphasis>
</taxonomicName>
was originally described from
<collectingCountry box="[656,760,382,403]" name="Uruguay" pageId="18" pageNumber="991">Uruguay</collectingCountry>
(
<bibRefCitation author="Harold E" box="[154,332,412,434]" pageId="18" pageNumber="991" pagination="80 - 86" refId="ref49890" refString="Harold E. 1868 a. Diagnosen neuer Coprophagen. Coleopterologische Hefte 3: 80 - 86." type="journal article" year="1868">Harold, 1868a</bibRefCitation>
) and subsequently recorded from
<collectingCountry box="[145,261,442,464]" name="Argentina" pageId="18" pageNumber="991">Argentina</collectingCountry>
(
<bibRefCitation author="Steinheil E &amp; Strobel P" box="[272,551,442,464]" pageId="18" pageNumber="991" pagination="554 - 578" refId="ref54656" refString="Steinheil E, Strobel P. 1872. Symbolae ad historiam Coleopterorum Argentiniae meridionalis, ossia enumerazione dei coleotteri raccolti dal prof. P. Strobel nell'Argentinia meridionale, e descrizione delle specie nuove, di Edoardo Steinheil (Traduzione dal manoscritto tedesco inedito per P. Strobel, coll'aggiunta delle notizie sulla dimora delle specie). Atti della Societa Italiana di Scienze Naturali 15: 554 - 578." type="journal article" year="1872">Steinheil &amp; Strobel, 1872</bibRefCitation>
),
<collectingCountry box="[569,649,442,464]" name="Bolivia" pageId="18" pageNumber="991">Bolivia</collectingCountry>
and
<collectingCountry box="[700,761,442,464]" name="Chile" pageId="18" pageNumber="991">Chile</collectingCountry>
(
<bibRefCitation author="d'Olsoufieff G" box="[153,350,473,495]" pageId="18" pageNumber="991" pagination="4 - 201" refId="ref52767" refString="d'Olsoufieff G. 1924. Les Phanaeides (Coleoptera - Lamellicornia), famille Scarabaeidae - tr. Coprini. Insecta 13: 4 - 201." type="journal article" year="1924">dOlsoufieff, 1924</bibRefCitation>
), with doubts from
<collectingCountry box="[574,628,474,495]" name="Peru" pageId="18" pageNumber="991">Peru</collectingCountry>
(
<bibRefCitation author="Balthasar V" pageId="18" pageNumber="991" pagination="337 - 358" refId="ref46125" refString="Balthasar V. 1941. Scarabaeidae Laparostictae (Col.). Beitrage Fauna Perus 1: 337 - 358." type="journal article" year="1941">Balthasar, 1941</bibRefCitation>
),
<collectingCountry box="[223,334,504,525]" name="Paraguay" pageId="18" pageNumber="991">Paraguay</collectingCountry>
(
<bibRefCitation author="Martinez A" box="[351,528,504,526]" pageId="18" pageNumber="991" pagination="1 - 126" refId="ref51548" refString="Martinez A. 1959. Catalogo de los Scarabaeidae Argentinos. Revista del Museo Argentino de Ciencias Naturales ' Bernardino Rivadavia', Ciencias Zoologicas 5: 1 - 126." type="journal article" year="1959">Martínez, 1959</bibRefCitation>
) and, finally,
<collectingCountry box="[689,760,504,526]" name="Brazil" pageId="18" pageNumber="991">Brazil</collectingCountry>
(
<bibRefCitation author="Vaz-de-Mello FZ &amp; Grossi PC" box="[154,496,534,556]" pageId="18" pageNumber="991" pagination="220" refId="ref55410" refString="Vaz-de-Mello FZ, Grossi PC. 2010. First report of Bolbites Harold (Scarabaeidae: Scarabaeinae: Phanaeini) in Brazil. The Coleopterists Bulletin 64: 220." type="journal article" year="2010">Vaz-de-Mello &amp; Grossi, 2010</bibRefCitation>
). dOlsoufieff s (1924) citation of the species for
<collectingCountry box="[455,518,565,587]" name="Chile" pageId="18" pageNumber="991">Chile</collectingCountry>
was likely based on two blue specimens deposited in the MNHN that are almost certainly mislabelled, for we have no evidence that the species occurs so high in the Andes as to be present in
<collectingCountry box="[275,340,688,710]" name="Chile" pageId="18" pageNumber="991">Chile</collectingCountry>
.
<bibRefCitation author="Balthasar V" box="[353,562,688,710]" pageId="18" pageNumber="991" pagination="337 - 358" refId="ref46125" refString="Balthasar V. 1941. Scarabaeidae Laparostictae (Col.). Beitrage Fauna Perus 1: 337 - 358." type="journal article" year="1941">Balthasars (1941</bibRefCitation>
,
<bibRefCitation author="Balthasar V" box="[577,633,688,710]" pageId="18" pageNumber="991" pagination="322 - 344" refId="ref46146" refString="Balthasar V. 1951. Scarabaeidae Laparostictae (Col.). Beitrage Fauna Perus 2: 322 - 344." type="journal article" year="1951">1951</bibRefCitation>
) reluctant inclusion of
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[284,417,718,740]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="991" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[284,417,718,740]" italics="true" pageId="18" pageNumber="991">B. onitoides</emphasis>
</taxonomicName>
in his list of the scarabaeines from
<collectingCountry box="[209,267,750,771]" name="Peru" pageId="18" pageNumber="991">Peru</collectingCountry>
, on the other hand, was based not on the examination of mislabelled specimens, but expressly on second-hand information from unspecified sources; he himself, Balthasar admitted, had never seen any evidence supporting that record. Indeed, no specimens reliably collected either in
<collectingCountry box="[447,501,903,924]" name="Peru" pageId="18" pageNumber="991">Peru</collectingCountry>
or in the neighbouring Bolivian Amazon region are known to exist, only in the Bolivian Chaco. The single potential record from
<collectingCountry box="[145,201,995,1016]" name="Peru" pageId="18" pageNumber="991">Peru</collectingCountry>
, that of an old RBINS red female labelled Pérou, is almost certainly a result of mislabelling. It is highly unlikely, therefore, that the species is indeed present in that country, and its absence from the most recent checklists of Peruvian dung beetles (
<bibRefCitation author="Figueroa L &amp; Edmonds WD &amp; Lujan NM" pageId="18" pageNumber="991" pagination="125 - 138" refId="ref48800" refString="Figueroa L, Edmonds WD, Lujan NM. 2014. La tribu Phanaeini (Coleoptera, Scarabaeidae, Scarabaeinae) en el Peru. Revista Peruana de Biologia 21: 125 - 138." type="journal article" year="2014">
Figueroa
<emphasis box="[695,754,1117,1139]" italics="true" pageId="18" pageNumber="991">et al.</emphasis>
, 2014
</bibRefCitation>
;
<bibRefCitation author="Ratcliffe BC &amp; Jameson ML &amp; Figueroa L &amp; Cave RD &amp; Paulsen MJ &amp; Cano EB &amp; Beza-Beza C &amp; Ferbans L &amp; Reyes-Castillo P" box="[220,474,1147,1170]" pageId="18" pageNumber="991" pagination="186 - 207" refId="ref53918" refString="Ratcliffe BC, Jameson ML, Figueroa L, Cave RD, Paulsen MJ, Cano EB, Beza-Beza C, Jimenez- Ferbans L, Reyes-Castillo P. 2015. Beetles (Coleoptera) of Peru: a survey of the families Scarabaeoidea. Journal of the Kansas Entomological Society 88: 186 - 207." type="journal article" year="2015">
Ratcliffe
<emphasis box="[336,399,1148,1169]" italics="true" pageId="18" pageNumber="991">et al.</emphasis>
, 2015
</bibRefCitation>
) is correct. Finally, its presence in
<collectingCountry box="[283,393,1178,1200]" name="Colombia" pageId="18" pageNumber="991">Colombia</collectingCountry>
, as a MNHN blue male labelled as coming from
<collectingRegion box="[335,417,1209,1231]" country="Colombia" name="Bogota" pageId="18" pageNumber="991">Bogotá</collectingRegion>
would suggest, is even more implausible. That specimen is certainly mislabelled.
</paragraph>
<paragraph blockId="18.[145,762,197,1875]" lastBlockId="18.[809,1426,197,1476]" pageId="18" pageNumber="991">
<bibRefCitation author="Steinheil E &amp; Strobel P" box="[169,480,1270,1292]" pageId="18" pageNumber="991" pagination="554 - 578" refId="ref54656" refString="Steinheil E, Strobel P. 1872. Symbolae ad historiam Coleopterorum Argentiniae meridionalis, ossia enumerazione dei coleotteri raccolti dal prof. P. Strobel nell'Argentinia meridionale, e descrizione delle specie nuove, di Edoardo Steinheil (Traduzione dal manoscritto tedesco inedito per P. Strobel, coll'aggiunta delle notizie sulla dimora delle specie). Atti della Societa Italiana di Scienze Naturali 15: 554 - 578." type="journal article" year="1872">Steinheil &amp; Strobel (1872)</bibRefCitation>
were the first to report the polychromatism of
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[418,556,1301,1323]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="991" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[418,556,1301,1323]" italics="true" pageId="18" pageNumber="991">B. onitoides</emphasis>
</taxonomicName>
. They mentioned the examination of reddish specimens from Concordia (
<collectingRegion box="[153,292,1362,1384]" country="Argentina" name="Entre Rios" pageId="18" pageNumber="991">Entre Ríos</collectingRegion>
, eastern
<collectingCountry box="[412,539,1362,1384]" name="Argentina" pageId="18" pageNumber="991">Argentina</collectingCountry>
) and
<collectingRegion box="[617,760,1362,1384]" country="Uruguay" name="Montevideo" pageId="18" pageNumber="991">Montevideo</collectingRegion>
(
<collectingCountry box="[153,261,1393,1415]" name="Uruguay" pageId="18" pageNumber="991">Uruguay</collectingCountry>
), and a blue-green female from Salvador near
<collectingRegion box="[202,302,1424,1446]" country="Argentina" name="San Luis" pageId="18" pageNumber="991">San Luis</collectingRegion>
(western
<collectingCountry box="[410,529,1424,1446]" name="Argentina" pageId="18" pageNumber="991">Argentina</collectingCountry>
). These observations are in line with the distribution of the colour variants observed by us. On the other hand, they also mentioned the study of 15 individuals from
<collectingRegion box="[546,653,1516,1538]" country="Argentina" name="Mendoza" pageId="18" pageNumber="991">Mendoza</collectingRegion>
(eastern
<collectingCountry box="[145,263,1546,1568]" name="Argentina" pageId="18" pageNumber="991">Argentina</collectingCountry>
) described by them as being entirely black and without any metallic sheen. This is odd as none of the 1204 specimens examined by us either is black or lacks metallic reflections. Also studying Argentinian specimens,
<bibRefCitation author="Burmeister H" box="[280,497,1669,1691]" pageId="18" pageNumber="991" pagination="120 - 133" refId="ref46788" refString="Burmeister H. 1874. Lamellicornia Argentina [Fortsetzung]. Entomologische Zeitung Stettin 35: 120 - 133." type="journal article" year="1874">Burmeister (1874)</bibRefCitation>
mentioned something equally strange. He examined individuals of the entire chromatic spectrum of the species, including the first report of the bicoloured variant. Nevertheless, Burmeister referred to likely blue specimens and the elytra of the bicoloured individuals as being black (
<emphasis box="[159,265,1853,1874]" italics="true" pageId="18" pageNumber="991">schwarze</emphasis>
in his original German). We re-examined the
<taxonomicName authorityName="Harold" authorityYear="1868" box="[852,943,197,218]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="991" phylum="Arthropoda" rank="genus">
<emphasis box="[852,943,197,218]" italics="true" pageId="18" pageNumber="991">Bolbites</emphasis>
</taxonomicName>
of the Burmeister collection at the MACN and found only six blue specimens and no individuals of the other colour variants. The MACN general collection itself has some old, undated, bicoloured and red specimens that may have been studied by
<bibRefCitation author="Burmeister H" box="[809,1020,350,372]" pageId="18" pageNumber="991" pagination="120 - 133" refId="ref46788" refString="Burmeister H. 1874. Lamellicornia Argentina [Fortsetzung]. Entomologische Zeitung Stettin 35: 120 - 133." type="journal article" year="1874">Burmeister (1874)</bibRefCitation>
, who worked in that museum from 1862 until his death in 1892 (
<bibRefCitation author="Ulrich W" box="[1179,1329,381,403]" pageId="18" pageNumber="991" pagination="1 - 22" refId="ref55076" refString="Ulrich W. 1972. Hermann Burmeister, 1807 to 1892. Annual Review of Entomology 17: 1 - 22." type="journal article" year="1972">Ulrich, 1972</bibRefCitation>
;
<bibRefCitation author="Bragg PE" pageId="18" pageNumber="991" pagination="1 - 4" refId="ref46515" refString="Bragg PE. 2007. Biographies of phasmatologists - 3. Hermann Burmeister. Phasmid Studies 16: 1 - 4." type="journal article" year="2007">Bragg, 2007</bibRefCitation>
); but none of the MACN specimens is black or has elytra of that colour. As for the specimens studied by
<bibRefCitation author="Steinheil E &amp; Strobel P" box="[809,1105,473,495]" pageId="18" pageNumber="991" pagination="554 - 578" refId="ref54656" refString="Steinheil E, Strobel P. 1872. Symbolae ad historiam Coleopterorum Argentiniae meridionalis, ossia enumerazione dei coleotteri raccolti dal prof. P. Strobel nell'Argentinia meridionale, e descrizione delle specie nuove, di Edoardo Steinheil (Traduzione dal manoscritto tedesco inedito per P. Strobel, coll'aggiunta delle notizie sulla dimora delle specie). Atti della Societa Italiana di Scienze Naturali 15: 554 - 578." type="journal article" year="1872">Steinheil &amp; Strobel (1872)</bibRefCitation>
, we reviewed the
<taxonomicName authorityName="Harold" authorityYear="1868" box="[1307,1398,473,494]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="991" phylum="Arthropoda" rank="genus">
<emphasis box="[1307,1398,473,494]" italics="true" pageId="18" pageNumber="991">Bolbites</emphasis>
</taxonomicName>
of the MNHN, where the Steinheil collection is currently housed (
<bibRefCitation author="Horn W &amp; Kahle I &amp; Friese G &amp; Gaedike R" box="[910,1125,534,556]" pageId="18" pageNumber="991" pagination="223 - 573" refId="ref50292" refString="Horn W, Kahle I, Friese G, Gaedike R. 1990 b. Collectiones entomologicae. Ein Kompendium uber den Verbleib entomologischer Sammlungen der Welt bis 1960. Teil II: L bis Z. Berlin: Akademie der Landwirtschaftswissenschaften der Deutschen Demokratischen Republik, 223 - 573." type="book chapter" year="1990">
Horn
<emphasis box="[979,1039,534,556]" italics="true" pageId="18" pageNumber="991">et al.</emphasis>
, 1990b
</bibRefCitation>
), and did not find any of the specimens mentioned in that work. It is likely they were all deposited in Strobels collection. According to
<bibRefCitation author="Weber K" box="[809,967,626,648]" pageId="18" pageNumber="991" refId="ref55600" refString="Weber K. 2016. [entry on] Strobel, Pellegrino de. In: Anonymous, ed. Biographies of the entomologists of the world. Available at: http: // sdei. senckenberg. de / biographies / (last accessed on 14 May 2020)." type="url" year="2016">Weber (2016)</bibRefCitation>
, Strobels Argentinian
<taxonomicName box="[1248,1377,626,648]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="991" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
are housed in the Museo di Storia Naturale dellUniversità di Parma,
<collectingCountry box="[926,979,687,709]" name="Italy" pageId="18" pageNumber="991">Italy</collectingCountry>
, and some other of his South American beetles were sent to the Museo Zoologico La Specola, Florence. Unfortunately, we did not have the chance to visit any Italian museums for this work to confirm
<bibRefCitation author="Steinheil E &amp; Strobel P" box="[809,1120,810,832]" pageId="18" pageNumber="991" pagination="554 - 578" refId="ref54656" refString="Steinheil E, Strobel P. 1872. Symbolae ad historiam Coleopterorum Argentiniae meridionalis, ossia enumerazione dei coleotteri raccolti dal prof. P. Strobel nell'Argentinia meridionale, e descrizione delle specie nuove, di Edoardo Steinheil (Traduzione dal manoscritto tedesco inedito per P. Strobel, coll'aggiunta delle notizie sulla dimora delle specie). Atti della Societa Italiana di Scienze Naturali 15: 554 - 578." type="journal article" year="1872">Steinheil &amp; Strobels (1872)</bibRefCitation>
information. However, like Burmeisters we suspect their specimens were actually dark blue instead of black as they described.
</paragraph>
<paragraph blockId="18.[809,1426,197,1476]" pageId="18" pageNumber="991">
<bibRefCitation author="Judulien F" box="[833,1023,902,924]" pageId="18" pageNumber="991" pagination="371 - 380" refId="ref50626" refString="Judulien F. 1899. Quelques notes sur plusieurs coprophages de Buenos Aires. Revista del Museo de La Plata 9: 371 - 380. [Frere Judulien was the religious name of the French- Argentinian entomologist Jean (Juan) Brethes (1871 - 1928) and it was with this name he signed his early works (Doello- Jurado, 1928)]." type="journal article" year="1899">Judulien (1899)</bibRefCitation>
illustrated and briefly described the larva of
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[948,1079,933,954]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="991" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[948,1079,933,954]" italics="true" pageId="18" pageNumber="991">B. onitoides</emphasis>
</taxonomicName>
, but the instar was not stated and cannot be inferred from the data provided. As we observed for the larva of
<taxonomicName authority="Harold, 1869" authorityName="Harold, Black" authorityYear="1869" class="Insecta" family="Scarabaeidae" genus="Gromphas" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="991" phylum="Arthropoda" rank="species" species="inermis">
<emphasis box="[1108,1328,994,1016]" italics="true" pageId="18" pageNumber="991">Gromphas inermis</emphasis>
Harold, 1869
</taxonomicName>
(see:
<bibRefCitation author="Cupello M &amp; Vaz-de-Mello FZ" box="[933,1282,1025,1047]" pageId="18" pageNumber="991" pagination="439 - 482" refId="ref47132" refString="Cupello M, Vaz-de-Mello FZ. 2013. Taxonomic revision of the South American dung beetle genus Gromphas Brulle, 1837 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini: Gromphadina). Zootaxa 3722: 439 - 482." type="journal article" year="2013">Cupello &amp; Vaz-de-Mello, 2013</bibRefCitation>
), Juduliens information that the
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[1062,1199,1055,1077]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="991" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[1062,1199,1055,1077]" italics="true" pageId="18" pageNumber="991">B. onitoides</emphasis>
</taxonomicName>
larva bears a fivearticulated antenna is contrary to
<bibRefCitation author="Edmonds WD &amp; Halffter G" pageId="18" pageNumber="991" pagination="307 - 331" refId="ref48185" refString="Edmonds WD, Halffter G. 1978. Taxonomic review of immature dung beetles of the subfamily Scarabaeinae (Coleoptera: Scarabaeidae). Systematic Entomology 3: 307 - 331." type="journal article" year="1978">Edmonds &amp; Halffters (1978)</bibRefCitation>
observation that the
<taxonomicName authorityName="Latreille" authorityYear="1802" box="[1265,1424,1117,1139]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1033" phylum="Arthropoda" rank="subFamily" subFamily="Scarabaeinae">Scarabaeinae</taxonomicName>
larvae have four antennomeres. Yet, under a close reading of Juduliens work, we see that he assigned a five-articulated antenna to almost all the scarabaeine larvae he described,
<taxonomicName authority="(Castelnau, 1840)" authorityName="Usually" baseAuthorityName="Castelnau" baseAuthorityYear="1840" class="Insecta" family="Scarabaeidae" genus="Sulcophanaeus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="991" phylum="Arthropoda" rank="species" species="menelas">
<emphasis box="[1102,1424,1239,1261]" italics="true" pageId="18" pageNumber="991">Sulcophanaeus menelas</emphasis>
(
<bibRefCitation author="de Castelnau, FL" box="[818,1032,1270,1292]" pageId="18" pageNumber="991" refId="ref47427" refString="de Castelnau, FL comte. 1840. Histoire naturelle des animaux articules, annelides, crustaces, arachnides, myriapodes et insectes. Histoire naturelle des insectes coleopteres. Tome deuxieme. Paris: Dumenil, 563 pp. + [1] + 38 pls." type="book" year="1840">Castelnau, 1840</bibRefCitation>
)
</taxonomicName>
(as
<taxonomicName authorityName="Pronotum" baseAuthorityName="Fabricius" baseAuthorityYear="1781" box="[1104,1407,1270,1291]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="991" phylum="Arthropoda" rank="species" species="splendidulus">
<emphasis box="[1104,1407,1270,1291]" italics="true" pageId="18" pageNumber="991">Phanaeus splendidulus</emphasis>
</taxonomicName>
),
<taxonomicName authority="Harold, 1867" authorityName="Harold" authorityYear="1867" box="[809,1200,1301,1323]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="991" phylum="Arthropoda" rank="species" species="muticus">
<emphasis box="[809,1025,1301,1322]" italics="true" pageId="18" pageNumber="991">Canthon muticus</emphasis>
Harold, 1867
</taxonomicName>
and
<emphasis box="[1269,1424,1301,1322]" italics="true" pageId="18" pageNumber="991">C. edentulus</emphasis>
Harold, 1868 being the only exceptions. Judging from his illustrations, we think it is possible that Judulien mistook the antennifer for the first antennomere, so miscounting the total number of articles in the antennae of the specimens he examined.
</paragraph>
<paragraph blockId="18.[809,1425,1517,1876]" lastBlockId="19.[163,780,197,1905]" lastPageId="19" lastPageNumber="992" pageId="18" pageNumber="991">
<emphasis box="[809,1146,1517,1538]" italics="true" pageId="18" pageNumber="991">
Ecology:
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[923,1146,1517,1538]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="991" phylum="Arthropoda" rank="species" species="onitoides">Bolbites onitoides</taxonomicName>
</emphasis>
inhabits natural and artificial grasslands, savannas and dry deciduous forests from about sea level to at least
<quantity box="[1263,1349,1578,1600]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="18" pageNumber="991" unit="m" value="1450.0">1450 m</quantity>
(
<bibRefCitation author="Kirk AA" pageId="18" pageNumber="991" pagination="45 - 54" refId="ref50693" refString="Kirk AA. 1992. Dung beetles (Coleoptera: Scarabaeidae) active in patchy forest and pasture habitats in Santa Cruz province, Bolivia, during spring. Folia Entomologica Mexicana 84: 45 - 54." type="journal article" year="1992">Kirk, 1992</bibRefCitation>
;
<bibRefCitation author="Hamel-Leigue AC &amp; Herzog SK &amp; Mann DJ &amp; Larsen TH &amp; Gill BD &amp; Edmonds WD &amp; Spector S" box="[876,1166,1609,1631]" pageId="18" pageNumber="991" pagination="43 - 95" refId="ref49799" refString="Hamel-Leigue AC, Herzog SK, Mann DJ, Larsen TH, Gill BD, Edmonds WD, Spector S. 2009. Distribucion e historia natural de escarabajos de la tribu Phanaeini (Coleoptera: Scarabaeidae: Scarabaeinae) en Bolivia. Kempffiana 5: 43 - 95." type="journal article" year="2009">
Hamel-Leigue
<emphasis box="[1043,1099,1609,1630]" italics="true" pageId="18" pageNumber="991">et al.</emphasis>
, 2009
</bibRefCitation>
;
<bibRefCitation author="Alonso CBG &amp; Zurita GA &amp; Bellocq MI" box="[1177,1376,1609,1631]" pageId="18" pageNumber="991" pagination="296 - 308" refId="ref45962" refString="Alonso CBG, Zurita GA, Bellocq MI. 2019. Livestock areas with canopy cover sustain dung beetle diversity in the humid subtropical Chaco forest. Insect Conservation and Diversity 12: 296 - 308." type="journal article" year="2019">
Alonso
<emphasis box="[1260,1315,1609,1630]" italics="true" pageId="18" pageNumber="991">et al.</emphasis>
2019
</bibRefCitation>
and label data). Label data indicate that adults are active throughout the rainiest half of the year, having been recorded from October to April. In
<collectingCountry box="[1188,1270,1701,1723]" name="Bolivia" pageId="18" pageNumber="991">Bolivia</collectingCountry>
, in particular,
<bibRefCitation author="Hamel-Leigue AC &amp; Herzog SK &amp; Mann DJ &amp; Larsen TH &amp; Gill BD &amp; Edmonds WD &amp; Spector S" box="[809,1118,1731,1753]" pageId="18" pageNumber="991" pagination="43 - 95" refId="ref49799" refString="Hamel-Leigue AC, Herzog SK, Mann DJ, Larsen TH, Gill BD, Edmonds WD, Spector S. 2009. Distribucion e historia natural de escarabajos de la tribu Phanaeini (Coleoptera: Scarabaeidae: Scarabaeinae) en Bolivia. Kempffiana 5: 43 - 95." type="journal article" year="2009">
Hamel-Leigue
<emphasis box="[980,1038,1731,1753]" italics="true" pageId="18" pageNumber="991">et al.</emphasis>
(2009)
</bibRefCitation>
recorded individuals from September to January, suggesting an earlier annual activity period there than in the examined southern part of the distribution. Also, according to Hamel-Leigue
<emphasis box="[894,951,1854,1876]" italics="true" pageId="18" pageNumber="991">et al.</emphasis>
(2009) and more recently
<bibRefCitation author="Cabrera Walsh G" lastPageId="19" lastPageNumber="992" pageId="18" pageNumber="991" pagination="760 - 762" refId="ref46808" refString="Cabrera Walsh G. 2020. Dung beetle (Coleoptera: Scarabaeidae: Scarabaeinae) coloration and foraging time. The Coleopterists Bulletin 74: 760 - 762." type="journal article" year="2020">Cabrera Walsh (2020)</bibRefCitation>
, the flight activity time of
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[541,673,197,219]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="992" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[541,673,197,219]" italics="true" pageId="19" pageNumber="992">B. onitoides</emphasis>
</taxonomicName>
seems to be diurnal; this can also be inferred from the metallic colouration of the species, which, at least among dung beetles, is usually associated with diurnal or crepuscular species (
<bibRefCitation author="Hernandez MIM" box="[463,675,320,342]" pageId="19" pageNumber="992" pagination="597 - 600" refId="ref50199" refString="Hernandez MIM. 2002. The night and day of dung beetles (Coleoptera, Scarabaeidae) in the Serra do Japi, Brazil: elytra colour related to daily activity. Revista Brasileira de Entomologia 46: 597 - 600." type="journal article" year="2002">Hernández, 2002</bibRefCitation>
;
<bibRefCitation author="Feer F &amp; Pincebourde S" pageId="19" pageNumber="992" pagination="21 - 30" refId="ref48664" refString="Feer F, Pincebourde S. 2005. Diel flight activity and ecological segregation within an assemblage of tropical forest dung and carrion beetles. Journal of Tropical Ecology 21: 21 - 30." type="journal article" year="2005">Feer &amp; Pincebourde, 2005</bibRefCitation>
;
<bibRefCitation author="Cabrera Walsh G" box="[384,621,350,372]" pageId="19" pageNumber="992" pagination="760 - 762" refId="ref46808" refString="Cabrera Walsh G. 2020. Dung beetle (Coleoptera: Scarabaeidae: Scarabaeinae) coloration and foraging time. The Coleopterists Bulletin 74: 760 - 762." type="journal article" year="2020">Cabrera Walsh, 2020</bibRefCitation>
).
</paragraph>
<paragraph blockId="19.[163,780,197,1905]" pageId="19" pageNumber="992">
Concerning food habits,
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[469,603,381,403]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="992" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[469,603,381,403]" italics="true" pageId="19" pageNumber="992">B. onitoides</emphasis>
</taxonomicName>
is known to be preferentially coprophagous, having been reported from cow, sheep, horse and human excrement (
<bibRefCitation author="Fabre J-H" pageId="19" pageNumber="992" refId="ref48519" refString="Fabre J-H. 1899. Souvenirs entomologiques (sixieme serie). Etudes sur l'instict et les moeurs des insectes. Paris: Librairie Ch. Delagrave, 418." type="book" year="1899">Fabre, 1899</bibRefCitation>
,
<bibRefCitation author="Fabre J-H" box="[235,292,473,494]" pageId="19" pageNumber="992" refId="ref48549" refString="Fabre J-H. 1919. The glow-worm and other beetles. Toronto: McClelland &amp; Stewart, 488." type="book" year="1919">1919</bibRefCitation>
;
<bibRefCitation author="Halffter G &amp; Matthews EG" box="[307,631,473,495]" pageId="19" pageNumber="992" pagination="1 - 312" refId="ref49650" refString="Halffter G, Matthews EG. 1966. The natural history of dung beetles of the subfamily Scarabaeinae (Coleoptera: Scarabaeidae). Folia Entomologica Mexicana 12 - 14: 1 - 312." type="journal article" year="1966">Halffter &amp; Matthews, 1966</bibRefCitation>
;
<bibRefCitation author="Kirk AA" box="[646,773,473,495]" pageId="19" pageNumber="992" pagination="45 - 54" refId="ref50693" refString="Kirk AA. 1992. Dung beetles (Coleoptera: Scarabaeidae) active in patchy forest and pasture habitats in Santa Cruz province, Bolivia, during spring. Folia Entomologica Mexicana 84: 45 - 54." type="journal article" year="1992">Kirk, 1992</bibRefCitation>
;
<bibRefCitation author="Cabrera Walsh G &amp; Gandolfo D" box="[163,537,504,526]" pageId="19" pageNumber="992" pagination="581 - 588" refId="ref46837" refString="Cabrera Walsh G, Gandolfo D. 1996. Nidification of thirteen common Argentine dung beetles (Scarabaeidae: Scarabaeinae). Annals of the Entomological Society of America 89: 581 - 588." type="journal article" year="1996">Cabrera Walsh &amp; Gandolfo, 1996</bibRefCitation>
; Morelli &amp; Gonzalez-Vainer, 1997;
<bibRefCitation author="Hamel-Leigue AC &amp; Herzog SK &amp; Mann DJ &amp; Larsen TH &amp; Gill BD &amp; Edmonds WD &amp; Spector S" box="[331,648,534,556]" pageId="19" pageNumber="992" pagination="43 - 95" refId="ref49799" refString="Hamel-Leigue AC, Herzog SK, Mann DJ, Larsen TH, Gill BD, Edmonds WD, Spector S. 2009. Distribucion e historia natural de escarabajos de la tribu Phanaeini (Coleoptera: Scarabaeidae: Scarabaeinae) en Bolivia. Kempffiana 5: 43 - 95." type="journal article" year="2009">
Hamel-Leigue
<emphasis box="[513,574,534,556]" italics="true" pageId="19" pageNumber="992">et al.</emphasis>
, 2009
</bibRefCitation>
; and label data). The Belgrano (
<collectingCountry box="[402,519,565,587]" name="Argentina" pageId="19" pageNumber="992">Argentina</collectingCountry>
) series studied for this work, which was collected in a trap baited with rotten chicken meat, shows that the species may also be at least occasionally necrophagous.
<bibRefCitation author="Fabre J-H" box="[557,698,657,679]" pageId="19" pageNumber="992" refId="ref48519" refString="Fabre J-H. 1899. Souvenirs entomologiques (sixieme serie). Etudes sur l'instict et les moeurs des insectes. Paris: Librairie Ch. Delagrave, 418." type="book" year="1899">Fabre (1899</bibRefCitation>
,
<bibRefCitation author="Fabre J-H" box="[713,769,657,678]" pageId="19" pageNumber="992" refId="ref48549" refString="Fabre J-H. 1919. The glow-worm and other beetles. Toronto: McClelland &amp; Stewart, 488." type="book" year="1919">1919</bibRefCitation>
) speculated that, before the European colonization of the New World and the introduction of Old World livestock,
<taxonomicName authorityName="Harold" authorityYear="1868" box="[283,378,749,770]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="992" phylum="Arthropoda" rank="genus">
<emphasis box="[283,378,749,770]" italics="true" pageId="19" pageNumber="992">Bolbites</emphasis>
</taxonomicName>
and the other large dung beetles of the South American grasslands relied mainly on ground sloth (Xenarthra: Folivora) dung for food. But since these giant mammals were extinct in South America for about 10 000 years before the arrival of the first European settlers (
<bibRefCitation author="Steadman DW &amp; Martin PS &amp; MacPhee RDE &amp; Jull AJT &amp; McDonald HG &amp; Woods CA &amp; Iturralde-Vinent M &amp; Hodgins GWL" box="[502,764,902,924]" pageId="19" pageNumber="992" pagination="11763 - 11768" refId="ref54603" refString="Steadman DW, Martin PS, MacPhee RDE, Jull AJT, McDonald HG, Woods CA, Iturralde-Vinent M, Hodgins GWL. 2005. Asynchronous extinction of late Quaternary sloths on continents and islands. Proceedings of the National Academy of Sciences of the USA 102: 11763 - 11768." type="journal article" year="2005">
Steadman
<emphasis box="[631,691,902,924]" italics="true" pageId="19" pageNumber="992">et al.</emphasis>
, 2005
</bibRefCitation>
), Fabres suggestion is, at most, only partly correct. The complete answer is probably much more trivial: between the extinction of the ground sloths and the arrival of Europeans,
<taxonomicName authorityName="Latreille" authorityYear="1802" box="[408,562,1025,1047]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="1033" phylum="Arthropoda" rank="subFamily" subFamily="Scarabaeinae">Scarabaeinae</taxonomicName>
communities must have consumed, as they likely had done before and certainly continue to do today, dung produced by other autochthonous mammalian fauna of those habitats, including animals such as tapirs, peccaries, giant armadillos, capybaras, monkeys, deer and, for the past 15 00011 000 years (
<bibRefCitation author="Dillehay TD" box="[479,646,1209,1231]" pageId="19" pageNumber="992" pagination="206 - 216" refId="ref47778" refString="Dillehay TD. 1999. The late Pleistocene cultures of South America. Evolutionary Anthropology 7: 206 - 216." type="journal article" year="1999">Dillehay, 1999</bibRefCitation>
;
<bibRefCitation author="Raghavan M &amp; Steinrucken M &amp; Harris K &amp; Schiffels S &amp; Rasmussen S &amp; DeGiorgio M &amp; Albrechtsen A &amp; Valdiosera C &amp; Avila-Arcos MC &amp; Malaspinas A-S &amp; Eriksson A &amp; Moltke I &amp; Metspalu M &amp; Homburger JR &amp; Wall J &amp; Cornejo OE &amp; Moreno-Mayar JV &amp; Korneliussen TS &amp; Pierre T &amp; Rasmussen M &amp; Campos PF &amp; Damgaard PB &amp; Allentoft ME &amp; Lindo J &amp; Metspalu E &amp; Rodriguez-Varela R &amp; Mansilla J &amp; Henrickson C &amp; Seguin-Orlando A &amp; Malmstrom H &amp; Stafford Jr T &amp; Shringarpure SS &amp; Moreno-Estrada A &amp; Karmin M &amp; Tambets K &amp; Bergstrom A &amp; Xue Y &amp; Warmuth V &amp; Friend AD &amp; Singarayer J &amp; Valdes P &amp; Balloux F &amp; Leboreiro I &amp; Vera JL &amp; Rangel-Villalobos H &amp; Pettener D &amp; Luiselli D &amp; Davis LG &amp; Heyer E &amp; Zollikofer CPE &amp; Ponce de Leon MS &amp; Smith CI &amp; Grimes V &amp; Pike K-A &amp; Deal M &amp; Fuller BT &amp; Arriaza B &amp; Standen V &amp; Luz MF &amp; Ricaut F &amp; Guidon N &amp; Osipova L &amp; Voevoda MI &amp; Posukh OL &amp; Balanovsky O &amp; Lavryashina M &amp; Bogunov Y &amp; Khusnutdinova E &amp; Gubina M &amp; Balanovska E &amp; Fedorova S &amp; Litvinov S &amp; Malyarchuk B &amp; Derenko M &amp; Mosher MJ &amp; Archer D &amp; Cybukski J &amp; Petzelt B &amp; Mitchell J &amp; Worl R &amp; Norman PJ &amp; Parham P &amp; Kemp BM &amp; Kivisild T &amp; Tyler-Smith C &amp; Sandhu MS &amp; Crawford M &amp; Villems R &amp; Smith DG &amp; Waters MR &amp; Goebel T &amp; Johnson JR &amp; Malhi RS &amp; Jakobsson M &amp; Meltzer DJ &amp; Manica A &amp; Durbin R &amp; Bustamante CD &amp; Song YS &amp; Nielsen R &amp; Willerslev E" pageId="19" pageNumber="992" pagination="10" refId="ref53592" refString="Raghavan M, Steinrucken M, Harris K, Schiffels S, Rasmussen S, DeGiorgio M, Albrechtsen A, Valdiosera C, Avila-Arcos MC, Malaspinas A-S, Eriksson A, Moltke I, Metspalu M, Homburger JR, Wall J, Cornejo OE, Moreno-Mayar JV, Korneliussen TS, Pierre T, Rasmussen M, Campos PF, Damgaard PB, Allentoft ME, Lindo J, Metspalu E, Rodriguez-Varela R, Mansilla J, Henrickson C, Seguin-Orlando A, Malmstrom H, Stafford Jr T, Shringarpure SS, Moreno-Estrada A, Karmin M, Tambets K, Bergstrom A, Xue Y, Warmuth V, Friend AD, Singarayer J, Valdes P, Balloux F, Leboreiro I, Vera JL, Rangel-Villalobos H, Pettener D, Luiselli D, Davis LG, Heyer E, Zollikofer CPE, Ponce de Leon MS, Smith CI, Grimes V, Pike K-A, Deal M, Fuller BT, Arriaza B, Standen V, Luz MF, Ricaut F, Guidon N, Osipova L, Voevoda MI, Posukh OL, Balanovsky O, Lavryashina M, Bogunov Y, Khusnutdinova E, Gubina M, Balanovska E, Fedorova S, Litvinov S, Malyarchuk B, Derenko M, Mosher MJ, Archer D, Cybukski J, Petzelt B, Mitchell J, Worl R, Norman PJ, Parham P, Kemp BM, Kivisild T, Tyler-Smith C, Sandhu MS, Crawford M, Villems R, Smith DG, Waters MR, Goebel T, Johnson JR, Malhi RS, Jakobsson M, Meltzer DJ, Manica A, Durbin R, Bustamante CD, Song YS, Nielsen R, Willerslev E. 2015. Genomic evidence for the Pleistocene and recent population history of Native Americans. Science 349: 10." type="journal article" year="2015">
Raghavan
<emphasis box="[163,219,1240,1261]" italics="true" pageId="19" pageNumber="992">et al.</emphasis>
, 2015
</bibRefCitation>
), humans. Naturally, this does not exclude the possibility that, when the ground sloths were still alive, South American dung beetles, including
<taxonomicName authorityName="Harold" authorityYear="1868" box="[688,779,1301,1322]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="992" phylum="Arthropoda" rank="genus">
<emphasis box="[688,779,1301,1322]" italics="true" pageId="19" pageNumber="992">Bolbites</emphasis>
</taxonomicName>
and its ancestors, did indeed include their excrement in their diet, as suggested by Fabre.
</paragraph>
<paragraph blockId="19.[163,780,197,1905]" lastBlockId="19.[827,1444,197,1323]" pageId="19" pageNumber="992">
The nesting behaviour of
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[531,685,1393,1415]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="992" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[531,685,1393,1415]" italics="true" pageId="19" pageNumber="992">B. onitoides</emphasis>
</taxonomicName>
has in part been directly observed and reviewed from the literature several times (
<bibRefCitation author="Fabre J-H" box="[543,692,1454,1476]" pageId="19" pageNumber="992" refId="ref48519" refString="Fabre J-H. 1899. Souvenirs entomologiques (sixieme serie). Etudes sur l'instict et les moeurs des insectes. Paris: Librairie Ch. Delagrave, 418." type="book" year="1899">Fabre, 1899</bibRefCitation>
,
<bibRefCitation author="Fabre J-H" box="[712,771,1454,1475]" pageId="19" pageNumber="992" refId="ref48549" refString="Fabre J-H. 1919. The glow-worm and other beetles. Toronto: McClelland &amp; Stewart, 488." type="book" year="1919">1919</bibRefCitation>
;
<bibRefCitation author="Judulien F" box="[163,344,1485,1507]" pageId="19" pageNumber="992" pagination="371 - 380" refId="ref50626" refString="Judulien F. 1899. Quelques notes sur plusieurs coprophages de Buenos Aires. Revista del Museo de La Plata 9: 371 - 380. [Frere Judulien was the religious name of the French- Argentinian entomologist Jean (Juan) Brethes (1871 - 1928) and it was with this name he signed his early works (Doello- Jurado, 1928)]." type="journal article" year="1899">Judulien, 1899</bibRefCitation>
;
<bibRefCitation author="Laloy L" box="[360,499,1485,1507]" pageId="19" pageNumber="992" pagination="169 - 171" refId="ref51055" refString="Laloy L. 1902. Moeurs des coleopteres coprophages, specialement de La Plata. Le Naturaliste, 2 e Serie 16: 169 - 171." type="journal article" year="1902">Laloy, 1902</bibRefCitation>
;
<bibRefCitation author="Halffter G &amp; Matthews EG" pageId="19" pageNumber="992" pagination="1 - 312" refId="ref49650" refString="Halffter G, Matthews EG. 1966. The natural history of dung beetles of the subfamily Scarabaeinae (Coleoptera: Scarabaeidae). Folia Entomologica Mexicana 12 - 14: 1 - 312." type="journal article" year="1966">Halffter &amp; Matthews, 1966</bibRefCitation>
;
<bibRefCitation author="Halffter G &amp; Edmonds WD" box="[236,554,1516,1538]" pageId="19" pageNumber="992" refId="ref49587" refString="Halffter G, Edmonds WD. 1982. The nesting behavior of dung beetles (Scarabaeinae). An ecological and evolutive approach. Mexico: Instituto de Ecologia, 176." type="book" year="1982">Halffter &amp; Edmonds, 1982</bibRefCitation>
;
<bibRefCitation author="Cabrera Walsh G &amp; Gandolfo D" pageId="19" pageNumber="992" pagination="581 - 588" refId="ref46837" refString="Cabrera Walsh G, Gandolfo D. 1996. Nidification of thirteen common Argentine dung beetles (Scarabaeidae: Scarabaeinae). Annals of the Entomological Society of America 89: 581 - 588." type="journal article" year="1996">Cabrera Walsh &amp; Gandolfo, 1996</bibRefCitation>
;
<bibRefCitation author="Cupello M &amp; Vaz-de-Mello FZ" box="[363,727,1546,1568]" pageId="19" pageNumber="992" pagination="439 - 482" refId="ref47132" refString="Cupello M, Vaz-de-Mello FZ. 2013. Taxonomic revision of the South American dung beetle genus Gromphas Brulle, 1837 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini: Gromphadina). Zootaxa 3722: 439 - 482." type="journal article" year="2013">Cupello &amp; Vaz-de-Mello, 2013</bibRefCitation>
). In summary,
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[290,433,1577,1599]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="992" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[290,433,1577,1599]" italics="true" pageId="19" pageNumber="992">B. onitoides</emphasis>
</taxonomicName>
builds pyriform brood balls typical of the Phanaeina and other dung beetles that conform to pattern II of Halffter &amp; Edmonds (1982) nesting behaviour classification.
<bibRefCitation author="Halffter G &amp; Edmonds WD" pageId="19" pageNumber="992" refId="ref49587" refString="Halffter G, Edmonds WD. 1982. The nesting behavior of dung beetles (Scarabaeinae). An ecological and evolutive approach. Mexico: Instituto de Ecologia, 176." type="book" year="1982">Halffter &amp; Edmonds (1982)</bibRefCitation>
, relying on information provided by G. T. Fincher, asserted that brood balls of
<taxonomicName authorityName="Harold" authorityYear="1868" box="[681,778,1730,1751]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="992" phylum="Arthropoda" rank="genus">
<emphasis box="[681,778,1730,1751]" italics="true" pageId="19" pageNumber="992">Bolbites</emphasis>
</taxonomicName>
apparently differ from those of the other Phanaeina by lacking a soil covering. Philips
<emphasis box="[541,589,1791,1813]" italics="true" pageId="19" pageNumber="992">et al</emphasis>
. (2004) repeated that information, but incorrectly mentioned
<bibRefCitation author="Cabrera Walsh G &amp; Gandolfo D" pageId="19" pageNumber="992" pagination="581 - 588" refId="ref46837" refString="Cabrera Walsh G, Gandolfo D. 1996. Nidification of thirteen common Argentine dung beetles (Scarabaeidae: Scarabaeinae). Annals of the Entomological Society of America 89: 581 - 588." type="journal article" year="1996">Cabrera Walsh &amp; Gandolfo (1996)</bibRefCitation>
as reference. In reality, all the authors who have observed
<taxonomicName authorityName="Harold" authorityYear="1868" box="[544,639,1883,1904]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="992" phylum="Arthropoda" rank="genus">
<emphasis box="[544,639,1883,1904]" italics="true" pageId="19" pageNumber="992">Bolbites</emphasis>
</taxonomicName>
brood balls at first-hand have stated exactly the opposite: that they are entirely covered by a thick layer of soil or cement like those of the other Phanaeina (Judulien apud:
<bibRefCitation author="Laloy L" box="[899,1030,289,311]" pageId="19" pageNumber="992" pagination="169 - 171" refId="ref51055" refString="Laloy L. 1902. Moeurs des coleopteres coprophages, specialement de La Plata. Le Naturaliste, 2 e Serie 16: 169 - 171." type="journal article" year="1902">Laloy, 1902</bibRefCitation>
;
<bibRefCitation author="Cabrera Walsh G &amp; Gandolfo D" box="[1044,1428,289,311]" pageId="19" pageNumber="992" pagination="581 - 588" refId="ref46837" refString="Cabrera Walsh G, Gandolfo D. 1996. Nidification of thirteen common Argentine dung beetles (Scarabaeidae: Scarabaeinae). Annals of the Entomological Society of America 89: 581 - 588." type="journal article" year="1996">Cabrera Walsh &amp; Gandolfo, 1996</bibRefCitation>
).
