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<document id="D303DDA693437FADC1D2D8F531BAABF5" ID-DOI="10.5852/ejt.2020.729.1187" ID-GBIF-Dataset="3ea2481a-7288-4dfc-9bae-ce104e5c0ebd" ID-ISSN="2118-9773" ID-Zenodo-Dep="4326611" ID-ZooBank="1277073A-03A6-4698-A614-3AD796B45AD1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="valdenar" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1608108559487" checkinUser="plazi" docAuthor="Granja-Fernández, Rebeca, Pineda-Enríquez, Tania, Solís-Marín, Francisco Alonso &amp; Laguarda-Figueras, Alfredo" docDate="2020" docId="03E8EB0C5321FFE1FDC89B70FEE3C3CB" docLanguage="en" docName="ejt-729.11.pdf" docOrigin="European Journal of Taxonomy 729" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ophioderma hendleri Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras 2020, sp. nov." docType="treatment" docVersion="10" lastPageNumber="23" masterDocId="FFD193745322FFEDFFAF9E62FFCEC423" masterDocTitle="Ophioderma hendleri sp. nov. (Echinodermata: Ophiuroidea: Ophiodermatidae) and its congeners from the Eastern Pacific" masterLastPageNumber="41" masterPageNumber="11" pageNumber="14" updateTime="1701998094450" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="5245B62FB257A454F1A657C745AAB4E1">Ophioderma hendleri sp. nov. (Echinodermata: Ophiuroidea: Ophiodermatidae) and its congeners from the Eastern Pacific</mods:title>
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<mods:namePart id="AEC1E4DBCA0DC4B009A1AD25245B5308">Granja-Fernández, Rebeca</mods:namePart>
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<mods:affiliation id="4785752DF821FBF34F5F550634D37299">Laboratorio de Ecología Marina. Centro Universitario de la Costa, Universidad de Guadalajara, 48280 Puerto Vallarta, Mexico.</mods:affiliation>
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<mods:namePart id="6DB5CB4BE4B84E68AF66A241690230EE">Pineda-Enríquez, Tania</mods:namePart>
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<mods:affiliation id="469C261127A9CE4A711214EBC4A19EAE">Department of Biology, Division of Invertebrate Zoology, Florida Museum of Natural History, University of Florida, Gainesville, 32611 Florida, USA. &amp; Natural History Museum of Los Angeles County, 900 Exposition Blvd., 90007 Los Angeles, California, USA.</mods:affiliation>
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<mods:namePart id="CA6D90BE2489A6D4D8425F4A94818D87">Solís-Marín, Francisco Alonso</mods:namePart>
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<mods:affiliation id="2B502C6918E20020A69278C75B1C4B6B">Colección Nacional de Equinodermos “ Ma. E. Caso Muñoz ”, Laboratorio de Sistemática y Ecología de Equinodermos, Instituto de Ciencias del Mar y Limnología (ICML), Universidad Nacional Autónoma de Mexico (UNAM), 04510 Ciudad de Mexico, Mexico.</mods:affiliation>
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<mods:namePart id="5010AACAE63ABABC97D9D3053296268D">Laguarda-Figueras, Alfredo</mods:namePart>
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<mods:affiliation id="18CB5021508DB24BAD2AE5FAF548152A">Colección Nacional de Equinodermos “ Ma. E. Caso Muñoz ”, Laboratorio de Sistemática y Ecología de Equinodermos, Instituto de Ciencias del Mar y Limnología (ICML), Universidad Nacional Autónoma de Mexico (UNAM), 04510 Ciudad de Mexico, Mexico.</mods:affiliation>
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<heading id="D0B6ED765321FFEEFDC89B70FC03C10E" box="[615,973,1298,1325]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="14" reason="2">
<taxonomicName id="4C4121995321FFEEFDC89B70FCA5C10E" authority="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras, 2020" authorityName="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras" authorityYear="2020" box="[615,875,1298,1325]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="3" pageNumber="14" phylum="Echinodermata" rank="species" species="hendleri" status="sp. nov.">
<emphasis id="B93586085321FFEEFDC89B70FCA5C10E" bold="true" box="[615,875,1298,1325]" italics="true" pageId="3" pageNumber="14">Ophioderma hendleri</emphasis>
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<figureCitation id="137A469F5321FFEEFD269B39FD3FC155" box="[649,753,1371,1398]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="5.[189,232,1893,1919]" captionStartId-1="6.[189,232,1752,1778]" captionStartId-2="7.[189,232,1929,1955]" captionStartId-3="8.[189,232,1928,1954]" captionTargetBox-0="[189,1398,291,1855]" captionTargetBox-1="[295,1292,265,1714]" captionTargetBox-2="[189,1398,327,1891]" captionTargetBox-3="[189,1398,325,1889]" captionTargetId-0="figure-12@5.[189,1398,291,1855]" captionTargetId-1="figure-170@6.[294,1293,265,1714]" captionTargetId-2="figure-11@7.[189,1398,327,1891]" captionTargetId-3="figure-17@8.[189,1398,325,1889]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="Fig. 1. Ophioderma hendleri sp. nov., holotype (ICML-UNAM 18315, DD = 14.8 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Structures of a jaw (according to Hendler 2018). E. Basal dorsal arm. F. Dorsal arm. G. Ventral arm. H. Lateral arm plates and arm spines. I. Dorsal view. See Material and methods for abbreviations." captionText-1="Fig. 2. SEM images of skeletal elements of Ophioderma hendleri sp. nov., paratype (ICML-UNAM 18323, DD = 15 mm). AB. Radial shield (external view), deepest pores marked with arrows. C. Center of the distal portion of the radial shield, open meshed pores. D. Dorsal arm plate, with one spur. E. Ventral arm plate, with one spur. F. Lateral arm plate, external face, with two spurs and one condyle. G. Lateral arm plate, lateral face, with spine articulations. H. Lateral arm plate, internal face. I. Outer surface of the LAP (F) with finely meshed and polygonal knobs. J. Spine articulations with a weak sigmoidal fold and tilted lobes. K. Arm vertebra, distal face. L. Arm vertebra, proximal face. M. Tentacle scale. N. Arm spine. O. Dental plate fragments. P. Oral plate, abradial face. Q. Oral plate, adradial face." captionText-2="Fig. 3. Ophioderma hendleri sp. nov., paratype (ICML-UNAM 10582, DD = 4.8 mm). A. Dorsal view. B. Basal dorsal arm. C. Dorsal arm. D. Ventral arm. E. Dorsal disc. F. Ventral disc. G. Interradii. H. Mouth." captionText-3="Fig. 4. Ophioderma hendleri sp. nov., paratype (ICML-UNAM 18319, DD = 2.2 mm). A. Dorsal view. B. Basal dorsal arm. C. Dorsal arm. D. Ventral arm. E. Dorsal disc. F. Ventral disc. G. Interradii. H. Mouth." figureDoi-0="http://doi.org/10.5281/zenodo.4326613" figureDoi-1="http://doi.org/10.5281/zenodo.4326617" figureDoi-2="http://doi.org/10.5281/zenodo.4326619" figureDoi-3="http://doi.org/10.5281/zenodo.4326621" httpUri-0="https://zenodo.org/record/4326613/files/figure.png" httpUri-1="https://zenodo.org/record/4326617/files/figure.png" httpUri-2="https://zenodo.org/record/4326619/files/figure.png" httpUri-3="https://zenodo.org/record/4326621/files/figure.png" pageId="3" pageNumber="14">Figs 14</figureCitation>
,
<figureCitation id="137A469F5321FFEEFD529B39FCC4C155" box="[765,778,1371,1398]" captionStart="Fig" captionStartId="9.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@9.[189,1398,291,1855]" captionTargetPageId="9" captionText="Fig. 5. AD. Ophioderma hendleri sp. nov. AB. In vivo coloration in natural habitat. CD. In vivo coloration of collected specimens. EJ. Ophioderma panamense Lütken, 1859, holotype (NHMD- 107679, DD = 21.9 mm). E. Dorsal disc. F. Ventral disc. G. Mouth. H. Dorsal arm. I. Ventral arm. J. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4326625" httpUri="https://zenodo.org/record/4326625/files/figure.png" pageId="3" pageNumber="14">5</figureCitation>
AD;
<tableCitation id="C6C36FA15321FFEEFCFB9B39FC65C155" box="[852,939,1371,1398]" captionStart="Table 1" captionStartId="10.[189,258,265,291]" captionTargetPageId="10" captionText="Table 1. Comparison of morphological characters between some species of Ophioderma from the Eastern Pacific." httpUri="http://table.plazi.org/id/DF3E0A925328FFE7FF129F6BFEBAC565" pageId="3" pageNumber="14" tableUuid="DF3E0A925328FFE7FF129F6BFEBAC565">Table 1</tableCitation>
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<paragraph id="8BFE5A1A5321FFEEFF129BFCFA92C19A" blockId="3.[189,1372,1438,1465]" box="[189,1372,1438,1465]" pageId="3" pageNumber="14">
<treatmentCitationGroup id="AB517D345321FFEEFF129BFCFA92C19A" box="[189,1372,1438,1465]" pageId="3" pageNumber="14">
<taxonomicName id="4C4121995321FFEEFF129BFCFEB4C19A" authorityName="Muller &amp; Troschel" authorityYear="1840" box="[189,378,1438,1465]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="3" pageNumber="14" phylum="Echinodermata" rank="species" species="sp.">
<emphasis id="B93586085321FFEEFF129BFCFE9FC19B" box="[189,337,1438,1464]" italics="true" pageId="3" pageNumber="14">Ophioderma</emphasis>
sp.
</taxonomicName>
<treatmentCitation id="0AE07C0B5321FFEEFE389BFCFCA6C19A" author="Granja-Fernandez R. &amp; Herrero-Perezrul M. D. &amp; Lopez-Perez R. A. &amp; Hernandez L. &amp; Rodriguez-Zaragoza F. A. &amp; Jones R. W. &amp; Pineda-Lopez R." box="[407,872,1438,1465]" page="135" pageId="3" pageNumber="14" year="2014">
<bibRefCitation id="EFD027EB5321FFEEFE389BFCFCA6C19A" author="Granja-Fernandez R. &amp; Herrero-Perezrul M. D. &amp; Lopez-Perez R. A. &amp; Hernandez L. &amp; Rodriguez-Zaragoza F. A. &amp; Jones R. W. &amp; Pineda-Lopez R." box="[407,872,1438,1465]" pageId="3" pageNumber="14" pagination="101 - 145" refId="ref15937" refString="Granja-Fernandez R., Herrero-Perezrul M. D., Lopez-Perez R. A., Hernandez L., Rodriguez-Zaragoza F. A., Jones R. W. &amp; Pineda-Lopez R. 2014. Ophiuroidea (Echinodermata) from coral reefs in the Mexican Pacific. Zookeys 406: 101 - 145. https: // doi. org / 10.3897 / zookeys. 406.6306" type="journal article" year="2014">
Granja-Fernández
<emphasis id="B93586085321FFEEFDDD9BFDFD68C19B" box="[626,678,1438,1464]" italics="true" pageId="3" pageNumber="14">et al</emphasis>
. 2014: 135137
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, fig. 7AF; 2015a: 4142; 2017: 172175.
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</paragraph>
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<subSubSection id="C35B09915321FFEEFF129B82FE9CC2B4" pageId="3" pageNumber="14" type="diagnosis">
<paragraph id="8BFE5A1A5321FFEEFF129B82FEF8C1D9" blockId="3.[189,310,1504,1530]" box="[189,310,1504,1530]" pageId="3" pageNumber="14">
<heading id="D0B6ED765321FFEEFF129B82FEF8C1D9" bold="true" box="[189,310,1504,1530]" fontSize="11" level="3" pageId="3" pageNumber="14" reason="3">
<emphasis id="B93586085321FFEEFF129B82FEF8C1D9" bold="true" box="[189,310,1504,1530]" pageId="3" pageNumber="14">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BFE5A1A5321FFEEFF129872FE9CC2B4" blockId="3.[189,1399,1552,1687]" pageId="3" pageNumber="14">Radial shields covered by granulation. Adoral shields naked, as long as broad. Up to 11 arm spines, dorsalmost the shortest and blunt, ventralmost the largest and robust. In vivo coloration, disc dark brown and mottled with small white spots, ventral margin beige and brown; arms dark brown with transverse white bands.</paragraph>
</subSubSection>
<subSubSection id="C35B09915321FFEEFF1298A2FC89C30D" pageId="3" pageNumber="14" type="etymology">
<paragraph id="8BFE5A1A5321FFEEFF1298A2FE8AC2F9" blockId="3.[189,324,1728,1754]" box="[189,324,1728,1754]" pageId="3" pageNumber="14">
<heading id="D0B6ED765321FFEEFF1298A2FE8AC2F9" bold="true" box="[189,324,1728,1754]" fontSize="11" level="3" pageId="3" pageNumber="14" reason="3">
<emphasis id="B93586085321FFEEFF1298A2FE8AC2F9" bold="true" box="[189,324,1728,1754]" pageId="3" pageNumber="14">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BFE5A1A5321FFEEFF12988DFC89C30D" blockId="3.[189,1399,1775,1838]" pageId="3" pageNumber="14">This species is named after Dr. Gordon Hendler, an eminent ophiuroid taxonomist, curator of echinoderms at the Natural History Museum of Los Angeles County.</paragraph>
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<subSubSection id="C35B09915321FFE9FF129934FDD6C0D9" lastPageId="4" lastPageNumber="15" pageId="3" pageNumber="14" type="materials_examined">
<paragraph id="8BFE5A1A5321FFEEFF129934FE64C353" blockId="3.[189,426,1878,1904]" box="[189,426,1878,1904]" pageId="3" pageNumber="14">
<heading id="D0B6ED765321FFEEFF129934FE64C353" bold="true" box="[189,426,1878,1904]" fontSize="11" level="3" pageId="3" pageNumber="14" reason="3">
<emphasis id="B93586085321FFEEFF129934FE64C353" bold="true" box="[189,426,1878,1904]" pageId="3" pageNumber="14">Material examined</emphasis>
</heading>
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<paragraph id="8BFE5A1A5321FFEEFF4599E4FE92C383" blockId="3.[189,1399,1926,2025]" box="[234,348,1926,1952]" pageId="3" pageNumber="14">
<emphasis id="B93586085321FFEEFF4599E4FE92C383" bold="true" box="[234,348,1926,1952]" pageId="3" pageNumber="14">
<typeStatus id="54FAE4B85321FFEEFF4599E4FE92C383" box="[234,348,1926,1952]" pageId="3" pageNumber="14" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BFE5A1A5321FFEEFF1299C8FD58C3CA" blockId="3.[189,1399,1926,2025]" pageId="3" pageNumber="14">
<materialsCitation id="3B2950475321FFEEFF1299C8FD58C3CA" ID-GBIF-Occurrence="2995182307" collectedFrom="under rocks" collectingDate="2011-11-05" collectionCode="R" collectorName="R. Granja" country="Mexico" elevation="21" latitude="16.030527" location="Chipehua" longLatPrecision="1" longitude="-95.35936" pageId="3" pageNumber="14" specimenCode="ICML-UNAM 18315" specimenCount="1" stateProvince="Oaxaca" typeStatus="holotype">
<collectingCountry id="F3561A8A5321FFEEFF1299C8FEFFC3E7" box="[189,305,1962,1988]" name="Mexico" pageId="3" pageNumber="14">MEXICO</collectingCountry>
<specimenCount id="9D4791935321FFEEFEE599C8FE6EC3E6" box="[330,416,1962,1989]" pageId="3" pageNumber="14" type="generic">1 spec.</specimenCount>
;
<location id="8E9E0CC15321FFEEFE0399C8FDD1C3E7" LSID="urn:lsid:plazi:treatment:03E8EB0C5321FFE1FDC89B70FEE3C3CB:8E9E0CC15321FFEEFE0399C8FDD1C3E7" box="[428,543,1962,1988]" country="Mexico" latitude="16.030527" longLatPrecision="1" longitude="-95.35936" name="Chipehua" pageId="3" pageNumber="14" stateProvince="Oaxaca">Chipehua</location>
,
<collectingRegion id="498594F85321FFEEFD8499C8FD48C3E7" box="[555,646,1962,1988]" country="Mexico" name="Oaxaca" pageId="3" pageNumber="14">Oaxaca</collectingRegion>
,
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,
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;
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;
<date id="FFFF7CDA5321FFEEFBCA99C8FB36C3E7" box="[1125,1272,1962,1989]" pageId="3" pageNumber="14" value="2011-11-05">
<collectingDate id="EFBB85325321FFEEFBCA99C8FB36C3E7" box="[1125,1272,1962,1989]" pageId="3" pageNumber="14" value="2011-11-05">5 Nov. 2011</collectingDate>
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;
<collectorName id="26B43FCC5321FFEEFAAB99C9FAB9C3E7" box="[1284,1399,1962,1989]" pageId="3" pageNumber="14">R. Granja</collectorName>
leg.;
<collectedFrom id="385D19E75321FFEEFF5799ACFE4BC3CB" box="[248,389,1998,2024]" pageId="3" pageNumber="14">under rocks</collectedFrom>
;
<specimenCode id="DBE7F2615321FFEEFE3E99ADFD5CC3CA" box="[401,658,1998,2025]" pageId="3" pageNumber="14">ICML-UNAM 18315</specimenCode>
.
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</paragraph>
<paragraph id="8BFE5A1A5326FFE9FF459F6BFEA9C500" blockId="4.[189,1342,265,327]" box="[234,359,265,291]" pageId="4" pageNumber="15">
<emphasis id="B93586085326FFE9FF459F6BFEA9C500" bold="true" box="[234,359,265,291]" pageId="4" pageNumber="15">
<typeStatus id="54FAE4B85326FFE9FF459F6BFEA9C500" box="[234,359,265,291]" pageId="4" pageNumber="15" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BFE5A1A5326FFE9FF129F4EFAF0C565" blockId="4.[189,1342,265,327]" box="[189,1342,300,327]" pageId="4" pageNumber="15">
<materialsCitation id="3B2950475326FFE9FF129F4EFAF7C565" ID-GBIF-Occurrence="2995182309" box="[189,1337,300,327]" collectingDate="1994-02-19" collectingDateMax="1994-02-28" collectingDateMin="1994-02-19" collectionCode="MZUCR-ECH" collectorName="Gordon Hendler" country="Costa Rica" location="Coco Island" pageId="4" pageNumber="15" specimenCode="MZUCR-ECH 321" specimenCount="6" typeStatus="paratype">
<collectingCountry id="F3561A8A5326FFE9FF129F4EFEABC565" box="[189,357,300,326]" name="Costa Rica" pageId="4" pageNumber="15">COSTA RICA</collectingCountry>
<specimenCount id="9D4791935326FFE9FED19F4EFE19C564" box="[382,471,300,327]" pageId="4" pageNumber="15">6 specs</specimenCount>
;
<location id="8E9E0CC15326FFE9FE4C9F4EFDBDC565" LSID="urn:lsid:plazi:treatment:03E8EB0C5321FFE1FDC89B70FEE3C3CB:8E9E0CC15326FFE9FE4C9F4EFDBDC565" box="[483,627,300,326]" country="Costa Rica" name="Coco Island" pageId="4" pageNumber="15">Coco Island</location>
;
<date id="FFFF7CDA5326FFE9FDD09F4EFC87C565" box="[639,841,300,326]" pageId="4" pageNumber="15" value="1994-02-19" valueMax="1994-02-28" valueMin="1994-02-19">
<collectingDate id="EFBB85325326FFE9FDD09F4EFC87C565" box="[639,841,300,326]" pageId="4" pageNumber="15" value="1994-02-19" valueMax="1994-02-28" valueMin="1994-02-19">1928 Feb. 1994</collectingDate>
</date>
;
<collectorName id="26B43FCC5326FFE9FCFA9F4EFBDAC565" box="[853,1044,300,326]" pageId="4" pageNumber="15">Gordon Hendler</collectorName>
leg.;
<specimenCode id="DBE7F2615326FFE9FBF99F4EFAF7C565" box="[1110,1337,300,326]" collectionCode="MZUCR-ECH" pageId="4" pageNumber="15">MZUCR-ECH 321</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BFE5A1A5326FFE9FF129F0CFAA7C071" blockId="4.[189,1399,366,1107]" pageId="4" pageNumber="15">
<materialsCitation id="3B2950475326FFE9FF129F0CFC91C58E" ID-GBIF-Occurrence="2995182315" collectedFrom="under rocks" collectingDate="2012-06-01" collectionCode="R" collectorName="R. Granja" country="Mexico" elevation="9" latitude="17.654167" location="Zacatoso" longLatPrecision="1" longitude="-101.62195" municipality="Zacatoso" pageId="4" pageNumber="15" specimenCode="ICML-UNAM 10582" specimenCount="2" stateProvince="Guerrero">
<collectingCountry id="F3561A8A5326FFE9FF129F0CFEFFC5AB" box="[189,305,366,392]" name="Mexico" pageId="4" pageNumber="15">MEXICO</collectingCountry>
<specimenCount id="9D4791935326FFE9FEFC9F0CFE7FC5AA" box="[339,433,366,393]" pageId="4" pageNumber="15">2 specs</specimenCount>
;
<location id="8E9E0CC15326FFE9FE6D9F0DFDE1C5AA" LSID="urn:lsid:plazi:treatment:03E8EB0C5321FFE1FDC89B70FEE3C3CB:8E9E0CC15326FFE9FE6D9F0DFDE1C5AA" box="[450,559,367,393]" country="Mexico" latitude="17.654167" longLatPrecision="1" longitude="-101.62195" municipality="Zacatoso" name="Zacatoso" pageId="4" pageNumber="15" stateProvince="Guerrero">Zacatoso</location>
,
<location id="8E9E0CC15326FFE9FD909F0DFD45C5AA" LSID="urn:lsid:plazi:treatment:03E8EB0C5321FFE1FDC89B70FEE3C3CB:8E9E0CC15326FFE9FD909F0DFD45C5AA" box="[575,651,367,393]" country="Mexico" latitude="17.654167" longLatPrecision="1" longitude="-101.62195" municipality="Zacatoso" name="Ixtapa" pageId="4" pageNumber="15" stateProvince="Guerrero">Ixtapa</location>
,
<collectingRegion id="498594F85326FFE9FD339F0CFCC4C5AB" box="[668,778,366,392]" country="Mexico" name="Guerrero" pageId="4" pageNumber="15">Guerrero</collectingRegion>
;
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,
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;
<quantity id="4CB9F7FF5326FFE9FB7F9F0CFAD4C5AA" box="[1232,1306,366,393]" metricMagnitude="0" metricUnit="m" metricValue="9.1" pageId="4" pageNumber="15" unit="m" value="9.1">