<bibRefCitation author="Fabre J-H" box="[827,972,319,341]" pageId="19" pageNumber="992" refId="ref48519" refString="Fabre J-H. 1899. Souvenirs entomologiques (sixieme serie). Etudes sur l'instict et les moeurs des insectes. Paris: Librairie Ch. Delagrave, 418." type="book" year="1899">Fabre (1899</bibRefCitation>
,
<bibRefCitation author="Fabre J-H" box="[988,1046,320,341]" pageId="19" pageNumber="992" refId="ref48549" refString="Fabre J-H. 1919. The glow-worm and other beetles. Toronto: McClelland &amp; Stewart, 488." type="book" year="1919">1919</bibRefCitation>
), utilizing information provided by Judulien (
<bibRefCitation author="Fabre J-H" box="[987,1175,350,372]" pageId="19" pageNumber="992" refId="ref48519" refString="Fabre J-H. 1899. Souvenirs entomologiques (sixieme serie). Etudes sur l'instict et les moeurs des insectes. Paris: Librairie Ch. Delagrave, 418." type="book" year="1899">Fabre, 1899: 70</bibRefCitation>
;
<bibRefCitation author="Fabre J-H" box="[1188,1307,350,372]" pageId="19" pageNumber="992" refId="ref48549" refString="Fabre J-H. 1919. The glow-worm and other beetles. Toronto: McClelland &amp; Stewart, 488." type="book" year="1919">1919: 238</bibRefCitation>
), described in vivid prose how this external layer and the rest of the brood ball are built by
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[1187,1328,411,433]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="992" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[1187,1328,411,433]" italics="true" pageId="19" pageNumber="992">B. onitoides</emphasis>
</taxonomicName>
.
<bibRefCitation author="Cabrera Walsh G &amp; Gandolfo D" pageId="19" pageNumber="992" pagination="581 - 588" refId="ref46837" refString="Cabrera Walsh G, Gandolfo D. 1996. Nidification of thirteen common Argentine dung beetles (Scarabaeidae: Scarabaeinae). Annals of the Entomological Society of America 89: 581 - 588." type="journal article" year="1996">Cabrera Walsh &amp; Gandolfo (1996)</bibRefCitation>
additionally found that all the
<taxonomicName box="[876,1002,473,495]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="1032" phylum="Arthropoda" rank="tribe" tribe="Phanaeini">Phanaeini</taxonomicName>
species studied by them, including
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[827,970,503,525]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="992" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[827,970,503,525]" italics="true" pageId="19" pageNumber="992">B. onitoides</emphasis>
</taxonomicName>
, leave a hole in the top of the cement covering of the brood ball egg chamber, an opening that presumably functions for aeration of the egg, something that had already been noted by
<bibRefCitation author="Judulien F" pageId="19" pageNumber="992" pagination="371 - 380" refId="ref50626" refString="Judulien F. 1899. Quelques notes sur plusieurs coprophages de Buenos Aires. Revista del Museo de La Plata 9: 371 - 380. [Frere Judulien was the religious name of the French- Argentinian entomologist Jean (Juan) Brethes (1871 - 1928) and it was with this name he signed his early works (Doello- Jurado, 1928)]." type="journal article" year="1899">Judulien (1899)</bibRefCitation>
and repeated by
<bibRefCitation author="Fabre J-H" box="[1141,1294,626,648]" pageId="19" pageNumber="992" refId="ref48519" refString="Fabre J-H. 1899. Souvenirs entomologiques (sixieme serie). Etudes sur l'instict et les moeurs des insectes. Paris: Librairie Ch. Delagrave, 418." type="book" year="1899">Fabre (1899</bibRefCitation>
,
<bibRefCitation author="Fabre J-H" box="[1315,1375,626,647]" pageId="19" pageNumber="992" refId="ref48549" refString="Fabre J-H. 1919. The glow-worm and other beetles. Toronto: McClelland &amp; Stewart, 488." type="book" year="1919">1919</bibRefCitation>
) and
<bibRefCitation author="Laloy L" box="[827,989,657,679]" pageId="19" pageNumber="992" pagination="169 - 171" refId="ref51055" refString="Laloy L. 1902. Moeurs des coleopteres coprophages, specialement de La Plata. Le Naturaliste, 2 e Serie 16: 169 - 171." type="journal article" year="1902">Laloy (1902)</bibRefCitation>
. The hole is subsequently covered with loose dung fibres and a thin layer of cement, but this porous, loose covering does not obstruct the air passage. This cement covering of the egg chamber and its top aeration hole are probable synapomorphies of the entire phanaeine clade (i.e. Phanaeina + Gromphadina) (
<bibRefCitation author="Cabrera Walsh G &amp; Gandolfo D" pageId="19" pageNumber="992" pagination="581 - 588" refId="ref46837" refString="Cabrera Walsh G, Gandolfo D. 1996. Nidification of thirteen common Argentine dung beetles (Scarabaeidae: Scarabaeinae). Annals of the Entomological Society of America 89: 581 - 588." type="journal article" year="1996">Cabrera Walsh &amp; Gandolfo, 1996</bibRefCitation>
). Unfortunately, no work has so far described the roles played by each sex of
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[1307,1443,902,924]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="992" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[1307,1443,902,924]" italics="true" pageId="19" pageNumber="992">B. onitoides</emphasis>
</taxonomicName>
during the construction of the nest, its provisioning with food and the preparation of the brood ball. Even so, it is reasonable to imagine that the species should follow the same pattern of the other phanaeines as discussed by
<bibRefCitation author="Halffter G &amp; Edmonds WD" box="[981,1300,1055,1077]" pageId="19" pageNumber="992" refId="ref49587" refString="Halffter G, Edmonds WD. 1982. The nesting behavior of dung beetles (Scarabaeinae). An ecological and evolutive approach. Mexico: Instituto de Ecologia, 176." type="book" year="1982">Halffter &amp; Edmonds (1982)</bibRefCitation>
.
</paragraph>
<paragraph blockId="19.[827,1444,197,1323]" pageId="19" pageNumber="992">
Finally,
<bibRefCitation author="Martinez A" box="[945,1132,1086,1108]" pageId="19" pageNumber="992" pagination="1 - 126" refId="ref51548" refString="Martinez A. 1959. Catalogo de los Scarabaeidae Argentinos. Revista del Museo Argentino de Ciencias Naturales ' Bernardino Rivadavia', Ciencias Zoologicas 5: 1 - 126." type="journal article" year="1959">Martínez (1959)</bibRefCitation>
reported that
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[1302,1437,1086,1108]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="992" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[1302,1437,1086,1108]" italics="true" pageId="19" pageNumber="992">B. onitoides</emphasis>
</taxonomicName>
, like many other dung beetles, is an intermediate host for the acanthocephalan worm
<emphasis italics="true" pageId="19" pageNumber="992">Macracanthorhynchus hirudinaceus</emphasis>
(Pallas, 1781), an intestinal parasite of mammals.
<bibRefCitation author="Krantz GW" box="[952,1112,1209,1231]" pageId="19" pageNumber="992" pagination="1 - 89" refId="ref50988" refString="Krantz GW. 1967. A review of the genus Holocelaeno Berlese, 1910 (Acarina: Macrochelidae). Acarologia 9 (Suppl.): 1 - 89." type="journal article" year="1967">Krantz (1967)</bibRefCitation>
, in turn, asserted that mites (Acari) of the species
<emphasis box="[1082,1341,1239,1260]" italics="true" pageId="19" pageNumber="992">Holocelaeno hypocrita</emphasis>
Berlese, 1917 and
<emphasis box="[944,1082,1270,1292]" italics="true" pageId="19" pageNumber="992">H. jugulans</emphasis>
Berlese, 1917 are phoretics of
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[827,958,1301,1322]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="992" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[827,958,1301,1322]" italics="true" pageId="19" pageNumber="992">B. onitoides</emphasis>
</taxonomicName>
individuals in
<collectingCountry box="[1128,1247,1301,1323]" name="Argentina" pageId="19" pageNumber="992">Argentina</collectingCountry>
.
</paragraph>
<paragraph blockId="19.[827,1442,1363,1416]" pageId="19" pageNumber="992">
<emphasis box="[827,1062,1363,1384]" italics="true" pageId="19" pageNumber="992">Material examined:</emphasis>
Altogether
<specimenCount box="[1212,1335,1363,1385]" pageId="19" pageNumber="992" type="male">698 males</specimenCount>
and
<specimenCount pageId="19" pageNumber="992" type="female">506 females</specimenCount>
.
</paragraph>
<paragraph blockId="19.[827,1443,1456,1908]" lastBlockId="20.[809,1426,197,587]" lastPageId="20" lastPageNumber="993" pageId="19" pageNumber="992">
<emphasis box="[827,1104,1457,1478]" italics="true" pageId="19" pageNumber="992">Bicoloured specimens:</emphasis>
<materialsCitation ID-GBIF-Occurrence="3704841462" collectionCode="MFNB" country="Argentina" location="ARGENTINA" pageId="19" pageNumber="992" specimenCount="1" specimenCount-female="1">
<emphasis bold="true" box="[1115,1301,1456,1478]" pageId="19" pageNumber="992">
<collectingCountry box="[1115,1295,1456,1478]" name="Argentina" pageId="19" pageNumber="992">ARGENTINA</collectingCountry>
:
</emphasis>
no further date
<specimenCount box="[910,1039,1487,1509]" pageId="19" pageNumber="992" type="female">one female</specimenCount>
(
<collectionCode box="[1056,1138,1487,1509]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="19" pageNumber="992">MFNB</collectionCode>
)
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3704841320" collectionCode="SDEI" country="Argentina" location="Kraatz" pageId="19" pageNumber="992" specimenCount="1" specimenCount-female="1">
and
<specimenCount box="[1208,1337,1487,1509]" pageId="19" pageNumber="992" type="female">one female</specimenCount>
(
<collectionCode box="[1355,1420,1487,1508]" pageId="19" pageNumber="992">SDEI</collectionCode>
ex
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC13FFEF3D6708E5D890F9F0" box="[863,946,1518,1539]" country="Argentina" name="Kraatz" pageId="19" pageNumber="992">Kraatz</location>
collection)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841354" collectionCode="MNHN" country="Argentina" location="BUENOS AIRES" pageId="19" pageNumber="992" specimenCount="1" specimenCount-male="1" stateProvince="Ciudad Autonoma de Buenos Aires">
<emphasis box="[1094,1308,1518,1539]" italics="true" pageId="19" pageNumber="992">
<collectingRegion box="[1094,1302,1518,1539]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="19" pageNumber="992">BUENOS AIRES</collectingRegion>
:
</emphasis>
no further data
<specimenCount box="[908,1011,1549,1571]" pageId="19" pageNumber="992" type="male">one male</specimenCount>
(
<collectionCode box="[1027,1116,1549,1571]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="19" pageNumber="992">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841408" collectionCode="MLPA" collectorName="Richter" country="Argentina" pageId="19" pageNumber="992" specimenCount="1" specimenCount-male="1">
no specific locality or date,
<collectorName box="[827,916,1579,1601]" pageId="19" pageNumber="992">Richter</collectorName>
leg.
<specimenCount box="[1000,1109,1579,1601]" pageId="19" pageNumber="992" type="male">one male</specimenCount>
(
<collectionCode box="[1126,1207,1579,1601]" pageId="19" pageNumber="992">MLPA</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841503" collectingDate="1942-02" collectionCode="BDGC" country="Argentina" county="Puan" location="Est. Felipe Sola" municipality="Felipe Sola" pageId="19" pageNumber="992" specimenCount="1" specimenCount-female="1">
<collectingCounty box="[1228,1293,1580,1601]" pageId="19" pageNumber="992">Puan</collectingCounty>
,
<collectingMunicipality box="[1307,1442,1579,1601]" pageId="19" pageNumber="992">Felipe Solá</collectingMunicipality>
(
<emphasis box="[841,1023,1610,1632]" italics="true" pageId="19" pageNumber="992">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC13FFEF3D710B40D8DDF9AC" box="[841,1023,1610,1632]" country="Argentina" county="Puan" municipality="Felipe Sola" name="Est. Felipe Sola" pageId="19" pageNumber="992">Est. Felipe Solá</location>
</emphasis>
),
<date box="[1052,1139,1610,1631]" pageId="19" pageNumber="992" value="1942-02">
<collectingDate box="[1052,1139,1610,1631]" pageId="19" pageNumber="992" value="1942-02">II.1942</collectingDate>
</date>
, no collector
<specimenCount box="[1316,1442,1610,1632]" pageId="19" pageNumber="992" type="female">one female</specimenCount>
(
<collectionCode box="[834,910,1641,1662]" pageId="19" pageNumber="992">BDGC</collectionCode>
ex
<collectorName box="[972,1111,1641,1663]" pageId="19" pageNumber="992">A. Martínez</collectorName>
and
<collectorName box="[1170,1243,1641,1662]" pageId="19" pageNumber="992">Henry</collectorName>
&amp;
<collectorName box="[1279,1443,1641,1663]" pageId="19" pageNumber="992">Anne Howden</collectorName>
collections)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841432" collectingDate="1941-02" collectionCode="DZUP" collectorName="F. Justus Jor" country="Argentina" county="San Isidro" location="Boulogne'" municipality="Boulogne Sur Mer" pageId="19" pageNumber="992" specimenCount="1" specimenCount-female="1">
<collectingCounty box="[965,1083,1671,1693]" pageId="19" pageNumber="992">San Isidro</collectingCounty>
,
<collectingMunicipality box="[1093,1299,1671,1693]" pageId="19" pageNumber="992">Boulogne Sur Mer</collectingMunicipality>
(
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC13FFEF3B1C0B8CDEB5F96E" box="[1316,1431,1671,1693]" country="Argentina" county="San Isidro" municipality="Boulogne Sur Mer" name="Boulogne'" pageId="19" pageNumber="992">Boulogne</location>
),
<date box="[827,910,1702,1723]" pageId="19" pageNumber="992" value="1941-02">
<collectingDate box="[827,910,1702,1723]" pageId="19" pageNumber="992" value="1941-02">II.1941</collectingDate>
</date>
, no collector
<specimenCount box="[1076,1198,1702,1724]" pageId="19" pageNumber="992" type="female">one female</specimenCount>
(
<collectionCode box="[1211,1283,1702,1723]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="19" pageNumber="992" type="University or college">DZUP</collectionCode>
ex
<collectorName pageId="19" pageNumber="992">F. Justus Jor</collectorName>
collection)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841525" collectionCode="MACN" collectorName="S. R. Castillo" country="Argentina" location="CIUDAD AUTONOMA DE BUENOS AIRES" pageId="19" pageNumber="992" specimenCount="1" specimenCount-female="1" stateProvince="Ciudad Autonoma de Buenos Aires">
<emphasis italics="true" pageId="19" pageNumber="992">
<collectingRegion country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="19" pageNumber="992">CIUDAD AUTONOMA DE BUENOS AIRES</collectingRegion>
:
</emphasis>
no specific locality or date,
<collectorName box="[1243,1395,1763,1785]" pageId="19" pageNumber="992">S. R. Castillo</collectorName>
leg.
<specimenCount box="[848,974,1794,1816]" pageId="19" pageNumber="992" type="female">one female</specimenCount>
(
<collectionCode box="[989,1073,1794,1816]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="19" pageNumber="992" type="Museum">MACN</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841415" collectingDate="1919-10-19" collectionCode="MLPA" country="Argentina" location="Nunez" pageId="19" pageNumber="992" specimenCount="1" specimenCount-female="1">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC13FFEF3A7C0A09DFB3F8EB" box="[1092,1169,1794,1816]" country="Argentina" name="Nunez" pageId="19" pageNumber="992">Núñez</location>
,
<date box="[1181,1298,1794,1815]" pageId="19" pageNumber="992" value="1919-10-19">
<collectingDate box="[1181,1298,1794,1815]" pageId="19" pageNumber="992" value="1919-10-19">19.X.1919</collectingDate>
</date>
, no collector
<specimenCount box="[849,975,1825,1847]" pageId="19" pageNumber="992" type="female">one female</specimenCount>
(
<collectionCode box="[991,1070,1825,1847]" pageId="19" pageNumber="992">MLPA</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841377" collectingDate="1993-01-13" collectionCode="CEMT" collectorName="Ugarte-Pena" country="Argentina" location="Charata" municipality="Chacabuco" pageId="19" pageNumber="992" specimenCount="1" specimenCount-male="1" stateProvince="Chaco">
<emphasis box="[1090,1191,1825,1846]" italics="true" pageId="19" pageNumber="992">
<collectingRegion box="[1090,1185,1825,1846]" country="Argentina" name="Chaco" pageId="19" pageNumber="992">CHACO</collectingRegion>
:
</emphasis>
<collectingMunicipality box="[1199,1327,1825,1847]" pageId="19" pageNumber="992">Chacabuco</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC13FFEF3B040A2ADEBCF8C4" box="[1340,1438,1825,1847]" country="Argentina" municipality="Chacabuco" name="Charata" pageId="19" pageNumber="992" stateProvince="Chaco">Charata</location>
,
<date box="[827,945,1855,1876]" pageId="19" pageNumber="992" value="1993-01-13">
<collectingDate box="[827,945,1855,1876]" pageId="19" pageNumber="992" value="1993-01-13">13.I.1993</collectingDate>
</date>
,
<collectorName box="[962,1120,1856,1877]" pageId="19" pageNumber="992">Ugarte-Peña</collectorName>
leg.
<specimenCount box="[1211,1325,1855,1877]" pageId="19" pageNumber="992" type="male">one male</specimenCount>
(
<collectionCode box="[1345,1429,1855,1877]" pageId="19" pageNumber="992">CEMT</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841479" collectionCode="BMNH" collectorName="S. Bolle" country="Argentina" lastPageId="20" lastPageNumber="993" pageId="19" pageNumber="992" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<collectorName box="[827,956,1886,1908]" pageId="19" pageNumber="992">Chacabuco</collectorName>
,
<collectorName box="[969,1068,1886,1908]" pageId="19" pageNumber="992">Charata</collectorName>
, 1995,
<collectorName box="[1153,1243,1886,1908]" pageId="19" pageNumber="992">S. Bolle</collectorName>
leg.
<specimenCount box="[1323,1442,1886,1908]" pageId="19" pageNumber="992" type="male">two males</specimenCount>
(both dissected) and
<specimenCount box="[379,515,197,219]" pageId="20" pageNumber="993" type="female">two females</specimenCount>
(
<collectionCode box="[529,614,197,219]" pageId="20" pageNumber="993">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841361" collectingDate="1995-01" collectionCode="NMPC, T" collectorName="Taylor &amp; S. Pokorny" country="Argentina" location="Charata" municipality="Chacabuco" pageId="20" pageNumber="993" specimenCount="1" specimenCount-male="1">
<collectingMunicipality box="[632,757,197,219]" pageId="20" pageNumber="993">Chacabuco</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC14FFE83EA90DEFDBD7FF09" box="[145,245,228,250]" country="Argentina" municipality="Chacabuco" name="Charata" pageId="20" pageNumber="993">Charata</location>
,
<date box="[258,335,228,250]" pageId="20" pageNumber="993" value="1995-01">
<collectingDate box="[258,335,228,250]" pageId="20" pageNumber="993" value="1995-01">I.1995</collectingDate>
</date>
, no collector
<specimenCount box="[517,625,228,250]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(dissected) (
<collectionCode box="[152,234,259,280]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="20" pageNumber="993">NMPC</collectionCode>
ex
<collectionCode box="[304,322,259,280]" name="Tavera, Department of Geology and Geophysics" pageId="20" pageNumber="993">T</collectionCode>
.
<collectorName box="[334,411,258,280]" pageId="20" pageNumber="993">Taylor</collectorName>
and
<collectorName box="[476,608,258,280]" pageId="20" pageNumber="993">S. Pokorný</collectorName>
collections)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841495" collectingDate="2001-12" collectionCode="BMNH" collectorName="S. Bolle &amp; J. Zidek" country="Argentina" location="Charata" municipality="Chacabuco" pageId="20" pageNumber="993" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<collectingMunicipality box="[145,275,289,311]" pageId="20" pageNumber="993">Chacabuco</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC14FFE83F180C2ADAA6FEC4" box="[288,388,289,311]" country="Argentina" municipality="Chacabuco" name="Charata" pageId="20" pageNumber="993">Charata</location>
,
<date box="[402,507,289,310]" pageId="20" pageNumber="993" value="2001-12">
<collectingDate box="[402,507,289,310]" pageId="20" pageNumber="993" value="2001-12">XII.2001</collectingDate>
</date>
,
<collectorName box="[521,612,289,311]" pageId="20" pageNumber="993">S. Bolle</collectorName>
leg.
<specimenCount pageId="20" pageNumber="993" type="male">seven males</specimenCount>
and
<specimenCount box="[264,403,320,342]" pageId="20" pageNumber="993" type="female">four females</specimenCount>
(
<collectionCode box="[414,497,320,341]" pageId="20" pageNumber="993">BMNH</collectionCode>
ex
<collectorName box="[549,635,320,342]" pageId="20" pageNumber="993">J. Zídek</collectorName>
collection)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841568" box="[145,709,349,373]" collectionCode="MACN" country="Argentina" location="CORDOBA" pageId="20" pageNumber="993" specimenCount="1" specimenCount-male="1" stateProvince="CORDOBA">
<emphasis box="[145,282,349,373]" italics="true" pageId="20" pageNumber="993">
<collectingRegion box="[145,277,349,373]" country="Argentina" name="CORDOBA" pageId="20" pageNumber="993">CÓRDOBA</collectingRegion>
:
</emphasis>
no further data
<specimenCount box="[500,605,350,372]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[620,703,351,373]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="20" pageNumber="993" type="Museum">MACN</collectionCode>
)
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3704841311" collectionCode="MLPA" country="Argentina" pageId="20" pageNumber="993" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
and
<specimenCount box="[145,249,381,403]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
and
<specimenCount box="[308,433,381,403]" pageId="20" pageNumber="993" type="female">one female</specimenCount>
(
<collectionCode box="[448,526,381,403]" pageId="20" pageNumber="993">MLPA</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841424" collectingDate="1896-12" collectionCode="MACN" collectorName="Carlos Bruch" country="Argentina" location="Carlos Bruch" pageId="20" pageNumber="993" specimenCount="1" specimenCount-male="1">
no specific locality,
<date box="[145,250,412,434]" pageId="20" pageNumber="993" value="1896-12">
<collectingDate box="[145,250,412,434]" pageId="20" pageNumber="993" value="1896-12">XII.1896</collectingDate>
</date>
,
<collectorName box="[262,420,412,434]" pageId="20" pageNumber="993">Carlos Bruch</collectorName>
leg.
<specimenCount box="[499,605,412,434]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[621,703,412,433]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="20" pageNumber="993" type="Museum">MACN</collectionCode>
ex
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC14FFE83EA90CB1DA13FE23" box="[145,305,442,464]" country="Argentina" name="Carlos Bruch" pageId="20" pageNumber="993">Carlos Bruch</location>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841343" collectingDate="1978-12-09" collectionCode="FSCA" collectorName="Km W Jaime Peter' &amp; Woodruff &amp; Runnacles" country="Argentina" county="Jaime Perez" location="Rt." municipality="Jaime Peter" pageId="20" pageNumber="993" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<collectingCounty box="[452,602,442,465]" pageId="20" pageNumber="993">Jaime Pérez</collectingCounty>
,
<collectingMunicipality box="[616,760,442,464]" pageId="20" pageNumber="993">Jaime Peter</collectingMunicipality>
(
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC14FFE83EA60CD2DBE3FE1D" box="[158,193,473,494]" country="Argentina" county="Jaime Perez" municipality="Jaime Peter" name="Rt." pageId="20" pageNumber="993">Rt.</location>
16, 6
<collectorName box="[254,482,473,495]" pageId="20" pageNumber="993">Km W Jaime Peter</collectorName>
),
<date box="[502,639,473,494]" pageId="20" pageNumber="993" value="1978-12-09">
<collectingDate box="[502,639,473,494]" pageId="20" pageNumber="993" value="1978-12-09">09.XII.1978</collectingDate>
</date>
,
<collectorName box="[650,757,473,495]" pageId="20" pageNumber="993">Woodruff</collectorName>
,
<collectorName box="[145,266,504,526]" pageId="20" pageNumber="993">Runnacles</collectorName>
and Cordo leg.
<specimenCount box="[471,576,504,526]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
and
<specimenCount box="[635,760,504,526]" pageId="20" pageNumber="993" type="female">one female</specimenCount>
(
<collectionCode box="[152,223,535,557]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="20" pageNumber="993" type="Museum">FSCA</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841409" collectingDate="1976-11-18" collectionCode="CMNC" collectorName="G. J. Williner" country="Argentina" county="Punilla" location="Francois Genier" municipality="San Esteban" pageId="20" pageNumber="993" specimenCount="1" specimenCount-male="1">
<collectingCounty box="[239,324,534,556]" pageId="20" pageNumber="993">Punilla</collectingCounty>
,
<collectingMunicipality box="[333,477,534,556]" pageId="20" pageNumber="993">San Esteban</collectingMunicipality>
,
<date box="[487,612,534,556]" pageId="20" pageNumber="993" value="1976-11-18">
<collectingDate box="[487,612,534,556]" pageId="20" pageNumber="993" value="1976-11-18">18.XI.1976</collectingDate>
</date>
,
<collectorName box="[621,761,534,556]" pageId="20" pageNumber="993">G.J. Williner</collectorName>
leg.
<specimenCount box="[207,308,565,587]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[319,399,565,586]" pageId="20" pageNumber="993">CMNC</collectionCode>
ex
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC14FFE83FFE0F3ED959FDB8" box="[454,635,565,587]" country="Argentina" county="Punilla" municipality="San Esteban" name="Francois Genier" pageId="20" pageNumber="993">François Génier</location>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841304" collectingDate="1974-02" collectionCode="CMNC" country="Argentina" location="San Jose de la Dormida" municipality="Tulumba" pageId="20" pageNumber="993" specimenCount="1" specimenCount-male="1">
<collectingMunicipality box="[145,251,596,618]" pageId="20" pageNumber="993">Tulumba</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC14FFE83F3C0F5FD932FD99" box="[260,528,596,618]" country="Argentina" municipality="Tulumba" name="San Jose de la Dormida" pageId="20" pageNumber="993">San José de la Dormida</location>
,
<date box="[538,622,596,617]" pageId="20" pageNumber="993" value="1974-02">
<collectingDate box="[538,622,596,617]" pageId="20" pageNumber="993" value="1974-02">II.1974</collectingDate>
</date>
, no collector
<specimenCount box="[171,288,626,648]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[308,399,627,649]" pageId="20" pageNumber="993">CMNC</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841367" collectingDate="1987-12" collectionCode="NMPC, T" collectorName="Taylor &amp; S. Pokorny" country="Argentina" location="Ituzaingo" pageId="20" pageNumber="993" specimenCount="1" specimenCount-male="1" stateProvince="Corrientes">
<emphasis box="[424,619,627,648]" italics="true" pageId="20" pageNumber="993">
<collectingRegion box="[424,613,627,648]" country="Argentina" name="Corrientes" pageId="20" pageNumber="993">CORRIENTES</collectingRegion>
:
</emphasis>
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC14FFE83C400F79D9D1FD7B" box="[632,755,626,648]" country="Argentina" name="Ituzaingo" pageId="20" pageNumber="993" stateProvince="Corrientes">Ituzaingó</location>
,
<date box="[145,245,657,678]" pageId="20" pageNumber="993" value="1987-12">
<collectingDate box="[145,245,657,678]" pageId="20" pageNumber="993" value="1987-12">XII.1987</collectingDate>
</date>
, no collector
<specimenCount box="[415,518,657,679]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(dissected) (
<collectionCode box="[661,740,657,678]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="20" pageNumber="993">NMPC</collectionCode>
ex
<collectionCode box="[175,192,688,709]" name="Tavera, Department of Geology and Geophysics" pageId="20" pageNumber="993">T</collectionCode>
.
<collectorName box="[200,273,688,710]" pageId="20" pageNumber="993">Taylor</collectorName>
and
<collectorName box="[328,448,688,710]" pageId="20" pageNumber="993">S. Pokorný</collectorName>
collections)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841395" collectingDate="1962-01" collectionCode="MLPA" country="Argentina" location="Pronunciamiento" municipality="Uruguay" pageId="20" pageNumber="993" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Entre Rios">
<emphasis box="[593,760,686,710]" italics="true" pageId="20" pageNumber="993">
<collectingRegion box="[593,754,686,710]" country="Argentina" name="Entre Rios" pageId="20" pageNumber="993">ENTRE RÍOS</collectingRegion>
:
</emphasis>
<collectingMunicipality box="[145,247,719,740]" pageId="20" pageNumber="993">Uruguay</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC14FFE83F3A0FC5DAE9FD17" box="[258,459,718,740]" country="Argentina" municipality="Uruguay" name="Pronunciamiento" pageId="20" pageNumber="993" stateProvince="Entre Rios">Pronunciamiento</location>
,
<date box="[472,546,718,740]" pageId="20" pageNumber="993" value="1962-01">
<collectingDate box="[472,546,718,740]" pageId="20" pageNumber="993" value="1962-01">I.1962</collectingDate>
</date>
, no collector
<specimenCount pageId="20" pageNumber="993" type="male">one male</specimenCount>
and
<specimenCount box="[260,385,749,771]" pageId="20" pageNumber="993" type="female">one female</specimenCount>
(
<collectionCode box="[400,479,749,771]" pageId="20" pageNumber="993">MLPA</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841385" collectingDate="1966-02" collectionCode="TAMU" collectorName="J. Foerster &amp; W. D. Edmonds" country="Argentina" location="Dos de Mayo" municipality="Cainguas" pageId="20" pageNumber="993" specimenCount="1" specimenCount-male="1" stateProvince="MISIONES">
<emphasis box="[496,638,749,770]" italics="true" pageId="20" pageNumber="993">
<collectingRegion box="[496,631,749,770]" country="Argentina" name="MISIONES" pageId="20" pageNumber="993">MISIONES</collectingRegion>
:
</emphasis>
<collectingMunicipality box="[646,756,749,771]" pageId="20" pageNumber="993">Cainguás</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC14FFE83EA90E07DA11FCD2" box="[145,307,780,802]" country="Argentina" municipality="Cainguas" name="Dos de Mayo" pageId="20" pageNumber="993" stateProvince="MISIONES">Dos de Mayo</location>
,
<date box="[322,413,780,802]" pageId="20" pageNumber="993" value="1966-02">
<collectingDate box="[322,413,780,802]" pageId="20" pageNumber="993" value="1966-02">II.1966</collectingDate>
</date>
,
<collectorName box="[428,561,780,801]" pageId="20" pageNumber="993">J. Foerster</collectorName>
leg.
<specimenCount box="[649,760,780,802]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[152,232,811,832]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="20" pageNumber="993" type="Herbarium">TAMU</collectionCode>
ex
<collectorName box="[295,465,810,832]" pageId="20" pageNumber="993">W.D. Edmonds</collectorName>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841372" collectingDate="1995-03" collectionCode="BDGC" country="Argentina" location="El Dorado'" municipality="Eldorado" pageId="20" pageNumber="993" specimenCount="1" specimenCount-male="1">
<collectingMunicipality box="[607,713,810,832]" pageId="20" pageNumber="993">Eldorado</collectingMunicipality>
(
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC14FFE83CE60E21DBC9FCAC" country="Argentina" municipality="Eldorado" name="El Dorado'" pageId="20" pageNumber="993">El Dorado</location>
),
<date box="[253,346,841,863]" pageId="20" pageNumber="993" value="1995-03">
<collectingDate box="[253,346,841,863]" pageId="20" pageNumber="993" value="1995-03">III.1995</collectingDate>
</date>
, no collector
<specimenCount box="[513,615,841,863]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[628,704,841,863]" pageId="20" pageNumber="993">BDGC</collectionCode>
)
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3704841558" collectionCode="FSCA" country="Argentina" pageId="20" pageNumber="993" specimenCount="1" specimenCount-female="1">
and
<specimenCount box="[145,267,872,894]" pageId="20" pageNumber="993" type="female">one female</specimenCount>
(
<collectionCode box="[280,351,872,894]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="20" pageNumber="993" type="Museum">FSCA</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841545" collectionCode="MFNB" collectorName="Brauer" country="Argentina" location="SALTA" pageId="20" pageNumber="993" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Salta">
<emphasis box="[369,458,872,893]" italics="true" pageId="20" pageNumber="993">
<collectingRegion box="[369,452,872,893]" country="Argentina" name="Salta" pageId="20" pageNumber="993">SALTA</collectingRegion>
:
</emphasis>
no specific locality or date,
<collectorName box="[145,229,903,924]" pageId="20" pageNumber="993">Brauer</collectorName>
leg.
<specimenCount box="[312,454,902,924]" pageId="20" pageNumber="993" type="male">three males</specimenCount>
and
<specimenCount box="[517,648,902,924]" pageId="20" pageNumber="993" type="female">one female</specimenCount>
(
<collectionCode box="[664,748,902,924]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="20" pageNumber="993">MFNB</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841538" collectingDate="1987-01" collectionCode="CMNC" country="Argentina" county="La Vina" location="Km" municipality="Ruta" pageId="20" pageNumber="993" specimenCount="1" specimenCount-male="1">
<collectingCounty box="[145,242,933,955]" pageId="20" pageNumber="993">La Viña</collectingCounty>
,
<collectingMunicipality box="[255,313,933,954]" pageId="20" pageNumber="993">Ruta</collectingMunicipality>
68,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC14FFE83F560EAEDABAFC49" box="[366,408,933,954]" country="Argentina" county="La Vina" municipality="Ruta" name="Km" pageId="20" pageNumber="993">Km</location>
89,
<date box="[461,537,933,954]" pageId="20" pageNumber="993" value="1987-01">
<collectingDate box="[461,537,933,954]" pageId="20" pageNumber="993" value="1987-01">I.1987</collectingDate>
</date>
, no collector
<specimenCount pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[217,302,964,986]" pageId="20" pageNumber="993">CMNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841472" collectingDate="1991-03" collectionCode="CMNC" country="Argentina" location="Ruta" municipality="La Vina" pageId="20" pageNumber="993" specimenCount="1" specimenCount-male="1">
<collectingMunicipality box="[322,416,964,986]" pageId="20" pageNumber="993">La Viña</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC14FFE83F950ECFDAC4FC2A" box="[429,486,964,985]" country="Argentina" municipality="La Vina" name="Ruta" pageId="20" pageNumber="993">Ruta</location>
68, km 98,
<date box="[624,720,964,985]" pageId="20" pageNumber="993" value="1991-03">
<collectingDate box="[624,720,964,985]" pageId="20" pageNumber="993" value="1991-03">III.1991</collectingDate>
</date>
, no collector
<specimenCount box="[269,372,994,1016]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[387,468,994,1016]" pageId="20" pageNumber="993">CMNC</collectionCode>
)
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3704841381" collectionCode="CNCI" country="Argentina" pageId="20" pageNumber="993" specimenCount="1" specimenCount-female="1">
and
<specimenCount box="[534,658,994,1016]" pageId="20" pageNumber="993" type="female">one female</specimenCount>
(
<collectionCode box="[673,740,994,1015]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="20" pageNumber="993">CNCI</collectionCode>
ex
<collectorName box="[175,309,1025,1047]" pageId="20" pageNumber="993">A. Martínez</collectorName>
and
<collectorName box="[362,434,1025,1046]" pageId="20" pageNumber="993">Henry</collectorName>
&amp;
<collectorName box="[464,623,1025,1047]" pageId="20" pageNumber="993">Anne Howden</collectorName>
collections)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841307" collectingDate="1911-12-18" collectionCode="BMNH" collectorName="G. E. Bryant" country="Argentina" location="San Javier River" municipality="Estancia La Noria" pageId="20" pageNumber="993" specimenCount="1" specimenCount-female="1" stateProvince="Santa Fe">
<emphasis box="[145,290,1056,1077]" italics="true" pageId="20" pageNumber="993">
<collectingRegion box="[145,285,1056,1077]" country="Argentina" name="Santa Fe" pageId="20" pageNumber="993">SANTA FE</collectingRegion>
:
</emphasis>
<collectingMunicipality box="[301,532,1056,1078]" pageId="20" pageNumber="993">Estancia La Noria</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC14FFE83C1B092BD9D1FBC5" box="[547,755,1056,1078]" country="Argentina" municipality="Estancia La Noria" name="San Javier River" pageId="20" pageNumber="993" stateProvince="Santa Fe">San Javier River</location>
,
<date box="[145,283,1086,1107]" pageId="20" pageNumber="993" value="1911-12-18">
<collectingDate box="[145,283,1086,1107]" pageId="20" pageNumber="993" value="1911-12-18">18.XII.1911</collectingDate>
</date>
,
<collectorName box="[296,443,1086,1108]" pageId="20" pageNumber="993">G. E. Bryant</collectorName>
leg.
<specimenCount box="[520,647,1086,1108]" pageId="20" pageNumber="993" type="female">one female</specimenCount>
(
<collectionCode box="[662,748,1086,1108]" pageId="20" pageNumber="993">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841411" collectingDate="1912-01-09" collectionCode="BMNH" collectorName="G. E. Bryant" country="Argentina" location="San Javier River" municipality="Estancia La Noria" pageId="20" pageNumber="993" specimenCount="1" specimenCount-female="1">
<collectingMunicipality box="[145,361,1117,1139]" pageId="20" pageNumber="993">Estancia La Noria</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC14FFE83F4D0956D918FB80" box="[373,570,1117,1139]" country="Argentina" municipality="Estancia La Noria" name="San Javier River" pageId="20" pageNumber="993">San Javier River</location>
,
<date box="[582,692,1117,1138]" pageId="20" pageNumber="993" value="1912-01-09">
<collectingDate box="[582,692,1117,1138]" pageId="20" pageNumber="993" value="1912-01-09">09.I.1912</collectingDate>
</date>
,
<collectorName pageId="20" pageNumber="993">G. E. Bryant</collectorName>
leg.
<specimenCount box="[306,433,1148,1170]" pageId="20" pageNumber="993" type="female">one female</specimenCount>
(
<collectionCode box="[449,536,1148,1170]" pageId="20" pageNumber="993">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841355" collectingDate="1980-11-19" collectingDateMax="1980-11-21" collectingDateMin="1980-11-19" collectionCode="FSCA" collectorName="R. Woodruff &amp; H. Cordo" country="Argentina" location="Arroyo Colastine" pageId="20" pageNumber="993" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC14FFE83C140977D9D6FB61" box="[556,756,1148,1170]" country="Argentina" name="Arroyo Colastine" pageId="20" pageNumber="993">Arroyo Colastine</location>
,
<locationDeviation box="[149,371,1178,1200]" location="Santa Fe'" pageId="20" pageNumber="993">
<quantity box="[149,224,1178,1200]" metricMagnitude="4" metricUnit="m" metricValue="5.3" pageId="20" pageNumber="993" unit="km" value="53.0">53 km</quantity>
S Santa Fe
</locationDeviation>
,
<date box="[383,555,1178,1199]" pageId="20" pageNumber="993" value="1980-11-19" valueMax="1980-11-21" valueMin="1980-11-19">
<collectingDate box="[383,555,1178,1199]" pageId="20" pageNumber="993" value="1980-11-19" valueMax="1980-11-21" valueMin="1980-11-19">1921.XI.1980</collectingDate>
</date>
,
<collectorName box="[569,708,1178,1200]" pageId="20" pageNumber="993">R. Woodruff</collectorName>
and
<collectorName box="[145,251,1209,1231]" pageId="20" pageNumber="993">H. Cordo</collectorName>
leg.
<specimenCount box="[332,454,1209,1231]" pageId="20" pageNumber="993" type="male">two males</specimenCount>
and
<specimenCount box="[516,658,1209,1231]" pageId="20" pageNumber="993" type="female">two females</specimenCount>
(
<collectionCode box="[675,748,1209,1231]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="20" pageNumber="993" type="Museum">FSCA</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841315" collectingDate="1924-10" collectionCode="BMNH" collectorName="K. J. Hayward" country="Argentina" location="Villa Ana" municipality="General Obligado" pageId="20" pageNumber="993" specimenCount="1" specimenCount-male="1">
<collectingMunicipality box="[145,357,1240,1262]" pageId="20" pageNumber="993">General Obligado</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC14FFE83F4A09D3DAC7FB1E" box="[370,485,1240,1262]" country="Argentina" municipality="General Obligado" name="Villa Ana" pageId="20" pageNumber="993">Villa Ana</location>
,
<date box="[499,583,1240,1261]" pageId="20" pageNumber="993" value="1924-10">
<collectingDate box="[499,583,1240,1261]" pageId="20" pageNumber="993" value="1924-10">X.1924</collectingDate>
</date>
,
<collectorName box="[597,760,1240,1262]" pageId="20" pageNumber="993">K.J. Hayward</collectorName>
leg.
<specimenCount box="[214,319,1270,1292]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[335,422,1270,1292]" pageId="20" pageNumber="993">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841430" collectionCode="SMF" collectorName="Rosario &amp; Laske" country="Argentina" pageId="20" pageNumber="993" specimenCount="1" specimenCount-female="1">
<collectorName box="[441,531,1270,1292]" pageId="20" pageNumber="993">Rosario</collectorName>
, no date,
<collectorName box="[644,714,1270,1292]" pageId="20" pageNumber="993">Laske</collectorName>
leg.
<specimenCount box="[167,296,1301,1323]" pageId="20" pageNumber="993" type="female">one female</specimenCount>
(
<collectionCode box="[313,374,1301,1323]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="20" pageNumber="993">SMF</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841510" collectionCode="MFNB" country="Argentina" location="SANTIAGO DEL ESTERO" pageId="20" pageNumber="993" specimenCount="1" specimenCount-male="1" stateProvince="Santiago del Estero">
<emphasis box="[395,723,1301,1323]" italics="true" pageId="20" pageNumber="993">
<collectingRegion box="[395,717,1301,1323]" country="Argentina" name="Santiago del Estero" pageId="20" pageNumber="993">SANTIAGO DEL ESTERO</collectingRegion>
:
</emphasis>
no further data
<specimenCount box="[310,412,1332,1354]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[426,507,1332,1354]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="20" pageNumber="993">MFNB</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841312" collectionCode="MLPA" collectorName="Wagner" country="Argentina" pageId="20" pageNumber="993" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
no specific locality or date,
<collectorName box="[206,294,1363,1384]" pageId="20" pageNumber="993">Wagner</collectorName>
leg.
<specimenCount box="[366,468,1362,1384]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
and
<specimenCount box="[524,659,1362,1384]" pageId="20" pageNumber="993" type="female">two females</specimenCount>
(
<collectionCode box="[673,750,1362,1384]" pageId="20" pageNumber="993">MLPA</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841302" collectingDate="1962-04" collectionCode="MLPA" country="Argentina" pageId="20" pageNumber="993" specimenCount="1" specimenCount-male="1">
no specific locality,
<date box="[373,464,1393,1415]" pageId="20" pageNumber="993" value="1962-04">
<collectingDate box="[373,464,1393,1415]" pageId="20" pageNumber="993" value="1962-04">IV.1962</collectingDate>
</date>
, no collector
<specimenCount box="[642,760,1393,1415]" pageId="20" pageNumber="993" type="male">two males</specimenCount>
(
<collectionCode box="[152,231,1424,1446]" pageId="20" pageNumber="993">MLPA</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841527" collectingDate="1943-03" collectionCode="DZUP" collectorName="F. Justus Jor" country="Argentina" location="Campo Gallo" municipality="Alberdi" pageId="20" pageNumber="993" specimenCount="1" specimenCount-male="1">
<collectingMunicipality box="[249,338,1424,1446]" pageId="20" pageNumber="993">Alberdi</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC14FFE83F67089BDADBFA55" box="[351,505,1424,1446]" country="Argentina" municipality="Alberdi" name="Campo Gallo" pageId="20" pageNumber="993">Campo Gallo</location>
,
<date box="[517,614,1424,1445]" pageId="20" pageNumber="993" value="1943-03">
<collectingDate box="[517,614,1424,1445]" pageId="20" pageNumber="993" value="1943-03">III.1943</collectingDate>
</date>
, no collector
<specimenCount box="[170,282,1454,1476]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[301,377,1454,1475]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="20" pageNumber="993" type="University or college">DZUP</collectionCode>
ex
<collectorName box="[452,616,1455,1476]" pageId="20" pageNumber="993">F. Justus Jor</collectorName>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841489" collectingDate="1943-03" collectionCode="CMNC" collectorName="Prosen" country="Argentina" location="Campo Gallo" municipality="Alberdi" pageId="20" pageNumber="993" specimenCount="1" specimenCount-male="1">
<collectingMunicipality box="[145,231,1485,1507]" pageId="20" pageNumber="993">Alberdi</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC14FFE83ECA08C6DAA5FA10" box="[242,391,1485,1507]" country="Argentina" municipality="Alberdi" name="Campo Gallo" pageId="20" pageNumber="993">Campo Gallo</location>
,
<date box="[402,495,1485,1506]" pageId="20" pageNumber="993" value="1943-03">
<collectingDate box="[402,495,1485,1506]" pageId="20" pageNumber="993" value="1943-03">III.1943</collectingDate>
</date>
,
<collectorName box="[507,586,1485,1506]" pageId="20" pageNumber="993">Prosen</collectorName>
leg.
<specimenCount box="[658,761,1485,1507]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[152,235,1516,1538]" pageId="20" pageNumber="993">CMNC</collectionCode>
)
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3704841325" collectionCode="DZUP, CMNC, ZSM" country="Argentina" pageId="20" pageNumber="993" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<specimenCount box="[252,374,1516,1538]" pageId="20" pageNumber="993" type="female">one female</specimenCount>
(
<collectionCode box="[387,459,1516,1537]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="20" pageNumber="993" type="University or college">DZUP</collectionCode>
ex
<collectionCode box="[517,597,1516,1538]" pageId="20" pageNumber="993">CMNC</collectionCode>
) and
<specimenCount box="[659,761,1516,1538]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
and
<specimenCount box="[195,320,1546,1568]" pageId="20" pageNumber="993" type="female">one female</specimenCount>
(
<collectionCode box="[335,393,1546,1568]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="20" pageNumber="993" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841425" collectingDate="1951-02" collectionCode="ZSM" collectorName="Prosen" country="Argentina" location="Campo Gallo" municipality="Alberdi" pageId="20" pageNumber="993" specimenCount="1" specimenCount-male="1">
<collectingMunicipality box="[411,498,1546,1568]" pageId="20" pageNumber="993">Alberdi</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC14FFE83FC60B01D9B7F9D3" box="[510,661,1546,1568]" country="Argentina" municipality="Alberdi" name="Campo Gallo" pageId="20" pageNumber="993">Campo Gallo</location>
,
<date box="[672,755,1546,1568]" pageId="20" pageNumber="993" value="1951-02">
<collectingDate box="[672,755,1546,1568]" pageId="20" pageNumber="993" value="1951-02">II.1951</collectingDate>
</date>
,
<collectorName box="[145,227,1577,1598]" pageId="20" pageNumber="993">Prosen</collectorName>
leg.
<specimenCount box="[307,415,1577,1599]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[432,492,1577,1599]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="20" pageNumber="993" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841423" collectingDate="2006-01" collectionCode="CEMT" collectorName="E. Abadie" country="Argentina" location="Frias" municipality="Choya" pageId="20" pageNumber="993" specimenCount="1" specimenCount-male="1">
<collectingMunicipality box="[512,589,1577,1599]" pageId="20" pageNumber="993">Choya</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC14FFE83C620B22D9B8F9CC" box="[602,666,1577,1599]" country="Argentina" municipality="Choya" name="Frias" pageId="20" pageNumber="993">Frías</location>
,
<date box="[680,756,1577,1599]" pageId="20" pageNumber="993" value="2006-01">
<collectingDate box="[680,756,1577,1599]" pageId="20" pageNumber="993" value="2006-01">I.2006</collectingDate>
</date>
,
<collectorName box="[145,265,1608,1630]" pageId="20" pageNumber="993">E. Abadie</collectorName>
leg.
<specimenCount box="[355,467,1608,1630]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[487,572,1608,1630]" pageId="20" pageNumber="993">CEMT</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841523" collectionCode="MLPA" collectorName="Wagner" country="Argentina" location="Salado River" pageId="20" pageNumber="993" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC14FFE83C6B0B43D9D1F9AD" box="[595,755,1608,1630]" country="Argentina" name="Salado River" pageId="20" pageNumber="993">Salado River</location>
, no date,
<collectorName box="[253,346,1639,1660]" pageId="20" pageNumber="993">Wagner</collectorName>
leg.
<specimenCount box="[434,546,1638,1660]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
and
<specimenCount box="[612,759,1638,1660]" pageId="20" pageNumber="993" type="female">two females</specimenCount>
(
<collectionCode box="[152,235,1669,1691]" pageId="20" pageNumber="993">MLPA</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841397" collectionCode="NHMB" collectorName="H. Marchand" country="Argentina" location="TUCUMAN" pageId="20" pageNumber="993" specimenCount="1" specimenCount-male="1" stateProvince="Tucuman">
<emphasis box="[257,408,1667,1691]" italics="true" pageId="20" pageNumber="993">
<collectingRegion box="[257,401,1667,1691]" country="Argentina" name="Tucuman" pageId="20" pageNumber="993">TUCUMÁN</collectingRegion>
:
</emphasis>
no further data
<specimenCount box="[649,760,1669,1691]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[152,236,1700,1721]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="20" pageNumber="993">NHMB</collectionCode>
ex
<collectorName box="[302,458,1700,1722]" pageId="20" pageNumber="993">H. Marchand</collectorName>
collection)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841548" collectingDate="1996-11" collectionCode="CNCI" country="Paraguay" location="Chaco" pageId="20" pageNumber="993" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<emphasis bold="true" box="[601,761,1699,1721]" pageId="20" pageNumber="993">
<collectingCountry box="[601,756,1699,1721]" name="Paraguay" pageId="20" pageNumber="993">PARAGUAY</collectingCountry>
:
</emphasis>
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC14FFE83EA90BC9DBFCF92B" box="[145,222,1730,1752]" country="Paraguay" name="Chaco" pageId="20" pageNumber="993">Chaco</location>
,
<date box="[238,338,1730,1752]" pageId="20" pageNumber="993" value="1996-11">
<collectingDate box="[238,338,1730,1752]" pageId="20" pageNumber="993" value="1996-11">XI.1996</collectingDate>
</date>
, no collector
<specimenCount box="[536,650,1730,1752]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
and
<specimenCount pageId="20" pageNumber="993" type="female">one female</specimenCount>
(
<collectionCode box="[240,311,1761,1783]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="20" pageNumber="993">CNCI</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841469" collectionCode="CEMT" collectorName="Trans-Chaco'" country="Paraguay" pageId="20" pageNumber="993" specimenCount="1" specimenCount-female="1">
<collectorName box="[335,494,1761,1783]" pageId="20" pageNumber="993">Trans-Chaco</collectorName>
, no further data
<specimenCount pageId="20" pageNumber="993" type="female">one female</specimenCount>
(
<collectionCode box="[245,329,1792,1814]" pageId="20" pageNumber="993">CEMT</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841466" collectingDate="1996-01-20" collectionCode="TAMU" collectorName="C. Aguilar &amp; W. D. Edmonds" country="Paraguay" location="Pioneros" municipality="Cruce" pageId="20" pageNumber="993" specimenCount="1" specimenCount-male="1" stateProvince="Presidente Hayes">
<emphasis box="[351,641,1792,1813]" italics="true" pageId="20" pageNumber="993">
<collectingRegion box="[351,634,1792,1813]" country="Paraguay" name="Presidente Hayes" pageId="20" pageNumber="993">PRESIDENTE HAYES</collectingRegion>
:
</emphasis>
<collectingMunicipality box="[652,723,1792,1813]" pageId="20" pageNumber="993">Cruce</collectingMunicipality>
de los
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC14FFE83E840A15DA05F8C7" box="[188,295,1822,1844]" country="Paraguay" municipality="Cruce" name="Pioneros" pageId="20" pageNumber="993" stateProvince="Presidente Hayes">Pioneros</location>
,
<date box="[310,425,1822,1844]" pageId="20" pageNumber="993" value="1996-01-20">
<collectingDate box="[310,425,1822,1844]" pageId="20" pageNumber="993" value="1996-01-20">20.I.1996</collectingDate>
</date>
,
<collectorName box="[440,565,1822,1844]" pageId="20" pageNumber="993">C. Aguilar</collectorName>
leg.
<specimenCount box="[651,760,1822,1844]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[152,232,1853,1874]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="20" pageNumber="993" type="Herbarium">TAMU</collectionCode>
ex
<collectorName box="[295,473,1853,1875]" pageId="20" pageNumber="993">W. D. Edmonds</collectorName>
collection)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841475" collectionCode="BMNH, MFNB" collectorName="Nevinson" country="Uruguay" location="URUGUAY" pageId="20" pageNumber="993" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<emphasis bold="true" box="[615,761,1853,1875]" pageId="20" pageNumber="993">
<collectingCountry box="[615,756,1853,1875]" name="Uruguay" pageId="20" pageNumber="993">URUGUAY</collectingCountry>
:
</emphasis>
no further data
<specimenCount box="[366,475,1883,1905]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[493,579,1884,1905]" pageId="20" pageNumber="993">BMNH</collectionCode>
ex
<collectorName box="[648,760,1883,1905]" pageId="20" pageNumber="993">Nevinson</collectorName>
collection) and
<specimenCount box="[981,1104,197,219]" pageId="20" pageNumber="993" type="female">one female</specimenCount>
(
<collectionCode box="[1119,1201,197,219]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="20" pageNumber="993">MFNB</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841514" collectingDate="1954-11-18" collectionCode="FSCA" collectorName="C. B. Carbonell &amp; E. N. Kellesvig-Waering" country="Uruguay" location="Arroyo de la Tavernada" pageId="20" pageNumber="993" specimenCount="1" specimenCount-male="1" stateProvince="Artigas">
<emphasis box="[1218,1341,197,218]" italics="true" pageId="20" pageNumber="993">
<collectingRegion box="[1218,1334,197,218]" country="Uruguay" name="Artigas" pageId="20" pageNumber="993">ARTIGAS</collectingRegion>
:
</emphasis>
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC14FFE83B7B0DCED8D5FF0A" country="Uruguay" name="Arroyo de la Tavernada" pageId="20" pageNumber="993" stateProvince="Artigas">Arroyo de la Tavernada</location>
(?),
<date box="[1076,1213,227,249]" pageId="20" pageNumber="993" value="1954-11-18">
<collectingDate box="[1076,1213,227,249]" pageId="20" pageNumber="993" value="1954-11-18">18.XI.1954</collectingDate>
</date>
,
<collectorName box="[1230,1423,227,249]" pageId="20" pageNumber="993">C. B. Carbonell</collectorName>
leg.
<specimenCount box="[880,987,258,280]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[1004,1074,258,279]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="20" pageNumber="993" type="Museum">FSCA</collectionCode>
ex
<collectorName box="[1140,1424,258,280]" pageId="20" pageNumber="993">E. N. Kellesvig-Waering</collectorName>
collection)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841520" collectionCode="MLPA" collectorName="Tremorelas" country="Uruguay" location="MONTEVIDEO" pageId="20" pageNumber="993" specimenCount="1" specimenCount-female="1" stateProvince="Montevideo">
<emphasis box="[936,1123,289,310]" italics="true" pageId="20" pageNumber="993">
<collectingRegion box="[936,1118,289,310]" country="Uruguay" name="Montevideo" pageId="20" pageNumber="993">MONTEVIDEO</collectingRegion>
:
</emphasis>
no specific locality or date,
<collectorName box="[809,947,319,341]" pageId="20" pageNumber="993">Tremorelas</collectorName>
leg.
<specimenCount box="[1032,1165,319,341]" pageId="20" pageNumber="993" type="female">one female</specimenCount>
(
<collectionCode box="[1182,1263,320,342]" pageId="20" pageNumber="993">MLPA</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841473" collectingDate="1908-02-26" collectionCode="MLPA" country="Uruguay" location="Costa Sur'" pageId="20" pageNumber="993" specimenCount="1" specimenCount-female="1">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC14FFE83B310C4BDEAEFEA6" box="[1289,1420,320,341]" country="Uruguay" name="Costa Sur'" pageId="20" pageNumber="993">Costa Sur</location>
,
<date box="[809,927,350,372]" pageId="20" pageNumber="993" value="1908-02-26">
<collectingDate box="[809,927,350,372]" pageId="20" pageNumber="993" value="1908-02-26">26.II.1908</collectingDate>
</date>
, no collector
<specimenCount box="[1091,1212,350,372]" pageId="20" pageNumber="993" type="female">one female</specimenCount>
(
<collectionCode box="[1226,1304,350,372]" pageId="20" pageNumber="993">MLPA</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841368" collectionCode="MLPA" country="Uruguay" pageId="20" pageNumber="993" specimenCount="1" specimenCount-female="1">
<emphasis bold="true" pageId="20" pageNumber="993">
<emphasis bold="true" italics="true" pageId="20" pageNumber="993">
<collectorName box="[1320,1425,350,372]" pageId="20" pageNumber="993">Dubious</collectorName>
data
</emphasis>
:
</emphasis>
<emphasis box="[885,1065,381,402]" italics="true" pageId="20" pageNumber="993">
<collectorName box="[885,1065,381,402]" pageId="20" pageNumber="993">Chacoand. Aula</collectorName>
</emphasis>
(or
<emphasis box="[1109,1162,381,402]" italics="true" pageId="20" pageNumber="993">
<collectorName box="[1109,1162,381,402]" pageId="20" pageNumber="993">Alba</collectorName>
</emphasis>
)
<specimenCount box="[1202,1323,381,403]" pageId="20" pageNumber="993" type="female">one female</specimenCount>
(
<collectionCode box="[1337,1415,381,403]" pageId="20" pageNumber="993">MLPA</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841532" collectionCode="MCZC, NHMW, OUMNH" country="Uruguay" location="No" pageId="20" pageNumber="993" specimenCount="4" specimenCount-female="2" specimenCount-male="2">
<emphasis bold="true" box="[809,917,411,433]" pageId="20" pageNumber="993">
<emphasis bold="true" box="[809,910,411,433]" italics="true" pageId="20" pageNumber="993">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC14FFE83D110C97D86EFE42" box="[809,844,412,433]" country="Uruguay" name="No" pageId="20" pageNumber="993">No</location>
data
</emphasis>
:
</emphasis>
<specimenCount box="[923,1026,411,433]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
[labelled with the
<emphasis box="[1238,1425,411,433]" italics="true" pageId="20" pageNumber="993">nomen in litteris</emphasis>
<emphasis box="[814,922,443,464]" italics="true" pageId="20" pageNumber="993">Escrapus</emphasis>
(?) /
<emphasis box="[983,1102,442,463]" italics="true" pageId="20" pageNumber="993">singularis</emphasis>
/
<emphasis box="[1127,1208,443,464]" italics="true" pageId="20" pageNumber="993">
<collectorName box="[1127,1208,443,464]" pageId="20" pageNumber="993">Ur. Rei</collectorName>
</emphasis>
] (
<collectionCode box="[1238,1315,442,463]" pageId="20" pageNumber="993">MCZC</collectionCode>
ex
<collectorName pageId="20" pageNumber="993">F.C. Bowditch</collectorName>
collection),
<specimenCount box="[1047,1148,473,495]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
and
<specimenCount box="[1202,1323,473,495]" pageId="20" pageNumber="993" type="female">one female</specimenCount>
(
<collectionCode box="[1335,1424,473,494]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="20" pageNumber="993" type="Museum">NHMW</collectionCode>
ex
<collectorName box="[865,995,503,525]" pageId="20" pageNumber="993">W. Zelenka</collectorName>
collection) and
<specimenCount box="[1186,1314,503,525]" pageId="20" pageNumber="993" type="female">one female</specimenCount>
(labelled with the
<emphasis box="[915,1108,534,556]" italics="true" pageId="20" pageNumber="993">nomen in litteris</emphasis>
<emphasis box="[1120,1322,534,556]" italics="true" pageId="20" pageNumber="993">
<collectorName box="[1120,1225,534,555]" pageId="20" pageNumber="993">Ascropus</collectorName>
cupreus
</emphasis>
<collectorName box="[1330,1417,534,556]" pageId="20" pageNumber="993">Olivier</collectorName>
) (
<collectionCode box="[816,920,565,586]" pageId="20" pageNumber="993">OUMNH</collectionCode>
ex
<collectorName box="[980,1038,565,587]" pageId="20" pageNumber="993">Hope</collectorName>
<collectorName box="[1052,1167,565,587]" pageId="20" pageNumber="993">Westwood</collectorName>
collection)
</materialsCitation>
.
</paragraph>
<paragraph blockId="20.[809,1425,627,1907]" lastBlockId="23.[827,1443,197,280]" lastPageId="23" lastPageNumber="996" pageId="20" pageNumber="993">
<emphasis box="[809,1114,627,649]" italics="true" pageId="20" pageNumber="993">Blue and green specimens:</emphasis>
<emphasis bold="true" box="[1122,1298,627,649]" pageId="20" pageNumber="993">
<collectingCountry box="[1122,1293,627,649]" name="Argentina" pageId="20" pageNumber="993">ARGENTINA</collectingCountry>
:
</emphasis>
no further data
<specimenCount box="[885,987,658,680]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[999,1075,658,679]" pageId="20" pageNumber="993">MCZC</collectionCode>
ex H.C. Fall collection),
<specimenCount pageId="20" pageNumber="993" type="female">three females</specimenCount>
(
<collectionCode box="[917,1001,689,711]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="20" pageNumber="993">MFNB</collectionCode>
),
<specimenCount box="[1022,1151,689,711]" pageId="20" pageNumber="993" type="female">one female</specimenCount>
(
<collectionCode box="[1167,1248,689,710]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="20" pageNumber="993">NMPC</collectionCode>
ex
<collectionCode box="[1315,1372,689,710]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="20" pageNumber="993">SMF</collectionCode>
and E. Witte collection),
<specimenCount box="[1044,1149,719,741]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[1164,1246,720,742]" pageId="20" pageNumber="993">PMOC</collectionCode>
),
<specimenCount box="[1266,1424,719,741]" pageId="20" pageNumber="993" type="female">three females</specimenCount>
(
<collectionCode box="[816,899,750,771]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="20" pageNumber="993" type="Museum">RMNH</collectionCode>
ex H. Gerth collection),
<specimenCount box="[1193,1327,750,772]" pageId="20" pageNumber="993" type="male">three males</specimenCount>
and
<specimenCount pageId="20" pageNumber="993" type="female">two females</specimenCount>
(
<collectionCode box="[913,977,781,802]" pageId="20" pageNumber="993">SDEI</collectionCode>
ex Kraatz collection),
<specimenCount box="[1257,1374,781,803]" pageId="20" pageNumber="993" type="male">two males</specimenCount>
and
<specimenCount box="[809,961,811,833]" pageId="20" pageNumber="993" type="female">two females</specimenCount>
(
<collectionCode box="[981,1063,812,834]" pageId="20" pageNumber="993">SMTD</collectionCode>
) and
<specimenCount box="[1141,1315,811,833]" pageId="20" pageNumber="993" type="female">three females</specimenCount>
(
<collectionCode box="[1336,1411,812,834]" pageId="20" pageNumber="993">SPPC</collectionCode>
); ambiguous data (
<collectingRegion box="[1030,1096,842,864]" country="Argentina" name="Salta" pageId="20" pageNumber="993">Salta</collectingRegion>
or Misiones?)
<specimenCount box="[1311,1423,842,864]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[816,905,873,895]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="20" pageNumber="993">MNHN</collectionCode>
); further data illegible
<specimenCount box="[1199,1305,873,895]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[1319,1403,873,894]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="20" pageNumber="993">NHMB</collectionCode>
ex Frey collection); Gran
<collectingRegion box="[1122,1197,903,925]" country="Argentina" name="Chaco" pageId="20" pageNumber="993">Chaco</collectingRegion>
, no further data
<specimenCount box="[809,949,934,956]" pageId="20" pageNumber="993" type="male">three males</specimenCount>
(
<collectionCode box="[965,1052,934,956]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="20" pageNumber="993" type="Museum">RBINS</collectionCode>
); Pampa Occidental, no further data
<specimenCount box="[893,1001,965,987]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[1017,1100,965,986]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="20" pageNumber="993" type="Museum">MACN</collectionCode>
ex Hermann Burmeister collection); Salado River, no further data
<specimenCount box="[1291,1424,995,1017]" pageId="20" pageNumber="993" type="male">three males</specimenCount>
and
<specimenCount box="[864,996,1026,1048]" pageId="20" pageNumber="993" type="female">one female</specimenCount>
(
<collectionCode box="[1014,1100,1026,1047]" pageId="20" pageNumber="993">BMNH</collectionCode>
ex Nevinson collection).
<emphasis box="[809,979,1057,1078]" italics="true" pageId="20" pageNumber="993">
<collectingRegion box="[809,973,1057,1078]" country="Argentina" name="Catamarca" pageId="20" pageNumber="993">CATAMARCA</collectingRegion>
:
</emphasis>
no further data
<specimenCount box="[1183,1317,1057,1079]" pageId="20" pageNumber="993" type="male">three males</specimenCount>
(
<collectionCode box="[1330,1414,1057,1079]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="20" pageNumber="993" type="Museum">MACN</collectionCode>
).
<emphasis box="[809,914,1088,1109]" italics="true" pageId="20" pageNumber="993">
<collectingRegion box="[809,908,1088,1109]" country="Argentina" name="Chaco" pageId="20" pageNumber="993">CHACO</collectingRegion>
:
</emphasis>
no further data
<specimenCount box="[1157,1270,1087,1109]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[1289,1372,1087,1109]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="20" pageNumber="993">NHRS</collectionCode>
); no specific locality,
<date box="[994,1085,1118,1139]" pageId="20" pageNumber="993" value="1908-11">XI.1908</date>
, Carlos Bruch leg.
<specimenCount box="[1322,1425,1118,1140]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[816,900,1149,1170]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="20" pageNumber="993" type="Museum">MACN</collectionCode>
ex Carlos Bruch collection); Chacabuco, Charata,
<date box="[924,1012,1179,1200]" pageId="20" pageNumber="993" value="1997-02">II.1997</date>
, no collector
<specimenCount box="[1197,1319,1179,1201]" pageId="20" pageNumber="993" type="male">two males</specimenCount>
and
<specimenCount pageId="20" pageNumber="993" type="female">two females</specimenCount>
(
<collectionCode box="[916,1000,1210,1232]" pageId="20" pageNumber="993">PMOC</collectionCode>
).
<emphasis box="[1019,1158,1209,1233]" italics="true" pageId="20" pageNumber="993">CÓRDOBA:</emphasis>
no further data
<specimenCount pageId="20" pageNumber="993" type="male">five males</specimenCount>
(
<collectionCode box="[900,991,1241,1263]" pageId="20" pageNumber="993">BMNH</collectionCode>
),
<specimenCount box="[1015,1127,1241,1263]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[1147,1234,1241,1262]" pageId="20" pageNumber="993">BMNH</collectionCode>
ex Nevinson collection),
<specimenCount box="[941,1047,1271,1293]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
and
<specimenCount box="[1106,1265,1271,1293]" pageId="20" pageNumber="993" type="female">three females</specimenCount>
(
<collectionCode box="[1280,1365,1271,1293]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="20" pageNumber="993" type="Museum">MACN</collectionCode>
),
<specimenCount pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[882,964,1302,1323]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="20" pageNumber="993" type="Museum">MACN</collectionCode>
ex Hermann Burmeister collection),
<specimenCount box="[809,921,1333,1355]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[939,1021,1333,1355]" pageId="20" pageNumber="993">MLPA</collectionCode>
),
<specimenCount box="[1045,1178,1333,1355]" pageId="20" pageNumber="993" type="male">four males</specimenCount>
(
<collectionCode box="[1196,1289,1333,1355]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="20" pageNumber="993">MNHN</collectionCode>
),
<specimenCount box="[1313,1424,1333,1355]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[816,900,1363,1385]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="20" pageNumber="993">NMPC</collectionCode>
),
<specimenCount box="[921,1032,1363,1385]" pageId="20" pageNumber="993" type="male">six males</specimenCount>
and
<specimenCount box="[1094,1228,1363,1385]" pageId="20" pageNumber="993" type="female">six females</specimenCount>
(
<collectionCode box="[1245,1315,1363,1385]" pageId="20" pageNumber="993">SPPC</collectionCode>
) and
<specimenCount pageId="20" pageNumber="993" type="male">one male</specimenCount>
and
<specimenCount box="[927,1056,1394,1416]" pageId="20" pageNumber="993" type="female">one female</specimenCount>
(
<collectionCode box="[1072,1152,1394,1416]" pageId="20" pageNumber="993">SMTD</collectionCode>
); no specific locality or date, Davis leg.
<specimenCount box="[1027,1136,1425,1447]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[1153,1225,1425,1447]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="20" pageNumber="993">CNCI</collectionCode>
),
<specimenCount box="[1248,1371,1425,1447]" pageId="20" pageNumber="993" type="male">five males</specimenCount>
and
<specimenCount box="[809,966,1455,1477]" pageId="20" pageNumber="993" type="female">three females</specimenCount>
(
<collectionCode box="[981,1060,1455,1477]" pageId="20" pageNumber="993">MCZC</collectionCode>
),
<specimenCount box="[1079,1183,1455,1477]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[1198,1277,1455,1476]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="20" pageNumber="993">NMPC</collectionCode>
ex Nickerl collection),
<specimenCount box="[941,1078,1486,1508]" pageId="20" pageNumber="993" type="male">three males</specimenCount>
and
<specimenCount box="[1138,1264,1486,1508]" pageId="20" pageNumber="993" type="female">one female</specimenCount>
(
<collectionCode box="[1279,1365,1486,1508]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="20" pageNumber="993" type="Museum">RBINS</collectionCode>
),
<specimenCount pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[882,968,1517,1539]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="20" pageNumber="993" type="Museum">RMNH</collectionCode>
) and
<specimenCount box="[1036,1164,1517,1539]" pageId="20" pageNumber="993" type="female">one female</specimenCount>
(
<collectionCode box="[1180,1237,1517,1538]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="20" pageNumber="993">SMF</collectionCode>
ex Pipitz and E. Witte collections); no specific locality or date, H. Richter leg.
<specimenCount box="[1025,1172,1578,1600]" pageId="20" pageNumber="993" type="female">two females</specimenCount>
(
<collectionCode box="[1190,1272,1578,1600]" pageId="20" pageNumber="993">MLPA</collectionCode>
); no specific locality or date, Weyenb. leg.
<specimenCount box="[1157,1280,1609,1631]" pageId="20" pageNumber="993" type="female">one female</specimenCount>
(
<collectionCode box="[1294,1380,1609,1631]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="20" pageNumber="993" type="Museum">RMNH</collectionCode>
); no specific locality, 1900s, C. Bruch leg.
<specimenCount box="[1292,1424,1639,1661]" pageId="20" pageNumber="993" type="female">one female</specimenCount>
(
<collectionCode box="[816,898,1670,1691]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="20" pageNumber="993" type="Museum">RBINS</collectionCode>
ex J. J. Gillet collection); no specific locality,
<date box="[809,942,1701,1722]" pageId="20" pageNumber="993" value="1904-11-07">07.XI.1904</date>
, C. Bruch leg.
<specimenCount box="[1148,1281,1701,1723]" pageId="20" pageNumber="993" type="female">one female</specimenCount>
(
<collectionCode box="[1298,1364,1701,1722]" pageId="20" pageNumber="993">SDEI</collectionCode>
ex Kraatz collection); no specific locality,
<date box="[1286,1419,1731,1752]" pageId="20" pageNumber="993" value="1904-11-10">10.XI.1904</date>
, C. Bruch leg.
<specimenCount box="[979,1082,1762,1784]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[1095,1159,1762,1783]" pageId="20" pageNumber="993">SDEI</collectionCode>
ex Kraatz collection); Calamuchita, El Sauce,
<date box="[1109,1215,1793,1814]" pageId="20" pageNumber="993" value="1940-12">XII.1940</date>
, Manuel J. Viana leg.
<specimenCount box="[871,972,1823,1845]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
and
<specimenCount box="[1025,1159,1823,1845]" pageId="20" pageNumber="993" type="female">two females</specimenCount>
(
<collectionCode box="[1171,1255,1823,1845]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="20" pageNumber="993" type="Museum">MACN</collectionCode>
); Calamuchita, Villa Yacanto de Calamuchita, no date, Breyer leg.
<specimenCount box="[809,914,1885,1907]" pageId="20" pageNumber="993" type="male">one male</specimenCount>
(
<collectionCode box="[930,1015,1885,1907]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="20" pageNumber="993" type="Museum">MACN</collectionCode>
); Ischilín, Copacabana,
<date box="[1294,1424,1885,1906]" pageId="20" pageNumber="993" value="1980-03-07">07.III.1980</date>
(cow dung), Woodruff leg.
<specimenCount box="[520,670,197,219]" pageId="21" pageNumber="994" type="female">two females</specimenCount>
(
<collectionCode box="[689,765,197,219]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="21" pageNumber="994" type="Museum">FSCA</collectionCode>
); Ischilín, near Copacabana,
<date box="[497,629,228,249]" pageId="21" pageNumber="994" value="1980-03-08">08.III.1980</date>
(pitfall with pig dung), Woodruff, Cordor and Fincher leg.
<specimenCount pageId="21" pageNumber="994" type="male">three males</specimenCount>
and
<specimenCount box="[303,445,289,311]" pageId="21" pageNumber="994" type="female">six females</specimenCount>
(
<collectionCode box="[465,542,289,311]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="21" pageNumber="994" type="Museum">FSCA</collectionCode>
); Ischilín, Quilino,
<date box="[163,269,320,342]" pageId="21" pageNumber="994" value="1965-12">XII.1965</date>
, Cichero leg.
<specimenCount box="[454,585,320,342]" pageId="21" pageNumber="994" type="female">one female</specimenCount>
(
<collectionCode box="[601,687,320,342]" pageId="21" pageNumber="994">CMNC</collectionCode>
); Jaime Pérez, Jaime Peter (Rt. 16, 6Km W Jaime Peter),
<date box="[163,300,381,402]" pageId="21" pageNumber="994" value="1978-12-09">09.XII.1978</date>
, Woodruff, Runnacles and Cordo leg.
<specimenCount pageId="21" pageNumber="994" type="male">22 males</specimenCount>
and
<specimenCount box="[297,427,412,434]" pageId="21" pageNumber="994" type="female">17 females</specimenCount>
(
<collectionCode box="[445,520,412,434]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="21" pageNumber="994" type="Museum">FSCA</collectionCode>
); Punilla, Capilla del Monte, no date, Hasseus (or Hosseus?) leg.
<specimenCount box="[676,779,442,464]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
and
<specimenCount box="[212,336,473,495]" pageId="21" pageNumber="994" type="female">one female</specimenCount>
(
<collectionCode box="[350,408,473,495]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="21" pageNumber="994" type="Museum">ZSM</collectionCode>
); Punilla, Valle Hermoso,
<date box="[701,774,473,494]" pageId="21" pageNumber="994" value="1942-01">I.1942</date>
, Viana leg.
<specimenCount box="[297,397,504,526]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[409,481,504,525]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="21" pageNumber="994" type="Museum">MZSP</collectionCode>
ex Ferraciolli collection); Río Cuarto, no further data
<specimenCount box="[515,621,534,556]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[636,721,535,557]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="21" pageNumber="994" type="Museum">MACN</collectionCode>
); Río Seco, no date, illegible collector name
<specimenCount box="[649,778,565,587]" pageId="21" pageNumber="994" type="female">one female</specimenCount>
(
<collectionCode box="[170,255,596,618]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="21" pageNumber="994" type="Museum">MACN</collectionCode>
); Royo Cortado,
<date box="[458,598,596,618]" pageId="21" pageNumber="994" value="1939-12-15">15.XII.1939</date>
, C.M.A.M. leg.
<specimenCount box="[183,286,626,648]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[299,376,627,649]" pageId="21" pageNumber="994">MLPA</collectionCode>
); San Alberto, Las Rabonas,
<date box="[701,774,626,648]" pageId="21" pageNumber="994" value="1964-01">I.1964</date>
, no collector
<specimenCount box="[339,450,657,679]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[468,550,657,679]" pageId="21" pageNumber="994">MLPA</collectionCode>
); San Alberto, Las Rabonas, Sierras Grandes,
<date box="[481,555,688,710]" pageId="21" pageNumber="994" value="1967-01">I.1967</date>
, no collector
<specimenCount pageId="21" pageNumber="994" type="male">four males</specimenCount>
and
<specimenCount box="[295,459,718,740]" pageId="21" pageNumber="994" type="female">three females</specimenCount>
(
<collectionCode box="[475,555,719,741]" pageId="21" pageNumber="994">MLPA</collectionCode>
); Sobremonte, San Francisco del Chañar, Rumi-Puca,
<date box="[570,679,749,771]" pageId="21" pageNumber="994" value="1966-01-15">15.I.1966</date>
, Cichero leg.
<specimenCount box="[241,368,780,802]" pageId="21" pageNumber="994" type="male">two males</specimenCount>
(
<collectionCode box="[387,476,780,802]" pageId="21" pageNumber="994">CMNC</collectionCode>
); Totoral,
<collectingRegion box="[605,735,780,801]" country="Argentina" name="La Pampa" pageId="21" pageNumber="994">La Pampa</collectingRegion>
, no further data
<specimenCount box="[340,468,810,832]" pageId="21" pageNumber="994" type="female">one female</specimenCount>
(
<collectionCode box="[484,575,811,833]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="21" pageNumber="994">MNHN</collectionCode>
).
<emphasis box="[594,778,811,832]" italics="true" pageId="21" pageNumber="994">
<collectingRegion box="[594,772,811,832]" country="Argentina" name="Corrientes" pageId="21" pageNumber="994">CORRIENTES</collectingRegion>
:
</emphasis>
General Paz, Itá Ibaté,
<date box="[453,550,841,862]" pageId="21" pageNumber="994" value="1979-09">IX.1979</date>
, no collector
<specimenCount pageId="21" pageNumber="994" type="female">one female</specimenCount>
(
<collectionCode box="[254,325,872,894]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="21" pageNumber="994" type="Museum">FSCA</collectionCode>
); Ituzaingó,
<date box="[465,567,872,893]" pageId="21" pageNumber="994" value="1987-12">XII.1987</date>
, S. Bolle leg.
<specimenCount pageId="21" pageNumber="994" type="male">two males</specimenCount>
(both dissected) and
<specimenCount box="[491,620,902,924]" pageId="21" pageNumber="994" type="female">one female</specimenCount>
(
<collectionCode box="[636,721,902,923]" pageId="21" pageNumber="994">BMNH</collectionCode>
ex J. Zídek collection).
<emphasis box="[448,643,932,956]" italics="true" pageId="21" pageNumber="994">
<collectingRegion box="[448,634,932,956]" country="Argentina" name="Entre Rios" pageId="21" pageNumber="994">ENTRE RÍOS</collectingRegion>
:
</emphasis>
Uruguay, Pronunciamiento,
<date box="[391,500,964,985]" pageId="21" pageNumber="994" value="1988-12">XII.1988</date>
, S. Bolle
<specimenCount box="[645,778,964,986]" pageId="21" pageNumber="994" type="female">one female</specimenCount>
(
<collectionCode box="[170,254,994,1015]" pageId="21" pageNumber="994">BMNH</collectionCode>
ex J. Zídek collection).
<emphasis box="[554,649,995,1016]" italics="true" pageId="21" pageNumber="994">
<collectingRegion box="[554,642,995,1016]" country="Argentina" name="Jujuy" pageId="21" pageNumber="994">JUJUY</collectingRegion>
:
</emphasis>
no further data
<specimenCount box="[243,380,1025,1047]" pageId="21" pageNumber="994" type="male">three males</specimenCount>
and
<specimenCount box="[438,576,1025,1047]" pageId="21" pageNumber="994" type="female">two females</specimenCount>
(
<collectionCode box="[591,673,1025,1047]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="21" pageNumber="994" type="Museum">MACN</collectionCode>
) and
<specimenCount pageId="21" pageNumber="994" type="female">one female</specimenCount>
(
<collectionCode box="[262,344,1056,1078]" pageId="21" pageNumber="994">MLPA</collectionCode>
).
<emphasis box="[366,511,1056,1077]" italics="true" pageId="21" pageNumber="994">
<collectingRegion box="[366,505,1056,1077]" country="Argentina" name="La Pampa" pageId="21" pageNumber="994">LA PAMPA</collectingRegion>
:
</emphasis>
Lihuel Calel, Parque Nacional Lihué Calel,
<date box="[432,555,1086,1108]" pageId="21" pageNumber="994" value="1986-02-12">12.II.1986</date>
, no collector
<specimenCount pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[233,313,1117,1139]" pageId="21" pageNumber="994">CEMT</collectionCode>
).
<emphasis box="[330,456,1117,1138]" italics="true" pageId="21" pageNumber="994">
<collectingRegion box="[330,450,1117,1138]" country="Argentina" name="La Rioja" pageId="21" pageNumber="994">LA RIOJA</collectingRegion>
:
</emphasis>
no further data
<specimenCount box="[664,778,1117,1139]" pageId="21" pageNumber="994" type="male">two males</specimenCount>
(
<collectionCode box="[170,254,1148,1170]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="21" pageNumber="994" type="Museum">MACN</collectionCode>
); no specific locality,
<date box="[494,568,1148,1169]" pageId="21" pageNumber="994" value="1894-01">I.1894</date>
, Carlos Bruch leg.
<specimenCount box="[185,291,1178,1200]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[306,388,1178,1199]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="21" pageNumber="994" type="Museum">MACN</collectionCode>
ex Carlos Bruch collection); no specific locality,
<date box="[364,442,1209,1230]" pageId="21" pageNumber="994" value="1904-01">I.1904</date>
, C. Bruch leg.
<specimenCount box="[647,778,1209,1231]" pageId="21" pageNumber="994" type="female">one female</specimenCount>
(
<collectionCode box="[170,247,1240,1262]" pageId="21" pageNumber="994">MLPA</collectionCode>
),
<specimenCount box="[265,388,1240,1262]" pageId="21" pageNumber="994" type="female">one female</specimenCount>
(
<collectionCode box="[402,484,1240,1261]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="21" pageNumber="994" type="Museum">RBINS</collectionCode>
ex J. J. Gillet collection) and
<specimenCount box="[214,320,1270,1292]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[335,399,1270,1291]" pageId="21" pageNumber="994">SDEI</collectionCode>
ex Kraatz collection); General San Martín, Ulapes,
<date box="[407,545,1301,1322]" pageId="21" pageNumber="994" value="2007-12-29">29.XII.2007</date>
, O. Boilly leg.
<specimenCount pageId="21" pageNumber="994" type="female">two females</specimenCount>
(
<collectionCode box="[271,349,1332,1354]" pageId="21" pageNumber="994">OBCL</collectionCode>
); Rosario Vera Peñaloza, Chepes (
<quantity metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="21" pageNumber="994" unit="km" value="6.0">6 km</quantity>
E Chepes),
<date box="[356,498,1362,1383]" pageId="21" pageNumber="994" value="1978-12-10">10.XII.1978</date>
[goat dung], Woodruff, Runnacles and Cordo leg.
<specimenCount box="[545,664,1393,1415]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[686,765,1393,1415]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="21" pageNumber="994" type="Museum">FSCA</collectionCode>
).
<emphasis box="[163,305,1424,1445]" italics="true" pageId="21" pageNumber="994">
<collectingRegion box="[163,299,1424,1445]" country="Argentina" name="Mendoza" pageId="21" pageNumber="994">MENDOZA</collectingRegion>
:
</emphasis>
no further data
<specimenCount box="[521,626,1424,1446]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[640,724,1424,1445]" pageId="21" pageNumber="994">BMNH</collectionCode>
ex Nevinson collection),
<specimenCount box="[439,559,1454,1476]" pageId="21" pageNumber="994" type="male">six males</specimenCount>
and
<specimenCount box="[627,777,1454,1476]" pageId="21" pageNumber="994" type="female">two females</specimenCount>
(
<collectionCode box="[171,257,1485,1507]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="21" pageNumber="994">MFNB</collectionCode>
),
<specimenCount box="[281,393,1485,1507]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[412,505,1485,1507]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="21" pageNumber="994">MNHN</collectionCode>
),
<specimenCount box="[528,655,1485,1507]" pageId="21" pageNumber="994" type="male">two males</specimenCount>
(
<collectionCode box="[673,765,1485,1507]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="21" pageNumber="994">MNHN</collectionCode>
),
<specimenCount box="[163,296,1516,1538]" pageId="21" pageNumber="994" type="male">three males</specimenCount>
and
<specimenCount box="[352,475,1516,1538]" pageId="21" pageNumber="994" type="female">one female</specimenCount>
(
<collectionCode box="[488,562,1516,1538]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="21" pageNumber="994" type="Museum">MZSP</collectionCode>
),
<specimenCount box="[580,682,1516,1538]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[696,779,1516,1537]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="21" pageNumber="994">NHMB</collectionCode>
ex H. Rolle and Frey collections),
<specimenCount box="[550,684,1546,1568]" pageId="21" pageNumber="994" type="female">two females</specimenCount>
(
<collectionCode box="[696,778,1546,1567]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="21" pageNumber="994" type="Museum">RBINS</collectionCode>
ex H. Rolle and Le Moult collections),
<specimenCount box="[668,778,1577,1599]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[170,258,1608,1630]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="21" pageNumber="994" type="Museum">RMNH</collectionCode>
),
<specimenCount box="[278,421,1608,1630]" pageId="21" pageNumber="994" type="male">seven males</specimenCount>
and
<specimenCount box="[482,622,1608,1630]" pageId="21" pageNumber="994" type="female">two females</specimenCount>
(
<collectionCode box="[639,710,1608,1630]" pageId="21" pageNumber="994">SPPC</collectionCode>
),
<specimenCount pageId="21" pageNumber="994" type="male">four males</specimenCount>
and
<specimenCount box="[289,427,1638,1660]" pageId="21" pageNumber="994" type="female">two females</specimenCount>
(
<collectionCode box="[442,519,1638,1660]" pageId="21" pageNumber="994">SMTD</collectionCode>
) and
<specimenCount box="[584,688,1638,1660]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[703,779,1638,1659]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="21" pageNumber="994" type="Museum">ZFMK</collectionCode>
ex H. Rolle and Fuhlrott collections); no specific locality or date, illegible collector name
<specimenCount box="[671,778,1700,1722]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[171,258,1730,1752]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="21" pageNumber="994">MFNB</collectionCode>
).
<emphasis box="[280,429,1731,1752]" italics="true" pageId="21" pageNumber="994">MISIONES:</emphasis>
no further data
<specimenCount box="[667,778,1730,1752]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[170,225,1761,1782]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="21" pageNumber="994" type="Museum">ZSM</collectionCode>
ex Juan Foerster collection); Capital, Posadas,
<date box="[163,264,1792,1813]" pageId="21" pageNumber="994" value="1997-03">III.1997</date>
, Lucio Coronel leg.
<specimenCount box="[538,672,1791,1813]" pageId="21" pageNumber="994" type="female">one female</specimenCount>
(
<collectionCode box="[691,766,1792,1814]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="21" pageNumber="994" type="Museum">FSCA</collectionCode>
); Eldorado, 9 de Julio, no further data
<specimenCount box="[666,777,1822,1844]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[170,254,1853,1875]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="21" pageNumber="994" type="Museum">MACN</collectionCode>
).
<emphasis box="[271,360,1853,1874]" italics="true" pageId="21" pageNumber="994">
<collectingRegion box="[271,354,1853,1874]" country="Argentina" name="Salta" pageId="21" pageNumber="994">SALTA</collectingRegion>
:
</emphasis>
no further data
<specimenCount box="[567,681,1853,1875]" pageId="21" pageNumber="994" type="male">two males</specimenCount>
and
<specimenCount pageId="21" pageNumber="994" type="female">two females</specimenCount>
(
<collectionCode box="[268,347,1884,1906]" pageId="21" pageNumber="994">MLPA</collectionCode>
),
<specimenCount box="[366,485,1883,1905]" pageId="21" pageNumber="994" type="male">two males</specimenCount>
and
<specimenCount box="[545,672,1883,1905]" pageId="21" pageNumber="994" type="female">one female</specimenCount>
(
<collectionCode box="[688,766,1884,1906]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="21" pageNumber="994">NHRS</collectionCode>
),
<specimenCount box="[827,956,197,219]" pageId="21" pageNumber="994" type="male">nine males</specimenCount>
and
<specimenCount box="[1016,1176,197,219]" pageId="21" pageNumber="994" type="female">three females</specimenCount>
(
<collectionCode box="[1193,1264,197,219]" pageId="21" pageNumber="994">SPPC</collectionCode>
), and
<specimenCount box="[1337,1443,197,219]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[834,913,228,250]" pageId="21" pageNumber="994">SMTD</collectionCode>
); no specific locality or date, Brauer leg.
<specimenCount pageId="21" pageNumber="994" type="male">two males</specimenCount>
and
<specimenCount box="[954,1080,258,280]" pageId="21" pageNumber="994" type="female">one female</specimenCount>
(
<collectionCode box="[1095,1177,258,280]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="21" pageNumber="994">MFNB</collectionCode>
); no specific locality or date, Hans Mayer leg.
<specimenCount box="[1094,1216,289,311]" pageId="21" pageNumber="994" type="female">one female</specimenCount>
(
<collectionCode box="[1228,1310,289,311]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="21" pageNumber="994">NMPC</collectionCode>
); no specific locality, 1905 (?), Steinbach leg.
<specimenCount box="[1223,1342,319,341]" pageId="21" pageNumber="994" type="male">two males</specimenCount>
and
<specimenCount pageId="21" pageNumber="994" type="female">two females</specimenCount>
(
<collectionCode box="[944,1032,350,372]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="21" pageNumber="994">MFNB</collectionCode>
); no specific locality,
<date box="[1315,1397,350,371]" pageId="21" pageNumber="994" value="1949-01">I.1949</date>
, no collector
<specimenCount box="[969,1082,381,403]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[1101,1178,381,403]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="21" pageNumber="994" type="Museum">FSCA</collectionCode>
); no specific locality,
<date box="[827,948,412,433]" pageId="21" pageNumber="994" value="1992-02-07">07.II.1992</date>
, no collector
<specimenCount box="[1126,1232,411,433]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[1248,1329,412,434]" pageId="21" pageNumber="994">CEMT</collectionCode>
); Capital,
<collectingRegion box="[827,896,442,464]" country="Argentina" name="Salta" pageId="21" pageNumber="994">Salta</collectingRegion>
, Cerro San Bernardo,
<quantity box="[1192,1289,442,464]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="21" pageNumber="994" unit="m" value="1450.0">1450 m</quantity>
,
<date box="[1305,1436,442,464]" pageId="21" pageNumber="994" value="1982-02-15">15.II.1982</date>
, H. Howden and A. Howden leg.
<specimenCount box="[1224,1330,473,495]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[1346,1431,473,495]" pageId="21" pageNumber="994">CMNC</collectionCode>
); Cerrillos, La Merced, no date, Raimoser leg.