<elevation id="006CBD295326FFE9FB7F9F0CFAD4C5AA" box="[1232,1306,366,393]" metricMagnitude="0" metricUnit="m" metricValue="9.1" pageId="4" pageNumber="15" unit="m" value="9.1">9.1 m</elevation>
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;
<date id="FFFF7CDA5326FFE9FA839F0CFF35C58F" pageId="4" pageNumber="15" value="2012-06-01">
<collectingDate id="EFBB85325326FFE9FA839F0CFF35C58F" pageId="4" pageNumber="15" value="2012-06-01">1 Jun. 2012</collectingDate>
</date>
;
<collectorName id="26B43FCC5326FFE9FEA69FF0FEB3C58F" box="[265,381,402,428]" pageId="4" pageNumber="15">R. Granja</collectorName>
leg.;
<collectedFrom id="385D19E75326FFE9FE6C9FF0FD9CC58F" box="[451,594,402,428]" pageId="4" pageNumber="15">under rocks</collectedFrom>
;
<specimenCode id="DBE7F2615326FFE9FDCF9FF0FC91C58E" box="[608,863,402,429]" pageId="4" pageNumber="15">ICML-UNAM 10582</specimenCode>
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<materialsCitation id="3B2950475326FFE9FCD39FF0FACDC5EC" ID-GBIF-Occurrence="2995182304" collectedFrom="under rocks" collectingDate="2017-01-14" collectionCode="R" collectorName="R. Granja" country="Mexico" elevation="7" latitude="15.750806" location="Chahue" longLatPrecision="1" longitude="-96.12318" pageId="4" pageNumber="15" specimenCode="ICML-UNAM 18316" specimenCount="1" stateProvince="Oaxaca">
<specimenCount id="9D4791935326FFE9FCD39FF0FC1DC58F" box="[892,979,402,428]" pageId="4" pageNumber="15" type="generic">1 spec.</specimenCount>
;
<location id="8E9E0CC15326FFE9FC4E9FF0FBF0C58F" LSID="urn:lsid:plazi:treatment:03E8EB0C5321FFE1FDC89B70FEE3C3CB:8E9E0CC15326FFE9FC4E9FF0FBF0C58F" box="[993,1086,402,428]" country="Mexico" latitude="15.750806" longLatPrecision="1" longitude="-96.12318" name="Chahue" pageId="4" pageNumber="15" stateProvince="Oaxaca">Chahué</location>
,
<collectingRegion id="498594F85326FFE9FBE49FF0FB69C58F" box="[1099,1191,402,428]" country="Mexico" name="Oaxaca" pageId="4" pageNumber="15">Oaxaca</collectingRegion>
;
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;
<quantity id="4CB9F7FF5326FFE9FE289FD7FE02C5F3" box="[391,460,437,464]" metricMagnitude="0" metricUnit="m" metricValue="6.5" pageId="4" pageNumber="15" unit="m" value="6.5">
<elevation id="006CBD295326FFE9FE289FD7FE02C5F3" box="[391,460,437,464]" metricMagnitude="0" metricUnit="m" metricValue="6.5" pageId="4" pageNumber="15" unit="m" value="6.5">6.5 m</elevation>
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;
<date id="FFFF7CDA5326FFE9FE789FD4FDA3C5F3" box="[471,621,438,464]" pageId="4" pageNumber="15" value="2017-01-14">
<collectingDate id="EFBB85325326FFE9FE789FD4FDA3C5F3" box="[471,621,438,464]" pageId="4" pageNumber="15" value="2017-01-14">14 Jan. 2017</collectingDate>
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;
<collectorName id="26B43FCC5326FFE9FDD89FD4FD26C5EC" box="[631,744,437,464]" pageId="4" pageNumber="15">R. Granja</collectorName>
leg.;
<collectedFrom id="385D19E75326FFE9FC889FD7FC7CC5EC" box="[807,946,437,463]" pageId="4" pageNumber="15">under rocks</collectedFrom>
, 27°C;
<specimenCode id="DBE7F2615326FFE9FBA79FD4FACDC5EC" box="[1032,1283,437,464]" pageId="4" pageNumber="15">ICML-UNAM 18316</specimenCode>
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<materialsCitation id="3B2950475326FFE9FAB59FD4FD69C634" ID-GBIF-Occurrence="2995182316" collectedFrom="under rocks" collectingDate="2016-01-29" collectionCode="R" collectorName="D. Rangel &amp; R. Granja" country="Mexico" elevation="9" latitude="15.687533" location="Tijera" longLatPrecision="1" longitude="-96.44208" pageId="4" pageNumber="15" specimenCode="ICML-UNAM 18317" specimenCount="2" stateProvince="Oaxaca">
<specimenCount id="9D4791935326FFE9FAB59FD4FABCC5F3" box="[1306,1394,438,464]" pageId="4" pageNumber="15">2 specs</specimenCount>
;
<location id="8E9E0CC15326FFE9FF129FBBFECBC5D0" LSID="urn:lsid:plazi:treatment:03E8EB0C5321FFE1FDC89B70FEE3C3CB:8E9E0CC15326FFE9FF129FBBFECBC5D0" box="[189,261,473,499]" country="Mexico" latitude="15.687533" longLatPrecision="1" longitude="-96.44208" name="Tijera" pageId="4" pageNumber="15" stateProvince="Oaxaca">Tijera</location>
,
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,
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;
<quantity id="4CB9F7FF5326FFE9FCB29FBBFC9EC5D7" box="[797,848,473,500]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="15" unit="m" value="9.0">
<elevation id="006CBD295326FFE9FCB29FBBFC9EC5D7" box="[797,848,473,500]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="15" unit="m" value="9.0">9 m</elevation>
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;
<date id="FFFF7CDA5326FFE9FCF09FBBFC33C5D0" box="[863,1021,473,500]" pageId="4" pageNumber="15" value="2016-01-29">
<collectingDate id="EFBB85325326FFE9FCF09FBBFC33C5D0" box="[863,1021,473,500]" pageId="4" pageNumber="15" value="2016-01-29">29 Jan. 2016</collectingDate>
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;
<collectorName id="26B43FCC5326FFE9FBA39FB8FB4BC5D0" box="[1036,1157,473,500]" pageId="4" pageNumber="15">D. Rangel</collectorName>
and
<collectorName id="26B43FCC5326FFE9FB6B9FB8FAF7C5D0" box="[1220,1337,473,500]" pageId="4" pageNumber="15">R. Granja</collectorName>
leg.;
<collectedFrom id="385D19E75326FFE9FF129F9FFE85C634" box="[189,331,509,535]" pageId="4" pageNumber="15">under rocks</collectedFrom>
, 27°C;
<specimenCode id="DBE7F2615326FFE9FE079F9FFD69C634" box="[424,679,509,535]" pageId="4" pageNumber="15">ICML-UNAM 18317</specimenCode>
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<materialsCitation id="3B2950475326FFE9FD6A9F9FFEC8C67D" ID-GBIF-Occurrence="2995182311" collectedFrom="sediment under rocks" collectingDate="2016-02-01" collectionCode="ICML-UNAM" collectorName="Rebeca Granja" country="Mexico" elevation="20" latitude="16.24174" location="Frente a Llano Grande" longLatPrecision="1" longitude="-98.45129" municipality="Frente" pageId="4" pageNumber="15" specimenCode="ICML-UNAM 18318" specimenCount="1" stateProvince="Oaxaca">
<specimenCount id="9D4791935326FFE9FD6A9F9FFCD2C634" box="[709,796,509,535]" pageId="4" pageNumber="15" type="generic">1 spec.</specimenCount>
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<location id="8E9E0CC15326FFE9FC849F9FFBF3C634" LSID="urn:lsid:plazi:treatment:03E8EB0C5321FFE1FDC89B70FEE3C3CB:8E9E0CC15326FFE9FC849F9FFBF3C634" box="[811,1085,509,535]" country="Mexico" latitude="16.24174" longLatPrecision="1" longitude="-98.45129" municipality="Frente" name="Frente a Llano Grande" pageId="4" pageNumber="15" stateProvince="Oaxaca">Frente a Llano Grande</location>
,
<collectingRegion id="498594F85326FFE9FBE49F9FFB69C634" box="[1099,1191,509,535]" country="Mexico" name="Oaxaca" pageId="4" pageNumber="15">Oaxaca</collectingRegion>
;
<geoCoordinate id="EE753CDD5326FFE9FB1B9F9FFABCC634" box="[1204,1394,509,535]" degrees="16" direction="north" minutes="14" orientation="latitude" pageId="4" pageNumber="15" precision="1" seconds="30.26" value="16.24174">16°14´30.26˝ N</geoCoordinate>
,
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;
<quantity id="4CB9F7FF5326FFE9FE229C43FE03C618" box="[397,461,545,571]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="15" unit="m" value="20.0">
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;
<date id="FFFF7CDA5326FFE9FE749C43FDA2C619" box="[475,620,544,571]" pageId="4" pageNumber="15" value="2016-02-01">
<collectingDate id="EFBB85325326FFE9FE749C43FDA2C619" box="[475,620,544,571]" pageId="4" pageNumber="15" value="2016-02-01">1 Feb. 2016</collectingDate>
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;
<collectorName id="26B43FCC5326FFE9FDD59C43FCE7C618" box="[634,809,545,571]" pageId="4" pageNumber="15">Rebeca Granja</collectorName>
leg.;
<collectedFrom id="385D19E75326FFE9FCC19C43FBA0C618" box="[878,1134,545,571]" pageId="4" pageNumber="15">sediment under rocks</collectedFrom>
, 27°C;
<specimenCode id="DBE7F2615326FFE9FB659C43FEC8C67D" collectionCode="ICML-UNAM" pageId="4" pageNumber="15">ICML-UNAM 18318</specimenCode>
</materialsCitation>
<materialsCitation id="3B2950475326FFE9FEB29C26FBACC6A1" ID-GBIF-Occurrence="2995182317" collectedFrom="sediment under rocks" collectingDate="2016-01-26" collectionCode="R" collectorName="D. Rangel &amp; R. Granja" country="Mexico" elevation="12" latitude="15.7197" location="La Jeringa" longLatPrecision="1" longitude="-96.16109" municipality="La Jeringa" pageId="4" pageNumber="15" specimenCode="ICML-UNAM 18319" specimenCount="1" stateProvince="Oaxaca">
<specimenCount id="9D4791935326FFE9FEB29C26FEBFC67C" box="[285,369,580,607]" pageId="4" pageNumber="15" type="generic">1 spec.</specimenCount>
;
<location id="8E9E0CC15326FFE9FED29C27FE34C67D" LSID="urn:lsid:plazi:treatment:03E8EB0C5321FFE1FDC89B70FEE3C3CB:8E9E0CC15326FFE9FED29C27FE34C67D" box="[381,506,580,607]" country="Mexico" latitude="15.7197" longLatPrecision="1" longitude="-96.16109" municipality="La Jeringa" name="La Jeringa" pageId="4" pageNumber="15" stateProvince="Oaxaca">La Jeringa</location>
,
<location id="8E9E0CC15326FFE9FDAB9C26FD72C67D" LSID="urn:lsid:plazi:treatment:03E8EB0C5321FFE1FDC89B70FEE3C3CB:8E9E0CC15326FFE9FDAB9C26FD72C67D" box="[516,700,580,606]" country="Mexico" latitude="15.7197" longLatPrecision="1" longitude="-96.16109" municipality="La Jeringa" name="Cacaluta Island" pageId="4" pageNumber="15" stateProvince="Oaxaca">Cacaluta Island</location>
,
<collectingRegion id="498594F85326FFE9FD689C26FCEDC67D" box="[711,803,580,606]" country="Mexico" name="Oaxaca" pageId="4" pageNumber="15">Oaxaca</collectingRegion>
;