<specimenCount pageId="21" pageNumber="994" type="male">three males</specimenCount>
and
<specimenCount box="[953,1097,534,556]" pageId="21" pageNumber="994" type="female">four females</specimenCount>
(
<collectionCode box="[1111,1204,534,556]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="21" pageNumber="994" type="Museum">NHMW</collectionCode>
); Chicoana, Ruta 68,
<date box="[827,915,565,586]" pageId="21" pageNumber="994" value="1984-02">II.1984</date>
, no collector
<specimenCount box="[1101,1210,565,587]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[1227,1313,565,587]" pageId="21" pageNumber="994">CMNC</collectionCode>
); El Vino,
<date box="[827,935,596,617]" pageId="21" pageNumber="994" value="2008-01-24">24.I.2008</date>
, O. Boilly leg.
<specimenCount box="[1119,1243,595,617]" pageId="21" pageNumber="994" type="female">one female</specimenCount>
(
<collectionCode box="[1257,1332,596,618]" pageId="21" pageNumber="994">OBCL</collectionCode>
); General José de San Martín, Embarcación,
<date box="[1214,1305,626,648]" pageId="21" pageNumber="994" value="1953-11">XI.1953</date>
, no collector
<specimenCount box="[851,962,657,679]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
(dissected) and
<specimenCount box="[1168,1301,657,679]" pageId="21" pageNumber="994" type="female">one female</specimenCount>
(dissected) (
<collectionCode box="[834,906,688,709]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="21" pageNumber="994" type="University or college">DZUP</collectionCode>
ex M. Alvarenga collection); General José de San Martín, Salvador Mazza (Pocitos),
<date box="[1356,1436,718,740]" pageId="21" pageNumber="994" value="1962-01">I.1962</date>
, A. Martínez leg.
<specimenCount box="[1032,1152,749,771]" pageId="21" pageNumber="994" type="male">four males</specimenCount>
and
<specimenCount box="[1208,1343,749,771]" pageId="21" pageNumber="994" type="female">two females</specimenCount>
(
<collectionCode box="[1357,1431,749,771]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="21" pageNumber="994" type="Museum">MZSP</collectionCode>
); La Viña,
<date box="[935,1019,780,801]" pageId="21" pageNumber="994" value="1984-10">X.1984</date>
, M. Viana leg.
<specimenCount box="[1218,1339,779,801]" pageId="21" pageNumber="994" type="male">five males</specimenCount>
and
<specimenCount pageId="21" pageNumber="994" type="female">five females</specimenCount>
(
<collectionCode box="[934,1007,810,832]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="21" pageNumber="994" type="Museum">FSCA</collectionCode>
); La Viña,
<date box="[1137,1270,810,831]" pageId="21" pageNumber="994" value="1991-03-01">01.III.1991</date>
, no collector
<specimenCount box="[827,943,841,863]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
and
<specimenCount box="[1015,1154,841,863]" pageId="21" pageNumber="994" type="female">one female</specimenCount>
(
<collectionCode box="[1175,1262,841,863]" pageId="21" pageNumber="994">CEMT</collectionCode>
); La Viña, El Carmen,
<date box="[944,1044,872,893]" pageId="21" pageNumber="994" value="1943-03">III.1943</date>
, no collector
<specimenCount box="[1229,1338,871,893]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
and
<specimenCount pageId="21" pageNumber="994" type="female">one female</specimenCount>
(
<collectionCode box="[920,1005,902,924]" pageId="21" pageNumber="994">CMNC</collectionCode>
); La Viña, Ruta 68, km 89,
<date box="[1327,1402,902,923]" pageId="21" pageNumber="994" value="1987-01">I.1987</date>
, no collector
<specimenCount box="[959,1068,933,955]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[1085,1171,933,955]" pageId="21" pageNumber="994">CMNC</collectionCode>
); Picharal, no further data
<specimenCount box="[905,1008,963,985]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[1022,1104,963,985]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="21" pageNumber="994">NMPC</collectionCode>
).
<emphasis box="[1121,1261,964,985]" italics="true" pageId="21" pageNumber="994">
<collectingRegion box="[1121,1255,964,985]" country="Argentina" name="San Juan" pageId="21" pageNumber="994">SAN JUAN</collectingRegion>
:
</emphasis>
no further data
<specimenCount box="[849,975,994,1016]" pageId="21" pageNumber="994" type="female">one female</specimenCount>
(
<collectionCode box="[991,1076,994,1016]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="21" pageNumber="994" type="Museum">MACN</collectionCode>
); Iglesia, Angualasto,
<date box="[1334,1438,994,1016]" pageId="21" pageNumber="994" value="1960-12">XII.1960</date>
, no collector
<specimenCount box="[1005,1139,1025,1047]" pageId="21" pageNumber="994" type="female">one female</specimenCount>
(
<collectionCode box="[1158,1241,1025,1047]" pageId="21" pageNumber="994">MLPA</collectionCode>
).
<emphasis box="[1262,1404,1025,1046]" italics="true" pageId="21" pageNumber="994">
<collectingRegion box="[1262,1397,1025,1046]" country="Argentina" name="San Luis" pageId="21" pageNumber="994">SAN LUIS</collectingRegion>
:
</emphasis>
no further data
<specimenCount box="[999,1116,1055,1077]" pageId="21" pageNumber="994" type="male">two males</specimenCount>
and
<specimenCount box="[1175,1333,1055,1077]" pageId="21" pageNumber="994" type="female">three females</specimenCount>
(
<collectionCode box="[1347,1431,1055,1077]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="21" pageNumber="994" type="Museum">MACN</collectionCode>
); no specific locality or date, Carlos Bruch leg.
<specimenCount pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[897,978,1117,1138]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="21" pageNumber="994" type="Museum">MACN</collectionCode>
ex Carlos Bruch collection); no specific locality or date, H. Richter leg.
<specimenCount box="[1203,1340,1147,1169]" pageId="21" pageNumber="994" type="female">two females</specimenCount>
(
<collectionCode box="[1354,1431,1147,1169]" pageId="21" pageNumber="994">MLPA</collectionCode>
); no specific locality, 1913, H. Gerth leg.
<specimenCount box="[1279,1393,1178,1200]" pageId="21" pageNumber="994" type="male">two males</specimenCount>
and
<specimenCount box="[827,968,1209,1231]" pageId="21" pageNumber="994" type="female">one female</specimenCount>
(
<collectionCode box="[990,1055,1209,1231]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="21" pageNumber="994">SMF</collectionCode>
); Belgrano, Fundo El Molle,
<geoCoordinate box="[827,978,1239,1260]" degrees="33" direction="south" minutes="02" orientation="latitude" pageId="21" pageNumber="994" precision="1" seconds="22.8" value="-33.039665">33°0222.8”S</geoCoordinate>
,
<geoCoordinate box="[992,1150,1239,1261]" degrees="66" direction="west" minutes="30" orientation="longitude" pageId="21" pageNumber="994" precision="1" seconds="47.6" value="-66.51322">66°3047.6”W</geoCoordinate>
,
<quantity box="[1163,1239,1239,1261]" metricMagnitude="2" metricUnit="m" metricValue="6.22" pageId="21" pageNumber="994" unit="m" value="622.0">622 m</quantity>
,
<date box="[1253,1374,1239,1260]" pageId="21" pageNumber="994" value="2017-02-07">07.II.2017</date>
(trap with chicken meat), G. Arriagada leg.
<specimenCount box="[1320,1442,1270,1292]" pageId="21" pageNumber="994" type="male">two males</specimenCount>
(both dissected) and
<specimenCount box="[1061,1196,1301,1323]" pageId="21" pageNumber="994" type="female">two females</specimenCount>
(
<collectionCode box="[1210,1290,1301,1323]" pageId="21" pageNumber="994">CEMT</collectionCode>
).
<emphasis box="[1307,1443,1301,1322]" italics="true" pageId="21" pageNumber="994">
<collectingRegion box="[1307,1436,1301,1322]" country="Argentina" name="Santa Fe" pageId="21" pageNumber="994">SANTA FE</collectingRegion>
:
</emphasis>
Estancia La Noria, San Javier River,
<date box="[1293,1436,1331,1353]" pageId="21" pageNumber="994" value="1911-12-15">15.XII.1911</date>
, G. E. Bryant leg.
<specimenCount box="[1035,1157,1362,1384]" pageId="21" pageNumber="994" type="female">one female</specimenCount>
(
<collectionCode box="[1169,1252,1362,1383]" pageId="21" pageNumber="994">BMNH</collectionCode>
ex G.E. Bryant collection); Estancia La Noria, San Javier River,
<date box="[827,963,1423,1444]" pageId="21" pageNumber="994" value="1911-12-18">18.XII.1911</date>
, G.E. Bryant leg.
<specimenCount box="[1188,1324,1423,1445]" pageId="21" pageNumber="994" type="male">three males</specimenCount>
(
<collectionCode box="[1338,1421,1423,1444]" pageId="21" pageNumber="994">BMNH</collectionCode>
ex G. E. Bryant collection); Estancia La Noria, San Javier River,
<date box="[978,1113,1485,1506]" pageId="21" pageNumber="994" value="1911-12-22">22.XII.1911</date>
, G. E. Bryant leg.
<specimenCount box="[1340,1443,1485,1507]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[834,920,1515,1536]" pageId="21" pageNumber="994">BMNH</collectionCode>
ex G.E. Bryant collection); Estancia La Noria, San Javier River,
<date box="[1110,1218,1546,1567]" pageId="21" pageNumber="994" value="1912-01-10">10.I.1912</date>
, G. E. Bryant leg.
<specimenCount box="[827,962,1577,1599]" pageId="21" pageNumber="994" type="female">one female</specimenCount>
(
<collectionCode box="[980,1067,1577,1598]" pageId="21" pageNumber="994">BMNH</collectionCode>
ex G. E. Bryant collection); Estancia La Noria, San Javier River,
<date box="[1256,1374,1607,1628]" pageId="21" pageNumber="994" value="1911-02-07">07.II.1911</date>
, G. E. Bryant leg.
<specimenCount box="[999,1110,1638,1660]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[1128,1214,1638,1659]" pageId="21" pageNumber="994">BMNH</collectionCode>
ex G. E. Bryant collection); General Obligado, Villa Ana,
<date box="[1298,1379,1669,1690]" pageId="21" pageNumber="994" value="1924-10">X.1924</date>
, K. J. Hayward leg.
<specimenCount box="[1012,1118,1699,1721]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[1134,1222,1699,1721]" pageId="21" pageNumber="994">BMNH</collectionCode>
).
<emphasis italics="true" pageId="21" pageNumber="994">
<collectingRegion country="Argentina" name="Santiago del Estero" pageId="21" pageNumber="994">SANTIAGO DEL ESTERO</collectingRegion>
:
</emphasis>
no further data no further data
<specimenCount box="[1323,1442,1730,1752]" pageId="21" pageNumber="994" type="male">four males</specimenCount>
and
<specimenCount box="[880,1022,1761,1783]" pageId="21" pageNumber="994" type="female">two females</specimenCount>
(
<collectionCode box="[1039,1125,1761,1783]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="21" pageNumber="994" type="Museum">MACN</collectionCode>
),
<specimenCount box="[1145,1253,1761,1783]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
and
<specimenCount box="[1314,1442,1761,1783]" pageId="21" pageNumber="994" type="female">11 females</specimenCount>
(
<collectionCode box="[835,922,1791,1813]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="21" pageNumber="994">MFNB</collectionCode>
),
<specimenCount box="[945,1058,1791,1813]" pageId="21" pageNumber="994" type="male">one male</specimenCount>
(
<collectionCode box="[1077,1160,1791,1813]" pageId="21" pageNumber="994">MLPA</collectionCode>
),
<specimenCount box="[1183,1310,1791,1813]" pageId="21" pageNumber="994" type="male">two males</specimenCount>
and
<specimenCount pageId="21" pageNumber="994" type="female">three females</specimenCount>
(
<collectionCode box="[932,1021,1822,1844]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="21" pageNumber="994">MNHN</collectionCode>
),
<specimenCount box="[1040,1158,1822,1844]" pageId="21" pageNumber="994" type="male">two males</specimenCount>
and
<specimenCount box="[1217,1343,1822,1844]" pageId="21" pageNumber="994" type="female">one female</specimenCount>
(
<collectionCode box="[1358,1434,1822,1844]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="21" pageNumber="994">NHRS</collectionCode>
) and
<specimenCount box="[880,1012,1852,1874]" pageId="21" pageNumber="994" type="male">nine males</specimenCount>
and
<specimenCount box="[1075,1205,1852,1874]" pageId="21" pageNumber="994" type="female">12 females</specimenCount>
(
<collectionCode box="[1222,1294,1853,1875]" pageId="21" pageNumber="994">SPPC</collectionCode>
); no specific locality or date, Wagner leg.
<specimenCount box="[1183,1293,1883,1905]" pageId="21" pageNumber="994" type="male">six males</specimenCount>
(
<collectionCode box="[1307,1384,1883,1905]" pageId="21" pageNumber="994">MLPA</collectionCode>
) and
<specimenCount box="[145,278,197,219]" pageId="22" pageNumber="995" type="female">one female</specimenCount>
(
<collectionCode box="[297,359,197,219]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="22" pageNumber="995">SMF</collectionCode>
); uncertain specific locality (La Palisa bei Bracho),
<collectingRegion box="[379,451,228,250]" country="Argentina" name="Chaco" pageId="22" pageNumber="995">Chaco</collectingRegion>
de Santiago, banks of the Salado River, no date, Wagner leg.
<specimenCount box="[561,664,258,280]" pageId="22" pageNumber="995" type="male">one male</specimenCount>
and
<specimenCount pageId="22" pageNumber="995" type="female">one female</specimenCount>
(
<collectionCode box="[242,338,289,311]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="22" pageNumber="995" type="Museum">NHMW</collectionCode>
); Alberdi, Campo Gallo, no further data
<specimenCount box="[235,348,320,342]" pageId="22" pageNumber="995" type="male">one male</specimenCount>
(
<collectionCode box="[367,460,320,342]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="22" pageNumber="995">MNHN</collectionCode>
); Alberdi, Campo Gallo,
<date box="[145,243,351,372]" pageId="22" pageNumber="995" value="1943-03">III.1943</date>
, Prosen leg.
<specimenCount box="[419,526,350,372]" pageId="22" pageNumber="995" type="male">one male</specimenCount>
(
<collectionCode box="[542,602,351,373]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="22" pageNumber="995" type="Museum">ZSM</collectionCode>
); Avellaneda, Colonia Dora,
<date box="[313,398,381,403]" pageId="22" pageNumber="995" value="1951-02">II.1951</date>
, Prosen leg.
<specimenCount box="[569,675,381,403]" pageId="22" pageNumber="995" type="male">one male</specimenCount>
(
<collectionCode box="[690,749,381,403]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="22" pageNumber="995" type="Museum">ZSM</collectionCode>
); Avellaneda, Icaño,
<date box="[370,475,412,433]" pageId="22" pageNumber="995" value="1991-12">XII.1991</date>
, no collector
<specimenCount box="[654,760,412,434]" pageId="22" pageNumber="995" type="male">one male</specimenCount>
(dissected) (
<collectionCode box="[294,376,443,464]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="22" pageNumber="995">NMPC</collectionCode>
ex.
<collectionCode box="[454,472,443,464]" name="Tavera, Department of Geology and Geophysics" pageId="22" pageNumber="995">T</collectionCode>
. Taylor and S. Pokorný collections); Avallaneda, Icaño,
<date box="[510,613,473,494]" pageId="22" pageNumber="995" value="1991-12">XII.1991</date>
, S. Bolle leg.
<specimenCount box="[166,281,504,526]" pageId="22" pageNumber="995" type="male">two males</specimenCount>
(both dissected) and
<specimenCount box="[526,663,504,526]" pageId="22" pageNumber="995" type="female">two females</specimenCount>
(
<collectionCode box="[678,761,504,525]" pageId="22" pageNumber="995">BMNH</collectionCode>
ex J. Zídek collection); Avellaneda, Icaño, Salado River, no date, Wagner leg.
<specimenCount box="[478,579,565,587]" pageId="22" pageNumber="995" type="male">28 males</specimenCount>
and
<specimenCount box="[638,761,565,587]" pageId="22" pageNumber="995" type="female">11 females</specimenCount>
(
<collectionCode box="[152,242,596,618]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="22" pageNumber="995">MNHN</collectionCode>
); Avellaneda, Icaño, Salado River, La Palma del Bracho, no further data
<specimenCount box="[516,639,626,648]" pageId="22" pageNumber="995" type="male">two males</specimenCount>
(
<collectionCode box="[657,748,627,649]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="22" pageNumber="995">MNHN</collectionCode>
); Avellaneda, Icaño, Mistol Paso, banks of the Salado River, III or
<date box="[296,401,688,709]" pageId="22" pageNumber="995" value="1909-12">XII.1909</date>
, E. R. Wagner leg.
<specimenCount box="[653,760,688,710]" pageId="22" pageNumber="995" type="male">one male</specimenCount>
(
<collectionCode box="[152,237,719,740]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="22" pageNumber="995">NHMB</collectionCode>
ex Frey collection); Avellaneda, southern
<collectingRegion box="[145,215,749,771]" country="Argentina" name="Chaco" pageId="22" pageNumber="995">Chaco</collectingRegion>
north of Icaño (
<collectingRegion box="[396,468,749,771]" country="Argentina" name="Chaco" pageId="22" pageNumber="995">Chaco</collectingRegion>
Austral au N dIcaño), no further data
<specimenCount box="[324,437,780,802]" pageId="22" pageNumber="995" type="male">six males</specimenCount>
and
<specimenCount box="[499,641,780,802]" pageId="22" pageNumber="995" type="female">two females</specimenCount>
(
<collectionCode box="[658,748,780,802]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="22" pageNumber="995">MNHN</collectionCode>
); Banda, La Banda,
<date box="[358,449,811,832]" pageId="22" pageNumber="995" value="1941-11">XI.1941</date>
, Alvaredo leg.
<specimenCount box="[637,761,810,832]" pageId="22" pageNumber="995" type="female">one female</specimenCount>
(
<collectionCode box="[152,236,841,863]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="22" pageNumber="995" type="Museum">MACN</collectionCode>
); Banda, La Banda,
<date box="[465,554,841,863]" pageId="22" pageNumber="995" value="1960-04">IV.1960</date>
, no collector
<specimenCount pageId="22" pageNumber="995" type="male">one male</specimenCount>
and
<specimenCount box="[269,403,872,894]" pageId="22" pageNumber="995" type="female">one female</specimenCount>
(
<collectionCode box="[421,503,872,893]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="22" pageNumber="995" type="Herbarium">TAMU</collectionCode>
ex W. D. Edmonds collection); Barrancas, no further data
<specimenCount box="[608,711,902,924]" pageId="22" pageNumber="995" type="male">one male</specimenCount>
and
<specimenCount box="[145,283,933,955]" pageId="22" pageNumber="995" type="female">two females</specimenCount>
(
<collectionCode box="[298,386,933,955]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="22" pageNumber="995">MNHN</collectionCode>
);
<collectingRegion box="[408,480,933,955]" country="Argentina" name="Chaco" pageId="22" pageNumber="995">Chaco</collectingRegion>
de Santiago, no further data
<specimenCount box="[228,368,964,986]" pageId="22" pageNumber="995" type="male">three males</specimenCount>
and
<specimenCount box="[429,558,964,986]" pageId="22" pageNumber="995" type="female">one female</specimenCount>
(
<collectionCode box="[574,651,964,986]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="22" pageNumber="995">NHRS</collectionCode>
) and
<specimenCount pageId="22" pageNumber="995" type="male">one male</specimenCount>
(
<collectionCode box="[219,300,994,1016]" pageId="22" pageNumber="995">SMTD</collectionCode>
);
<collectingRegion box="[323,397,994,1016]" country="Argentina" name="Chaco" pageId="22" pageNumber="995">Chaco</collectingRegion>
de Santiago, Salado River, no further data
<specimenCount box="[314,435,1025,1047]" pageId="22" pageNumber="995" type="male">four males</specimenCount>
and
<specimenCount box="[493,649,1025,1047]" pageId="22" pageNumber="995" type="female">three females</specimenCount>
(
<collectionCode box="[664,749,1025,1047]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="22" pageNumber="995" type="Museum">RBINS</collectionCode>
),
<specimenCount box="[145,297,1056,1078]" pageId="22" pageNumber="995" type="female">four females</specimenCount>
(
<collectionCode box="[315,401,1056,1077]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="22" pageNumber="995" type="Museum">RBINS</collectionCode>
ex A. Janssens collection),
<specimenCount box="[145,289,1086,1108]" pageId="22" pageNumber="995" type="male">seven males</specimenCount>
and
<specimenCount box="[351,485,1086,1108]" pageId="22" pageNumber="995" type="female">six females</specimenCount>
(
<collectionCode box="[501,586,1086,1107]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="22" pageNumber="995" type="Museum">RBINS</collectionCode>
ex Le Moult collection) and
<specimenCount box="[314,413,1117,1139]" pageId="22" pageNumber="995" type="male">48 males</specimenCount>
(two dissected) and
<specimenCount box="[640,761,1117,1139]" pageId="22" pageNumber="995" type="female">39 females</specimenCount>
(
<collectionCode box="[152,234,1148,1169]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="22" pageNumber="995" type="Museum">RBINS</collectionCode>
ex Wagner and Le Moult collections);
<collectingRegion box="[689,760,1148,1170]" country="Argentina" name="Chaco" pageId="22" pageNumber="995">Chaco</collectingRegion>
de Santiago, Salado River, I.(no year), no further data
<specimenCount box="[164,287,1209,1231]" pageId="22" pageNumber="995" type="female">one female</specimenCount>
(
<collectionCode box="[300,382,1209,1230]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="22" pageNumber="995" type="Museum">RBINS</collectionCode>
ex Le Moult collection) and
<specimenCount pageId="22" pageNumber="995" type="male">two males</specimenCount>
(one dissected) and
<specimenCount box="[460,603,1240,1262]" pageId="22" pageNumber="995" type="female">five females</specimenCount>
(
<collectionCode box="[619,704,1240,1261]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="22" pageNumber="995" type="Museum">RBINS</collectionCode>
ex Wagner and Le Moult collections);
<collectingRegion box="[539,610,1270,1292]" country="Argentina" name="Chaco" pageId="22" pageNumber="995">Chaco</collectingRegion>
de Santiago, Salado River, II.(no year), no further data
<specimenCount box="[638,761,1301,1323]" pageId="22" pageNumber="995" type="female">one female</specimenCount>
(
<collectionCode box="[152,234,1332,1353]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="22" pageNumber="995" type="Museum">RBINS</collectionCode>
ex Le Moult collection) and
<specimenCount box="[576,711,1332,1354]" pageId="22" pageNumber="995" type="male">three males</specimenCount>
and
<specimenCount box="[145,297,1362,1384]" pageId="22" pageNumber="995" type="female">two females</specimenCount>
(
<collectionCode box="[317,406,1362,1383]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="22" pageNumber="995" type="Museum">RBINS</collectionCode>
ex Wagner and Le Moult collections);
<collectingRegion box="[291,363,1393,1415]" country="Argentina" name="Chaco" pageId="22" pageNumber="995">Chaco</collectingRegion>
de Santiago, Salado River, III.(no year), no further data
<specimenCount box="[436,564,1424,1446]" pageId="22" pageNumber="995" type="female">one female</specimenCount>
(
<collectionCode box="[580,665,1424,1445]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="22" pageNumber="995" type="Museum">RBINS</collectionCode>
ex Le Moult collection);
<collectingRegion box="[362,435,1454,1476]" country="Argentina" name="Chaco" pageId="22" pageNumber="995">Chaco</collectingRegion>
de Santiago, Salado River, IV.(no year), no further data
<specimenCount box="[530,654,1485,1507]" pageId="22" pageNumber="995" type="male">two males</specimenCount>
and
<specimenCount pageId="22" pageNumber="995" type="female">one female</specimenCount>
(
<collectionCode box="[235,317,1516,1537]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="22" pageNumber="995" type="Museum">RBINS</collectionCode>
ex Wagner and Le Moult collections);
<collectingRegion box="[149,224,1546,1568]" country="Argentina" name="Chaco" pageId="22" pageNumber="995">Chaco</collectingRegion>
de Santiago, Salado River, IX.(no year), no further data
<specimenCount box="[318,423,1577,1599]" pageId="22" pageNumber="995" type="male">one male</specimenCount>
(
<collectionCode box="[438,521,1577,1598]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="22" pageNumber="995" type="Museum">RBINS</collectionCode>
ex Wagner and Le Moult collections);
<collectingRegion box="[369,441,1608,1630]" country="Argentina" name="Chaco" pageId="22" pageNumber="995">Chaco</collectingRegion>
de Santiago, Salado River, XII.(no year), no further data
<specimenCount box="[507,610,1638,1660]" pageId="22" pageNumber="995" type="male">one male</specimenCount>
(
<collectionCode box="[625,707,1638,1659]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="22" pageNumber="995" type="Museum">RBINS</collectionCode>
ex Le Moult collection) and
<specimenCount box="[440,579,1669,1691]" pageId="22" pageNumber="995" type="male">three males</specimenCount>
(one dissected) and
<specimenCount box="[197,342,1700,1722]" pageId="22" pageNumber="995" type="female">four females</specimenCount>
(
<collectionCode box="[358,442,1700,1721]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="22" pageNumber="995" type="Museum">RBINS</collectionCode>
ex Wagner and Le Moult collections);
<collectingRegion box="[294,367,1730,1752]" country="Argentina" name="Chaco" pageId="22" pageNumber="995">Chaco</collectingRegion>
de Santiago, Salado River, 1909, no collector
<specimenCount box="[307,416,1761,1783]" pageId="22" pageNumber="995" type="male">six males</specimenCount>
and
<specimenCount box="[476,607,1761,1783]" pageId="22" pageNumber="995" type="female">six females</specimenCount>
(
<collectionCode box="[622,706,1761,1782]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="22" pageNumber="995" type="Museum">RBINS</collectionCode>
ex Le Moult collection); Choya,
<date box="[490,587,1791,1813]" pageId="22" pageNumber="995" value="1962-03">III.1962</date>
, no collector
<specimenCount box="[145,283,1822,1844]" pageId="22" pageNumber="995" type="male">three males</specimenCount>
and
<specimenCount box="[343,469,1822,1844]" pageId="22" pageNumber="995" type="female">one female</specimenCount>
(
<collectionCode box="[484,563,1822,1844]" pageId="22" pageNumber="995">MLPA</collectionCode>
); Choya,
<date box="[670,756,1822,1844]" pageId="22" pageNumber="995" value="1963-02">II.1963</date>
, Andrae (?) leg.
<specimenCount box="[350,456,1853,1875]" pageId="22" pageNumber="995" type="male">one male</specimenCount>
(
<collectionCode box="[473,559,1853,1875]" pageId="22" pageNumber="995">CMNC</collectionCode>
); Choya,
<date box="[669,756,1853,1875]" pageId="22" pageNumber="995" value="1963-02">II.1963</date>
, Forster leg.
<specimenCount box="[308,414,1883,1905]" pageId="22" pageNumber="995" type="male">one male</specimenCount>
(
<collectionCode box="[430,515,1884,1906]" pageId="22" pageNumber="995">CMNC</collectionCode>
); Choya,
<date box="[623,719,1884,1905]" pageId="22" pageNumber="995" value="1974-03">III.1974</date>
, no collector
<specimenCount box="[936,1042,197,219]" pageId="22" pageNumber="995" type="male">one male</specimenCount>
and
<specimenCount box="[1101,1227,197,219]" pageId="22" pageNumber="995" type="female">one female</specimenCount>
(
<collectionCode box="[1243,1326,197,219]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="22" pageNumber="995" type="Herbarium">TAMU</collectionCode>
); Choya,
<date box="[809,928,228,249]" pageId="22" pageNumber="995" value="1974-08">VIII.1974</date>
, no collector
<specimenCount box="[1117,1227,227,249]" pageId="22" pageNumber="995" type="male">one male</specimenCount>
(
<collectionCode box="[1245,1320,228,250]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="22" pageNumber="995" type="Museum">FSCA</collectionCode>
); Choya, Frías,
<date box="[887,994,258,280]" pageId="22" pageNumber="995" value="1952-12">XII.1952</date>
, no collector
<specimenCount box="[1179,1310,258,280]" pageId="22" pageNumber="995" type="female">one female</specimenCount>
(
<collectionCode box="[1327,1413,258,280]" pageId="22" pageNumber="995">CMNC</collectionCode>
); Choya, Frías,
<date box="[985,1065,289,311]" pageId="22" pageNumber="995" value="2006-01">I.2006</date>
, E. Abadie leg.
<specimenCount box="[1290,1424,289,311]" pageId="22" pageNumber="995" type="female">one female</specimenCount>
(
<collectionCode box="[816,896,320,342]" pageId="22" pageNumber="995">CEMT</collectionCode>
); Choya, Villa la Punta, no further data
<specimenCount pageId="22" pageNumber="995" type="female">one female</specimenCount>
(
<collectionCode box="[898,982,350,372]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="22" pageNumber="995" type="Museum">MACN</collectionCode>
); Jiménez, El Charco,
<date box="[1230,1314,350,372]" pageId="22" pageNumber="995" value="1960-02">II.1960</date>
, Berthold leg.
<specimenCount box="[888,1014,381,403]" pageId="22" pageNumber="995" type="male">two males</specimenCount>
and
<specimenCount box="[1083,1233,381,403]" pageId="22" pageNumber="995" type="female">two females</specimenCount>
(
<collectionCode box="[1252,1314,381,403]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="22" pageNumber="995" type="Museum">ZSM</collectionCode>
); Loreto,
<date box="[809,897,411,433]" pageId="22" pageNumber="995" value="1963-04">IV.1963</date>
, no collector
<specimenCount box="[1064,1187,411,433]" pageId="22" pageNumber="995" type="female">one female</specimenCount>
(
<collectionCode box="[1200,1277,412,434]" pageId="22" pageNumber="995">MLPA</collectionCode>
); Mitre, Villa Unión, no further data
<specimenCount box="[1103,1208,442,464]" pageId="22" pageNumber="995" type="male">one male</specimenCount>
(
<collectionCode box="[1223,1312,442,464]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="22" pageNumber="995">MNHN</collectionCode>
); Moreno, Puna,
<date box="[888,1032,473,494]" pageId="22" pageNumber="995" value="1939-12-11">11.XII.1939</date>
, Biraben-Bezzi leg.
<specimenCount box="[1302,1425,473,495]" pageId="22" pageNumber="995" type="male">two males</specimenCount>
(
<collectionCode box="[816,893,504,526]" pageId="22" pageNumber="995">MLPA</collectionCode>
); Ojo de Água,
<date box="[1062,1146,504,525]" pageId="22" pageNumber="995" value="1974-02">II.1974</date>
, A. Martínez leg.
<specimenCount pageId="22" pageNumber="995" type="male">three males</specimenCount>
and
<specimenCount box="[937,1083,534,556]" pageId="22" pageNumber="995" type="female">four females</specimenCount>
(
<collectionCode box="[1099,1178,534,556]" pageId="22" pageNumber="995">BDGC</collectionCode>
); Ojo de Água, Dique de Baez,
<date box="[911,994,565,586]" pageId="22" pageNumber="995" value="1974-02">II.1974</date>
, no collector
<specimenCount box="[1162,1277,565,587]" pageId="22" pageNumber="995" type="male">two males</specimenCount>
(
<collectionCode box="[1292,1372,565,586]" pageId="22" pageNumber="995">CMNC</collectionCode>
ex A. Martínez and Henry &amp; Anne Howden collections) and
<specimenCount box="[859,962,626,648]" pageId="22" pageNumber="995" type="male">one male</specimenCount>
(dissected) (
<collectionCode box="[1105,1177,626,647]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="22" pageNumber="995" type="University or college">DZUP</collectionCode>
ex A. Martínez and Henry &amp; Anne Howden collections, and
<collectionCode box="[1282,1366,657,679]" pageId="22" pageNumber="995">CMNC</collectionCode>
); Ojo de Água, Dique de Baez (
<emphasis box="[1108,1297,687,709]" italics="true" pageId="22" pageNumber="995">Represa de Baez</emphasis>
),
<date box="[1325,1420,688,709]" pageId="22" pageNumber="995" value="1974-03">III.1974</date>
, no collector
<specimenCount box="[990,1104,718,740]" pageId="22" pageNumber="995" type="male">one male</specimenCount>
(
<collectionCode box="[1122,1202,718,739]" pageId="22" pageNumber="995">BDGC</collectionCode>
ex A. Martínez collection); Salado River, no date, Wagner leg.
<specimenCount pageId="22" pageNumber="995" type="male">one male</specimenCount>
(
<collectionCode box="[878,962,780,802]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="22" pageNumber="995" type="Museum">MACN</collectionCode>
),
<specimenCount box="[978,1112,779,801]" pageId="22" pageNumber="995" type="male">three males</specimenCount>
and
<specimenCount box="[1167,1290,779,801]" pageId="22" pageNumber="995" type="female">one female</specimenCount>
(
<collectionCode box="[1303,1380,780,802]" pageId="22" pageNumber="995">MLPA</collectionCode>
),
<specimenCount pageId="22" pageNumber="995" type="male">15 males</specimenCount>
and
<specimenCount box="[934,1056,810,832]" pageId="22" pageNumber="995" type="female">15 females</specimenCount>
(
<collectionCode box="[1072,1139,810,832]" country="Belgium" httpUri="http://grbio.org/cool/wzbt-fxwr" name="Institut Royal des Sciences Naturelles de Belgique" pageId="22" pageNumber="995">IRSN</collectionCode>
).
<emphasis box="[1157,1300,809,833]" italics="true" pageId="22" pageNumber="995">
<collectingRegion box="[1157,1293,809,833]" country="Argentina" name="Tucuman" pageId="22" pageNumber="995">TUCUMÁN</collectingRegion>
:
</emphasis>
no further data
<specimenCount box="[887,990,841,863]" pageId="22" pageNumber="995" type="male">one male</specimenCount>
(
<collectionCode box="[1004,1087,841,862]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="22" pageNumber="995">NHMB</collectionCode>
ex H. Marchand collection); further data illegible
<specimenCount box="[1071,1194,871,893]" pageId="22" pageNumber="995" type="female">one female</specimenCount>
(
<collectionCode box="[1207,1291,872,894]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="22" pageNumber="995" type="Museum">MACN</collectionCode>
); no specific locality,
<date box="[913,1014,902,924]" pageId="22" pageNumber="995" value="1950-03">III.1950</date>
, no collector
<specimenCount box="[1205,1316,902,924]" pageId="22" pageNumber="995" type="male">one male</specimenCount>
(
<collectionCode box="[1334,1412,902,924]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="22" pageNumber="995" type="Museum">MZSP</collectionCode>
); Burruyacú,
<date box="[944,1038,933,955]" pageId="22" pageNumber="995" value="1946-03">III.1946</date>
, Martínez leg.
<specimenCount box="[1224,1326,933,955]" pageId="22" pageNumber="995" type="male">one male</specimenCount>
(
<collectionCode box="[1339,1413,933,955]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="22" pageNumber="995" type="Museum">MZSP</collectionCode>
); Burruyacú, Burruyacú,
<date box="[1099,1198,963,985]" pageId="22" pageNumber="995" value="1946-03">III.1946</date>
, no collector
<specimenCount pageId="22" pageNumber="995" type="male">one male</specimenCount>
(
<collectionCode box="[878,961,994,1016]" pageId="22" pageNumber="995">CMNC</collectionCode>
); Cerro San Javier,
<date box="[1179,1281,994,1015]" pageId="22" pageNumber="995" value="1948-12">XII.1948</date>
, A. Martínez leg.
<specimenCount box="[879,985,1025,1047]" pageId="22" pageNumber="995" type="male">one male</specimenCount>
(
<collectionCode box="[1001,1077,1025,1047]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="22" pageNumber="995" type="Museum">MZSP</collectionCode>
); Tinolina, no further data
<specimenCount box="[809,911,1055,1077]" pageId="22" pageNumber="995" type="male">one male</specimenCount>
(
<collectionCode box="[924,1012,1055,1077]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="22" pageNumber="995">MNHN</collectionCode>
); Trancas, Tapia,
<date box="[1209,1317,1055,1077]" pageId="22" pageNumber="995" value="2009-01-05">05.I.2009</date>
, O. Boilly leg.
<specimenCount box="[881,990,1086,1108]" pageId="22" pageNumber="995" type="male">one male</specimenCount>
(
<collectionCode box="[1006,1085,1086,1108]" pageId="22" pageNumber="995">OBCL</collectionCode>
).
<emphasis bold="true" box="[1105,1234,1086,1108]" pageId="22" pageNumber="995">
<collectingCountry box="[1105,1229,1086,1108]" name="Bolivia" pageId="22" pageNumber="995">BOLIVIA</collectingCountry>
:
</emphasis>
<emphasis box="[1243,1425,1086,1107]" italics="true" pageId="22" pageNumber="995">
<collectingRegion box="[1243,1419,1086,1107]" country="Bolivia" name="Santa Cruz" pageId="22" pageNumber="995">SANTA CRUZ</collectingRegion>
:
</emphasis>
Andrés Ibáñez, between Guayabas and Basilio,
<geoCoordinate box="[809,944,1147,1169]" degrees="18" direction="south" minutes="01" orientation="latitude" pageId="22" pageNumber="995" precision="15" seconds="06" value="-18.018333">18°0106”S</geoCoordinate>
,
<geoCoordinate box="[960,1101,1147,1169]" degrees="63" direction="west" minutes="11" orientation="longitude" pageId="22" pageNumber="995" precision="15" seconds="24" value="-63.190002">63°1124”W</geoCoordinate>
,
<quantity box="[1117,1197,1147,1169]" metricMagnitude="2" metricUnit="m" metricValue="4.57" pageId="22" pageNumber="995" unit="m" value="457.0">457 m</quantity>
,
<date box="[1212,1330,1147,1169]" pageId="22" pageNumber="995" value="2005-01-01">01.I.2005</date>
, Mann, Hamel and Herzog leg.
<specimenCount box="[1140,1277,1178,1200]" pageId="22" pageNumber="995" type="female">one female</specimenCount>
(
<collectionCode box="[1297,1411,1178,1200]" pageId="22" pageNumber="995">OUMNH</collectionCode>
);
<collectingRegion box="[809,934,1209,1231]" country="Paraguay" name="Cordillera" pageId="22" pageNumber="995">Cordillera</collectingRegion>
, Camiri,
<geoCoordinate box="[1048,1179,1208,1230]" degrees="20" direction="south" minutes="05" orientation="latitude" pageId="22" pageNumber="995" precision="15" seconds="45" value="-20.095835">20°0545”S</geoCoordinate>
,
<geoCoordinate box="[1193,1330,1209,1231]" degrees="63" direction="west" minutes="30" orientation="longitude" pageId="22" pageNumber="995" precision="15" seconds="10" value="-63.502777">63°3010”W</geoCoordinate>
,
<quantity box="[1344,1420,1209,1231]" metricMagnitude="2" metricUnit="m" metricValue="7.859999999999999" pageId="22" pageNumber="995" unit="m" value="786.0">786 m</quantity>
,
<date box="[809,917,1239,1261]" pageId="22" pageNumber="995" value="2005-01-02">02.I.2005</date>
, Mann and Hamel leg.
<specimenCount box="[1195,1309,1239,1261]" pageId="22" pageNumber="995" type="male">five males</specimenCount>
and
<specimenCount pageId="22" pageNumber="995" type="female">three females</specimenCount>
(
<collectionCode box="[912,1020,1270,1292]" pageId="22" pageNumber="995">OUMNH</collectionCode>
);
<collectingRegion box="[1039,1158,1270,1292]" country="Paraguay" name="Cordillera" pageId="22" pageNumber="995">Cordillera</collectingRegion>
, San Antonio,
<date box="[1326,1420,1270,1292]" pageId="22" pageNumber="995" value="1986-03">III.1986</date>
, Malkin (?) leg.
<specimenCount box="[1036,1151,1301,1323]" pageId="22" pageNumber="995" type="male">one male</specimenCount>
(
<collectionCode box="[1172,1262,1301,1323]" pageId="22" pageNumber="995">CMNC</collectionCode>
);
<collectingRegion box="[1287,1418,1301,1323]" country="Paraguay" name="Cordillera" pageId="22" pageNumber="995">Cordillera</collectingRegion>
, Tatarenda, Chaco, no further data
<specimenCount box="[1223,1323,1331,1353]" pageId="22" pageNumber="995" type="male">one male</specimenCount>
(
<collectionCode box="[1336,1414,1331,1353]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="22" pageNumber="995">NHRS</collectionCode>
).
<emphasis box="[809,913,1362,1383]" italics="true" pageId="22" pageNumber="995">
<collectingRegion box="[809,907,1362,1383]" country="Bolivia" name="Tarija" pageId="22" pageNumber="995">TARIJA</collectingRegion>
:
</emphasis>
Gran Chaco, El Bagual (between Caiza &amp; Creveux),
<geoCoordinate box="[935,1063,1392,1414]" degrees="21" direction="south" minutes="50" orientation="latitude" pageId="22" pageNumber="995" precision="15" seconds="19" value="-21.838612">21°5019”S</geoCoordinate>
, 63°2468”W,
<quantity box="[1222,1296,1393,1414]" metricMagnitude="2" metricUnit="m" metricValue="4.87" pageId="22" pageNumber="995" unit="m" value="487.0">487 m</quantity>
,
<date box="[1309,1420,1392,1414]" pageId="22" pageNumber="995" value="2005-01-04">04.I.2005</date>
, Mann, Hamel and Herzog leg.
<specimenCount box="[1219,1330,1423,1445]" pageId="22" pageNumber="995" type="male">15 males</specimenCount>
and
<specimenCount pageId="22" pageNumber="995" type="female">11 females</specimenCount>
(
<collectionCode box="[913,1021,1454,1476]" pageId="22" pageNumber="995">OUMNH</collectionCode>
); Gran Chaco, El Bagual (between Caiza &amp; Creveux),
<geoCoordinate box="[1047,1178,1484,1506]" degrees="21" direction="south" minutes="50" orientation="latitude" pageId="22" pageNumber="995" precision="15" seconds="19" value="-21.838612">21°5019”S</geoCoordinate>
, 63°2468”W,
<quantity box="[1343,1420,1485,1506]" metricMagnitude="2" metricUnit="m" metricValue="4.87" pageId="22" pageNumber="995" unit="m" value="487.0">487 m</quantity>
,
<date box="[809,967,1515,1537]" pageId="22" pageNumber="995" value="2005-01-04" valueMax="2005-01-05" valueMin="2005-01-04">0405.I.2005</date>
, Mann, Hamel and Herzog leg.
<specimenCount pageId="22" pageNumber="995" type="male">two males</specimenCount>
and
<specimenCount box="[952,1109,1546,1568]" pageId="22" pageNumber="995" type="female">two females</specimenCount>
; Gran Chaco, El Bagual (between Caiza &amp; Creveux),
<geoCoordinate box="[1153,1279,1576,1598]" degrees="21" direction="south" minutes="50" orientation="latitude" pageId="22" pageNumber="995" precision="15" seconds="19" value="-21.838612">21°5019”S</geoCoordinate>
, 63°2468”W [
<emphasis box="[817,848,1608,1629]" italics="true" pageId="22" pageNumber="995">sic</emphasis>
],
<quantity box="[870,944,1607,1629]" metricMagnitude="2" metricUnit="m" metricValue="4.87" pageId="22" pageNumber="995" unit="m" value="487.0">487 m</quantity>
,
<date box="[955,1065,1607,1629]" pageId="22" pageNumber="995" value="2005-01-06">06.I.2005</date>
, Mann, Hamel and Herzog leg.
<specimenCount box="[831,936,1638,1660]" pageId="22" pageNumber="995" type="male">one male</specimenCount>
and
<specimenCount box="[996,1136,1638,1660]" pageId="22" pageNumber="995" type="female">two females</specimenCount>
(
<collectionCode box="[1151,1260,1638,1660]" pageId="22" pageNumber="995">OUMNH</collectionCode>
); Gran Chaco, Villamontes,
<date box="[959,1033,1669,1690]" pageId="22" pageNumber="995" value="1931-01">I.1931</date>
, Eisennhauty leg.
<specimenCount box="[1266,1373,1669,1691]" pageId="22" pageNumber="995" type="male">six males</specimenCount>
and
<specimenCount box="[809,950,1699,1721]" pageId="22" pageNumber="995" type="female">four females</specimenCount>
(
<collectionCode box="[965,1046,1699,1721]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="22" pageNumber="995">MFNB</collectionCode>
); Gran Chaco, Yacuiba, Yaguacua (Sachapera-Yaguacua),
<quantity box="[1089,1161,1730,1752]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="22" pageNumber="995" unit="m" value="680.0">680 m</quantity>
,
<date box="[1173,1247,1730,1752]" pageId="22" pageNumber="995" value="2005-01">I.2005</date>
, D. Mann leg.
<specimenCount box="[809,909,1761,1783]" pageId="22" pageNumber="995" type="male">one male</specimenCount>
(dissected) (
<collectionCode box="[1047,1126,1761,1782]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="22" pageNumber="995">NMPC</collectionCode>
ex S. Pokorný collection); Gran Chaco, Yacuiba, Yaguacua (Sachapera-Yaguacua) [transition between Chaco and Chaco Serrano zone;
<emphasis box="[974,1051,1853,1874]" italics="true" pageId="22" pageNumber="995">Acacia</emphasis>
scrub, open habitat; loamy soil],
<geoCoordinate box="[809,943,1883,1904]" degrees="21" direction="south" minutes="41" orientation="latitude" pageId="22" pageNumber="995" precision="15" seconds="09" value="-21.685833">21°4109”S</geoCoordinate>
,
<geoCoordinate box="[960,1101,1883,1905]" degrees="63" direction="west" minutes="33" orientation="longitude" pageId="22" pageNumber="995" precision="15" seconds="10" value="-63.552776">63°3310”W</geoCoordinate>
,
<quantity box="[1117,1197,1883,1905]" metricMagnitude="2" metricUnit="m" metricValue="6.76" pageId="22" pageNumber="995" unit="m" value="676.0">676 m</quantity>
,
<date box="[1212,1331,1883,1905]" pageId="22" pageNumber="995" value="2005-01-03">03.I.2005</date>
, Mann, Hamel and Herzog leg.