<geoCoordinate id="EE753CDD5326FFE9FC829C26FC26C67C" box="[813,1000,580,607]" degrees="15" direction="north" minutes="43" orientation="latitude" pageId="4" pageNumber="15" precision="1" seconds="10.92" value="15.7197">15°43´10.92˝ N</geoCoordinate>
,
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;
<quantity id="4CB9F7FF5326FFE9FB119C26FAD7C67C" box="[1214,1305,580,607]" metricMagnitude="1" metricUnit="m" metricValue="1.15" metricValueMax="1.4" metricValueMin="0.9" pageId="4" pageNumber="15" unit="m" value="11.5" valueMax="14.0" valueMin="9.0">
<elevation id="006CBD295326FFE9FB119C26FAD7C67C" box="[1214,1305,580,607]" metricMagnitude="1" metricUnit="m" metricValue="1.15" metricValueMax="1.4" metricValueMin="0.9" pageId="4" pageNumber="15" unit="m" value="11.5" valueMax="14.0" valueMin="9.0">914 m</elevation>
</quantity>
;
<date id="FFFF7CDA5326FFE9FA8B9C26FF35C6A1" pageId="4" pageNumber="15" value="2016-01-26">
<collectingDate id="EFBB85325326FFE9FA8B9C26FF35C6A1" pageId="4" pageNumber="15" value="2016-01-26">26 Jan. 2016</collectingDate>
</date>
;
<collectorName id="26B43FCC5326FFE9FEA99C0AFEB8C6A1" box="[262,374,616,642]" pageId="4" pageNumber="15">D.Rangel</collectorName>
and
<collectorName id="26B43FCC5326FFE9FE029C0BFDD0C6A1" box="[429,542,616,643]" pageId="4" pageNumber="15">R. Granja</collectorName>
leg.;
<collectedFrom id="385D19E75326FFE9FDF19C0AFC95C6A1" box="[606,859,616,642]" pageId="4" pageNumber="15">sediment under rocks</collectedFrom>
;
<specimenCode id="DBE7F2615326FFE9FCC99C0AFBACC6A1" box="[870,1122,616,642]" pageId="4" pageNumber="15">ICML-UNAM 18319</specimenCode>
</materialsCitation>
<materialsCitation id="3B2950475326FFE9FBD69C0AFC80C6E9" ID-GBIF-Occurrence="2995182306" collectedFrom="under rocks" collectingDate="2016-10-28" collectionCode="R" collectorName="R. Granja" country="Mexico" county="Punta Lobos" elevation="16" latitude="24.47474" location="Espiritu Santo Island" longLatPrecision="1" longitude="-110.28907" municipality="Espiritu Santo Island" pageId="4" pageNumber="15" specimenCode="ICML-UNAM 18320" specimenCount="2">
<specimenCount id="9D4791935326FFE9FBD69C0AFB1FC6A0" box="[1145,1233,616,643]" pageId="4" pageNumber="15">2 specs</specimenCount>
;
<collectingCounty id="629F22965326FFE9FB739C0BFABCC6A1" box="[1244,1394,616,643]" pageId="4" pageNumber="15">Punta Lobos</collectingCounty>
,
<location id="8E9E0CC15326FFE9FF129CEEFE74C685" LSID="urn:lsid:plazi:treatment:03E8EB0C5321FFE1FDC89B70FEE3C3CB:8E9E0CC15326FFE9FF129CEEFE74C685" box="[189,442,652,678]" country="Mexico" county="Punta Lobos" latitude="24.47474" longLatPrecision="1" longitude="-110.28907" municipality="Espiritu Santo Island" name="Espiritu Santo Island" pageId="4" pageNumber="15">Espíritu Santo Island</location>
,
<location id="8E9E0CC15326FFE9FE689CEEFD6CC685" LSID="urn:lsid:plazi:treatment:03E8EB0C5321FFE1FDC89B70FEE3C3CB:8E9E0CC15326FFE9FE689CEEFD6CC685" box="[455,674,652,678]" country="Mexico" county="Punta Lobos" latitude="24.47474" longLatPrecision="1" longitude="-110.28907" municipality="Espiritu Santo Island" name="Gulf of California" pageId="4" pageNumber="15">Gulf of California</location>
;
<geoCoordinate id="EE753CDD5326FFE9FD009CEEFCA2C685" box="[687,876,652,678]" degrees="24" direction="north" minutes="28" orientation="latitude" pageId="4" pageNumber="15" precision="1" seconds="29.06" value="24.47474">24°28´29.06˝ N</geoCoordinate>
,
<geoCoordinate id="EE753CDD5326FFE9FCD59CE9FB84C685" box="[890,1098,651,678]" degrees="110" direction="west" minutes="17" orientation="longitude" pageId="4" pageNumber="15" precision="1" seconds="20.65" value="-110.28907">110°17´20.65˝ W</geoCoordinate>
;
<quantity id="4CB9F7FF5326FFE9FBF79CE9FB0AC685" box="[1112,1220,651,678]" metricMagnitude="1" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="4" pageNumber="15" unit="m" value="16.0" valueMax="17.0" valueMin="15.0">
<elevation id="006CBD295326FFE9FBF79CE9FB0AC685" box="[1112,1220,651,678]" metricMagnitude="1" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="4" pageNumber="15" unit="m" value="16.0" valueMax="17.0" valueMin="15.0">1517 m</elevation>
</quantity>
;
<date id="FFFF7CDA5326FFE9FB7D9CEEFABCC685" box="[1234,1394,652,678]" pageId="4" pageNumber="15" value="2016-10-28">
<collectingDate id="EFBB85325326FFE9FB7D9CEEFABCC685" box="[1234,1394,652,678]" pageId="4" pageNumber="15" value="2016-10-28">28 Oct. 2016</collectingDate>
</date>
;
<collectorName id="26B43FCC5326FFE9FF129CD2FEE1C6EA" box="[189,303,687,714]" pageId="4" pageNumber="15">R. Granja</collectorName>
leg.;
<collectedFrom id="385D19E75326FFE9FEC09CCDFE35C6EA" box="[367,507,687,713]" pageId="4" pageNumber="15">under rocks</collectedFrom>
, 28°C;
<specimenCode id="DBE7F2615326FFE9FDFD9CD2FC80C6E9" box="[594,846,688,714]" pageId="4" pageNumber="15">ICML-UNAM 18320</specimenCode>
</materialsCitation>
<materialsCitation id="3B2950475326FFE9FCC99CD2FEA8C732" ID-GBIF-Occurrence="2995182302" collectedFrom="under rocks" collectingDate="2016-10-29" collectionCode="R, ICML-" collectorName="R. Granja" country="Mexico" elevation="13" latitude="25.2043" location="Gulf of California" longLatPrecision="1" longitude="-110.6949" municipality="San Diego Norte" pageId="4" pageNumber="15" specimenCode="ICML- UNAM 18321" specimenCount="3">
<specimenCount id="9D4791935326FFE9FCC99CD2FC70C6E9" box="[870,958,688,714]" pageId="4" pageNumber="15">3 specs</specimenCount>
;
<collectingMunicipality id="6B9AC0605326FFE9FC659CD2FB5FC6E9" box="[970,1169,687,714]" pageId="4" pageNumber="15">San Diego Norte</collectingMunicipality>
,
<location id="8E9E0CC15326FFE9FB339CCDFABCC6EA" LSID="urn:lsid:plazi:treatment:03E8EB0C5321FFE1FDC89B70FEE3C3CB:8E9E0CC15326FFE9FB339CCDFABCC6EA" box="[1180,1394,687,713]" country="Mexico" latitude="25.2043" longLatPrecision="1" longitude="-110.6949" municipality="San Diego Norte" name="Gulf of California" pageId="4" pageNumber="15">Gulf of California</location>
;
<geoCoordinate id="EE753CDD5326FFE9FF129CB1FEB2C6CD" box="[189,380,723,750]" degrees="25" direction="north" minutes="12" orientation="latitude" pageId="4" pageNumber="15" precision="1" seconds="15.48" value="25.2043">25°12´15.48˝ N</geoCoordinate>
,
<geoCoordinate id="EE753CDD5326FFE9FE239CB1FD90C6CD" box="[396,606,723,750]" degrees="110" direction="west" minutes="41" orientation="longitude" pageId="4" pageNumber="15" precision="1" seconds="41.64" value="-110.6949">110°41´41.64˝ W</geoCoordinate>
;
<quantity id="4CB9F7FF5326FFE9FDC19CB1FD7EC6CD" box="[622,688,723,750]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="15" unit="m" value="13.0">
<elevation id="006CBD295326FFE9FDC19CB1FD7EC6CD" box="[622,688,723,750]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="15" unit="m" value="13.0">13 m</elevation>
</quantity>
;
<date id="FFFF7CDA5326FFE9FD6F9CB1FCAAC6CE" box="[704,868,723,749]" pageId="4" pageNumber="15" value="2016-10-29">
<collectingDate id="EFBB85325326FFE9FD6F9CB1FCAAC6CE" box="[704,868,723,749]" pageId="4" pageNumber="15" value="2016-10-29">29 Oct. 2016</collectingDate>
</date>
;
<collectorName id="26B43FCC5326FFE9FCDB9CB6FC24C6CE" box="[884,1002,723,750]" pageId="4" pageNumber="15">R. Granja</collectorName>
leg.;
<collectedFrom id="385D19E75326FFE9FB9C9CB1FB0DC6CE" box="[1075,1219,723,749]" pageId="4" pageNumber="15">under rocks</collectedFrom>
, 29°C;
<specimenCode id="DBE7F2615326FFE9FA8C9CB1FEA8C732" pageId="4" pageNumber="15">ICML- UNAM 18321</specimenCode>
</materialsCitation>
<materialsCitation id="3B2950475326FFE9FED19C95FC37C716" ID-GBIF-Occurrence="2995182314" collectedFrom="under rocks" collectingDate="2016-10-31" collectionCode="R" collectorName="R. Granja" country="Mexico" elevation="12" latitude="25.378761" location="Gulf of California" longLatPrecision="1" longitude="-110.99284" municipality="San Mateo Norte" pageId="4" pageNumber="15" specimenCode="ICML-UNAM 18322" specimenCount="1">
<specimenCount id="9D4791935326FFE9FED19C95FE1DC732" box="[382,467,759,785]" pageId="4" pageNumber="15" type="generic">1 spec.</specimenCount>
;
<collectingMunicipality id="6B9AC0605326FFE9FE719C95FD66C732" box="[478,680,759,785]" pageId="4" pageNumber="15">San Mateo Norte</collectingMunicipality>
,
<location id="8E9E0CC15326FFE9FD1C9C95FC44C732" LSID="urn:lsid:plazi:treatment:03E8EB0C5321FFE1FDC89B70FEE3C3CB:8E9E0CC15326FFE9FD1C9C95FC44C732" box="[691,906,759,785]" country="Mexico" latitude="25.378761" longLatPrecision="1" longitude="-110.99284" municipality="San Mateo Norte" name="Gulf of California" pageId="4" pageNumber="15">Gulf of California</location>
;
<geoCoordinate id="EE753CDD5326FFE9FC3B9C95FB81C732" box="[916,1103,759,786]" degrees="25" direction="north" minutes="22" orientation="latitude" pageId="4" pageNumber="15" precision="1" seconds="43.54" value="25.378761">25°22´43.54˝ N</geoCoordinate>
,
<geoCoordinate id="EE753CDD5326FFE9FBF59C95FAE6C732" box="[1114,1320,759,786]" degrees="110" direction="west" minutes="59" orientation="longitude" pageId="4" pageNumber="15" precision="1" seconds="34.26" value="-110.99284">110°59´34.26˝ W</geoCoordinate>
;
<quantity id="4CB9F7FF5326FFE9FA9C9C95FABFC732" box="[1331,1393,759,785]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="15" unit="m" value="12.0">
<elevation id="006CBD295326FFE9FA9C9C95FABFC732" box="[1331,1393,759,785]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="15" unit="m" value="12.0">12 m</elevation>
</quantity>
;
<date id="FFFF7CDA5326FFE9FF129D79FE96C717" box="[189,344,794,821]" pageId="4" pageNumber="15" value="2016-10-31">
<collectingDate id="EFBB85325326FFE9FF129D79FE96C717" box="[189,344,794,821]" pageId="4" pageNumber="15" value="2016-10-31">31 Oct. 2016</collectingDate>
</date>
;
<collectorName id="26B43FCC5326FFE9FECB9D79FE19C717" box="[356,471,794,821]" pageId="4" pageNumber="15">R. Granja</collectorName>
leg.;
<collectedFrom id="385D19E75326FFE9FDB79D78FD6AC717" box="[536,676,794,820]" pageId="4" pageNumber="15">under rocks</collectedFrom>
, 29°C;
<specimenCode id="DBE7F2615326FFE9FD539D79FC37C716" box="[764,1017,795,821]" pageId="4" pageNumber="15">ICML-UNAM 18322</specimenCode>
</materialsCitation>
<materialsCitation id="3B2950475326FFE9FBBD9D79FD59C75F" ID-GBIF-Occurrence="2995182310" collectedFrom="under rocks" collectingDate="2008-08-31" collectionCode="ICML-UNAM" collectorName="F. Solis &amp; Solis-Wolfowitz" country="Mexico" elevation="11" latitude="25.20355" location="Gulf of California" longLatPrecision="1" longitude="-110.69448" municipality="Isla San Diego" pageId="4" pageNumber="15" specimenCode="ICML-UNAM 18323" specimenCount="10">
<specimenCount id="9D4791935326FFE9FBBD9D79FBB4C716" box="[1042,1146,795,821]" pageId="4" pageNumber="15">10 specs</specimenCount>
;
<collectingMunicipality id="6B9AC0605326FFE9FB299D78FAFBC717" box="[1158,1333,794,821]" pageId="4" pageNumber="15">Isla San Diego</collectingMunicipality>
,
<location id="8E9E0CC15326FFE9FAEE9D78FE96C77B" LSID="urn:lsid:plazi:treatment:03E8EB0C5321FFE1FDC89B70FEE3C3CB:8E9E0CC15326FFE9FAEE9D78FE96C77B" country="Mexico" latitude="25.20355" longLatPrecision="1" longitude="-110.69448" municipality="Isla San Diego" name="Gulf of California" pageId="4" pageNumber="15">Gulf of California</location>
;
<geoCoordinate id="EE753CDD5326FFE9FECC9D5CFDD0C77A" box="[355,542,830,857]" degrees="25" direction="north" minutes="12" orientation="latitude" pageId="4" pageNumber="15" precision="1" seconds="12.78" value="25.20355">25°12´12.78˝ N</geoCoordinate>
,
<geoCoordinate id="EE753CDD5326FFE9FD859D5CFD36C77A" box="[554,760,830,857]" degrees="110" direction="west" minutes="41" orientation="longitude" pageId="4" pageNumber="15" precision="1" seconds="40.14" value="-110.69448">110°41´40.14˝ W</geoCoordinate>
;
<quantity id="4CB9F7FF5326FFE9FCAB9D5CFCA3C77A" box="[772,877,830,857]" metricMagnitude="1" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="4" pageNumber="15" unit="m" value="10.5" valueMax="11.0" valueMin="10.0">
<elevation id="006CBD295326FFE9FCAB9D5CFCA3C77A" box="[772,877,830,857]" metricMagnitude="1" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="4" pageNumber="15" unit="m" value="10.5" valueMax="11.0" valueMin="10.0">1011 m</elevation>
</quantity>
;
<date id="FFFF7CDA5326FFE9FCD69D5CFBD5C77B" box="[889,1051,830,856]" pageId="4" pageNumber="15" value="2008-08-31">
<collectingDate id="EFBB85325326FFE9FCD69D5CFBD5C77B" box="[889,1051,830,856]" pageId="4" pageNumber="15" value="2008-08-31">31 Aug. 2008</collectingDate>
</date>
;
<collectorName id="26B43FCC5326FFE9FB889D5DFBB1C77B" box="[1063,1151,830,857]" pageId="4" pageNumber="15">F. Solís</collectorName>
and
<collectorName id="26B43FCC5326FFE9FB189D5CFAB9C77B" box="[1207,1399,830,856]" pageId="4" pageNumber="15">Solís-Wolfowitz</collectorName>
leg.;
<collectedFrom id="385D19E75326FFE9FF559D00FE47C75F" box="[250,393,866,892]" pageId="4" pageNumber="15">under rocks</collectedFrom>
;
<specimenCode id="DBE7F2615326FFE9FE389D00FD59C75F" box="[407,663,866,892]" collectionCode="ICML-UNAM" pageId="4" pageNumber="15">ICML-UNAM 18323</specimenCode>
</materialsCitation>
<materialsCitation id="3B2950475326FFE9FD1B9D00FEC8C7E0" ID-GBIF-Occurrence="2995182318" collectedFrom="under rocks" collectingDate="2011-08-12" collectionCode="ICML-UNAM" collectorName="C. Sanchez" country="Mexico" county="El Peruano" elevation="12" latitude="27.90835" location="Gulf of California" longLatPrecision="1" longitude="-110.969635" municipality="Guaymas" pageId="4" pageNumber="15" specimenCode="ICML-UNAM 18324" specimenCount="5" stateProvince="Sonora">
<specimenCount id="9D4791935326FFE9FD1B9D00FCDEC75F" box="[692,784,866,893]" pageId="4" pageNumber="15">5 specs</specimenCount>
;
<collectingCounty id="629F22965326FFE9FCB19D00FC6AC75F" box="[798,932,866,892]" pageId="4" pageNumber="15">El Peruano</collectingCounty>
,
<collectingMunicipality id="6B9AC0605326FFE9FC1D9D00FBEAC75F" box="[946,1060,866,892]" pageId="4" pageNumber="15">Guaymas</collectingMunicipality>
,
<collectingRegion id="498594F85326FFE9FB9D9D00FB46C75F" box="[1074,1160,866,892]" country="Mexico" name="Sonora" pageId="4" pageNumber="15">Sonora</collectingRegion>
,
<location id="8E9E0CC15326FFE9FB399D00FABCC75F" LSID="urn:lsid:plazi:treatment:03E8EB0C5321FFE1FDC89B70FEE3C3CB:8E9E0CC15326FFE9FB399D00FABCC75F" box="[1174,1394,866,892]" country="Mexico" county="El Peruano" latitude="27.90835" longLatPrecision="1" longitude="-110.969635" municipality="Guaymas" name="Gulf of California" pageId="4" pageNumber="15" stateProvince="Sonora">Gulf of California</location>
;
<geoCoordinate id="EE753CDD5326FFE9FF129DE7FEB4C783" box="[189,378,901,928]" degrees="27" direction="north" minutes="54" orientation="latitude" pageId="4" pageNumber="15" precision="1" seconds="30.06" value="27.90835">27°54´30.06˝ N</geoCoordinate>
,
<geoCoordinate id="EE753CDD5326FFE9FE289DE7FD99C783" box="[391,599,901,928]" degrees="110" direction="west" minutes="58" orientation="longitude" pageId="4" pageNumber="15" precision="1" seconds="10.68" value="-110.969635">110°58´10.68˝ W</geoCoordinate>
;
<quantity id="4CB9F7FF5326FFE9FDCB9DE4FD6BC783" box="[612,677,902,928]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="15" unit="m" value="12.0">
<elevation id="006CBD295326FFE9FDCB9DE4FD6BC783" box="[612,677,902,928]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="15" unit="m" value="12.0">12 m</elevation>
</quantity>
;
<date id="FFFF7CDA5326FFE9FD1D9DE4FC99C783" box="[690,855,902,928]" pageId="4" pageNumber="15" value="2011-08-12">
<collectingDate id="EFBB85325326FFE9FD1D9DE4FC99C783" box="[690,855,902,928]" pageId="4" pageNumber="15" value="2011-08-12">12 Aug. 2011</collectingDate>
</date>
;
<collectorName id="26B43FCC5326FFE9FCCB9DE4FC24C783" box="[868,1002,902,928]" pageId="4" pageNumber="15">C. Sánchez</collectorName>
leg.;
<collectedFrom id="385D19E75326FFE9FB819DE4FB72C783" box="[1070,1212,902,928]" pageId="4" pageNumber="15">under rocks</collectedFrom>
,
<specimenCode id="DBE7F2615326FFE9FB659DE4FEC8C7E0" collectionCode="ICML-UNAM" pageId="4" pageNumber="15">ICML-UNAM 18324</specimenCode>
</materialsCitation>
<materialsCitation id="3B2950475326FFE9FE8B9DCBFC07C7CB" ID-GBIF-Occurrence="2995182301" collectedFrom="under rocks" collectingDate="2010-03-02" collectionCode="ICML-UNAM" collectorName="F. Solis" country="Mexico" county="La Lobera" elevation="7" latitude="24.615664" location="La Lobera" longLatPrecision="1" longitude="-110.40016" municipality="Bahia de La Paz" pageId="4" pageNumber="15" specimenCode="ICML-UNAM 18325" specimenCount="1">
<specimenCount id="9D4791935326FFE9FE8B9DCBFEB5C7E7" box="[292,379,937,964]" pageId="4" pageNumber="15" type="generic">1 spec.</specimenCount>
;
<location id="8E9E0CC15326FFE9FE269DC8FDC9C7E0" LSID="urn:lsid:plazi:treatment:03E8EB0C5321FFE1FDC89B70FEE3C3CB:8E9E0CC15326FFE9FE269DC8FDC9C7E0" box="[393,519,937,964]" country="Mexico" county="La Lobera" latitude="24.615664" longLatPrecision="1" longitude="-110.40016" municipality="Bahia de La Paz" name="La Lobera" pageId="4" pageNumber="15">La Lobera</location>
,
<location id="8E9E0CC15326FFE9FDBA9DCBFD10C7E7" LSID="urn:lsid:plazi:treatment:03E8EB0C5321FFE1FDC89B70FEE3C3CB:8E9E0CC15326FFE9FDBA9DCBFD10C7E7" box="[533,734,937,964]" country="Mexico" county="La Lobera" latitude="24.615664" longLatPrecision="1" longitude="-110.40016" municipality="Bahia de La Paz" name="Bahia de La Paz" pageId="4" pageNumber="15">Bahía de La Paz</location>
,
<location id="8E9E0CC15326FFE9FD439DCBFC06C7E0" LSID="urn:lsid:plazi:treatment:03E8EB0C5321FFE1FDC89B70FEE3C3CB:8E9E0CC15326FFE9FD439DCBFC06C7E0" box="[748,968,937,963]" country="Mexico" county="La Lobera" latitude="24.615664" longLatPrecision="1" longitude="-110.40016" municipality="Bahia de La Paz" name="Gulf of California" pageId="4" pageNumber="15">Gulf of California</location>
;
<geoCoordinate id="EE753CDD5326FFE9FC799DCBFB5DC7E7" box="[982,1171,937,964]" degrees="24" direction="north" minutes="36" orientation="latitude" pageId="4" pageNumber="15" precision="1" seconds="56.39" value="24.615664">24°36´56.39˝ N</geoCoordinate>
,
<geoCoordinate id="EE753CDD5326FFE9FB0E9DCBFABCC7E7" box="[1185,1394,937,964]" degrees="110" direction="west" minutes="24" orientation="longitude" pageId="4" pageNumber="15" precision="1" seconds="00.57" value="-110.40016">110°24´00.57˝ W</geoCoordinate>
;
<quantity id="4CB9F7FF5326FFE9FF129DACFF23C7CB" box="[189,237,974,1000]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="15" unit="m" value="7.0">
<elevation id="006CBD295326FFE9FF129DACFF23C7CB" box="[189,237,974,1000]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="15" unit="m" value="7.0">7 m</elevation>
</quantity>
;
<date id="FFFF7CDA5326FFE9FF559DAFFE42C7C4" box="[250,396,973,1000]" pageId="4" pageNumber="15" value="2010-03-02">
<collectingDate id="EFBB85325326FFE9FF559DAFFE42C7C4" box="[250,396,973,1000]" pageId="4" pageNumber="15" value="2010-03-02">2 Mar. 2010</collectingDate>
</date>
;
<collectorName id="26B43FCC5326FFE9FE369DACFE3EC7C4" box="[409,496,973,1000]" pageId="4" pageNumber="15">F. Solís</collectorName>
leg.;