<specimenCount box="[475,584,197,219]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
and
<specimenCount box="[648,777,197,219]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[170,251,228,250]" pageId="23" pageNumber="996">BDGC</collectionCode>
),
<specimenCount box="[272,419,228,250]" pageId="23" pageNumber="996" type="male">seven males</specimenCount>
and
<specimenCount box="[482,646,228,250]" pageId="23" pageNumber="996" type="female">three females</specimenCount>
(
<collectionCode box="[662,770,228,250]" pageId="23" pageNumber="996">OUMNH</collectionCode>
) and
<specimenCount box="[213,316,258,280]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
(
<collectionCode box="[331,401,259,281]" pageId="23" pageNumber="996">SPPC</collectionCode>
).
<emphasis bold="true" box="[419,576,258,280]" pageId="23" pageNumber="996">
<collectingCountry box="[419,571,258,280]" name="Paraguay" pageId="23" pageNumber="996">PARAGUAY</collectingCountry>
:
</emphasis>
no further data
<specimenCount box="[163,268,289,311]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
(
<collectionCode box="[284,357,289,311]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="23" pageNumber="996" type="Museum">FSCA</collectionCode>
),
<specimenCount box="[376,482,289,311]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
and
<specimenCount box="[541,668,289,311]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[684,771,289,311]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="996">MNHN</collectionCode>
) and
<specimenCount box="[220,358,320,342]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[378,466,320,341]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="23" pageNumber="996">NHMB</collectionCode>
ex Nonfried and Frey collections);
<collectingRegion box="[311,385,350,372]" country="Argentina" name="Chaco" pageId="23" pageNumber="996">Chaco</collectingRegion>
, no date, S. Bolle leg.
<specimenCount box="[672,778,350,372]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
(dissected) (
<collectionCode box="[305,390,381,402]" pageId="23" pageNumber="996">BMNH</collectionCode>
ex J. Zídek collection); Trans-Chaco, no further data
<specimenCount box="[470,577,412,434]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
(
<collectionCode box="[594,675,412,434]" pageId="23" pageNumber="996">CEMT</collectionCode>
); Trans-Chaco, no date, T. Taylor leg.
<specimenCount box="[523,626,442,464]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
(
<collectionCode box="[641,724,443,464]" pageId="23" pageNumber="996">BMNH</collectionCode>
ex J. Zídek collection).
<emphasis box="[448,625,472,496]" italics="true" pageId="23" pageNumber="996">
<collectingRegion box="[448,616,472,496]" country="Paraguay" name="Boqueron" pageId="23" pageNumber="996">BOQUERÓN</collectingRegion>
:
</emphasis>
Filadelfia, Transchaco (Ruta 9) km 430, Drechsel leg.
<specimenCount pageId="23" pageNumber="996" type="female">two females</specimenCount>
(
<collectionCode box="[284,367,535,556]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="23" pageNumber="996" type="Museum">ZFMK</collectionCode>
ex Franz collection); Fortín Guachalla (Guachallas, Alto Pilcomayo),
<date box="[687,773,565,587]" pageId="23" pageNumber="996" value="1946-10">X.1946</date>
, P. Willim leg.
<specimenCount box="[360,471,596,618]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
(
<collectionCode box="[489,577,596,618]" pageId="23" pageNumber="996">CMNC</collectionCode>
).
<emphasis italics="true" pageId="23" pageNumber="996">
<collectingRegion country="Paraguay" name="Presidente Hayes" pageId="23" pageNumber="996">PRESIDENTE HAYES</collectingRegion>
:
</emphasis>
Cruce de los Pioneros,
<date box="[535,646,626,648]" pageId="23" pageNumber="996" value="1996-01-20">20.I.1996</date>
, C. Aguilar leg.
<specimenCount box="[225,338,657,679]" pageId="23" pageNumber="996" type="male">two males</specimenCount>
(
<collectionCode box="[351,430,657,678]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="23" pageNumber="996" type="Herbarium">TAMU</collectionCode>
ex W. D. Edmonds collection); Cruce de los Pioneros,
<date box="[425,533,688,710]" pageId="23" pageNumber="996" value="2003-01-05">05.I.2003</date>
, C. Aguilar leg.
<specimenCount pageId="23" pageNumber="996" type="male">one male</specimenCount>
(
<collectionCode box="[232,311,719,740]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="23" pageNumber="996" type="Herbarium">TAMU</collectionCode>
ex W. D. Edmonds collection); Cruce de los Pioneros,
<date box="[310,447,749,770]" pageId="23" pageNumber="996" value="2003-12-20">20.XII.2003</date>
, no collector
<specimenCount box="[610,729,749,771]" pageId="23" pageNumber="996" type="male">four males</specimenCount>
and
<specimenCount box="[163,320,780,802]" pageId="23" pageNumber="996" type="female">three females</specimenCount>
(
<collectionCode box="[334,413,780,801]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="23" pageNumber="996" type="Herbarium">TAMU</collectionCode>
ex W. D. Edmonds collection); Pozo Colorado,Trans-Chaco,
<date box="[483,572,810,832]" pageId="23" pageNumber="996" value="1995-04">IV.1995</date>
, J. C. DAlessandro leg.
<specimenCount box="[252,388,841,863]" pageId="23" pageNumber="996" type="male">two males</specimenCount>
and
<specimenCount box="[466,652,841,863]" pageId="23" pageNumber="996" type="female">three females</specimenCount>
(
<collectionCode box="[678,759,841,863]" pageId="23" pageNumber="996">CEMT</collectionCode>
).
<emphasis bold="true" box="[163,379,871,893]" pageId="23" pageNumber="996">
<emphasis bold="true" box="[163,372,871,893]" italics="true" pageId="23" pageNumber="996">Ambiguous data</emphasis>
:
</emphasis>
1908
<specimenCount box="[479,603,872,894]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[617,701,872,894]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="23" pageNumber="996" type="Museum">MACN</collectionCode>
); Gran
<collectingRegion box="[163,234,902,924]" country="Argentina" name="Chaco" pageId="23" pageNumber="996">Chaco</collectingRegion>
(Bolivian province of Gran
<collectingRegion box="[556,630,902,924]" country="Argentina" name="Chaco" pageId="23" pageNumber="996">Chaco</collectingRegion>
?), no further data
<specimenCount box="[241,343,933,955]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
(
<collectionCode box="[357,440,933,954]" pageId="23" pageNumber="996">BMNH</collectionCode>
ex Nevinson collection) and
<specimenCount box="[163,268,964,986]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
and
<specimenCount box="[327,453,964,986]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[469,547,964,986]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="23" pageNumber="996">NHRS</collectionCode>
); Jacanto, 1930, no collector
<specimenCount box="[285,387,994,1016]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
(
<collectionCode box="[401,485,994,1016]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="23" pageNumber="996" type="Museum">MACN</collectionCode>
); J. Caño E W
<specimenCount box="[676,779,994,1016]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
and
<specimenCount box="[215,343,1025,1047]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[358,437,1025,1047]" pageId="23" pageNumber="996">MLPA</collectionCode>
);
<emphasis box="[461,554,1026,1047]" italics="true" pageId="23" pageNumber="996">Pampas</emphasis>
, no further data
<specimenCount box="[163,286,1056,1078]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[300,382,1056,1077]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="23" pageNumber="996" type="Museum">RBINS</collectionCode>
ex E. Candèze collection);
<emphasis box="[708,768,1056,1077]" italics="true" pageId="23" pageNumber="996">Mont</emphasis>
. (Montevideo or Monte, Argentina?), no further data
<specimenCount box="[163,269,1117,1139]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
(
<collectionCode box="[285,365,1117,1138]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="23" pageNumber="996">NMPC</collectionCode>
ex
<collectionCode box="[430,486,1117,1138]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="23" pageNumber="996">SMF</collectionCode>
and E. Witte collection); Parana. Febr. (province of
<collectingRegion box="[503,665,1148,1170]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="23" pageNumber="996">Buenos Aires</collectingRegion>
or
<collectingRegion country="Argentina" name="Entre Rios" pageId="23" pageNumber="996">Entre Ríos</collectingRegion>
?), no further data
<specimenCount box="[481,616,1178,1200]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[633,717,1178,1199]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="23" pageNumber="996" type="Museum">MACN</collectionCode>
ex Hermann Burmeister collection); Parana. Mrz. (province of
<collectingRegion box="[307,460,1240,1262]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="23" pageNumber="996">Buenos Aires</collectingRegion>
or
<collectingRegion box="[500,629,1240,1262]" country="Argentina" name="Entre Rios" pageId="23" pageNumber="996">Entre Ríos</collectingRegion>
?), no further data
<specimenCount box="[247,355,1270,1292]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
(
<collectionCode box="[371,454,1270,1291]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="23" pageNumber="996" type="Museum">MACN</collectionCode>
ex Hermann Burmeister collection); S. Corral,
<date box="[415,507,1301,1322]" pageId="23" pageNumber="996" value="1907-11">XI.1907</date>
, collector illegible
<specimenCount pageId="23" pageNumber="996" type="male">one male</specimenCount>
(
<collectionCode box="[234,318,1332,1354]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="23" pageNumber="996" type="Museum">MACN</collectionCode>
); S. Curral,
<date box="[467,569,1332,1353]" pageId="23" pageNumber="996" value="1907-12">XII.1907</date>
, illegible collector
<specimenCount box="[185,329,1362,1384]" pageId="23" pageNumber="996" type="female">two females</specimenCount>
(
<collectionCode box="[345,432,1362,1384]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="23" pageNumber="996" type="Museum">MACN</collectionCode>
).
<emphasis bold="true" box="[452,640,1362,1384]" pageId="23" pageNumber="996">
<emphasis bold="true" box="[452,632,1362,1384]" italics="true" pageId="23" pageNumber="996">Illegible data</emphasis>
:
</emphasis>
<specimenCount box="[649,779,1362,1384]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[170,255,1393,1415]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="23" pageNumber="996" type="Museum">MACN</collectionCode>
).
<emphasis bold="true" box="[273,779,1392,1415]" pageId="23" pageNumber="996">
<emphasis bold="true" box="[273,590,1392,1414]" italics="true" pageId="23" pageNumber="996">Probably erroneous data</emphasis>
:
<collectingCountry box="[603,773,1393,1415]" name="Argentina" pageId="23" pageNumber="996">ARGENTINA</collectingCountry>
:
</emphasis>
<emphasis box="[163,371,1424,1445]" italics="true" pageId="23" pageNumber="996">
<collectingRegion box="[163,365,1424,1445]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="23" pageNumber="996">BUENOS AIRES</collectingRegion>
:
</emphasis>
no further data
<specimenCount box="[589,728,1424,1446]" pageId="23" pageNumber="996" type="male">three males</specimenCount>
and
<specimenCount box="[163,308,1454,1476]" pageId="23" pageNumber="996" type="female">two females</specimenCount>
(
<collectionCode box="[326,409,1454,1476]" pageId="23" pageNumber="996">SMTD</collectionCode>
).
<emphasis bold="true" box="[429,545,1454,1475]" pageId="23" pageNumber="996">
<collectingCountry box="[429,539,1454,1475]" name="Brazil" pageId="23" pageNumber="996">BRAZIL</collectingCountry>
:
</emphasis>
<emphasis italics="true" pageId="23" pageNumber="996">
<collectingRegion country="Brazil" name="Rio Grande do Sul" pageId="23" pageNumber="996">RIO GRANDE DO SUL</collectingRegion>
:
</emphasis>
Porto Alegre, no further data
<specimenCount box="[650,777,1485,1507]" pageId="23" pageNumber="996" type="male">two males</specimenCount>
(
<collectionCode box="[171,263,1516,1538]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="23" pageNumber="996" type="Museum">MHNG</collectionCode>
).
<emphasis bold="true" box="[284,434,1515,1537]" pageId="23" pageNumber="996">
<collectingCountry box="[284,429,1515,1537]" name="Uruguay" pageId="23" pageNumber="996">URUGUAY</collectingCountry>
:
</emphasis>
<emphasis box="[444,641,1516,1537]" italics="true" pageId="23" pageNumber="996">
<collectingRegion box="[444,635,1516,1537]" country="Uruguay" name="Montevideo" pageId="23" pageNumber="996">MONTEVIDEO</collectingRegion>
:
</emphasis>
no further data
<specimenCount box="[247,354,1546,1568]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
(
<collectionCode box="[370,455,1546,1567]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="23" pageNumber="996" type="Museum">RBINS</collectionCode>
ex J. J. Gillet collection).
<emphasis bold="true" box="[163,469,1576,1599]" pageId="23" pageNumber="996">
<emphasis bold="true" box="[163,360,1576,1598]" italics="true" pageId="23" pageNumber="996">Erroneous data</emphasis>
:
<collectingCountry box="[373,463,1577,1599]" name="Chile" pageId="23" pageNumber="996">CHILE</collectingCountry>
:
</emphasis>
no further data
<specimenCount box="[676,779,1577,1599]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
(
<collectionCode box="[170,255,1608,1630]" pageId="23" pageNumber="996">BMNH</collectionCode>
) and
<specimenCount box="[321,439,1608,1630]" pageId="23" pageNumber="996" type="male">two males</specimenCount>
(
<collectionCode box="[454,543,1608,1630]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="996">MNHN</collectionCode>
); no specific locality, 1870, Philippi leg.
<specimenCount box="[394,496,1638,1660]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
(
<collectionCode box="[510,603,1638,1660]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="23" pageNumber="996" type="Museum">NHMW</collectionCode>
).
<emphasis bold="true" box="[620,779,1638,1660]" pageId="23" pageNumber="996">
<collectingCountry box="[620,774,1638,1660]" name="Colombia" pageId="23" pageNumber="996">COLOMBIA</collectingCountry>
:
</emphasis>
<emphasis box="[163,423,1669,1690]" italics="true" pageId="23" pageNumber="996">DISTRITO CAPITAL:</emphasis>
<collectingRegion box="[431,514,1669,1691]" country="Colombia" name="Bogota" pageId="23" pageNumber="996">Bogotá</collectingRegion>
, no further data
<specimenCount pageId="23" pageNumber="996" type="male">one male</specimenCount>
(
<collectionCode box="[240,333,1700,1722]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="996">MNHN</collectionCode>
).
<emphasis bold="true" box="[355,472,1699,1721]" pageId="23" pageNumber="996">
<emphasis bold="true" box="[355,464,1699,1721]" italics="true" pageId="23" pageNumber="996">No data</emphasis>
:
</emphasis>
<specimenCount box="[482,613,1700,1722]" pageId="23" pageNumber="996" type="male">four males</specimenCount>
(
<collectionCode box="[631,717,1700,1721]" pageId="23" pageNumber="996">BMNH</collectionCode>
ex Nevinson collection),
<specimenCount box="[428,577,1730,1752]" pageId="23" pageNumber="996" type="male">seven males</specimenCount>
and
<specimenCount box="[641,778,1730,1752]" pageId="23" pageNumber="996" type="female">six females</specimenCount>
(
<collectionCode box="[171,261,1761,1783]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="23" pageNumber="996" type="Museum">MACN</collectionCode>
),
<specimenCount box="[285,400,1761,1783]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
and
<specimenCount box="[470,609,1761,1783]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[628,714,1761,1782]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="23" pageNumber="996" type="Museum">MACN</collectionCode>
ex Hermann Burmeister collection),
<specimenCount box="[565,671,1791,1813]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
(
<collectionCode box="[688,767,1792,1814]" pageId="23" pageNumber="996">MLPA</collectionCode>
),
<specimenCount box="[163,288,1822,1844]" pageId="23" pageNumber="996" type="male">four males</specimenCount>
and
<specimenCount box="[350,512,1822,1844]" pageId="23" pageNumber="996" type="female">three females</specimenCount>
(
<collectionCode box="[528,618,1822,1844]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="996">MNHN</collectionCode>
),
<specimenCount box="[639,779,1822,1844]" pageId="23" pageNumber="996" type="male">three males</specimenCount>
and
<specimenCount box="[214,352,1853,1875]" pageId="23" pageNumber="996" type="female">two females</specimenCount>
(
<collectionCode box="[368,458,1853,1874]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="23" pageNumber="996" type="Museum">NHMW</collectionCode>
ex W. Zelenka collection),
<specimenCount box="[163,292,1883,1905]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[309,389,1884,1906]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="23" pageNumber="996">NHRS</collectionCode>
),
<specimenCount box="[410,538,1883,1905]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[555,636,1884,1905]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="23" pageNumber="996">NMPC</collectionCode>
ex Dohon collection),
<specimenCount box="[958,1064,197,219]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
(
<collectionCode box="[1079,1165,197,219]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="23" pageNumber="996" type="Museum">RBINS</collectionCode>
),
<specimenCount box="[1184,1289,197,219]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
(
<collectionCode box="[1304,1387,197,218]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="23" pageNumber="996" type="Museum">RBINS</collectionCode>
ex E. Candèze collection),
<specimenCount box="[1111,1242,227,249]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[1259,1345,228,249]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="23" pageNumber="996" type="Museum">RBINS</collectionCode>
ex Le Moult collection) and
<specimenCount box="[1075,1199,258,280]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[1213,1300,258,280]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="23" pageNumber="996" type="Museum">RMNH</collectionCode>
).
</paragraph>
<paragraph blockId="23.[827,1443,321,1907]" lastBlockId="24.[809,1425,197,464]" lastPageId="24" lastPageNumber="997" pageId="23" pageNumber="996">
<emphasis box="[827,1051,321,342]" italics="true" pageId="23" pageNumber="996">Coppery specimens:</emphasis>
<materialsCitation ID-GBIF-Occurrence="3704841356" collectionCode="SMTD" country="Argentina" location="Felsche" pageId="23" pageNumber="996" specimenCount="1" specimenCount-male="1">
<emphasis bold="true" box="[1059,1235,321,343]" pageId="23" pageNumber="996">
<collectingCountry box="[1059,1230,321,343]" name="Argentina" pageId="23" pageNumber="996">ARGENTINA</collectingCountry>
:
</emphasis>
no further data
<specimenCount box="[827,929,351,373]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
(
<collectionCode box="[942,1018,352,373]" pageId="23" pageNumber="996">SMTD</collectionCode>
ex C.
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC17FFEB3A690C54DF85FE86" box="[1105,1191,351,373]" country="Argentina" name="Felsche" pageId="23" pageNumber="996">Felsche</location>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841526" collectionCode="BMNH" collectorName="O. W. Thomas" country="Argentina" pageId="23" pageNumber="996" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
no specific locality or date,
<collectorName box="[1013,1165,382,404]" pageId="23" pageNumber="996">O.W. Thomas</collectorName>
leg.
<specimenCount box="[1240,1344,382,404]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
and
<specimenCount pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[924,1013,413,435]" pageId="23" pageNumber="996">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841371" collectionCode="BMNH" collectorName="Taylor" country="Argentina" pageId="23" pageNumber="996" specimenCount="1" specimenCount-male="1">
no specific locality or date,
<collectorName box="[1366,1442,413,435]" pageId="23" pageNumber="996">Taylor</collectorName>
leg.
<specimenCount box="[893,995,443,465]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
(
<collectionCode box="[1010,1096,444,466]" pageId="23" pageNumber="996">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841388" collectionCode="OUMNH, SMF" collectorName="Hope-Westwood" country="Argentina" location="Albert Hepp" municipality="Escrapus" pageId="23" pageNumber="996" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Ciudad Autonoma de Buenos Aires">
<emphasis box="[1114,1319,444,465]" italics="true" pageId="23" pageNumber="996">
<collectingRegion box="[1114,1313,444,465]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="23" pageNumber="996">BUENOS AIRES</collectingRegion>
:
</emphasis>
no further data
<specimenCount box="[913,1022,474,496]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
and
<specimenCount box="[1086,1217,474,496]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(labelled with the
<emphasis box="[827,1016,505,526]" italics="true" pageId="23" pageNumber="996">nomen in litteris</emphasis>
<emphasis box="[1027,1256,505,526]" italics="true" pageId="23" pageNumber="996">
<collectingMunicipality box="[1027,1133,505,526]" pageId="23" pageNumber="996">Escrapus</collectingMunicipality>
singularis
</emphasis>
) (
<collectionCode box="[1284,1389,505,526]" pageId="23" pageNumber="996">OUMNH</collectionCode>
ex
<collectorName box="[827,1010,535,557]" pageId="23" pageNumber="996">Hope-Westwood</collectorName>
collection) and
<specimenCount box="[1192,1317,535,557]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[1332,1388,536,557]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="23" pageNumber="996">SMF</collectionCode>
ex
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC17FFEB3D030F3DD8EFFDBF" box="[827,973,566,588]" country="Argentina" municipality="Escrapus" name="Albert Hepp" pageId="23" pageNumber="996" stateProvince="Ciudad Autonoma de Buenos Aires">Albert Hepp</location>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841390" collectingDate="1941-02" collectionCode="FSCA" country="Argentina" pageId="23" pageNumber="996" specimenCount="1" specimenCount-male="1" stateProvince="Ciudad Autonoma de Buenos Aires">
no specific locality,
<date box="[1351,1437,566,587]" pageId="23" pageNumber="996" value="1941-02">
<collectingDate box="[1351,1437,566,587]" pageId="23" pageNumber="996" value="1941-02">II.1941</collectingDate>
</date>
, no collector
<specimenCount box="[997,1105,597,619]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
(
<collectionCode box="[1122,1196,597,619]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="23" pageNumber="996" type="Museum">FSCA</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841389" collectingDate="1942-02" collectionCode="FSCA" country="Argentina" pageId="23" pageNumber="996" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Ciudad Autonoma de Buenos Aires">
no specific locality,
<date box="[827,911,628,649]" pageId="23" pageNumber="996" value="1942-02">
<collectingDate box="[827,911,628,649]" pageId="23" pageNumber="996" value="1942-02">II.1942</collectingDate>
</date>
, no collector
<specimenCount box="[1069,1182,627,649]" pageId="23" pageNumber="996" type="male">two males</specimenCount>
and
<specimenCount box="[1233,1355,627,649]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[1366,1436,628,650]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="23" pageNumber="996" type="Museum">FSCA</collectionCode>
)
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3704841419" box="[827,1302,658,680]" collectionCode="MZSP" country="Argentina" pageId="23" pageNumber="996" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Ciudad Autonoma de Buenos Aires">
and
<specimenCount box="[881,1003,658,680]" pageId="23" pageNumber="996" type="male">two males</specimenCount>
and
<specimenCount box="[1068,1198,658,680]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[1216,1294,658,680]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="23" pageNumber="996" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841453" collectingDate="1957-01" collectionCode="MHNG" collectorName="Chassot" country="Argentina" pageId="23" pageNumber="996" specimenCount="1" specimenCount-male="1" stateProvince="Ciudad Autonoma de Buenos Aires">
no specific locality,
<date box="[927,1004,689,711]" pageId="23" pageNumber="996" value="1957-01">
<collectingDate box="[927,1004,689,711]" pageId="23" pageNumber="996" value="1957-01">I.1957</collectingDate>
</date>
,
<collectorName box="[1019,1115,689,711]" pageId="23" pageNumber="996">Chassot</collectorName>
leg.
<specimenCount box="[1196,1324,689,711]" pageId="23" pageNumber="996" type="male">four males</specimenCount>
(
<collectionCode box="[1341,1431,689,711]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="23" pageNumber="996" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841324" collectionCode="TAMU" collectorName="W. D. Edmonds" country="Argentina" location="Bahia Blanca" pageId="23" pageNumber="996" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Ciudad Autonoma de Buenos Aires">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC17FFEB3D030FC4D8FBFD16" box="[827,985,719,741]" country="Argentina" name="Bahia Blanca" pageId="23" pageNumber="996" stateProvince="Ciudad Autonoma de Buenos Aires">Bahía Blanca</location>
, no further data
<specimenCount box="[1205,1323,719,741]" pageId="23" pageNumber="996" type="male">two males</specimenCount>
and
<specimenCount pageId="23" pageNumber="996" type="female">three females</specimenCount>
(
<collectionCode box="[929,1008,750,771]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="23" pageNumber="996" type="Herbarium">TAMU</collectionCode>
ex
<collectorName box="[1066,1239,750,772]" pageId="23" pageNumber="996">W. D. Edmonds</collectorName>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841418" collectingDate="1941-02" collectionCode="CEIOC, MZSP" collectorName="Zikan" country="Argentina" location="Bahia Blanca" pageId="23" pageNumber="996" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Ciudad Autonoma de Buenos Aires">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC17FFEB3B660FE5D8B2FCD0" country="Argentina" name="Bahia Blanca" pageId="23" pageNumber="996" stateProvince="Ciudad Autonoma de Buenos Aires">Bahía Blanca</location>
,
<date box="[925,1013,781,802]" pageId="23" pageNumber="996" value="1941-02">
<collectingDate box="[925,1013,781,802]" pageId="23" pageNumber="996" value="1941-02">II.1941</collectingDate>
</date>
, no collector
<specimenCount box="[1198,1306,781,803]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
(dissected) (
<collectionCode box="[834,918,812,833]" pageId="23" pageNumber="996">CEIOC</collectionCode>
ex J. F.
<collectorName box="[1030,1097,811,833]" pageId="23" pageNumber="996">Zikan</collectorName>
collection) and
<specimenCount box="[1277,1393,811,833]" pageId="23" pageNumber="996" type="male">two males</specimenCount>
and
<specimenCount box="[827,976,842,864]" pageId="23" pageNumber="996" type="female">four females</specimenCount>
(
<collectionCode box="[993,1070,842,864]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="23" pageNumber="996" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841497" collectionCode="CNCI" collectorName="Patagonia' &amp; E. Weiske" country="Argentina" location="Bahia Blanca" pageId="23" pageNumber="996" specimenCount="1" specimenCount-male="1" stateProvince="Ciudad Autonoma de Buenos Aires">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC17FFEB3A7B0E41DFCAFC93" box="[1091,1256,842,864]" country="Argentina" name="Bahia Blanca" pageId="23" pageNumber="996" stateProvince="Ciudad Autonoma de Buenos Aires">Bahía Blanca</location>
,
<collectorName box="[1272,1401,842,864]" pageId="23" pageNumber="996">Patagonia</collectorName>
, no date,
<collectorName box="[894,1012,873,895]" pageId="23" pageNumber="996">E. Weiske</collectorName>
leg.
<specimenCount box="[1094,1202,873,895]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
(
<collectionCode box="[1219,1291,873,895]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="23" pageNumber="996">CNCI</collectionCode>
)
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3704841340" collectionCode="NHMB, RMNH, SDEI, SMF" country="Argentina" pageId="23" pageNumber="996" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Ciudad Autonoma de Buenos Aires">
<specimenCount box="[1312,1442,873,895]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[834,917,904,925]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="23" pageNumber="996">NHMB</collectionCode>
ex
<collectorName box="[978,1031,904,925]" pageId="23" pageNumber="996">Frey</collectorName>
collection),
<specimenCount box="[1168,1292,903,925]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[1306,1389,904,925]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="23" pageNumber="996" type="Museum">RMNH</collectionCode>
ex
<collectorName box="[827,1008,934,956]" pageId="23" pageNumber="996">Valck Lucassen</collectorName>
collection),
<specimenCount box="[1150,1255,934,956]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
and
<specimenCount box="[1315,1442,934,956]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[834,898,965,986]" pageId="23" pageNumber="996">SDEI</collectionCode>
ex
<collectorName box="[962,1152,965,987]" pageId="23" pageNumber="996">Franklin Müller</collectorName>
collection) and
<specimenCount box="[1337,1442,965,987]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
and
<specimenCount box="[880,1008,995,1017]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[1025,1083,996,1017]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="23" pageNumber="996">SMF</collectionCode>
ex
<collectorName box="[1149,1307,995,1017]" pageId="23" pageNumber="996">B. Schwarzer</collectorName>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841528" collectingDate="1926-02-02" collectionCode="NMPC" collectorName="Brida" country="Argentina" location="Pontevedra" municipality="Merlo" pageId="23" pageNumber="996" specimenCount="1" specimenCount-female="1" stateProvince="Ciudad Autonoma de Buenos Aires">
<collectingMunicipality box="[827,896,1026,1048]" pageId="23" pageNumber="996">Merlo</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC17FFEB3DB30909DF2DFBEB" box="[907,1039,1026,1048]" country="Argentina" municipality="Merlo" name="Pontevedra" pageId="23" pageNumber="996" stateProvince="Ciudad Autonoma de Buenos Aires">Pontevedra</location>
,
<date box="[1050,1168,1026,1048]" pageId="23" pageNumber="996" value="1926-02-02">
<collectingDate box="[1050,1168,1026,1048]" pageId="23" pageNumber="996" value="1926-02-02">02.II.1926</collectingDate>
</date>
,
<collectorName box="[1180,1245,1026,1048]" pageId="23" pageNumber="996">Brida</collectorName>
leg.
<specimenCount box="[1319,1443,1026,1048]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[834,916,1057,1079]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="23" pageNumber="996">NMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841406" collectingDate="1940-11" collectionCode="CMNC" country="Argentina" location="Felipe Sola" municipality="Puan" pageId="23" pageNumber="996" specimenCount="1" specimenCount-male="1" stateProvince="Ciudad Autonoma de Buenos Aires">
<collectingMunicipality box="[934,996,1057,1078]" pageId="23" pageNumber="996">Puan</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC17FFEB3DD6092ADF4CFBC4" box="[1006,1134,1057,1079]" country="Argentina" municipality="Puan" name="Felipe Sola" pageId="23" pageNumber="996" stateProvince="Ciudad Autonoma de Buenos Aires">Felipe Solá</location>
,
<date box="[1144,1236,1057,1078]" pageId="23" pageNumber="996" value="1940-11">
<collectingDate box="[1144,1236,1057,1078]" pageId="23" pageNumber="996" value="1940-11">XI.1940</collectingDate>
</date>
, no collector
<specimenCount pageId="23" pageNumber="996" type="male">two males</specimenCount>
(
<collectionCode box="[908,991,1087,1109]" pageId="23" pageNumber="996">CMNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841452" collectingDate="1942-02" collectionCode="CMNC" country="Argentina" location="Felipe Sola" municipality="Puan" pageId="23" pageNumber="996" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Ciudad Autonoma de Buenos Aires">
<collectingMunicipality box="[1008,1070,1088,1109]" pageId="23" pageNumber="996">Puan</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC17FFEB3A010934DF9BFBA6" box="[1081,1209,1087,1109]" country="Argentina" municipality="Puan" name="Felipe Sola" pageId="23" pageNumber="996" stateProvince="Ciudad Autonoma de Buenos Aires">Felipe Solá</location>
,
<date box="[1219,1303,1087,1108]" pageId="23" pageNumber="996" value="1942-02">
<collectingDate box="[1219,1303,1087,1108]" pageId="23" pageNumber="996" value="1942-02">II.1942</collectingDate>
</date>
, no collector
<specimenCount box="[850,960,1118,1140]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
and
<specimenCount box="[1025,1158,1118,1140]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[1175,1260,1118,1140]" pageId="23" pageNumber="996">CMNC</collectionCode>
)
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3704841301" collectionCode="DZUP" country="Argentina" location="Justus Jor" pageId="23" pageNumber="996" specimenCount="1" specimenCount-male="1" stateProvince="Ciudad Autonoma de Buenos Aires">
and
<specimenCount box="[1333,1442,1118,1140]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
(dissected) (
<collectionCode box="[962,1034,1149,1170]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="23" pageNumber="996" type="University or college">DZUP</collectionCode>
ex F.
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC17FFEB3A580976DFFAFB61" box="[1120,1240,1149,1170]" country="Argentina" name="Justus Jor" pageId="23" pageNumber="996" stateProvince="Ciudad Autonoma de Buenos Aires">Justus Jor</location>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841496" collectingDate="1942-02" collectionCode="MZSP" collectorName="B. Pohl" country="Argentina" location="Felipe Sola" municipality="Puan" pageId="23" pageNumber="996" specimenCount="1" specimenCount-male="1" stateProvince="Ciudad Autonoma de Buenos Aires">
<collectingMunicipality box="[1376,1438,1149,1170]" pageId="23" pageNumber="996">Puan</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC17FFEB3D030990D8E2FB42" box="[827,960,1179,1201]" country="Argentina" municipality="Puan" name="Felipe Sola" pageId="23" pageNumber="996" stateProvince="Ciudad Autonoma de Buenos Aires">Felipe Solá</location>
,
<date box="[972,1058,1179,1200]" pageId="23" pageNumber="996" value="1942-02">
<collectingDate box="[972,1058,1179,1200]" pageId="23" pageNumber="996" value="1942-02">II.1942</collectingDate>
</date>
,
<collectorName box="[1071,1155,1179,1201]" pageId="23" pageNumber="996">B. Pohl</collectorName>
leg.
<specimenCount box="[1233,1339,1179,1201]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
(
<collectionCode box="[1355,1431,1179,1201]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="23" pageNumber="996" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841487" collectingDate="1942-10" collectionCode="MZSP" collectorName="B. Pohl" country="Argentina" location="Felipe Sola" municipality="Puan" pageId="23" pageNumber="996" specimenCount="1" specimenCount-female="1" stateProvince="Ciudad Autonoma de Buenos Aires">
<collectingMunicipality box="[827,890,1210,1231]" pageId="23" pageNumber="996">Puan</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC17FFEB3DBE09B1DF28FB23" box="[902,1034,1210,1232]" country="Argentina" municipality="Puan" name="Felipe Sola" pageId="23" pageNumber="996" stateProvince="Ciudad Autonoma de Buenos Aires">Felipe Solá</location>
,
<date box="[1047,1130,1210,1231]" pageId="23" pageNumber="996" value="1942-10">
<collectingDate box="[1047,1130,1210,1231]" pageId="23" pageNumber="996" value="1942-10">X.1942</collectingDate>
</date>
,
<collectorName box="[1142,1225,1210,1232]" pageId="23" pageNumber="996">B. Pohl</collectorName>
leg.
<specimenCount box="[1303,1443,1210,1232]" pageId="23" pageNumber="996" type="female">two females</specimenCount>
(
<collectionCode box="[835,913,1241,1263]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="23" pageNumber="996" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841414" collectingDate="1943-12" collectionCode="MNHN" country="Argentina" county="Puan" location="Estacion Felipe Sola'" municipality="Felipe Sola" pageId="23" pageNumber="996" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Ciudad Autonoma de Buenos Aires">
<collectingCounty box="[934,999,1241,1262]" pageId="23" pageNumber="996">Puan</collectingCounty>
,
<collectingMunicipality box="[1013,1148,1241,1263]" pageId="23" pageNumber="996">Felipe Solá</collectingMunicipality>
(
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC17FFEB3AAA09D2DEB4FB1C" box="[1170,1430,1241,1263]" country="Argentina" county="Puan" municipality="Felipe Sola" name="Estacion Felipe Sola'" pageId="23" pageNumber="996" stateProvince="Ciudad Autonoma de Buenos Aires">Estación Felipe Sola</location>
),
<date box="[827,935,1271,1292]" pageId="23" pageNumber="996" value="1943-12">
<collectingDate box="[827,935,1271,1292]" pageId="23" pageNumber="996" value="1943-12">XII.1943</collectingDate>
</date>
, no collector
<specimenCount box="[1123,1246,1271,1293]" pageId="23" pageNumber="996" type="male">two males</specimenCount>
and
<specimenCount box="[1311,1442,1271,1293]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[834,921,1302,1323]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="996">MNHN</collectionCode>
ex
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC17FFEB3DE5081DDF15FADF" box="[989,1079,1302,1324]" country="Argentina" county="Puan" municipality="Felipe Sola" name="Coiffait" pageId="23" pageNumber="996" stateProvince="Ciudad Autonoma de Buenos Aires">Coiffait</location>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841468" collectingDate="1944-02" collectionCode="NHMB" country="Argentina" county="Puan" location="Frey" municipality="Felipe Sola" pageId="23" pageNumber="996" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Ciudad Autonoma de Buenos Aires">
<collectingCounty box="[1225,1289,1302,1323]" pageId="23" pageNumber="996">Puan</collectingCounty>
,
<collectingMunicipality box="[1303,1438,1302,1324]" pageId="23" pageNumber="996">Felipe Solá</collectingMunicipality>
,
<date box="[827,911,1333,1354]" pageId="23" pageNumber="996" value="1944-02">
<collectingDate box="[827,911,1333,1354]" pageId="23" pageNumber="996" value="1944-02">II.1944</collectingDate>
</date>
, no collector
<specimenCount box="[1072,1173,1333,1355]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
and
<specimenCount box="[1226,1348,1333,1355]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[1360,1443,1333,1354]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="23" pageNumber="996">NHMB</collectionCode>
ex
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC17FFEB3D4D085FD88EFA9A" box="[885,940,1364,1385]" country="Argentina" county="Puan" municipality="Felipe Sola" name="Frey" pageId="23" pageNumber="996" stateProvince="Ciudad Autonoma de Buenos Aires">Frey</location>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841518" collectingDate="1950-02" collectionCode="CMNC" country="Argentina" location="Felipe Sola" municipality="Puan" pageId="23" pageNumber="996" specimenCount="1" specimenCount-female="1" stateProvince="Ciudad Autonoma de Buenos Aires">
<collectingMunicipality box="[1086,1150,1364,1385]" pageId="23" pageNumber="996">Puan</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC17FFEB3AB40858DE36FA9A" box="[1164,1300,1363,1385]" country="Argentina" municipality="Puan" name="Felipe Sola" pageId="23" pageNumber="996" stateProvince="Ciudad Autonoma de Buenos Aires">Felipe Solá</location>
,
<date box="[1313,1401,1363,1385]" pageId="23" pageNumber="996" value="1950-02">
<collectingDate box="[1313,1401,1363,1385]" pageId="23" pageNumber="996" value="1950-02">II.1950</collectingDate>
</date>
, no collector
<specimenCount box="[957,1086,1394,1416]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[1102,1184,1394,1415]" pageId="23" pageNumber="996">CMNC</collectionCode>
ex
<collectorName box="[1249,1390,1394,1416]" pageId="23" pageNumber="996">A. Martínez</collectorName>
and
<collectorName box="[827,900,1425,1446]" pageId="23" pageNumber="996">Henry</collectorName>
&amp;
<collectorName box="[931,1092,1425,1447]" pageId="23" pageNumber="996">Anne Howden</collectorName>
collections)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841313" collectingDate="1959-11" collectionCode="MZSP" collectorName="A. Martinez" country="Argentina" location="Felipe Sola" municipality="Puan" pageId="23" pageNumber="996" specimenCount="1" specimenCount-female="1" stateProvince="Ciudad Autonoma de Buenos Aires">
<collectingMunicipality box="[1238,1300,1425,1446]" pageId="23" pageNumber="996">Puan</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC17FFEB3B27089ADEBCFA54" box="[1311,1438,1425,1447]" country="Argentina" municipality="Puan" name="Felipe Sola" pageId="23" pageNumber="996" stateProvince="Ciudad Autonoma de Buenos Aires">Felipe Solá</location>
,
<date box="[827,919,1455,1477]" pageId="23" pageNumber="996" value="1959-11">
<collectingDate box="[827,919,1455,1477]" pageId="23" pageNumber="996" value="1959-11">XI.1959</collectingDate>
</date>
,
<collectorName box="[930,1067,1455,1477]" pageId="23" pageNumber="996">A. Martínez</collectorName>
leg.
<specimenCount box="[1142,1267,1455,1477]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[1282,1357,1455,1477]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="23" pageNumber="996" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841322" collectingDate="1961-02" collectionCode="MZSP" collectorName="A. Martinez" country="Argentina" location="Felipe Sola" municipality="Puan" pageId="23" pageNumber="996" specimenCount="1" specimenCount-female="1" stateProvince="Ciudad Autonoma de Buenos Aires">
<collectingMunicipality box="[1376,1438,1456,1477]" pageId="23" pageNumber="996">Puan</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC17FFEB3D0308C5D8E6FA17" box="[827,964,1486,1508]" country="Argentina" municipality="Puan" name="Felipe Sola" pageId="23" pageNumber="996" stateProvince="Ciudad Autonoma de Buenos Aires">Felipe Solá</location>
,
<date box="[979,1068,1486,1508]" pageId="23" pageNumber="996" value="1961-02">
<collectingDate box="[979,1068,1486,1508]" pageId="23" pageNumber="996" value="1961-02">II.1961</collectingDate>
</date>
,
<collectorName box="[1082,1227,1486,1508]" pageId="23" pageNumber="996">A. Martínez</collectorName>
leg.
<specimenCount box="[1311,1442,1486,1508]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[834,909,1517,1539]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="23" pageNumber="996" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841308" collectingDate="1961-12" collectionCode="MZSP" collectorName="A. Martinez" country="Argentina" location="Felipe Sola" municipality="Puan" pageId="23" pageNumber="996" specimenCount="1" specimenCount-male="1" stateProvince="Ciudad Autonoma de Buenos Aires">
<collectingMunicipality box="[928,991,1517,1538]" pageId="23" pageNumber="996">Puan</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC17FFEB3DD308E6DF4FF9F0" box="[1003,1133,1517,1539]" country="Argentina" municipality="Puan" name="Felipe Sola" pageId="23" pageNumber="996" stateProvince="Ciudad Autonoma de Buenos Aires">Felipe Solá</location>
,
<date box="[1145,1247,1517,1539]" pageId="23" pageNumber="996" value="1961-12">
<collectingDate box="[1145,1247,1517,1539]" pageId="23" pageNumber="996" value="1961-12">XII.1961</collectingDate>
</date>
,
<collectorName box="[1258,1396,1517,1539]" pageId="23" pageNumber="996">A. Martínez</collectorName>
leg.
<specimenCount box="[847,950,1547,1569]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
(
<collectionCode box="[965,1039,1547,1569]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="23" pageNumber="996" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841553" collectingDate="1888-05" collectionCode="SMF" collectorName="F. Ohaus" country="Argentina" location="San Nicolas" pageId="23" pageNumber="996" specimenCount="1" specimenCount-male="1" stateProvince="Ciudad Autonoma de Buenos Aires">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC17FFEB3A1A0B00DF89F9D2" box="[1058,1195,1547,1569]" country="Argentina" name="San Nicolas" pageId="23" pageNumber="996" stateProvince="Ciudad Autonoma de Buenos Aires">San Nicolás</location>
,
<date box="[1205,1284,1547,1568]" pageId="23" pageNumber="996" value="1888-05">
<collectingDate box="[1205,1284,1547,1568]" pageId="23" pageNumber="996" value="1888-05">V.1888</collectingDate>
</date>
,
<collectorName box="[1295,1397,1547,1569]" pageId="23" pageNumber="996">F. Ohaus</collectorName>
leg.
<specimenCount box="[848,952,1578,1600]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
(dissected) (
<collectionCode box="[1098,1158,1578,1600]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="23" pageNumber="996">SMF</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841570" collectingDate="1942-01" collectionCode="DZUP" collectorName="F. Justus Jor" country="Argentina" location="CIUDAD AUTONOMA DE BUENOS AIRES" pageId="23" pageNumber="996" specimenCount="1" specimenCount-female="1" stateProvince="Ciudad Autonoma de Buenos Aires">
<emphasis italics="true" pageId="23" pageNumber="996">
<collectingRegion country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="23" pageNumber="996">CIUDAD AUTÓNOMA DE BUENOS AIRES</collectingRegion>
:
</emphasis>
no specific locality,
<date box="[1357,1437,1609,1630]" pageId="23" pageNumber="996" value="1942-01">
<collectingDate box="[1357,1437,1609,1630]" pageId="23" pageNumber="996" value="1942-01">I.1942</collectingDate>
</date>
, no collector
<specimenCount box="[997,1128,1639,1661]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[1144,1219,1639,1660]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="23" pageNumber="996" type="University or college">DZUP</collectionCode>
ex
<collectorName box="[1287,1442,1640,1661]" pageId="23" pageNumber="996">F. Justus Jor</collectorName>
collection)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841400" collectingDate="1938-12" collectionCode="MNRJ" collectorName="M. Viana" country="Argentina" location="El Jouer'" pageId="23" pageNumber="996" specimenCount="1" specimenCount-female="1" stateProvince="Ciudad Autonoma de Buenos Aires">
<emphasis box="[978,1126,1668,1692]" italics="true" pageId="23" pageNumber="996">CÓRDOBA:</emphasis>
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC17FFEB3A4E0B8DDFCFF968" box="[1142,1261,1670,1692]" country="Argentina" name="El Jouer'" pageId="23" pageNumber="996" stateProvince="Ciudad Autonoma de Buenos Aires">El Jouer</location>
(?),
<date box="[1325,1436,1670,1691]" pageId="23" pageNumber="996" value="1938-12">
<collectingDate box="[1325,1436,1670,1691]" pageId="23" pageNumber="996" value="1938-12">XII.1938</collectingDate>
</date>
,
<collectorName box="[827,932,1701,1723]" pageId="23" pageNumber="996">M. Viana</collectorName>
leg.
<specimenCount box="[1008,1134,1701,1723]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[1149,1228,1701,1723]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="23" pageNumber="996">MNRJ</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841366" collectingDate="1965-12" collectionCode="CMNC" collectorName="Cichero" country="Argentina" location="Quilino" municipality="Ischilin" pageId="23" pageNumber="996" specimenCount="1" specimenCount-female="1" stateProvince="Ciudad Autonoma de Buenos Aires">
<collectingMunicipality box="[1248,1338,1701,1723]" pageId="23" pageNumber="996">Ischilín</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC17FFEB3B7E0BAEDEBCF948" box="[1350,1438,1701,1723]" country="Argentina" municipality="Ischilin" name="Quilino" pageId="23" pageNumber="996" stateProvince="Ciudad Autonoma de Buenos Aires">Quilino</location>
,
<date box="[827,929,1731,1753]" pageId="23" pageNumber="996" value="1965-12">
<collectingDate box="[827,929,1731,1753]" pageId="23" pageNumber="996" value="1965-12">XII.1965</collectingDate>
</date>
,
<collectorName box="[942,1031,1731,1753]" pageId="23" pageNumber="996">Cichero</collectorName>
leg.