<collectedFrom id="385D19E75326FFE9FD9C9DAFFD71C7C4" box="[563,703,973,999]" pageId="4" pageNumber="15">under rocks</collectedFrom>
,
<specimenCode id="DBE7F2615326FFE9FD649DAFFC07C7CB" box="[715,969,973,1000]" collectionCode="ICML-UNAM" pageId="4" pageNumber="15">ICML-UNAM 18325</specimenCode>
</materialsCitation>
<materialsCitation id="3B2950475326FFE9FC4C9DAFFDC6C00D" ID-GBIF-Occurrence="2995182320" collectedFrom="under rocks" collectingDate="1996-03-22" collectionCode="ICML-UNAM" collectorName="F. Solis &amp; C. Nepote" country="Mexico" county="Isla Larga" elevation="9" latitude="20.685555" location="Isla Larga" longLatPrecision="1" longitude="-105.566666" municipality="Este" pageId="4" pageNumber="15" specimenCode="ICML-UNAM 18326" specimenCount="1">
<specimenCount id="9D4791935326FFE9FC4C9DAFFBF6C7CB" box="[995,1080,973,1000]" pageId="4" pageNumber="15" type="generic">1 spec.</specimenCount>
;
<location id="8E9E0CC15326FFE9FBEB9DAFFB74C7CB" LSID="urn:lsid:plazi:treatment:03E8EB0C5321FFE1FDC89B70FEE3C3CB:8E9E0CC15326FFE9FBEB9DAFFB74C7CB" box="[1092,1210,973,1000]" country="Mexico" county="Isla Larga" latitude="20.685555" longLatPrecision="1" longitude="-105.566666" municipality="Este" name="Isla Larga" pageId="4" pageNumber="15">Isla Larga</location>
(estación Este),
<location id="8E9E0CC15326FFE9FF129D93FEB1C028" LSID="urn:lsid:plazi:treatment:03E8EB0C5321FFE1FDC89B70FEE3C3CB:8E9E0CC15326FFE9FF129D93FEB1C028" box="[189,383,1009,1035]" country="Mexico" county="Isla Larga" latitude="20.685555" longLatPrecision="1" longitude="-105.566666" municipality="Este" name="Marietas Islands" pageId="4" pageNumber="15">Marietas Islands</location>
;
<geoCoordinate id="EE753CDD5326FFE9FE279D93FD8FC028" box="[392,577,1009,1035]" degrees="20" direction="north" minutes="41" orientation="latitude" pageId="4" pageNumber="15" precision="1" seconds="08.00" value="20.685555">20°41´08.00˝ N</geoCoordinate>
,
<geoCoordinate id="EE753CDD5326FFE9FDE59D92FCD8C028" box="[586,790,1008,1035]" degrees="105" direction="west" minutes="34" orientation="longitude" pageId="4" pageNumber="15" precision="1" seconds="00.00" value="-105.566666">105°34´00.00˝ W</geoCoordinate>
;
<quantity id="4CB9F7FF5326FFE9FC8F9D93FC83C028" box="[800,845,1009,1035]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="15" unit="m" value="9.0">
<elevation id="006CBD295326FFE9FC8F9D93FC83C028" box="[800,845,1009,1035]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="15" unit="m" value="9.0">9 m</elevation>
</quantity>
;
<date id="FFFF7CDA5326FFE9FCF99D93FC3CC028" box="[854,1010,1009,1035]" pageId="4" pageNumber="15" value="1996-03-22">
<collectingDate id="EFBB85325326FFE9FCF99D93FC3CC028" box="[854,1010,1009,1035]" pageId="4" pageNumber="15" value="1996-03-22">22 Mar. 1996</collectingDate>
</date>
;
<collectorName id="26B43FCC5326FFE9FC549D93FB9EC028" box="[1019,1104,1009,1035]" pageId="4" pageNumber="15">F. Solís</collectorName>
and
<collectorName id="26B43FCC5326FFE9FB2C9D93FB39C028" box="[1155,1271,1009,1035]" pageId="4" pageNumber="15">C. Nepote</collectorName>
leg.;
<collectedFrom id="385D19E75326FFE9FA9B9D93FECEC00D" pageId="4" pageNumber="15">under rocks</collectedFrom>
;
<specimenCode id="DBE7F2615326FFE9FEA49A77FDC6C00D" box="[267,520,1044,1071]" collectionCode="ICML-UNAM" pageId="4" pageNumber="15">ICML-UNAM 18326</specimenCode>
</materialsCitation>
<materialsCitation id="3B2950475326FFE9FDB09A77FAABC071" ID-GBIF-Occurrence="2995182324" collectedFrom="sediment under rocks" collectingDate="1997-04-28" collectionCode="ICML-UNAM" collectorName="H. Cortes" country="Mexico" county="Los Islotes" elevation="18" latitude="24.583334" location="Los Islotes" longLatPrecision="1" longitude="-110.416664" municipality="Bahia de La Paz" pageId="4" pageNumber="15" specimenCode="ICML-UNAM 18327" specimenCount="1">
<specimenCount id="9D4791935326FFE9FDB09A77FDBDC00C" box="[543,627,1045,1071]" pageId="4" pageNumber="15" type="generic">1 spec.</specimenCount>
;
<location id="8E9E0CC15326FFE9FDD09A77FCCEC00D" LSID="urn:lsid:plazi:treatment:03E8EB0C5321FFE1FDC89B70FEE3C3CB:8E9E0CC15326FFE9FDD09A77FCCEC00D" box="[639,768,1044,1071]" country="Mexico" county="Los Islotes" latitude="24.583334" longLatPrecision="1" longitude="-110.416664" municipality="Bahia de La Paz" name="Los Islotes" pageId="4" pageNumber="15">Los Islotes</location>
,
<location id="8E9E0CC15326FFE9FCA39A76FC05C00C" LSID="urn:lsid:plazi:treatment:03E8EB0C5321FFE1FDC89B70FEE3C3CB:8E9E0CC15326FFE9FCA39A76FC05C00C" box="[780,971,1044,1071]" country="Mexico" county="Los Islotes" latitude="24.583334" longLatPrecision="1" longitude="-110.416664" municipality="Bahia de La Paz" name="Bahia de La Paz" pageId="4" pageNumber="15">Bahía de La Paz</location>
,
<location id="8E9E0CC15326FFE9FC789A76FB63C00D" LSID="urn:lsid:plazi:treatment:03E8EB0C5321FFE1FDC89B70FEE3C3CB:8E9E0CC15326FFE9FC789A76FB63C00D" box="[983,1197,1044,1070]" country="Mexico" county="Los Islotes" latitude="24.583334" longLatPrecision="1" longitude="-110.416664" municipality="Bahia de La Paz" name="Gulf of California" pageId="4" pageNumber="15">Gulf of California</location>
;
<geoCoordinate id="EE753CDD5326FFE9FB189A76FABCC00C" box="[1207,1394,1044,1071]" degrees="24" direction="north" minutes="35" orientation="latitude" pageId="4" pageNumber="15" precision="1" seconds="00.00" value="24.583334">24°35´00.00˝ N</geoCoordinate>
,
<geoCoordinate id="EE753CDD5326FFE9FF129A5AFE45C070" box="[189,395,1080,1107]" degrees="110" direction="west" minutes="25" orientation="longitude" pageId="4" pageNumber="15" precision="1" seconds="00.00" value="-110.416664">110°25´00.00˝ W</geoCoordinate>
;
<quantity id="4CB9F7FF5326FFE9FE389A5AFE22C070" box="[407,492,1080,1107]" metricMagnitude="1" metricUnit="m" metricValue="1.83" pageId="4" pageNumber="15" unit="m" value="18.3">
<elevation id="006CBD295326FFE9FE389A5AFE22C070" box="[407,492,1080,1107]" metricMagnitude="1" metricUnit="m" metricValue="1.83" pageId="4" pageNumber="15" unit="m" value="18.3">18.3 m</elevation>
</quantity>
;
<date id="FFFF7CDA5326FFE9FE579A5AFD55C071" box="[504,667,1080,1106]" pageId="4" pageNumber="15" value="1997-04-28">
<collectingDate id="EFBB85325326FFE9FE579A5AFD55C071" box="[504,667,1080,1106]" pageId="4" pageNumber="15" value="1997-04-28">28 Apr. 1997</collectingDate>
</date>
;
<collectorName id="26B43FCC5326FFE9FD079A5BFCD6C071" box="[680,792,1080,1107]" pageId="4" pageNumber="15">H. Cortés</collectorName>
leg.;
<collectedFrom id="385D19E75326FFE9FCF59A5AFB96C071" box="[858,1112,1080,1106]" pageId="4" pageNumber="15">sediment under rocks</collectedFrom>
;
<specimenCode id="DBE7F2615326FFE9FBCB9A5AFAABC071" box="[1124,1381,1080,1106]" collectionCode="ICML-UNAM" pageId="4" pageNumber="15">ICML-UNAM 18327</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BFE5A1A5326FFE9FF129A18FACEC0B7" blockId="4.[189,1280,1146,1173]" box="[189,1280,1146,1173]" pageId="4" pageNumber="15">
<materialsCitation id="3B2950475326FFE9FF129A18FB32C0B7" ID-GBIF-Occurrence="2995182303" box="[189,1276,1146,1173]" collectingDate="1872-06-24" collectingDateMax="1872-07-23" collectingDateMin="1872-06-24" collectionCode="MCZ" country="Panama" location="Panama" pageId="4" pageNumber="15" specimenCode="MCZ OPH-116" specimenCount="2" stateProvince="Panama">
<collectingCountry id="F3561A8A5326FFE9FF129A18FEF6C0B7" box="[189,312,1146,1172]" name="Panama" pageId="4" pageNumber="15">PANAMA</collectingCountry>
<specimenCount id="9D4791935326FFE9FEFE9A18FE64C0B6" box="[337,426,1146,1173]" pageId="4" pageNumber="15">2 specs</specimenCount>
;
<collectingRegion id="498594F85326FFE9FE199A19FDD9C0B6" box="[438,535,1147,1173]" country="Panama" name="Panama" pageId="4" pageNumber="15">Panama</collectingRegion>
;
<date id="FFFF7CDA5326FFE9FD8D9A18FCD7C0B7" box="[546,793,1146,1172]" pageId="4" pageNumber="15" value="1872-06-24" valueMax="1872-07-23" valueMin="1872-06-24">
<collectingDate id="EFBB85325326FFE9FD8D9A18FCD7C0B7" box="[546,793,1146,1172]" pageId="4" pageNumber="15" value="1872-06-24" valueMax="1872-07-23" valueMin="1872-06-24">24 Jun.23 Jul. 1872</collectingDate>
</date>
; Hassler expedition leg.;
<specimenCode id="DBE7F2615326FFE9FBED9A18FB32C0B7" box="[1090,1276,1146,1172]" pageId="4" pageNumber="15">MCZ OPH-116</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BFE5A1A5326FFE9FF459ADEFE68C0F5" blockId="4.[189,536,1212,1274]" box="[234,422,1212,1238]" pageId="4" pageNumber="15">
<emphasis id="B93586085326FFE9FF459ADEFE68C0F5" bold="true" box="[234,422,1212,1238]" pageId="4" pageNumber="15">Other material</emphasis>
</paragraph>
<paragraph id="8BFE5A1A5326FFE9FF129A82FDD6C0D9" blockId="4.[189,536,1212,1274]" box="[189,536,1248,1274]" pageId="4" pageNumber="15">See: Supplementary material.</paragraph>
</subSubSection>
<subSubSection id="C35B09915326FFE1FF129B40FBECC5B7" lastPageId="12" lastPageNumber="23" pageId="4" pageNumber="15" type="description">
<paragraph id="8BFE5A1A5326FFE9FF129B40FE0CC118" blockId="4.[189,450,1313,1340]" box="[189,450,1313,1340]" pageId="4" pageNumber="15">
<heading id="D0B6ED765326FFE9FF129B40FE0CC118" bold="true" box="[189,450,1313,1340]" fontSize="11" level="3" pageId="4" pageNumber="15" reason="3">
<emphasis id="B93586085326FFE9FF129B40FE0CC118" bold="true" box="[189,450,1313,1340]" pageId="4" pageNumber="15">
<typeStatus id="54FAE4B85326FFE9FF129B40FEE1C11F" box="[189,303,1314,1340]" pageId="4" pageNumber="15" type="holotype">Holotype</typeStatus>
description
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFE5A1A5326FFE9FF129B33FD7EC246" blockId="4.[189,1399,1360,1637]" pageId="4" pageNumber="15">