<specimenCount box="[1107,1233,1731,1753]" pageId="23" pageNumber="996" type="female">one female</specimenCount>
(
<collectionCode box="[1248,1332,1731,1753]" pageId="23" pageNumber="996">CMNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841547" collectingDate="1976-11-18" collectionCode="CMNC" collectorName="G. J. Williner" country="Argentina" location="San Esteban" municipality="Punilla" pageId="23" pageNumber="996" specimenCount="1" specimenCount-male="1" stateProvince="Ciudad Autonoma de Buenos Aires">
<collectingMunicipality box="[1351,1438,1731,1753]" pageId="23" pageNumber="996">Punilla</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC17FFEB3D030BE9D8FBF90B" box="[827,985,1762,1784]" country="Argentina" municipality="Punilla" name="San Esteban" pageId="23" pageNumber="996" stateProvince="Ciudad Autonoma de Buenos Aires">San Esteban</location>
,
<date box="[1000,1136,1762,1784]" pageId="23" pageNumber="996" value="1976-11-18">
<collectingDate box="[1000,1136,1762,1784]" pageId="23" pageNumber="996" value="1976-11-18">18.XI.1976</collectingDate>
</date>
,
<collectorName box="[1150,1314,1762,1784]" pageId="23" pageNumber="996">G. J. Williner</collectorName>
leg.
<specimenCount pageId="23" pageNumber="996" type="male">one male</specimenCount>
(
<collectionCode box="[899,984,1793,1815]" pageId="23" pageNumber="996">CMNC</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841328" collectionCode="BMNH" country="Argentina" location="Reiche'" municipality="Aeschropus" pageId="23" pageNumber="996" specimenCount="1" specimenCount-male="1" stateProvince="Corrientes">
<emphasis box="[1002,1185,1793,1814]" italics="true" pageId="23" pageNumber="996">
<collectingRegion box="[1002,1178,1793,1814]" country="Argentina" name="Corrientes" pageId="23" pageNumber="996">CORRIENTES</collectingRegion>
:
</emphasis>
no further data
<specimenCount pageId="23" pageNumber="996" type="male">one male</specimenCount>
(labelled with the
<emphasis box="[1106,1299,1823,1845]" italics="true" pageId="23" pageNumber="996">nomen in litteris</emphasis>
<emphasis italics="true" pageId="23" pageNumber="996">
<collectingMunicipality box="[1311,1442,1823,1844]" pageId="23" pageNumber="996">Aeschropus</collectingMunicipality>
singularis
</emphasis>
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC17FFEB3D8E0A35DF2BF8A7" box="[950,1033,1854,1876]" country="Argentina" municipality="Aeschropus" name="Reiche'" pageId="23" pageNumber="996" stateProvince="Corrientes">Reiche</location>
) (
<collectionCode box="[1056,1142,1854,1876]" pageId="23" pageNumber="996">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841554" collectingDate="1988-12" collectionCode="BMNH" collectorName="S. Bolle &amp; J. Zidek" country="Uruguay" lastPageId="24" lastPageNumber="997" location="Pronunciamiento" pageId="23" pageNumber="996" specimenCount="1" specimenCount-male="1" stateProvince="Entre Rios">
<emphasis box="[1160,1330,1852,1876]" italics="true" pageId="23" pageNumber="996">
<collectingRegion box="[1160,1324,1852,1876]" country="Argentina" name="Entre Rios" pageId="23" pageNumber="996">ENTRE RÍOS</collectingRegion>
:
</emphasis>
<collectingCountry box="[1338,1439,1854,1875]" name="Uruguay" pageId="23" pageNumber="996">Uruguay</collectingCountry>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC17FFEB3D030A56DF28F880" box="[827,1034,1885,1907]" country="Uruguay" name="Pronunciamiento" pageId="23" pageNumber="996" stateProvince="Entre Rios">Pronunciamiento</location>
,
<date box="[1046,1152,1885,1906]" pageId="23" pageNumber="996" value="1988-12">
<collectingDate box="[1046,1152,1885,1906]" pageId="23" pageNumber="996" value="1988-12">XII.1988</collectingDate>
</date>
,
<collectorName box="[1165,1255,1885,1907]" pageId="23" pageNumber="996">S. Bolle</collectorName>
leg.
<specimenCount box="[1336,1442,1885,1907]" pageId="23" pageNumber="996" type="male">one male</specimenCount>
(dissected) (
<collectionCode box="[281,364,197,218]" pageId="24" pageNumber="997">BMNH</collectionCode>
ex
<collectorName box="[426,515,197,219]" pageId="24" pageNumber="997">J. Zídek</collectorName>
leg.)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841359" collectionCode="NMPC" country="Uruguay" location="Rolle" pageId="24" pageNumber="997" specimenCount="1" specimenCount-male="1" stateProvince="Entre Rios">
<emphasis box="[585,726,197,218]" italics="true" pageId="24" pageNumber="997">MENDOZA:</emphasis>
no further data
<specimenCount box="[305,404,228,250]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(
<collectionCode box="[416,495,228,249]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="24" pageNumber="997">NMPC</collectionCode>
ex H.
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43C780DEFD959FF09" box="[576,635,228,250]" country="Uruguay" name="Rolle" pageId="24" pageNumber="997" stateProvince="Entre Rios">Rolle</location>
collection)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841316" collectingDate="1995-03" collectionCode="FSCA" country="Uruguay" location="Eldorado" pageId="24" pageNumber="997" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Entre Rios">
<emphasis box="[145,293,259,280]" italics="true" pageId="24" pageNumber="997">MISIONES:</emphasis>
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43F170C09DABDFEEB" box="[303,415,258,280]" country="Uruguay" name="Eldorado" pageId="24" pageNumber="997" stateProvince="Entre Rios">Eldorado</location>
,
<date box="[429,530,258,280]" pageId="24" pageNumber="997" value="1995-03">
<collectingDate box="[429,530,258,280]" pageId="24" pageNumber="997" value="1995-03">III.1995</collectingDate>
</date>
, no collector
<specimenCount pageId="24" pageNumber="997" type="male">one male</specimenCount>
and
<specimenCount box="[264,392,289,311]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[409,483,289,311]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="24" pageNumber="997" type="Museum">FSCA</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841436" collectingDate="2017-02-07" collectionCode="CEMT" collectorName="G. Arriagada" country="Uruguay" elevation="622" latitude="-33.039665" location="Fundo El Molle'" longLatPrecision="1" longitude="-66.51322" municipality="Belgrano" pageId="24" pageNumber="997" specimenCount="1" specimenCount-male="1" stateProvince="Entre Rios">
<emphasis box="[503,638,289,310]" italics="true" pageId="24" pageNumber="997">SAN LUIS:</emphasis>
<collectingMunicipality box="[648,756,289,311]" pageId="24" pageNumber="997">Belgrano</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43EAD0C4BDA77FEA5" box="[149,341,320,342]" country="Uruguay" latitude="-33.039665" longLatPrecision="1" longitude="-66.51322" municipality="Belgrano" name="Fundo El Molle'" pageId="24" pageNumber="997" stateProvince="Entre Rios">Fundo El Molle</location>
,
<geoCoordinate box="[353,502,320,341]" degrees="33" direction="south" minutes="02" orientation="latitude" pageId="24" pageNumber="997" precision="1" seconds="22.8" value="-33.039665">33°0222.8”S</geoCoordinate>
,
<geoCoordinate box="[515,670,320,342]" degrees="66" direction="west" minutes="30" orientation="longitude" pageId="24" pageNumber="997" precision="1" seconds="47.6" value="-66.51322">66°3047.6”W</geoCoordinate>
,
<quantity box="[683,757,320,342]" metricMagnitude="2" metricUnit="m" metricValue="6.22" pageId="24" pageNumber="997" unit="m" value="622.0">
<elevation box="[683,757,320,342]" metricMagnitude="2" metricUnit="m" metricValue="6.22" pageId="24" pageNumber="997" unit="m" value="622.0">622 m</elevation>
</quantity>
,
<date box="[145,270,351,372]" pageId="24" pageNumber="997" value="2017-02-07">
<collectingDate box="[145,270,351,372]" pageId="24" pageNumber="997" value="2017-02-07">07.II.2017</collectingDate>
</date>
(trap with chicken meat),
<collectorName box="[603,760,350,372]" pageId="24" pageNumber="997">G. Arriagada</collectorName>
leg.
<specimenCount box="[224,338,381,403]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(dissected) (
<collectionCode box="[503,589,381,403]" pageId="24" pageNumber="997">CEMT</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841374" collectingDate="1980-11-19" collectingDateMax="1980-11-21" collectingDateMin="1980-11-19" collectionCode="FSCA" collectorName="R. Woodruff &amp; H. Cordo" country="Uruguay" location="Arroyo Colastine" pageId="24" pageNumber="997" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Entre Rios">
<emphasis box="[612,760,381,403]" italics="true" pageId="24" pageNumber="997">SANTA FE:</emphasis>
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43EA90C97DA7AFE41" box="[145,344,412,434]" country="Uruguay" name="Arroyo Colastine" pageId="24" pageNumber="997" stateProvince="Entre Rios">Arroyo Colastine</location>
,
<locationDeviation box="[359,575,412,434]" location="Santa Fe'" pageId="24" pageNumber="997">
<quantity box="[359,432,412,434]" metricMagnitude="4" metricUnit="m" metricValue="5.3" pageId="24" pageNumber="997" unit="km" value="53.0">53 km</quantity>
S Santa Fe
</locationDeviation>
,
<date box="[587,756,412,433]" pageId="24" pageNumber="997" value="1980-11-19" valueMax="1980-11-21" valueMin="1980-11-19">
<collectingDate box="[587,756,412,433]" pageId="24" pageNumber="997" value="1980-11-19" valueMax="1980-11-21" valueMin="1980-11-19">1921.XI.1980</collectingDate>
</date>
,
<collectorName box="[145,285,442,464]" pageId="24" pageNumber="997">R. Woodruff</collectorName>
and
<collectorName box="[348,454,442,464]" pageId="24" pageNumber="997">H. Cordo</collectorName>
leg.
<specimenCount box="[535,657,442,464]" pageId="24" pageNumber="997" type="male">two males</specimenCount>
and
<specimenCount pageId="24" pageNumber="997" type="female">two females</specimenCount>
(
<collectionCode box="[251,324,473,495]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="24" pageNumber="997" type="Museum">FSCA</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841334" collectingDate="1888-05" collectionCode="NMPC, SMF" collectorName="F. Ohaus" country="Uruguay" location="Rosario" pageId="24" pageNumber="997" specimenCount="1" specimenCount-male="1" stateProvince="Entre Rios">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43F6F0CD2DA90FE1C" box="[343,434,473,495]" country="Uruguay" name="Rosario" pageId="24" pageNumber="997" stateProvince="Entre Rios">Rosario</location>
,
<date box="[445,526,473,494]" pageId="24" pageNumber="997" value="1888-05">
<collectingDate box="[445,526,473,494]" pageId="24" pageNumber="997" value="1888-05">V.1888</collectingDate>
</date>
,
<collectorName box="[538,642,473,495]" pageId="24" pageNumber="997">F. Ohaus</collectorName>
leg.
<specimenCount pageId="24" pageNumber="997" type="male">one male</specimenCount>
(
<collectionCode box="[223,306,504,525]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="24" pageNumber="997">NMPC</collectionCode>
ex
<collectionCode box="[381,443,504,526]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="24" pageNumber="997">SMF</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841445" collectionCode="NHMB" collectorName="Hersilia &amp; La Hersilia'" country="Saint Kitts and Nevis" location="Frey" municipality="Hubrich" pageId="24" pageNumber="997" specimenCount="1" specimenCount-female="1">
<collectingCountry box="[467,644,504,526]" name="Saint Kitts and Nevis" pageId="24" pageNumber="997">San Cristóbal</collectingCountry>
,
<collectorName box="[659,759,504,526]" pageId="24" pageNumber="997">Hersilia</collectorName>
(
<collectorName box="[157,301,534,556]" pageId="24" pageNumber="997">La Hersilia</collectorName>
), no further data
<specimenCount box="[534,660,534,556]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[677,761,535,556]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="24" pageNumber="997">NHMB</collectionCode>
ex
<collectingMunicipality box="[202,299,565,587]" pageId="24" pageNumber="997">Hubrich</collectingMunicipality>
and
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43F500F3DDABCFDB8" box="[360,414,566,587]" country="Saint Kitts and Nevis" municipality="Hubrich" name="Frey" pageId="24" pageNumber="997">Frey</location>
collection)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841563" collectionCode="RBINS" collectorName="Le Moult" country="Saint Kitts and Nevis" location="Salado River" municipality="Chaco de Santiago del Estero'" pageId="24" pageNumber="997" specimenCount="1" specimenCount-male="1">
<emphasis italics="true" pageId="24" pageNumber="997">SANTIAGO DEL ESTERO:</emphasis>
<collectingMunicipality box="[279,662,596,618]" pageId="24" pageNumber="997">Chaco de Santiago del Estero</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43C9E0F5FDBF1FD7B" country="Saint Kitts and Nevis" municipality="Chaco de Santiago del Estero'" name="Salado River" pageId="24" pageNumber="997">Salado River</location>
, no further data
<specimenCount box="[448,557,626,648]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(
<collectionCode box="[575,661,627,648]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="24" pageNumber="997" type="Museum">RBINS</collectionCode>
ex
<collectorName pageId="24" pageNumber="997">Le Moult</collectorName>
collection)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841464" collectingDate="1945-01" collectionCode="MGPA" country="Brazil" location="Guarai" pageId="24" pageNumber="997" specimenCount="1" specimenCount-female="1" stateProvince="Rio Grande do Sul">
<emphasis bold="true" box="[355,467,657,678]" pageId="24" pageNumber="997">
<collectingCountry box="[355,462,657,678]" name="Brazil" pageId="24" pageNumber="997">BRAZIL</collectingCountry>
:
</emphasis>
<emphasis box="[475,760,657,678]" italics="true" pageId="24" pageNumber="997">
<collectingRegion box="[475,754,657,678]" country="Brazil" name="Rio Grande do Sul" pageId="24" pageNumber="997">RIO GRANDE DO SUL</collectingRegion>
:
</emphasis>
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43EA90FBBDBCAFD35" box="[145,232,688,710]" country="Brazil" name="Guarai" pageId="24" pageNumber="997" stateProvince="Rio Grande do Sul">Guaraí</location>
,
<date box="[246,324,688,710]" pageId="24" pageNumber="997" value="1945-01">
<collectingDate box="[246,324,688,710]" pageId="24" pageNumber="997" value="1945-01">I.1945</collectingDate>
</date>
, no collector
<specimenCount box="[510,656,688,710]" pageId="24" pageNumber="997" type="female">two females</specimenCount>
(
<collectionCode box="[672,753,688,710]" pageId="24" pageNumber="997">MGPA</collectionCode>
)
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3704841505" box="[145,449,718,741]" collectionCode="MZSP" country="Brazil" pageId="24" pageNumber="997" specimenCount="1" specimenCount-female="1" stateProvince="Rio Grande do Sul">
and
<specimenCount box="[202,340,718,740]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[360,441,719,741]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="24" pageNumber="997" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841403" collectionCode="MFNB" country="Uruguay" location="URUGUAY" pageId="24" pageNumber="997" specimenCount="1" specimenCount-female="1">
<emphasis bold="true" box="[465,618,718,740]" pageId="24" pageNumber="997">
<collectingCountry box="[465,613,718,740]" name="Uruguay" pageId="24" pageNumber="997">URUGUAY</collectingCountry>
:
</emphasis>
no further data
<specimenCount box="[233,365,749,771]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[383,468,749,771]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="24" pageNumber="997">MFNB</collectionCode>
)
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3704841348" box="[491,705,749,771]" collectionCode="MZSP" country="Uruguay" isTailEnumeration="true" location="URUGUAY" pageId="24" pageNumber="997" specimenCount="1" specimenCount-male="1">
<specimenCount box="[491,601,749,771]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(
<collectionCode box="[619,697,749,771]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="24" pageNumber="997" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3704841484" collectionCode="NMPC" country="Uruguay" pageId="24" pageNumber="997" specimenCount="1" specimenCount-female="1">
<specimenCount pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[249,338,780,802]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="24" pageNumber="997">NMPC</collectionCode>
)
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3704841451" collectionCode="OUMNH, RBINS" collectorName="Westwood &amp; Le Moult" country="Uruguay" pageId="24" pageNumber="997" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount box="[362,478,780,802]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(
<collectionCode box="[498,609,780,801]" pageId="24" pageNumber="997">OUMNH</collectionCode>
ex
<collectorName box="[145,267,810,832]" pageId="24" pageNumber="997">Hope-Westwood</collectorName>
collection),
<specimenCount box="[420,554,810,832]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[572,659,811,832]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="24" pageNumber="997" type="Museum">RBINS</collectionCode>
ex
<collectorName pageId="24" pageNumber="997">Le Moult</collectorName>
collection) and
<specimenCount box="[401,507,841,863]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(dissected) (
<collectionCode box="[655,739,841,862]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="24" pageNumber="997" type="Museum">RBINS</collectionCode>
ex
<collectorName box="[177,282,872,894]" pageId="24" pageNumber="997">Le Moult</collectorName>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841434" collectionCode="ZMFK" collectorName="Lehr" country="Uruguay" location="Fuhlrott" pageId="24" pageNumber="997" specimenCount="1" specimenCount-female="1">
no specific locality, 1885,
<collectorName box="[706,761,872,894]" pageId="24" pageNumber="997">Lehr</collectorName>
leg.
<specimenCount box="[220,352,902,924]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[369,447,902,923]" pageId="24" pageNumber="997">ZMFK</collectionCode>
ex
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43C3D0E8DD94EFC6F" box="[517,620,902,924]" country="Uruguay" name="Fuhlrott" pageId="24" pageNumber="997">Fuhlrott</location>
collection)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841541" collectingDate="1954-11-18" collectionCode="FSCA" collectorName="C. B. Carbonell &amp; E. N. Kellesvig-Waering" country="Uruguay" location="Arroyo de la Tavernada" pageId="24" pageNumber="997" specimenCount="1" specimenCount-female="1" stateProvince="Artigas">
<emphasis box="[145,273,933,954]" italics="true" pageId="24" pageNumber="997">
<collectingRegion box="[145,267,933,954]" country="Uruguay" name="Artigas" pageId="24" pageNumber="997">ARTIGAS</collectingRegion>
:
</emphasis>
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43F210EAED91AFC48" box="[281,568,933,955]" country="Uruguay" name="Arroyo de la Tavernada" pageId="24" pageNumber="997" stateProvince="Artigas">Arroyo de la Tavernada</location>
(?),
<date box="[623,755,933,955]" pageId="24" pageNumber="997" value="1954-11-18">
<collectingDate box="[623,755,933,955]" pageId="24" pageNumber="997" value="1954-11-18">18.XI.1954</collectingDate>
</date>
,
<collectorName box="[145,329,964,986]" pageId="24" pageNumber="997">C. B. Carbonell</collectorName>
leg.
<specimenCount box="[413,544,964,986]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[561,632,964,985]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="24" pageNumber="997" type="Museum">FSCA</collectionCode>
ex
<collectorName pageId="24" pageNumber="997">E. N. Kellesvig-Waering</collectorName>
collection)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841305" collectionCode="MNHN, R, RBINS" country="Uruguay" location="Edgar" pageId="24" pageNumber="997" specimenCount="2" specimenCount-male="2" stateProvince="Montevideo" typeStatus="lectotype">
<emphasis box="[524,721,995,1016]" italics="true" pageId="24" pageNumber="997">
<collectingRegion box="[524,716,995,1016]" country="Uruguay" name="Montevideo" pageId="24" pageNumber="997">MONTEVIDEO</collectingRegion>
:
</emphasis>
no further data
<specimenCount box="[307,407,1025,1047]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(
<typeStatus box="[419,523,1025,1047]" pageId="24" pageNumber="997">lectotype</typeStatus>
of
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[553,753,1025,1046]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="997" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[553,753,1025,1046]" italics="true" pageId="24" pageNumber="997">Bolbites onitoides</emphasis>
</taxonomicName>
) (
<collectionCode box="[152,242,1056,1077]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="24" pageNumber="997">MNHN</collectionCode>
ex
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43F04092BDAA5FBC5" box="[316,391,1056,1078]" country="Uruguay" name="Edgar" pageId="24" pageNumber="997" stateProvince="Montevideo">Edgar</location>
von
<collectorName box="[456,542,1056,1078]" pageId="24" pageNumber="997">Harold</collectorName>
and
<collectionCode box="[609,631,1056,1077]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="24" pageNumber="997">R</collectionCode>
.
<collectorName box="[645,759,1056,1078]" pageId="24" pageNumber="997">Oberthür</collectorName>
collections) and
<specimenCount box="[333,438,1086,1108]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(
<collectionCode box="[453,536,1086,1107]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="24" pageNumber="997" type="Museum">RBINS</collectionCode>
ex
<collectorName box="[599,760,1086,1108]" pageId="24" pageNumber="997">P. de Moffarts</collectorName>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841420" collectionCode="BMNH" country="Uruguay" location="Nevinson" pageId="24" pageNumber="997" specimenCount="1" specimenCount-female="1" stateProvince="Montevideo" typeStatus="lectotype">
no specific locality, 1901, no collector
<specimenCount pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[239,323,1148,1169]" pageId="24" pageNumber="997">BMNH</collectionCode>
ex
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43FBB0977DAD3FB61" box="[387,497,1148,1170]" country="Uruguay" name="Nevinson" pageId="24" pageNumber="997" stateProvince="Montevideo">Nevinson</location>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841330" collectingDate="1907-12-25" collectionCode="RBINS" collectorName="Gillet" country="Uruguay" pageId="24" pageNumber="997" specimenCount="1" specimenCount-male="1" stateProvince="Montevideo" typeStatus="lectotype">
no specific locality,
<date box="[242,380,1178,1200]" pageId="24" pageNumber="997" value="1907-12-25">
<collectingDate box="[242,380,1178,1200]" pageId="24" pageNumber="997" value="1907-12-25">25.XII.1907</collectingDate>
</date>
, no collector
<specimenCount box="[557,662,1178,1200]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(
<collectionCode box="[677,760,1178,1199]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="24" pageNumber="997" type="Museum">RBINS</collectionCode>
ex J. J.
<collectorName box="[250,315,1209,1231]" pageId="24" pageNumber="997">Gillet</collectorName>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841555" collectingDate="1952-10" collectionCode="DZUP" collectorName="Alvarenga" country="Uruguay" pageId="24" pageNumber="997" specimenCount="1" specimenCount-female="1" stateProvince="Montevideo" typeStatus="lectotype">
no specific locality,
<date box="[675,756,1209,1231]" pageId="24" pageNumber="997" value="1952-10">
<collectingDate box="[675,756,1209,1231]" pageId="24" pageNumber="997" value="1952-10">X.1952</collectingDate>
</date>
, no collector
<specimenCount box="[313,442,1240,1262]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[459,533,1240,1261]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="24" pageNumber="997" type="University or college">DZUP</collectionCode>
ex M.
<collectorName box="[638,760,1240,1262]" pageId="24" pageNumber="997">Alvarenga</collectorName>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841327" collectingDate="1932-10-12" collectionCode="CEMT" country="Uruguay" location="Cerro" pageId="24" pageNumber="997" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Montevideo" typeStatus="lectotype">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43F2A09FDDA71FAF8" box="[274,339,1270,1291]" country="Uruguay" name="Cerro" pageId="24" pageNumber="997" stateProvince="Montevideo">Cerro</location>
de
<collectingRegion box="[379,514,1270,1292]" country="Uruguay" name="Montevideo" pageId="24" pageNumber="997">Montevideo</collectingRegion>
,
<date box="[525,639,1270,1291]" pageId="24" pageNumber="997" value="1932-10-12">
<collectingDate box="[525,639,1270,1291]" pageId="24" pageNumber="997" value="1932-10-12">12.X.1932</collectingDate>
</date>
(on dung), no collector
<specimenCount box="[302,404,1301,1323]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
and
<specimenCount box="[461,584,1301,1323]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[598,677,1301,1323]" pageId="24" pageNumber="997">CEMT</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841346" collectingDate="1935-01-18" collectionCode="RMNH" collectorName="Van Doesburg" country="Uruguay" location="Cerro" municipality="Cerro" pageId="24" pageNumber="997" specimenCount="1" specimenCount-male="1" stateProvince="Montevideo" typeStatus="lectotype">
<collectingMunicipality box="[696,761,1301,1322]" pageId="24" pageNumber="997">Cerro</collectingMunicipality>
de
<collectingRegion box="[180,316,1332,1354]" country="Uruguay" name="Montevideo" pageId="24" pageNumber="997">Montevideo</collectingRegion>
(
<emphasis box="[338,403,1332,1353]" italics="true" pageId="24" pageNumber="997">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43F6A083FDAB1FABA" box="[338,403,1332,1353]" country="Uruguay" municipality="Cerro" name="Cerro" pageId="24" pageNumber="997" stateProvince="Montevideo">Cerro</location>
</emphasis>
),
<date box="[432,543,1331,1353]" pageId="24" pageNumber="997" value="1935-01-18">
<collectingDate box="[432,543,1331,1353]" pageId="24" pageNumber="997" value="1935-01-18">18.I.1935</collectingDate>
</date>
, no collector
<specimenCount pageId="24" pageNumber="997" type="male">one male</specimenCount>
(
<collectionCode box="[219,304,1362,1383]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="24" pageNumber="997" type="Museum">RMNH</collectionCode>
ex
<collectorName box="[371,539,1362,1384]" pageId="24" pageNumber="997">Van Doesburg</collectorName>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841474" collectionCode="BMNH" collectorName="Rio de la Plata" country="Uruguay" location="Fry" pageId="24" pageNumber="997" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Montevideo" typeStatus="lectotype">
<collectorName pageId="24" pageNumber="997">Río de la Plata</collectorName>
, no further data
<specimenCount box="[466,572,1393,1415]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
and
<specimenCount box="[633,760,1393,1415]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[152,235,1424,1445]" pageId="24" pageNumber="997">BMNH</collectionCode>
ex
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43F1D089BDA6CFA56" box="[293,334,1424,1445]" country="Uruguay" name="Fry" pageId="24" pageNumber="997" stateProvince="Montevideo">Fry</location>
collection)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841398" collectingDate="1963-01-24" collectionCode="MZSP" collectorName="A. Toccheton" country="Uruguay" location="5 km NW Belen'" pageId="24" pageNumber="997" specimenCount="1" specimenCount-female="1" stateProvince="Salto" typeStatus="lectotype">
<emphasis box="[468,560,1424,1445]" italics="true" pageId="24" pageNumber="997">
<collectingRegion box="[468,554,1424,1445]" country="Uruguay" name="Salto" pageId="24" pageNumber="997">SALTO</collectingRegion>
:
</emphasis>
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43C020884D9D7FA55" box="[570,757,1423,1446]" country="Uruguay" name="5 km NW Belen'" pageId="24" pageNumber="997" stateProvince="Salto">
<quantity box="[570,627,1423,1446]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="997" unit="km" value="5.0">5 km</quantity>
NW Belén
</location>
,
<date box="[145,260,1454,1476]" pageId="24" pageNumber="997" value="1963-01-24">
<collectingDate box="[145,260,1454,1476]" pageId="24" pageNumber="997" value="1963-01-24">24.I.1963</collectingDate>
</date>
,
<collectorName box="[275,432,1454,1476]" pageId="24" pageNumber="997">A. Toccheton</collectorName>
leg.
<specimenCount box="[518,651,1454,1476]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[670,749,1454,1476]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="24" pageNumber="997" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841470" collectionCode="MNHG" country="Uruguay" location="SAN JOSE" pageId="24" pageNumber="997" specimenCount="1" specimenCount-female="1" stateProvince="San Jose" typeStatus="lectotype">
<emphasis box="[145,285,1483,1507]" italics="true" pageId="24" pageNumber="997">
<collectingRegion box="[145,278,1483,1507]" country="Uruguay" name="San Jose" pageId="24" pageNumber="997">SAN JOSÉ</collectingRegion>
:
</emphasis>
<emphasis box="[297,421,1485,1507]" italics="true" pageId="24" pageNumber="997">
<collectorName box="[297,421,1485,1507]" pageId="24" pageNumber="997">Mr. Claras</collectorName>
</emphasis>
,
<collectorName box="[441,616,1485,1507]" pageId="24" pageNumber="997">Río de la Plata</collectorName>
(
<emphasis box="[638,740,1485,1507]" italics="true" pageId="24" pageNumber="997">
<collectorName box="[638,740,1485,1507]" pageId="24" pageNumber="997">La Plata</collectorName>
</emphasis>
), no further data
<specimenCount box="[360,489,1516,1538]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[506,596,1516,1538]" pageId="24" pageNumber="997">MNHG</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841493" collectionCode="RBINS" collectorName="Parry" country="Uruguay" location="Probably" pageId="24" pageNumber="997" specimenCount="1" specimenCount-female="1" stateProvince="San Jose" typeStatus="lectotype">
<emphasis bold="true" pageId="24" pageNumber="997">
<emphasis bold="true" italics="true" pageId="24" pageNumber="997">
<collectorName box="[616,652,1516,1537]" pageId="24" pageNumber="997">No</collectorName>
locality data
</emphasis>
:
</emphasis>
no date,
<collectorName box="[326,393,1547,1568]" pageId="24" pageNumber="997">Parry</collectorName>
leg.
<specimenCount box="[473,602,1546,1568]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[619,704,1546,1567]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="24" pageNumber="997" type="Museum">RBINS</collectionCode>
ex
<collectorName box="[145,307,1577,1599]" pageId="24" pageNumber="997">J. La Fontaine</collectorName>
collection).
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43F810B23D90EF9CD" box="[441,556,1576,1598]" country="Uruguay" name="Probably" pageId="24" pageNumber="997" stateProvince="San Jose">Probably</location>
erroneous data
</materialsCitation>
<emphasis bold="true" box="[754,761,1577,1598]" pageId="24" pageNumber="997">:</emphasis>
<materialsCitation ID-GBIF-Occurrence="3704841318" collectionCode="MHNG" collectorName="Porto Alegre" country="Brazil" location="RIO GRANDE DO SUL" pageId="24" pageNumber="997" specimenCount="1" specimenCount-female="1" stateProvince="Rio Grande do Sul" typeStatus="lectotype">
<emphasis bold="true" box="[145,259,1607,1628]" pageId="24" pageNumber="997">
<collectingCountry box="[145,254,1607,1628]" name="Brazil" pageId="24" pageNumber="997">BRAZIL</collectingCountry>
:
</emphasis>
<emphasis box="[268,560,1608,1629]" italics="true" pageId="24" pageNumber="997">
<collectingRegion box="[268,553,1608,1629]" country="Brazil" name="Rio Grande do Sul" pageId="24" pageNumber="997">RIO GRANDE DO SUL</collectingRegion>
:
</emphasis>
<collectorName box="[569,720,1608,1630]" pageId="24" pageNumber="997">Porto Alegre</collectorName>
, no further data
<specimenCount box="[308,442,1638,1660]" pageId="24" pageNumber="997" type="female">two females</specimenCount>
(
<collectionCode box="[455,543,1638,1660]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="24" pageNumber="997" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841481" collectionCode="RBINS" collectorName="Uncertain" country="Brazil" pageId="24" pageNumber="997" specimenCount="1" specimenCount-male="1" stateProvince="Rio Grande do Sul" typeStatus="lectotype">
<emphasis bold="true" box="[559,761,1638,1660]" pageId="24" pageNumber="997">
<emphasis bold="true" box="[559,754,1638,1660]" italics="true" pageId="24" pageNumber="997">
<collectorName box="[559,689,1638,1659]" pageId="24" pageNumber="997">Uncertain</collectorName>
data
</emphasis>
:
</emphasis>
10, no further data
<specimenCount box="[413,519,1669,1691]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(
<collectionCode box="[536,623,1669,1691]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="24" pageNumber="997" type="Museum">RBINS</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841500" collectingDate="1925-11-13" collectionCode="NMPC" collectorName="Bridarolli" country="Brazil" location="Martinez" pageId="24" pageNumber="997" specimenCount="1" specimenCount-male="1" stateProvince="Rio Grande do Sul" typeStatus="lectotype">
<emphasis box="[649,755,1670,1691]" italics="true" pageId="24" pageNumber="997">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43CB10B8DD9D1F968" box="[649,755,1670,1691]" country="Brazil" name="Martinez" pageId="24" pageNumber="997" stateProvince="Rio Grande do Sul">Martinez</location>
</emphasis>
(?),
<date box="[190,322,1699,1721]" pageId="24" pageNumber="997" value="1925-11-13">
<collectingDate box="[190,322,1699,1721]" pageId="24" pageNumber="997" value="1925-11-13">13.XI.1925</collectingDate>
</date>
,
<collectorName box="[337,455,1700,1722]" pageId="24" pageNumber="997">Bridarolli</collectorName>
leg.
<specimenCount box="[538,647,1700,1722]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(
<collectionCode box="[664,749,1700,1722]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="24" pageNumber="997">NMPC</collectionCode>
)
</materialsCitation>
.
<emphasis bold="true" box="[145,349,1730,1752]" pageId="24" pageNumber="997">
<emphasis bold="true" box="[145,343,1730,1752]" italics="true" pageId="24" pageNumber="997">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43EA90BC8DA36F92B" box="[145,276,1731,1752]" country="Brazil" name="Erroneous" pageId="24" pageNumber="997" stateProvince="Rio Grande do Sul">Erroneous</location>
data
</emphasis>
:
</emphasis>
<materialsCitation ID-GBIF-Occurrence="3704841552" collectionCode="RMNH" country="Argentina" location="ARGENTINA" pageId="24" pageNumber="997" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<emphasis bold="true" box="[356,531,1730,1752]" pageId="24" pageNumber="997">
<collectingCountry box="[356,526,1730,1752]" name="Argentina" pageId="24" pageNumber="997">ARGENTINA</collectingCountry>
:
</emphasis>
<collectorName box="[538,654,1730,1752]" pageId="24" pageNumber="997">Patagonia</collectorName>
(
<emphasis box="[674,752,1731,1752]" italics="true" pageId="24" pageNumber="997">
<collectorName box="[674,752,1731,1752]" pageId="24" pageNumber="997">Patago</collectorName>
</emphasis>
- /
<emphasis box="[160,195,1761,1782]" italics="true" pageId="24" pageNumber="997">nie</emphasis>
), no further data
<specimenCount box="[438,545,1761,1783]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(
<collectionCode box="[561,646,1761,1782]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="24" pageNumber="997" type="Museum">RMNH</collectionCode>
ex
<collectorName pageId="24" pageNumber="997">A. J. Buis</collectorName>
and
<collectorName box="[261,431,1791,1813]" pageId="24" pageNumber="997">F. C. Drescher</collectorName>
collections)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841399" collectionCode="NHMW" country="Australia" location="Zelenka" pageId="24" pageNumber="997" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="lectotype">
<emphasis bold="true" box="[588,761,1791,1813]" pageId="24" pageNumber="997">
<collectingCountry box="[588,755,1791,1813]" name="Australia" pageId="24" pageNumber="997">AUSTRALIA</collectingCountry>
:
</emphasis>
no further data
<specimenCount box="[358,465,1822,1844]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
and
<specimenCount box="[526,654,1822,1844]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[669,760,1822,1843]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="24" pageNumber="997" type="Museum">NHMW</collectionCode>
ex W.
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43EC80A36DA72F8A0" box="[240,336,1853,1875]" country="Australia" name="Zelenka" pageId="24" pageNumber="997">Zelenka</location>
collection)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841347" collectionCode="BMNH, MNHN, R, NMPC" collectorName="Edgar von Harold &amp; Oberthur" country="Australia" county="No" location="Escrapus" municipality="Coelatocollis Reiche'" pageId="24" pageNumber="997" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="paralectotype">
<emphasis bold="true" box="[485,600,1852,1874]" pageId="24" pageNumber="997">
<emphasis bold="true" box="[485,593,1852,1874]" italics="true" pageId="24" pageNumber="997">
<collectingCounty box="[485,521,1853,1874]" pageId="24" pageNumber="997">No</collectingCounty>
data
</emphasis>
:
</emphasis>
<specimenCount box="[610,760,1853,1875]" pageId="24" pageNumber="997" type="female">four females</specimenCount>
(one labelled with the
<emphasis box="[410,605,1883,1905]" italics="true" pageId="24" pageNumber="997">nomen in litteris</emphasis>
<collectingMunicipality box="[617,894,197,1904]" pageId="24" pageNumber="997">
<emphasis box="[617,761,1883,1904]" italics="true" pageId="24" pageNumber="997">Coelatocollis</emphasis>
Reiche
</collectingMunicipality>
, another with the
<emphasis box="[1111,1305,197,219]" italics="true" pageId="24" pageNumber="997">nomen in litteris</emphasis>
<emphasis italics="true" pageId="24" pageNumber="997">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43B1D0DCDDEB2FF28" box="[1317,1424,198,219]" country="Australia" county="No" municipality="Coelatocollis Reiche'" name="Escrapus" pageId="24" pageNumber="997">Escrapus</location>
singularis
</emphasis>
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43D9D0DE8D8DBFF0A" box="[933,1017,227,249]" country="Australia" county="No" municipality="Coelatocollis Reiche'" name="Reiche'" pageId="24" pageNumber="997">Reiche</location>
) (
<collectionCode box="[1039,1123,228,249]" pageId="24" pageNumber="997">BMNH</collectionCode>
ex Nevinson collection),
<specimenCount box="[809,917,258,280]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
and
<specimenCount box="[981,1112,258,280]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(potential
<typeStatus box="[1248,1423,258,280]" pageId="24" pageNumber="997">paralectotypes</typeStatus>
of
<taxonomicName authorityName="Harold des Cupello" authorityYear="1868" box="[839,978,289,311]" class="Insecta" family="Scarabaeidae" genus="Bolbites" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="997" phylum="Arthropoda" rank="species" species="onitoides">
<emphasis box="[839,978,289,311]" italics="true" pageId="24" pageNumber="997">B. onitoides</emphasis>
</taxonomicName>
) (
<collectionCode box="[1003,1090,289,310]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="24" pageNumber="997">MNHN</collectionCode>
ex
<collectorName box="[1157,1372,289,311]" pageId="24" pageNumber="997">Edgar von Harold</collectorName>
and
<collectionCode box="[809,831,320,341]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="24" pageNumber="997">R</collectionCode>
.
<collectorName box="[847,965,319,341]" pageId="24" pageNumber="997">Oberthür</collectorName>
collection),
<specimenCount box="[1129,1246,319,341]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(
<collectionCode box="[1266,1355,320,342]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="24" pageNumber="997">NMPC</collectionCode>
)
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3704841382" collectionCode="RBINS" collectorName="Edgar von Harold &amp; Oberthur" country="Australia" county="No" isTailEnumeration="true" location="Escrapus" municipality="Coelatocollis Reiche'" pageId="24" pageNumber="997" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<specimenCount pageId="24" pageNumber="997" type="female">two females</specimenCount>
(
<collectionCode box="[911,996,350,372]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="24" pageNumber="997" type="Museum">RBINS</collectionCode>
)
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3704841559" collectionCode="RBINS" collectorName="A. Janssens &amp; E. Candeze" country="Australia" location="Le Moult" pageId="24" pageNumber="997" specimenCount="4" specimenCount-female="3" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount box="[1014,1137,350,372]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[1151,1234,350,371]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="24" pageNumber="997" type="Museum">RBINS</collectionCode>
ex
<collectorName box="[1290,1424,350,372]" pageId="24" pageNumber="997">A. Janssens</collectorName>
collection),
<specimenCount box="[958,1096,381,403]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[1116,1205,381,402]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="24" pageNumber="997" type="Museum">RBINS</collectionCode>
ex
<collectorName box="[1281,1424,381,403]" pageId="24" pageNumber="997">E. Candèze</collectorName>
collection) and
<specimenCount box="[984,1088,411,433]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
and
<specimenCount box="[1147,1272,411,433]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[1287,1370,412,433]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="24" pageNumber="997" type="Museum">RBINS</collectionCode>
ex
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43D110CB0D8B0FE23" box="[809,914,442,464]" country="Australia" name="Le Moult" pageId="24" pageNumber="997">Le Moult</location>
collection)
</materialsCitation>
.
</paragraph>
<paragraph blockId="24.[809,1425,529,1900]" lastBlockId="25.[827,1443,197,1874]" lastPageId="25" lastPageNumber="998" pageId="24" pageNumber="997">
<emphasis box="[809,994,529,550]" italics="true" pageId="24" pageNumber="997">Red specimens:</emphasis>
<materialsCitation ID-GBIF-Occurrence="3704841358" collectionCode="BMNH" collectorName="O. W. Thomas" country="Argentina" location="ARGENTINA" pageId="24" pageNumber="997" specimenCount="1" specimenCount-female="1">
<emphasis bold="true" box="[1004,1186,529,551]" pageId="24" pageNumber="997">
<collectingCountry box="[1004,1181,529,551]" name="Argentina" pageId="24" pageNumber="997">ARGENTINA</collectingCountry>
:
</emphasis>
no specific locality or date,
<collectorName box="[914,1082,559,581]" pageId="24" pageNumber="997">O. W. Thomas</collectorName>
leg.
<specimenCount box="[1169,1304,559,581]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[1322,1412,560,582]" pageId="24" pageNumber="997">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841303" collectionCode="BMNH" collectorName="Taylor" country="Argentina" pageId="24" pageNumber="997" specimenCount="1" specimenCount-male="1">
no specific locality or date,
<collectorName box="[1151,1229,590,612]" pageId="24" pageNumber="997">Taylor</collectorName>
leg.
<specimenCount box="[1314,1424,590,612]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(
<collectionCode box="[817,908,621,643]" pageId="24" pageNumber="997">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841507" collectionCode="MACN" country="Argentina" location="BUENOS AIRES" pageId="24" pageNumber="997" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Ciudad Autonoma de Buenos Aires">
<emphasis box="[929,1148,621,642]" italics="true" pageId="24" pageNumber="997">
<collectingRegion box="[929,1140,621,642]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="24" pageNumber="997">BUENOS AIRES</collectingRegion>
:
</emphasis>
no further data
<specimenCount pageId="24" pageNumber="997" type="male">one male</specimenCount>
and
<specimenCount box="[936,1108,651,673]" pageId="24" pageNumber="997" type="female">three females</specimenCount>
(
<collectionCode box="[1127,1214,652,674]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="24" pageNumber="997" type="Museum">MACN</collectionCode>
)
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3704841573" collectionCode="MNHN" country="Argentina" location="Granger" pageId="24" pageNumber="997" specimenCount="1" specimenCount-female="1" stateProvince="Ciudad Autonoma de Buenos Aires">
and
<specimenCount box="[1289,1423,651,673]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[816,902,682,703]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="24" pageNumber="997">MNHN</collectionCode>
ex
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43D850FA1DF3CFD4C" box="[957,1054,682,703]" country="Argentina" name="Granger" pageId="24" pageNumber="997" stateProvince="Ciudad Autonoma de Buenos Aires">Granger</location>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841441" collectionCode="MACN" collectorName="Carlos Bruch" country="Argentina" location="Carlos Bruch" pageId="24" pageNumber="997" specimenCount="1" specimenCount-male="1" stateProvince="Ciudad Autonoma de Buenos Aires">
no specific locality or date,
<collectorName box="[871,1023,713,735]" pageId="24" pageNumber="997">Carlos Bruch</collectorName>
leg.
<specimenCount box="[1094,1197,713,735]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(
<collectionCode box="[1210,1291,713,734]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="24" pageNumber="997" type="Museum">MACN</collectionCode>
ex
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43B7D0FC2D852FD0E" country="Argentina" name="Carlos Bruch" pageId="24" pageNumber="997" stateProvince="Ciudad Autonoma de Buenos Aires">Carlos Bruch</location>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841309" collectionCode="MLPA" collectorName="J. Bosq" country="Argentina" pageId="24" pageNumber="997" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Ciudad Autonoma de Buenos Aires">
no specific locality or date,
<collectorName box="[1342,1424,744,765]" pageId="24" pageNumber="997">J. Bosq</collectorName>
leg.
<specimenCount box="[881,990,774,796]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
and
<specimenCount box="[1052,1182,774,796]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[1198,1278,774,796]" pageId="24" pageNumber="997">MLPA</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841310" collectingDate="1894-01" collectionCode="MLPA" country="Argentina" pageId="24" pageNumber="997" specimenCount="1" specimenCount-female="1" stateProvince="Ciudad Autonoma de Buenos Aires">
no specific locality,
<date box="[912,992,805,826]" pageId="24" pageNumber="997" value="1894-01">
<collectingDate box="[912,992,805,826]" pageId="24" pageNumber="997" value="1894-01">I.1894</collectingDate>
</date>
, no collector
<specimenCount box="[1181,1313,805,827]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[1331,1412,805,827]" pageId="24" pageNumber="997">MLPA</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841380" collectingDate="1993-01-13" collectionCode="CEMT" collectorName="Ugarte-Pena" country="Argentina" location="Charata" municipality="Chacabuco" pageId="24" pageNumber="997" specimenCount="1" specimenCount-male="1" stateProvince="Chaco">
<emphasis box="[809,909,836,857]" italics="true" pageId="24" pageNumber="997">
<collectingRegion box="[809,903,836,857]" country="Argentina" name="Chaco" pageId="24" pageNumber="997">CHACO</collectingRegion>
:
</emphasis>
<collectingMunicipality box="[914,1041,835,857]" pageId="24" pageNumber="997">Chacabuco</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43A230E48DF5FFCAA" box="[1051,1149,835,857]" country="Argentina" municipality="Chacabuco" name="Charata" pageId="24" pageNumber="997" stateProvince="Chaco">Charata</location>
,
<date box="[1158,1267,836,857]" pageId="24" pageNumber="997" value="1993-01-13">
<collectingDate box="[1158,1267,836,857]" pageId="24" pageNumber="997" value="1993-01-13">13.I.1993</collectingDate>
</date>
,
<collectorName box="[1277,1424,836,857]" pageId="24" pageNumber="997">Ugarte-Peña</collectorName>
leg.