DD =
<quantity id="4CB9F7FF5326FFE9FEAC9B33FEA2C148" box="[259,364,1361,1387]" metricMagnitude="-2" metricUnit="m" metricValue="1.48" pageId="4" pageNumber="15" unit="mm" value="14.8">14.8 mm</quantity>
; 5 arms, AL =
<quantity id="4CB9F7FF5326FFE9FDBF9B33FDB7C148" box="[528,633,1361,1387]" metricMagnitude="-2" metricUnit="m" metricValue="6.909999999999999" pageId="4" pageNumber="15" unit="mm" value="69.1">69.1 mm</quantity>
. Disc flat, nearly circular. Dorsal disc densely covered by rounded, small granules, slightly separated from each other; granule density 113
<superScript id="7C34F7525326FFE9FBA49B17FB88C1A1" attach="left" box="[1035,1094,1396,1423]" fontSize="6" pageId="4" pageNumber="15">mm-2</superScript>
. Granules at periphery of disc and on base of arms slightly larger than on central part of disc. Radial shields covered by granules (
<figureCitation id="137A469F5326FFE9FF699BDEFEEBC1F5" box="[198,293,1468,1494]" captionStart="Fig" captionStartId="5.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@5.[189,1398,291,1855]" captionTargetPageId="5" captionText="Fig. 1. Ophioderma hendleri sp. nov., holotype (ICML-UNAM 18315, DD = 14.8 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Structures of a jaw (according to Hendler 2018). E. Basal dorsal arm. F. Dorsal arm. G. Ventral arm. H. Lateral arm plates and arm spines. I. Dorsal view. See Material and methods for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4326613" httpUri="https://zenodo.org/record/4326613/files/figure.png" pageId="4" pageNumber="15">Fig. 1A</figureCitation>
). Interradii covered with granulation, similar to periphery and base of arms (
<figureCitation id="137A469F5326FFE9FB679BDEFAE8C1F5" box="[1224,1318,1468,1494]" captionStart="Fig" captionStartId="5.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@5.[189,1398,291,1855]" captionTargetPageId="5" captionText="Fig. 1. Ophioderma hendleri sp. nov., holotype (ICML-UNAM 18315, DD = 14.8 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Structures of a jaw (according to Hendler 2018). E. Basal dorsal arm. F. Dorsal arm. G. Ventral arm. H. Lateral arm plates and arm spines. I. Dorsal view. See Material and methods for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4326613" httpUri="https://zenodo.org/record/4326613/files/figure.png" pageId="4" pageNumber="15">Fig. 1B</figureCitation>
). Four genital slits in each interradius (
<figureCitation id="137A469F5326FFE9FD969B82FD5AC1D9" box="[569,660,1504,1530]" captionStart="Fig" captionStartId="5.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@5.[189,1398,291,1855]" captionTargetPageId="5" captionText="Fig. 1. Ophioderma hendleri sp. nov., holotype (ICML-UNAM 18315, DD = 14.8 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Structures of a jaw (according to Hendler 2018). E. Basal dorsal arm. F. Dorsal arm. G. Ventral arm. H. Lateral arm plates and arm spines. I. Dorsal view. See Material and methods for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4326613" httpUri="https://zenodo.org/record/4326613/files/figure.png" pageId="4" pageNumber="15">Fig. 1B</figureCitation>
); proximal genital slits oval, in contact with distal part of oral shield and with 1
<superScript id="7C34F7525326FFE9FE289861FE5CC232" attach="left" box="[391,402,1539,1553]" fontSize="6" pageId="4" pageNumber="15">st</superScript>
LAP; distal genital slits rounded, elongated, placed between 5
<superScript id="7C34F7525326FFE9FBDB9861FB4FC232" attach="left" box="[1140,1153,1539,1553]" fontSize="6" pageId="4" pageNumber="15">th</superScript>
and 6
<superScript id="7C34F7525326FFE9FB659861FB19C232" attach="left" box="[1226,1239,1539,1553]" fontSize="6" pageId="4" pageNumber="15">th</superScript>
arm segment and close to margin of disc, surrounded by granulation and numerous, elongated and imbricated scales next to side of lateral arm plates (
<figureCitation id="137A469F5326FFE9FDEA9829FD6EC246" box="[581,672,1611,1637]" captionStart="Fig" captionStartId="5.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@5.[189,1398,291,1855]" captionTargetPageId="5" captionText="Fig. 1. Ophioderma hendleri sp. nov., holotype (ICML-UNAM 18315, DD = 14.8 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Structures of a jaw (according to Hendler 2018). E. Basal dorsal arm. F. Dorsal arm. G. Ventral arm. H. Lateral arm plates and arm spines. I. Dorsal view. See Material and methods for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4326613" httpUri="https://zenodo.org/record/4326613/files/figure.png" pageId="4" pageNumber="15">Fig. 1B</figureCitation>
).
</paragraph>
<paragraph id="8BFE5A1A5326FFE9FF1298EFFAD6C3CB" blockId="4.[189,1399,1677,2025]" pageId="4" pageNumber="15">
Oral shields slightly broader than long, rounded triangular, with convex obtuse proximal angle, straight to convex distal edge, obtuse lateral angles. Madreporite with central, circular, shallow depression, located in distal part of oral shield. Adoral shields naked, as long as broad, triangular and completely separated from each other. Jaws bear 89 oral papillae on each side: LyOs small, 2 × as long as broad, angled upwards; AdShSp largest, rounded; 2°AdShSp elongated and narrow; LOPas 45 conical and pointed, slightly separated; IPa reduced, elongated and pointed; TPa pair at apex of jaw, elongated, robust and pointed, slightly separated. vT similar in shape and slightly larger than TPa. Four teeth: ventralmost broader than long, quadrangular but with rounded borders; dorsalmost pointed, slender and smallest. One OPRSp, large and pointed at each side, visible within buccal cavity. Granules covering oral plates and below adoral shields larger than those on dorsal disc and interradii (
<figureCitation id="137A469F5326FFE9FB2599ACFB05C3CB" box="[1162,1227,1998,2024]" captionStart="Fig" captionStartId="5.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@5.[189,1398,291,1855]" captionTargetPageId="5" captionText="Fig. 1. Ophioderma hendleri sp. nov., holotype (ICML-UNAM 18315, DD = 14.8 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Structures of a jaw (according to Hendler 2018). E. Basal dorsal arm. F. Dorsal arm. G. Ventral arm. H. Lateral arm plates and arm spines. I. Dorsal view. See Material and methods for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4326613" httpUri="https://zenodo.org/record/4326613/files/figure.png" pageId="4" pageNumber="15">Fig. 1</figureCitation>
CD).
</paragraph>
<caption id="DF3E0A925327FFE8FF129907FE39C3CA" ID-DOI="http://doi.org/10.5281/zenodo.4326613" ID-Zenodo-Dep="4326613" httpUri="https://zenodo.org/record/4326613/files/figure.png" pageId="5" pageNumber="16" startId="5.[189,232,1893,1919]" targetBox="[189,1398,291,1855]" targetPageId="5">
<paragraph id="8BFE5A1A5327FFE8FF129907FE39C3CA" blockId="5.[189,1399,1893,2025]" pageId="5" pageNumber="16">
<emphasis id="B93586085327FFE8FF129907FEC4C35C" bold="true" box="[189,266,1893,1919]" pageId="5" pageNumber="16">Fig. 1.</emphasis>
<taxonomicName id="4C4121995327FFE8FEBE9907FDC0C35C" authority="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras, 2020" authorityName="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras" authorityYear="2020" box="[273,526,1893,1919]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="5" pageNumber="16" phylum="Echinodermata" rank="species" species="hendleri" status="sp. nov.">
<emphasis id="B93586085327FFE8FEBE9907FDC0C35C" box="[273,526,1893,1919]" italics="true" pageId="5" pageNumber="16">Ophioderma hendleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2063B735327FFE8FDBA9904FDBFC3A3" box="[533,625,1894,1920]" pageId="5" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (ICML-UNAM 18315, DD = 14.8 mm).
<emphasis id="B93586085327FFE8FB689907FB12C35C" bold="true" box="[1223,1244,1893,1919]" pageId="5" pageNumber="16">A</emphasis>
. Dorsal disc.
<emphasis id="B93586085327FFE8FF1299EBFF1FC380" bold="true" box="[189,209,1929,1955]" pageId="5" pageNumber="16">B</emphasis>
. Ventral disc.
<emphasis id="B93586085327FFE8FE2F99EAFE5BC381" bold="true" box="[384,405,1928,1954]" pageId="5" pageNumber="16">C</emphasis>
. Mouth.
<emphasis id="B93586085327FFE8FDA999EBFDD5C380" bold="true" box="[518,539,1929,1955]" pageId="5" pageNumber="16">D</emphasis>
. Structures of a jaw (according to
<bibRefCitation id="EFD027EB5327FFE8FC6D99EBFBA5C380" author="Hendler G." box="[962,1131,1929,1955]" pageId="5" pageNumber="16" pagination="189 - 311" refId="ref16304" refString="Hendler G. 2018. Armed to the teeth: a new paradigm for the buccal skeleton of brittle stars (Echinodermata: Ophiuroidea). Contributions in Science 526: 189 - 311." type="journal article" year="2018">Hendler 2018</bibRefCitation>
).
<emphasis id="B93586085327FFE8FB2F99EBFB5AC380" bold="true" box="[1152,1172,1929,1955]" pageId="5" pageNumber="16">E</emphasis>
. Basal dorsal arm.
<emphasis id="B93586085327FFE8FF1299CEFF01C3E5" bold="true" box="[189,207,1964,1990]" pageId="5" pageNumber="16">F</emphasis>
. Dorsal arm.
<emphasis id="B93586085327FFE8FEDD99CEFE47C3E5" bold="true" box="[370,393,1964,1990]" pageId="5" pageNumber="16">G</emphasis>
. Ventral arm.
<emphasis id="B93586085327FFE8FD9E99CEFD86C3E5" bold="true" box="[561,584,1964,1990]" pageId="5" pageNumber="16">H</emphasis>
. Lateral arm plates and arm spines.
<emphasis id="B93586085327FFE8FC5999CEFBCFC3E5" bold="true" box="[1014,1025,1964,1990]" pageId="5" pageNumber="16">I</emphasis>
. Dorsal view. See Material and methods for abbreviations.