<specimenCount box="[877,982,866,888]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(
<collectionCode box="[997,1077,866,888]" pageId="24" pageNumber="997">CEMT</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841508" collectingDate="1995-01" collectionCode="NMPC, T" collectorName="S. Pokorny" country="Argentina" county="Chacabuco" location="Taylor" municipality="Charata" pageId="24" pageNumber="997" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Chaco">
<collectingCounty box="[1096,1223,866,888]" pageId="24" pageNumber="997">Chacabuco</collectingCounty>
,
<collectingMunicipality box="[1235,1333,866,888]" pageId="24" pageNumber="997">Charata</collectingMunicipality>
,
<date box="[1345,1420,866,888]" pageId="24" pageNumber="997" value="1995-01">
<collectingDate box="[1345,1420,866,888]" pageId="24" pageNumber="997" value="1995-01">I.1995</collectingDate>
</date>
, no collection
<specimenCount box="[990,1098,897,919]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(dissected) and
<specimenCount box="[1295,1424,897,919]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[816,898,928,949]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="24" pageNumber="997">NMPC</collectionCode>
ex
<collectionCode box="[968,986,928,949]" name="Tavera, Department of Geology and Geophysics" pageId="24" pageNumber="997">T</collectionCode>
.
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43DDD0E94DF11FC46" box="[997,1075,927,949]" country="Argentina" county="Chacabuco" municipality="Charata" name="Taylor" pageId="24" pageNumber="997" stateProvince="Chaco">Taylor</location>
and
<collectorName box="[1139,1271,927,949]" pageId="24" pageNumber="997">S. Pokorný</collectorName>
collections)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841542" collectingDate="2001-12" collectionCode="BMNH" collectorName="S. Bolle &amp; J. Zidek" country="Argentina" location="Charata" municipality="Chacabuco" pageId="24" pageNumber="997" specimenCount="1" specimenCount-male="1" stateProvince="Chaco">
<collectingMunicipality box="[809,944,958,980]" pageId="24" pageNumber="997">Chacabuco</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43D860EB5DF04FC27" box="[958,1062,958,980]" country="Argentina" municipality="Chacabuco" name="Charata" pageId="24" pageNumber="997" stateProvince="Chaco">Charata</location>
,
<date box="[1077,1186,958,979]" pageId="24" pageNumber="997" value="2001-12">
<collectingDate box="[1077,1186,958,979]" pageId="24" pageNumber="997" value="2001-12">XII.2001</collectingDate>
</date>
,
<collectorName box="[1201,1295,958,980]" pageId="24" pageNumber="997">S. Bolle</collectorName>
leg.
<specimenCount pageId="24" pageNumber="997" type="male">one male</specimenCount>
(
<collectionCode box="[882,967,989,1010]" pageId="24" pageNumber="997">BMNH</collectionCode>
ex
<collectorName box="[1032,1125,989,1011]" pageId="24" pageNumber="997">J. Zídek</collectorName>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841331" collectingDate="1939-03-03" collectionCode="MLPA" country="Argentina" county="Liberdad" location="La Chanza'" municipality="Puerto Tirol'" pageId="24" pageNumber="997" specimenCount="1" specimenCount-male="1" stateProvince="Chaco">
<collectingCounty box="[1270,1379,989,1011]" pageId="24" pageNumber="997">Liberdad</collectingCounty>
,
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="24" pageNumber="997" unit="km" value="15.0">15 km</quantity>
from
<collectingMunicipality box="[920,1075,1019,1041]" pageId="24" pageNumber="997">Puerto Tirol</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43A7D0EF7DFF3FBE2" box="[1093,1233,1019,1041]" country="Argentina" county="Liberdad" municipality="Puerto Tirol'" name="La Chanza'" pageId="24" pageNumber="997" stateProvince="Chaco">La Chanza</location>
,
<date box="[1247,1383,1020,1041]" pageId="24" pageNumber="997" value="1939-03-03">
<collectingDate box="[1247,1383,1020,1041]" pageId="24" pageNumber="997" value="1939-03-03">03.III.1939</collectingDate>
</date>
, no collector
<specimenCount box="[937,1042,1050,1072]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(
<collectionCode box="[1058,1138,1050,1072]" pageId="24" pageNumber="997">MLPA</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841454" collectionCode="MCZC" collectorName="Davis" country="Argentina" location="CORDOBA" pageId="24" pageNumber="997" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="CORDOBA">
<emphasis box="[1156,1295,1049,1073]" italics="true" pageId="24" pageNumber="997">
<collectingRegion box="[1156,1288,1049,1073]" country="Argentina" name="CORDOBA" pageId="24" pageNumber="997">CÓRDOBA</collectingRegion>
:
</emphasis>
no locality or date,
<collectorName box="[900,966,1081,1103]" pageId="24" pageNumber="997">Davis</collectorName>
col.
<specimenCount box="[1034,1136,1081,1103]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
and
<specimenCount box="[1191,1327,1081,1103]" pageId="24" pageNumber="997" type="female">two females</specimenCount>
(
<collectionCode box="[1340,1417,1081,1103]" pageId="24" pageNumber="997">MCZC</collectionCode>
)
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3704841421" box="[809,1086,1111,1133]" collectionCode="NMPC" country="Argentina" pageId="24" pageNumber="997" specimenCount="1" specimenCount-male="1" stateProvince="CORDOBA">
and
<specimenCount box="[864,974,1111,1133]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(
<collectionCode box="[992,1079,1111,1133]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="24" pageNumber="997">NMPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841485" collectionCode="MLPA" country="Argentina" location="CORRIENTES" pageId="24" pageNumber="997" specimenCount="1" specimenCount-male="1" stateProvince="Corrientes">
<emphasis box="[1100,1289,1112,1133]" italics="true" pageId="24" pageNumber="997">
<collectingRegion box="[1100,1283,1112,1133]" country="Argentina" name="Corrientes" pageId="24" pageNumber="997">CORRIENTES</collectingRegion>
:
</emphasis>
no further data
<specimenCount box="[900,1013,1142,1164]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(
<collectionCode box="[1031,1114,1142,1164]" pageId="24" pageNumber="997">MLPA</collectionCode>
)
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3704841486" box="[1137,1368,1142,1164]" collectionCode="MNHN" country="Argentina" isTailEnumeration="true" location="CORRIENTES" pageId="24" pageNumber="997" specimenCount="1" specimenCount-male="1" stateProvince="Corrientes">
<specimenCount box="[1137,1250,1142,1164]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(
<collectionCode box="[1269,1360,1142,1164]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="24" pageNumber="997">MNHN</collectionCode>
)
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3704841556" collectionCode="ZSM" country="Argentina" pageId="24" pageNumber="997" specimenCount="1" specimenCount-female="1" stateProvince="Corrientes">
and
<specimenCount box="[809,939,1173,1195]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[957,1018,1173,1195]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="24" pageNumber="997" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841396" collectingDate="1925-10" collectionCode="ZSM" country="Argentina" county="Colon" location="Chaco" municipality="San Jose" pageId="24" pageNumber="997" specimenCount="1" specimenCount-male="1" stateProvince="Entre Rios">
<emphasis box="[1039,1217,1171,1195]" italics="true" pageId="24" pageNumber="997">
<collectingRegion box="[1039,1210,1171,1195]" country="Argentina" name="Entre Rios" pageId="24" pageNumber="997">ENTRE RÍOS</collectingRegion>
:
</emphasis>
<collectingCounty box="[1226,1297,1173,1195]" pageId="24" pageNumber="997">Colón</collectingCounty>
,
<collectingMunicipality box="[1311,1419,1173,1195]" pageId="24" pageNumber="997">San José</collectingMunicipality>
,
<date box="[809,894,1203,1225]" pageId="24" pageNumber="997" value="1925-10">
<collectingDate box="[809,894,1203,1225]" pageId="24" pageNumber="997" value="1925-10">X.1925</collectingDate>
</date>
, D.
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43D9509B8D8D4FB3A" box="[941,1014,1203,1225]" country="Argentina" county="Colon" municipality="San Jose" name="Chaco" pageId="24" pageNumber="997" stateProvince="Entre Rios">Chaco</location>
expedition leg.
<specimenCount box="[1228,1336,1203,1225]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(
<collectionCode box="[1353,1413,1203,1225]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="24" pageNumber="997" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841405" collectingDate="1962-01" collectionCode="MLPA" country="Argentina" location="Pronunciamiento" municipality="Uruguay" pageId="24" pageNumber="997" specimenCount="1" specimenCount-male="1" stateProvince="Entre Rios">
<collectingMunicipality box="[809,911,1234,1255]" pageId="24" pageNumber="997">Uruguay</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43DA209D9DF41FB1B" box="[922,1123,1234,1256]" country="Argentina" municipality="Uruguay" name="Pronunciamiento" pageId="24" pageNumber="997" stateProvince="Entre Rios">Pronunciamiento</location>
,
<date box="[1135,1210,1234,1256]" pageId="24" pageNumber="997" value="1962-01">
<collectingDate box="[1135,1210,1234,1256]" pageId="24" pageNumber="997" value="1962-01">I.1962</collectingDate>
</date>
, no collector
<specimenCount pageId="24" pageNumber="997" type="male">two males</specimenCount>
(
<collectionCode box="[892,970,1265,1287]" pageId="24" pageNumber="997">MLPA</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841422" collectingDate="1988-12" collectionCode="NMPC, T" collectorName="S. Pokorny" country="Argentina" county="Uruguay" location="Taylor" municipality="Pronunciamiento" pageId="24" pageNumber="997" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Entre Rios">
<collectingCounty box="[989,1091,1265,1286]" pageId="24" pageNumber="997">Uruguay</collectingCounty>
,
<collectingMunicipality box="[1103,1305,1265,1287]" pageId="24" pageNumber="997">Pronunciamiento</collectingMunicipality>
,
<date box="[1317,1420,1265,1286]" pageId="24" pageNumber="997" value="1988-12">
<collectingDate box="[1317,1420,1265,1286]" pageId="24" pageNumber="997" value="1988-12">XII.1988</collectingDate>
</date>
, no collector
<specimenCount box="[981,1090,1295,1317]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(dissected) and
<specimenCount box="[1293,1424,1295,1317]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[816,899,1326,1347]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="24" pageNumber="997">NMPC</collectionCode>
ex
<collectionCode box="[971,989,1326,1347]" name="Tavera, Department of Geology and Geophysics" pageId="24" pageNumber="997">T</collectionCode>
.
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43DD10825DF1AFAB7" box="[1001,1080,1326,1348]" country="Argentina" county="Uruguay" municipality="Pronunciamiento" name="Taylor" pageId="24" pageNumber="997" stateProvince="Entre Rios">Taylor</location>
and
<collectorName box="[1147,1280,1326,1348]" pageId="24" pageNumber="997">S. Pokorný</collectorName>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841531" collectingDate="1988-12" collectionCode="BMNH" collectorName="S. Bolle &amp; J. Zidek" country="Argentina" location="Pronunciamiento" municipality="Uruguay" pageId="24" pageNumber="997" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Entre Rios">
<collectingMunicipality box="[809,913,1357,1378]" pageId="24" pageNumber="997">Uruguay</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43DA50846DF4EFA90" box="[925,1132,1357,1379]" country="Argentina" municipality="Uruguay" name="Pronunciamiento" pageId="24" pageNumber="997" stateProvince="Entre Rios">Pronunciamiento</location>
,
<date box="[1145,1250,1357,1378]" pageId="24" pageNumber="997" value="1988-12">
<collectingDate box="[1145,1250,1357,1378]" pageId="24" pageNumber="997" value="1988-12">XII.1988</collectingDate>
</date>
,
<collectorName box="[1263,1353,1357,1379]" pageId="24" pageNumber="997">S. Bolle</collectorName>
leg.
<specimenCount box="[809,930,1387,1409]" pageId="24" pageNumber="997" type="male">two males</specimenCount>
(both dissected) and
<specimenCount box="[1193,1323,1387,1409]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[1339,1424,1387,1408]" pageId="24" pageNumber="997">BMNH</collectionCode>
ex
<collectorName box="[871,969,1418,1440]" pageId="24" pageNumber="997">J. Zídek</collectorName>
leg.)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841351" collectingDate="1949-02" collectionCode="MZSP" collectorName="A. Martinez" country="Argentina" location="Formosa" pageId="24" pageNumber="997" specimenCount="1" specimenCount-female="1" stateProvince="Formosa">
<emphasis box="[1050,1195,1418,1439]" italics="true" pageId="24" pageNumber="997">
<collectingRegion box="[1050,1189,1418,1439]" country="Argentina" name="Formosa" pageId="24" pageNumber="997">FORMOSA</collectingRegion>
:
</emphasis>
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43A8E0881DE00FA6C" box="[1206,1314,1418,1439]" country="Argentina" name="Formosa" pageId="24" pageNumber="997" stateProvince="Formosa">Formosa</location>
,
<date box="[1329,1418,1418,1439]" pageId="24" pageNumber="997" value="1949-02">
<collectingDate box="[1329,1418,1418,1439]" pageId="24" pageNumber="997" value="1949-02">II.1949</collectingDate>
</date>
,
<collectorName box="[809,952,1449,1471]" pageId="24" pageNumber="997">A. Martínez</collectorName>
leg.
<specimenCount box="[1034,1164,1449,1471]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[1181,1259,1449,1471]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="24" pageNumber="997" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841457" collectingDate="1989-07-07" collectionCode="FSCA" country="Argentina" location="Eldorado" pageId="24" pageNumber="997" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="MISIONES">
<emphasis box="[1279,1425,1449,1470]" italics="true" pageId="24" pageNumber="997">
<collectingRegion box="[1279,1418,1449,1470]" country="Argentina" name="MISIONES" pageId="24" pageNumber="997">MISIONES</collectingRegion>
:
</emphasis>
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43D1108CCD8BAFA2E" box="[809,920,1479,1501]" country="Argentina" name="Eldorado" pageId="24" pageNumber="997" stateProvince="MISIONES">Eldorado</location>
,
<date box="[933,1077,1479,1500]" pageId="24" pageNumber="997" value="1989-07-07">
<collectingDate box="[933,1077,1479,1500]" pageId="24" pageNumber="997" value="1989-07-07">07.VII.1989</collectingDate>
</date>
, no collector
<specimenCount box="[1262,1371,1479,1501]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
and
<specimenCount box="[809,939,1510,1532]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[957,1032,1510,1532]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="24" pageNumber="997" type="Museum">FSCA</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841342" collectingDate="1924-10" collectionCode="BMNH" collectorName="K. J. Hayward" country="Argentina" location="Villa Ana" municipality="General Obligado" pageId="24" pageNumber="997" specimenCount="1" specimenCount-male="1" stateProvince="Santa Fe">
<emphasis box="[1053,1196,1510,1532]" italics="true" pageId="24" pageNumber="997">
<collectingRegion box="[1053,1190,1510,1532]" country="Argentina" name="Santa Fe" pageId="24" pageNumber="997">SANTA FE</collectingRegion>
:
</emphasis>
<collectingMunicipality box="[1205,1420,1510,1532]" pageId="24" pageNumber="997">General Obligado</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43D110B0ED883F9E9" box="[809,929,1541,1563]" country="Argentina" municipality="General Obligado" name="Villa Ana" pageId="24" pageNumber="997" stateProvince="Santa Fe">Villa Ana</location>
,
<date box="[944,1032,1541,1562]" pageId="24" pageNumber="997" value="1924-10">
<collectingDate box="[944,1032,1541,1562]" pageId="24" pageNumber="997" value="1924-10">X.1924</collectingDate>
</date>
,
<collectorName box="[1047,1225,1541,1563]" pageId="24" pageNumber="997">K. J. Hayward</collectorName>
leg.
<specimenCount box="[1313,1424,1541,1563]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(
<collectionCode box="[817,909,1571,1593]" pageId="24" pageNumber="997">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841490" collectingDate="1926-03" collectionCode="BMNH" collectorName="K. J. Hayward" country="Argentina" location="Villa Ana" municipality="General Obligado" pageId="24" pageNumber="997" specimenCount="1" specimenCount-male="1" stateProvince="Santa Fe">
<collectingMunicipality box="[933,1160,1571,1593]" pageId="24" pageNumber="997">General Obligado</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43AAF0B28DE30F9CB" box="[1175,1298,1571,1593]" country="Argentina" municipality="General Obligado" name="Villa Ana" pageId="24" pageNumber="997" stateProvince="Santa Fe">Villa Ana</location>
,
<date box="[1315,1419,1571,1593]" pageId="24" pageNumber="997" value="1926-03">
<collectingDate box="[1315,1419,1571,1593]" pageId="24" pageNumber="997" value="1926-03">III.1926</collectingDate>
</date>
,
<collectorName box="[809,979,1602,1624]" pageId="24" pageNumber="997">K. J. Hayward</collectorName>
leg.
<specimenCount box="[1059,1167,1602,1624]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(
<collectionCode box="[1183,1271,1602,1624]" pageId="24" pageNumber="997">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841572" collectionCode="SMF" collectorName="Rosario &amp; J. Hubrich" country="Argentina" pageId="24" pageNumber="997" specimenCount="1" specimenCount-male="1" stateProvince="Santa Fe">
<collectorName box="[1292,1384,1602,1624]" pageId="24" pageNumber="997">Rosario</collectorName>
, no date,
<collectorName box="[877,1004,1633,1655]" pageId="24" pageNumber="997">J. Hubrich</collectorName>
leg.
<specimenCount box="[1086,1195,1633,1655]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(
<collectionCode box="[1212,1274,1633,1655]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="24" pageNumber="997">SMF</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841498" collectionCode="BMNH, MNHN" collectorName="Dejean" country="Bolivia" location="Reiche'" pageId="24" pageNumber="997" specimenCount="2" specimenCount-male="2">
<emphasis bold="true" box="[1295,1425,1632,1654]" pageId="24" pageNumber="997">
<collectingCountry box="[1295,1419,1632,1654]" name="Bolivia" pageId="24" pageNumber="997">BOLIVIA</collectingCountry>
:
</emphasis>
no further data
<specimenCount box="[1019,1125,1663,1685]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(labelled with the
<emphasis italics="true" pageId="24" pageNumber="997">nomen in litteris</emphasis>
<emphasis box="[943,1211,1694,1715]" italics="true" pageId="24" pageNumber="997">AEschropus singularis</emphasis>
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43AFD0B95DE3DF947" box="[1221,1311,1694,1716]" country="Bolivia" name="Reiche'" pageId="24" pageNumber="997">Reiche</location>
) (
<collectionCode box="[1337,1423,1694,1715]" pageId="24" pageNumber="997">BMNH</collectionCode>
ex
<collectorName box="[875,963,1725,1747]" pageId="24" pageNumber="997">Dejean</collectorName>
collection) and
<specimenCount box="[1179,1296,1725,1747]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
(
<collectionCode box="[1316,1411,1725,1747]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="24" pageNumber="997">MNHN</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841482" collectionCode="RBINS" collectorName="Parry" country="Brazil" location="La Fontaine" pageId="24" pageNumber="997" specimenCount="1" specimenCount-male="1">
<emphasis bold="true" box="[809,920,1755,1776]" pageId="24" pageNumber="997">
<collectingCountry box="[809,916,1755,1776]" name="Brazil" pageId="24" pageNumber="997">BRAZIL</collectingCountry>
:
</emphasis>
no specific locality or date,
<collectorName box="[1243,1308,1756,1777]" pageId="24" pageNumber="997">Parry</collectorName>
leg.
<specimenCount pageId="24" pageNumber="997" type="male">one male</specimenCount>
(
<collectionCode box="[882,967,1786,1807]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="24" pageNumber="997" type="Museum">RBINS</collectionCode>
ex J.
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43A1B0BF1DF91F8E3" box="[1059,1203,1786,1808]" country="Brazil" name="La Fontaine" pageId="24" pageNumber="997">La Fontaine</location>
collection)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841521" collectingDate="2008-01-23" collectionCode="CEMT" collectorName="F. R. Fernandes" country="Brazil" latitude="-21.697498" location="Porto Murtinho" longLatPrecision="21" longitude="-57.88889" pageId="24" pageNumber="997" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Mato Grosso do Sul">
<emphasis italics="true" pageId="24" pageNumber="997">
<collectingRegion country="Brazil" name="Mato Grosso do Sul" pageId="24" pageNumber="997">MATO GROSSO DO SUL</collectingRegion>
:
</emphasis>
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE43A110A12DFD0F8DC" box="[1065,1266,1817,1839]" country="Brazil" latitude="-21.697498" longLatPrecision="21" longitude="-57.88889" name="Porto Murtinho" pageId="24" pageNumber="997" stateProvince="Mato Grosso do Sul">Porto Murtinho</location>
,
<geoCoordinate box="[1282,1419,1816,1838]" degrees="21" direction="south" minutes="41" orientation="latitude" pageId="24" pageNumber="997" precision="15" seconds="51" value="-21.697498">21°4151”S</geoCoordinate>
,
<geoCoordinate box="[809,941,1847,1869]" degrees="57" direction="west" minutes="53" orientation="longitude" pageId="24" pageNumber="997" precision="15" seconds="20" value="-57.88889">57°5320”W</geoCoordinate>
,
<date box="[951,1061,1847,1868]" pageId="24" pageNumber="997" value="2008-01-23">
<collectingDate box="[951,1061,1847,1868]" pageId="24" pageNumber="997" value="2008-01-23">23.I.2008</collectingDate>
</date>
,
<collectorName box="[1072,1250,1847,1869]" pageId="24" pageNumber="997">F. R. Fernandes</collectorName>
leg.
<specimenCount box="[1322,1424,1847,1869]" pageId="24" pageNumber="997" type="male">one male</specimenCount>
and
<specimenCount box="[860,985,1878,1900]" pageId="24" pageNumber="997" type="female">one female</specimenCount>
(
<collectionCode box="[1000,1080,1878,1900]" pageId="24" pageNumber="997">CEMT</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841393" collectingDate="2008-01-25" collectionCode="CEMT" collectorName="F. R. Fernandes" country="Brazil" lastPageId="25" lastPageNumber="998" location="Pao de Acucar" municipality="Porto Murtinho" pageId="24" pageNumber="997" specimenCount="1" specimenCount-male="1" stateProvince="Mato Grosso do Sul">
<collectingMunicipality box="[1099,1280,1878,1900]" pageId="24" pageNumber="997">Porto Murtinho</collectingMunicipality>
, road to
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC18FFE53B5E0A5DDA34FF28" country="Brazil" lastPageId="25" lastPageNumber="998" municipality="Porto Murtinho" name="Pao de Acucar" pageId="24" pageNumber="997" stateProvince="Mato Grosso do Sul">Pão de Açúcar</location>
,
<date box="[290,400,197,219]" pageId="25" pageNumber="998" value="2008-01-25">
<collectingDate box="[290,400,197,219]" pageId="25" pageNumber="998" value="2008-01-25">25.I.2008</collectingDate>
</date>
,
<collectorName box="[413,596,197,219]" pageId="25" pageNumber="998">F. R. Fernandes</collectorName>
leg.
<specimenCount box="[673,778,197,219]" pageId="25" pageNumber="998" type="male">one male</specimenCount>
(
<collectionCode box="[170,253,228,250]" pageId="25" pageNumber="998">CEMT</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841360" collectingDate="2011-11-08" collectingDateMax="2011-11-14" collectingDateMin="2011-11-08" collectionCode="CEMT" collectorName="R. M. Moraes" country="Brazil" latitude="-31.458334" location="Fazenda Sentinela" longLatPrecision="20" longitude="-54.355" municipality="Acegua" pageId="25" pageNumber="998" specimenCount="1" specimenCount-male="1" stateProvince="Rio Grande do Sul">
<emphasis box="[274,568,228,249]" italics="true" pageId="25" pageNumber="998">
<collectingRegion box="[274,560,228,249]" country="Brazil" name="Rio Grande do Sul" pageId="25" pageNumber="998">RIO GRANDE DO SUL</collectingRegion>
:
</emphasis>
<collectingMunicipality box="[576,665,228,250]" pageId="25" pageNumber="998">Aceguá</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53C9E0DEFDA39FEEB" country="Brazil" latitude="-31.458334" longLatPrecision="20" longitude="-54.355" municipality="Acegua" name="Fazenda Sentinela" pageId="25" pageNumber="998" stateProvince="Rio Grande do Sul">Fazenda Sentinela</location>
,
<geoCoordinate box="[298,431,259,280]" degrees="31" direction="south" minutes="27" orientation="latitude" pageId="25" pageNumber="998" precision="15" seconds="30" value="-31.458334">31°2730”S</geoCoordinate>
,
<geoCoordinate box="[446,587,258,280]" degrees="54" direction="west" minutes="21" orientation="longitude" pageId="25" pageNumber="998" precision="15" seconds="18" value="-54.355">54°2118”W</geoCoordinate>
,
<date box="[601,778,259,280]" pageId="25" pageNumber="998" value="2011-11-08" valueMax="2011-11-14" valueMin="2011-11-08">
<collectingDate box="[601,778,259,280]" pageId="25" pageNumber="998" value="2011-11-08" valueMax="2011-11-14" valueMin="2011-11-08">0814.XI.2011</collectingDate>
</date>
(
<collectingMethod box="[170,437,289,311]" pageId="25" pageNumber="998">flight interception trap</collectingMethod>
),
<collectorName box="[456,606,290,311]" pageId="25" pageNumber="998">R. M. Moraes</collectorName>
leg.
<specimenCount box="[676,778,289,311]" pageId="25" pageNumber="998" type="male">one male</specimenCount>
(
<collectionCode box="[170,252,320,342]" pageId="25" pageNumber="998">CEMT</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841407" collectingDate="2011-11-08" collectingDateMax="2011-11-14" collectingDateMin="2011-11-08" collectionCode="CEMT" collectorName="R. M. Moraes" country="Brazil" latitude="-31.455393" location="Fazenda Sentinela" longLatPrecision="1" longitude="-54.352863" municipality="Acegua" pageId="25" pageNumber="998" specimenCount="1" specimenCount-female="1" stateProvince="Rio Grande do Sul">
<collectingMunicipality box="[272,360,320,342]" pageId="25" pageNumber="998">Aceguá</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53F4D0C4BD977FEA5" box="[373,597,320,342]" country="Brazil" latitude="-31.455393" longLatPrecision="1" longitude="-54.352863" municipality="Acegua" name="Fazenda Sentinela" pageId="25" pageNumber="998" stateProvince="Rio Grande do Sul">Fazenda Sentinela</location>
,
<geoCoordinate box="[610,774,320,341]" degrees="31" direction="south" minutes="27" orientation="latitude" pageId="25" pageNumber="998" precision="1" seconds="19.41" value="-31.455393">31°2719.41”S</geoCoordinate>
,
<geoCoordinate box="[163,340,350,372]" degrees="54" direction="west" minutes="21" orientation="longitude" pageId="25" pageNumber="998" precision="1" seconds="10.32" value="-54.352863">54°2110.32”W</geoCoordinate>
,
<date box="[355,532,351,372]" pageId="25" pageNumber="998" value="2011-11-08" valueMax="2011-11-14" valueMin="2011-11-08">
<collectingDate box="[355,532,351,372]" pageId="25" pageNumber="998" value="2011-11-08" valueMax="2011-11-14" valueMin="2011-11-08">0814.XI.2011</collectingDate>
</date>
(
<collectingMethod box="[551,627,350,372]" pageId="25" pageNumber="998">pitfall</collectingMethod>
baited with human and pig excrement),
<collectorName box="[500,658,382,403]" pageId="25" pageNumber="998">R. M. Moraes</collectorName>
leg.
<specimenCount pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[258,340,412,434]" pageId="25" pageNumber="998">CEMT</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841391" collectingDate="1945-01" collectionCode="MGPA" country="Brazil" location="Guarai" pageId="25" pageNumber="998" specimenCount="1" specimenCount-male="1" stateProvince="Rio Grande do Sul">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53F500C97DA9FFE41" box="[360,445,412,434]" country="Brazil" name="Guarai" pageId="25" pageNumber="998" stateProvince="Rio Grande do Sul">Guaraí</location>
,
<date box="[459,536,412,434]" pageId="25" pageNumber="998" value="1945-01">
<collectingDate box="[459,536,412,434]" pageId="25" pageNumber="998" value="1945-01">I.1945</collectingDate>
</date>
, no collector
<specimenCount pageId="25" pageNumber="998" type="male">three males</specimenCount>
(
<collectionCode box="[243,322,443,465]" pageId="25" pageNumber="998">MGPA</collectionCode>
)
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3704841362" box="[334,580,442,465]" collectionCode="MZSP" country="Brazil" pageId="25" pageNumber="998" specimenCount="1" specimenCount-male="1" stateProvince="Rio Grande do Sul">
and
<specimenCount box="[382,484,442,464]" pageId="25" pageNumber="998" type="male">one male</specimenCount>
(
<collectionCode box="[497,573,443,465]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="25" pageNumber="998" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841306" collectionCode="MNHN" country="Paraguay" location="PARAGUAY" pageId="25" pageNumber="998" specimenCount="1" specimenCount-male="1">
<emphasis bold="true" box="[588,746,442,464]" pageId="25" pageNumber="998">
<collectingCountry box="[588,741,442,464]" name="Paraguay" pageId="25" pageNumber="998">PARAGUAY</collectingCountry>
:
</emphasis>
no further data
<specimenCount box="[338,444,473,495]" pageId="25" pageNumber="998" type="male">one male</specimenCount>
(
<collectionCode box="[460,550,473,495]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="998">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841477" collectingDate="1925-10-28" collectionCode="MACN" collectorName="F. Felippone" country="Paraguay" location="Chaco" pageId="25" pageNumber="998" specimenCount="1" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53C020CD2D9A6FE1C" box="[570,644,473,495]" country="Paraguay" name="Chaco" pageId="25" pageNumber="998">Chaco</location>
,
<date box="[655,774,473,495]" pageId="25" pageNumber="998" value="1925-10-28">
<collectingDate box="[655,774,473,495]" pageId="25" pageNumber="998" value="1925-10-28">28.X.1925</collectingDate>
</date>
,
<collectorName box="[163,314,504,526]" pageId="25" pageNumber="998">F. Felippone</collectorName>
leg.
<specimenCount box="[401,527,504,526]" pageId="25" pageNumber="998" type="male">two males</specimenCount>
(
<collectionCode box="[545,634,504,526]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="25" pageNumber="998" type="Museum">MACN</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841333" collectingDate="1928-02" collectionCode="CEMT" collectorName="F. Schada" country="Paraguay" location="Nape'" pageId="25" pageNumber="998" specimenCount="1" specimenCount-female="1">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53CAC0CF3D9FEFDFD" box="[660,732,504,526]" country="Paraguay" name="Nape'" pageId="25" pageNumber="998">Napé</location>
(?),
<date box="[163,251,535,556]" pageId="25" pageNumber="998" value="1928-02">
<collectingDate box="[163,251,535,556]" pageId="25" pageNumber="998" value="1928-02">II.1928</collectingDate>
</date>
,
<collectorName box="[264,379,534,556]" pageId="25" pageNumber="998">F. Schada</collectorName>
leg.
<specimenCount box="[460,590,534,556]" pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[606,689,535,557]" pageId="25" pageNumber="998">CEMT</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841379" collectingDate="1949-10" collectionCode="MACN" collectorName="Giai" country="Paraguay" location="ALTO PARAGUAY" pageId="25" pageNumber="998" specimenCount="1" specimenCount-male="1" stateProvince="Alto Paraguay">
<emphasis italics="true" pageId="25" pageNumber="998">
<collectingRegion country="Paraguay" name="Alto Paraguay" pageId="25" pageNumber="998">ALTO PARAGUAY</collectingRegion>
:
</emphasis>
no specific locality,
<date box="[531,613,565,586]" pageId="25" pageNumber="998" value="1949-10">
<collectingDate box="[531,613,565,586]" pageId="25" pageNumber="998" value="1949-10">X.1949</collectingDate>
</date>
, A.
<collectorName box="[652,701,565,587]" pageId="25" pageNumber="998">Giai</collectorName>
(?) leg.
<specimenCount box="[184,289,596,618]" pageId="25" pageNumber="998" type="male">one male</specimenCount>
(
<collectionCode box="[304,389,596,618]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="25" pageNumber="998" type="Museum">MACN</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841465" collectingDate="1950-11" collectionCode="MACN" collectorName="Giai" country="Paraguay" location="Puerto Casado" pageId="25" pageNumber="998" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Alto Paraguay">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53FA00F5FD966FD99" box="[408,580,596,618]" country="Paraguay" name="Puerto Casado" pageId="25" pageNumber="998" stateProvince="Alto Paraguay">Puerto Casado</location>
,
<date box="[593,686,596,618]" pageId="25" pageNumber="998" value="1950-11">
<collectingDate box="[593,686,596,618]" pageId="25" pageNumber="998" value="1950-11">XI.1950</collectingDate>
</date>
, A.
<collectorName box="[729,778,596,618]" pageId="25" pageNumber="998">Giai</collectorName>
(?) leg.
<specimenCount box="[267,384,626,648]" pageId="25" pageNumber="998" type="male">two males</specimenCount>
and
<specimenCount box="[444,582,626,648]" pageId="25" pageNumber="998" type="female">two females</specimenCount>
(
<collectionCode box="[597,681,627,649]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="25" pageNumber="998" type="Museum">MACN</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841349" collectingDate="2006-12-04" collectingDateMax="2006-12-09" collectingDateMin="2006-12-04" collectionCode="TAMU" collectorName="B. Garcete &amp; W. D. Edmonds" country="Paraguay" elevation="78" latitude="-22.294443" location="Pto. La Victoria'" longLatPrecision="21" longitude="-57.9375" municipality="Puerto Casado" pageId="25" pageNumber="998" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Alto Paraguay">
<collectingMunicipality pageId="25" pageNumber="998">Puerto Casado</collectingMunicipality>
(1.7 km SE
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53FA60F99D949FD54" box="[414,619,657,679]" country="Paraguay" latitude="-22.294443" longLatPrecision="21" longitude="-57.9375" municipality="Puerto Casado" name="Pto. La Victoria'" pageId="25" pageNumber="998" stateProvince="Alto Paraguay">Pto. La Victoria</location>
),
<geoCoordinate box="[641,773,657,678]" degrees="22" direction="south" minutes="17" orientation="latitude" pageId="25" pageNumber="998" precision="15" seconds="40" value="-22.294443">22°1740”S</geoCoordinate>
,
<geoCoordinate box="[163,299,688,710]" degrees="57" direction="west" minutes="56" orientation="longitude" pageId="25" pageNumber="998" precision="15" seconds="15" value="-57.9375">57°5615”W</geoCoordinate>
,
<quantity box="[311,372,688,709]" metricMagnitude="1" metricUnit="m" metricValue="7.8" pageId="25" pageNumber="998" unit="m" value="78.0">
<elevation box="[311,372,688,709]" metricMagnitude="1" metricUnit="m" metricValue="7.8" pageId="25" pageNumber="998" unit="m" value="78.0">78 m</elevation>
</quantity>
,
<date box="[385,570,688,710]" pageId="25" pageNumber="998" value="2006-12-04" valueMax="2006-12-09" valueMin="2006-12-04">
<collectingDate box="[385,570,688,710]" pageId="25" pageNumber="998" value="2006-12-04" valueMax="2006-12-09" valueMin="2006-12-04">0409.XII.2006</collectingDate>
</date>
,
<collectorName box="[582,708,688,709]" pageId="25" pageNumber="998">B. Garcete</collectorName>
leg.
<specimenCount box="[163,266,718,740]" pageId="25" pageNumber="998" type="male">one male</specimenCount>
and
<specimenCount box="[325,450,718,740]" pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[464,543,719,740]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="25" pageNumber="998" type="Herbarium">TAMU</collectionCode>
ex
<collectorName box="[604,778,718,740]" pageId="25" pageNumber="998">W. D. Edmonds</collectorName>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841369" collectingDate="2006-12-02" collectionCode="TAMU" collectorName="W. D. Edmonds" country="Paraguay" location="Puerto Esperanza" pageId="25" pageNumber="998" specimenCount="1" specimenCount-female="1" stateProvince="Alto Paraguay">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53F060FE5D90FFCF0" box="[318,557,750,771]" country="Paraguay" name="Puerto Esperanza" pageId="25" pageNumber="998" stateProvince="Alto Paraguay">Puerto Esperanza</location>
,
<date box="[575,731,749,771]" pageId="25" pageNumber="998" value="2006-12-02">
<collectingDate box="[575,731,749,771]" pageId="25" pageNumber="998" value="2006-12-02">02.XII.2006</collectingDate>
</date>
, no collector
<specimenCount box="[297,428,780,802]" pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[445,527,780,801]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="25" pageNumber="998" type="Herbarium">TAMU</collectionCode>
ex
<collectorName box="[595,778,780,802]" pageId="25" pageNumber="998">W. D. Edmonds</collectorName>
collection)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841506" collectionCode="SMTD" country="Paraguay" location="Paraguay River" pageId="25" pageNumber="998" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Concepcion">
<emphasis box="[305,499,809,833]" italics="true" pageId="25" pageNumber="998">
<collectingRegion box="[305,492,809,833]" country="Paraguay" name="Concepcion" pageId="25" pageNumber="998">CONCEPCIÓN</collectingRegion>
:
</emphasis>
<collectorName box="[509,656,811,832]" pageId="25" pageNumber="998">Puerto Max</collectorName>
,
<collectorName pageId="25" pageNumber="998">Estáncia Postillon</collectorName>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53F2C0E41DAE5FCAC" box="[276,455,841,863]" country="Paraguay" name="Paraguay River" pageId="25" pageNumber="998" stateProvince="Concepcion">Paraguay River</location>
, no further data
<specimenCount box="[663,778,841,863]" pageId="25" pageNumber="998" type="male">two males</specimenCount>
and
<specimenCount box="[214,373,872,894]" pageId="25" pageNumber="998" type="female">three females</specimenCount>
(
<collectionCode box="[388,465,872,893]" pageId="25" pageNumber="998">SMTD</collectionCode>
ex
<collectorName box="[528,646,872,894]" pageId="25" pageNumber="998">C. Felsche</collectorName>
collection)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841352" collectingDate="1926-02" collectionCode="RMNH" collectorName="Valck Lucassen" country="Paraguay" location="Itape" pageId="25" pageNumber="998" specimenCount="1" specimenCount-female="1" stateProvince="Guaira">
<emphasis box="[163,277,901,925]" italics="true" pageId="25" pageNumber="998">
<collectingRegion box="[163,271,901,925]" country="Paraguay" name="Guaira" pageId="25" pageNumber="998">GUAIRÁ</collectingRegion>
:
</emphasis>
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53F270E8DDA43FC6F" box="[287,353,902,924]" country="Paraguay" name="Itape" pageId="25" pageNumber="998" stateProvince="Guaira">Itapé</location>
,
<date box="[367,457,902,924]" pageId="25" pageNumber="998" value="1926-02">
<collectingDate box="[367,457,902,924]" pageId="25" pageNumber="998" value="1926-02">II.1926</collectingDate>
</date>
, no collector
<specimenCount box="[646,777,902,924]" pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[170,255,933,954]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="25" pageNumber="998" type="Museum">RMNH</collectionCode>
ex
<collectorName box="[320,505,933,955]" pageId="25" pageNumber="998">Valck Lucassen</collectorName>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841494" collectingDate="1946-09-27" collectionCode="BDGC" collectorName="Schade " country="Paraguay" location="Mbocayaty del Guaira" pageId="25" pageNumber="998" specimenCount="1" specimenCount-male="1" stateProvince="Guaira">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53CB20EAEDA3FFC29" country="Paraguay" name="Mbocayaty del Guaira" pageId="25" pageNumber="998" stateProvince="Guaira">Mbocayaty del Guairá</location>
(
<emphasis box="[305,468,964,985]" italics="true" pageId="25" pageNumber="998">MBUCAYATY</emphasis>
),
<date box="[493,621,964,986]" pageId="25" pageNumber="998" value="1946-09-27">
<collectingDate box="[493,621,964,986]" pageId="25" pageNumber="998" value="1946-09-27">27.IX.1946</collectingDate>
</date>
,
<collectorName box="[631,714,964,986]" pageId="25" pageNumber="998">Schade</collectorName>
leg.
<specimenCount box="[163,266,994,1016]" pageId="25" pageNumber="998" type="male">one male</specimenCount>
(
<collectionCode box="[281,357,994,1015]" pageId="25" pageNumber="998">BDGC</collectionCode>
ex
<collectorName box="[416,553,994,1016]" pageId="25" pageNumber="998">A. Martínez</collectorName>
and
<collectorName box="[611,684,995,1016]" pageId="25" pageNumber="998">Henry</collectorName>
&amp;
<collectorName pageId="25" pageNumber="998">Anne Howden</collectorName>
collections)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841534" collectionCode="MNHN" collectorName="Villarrica" country="Paraguay" pageId="25" pageNumber="998" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Guaira">
<collectorName box="[407,520,1025,1047]" pageId="25" pageNumber="998">Villarrica</collectorName>
, no further data
<specimenCount pageId="25" pageNumber="998" type="male">two males</specimenCount>
and
<specimenCount box="[288,412,1056,1078]" pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[426,513,1056,1078]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="998">MNHN</collectionCode>
)
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3704841439" collectionCode="NHMB" country="Paraguay" location="Frey" pageId="25" pageNumber="998" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Guaira">
and
<specimenCount box="[577,680,1056,1078]" pageId="25" pageNumber="998" type="male">one male</specimenCount>
and
<specimenCount pageId="25" pageNumber="998" type="female">two females</specimenCount>
(
<collectionCode box="[269,354,1086,1107]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="25" pageNumber="998">NHMB</collectionCode>
ex
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53F9B0934DAFBFBA7" box="[419,473,1087,1108]" country="Paraguay" name="Frey" pageId="25" pageNumber="998" stateProvince="Guaira">Frey</location>
collection)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841394" collectingDate="1948-10" collectionCode="CMNC" collectorName="F. H. Schade" country="Paraguay" county="San Cosme" location="Cosme'" municipality="Damian" pageId="25" pageNumber="998" specimenCount="1" specimenCount-male="1" stateProvince="Itapua">
<emphasis box="[617,725,1085,1109]" italics="true" pageId="25" pageNumber="998">
<collectingRegion box="[617,719,1085,1109]" country="Paraguay" name="Itapua" pageId="25" pageNumber="998">ITAPUÁ</collectingRegion>
:
</emphasis>
<collectingCounty pageId="25" pageNumber="998">San Cosme</collectingCounty>
y
<collectingMunicipality box="[265,358,1117,1139]" pageId="25" pageNumber="998">Damián</collectingMunicipality>
(
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53F4F0956DAECFB80" box="[375,462,1117,1139]" country="Paraguay" county="San Cosme" municipality="Damian" name="Cosme'" pageId="25" pageNumber="998" stateProvince="Itapua">Cosme</location>
),
<date box="[480,562,1117,1138]" pageId="25" pageNumber="998" value="1948-10">
<collectingDate box="[480,562,1117,1138]" pageId="25" pageNumber="998" value="1948-10">X.1948</collectingDate>
</date>
,
<collectorName box="[572,714,1117,1139]" pageId="25" pageNumber="998">F. H. Schade</collectorName>
leg.
<specimenCount box="[163,268,1148,1170]" pageId="25" pageNumber="998" type="male">one male</specimenCount>
(
<collectionCode box="[284,369,1148,1170]" pageId="25" pageNumber="998">CMNC</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841383" collectingDate="1952-11" collectionCode="FSCA" collectorName="E. N. Kellesvig-Waering" country="Paraguay" location="PARAGUARI" pageId="25" pageNumber="998" specimenCount="1" specimenCount-female="1" stateProvince="Paraguari">
<emphasis box="[388,551,1146,1170]" italics="true" pageId="25" pageNumber="998">
<collectingRegion box="[388,545,1146,1170]" country="Paraguay" name="Paraguari" pageId="25" pageNumber="998">PARAGUARÍ</collectingRegion>
:
</emphasis>
no specific locality,
<date box="[163,257,1178,1200]" pageId="25" pageNumber="998" value="1952-11">
<collectingDate box="[163,257,1178,1200]" pageId="25" pageNumber="998" value="1952-11">XI.1952</collectingDate>
</date>
, no collector
<specimenCount box="[439,567,1178,1200]" pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[584,655,1178,1199]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="25" pageNumber="998" type="Museum">FSCA</collectionCode>
ex
<collectorName pageId="25" pageNumber="998">E. N. Kellesvig-Waering</collectorName>
collection)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841435" collectingDate="2003-01-05" collectionCode="TAMU" collectorName="C. Aguilar &amp; W. D. Edmonds" country="Paraguay" location="Pioneros" municipality="Cruce" pageId="25" pageNumber="998" specimenCount="1" specimenCount-female="1" stateProvince="PRESIDENTE HAYAS">
<emphasis box="[508,779,1209,1230]" italics="true" pageId="25" pageNumber="998">
<collectingRegion box="[508,773,1209,1230]" country="Paraguay" name="PRESIDENTE HAYAS" pageId="25" pageNumber="998">PRESIDENTE HAYAS</collectingRegion>
:
</emphasis>
<collectingMunicipality box="[163,234,1240,1261]" pageId="25" pageNumber="998">Cruce</collectingMunicipality>
de los
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53F7D09D3DA93FB1D" box="[325,433,1240,1262]" country="Paraguay" municipality="Cruce" name="Pioneros" pageId="25" pageNumber="998" stateProvince="PRESIDENTE HAYAS">Pioneros</location>
,
<date box="[447,563,1239,1261]" pageId="25" pageNumber="998" value="2003-01-05">
<collectingDate box="[447,563,1239,1261]" pageId="25" pageNumber="998" value="2003-01-05">05.I.2003</collectingDate>
</date>
,
<collectorName box="[578,703,1240,1262]" pageId="25" pageNumber="998">C. Aguilar</collectorName>
leg.
<specimenCount box="[163,293,1270,1292]" pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[310,391,1270,1291]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="25" pageNumber="998" type="Herbarium">TAMU</collectionCode>
ex
<collectorName box="[458,640,1270,1292]" pageId="25" pageNumber="998">W. D. Edmonds</collectorName>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841524" collectingDate="2003-12-20" collectionCode="TAMU" collectorName="C. Aguilar &amp; W. D. Edmonds" country="Paraguay" location="Pioneros" municipality="Cruce" pageId="25" pageNumber="998" specimenCount="1" specimenCount-female="1" stateProvince="PRESIDENTE HAYAS">
<collectingMunicipality box="[163,232,1301,1322]" pageId="25" pageNumber="998">Cruce</collectingMunicipality>
de los
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53F05081EDA87FAD8" box="[317,421,1301,1323]" country="Paraguay" municipality="Cruce" name="Pioneros" pageId="25" pageNumber="998" stateProvince="PRESIDENTE HAYAS">Pioneros</location>
,
<date box="[434,575,1301,1322]" pageId="25" pageNumber="998" value="2003-12-20">
<collectingDate box="[434,575,1301,1322]" pageId="25" pageNumber="998" value="2003-12-20">20.XII.2003</collectingDate>
</date>
,
<collectorName box="[587,708,1301,1323]" pageId="25" pageNumber="998">C. Aguilar</collectorName>
leg.
<specimenCount box="[163,293,1332,1354]" pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[310,391,1332,1353]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="25" pageNumber="998" type="Herbarium">TAMU</collectionCode>
ex
<collectorName box="[458,640,1332,1354]" pageId="25" pageNumber="998">W. D. Edmonds</collectorName>
collection)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841511" collectionCode="MNHN" country="Uruguay" location="URUGUAY" pageId="25" pageNumber="998" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<emphasis bold="true" box="[163,310,1362,1384]" pageId="25" pageNumber="998">
<collectingCountry box="[163,305,1362,1384]" name="Uruguay" pageId="25" pageNumber="998">URUGUAY</collectingCountry>
:
</emphasis>
no further data
<specimenCount box="[534,655,1362,1384]" pageId="25" pageNumber="998" type="male">two males</specimenCount>
and
<specimenCount pageId="25" pageNumber="998" type="female">three females</specimenCount>
(
<collectionCode box="[271,360,1393,1415]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="998">MNHN</collectionCode>
)
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3704841480" box="[377,640,1393,1415]" collectionCode="MZSP" country="Uruguay" pageId="25" pageNumber="998" specimenCount="1" specimenCount-male="1">
and
<specimenCount box="[430,538,1393,1415]" pageId="25" pageNumber="998" type="male">one male</specimenCount>
(
<collectionCode box="[555,632,1393,1415]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="25" pageNumber="998" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841467" collectionCode="NHMB" collectorName="Fruhstorfer" country="Uruguay" location="Frey" pageId="25" pageNumber="998" specimenCount="1" specimenCount-female="1">
no specific locality or date,
<collectorName box="[348,484,1424,1446]" pageId="25" pageNumber="998">Fruhstorfer</collectorName>
leg.
<specimenCount box="[556,681,1424,1446]" pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[695,778,1424,1445]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="25" pageNumber="998">NHMB</collectionCode>
ex
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53EEF08A4DA2EFA37" box="[215,268,1455,1476]" country="Uruguay" name="Frey" pageId="25" pageNumber="998">Frey</location>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841329" collectionCode="RBINS" collectorName="Parry" country="Uruguay" location="La Fontaine" pageId="25" pageNumber="998" specimenCount="1" specimenCount-female="1">
no specific locality or date,
<collectorName box="[713,778,1455,1476]" pageId="25" pageNumber="998">Parry</collectorName>
leg.
<specimenCount box="[230,356,1485,1507]" pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[370,453,1485,1506]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="25" pageNumber="998" type="Museum">RBINS</collectionCode>
J.
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53FC308C6D9AAFA10" box="[507,648,1485,1507]" country="Uruguay" name="La Fontaine" pageId="25" pageNumber="998">La Fontaine</location>
collection)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841416" collectingDate="1954-12-24" collectionCode="CNCI" collectorName="C. S. Carbonell" country="Uruguay" location="Empalme Rutas" pageId="25" pageNumber="998" specimenCount="1" specimenCount-male="1" stateProvince="Artigas">
<emphasis box="[163,287,1516,1537]" italics="true" pageId="25" pageNumber="998">
<collectingRegion box="[163,281,1516,1537]" country="Uruguay" name="Artigas" pageId="25" pageNumber="998">ARTIGAS</collectingRegion>
:
</emphasis>
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53F1208E7DAC5F9F2" box="[298,487,1516,1538]" country="Uruguay" name="Empalme Rutas" pageId="25" pageNumber="998" stateProvince="Artigas">Empalme Rutas</location>
30 y 3,
<date box="[584,722,1515,1537]" pageId="25" pageNumber="998" value="1954-12-24">
<collectingDate box="[584,722,1515,1537]" pageId="25" pageNumber="998" value="1954-12-24">24.XII.1954</collectingDate>
</date>
,
<collectorName pageId="25" pageNumber="998">C.S. Carbonell</collectorName>
leg.
<specimenCount box="[351,468,1546,1568]" pageId="25" pageNumber="998" type="male">two males</specimenCount>
(
<collectionCode box="[483,554,1546,1568]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="25" pageNumber="998">CNCI</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841446" collectionCode="MLPA" country="Uruguay" location="CANELONES" pageId="25" pageNumber="998" specimenCount="1" specimenCount-male="1" stateProvince="Canelones">
<emphasis box="[572,744,1547,1568]" italics="true" pageId="25" pageNumber="998">
<collectingRegion box="[572,738,1547,1568]" country="Uruguay" name="Canelones" pageId="25" pageNumber="998">CANELONES</collectingRegion>
:
</emphasis>
no further data
<specimenCount box="[335,440,1577,1599]" pageId="25" pageNumber="998" type="male">one male</specimenCount>
(
<collectionCode box="[455,533,1577,1599]" pageId="25" pageNumber="998">MLPA</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841353" collectingDate="1907-02-19" collectionCode="MLPA" collectorName="Hermano Mario" country="Uruguay" location="Los Brejos'" municipality="Canelon Chico" pageId="25" pageNumber="998" specimenCount="1" specimenCount-male="1" stateProvince="Canelones">
<collectingMunicipality box="[552,722,1577,1599]" pageId="25" pageNumber="998">Canelón Chico</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53CD90B22DBD5F9AD" country="Uruguay" municipality="Canelon Chico" name="Los Brejos'" pageId="25" pageNumber="998" stateProvince="Canelones">Los Brejos</location>
,
<date box="[260,384,1608,1629]" pageId="25" pageNumber="998" value="1907-02-19">
<collectingDate box="[260,384,1608,1629]" pageId="25" pageNumber="998" value="1907-02-19">19.II.1907</collectingDate>
</date>
,
<collectorName box="[398,589,1608,1630]" pageId="25" pageNumber="998">Hermano Mario</collectorName>
leg.
<specimenCount box="[671,778,1608,1630]" pageId="25" pageNumber="998" type="male">one male</specimenCount>
(
<collectionCode box="[170,251,1638,1660]" pageId="25" pageNumber="998">MLPA</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841491" collectionCode="MLPA" collectorName="Canelones &amp; Juanico" country="Uruguay" pageId="25" pageNumber="998" specimenCount="1" specimenCount-female="1" stateProvince="Canelones">
<collectorName box="[271,397,1638,1660]" pageId="25" pageNumber="998">Canelones</collectorName>
,
<collectorName box="[410,505,1638,1660]" pageId="25" pageNumber="998">Juanicó</collectorName>
, no further data
<specimenCount pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[256,335,1669,1691]" pageId="25" pageNumber="998">MLPA</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841410" collectionCode="MACN" collectorName="J. G. Hosmann" country="Uruguay" location="CERRO LARGO" pageId="25" pageNumber="998" specimenCount="1" stateProvince="Cerro Largo">
<emphasis box="[354,555,1669,1690]" italics="true" pageId="25" pageNumber="998">
<collectingRegion box="[354,548,1669,1690]" country="Uruguay" name="Cerro Largo" pageId="25" pageNumber="998">CERRO LARGO</collectingRegion>
:
</emphasis>
no specific locality nor date,
<collectorName box="[273,444,1700,1721]" pageId="25" pageNumber="998">J. G. Hosmann</collectorName>
leg. one unsexed specimen (
<collectionCode box="[170,255,1730,1752]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="25" pageNumber="998" type="Museum">MACN</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841335" collectionCode="MACN" country="Uruguay" location="COLONIA" pageId="25" pageNumber="998" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Colonia">
<emphasis box="[272,401,1731,1752]" italics="true" pageId="25" pageNumber="998">
<collectingRegion box="[272,395,1731,1752]" country="Uruguay" name="Colonia" pageId="25" pageNumber="998">COLONIA</collectingRegion>
:
</emphasis>
no further data
<specimenCount box="[611,728,1730,1752]" pageId="25" pageNumber="998" type="male">five males</specimenCount>
and
<specimenCount box="[163,288,1761,1783]" pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[302,387,1761,1783]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="25" pageNumber="998" type="Museum">MACN</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841557" collectionCode="BMNH, MNHN" collectorName="Nevinson" country="Uruguay" location="MONTEVIDEO" pageId="25" pageNumber="998" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Montevideo">
<emphasis box="[404,594,1761,1782]" italics="true" pageId="25" pageNumber="998">
<collectingRegion box="[404,588,1761,1782]" country="Uruguay" name="Montevideo" pageId="25" pageNumber="998">MONTEVIDEO</collectingRegion>
:
</emphasis>
no further data
<specimenCount box="[185,306,1791,1813]" pageId="25" pageNumber="998" type="male">two males</specimenCount>
(
<collectionCode box="[322,407,1792,1813]" pageId="25" pageNumber="998">BMNH</collectionCode>
ex
<collectorName box="[473,585,1791,1813]" pageId="25" pageNumber="998">Nevinson</collectorName>
collection),
<specimenCount pageId="25" pageNumber="998" type="male">four males</specimenCount>
and
<specimenCount box="[290,448,1822,1844]" pageId="25" pageNumber="998" type="female">three females</specimenCount>
(
<collectionCode box="[464,553,1822,1844]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="998">MNHN</collectionCode>
)
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3704841502" box="[572,775,1822,1844]" collectionCode="MZSP" collectorName="Nevinson" country="Uruguay" isTailEnumeration="true" location="MONTEVIDEO" pageId="25" pageNumber="998" specimenCount="1" specimenCount-male="1" stateProvince="Montevideo">
<specimenCount box="[572,677,1822,1844]" pageId="25" pageNumber="998" type="male">one male</specimenCount>
(
<collectionCode box="[692,767,1822,1844]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="25" pageNumber="998" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3704841365" collectionCode="RBINS, SMTD" country="Uruguay" location="Felsche" pageId="25" pageNumber="998" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Montevideo">
<specimenCount box="[163,280,1853,1875]" pageId="25" pageNumber="998" type="male">one male</specimenCount>
(dissected) (
<collectionCode box="[454,544,1853,1874]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="25" pageNumber="998" type="Museum">RBINS</collectionCode>
ex A.
<collectorName box="[662,776,1853,1874]" pageId="25" pageNumber="998">Janssens</collectorName>
collection),
<specimenCount box="[967,1099,197,219]" pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[1117,1203,197,218]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="25" pageNumber="998" type="Museum">RBINS</collectionCode>
ex
<collectorName box="[1272,1442,197,219]" pageId="25" pageNumber="998">P. de Moffarts</collectorName>
collection),
<specimenCount box="[971,1083,227,249]" pageId="25" pageNumber="998" type="male">one male</specimenCount>
(dissected) and
<specimenCount box="[1294,1442,227,249]" pageId="25" pageNumber="998" type="female">two females</specimenCount>
(
<collectionCode box="[834,921,258,279]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="25" pageNumber="998" type="Museum">RBINS</collectionCode>
ex
<collectorName box="[991,1118,258,280]" pageId="25" pageNumber="998">J. J. Gillet</collectorName>
collection) and
<specimenCount box="[1318,1442,258,280]" pageId="25" pageNumber="998" type="male">two males</specimenCount>
(
<collectionCode box="[834,910,289,310]" pageId="25" pageNumber="998">SMTD</collectionCode>
ex C.
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53DD40C2ADF60FEC4" box="[1004,1090,289,311]" country="Uruguay" name="Felsche" pageId="25" pageNumber="998" stateProvince="Montevideo">Felsche</location>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841440" collectionCode="MACN" collectorName="J. G. Hosmann" country="Uruguay" pageId="25" pageNumber="998" specimenCount="1" specimenCount-female="1" stateProvince="Montevideo">
no specific locality or date,
<collectorName box="[930,1109,320,341]" pageId="25" pageNumber="998">J. G. Hosmann</collectorName>
leg.
<specimenCount box="[1194,1326,319,341]" pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[1343,1430,320,342]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="25" pageNumber="998" type="Museum">MACN</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841513" collectingDate="1883-02" collectionCode="RBINS" collectorName="Le Moult" country="Uruguay" pageId="25" pageNumber="998" specimenCount="1" specimenCount-female="1" stateProvince="Montevideo">
no specific locality,
<date box="[1054,1140,350,371]" pageId="25" pageNumber="998" value="1883-02">
<collectingDate box="[1054,1140,350,371]" pageId="25" pageNumber="998" value="1883-02">II.1883</collectingDate>
</date>
, no collector
<specimenCount box="[1316,1443,350,372]" pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[834,918,381,402]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="25" pageNumber="998" type="Museum">RBINS</collectionCode>
ex
<collectorName box="[981,1087,381,403]" pageId="25" pageNumber="998">Le Moult</collectorName>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841345" collectionCode="MHNG" collectorName="Zufferey" country="Uruguay" pageId="25" pageNumber="998" specimenCount="1" specimenCount-female="1" stateProvince="Montevideo">
no specific locality or date,
<collectorName box="[937,1043,411,433]" pageId="25" pageNumber="998">Zufferey</collectorName>
leg.
<specimenCount box="[1136,1275,411,433]" pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[1295,1389,412,434]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="25" pageNumber="998" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841539" collectingDate="1907-12-25" collectionCode="RBINS" collectorName="Gillet" country="Uruguay" pageId="25" pageNumber="998" specimenCount="1" specimenCount-female="1" stateProvince="Montevideo">
no specific locality,
<date box="[1013,1151,442,464]" pageId="25" pageNumber="998" value="1907-12-25">
<collectingDate box="[1013,1151,442,464]" pageId="25" pageNumber="998" value="1907-12-25">25.XII.1907</collectingDate>
</date>
, no collector
<specimenCount box="[1319,1442,442,464]" pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[834,917,473,494]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="25" pageNumber="998" type="Museum">RBINS</collectionCode>
ex J. J.
<collectorName box="[1026,1092,473,495]" pageId="25" pageNumber="998">Gillet</collectorName>
collection)
</materialsCitation>
; no specific locality,
<date box="[827,972,503,525]" pageId="25" pageNumber="998" value="1909-12-25">25.XII.1909</date>
, no collector
<specimenCount box="[1163,1299,503,525]" pageId="25" pageNumber="998" type="female">one female</specimenCount>
;
<materialsCitation ID-GBIF-Occurrence="3704841357" collectingDate="1952-10" collectionCode="DZUP" collectorName="Alvarenga" country="Uruguay" pageId="25" pageNumber="998" specimenCount="1" specimenCount-male="1" stateProvince="Montevideo">
no specific locality,
<date box="[928,1012,534,556]" pageId="25" pageNumber="998" value="1952-10">
<collectingDate box="[928,1012,534,556]" pageId="25" pageNumber="998" value="1952-10">X.1952</collectingDate>
</date>
, no collector
<specimenCount box="[1197,1305,534,556]" pageId="25" pageNumber="998" type="male">one male</specimenCount>
(dissected) (
<collectionCode box="[834,907,565,586]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="25" pageNumber="998" type="University or college">DZUP</collectionCode>
ex M.
<collectorName box="[1002,1122,565,587]" pageId="25" pageNumber="998">Alvarenga</collectorName>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841575" collectionCode="NHMB" collectorName="Montevideo &amp; Lasseaux" country="Uruguay" location="Frey" pageId="25" pageNumber="998" specimenCount="1" specimenCount-male="1" stateProvince="Montevideo">
surroundings of
<collectorName box="[827,966,595,617]" pageId="25" pageNumber="998">Montevideo</collectorName>
, 1867,
<collectorName box="[1048,1159,596,617]" pageId="25" pageNumber="998">Lasseaux</collectorName>
leg.
<specimenCount box="[1237,1343,595,617]" pageId="25" pageNumber="998" type="male">one male</specimenCount>
(
<collectionCode box="[1359,1443,596,617]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="25" pageNumber="998">NHMB</collectionCode>
ex
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53D490F78D884FD7B" box="[881,934,627,648]" country="Uruguay" name="Frey" pageId="25" pageNumber="998" stateProvince="Montevideo">Frey</location>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841373" collectingDate="1932-10-12" collectionCode="BDGC" country="Uruguay" location="Cerro de Montevideo" pageId="25" pageNumber="998" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Montevideo">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53A090F79DE01FD7B" box="[1073,1315,626,648]" country="Uruguay" name="Cerro de Montevideo" pageId="25" pageNumber="998" stateProvince="Montevideo">Cerro de Montevideo</location>
,
<date box="[1327,1443,626,647]" pageId="25" pageNumber="998" value="1932-10-12">
<collectingDate box="[1327,1443,626,647]" pageId="25" pageNumber="998" value="1932-10-12">12.X.1932</collectingDate>
</date>
(on unspecified dung), no collector
<specimenCount box="[1243,1346,657,679]" pageId="25" pageNumber="998" type="male">one male</specimenCount>
and
<specimenCount pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[921,1001,688,710]" pageId="25" pageNumber="998">BDGC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841517" collectingDate="1932-10-12" collectionCode="MLPA" country="Uruguay" location="Cerro de Montevideo" pageId="25" pageNumber="998" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Montevideo">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53DC50FBBDFD7FD36" box="[1021,1269,687,709]" country="Uruguay" name="Cerro de Montevideo" pageId="25" pageNumber="998" stateProvince="Montevideo">Cerro de Montevideo</location>
,
<date box="[1281,1398,688,709]" pageId="25" pageNumber="998" value="1932-10-12">
<collectingDate box="[1281,1398,688,709]" pageId="25" pageNumber="998" value="1932-10-12">12.X.1932</collectingDate>
</date>
(on horse and cow dung), no collector
<specimenCount box="[1265,1388,718,740]" pageId="25" pageNumber="998" type="male">five males</specimenCount>
and
<specimenCount box="[827,969,749,771]" pageId="25" pageNumber="998" type="female">five females</specimenCount>
(
<collectionCode box="[985,1065,749,771]" pageId="25" pageNumber="998">MLPA</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841515" collectingDate="1935-01-18" collectionCode="RMNH" collectorName="Van Doesburg" country="Uruguay" location="Cerro" municipality="Cerro de Montevideo" pageId="25" pageNumber="998" specimenCount="1" specimenCount-male="1" stateProvince="Montevideo">
<collectingMunicipality box="[1085,1334,749,771]" pageId="25" pageNumber="998">Cerro de Montevideo</collectingMunicipality>
(
<emphasis box="[1356,1421,749,770]" italics="true" pageId="25" pageNumber="998">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53B740FE6DEAFFCF1" box="[1356,1421,749,770]" country="Uruguay" municipality="Cerro de Montevideo" name="Cerro" pageId="25" pageNumber="998" stateProvince="Montevideo">Cerro</location>
</emphasis>
),
<date box="[827,936,779,801]" pageId="25" pageNumber="998" value="1935-01-18">
<collectingDate box="[827,936,779,801]" pageId="25" pageNumber="998" value="1935-01-18">18.I.1935</collectingDate>
</date>
, no collector
<specimenCount box="[1106,1241,779,801]" pageId="25" pageNumber="998" type="male">three males</specimenCount>
(
<collectionCode box="[1255,1339,780,801]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="25" pageNumber="998" type="Museum">RMNH</collectionCode>
ex
<collectorName pageId="25" pageNumber="998">Van Doesburg</collectorName>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841404" collectingDate="1932-12-12" collectionCode="RBINS" collectorName="Le Moult" country="Uruguay" location="Pt Yeguas" municipality="Playa Punta Yeguas" pageId="25" pageNumber="998" specimenCount="1" specimenCount-female="1" stateProvince="Montevideo">
<collectingMunicipality box="[1069,1296,810,832]" pageId="25" pageNumber="998">Playa Punta Yeguas</collectingMunicipality>
(
<emphasis box="[1314,1423,811,832]" italics="true" pageId="25" pageNumber="998">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53B1A0E20DEADFCB3" box="[1314,1423,811,832]" country="Uruguay" municipality="Playa Punta Yeguas" name="Pt Yeguas" pageId="25" pageNumber="998" stateProvince="Montevideo">Pt Yeguas</location>
</emphasis>
),
<date box="[827,964,841,862]" pageId="25" pageNumber="998" value="1932-12-12">
<collectingDate box="[827,964,841,862]" pageId="25" pageNumber="998" value="1932-12-12">12.XII.1932</collectingDate>
</date>
, no collector
<specimenCount box="[1127,1263,841,863]" pageId="25" pageNumber="998" type="female">two females</specimenCount>
(
<collectionCode box="[1276,1359,841,862]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="25" pageNumber="998" type="Museum">RBINS</collectionCode>
ex
<collectorName pageId="25" pageNumber="998">Le Moult</collectorName>
collection)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841402" collectionCode="NMPC, R" country="Uruguay" location="Oberthur" pageId="25" pageNumber="998" specimenCount="1" specimenCount-male="1" stateProvince="Paysandu">
<emphasis box="[1036,1186,870,894]" italics="true" pageId="25" pageNumber="998">
<collectingRegion box="[1036,1181,870,894]" country="Uruguay" name="Paysandu" pageId="25" pageNumber="998">PAYSANDÚ</collectingRegion>
:
</emphasis>
no further data
<specimenCount pageId="25" pageNumber="998" type="male">one male</specimenCount>
(
<collectionCode box="[898,978,902,923]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="25" pageNumber="998">NMPC</collectionCode>
ex
<collectionCode box="[1041,1062,903,924]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="25" pageNumber="998">R</collectionCode>
.
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53A090E8DDFBCFC6F" box="[1073,1182,902,924]" country="Uruguay" name="Oberthur" pageId="25" pageNumber="998" stateProvince="Paysandu">Oberthür</location>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841370" collectingDate="1928-12-20" collectionCode="MLPA" country="Uruguay" county="Paysandu" location="Sacra'" municipality="Arroyo Sacra" pageId="25" pageNumber="998" specimenCount="1" specimenCount-male="1" stateProvince="Paysandu">
<collectingCounty box="[1323,1439,902,924]" pageId="25" pageNumber="998">Paysandú</collectingCounty>
,
<collectingMunicipality box="[827,979,933,954]" pageId="25" pageNumber="998">Arroyo Sacra</collectingMunicipality>
(
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53DDF0EAEDF13FC48" box="[999,1073,933,955]" country="Uruguay" county="Paysandu" municipality="Arroyo Sacra" name="Sacra'" pageId="25" pageNumber="998" stateProvince="Paysandu">Sacra</location>
),
<date box="[1093,1230,933,954]" pageId="25" pageNumber="998" value="1928-12-20">
<collectingDate box="[1093,1230,933,954]" pageId="25" pageNumber="998" value="1928-12-20">20.XII.1928</collectingDate>
</date>
, no collector
<specimenCount pageId="25" pageNumber="998" type="male">one male</specimenCount>
(
<collectionCode box="[899,978,963,985]" pageId="25" pageNumber="998">MLPA</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841483" collectingDate="1961-10-23" collectionCode="FSCA" collectorName="N. A. Monn &amp; E. N. Kellesvig-Waering" country="Uruguay" location="Arroyo Coronilla" municipality="Vichadero" pageId="25" pageNumber="998" specimenCount="1" specimenCount-male="1" stateProvince="Rivera">
<emphasis box="[996,1104,964,985]" italics="true" pageId="25" pageNumber="998">
<collectingRegion box="[996,1098,964,985]" country="Uruguay" name="Rivera" pageId="25" pageNumber="998">RIVERA</collectingRegion>
:
</emphasis>
<collectingMunicipality box="[1111,1230,963,985]" pageId="25" pageNumber="998">Vichadero</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53AE10ECFDEBCFC2A" box="[1241,1438,963,985]" country="Uruguay" municipality="Vichadero" name="Arroyo Coronilla" pageId="25" pageNumber="998" stateProvince="Rivera">Arroyo Coronilla</location>
,
<date box="[827,950,994,1016]" pageId="25" pageNumber="998" value="1961-10-23">
<collectingDate box="[827,950,994,1016]" pageId="25" pageNumber="998" value="1961-10-23">23.X.1961</collectingDate>
</date>
,
<collectorName box="[965,1104,994,1015]" pageId="25" pageNumber="998">N. A. Monn</collectorName>
leg
<specimenCount box="[1183,1329,994,1016]" pageId="25" pageNumber="998" type="male">three males</specimenCount>
(
<collectionCode box="[1347,1419,994,1015]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="25" pageNumber="998" type="Museum">FSCA</collectionCode>
ex
<collectorName box="[864,1161,1025,1047]" pageId="25" pageNumber="998">E. N. Kellesvig-Waering</collectorName>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841478" collectingDate="2011-10-03" collectingDateMax="2011-10-11" collectingDateMin="2011-10-03" collectionCode="CEMT" collectorName="R. M. Moraes" country="Uruguay" latitude="-31.684605" location="Santa" longLatPrecision="1" longitude="-54.623695" municipality="Vichadero" pageId="25" pageNumber="998" specimenCount="1" specimenCount-female="1" stateProvince="Rivera">
<collectingMunicipality box="[1313,1437,1025,1047]" pageId="25" pageNumber="998">Vichadero</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53D030914D8A2FBC7" box="[827,896,1055,1076]" country="Uruguay" latitude="-31.684605" longLatPrecision="1" longitude="-54.623695" municipality="Vichadero" name="Santa" pageId="25" pageNumber="998" stateProvince="Rivera">Santa</location>
Adelaida Farm,
<geoCoordinate box="[1093,1256,1055,1077]" degrees="31" direction="south" minutes="41" orientation="latitude" pageId="25" pageNumber="998" precision="1" seconds="04.58" value="-31.684605">31°4104.58”S</geoCoordinate>
,
<geoCoordinate box="[1269,1438,1055,1077]" degrees="54" direction="west" minutes="37" orientation="longitude" pageId="25" pageNumber="998" precision="1" seconds="25.31" value="-54.623695">54°3725.31”W</geoCoordinate>
,
<date box="[827,994,1086,1107]" pageId="25" pageNumber="998" value="2011-10-03" valueMax="2011-10-11" valueMin="2011-10-03">
<collectingDate box="[827,994,1086,1107]" pageId="25" pageNumber="998" value="2011-10-03" valueMax="2011-10-11" valueMin="2011-10-03">0311.X.2011</collectingDate>
</date>
(
<collectingMethod box="[1012,1089,1086,1108]" pageId="25" pageNumber="998">pitfall</collectingMethod>
baited with human and pig excrement),
<collectorName box="[972,1128,1117,1138]" pageId="25" pageNumber="998">R. M. Moraes</collectorName>
leg.
<specimenCount box="[1207,1334,1117,1139]" pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[1350,1431,1117,1139]" pageId="25" pageNumber="998">CEMT</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841492" collectingDate="1963-01-24" collectionCode="MZSP" collectorName="A. Toccheton" country="Uruguay" location="5 km NW Belen'" pageId="25" pageNumber="998" specimenCount="1" specimenCount-female="1" stateProvince="Salto">
<emphasis box="[827,922,1148,1169]" italics="true" pageId="25" pageNumber="998">
<collectingRegion box="[827,917,1148,1169]" country="Uruguay" name="Salto" pageId="25" pageNumber="998">SALTO</collectingRegion>
:
</emphasis>
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53D910970DF55FB62" box="[937,1143,1147,1169]" country="Uruguay" name="5 km NW Belen'" pageId="25" pageNumber="998" stateProvince="Salto">
<quantity box="[937,1000,1147,1169]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="998" unit="km" value="5.0">5 km</quantity>
NW Belén
</location>
,
<date box="[1157,1273,1147,1169]" pageId="25" pageNumber="998" value="1963-01-24">
<collectingDate box="[1157,1273,1147,1169]" pageId="25" pageNumber="998" value="1963-01-24">24.I.1963</collectingDate>
</date>
,
<collectorName box="[1287,1441,1147,1169]" pageId="25" pageNumber="998">A. Toccheton</collectorName>
leg.
<specimenCount box="[897,1025,1178,1200]" pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[1040,1117,1178,1200]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="25" pageNumber="998" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841443" collectingDate="1908-04" collectionCode="MLPA" country="Uruguay" location="Zapucay" pageId="25" pageNumber="998" specimenCount="1" specimenCount-female="1" stateProvince="Tacuarembo">
<emphasis box="[1136,1333,1177,1201]" italics="true" pageId="25" pageNumber="998">
<collectingRegion box="[1136,1326,1177,1201]" country="Uruguay" name="Tacuarembo" pageId="25" pageNumber="998">TACUAREMBÓ</collectingRegion>
:
</emphasis>
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53B050991DEBCFB5C" box="[1341,1438,1178,1199]" country="Uruguay" name="Zapucay" pageId="25" pageNumber="998" stateProvince="Tacuarembo">Zapucay</location>
,
<date box="[827,917,1209,1230]" pageId="25" pageNumber="998" value="1908-04">
<collectingDate box="[827,917,1209,1230]" pageId="25" pageNumber="998" value="1908-04">IV.1908</collectingDate>
</date>
, no collector
<specimenCount box="[1090,1216,1209,1231]" pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[1230,1309,1209,1231]" pageId="25" pageNumber="998">MLPA</collectionCode>
)
</materialsCitation>
.
<emphasis bold="true" pageId="25" pageNumber="998">
<materialsCitation ID-GBIF-Occurrence="3704841564" country="Uruguay" location="Probably" pageId="25" pageNumber="998" specimenCount="1" stateProvince="Tacuarembo">
<emphasis bold="true" italics="true" pageId="25" pageNumber="998">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53B1709B3DE80FB3D" box="[1327,1442,1208,1230]" country="Uruguay" name="Probably" pageId="25" pageNumber="998" stateProvince="Tacuarembo">Probably</location>
erroneous data
</emphasis>
</materialsCitation>
:
</emphasis>
<materialsCitation ID-GBIF-Occurrence="3704841562" collectionCode="MNHB" collectorName="H. Marchand" country="Argentina" location="TUCUMAN" pageId="25" pageNumber="998" specimenCount="1" specimenCount-male="1" stateProvince="Tucuman">
<emphasis bold="true" box="[1055,1240,1239,1261]" pageId="25" pageNumber="998">
<collectingCountry box="[1055,1235,1239,1261]" name="Argentina" pageId="25" pageNumber="998">ARGENTINA</collectingCountry>
:
</emphasis>
<emphasis box="[1252,1403,1238,1262]" italics="true" pageId="25" pageNumber="998">
<collectingRegion box="[1252,1397,1238,1262]" country="Argentina" name="Tucuman" pageId="25" pageNumber="998">TUCUMÁN</collectingRegion>
:
</emphasis>
no further data
<specimenCount box="[1007,1116,1270,1292]" pageId="25" pageNumber="998" type="male">one male</specimenCount>
(
<collectionCode box="[1132,1217,1270,1291]" pageId="25" pageNumber="998">MNHB</collectionCode>
ex
<collectorName box="[1285,1442,1270,1292]" pageId="25" pageNumber="998">H. Marchand</collectorName>
collection)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841463" collectionCode="MHNG" collectorName="Porto Alegre" country="Brazil" location="RIO GRANDE DO SUL" pageId="25" pageNumber="998" specimenCount="1" specimenCount-male="1" stateProvince="Rio Grande do Sul">
<emphasis bold="true" box="[984,1107,1300,1322]" pageId="25" pageNumber="998">
<collectingCountry box="[984,1098,1300,1321]" name="Brazil" pageId="25" pageNumber="998">BRAZIL</collectingCountry>
:
</emphasis>
<emphasis box="[1121,1443,1301,1322]" italics="true" pageId="25" pageNumber="998">
<collectingRegion box="[1121,1436,1301,1322]" country="Brazil" name="Rio Grande do Sul" pageId="25" pageNumber="998">RIO GRANDE DO SUL</collectingRegion>
:
</emphasis>
<collectorName box="[827,976,1331,1353]" pageId="25" pageNumber="998">Porto Alegre</collectorName>
, no further data
<specimenCount box="[1204,1324,1331,1353]" pageId="25" pageNumber="998" type="male">five males</specimenCount>
(
<collectionCode box="[1341,1431,1331,1353]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="25" pageNumber="998" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841569" collectingDate="1925-11-13" collectionCode="NMPC" collectorName="Bridarolli" country="Brazil" location="Martinez" municipality="Uncertain" pageId="25" pageNumber="998" specimenCount="1" specimenCount-male="1" stateProvince="Rio Grande do Sul">
<emphasis bold="true" box="[827,1097,1362,1384]" pageId="25" pageNumber="998">
<emphasis bold="true" box="[827,1088,1362,1384]" italics="true" pageId="25" pageNumber="998">
<collectingMunicipality box="[827,969,1362,1383]" pageId="25" pageNumber="998">Uncertain</collectingMunicipality>
locality
</emphasis>
:
</emphasis>
<emphasis box="[1115,1229,1363,1384]" italics="true" pageId="25" pageNumber="998">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53A630858DFEFFA9B" box="[1115,1229,1363,1384]" country="Brazil" municipality="Uncertain" name="Martinez" pageId="25" pageNumber="998" stateProvince="Rio Grande do Sul">Martinez</location>
</emphasis>
(?),
<date box="[1297,1436,1362,1384]" pageId="25" pageNumber="998" value="1925-11-13">
<collectingDate box="[1297,1436,1362,1384]" pageId="25" pageNumber="998" value="1925-11-13">13.XI.1925</collectingDate>
</date>
,
<collectorName box="[827,940,1393,1415]" pageId="25" pageNumber="998">Bridarolli</collectorName>
leg.
<specimenCount box="[1012,1115,1393,1415]" pageId="25" pageNumber="998" type="male">one male</specimenCount>
(
<collectionCode box="[1129,1212,1393,1415]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="25" pageNumber="998">NMPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841522" collectingDate="1943-03" collectionCode="MACN" country="Brazil" location="No" pageId="25" pageNumber="998" specimenCount="1" specimenCount-male="1" stateProvince="Rio Grande do Sul">
<emphasis bold="true" box="[1230,1443,1392,1414]" pageId="25" pageNumber="998">
<emphasis bold="true" box="[1230,1435,1392,1414]" italics="true" pageId="25" pageNumber="998">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53AF6087ADFD3FA75" box="[1230,1265,1393,1414]" country="Brazil" name="No" pageId="25" pageNumber="998" stateProvince="Rio Grande do Sul">No</location>
locality data
</emphasis>
:
</emphasis>
<date box="[832,933,1423,1444]" pageId="25" pageNumber="998" value="1943-03">
<collectingDate box="[832,933,1423,1444]" pageId="25" pageNumber="998" value="1943-03">III.1943</collectingDate>
</date>
<specimenCount box="[972,1084,1423,1445]" pageId="25" pageNumber="998" type="male">one male</specimenCount>
(
<collectionCode box="[1102,1191,1423,1445]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="25" pageNumber="998" type="Museum">MACN</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841459" collectingDate="1932-11" collectionCode="RBINS" collectorName="Le Moult" country="Brazil" location="Cerro" municipality="Ambiguous" pageId="25" pageNumber="998" specimenCount="1" specimenCount-female="1" stateProvince="Rio Grande do Sul">
<emphasis bold="true" box="[1212,1443,1423,1445]" pageId="25" pageNumber="998">
<emphasis bold="true" box="[1212,1435,1423,1445]" italics="true" pageId="25" pageNumber="998">
<collectingMunicipality box="[1212,1362,1423,1445]" pageId="25" pageNumber="998">Ambiguous</collectingMunicipality>
data
</emphasis>
:
</emphasis>
<emphasis box="[833,900,1454,1475]" italics="true" pageId="25" pageNumber="998">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53D7908A5D8A6FA30" box="[833,900,1454,1475]" country="Brazil" municipality="Ambiguous" name="Cerro" pageId="25" pageNumber="998" stateProvince="Rio Grande do Sul">Cerro</location>
</emphasis>
,
<date box="[922,1018,1454,1475]" pageId="25" pageNumber="998" value="1932-11">
<collectingDate box="[922,1018,1454,1475]" pageId="25" pageNumber="998" value="1932-11">XI.1932</collectingDate>
</date>
, no collector
<specimenCount box="[1207,1339,1454,1476]" pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[1357,1442,1454,1475]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="25" pageNumber="998" type="Museum">RBINS</collectionCode>
ex
<collectorName box="[889,1003,1485,1507]" pageId="25" pageNumber="998">Le Moult</collectorName>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841433" collectionCode="NMPC, SMF" country="Brazil" pageId="25" pageNumber="998" specimenCount="1" specimenCount-male="1" stateProvince="Rio Grande do Sul">
<collectorName box="[1166,1235,1485,1506]" pageId="25" pageNumber="998">
<emphasis box="[1166,1229,1485,1506]" italics="true" pageId="25" pageNumber="998">Mont</emphasis>
.
</collectorName>
(
<collectorName box="[1262,1407,1485,1507]" pageId="25" pageNumber="998">Montevideo</collectorName>
or
<collectorName box="[827,906,1516,1537]" pageId="25" pageNumber="998">Monte</collectorName>
,
<collectorName box="[920,1046,1515,1537]" pageId="25" pageNumber="998">Argentina</collectorName>
?)
<specimenCount box="[1101,1212,1515,1537]" pageId="25" pageNumber="998" type="male">one male</specimenCount>
(
<collectionCode box="[1230,1313,1515,1536]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="25" pageNumber="998">NMPC</collectionCode>
ex
<collectionCode box="[1385,1441,1515,1536]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="25" pageNumber="998">SMF</collectionCode>
and
<collectorName box="[879,975,1546,1568]" pageId="25" pageNumber="998">E. Witte</collectorName>
collection)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841530" collectionCode="MLPA" collectorName="Menatra" country="Brazil" location="Rio Negro" pageId="25" pageNumber="998" specimenCount="1" specimenCount-female="1" stateProvince="Rio Grande do Sul">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53A590B01DFFAF9EC" box="[1121,1240,1546,1568]" country="Brazil" name="Rio Negro" pageId="25" pageNumber="998" stateProvince="Rio Grande do Sul">Río Negro</location>
(
<collectorName box="[1256,1360,1546,1568]" pageId="25" pageNumber="998">Menatra</collectorName>
)
<specimenCount pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[919,997,1577,1599]" pageId="25" pageNumber="998">MLPA</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3704841386" box="[1020,1439,1577,1599]" collectionCode="MACN" collectorName="Yventario'" country="Brazil" pageId="25" pageNumber="998" specimenCount="1" specimenCount-female="1" stateProvince="Rio Grande do Sul">
<collectorName box="[1020,1141,1577,1599]" pageId="25" pageNumber="998">Yventario</collectorName>
(?)
<specimenCount box="[1206,1332,1577,1599]" pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[1347,1432,1577,1599]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="25" pageNumber="998" type="Museum">MACN</collectionCode>
)
</materialsCitation>
.
<emphasis bold="true" box="[827,1037,1607,1629]" pageId="25" pageNumber="998">
<materialsCitation ID-GBIF-Occurrence="3704841326" box="[827,1030,1607,1629]" country="Brazil" location="Erroneous" pageId="25" pageNumber="998" specimenCount="1" stateProvince="Rio Grande do Sul">
<emphasis bold="true" box="[827,1030,1607,1629]" italics="true" pageId="25" pageNumber="998">
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53D030B43D8E3F9AE" box="[827,961,1608,1629]" country="Brazil" name="Erroneous" pageId="25" pageNumber="998" stateProvince="Rio Grande do Sul">Erroneous</location>
data
</emphasis>
</materialsCitation>
:
</emphasis>
<materialsCitation ID-GBIF-Occurrence="3704841549" collectionCode="RBINS" collectorName="Perou' &amp; Popelaire'" country="Peru" location="PERU" pageId="25" pageNumber="998" specimenCount="1" specimenCount-female="1">
<emphasis bold="true" box="[1046,1132,1607,1628]" pageId="25" pageNumber="998">
<collectingCountry box="[1046,1128,1607,1628]" name="Peru" pageId="25" pageNumber="998">PERU</collectingCountry>
:
</emphasis>
<collectorName box="[1144,1222,1607,1629]" pageId="25" pageNumber="998">Pérou</collectorName>
,
<collectorName box="[1238,1358,1607,1629]" pageId="25" pageNumber="998">Popelaire</collectorName>
, 5717, no further data
<specimenCount box="[1056,1191,1638,1660]" pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[1210,1302,1638,1660]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="25" pageNumber="998" type="Museum">RBINS</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3704841431" collectionCode="BMNH" collectorName="No" country="Peru" pageId="25" pageNumber="998" specimenCount="1" specimenCount-male="1">
<emphasis bold="true" box="[1324,1443,1637,1659]" pageId="25" pageNumber="998">
<emphasis bold="true" box="[1324,1435,1637,1659]" italics="true" pageId="25" pageNumber="998">
<collectorName box="[1324,1360,1638,1659]" pageId="25" pageNumber="998">No</collectorName>
data
</emphasis>
:
</emphasis>
<specimenCount box="[827,933,1669,1691]" pageId="25" pageNumber="998" type="male">one male</specimenCount>
(
<collectionCode box="[950,1038,1669,1691]" pageId="25" pageNumber="998">BMNH</collectionCode>
)
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3704841544" collectionCode="BMNH, MACN" collectorName="Nevinson" country="Peru" pageId="25" pageNumber="998" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount box="[1058,1165,1669,1691]" pageId="25" pageNumber="998" type="male">one male</specimenCount>
(
<collectionCode box="[1181,1266,1669,1690]" pageId="25" pageNumber="998">BMNH</collectionCode>
ex
<collectorName box="[1331,1442,1669,1691]" pageId="25" pageNumber="998">Nevinson</collectorName>
collection),
<specimenCount box="[964,1103,1699,1721]" pageId="25" pageNumber="998" type="male">eight males</specimenCount>
and
<specimenCount box="[1165,1329,1699,1721]" pageId="25" pageNumber="998" type="female">three females</specimenCount>
(
<collectionCode box="[1345,1431,1699,1721]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="25" pageNumber="998" type="Museum">MACN</collectionCode>
)
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3704841460" box="[827,1292,1730,1752]" collectionCode="MLPA" collectorName="Nevinson" country="Peru" isTailEnumeration="true" pageId="25" pageNumber="998" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount box="[827,941,1730,1752]" pageId="25" pageNumber="998" type="male">one male</specimenCount>
and
<specimenCount box="[1009,1182,1730,1752]" pageId="25" pageNumber="998" type="female">three females</specimenCount>
(
<collectionCode box="[1201,1284,1730,1752]" pageId="25" pageNumber="998">MLPA</collectionCode>
)
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3704841339" collectionCode="MNHN" country="Peru" pageId="25" pageNumber="998" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount box="[1308,1441,1730,1752]" pageId="25" pageNumber="998" type="male">four males</specimenCount>
and
<specimenCount box="[879,1021,1761,1783]" pageId="25" pageNumber="998" type="female">two females</specimenCount>
(
<collectionCode box="[1037,1127,1761,1783]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="998">MNHN</collectionCode>
)
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3704841535" collectionCode="RBINS, SMF" collectorName="E. Candeze &amp; Le Moult" country="Peru" location="Albert Hepp" pageId="25" pageNumber="998" specimenCount="5" specimenCount-female="2" specimenCount-male="3">
<specimenCount box="[1147,1253,1761,1783]" pageId="25" pageNumber="998" type="male">one male</specimenCount>
and
<specimenCount box="[1315,1442,1761,1783]" pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[834,919,1791,1812]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="25" pageNumber="998" type="Museum">RBINS</collectionCode>
ex
<collectorName box="[987,1121,1791,1813]" pageId="25" pageNumber="998">E. Candèze</collectorName>
collection),
<specimenCount box="[1268,1389,1791,1813]" pageId="25" pageNumber="998" type="male">two males</specimenCount>
and
<specimenCount box="[827,952,1822,1844]" pageId="25" pageNumber="998" type="female">one female</specimenCount>
(
<collectionCode box="[967,1050,1822,1843]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="25" pageNumber="998" type="Museum">RBINS</collectionCode>
ex
<collectorName box="[1113,1218,1822,1844]" pageId="25" pageNumber="998">Le Moult</collectorName>
collection) and
<specimenCount pageId="25" pageNumber="998" type="male">one male</specimenCount>
(
<collectionCode box="[898,955,1853,1874]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="25" pageNumber="998">SMF</collectionCode>
ex
<location LSID="urn:lsid:plazi:treatment:27085036DC08FFE53ECE08B8DE29F8A1:AA7EB7FBDC19FFE53DCF0A37DFA4F8A1" box="[1015,1158,1852,1874]" country="Peru" name="Albert Hepp" pageId="25" pageNumber="998">Albert Hepp</location>
collection)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>