</paragraph>
</caption>
<caption id="DF3E0A925324FFEBFF1298BAFB17C3CB" ID-DOI="http://doi.org/10.5281/zenodo.4326617" ID-Zenodo-Dep="4326617" httpUri="https://zenodo.org/record/4326617/files/figure.png" pageId="6" pageNumber="17" startId="6.[189,232,1752,1778]" targetBox="[295,1292,265,1714]" targetPageId="6">
<paragraph id="8BFE5A1A5324FFEBFF1298BAFB17C3CB" blockId="6.[189,1399,1752,2025]" pageId="6" pageNumber="17">
<emphasis id="B93586085324FFEBFF1298BAFEC3C2D1" bold="true" box="[189,269,1752,1778]" pageId="6" pageNumber="17">Fig. 2.</emphasis>
SEM images of skeletal elements of
<taxonomicName id="4C4121995324FFEBFD7898BAFC16C2D1" authority="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras, 2020" authorityName="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras" authorityYear="2020" box="[727,984,1752,1778]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="6" pageNumber="17" phylum="Echinodermata" rank="species" species="hendleri" status="sp. nov.">
<emphasis id="B93586085324FFEBFD7898BAFC16C2D1" box="[727,984,1752,1778]" italics="true" pageId="6" pageNumber="17">Ophioderma hendleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2063B735324FFEBFC4D98BAFB8CC2D1" box="[994,1090,1752,1778]" pageId="6" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
, paratype (ICML-UNAM 18323, DD = 15 mm).
<emphasis id="B93586085324FFEBFE6A9899FE32C336" bold="true" box="[453,508,1787,1813]" pageId="6" pageNumber="17">AB</emphasis>
. Radial shield (external view), deepest pores marked with arrows.
<emphasis id="B93586085324FFEBFAAA9899FAD4C336" bold="true" box="[1285,1306,1787,1813]" pageId="6" pageNumber="17">C</emphasis>
. Center of the distal portion of the radial shield, open meshed pores.
<emphasis id="B93586085324FFEBFCC1997CFC4DC31B" bold="true" box="[878,899,1822,1848]" pageId="6" pageNumber="17">D</emphasis>
. Dorsal arm plate, with one spur.
<emphasis id="B93586085324FFEBFAAD997CFAD8C31B" bold="true" box="[1282,1302,1822,1848]" pageId="6" pageNumber="17">E</emphasis>
. Ventral arm plate, with one spur.
<emphasis id="B93586085324FFEBFE4B9920FE38C37F" bold="true" box="[484,502,1858,1884]" pageId="6" pageNumber="17">F</emphasis>
. Lateral arm plate, external face, with two spurs and one condyle.
<emphasis id="B93586085324FFEBFB519923FADBC378" bold="true" box="[1278,1301,1857,1883]" pageId="6" pageNumber="17">G</emphasis>
. Lateral arm plate, lateral face, with spine articulations.
<emphasis id="B93586085324FFEBFD5B9907FCC5C35C" bold="true" box="[756,779,1893,1919]" pageId="6" pageNumber="17">H</emphasis>
. Lateral arm plate, internal face.
<emphasis id="B93586085324FFEBFB369907FB6AC35C" bold="true" box="[1177,1188,1893,1919]" pageId="6" pageNumber="17">I</emphasis>
. Outer surface of the LAP (F) with finely meshed and polygonal knobs.
<emphasis id="B93586085324FFEBFC9699EAFC86C381" bold="true" box="[825,840,1928,1954]" pageId="6" pageNumber="17">J</emphasis>
. Spine articulations with a weak sigmoidal fold and tilted lobes.
<emphasis id="B93586085324FFEBFED399C9FE5DC3E6" bold="true" box="[380,403,1963,1989]" pageId="6" pageNumber="17">K</emphasis>
. Arm vertebra, distal face.
<emphasis id="B93586085324FFEBFD6299C9FD2FC3E6" bold="true" box="[717,737,1963,1989]" pageId="6" pageNumber="17">L</emphasis>
. Arm vertebra, proximal face.
<emphasis id="B93586085324FFEBFBEA99C9FBAFC3E6" bold="true" box="[1093,1121,1963,1989]" pageId="6" pageNumber="17">M</emphasis>
. Tentacle scale.
<emphasis id="B93586085324FFEBFAB099C9FAFAC3E6" bold="true" box="[1311,1332,1963,1989]" pageId="6" pageNumber="17">N</emphasis>
. Arm spine.
<emphasis id="B93586085324FFEBFEA599ACFEEFC3CB" bold="true" box="[266,289,1998,2024]" pageId="6" pageNumber="17">O</emphasis>
. Dental plate fragments.
<emphasis id="B93586085324FFEBFDE699ACFD95C3CB" bold="true" box="[585,603,1998,2024]" pageId="6" pageNumber="17">P</emphasis>
. Oral plate, abradial face.
<emphasis id="B93586085324FFEBFC3D99ACFC67C3CB" bold="true" box="[914,937,1998,2024]" pageId="6" pageNumber="17">Q</emphasis>
. Oral plate, adradial face.
</paragraph>
</caption>
<caption id="DF3E0A925325FFEAFF1299EBFEF7C3C9" ID-DOI="http://doi.org/10.5281/zenodo.4326619" ID-Zenodo-Dep="4326619" httpUri="https://zenodo.org/record/4326619/files/figure.png" pageId="7" pageNumber="18" startId="7.[189,232,1929,1955]" targetBox="[189,1398,327,1891]" targetPageId="7">
<paragraph id="8BFE5A1A5325FFEAFF1299EBFEF7C3C9" blockId="7.[189,1399,1929,2027]" pageId="7" pageNumber="18">
<emphasis id="B93586085325FFEAFF1299EBFEC4C387" bold="true" box="[189,266,1929,1956]" pageId="7" pageNumber="18">Fig. 3.</emphasis>
<taxonomicName id="4C4121995325FFEAFEBD99E8FDDEC387" authority="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras, 2020" authorityName="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras" authorityYear="2020" box="[274,528,1930,1956]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="7" pageNumber="18" phylum="Echinodermata" rank="species" species="hendleri" status="sp. nov.">
<emphasis id="B93586085325FFEAFEBD99E8FDDEC387" box="[274,528,1930,1956]" italics="true" pageId="7" pageNumber="18">Ophioderma hendleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2063B735325FFEAFDB899E8FDBAC387" box="[535,628,1930,1956]" pageId="7" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
, paratype (ICML-UNAM 10582, DD = 4.8 mm).
<emphasis id="B93586085325FFEAFB1199EBFB1DC380" bold="true" box="[1214,1235,1929,1955]" pageId="7" pageNumber="18">A</emphasis>
. Dorsal view.
<emphasis id="B93586085325FFEAFF1299CFFF1FC3E4" bold="true" box="[189,209,1965,1991]" pageId="7" pageNumber="18">B</emphasis>
. Basal dorsal arm.
<emphasis id="B93586085325FFEAFE6B99CFFE17C3E4" bold="true" box="[452,473,1965,1991]" pageId="7" pageNumber="18">C</emphasis>
. Dorsal arm.
<emphasis id="B93586085325FFEAFD2999CFFD55C3E4" bold="true" box="[646,667,1965,1991]" pageId="7" pageNumber="18">D</emphasis>
. Ventral arm.
<emphasis id="B93586085325FFEAFCE099CFFCADC3E4" bold="true" box="[847,867,1965,1991]" pageId="7" pageNumber="18">E</emphasis>
. Dorsal disc.
<emphasis id="B93586085325FFEAFBBE99CFFBEDC3E4" bold="true" box="[1041,1059,1965,1991]" pageId="7" pageNumber="18">F</emphasis>
. Ventral disc.
<emphasis id="B93586085325FFEAFB7799CFFB21C3E4" bold="true" box="[1240,1263,1965,1991]" pageId="7" pageNumber="18">G</emphasis>
. Interradii.
<emphasis id="B93586085325FFEAFF1299B2FF1AC3C9" bold="true" box="[189,212,2000,2026]" pageId="7" pageNumber="18">H</emphasis>
. Mouth.
</paragraph>
</caption>
<caption id="DF3E0A92532AFFE5FF1299EAFEF7C3CB" ID-DOI="http://doi.org/10.5281/zenodo.4326621" ID-Zenodo-Dep="4326621" httpUri="https://zenodo.org/record/4326621/files/figure.png" pageId="8" pageNumber="19" startId="8.[189,232,1928,1954]" targetBox="[189,1398,325,1889]" targetPageId="8">
<paragraph id="8BFE5A1A532AFFE5FF1299EAFEF7C3CB" blockId="8.[189,1399,1928,2025]" pageId="8" pageNumber="19">
<emphasis id="B9358608532AFFE5FF1299EAFEC5C381" bold="true" box="[189,267,1928,1954]" pageId="8" pageNumber="19">Fig. 4.</emphasis>
<taxonomicName id="4C412199532AFFE5FEBD99EAFDDEC381" authority="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras, 2020" authorityName="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras" authorityYear="2020" box="[274,528,1928,1954]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="8" pageNumber="19" phylum="Echinodermata" rank="species" species="hendleri" status="sp. nov.">
<emphasis id="B9358608532AFFE5FEBD99EAFDDEC381" box="[274,528,1928,1954]" italics="true" pageId="8" pageNumber="19">Ophioderma hendleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2063B73532AFFE5FDB899EBFDBAC380" box="[535,628,1929,1955]" pageId="8" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
, paratype (ICML-UNAM 18319, DD = 2.2 mm).
<emphasis id="B9358608532AFFE5FB1199EAFB1DC381" bold="true" box="[1214,1235,1928,1954]" pageId="8" pageNumber="19">A</emphasis>
. Dorsal view.
<emphasis id="B9358608532AFFE5FF1299C9FF1FC3E6" bold="true" box="[189,209,1963,1989]" pageId="8" pageNumber="19">B</emphasis>
. Basal dorsal arm.
<emphasis id="B9358608532AFFE5FE6B99C9FE17C3E6" bold="true" box="[452,473,1963,1989]" pageId="8" pageNumber="19">C</emphasis>
. Dorsal arm.
<emphasis id="B9358608532AFFE5FD2999C9FD55C3E6" bold="true" box="[646,667,1963,1989]" pageId="8" pageNumber="19">D</emphasis>
. Ventral arm.
<emphasis id="B9358608532AFFE5FCE099C9FCADC3E6" bold="true" box="[847,867,1963,1989]" pageId="8" pageNumber="19">E</emphasis>
. Dorsal disc.
<emphasis id="B9358608532AFFE5FBBE99C9FBEDC3E6" bold="true" box="[1041,1059,1963,1989]" pageId="8" pageNumber="19">F</emphasis>
. Ventral disc.
<emphasis id="B9358608532AFFE5FB7799C9FB21C3E6" bold="true" box="[1240,1263,1963,1989]" pageId="8" pageNumber="19">G</emphasis>
. Interradii.
<emphasis id="B9358608532AFFE5FF1299ACFF1AC3CB" bold="true" box="[189,212,1998,2024]" pageId="8" pageNumber="19">H</emphasis>
. Mouth.
</paragraph>
</caption>
<caption id="DF3E0A92532BFFE4FF129907FEA1C3CA" ID-DOI="http://doi.org/10.5281/zenodo.4326625" ID-Zenodo-Dep="4326625" httpUri="https://zenodo.org/record/4326625/files/figure.png" pageId="9" pageNumber="20" startId="9.[189,232,1893,1919]" targetBox="[189,1398,291,1855]" targetPageId="9">
<paragraph id="8BFE5A1A532BFFE4FF129907FEA1C3CA" blockId="9.[189,1399,1893,2026]" pageId="9" pageNumber="20">
<emphasis id="B9358608532BFFE4FF129907FE81C35C" bold="true" box="[189,335,1893,1920]" pageId="9" pageNumber="20">Fig. 5. AD</emphasis>
.
<taxonomicName id="4C412199532BFFE4FECE9907FDAFC35C" authority="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras, 2020" authorityName="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras" authorityYear="2020" box="[353,609,1893,1919]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="9" pageNumber="20" phylum="Echinodermata" rank="species" species="hendleri" status="sp. nov.">
<emphasis id="B9358608532BFFE4FECE9907FDAFC35C" box="[353,609,1893,1919]" italics="true" pageId="9" pageNumber="20">Ophioderma hendleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2063B73532BFFE4FDC59904FD09C3A3" box="[618,711,1894,1920]" pageId="9" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B9358608532BFFE4FD7F9907FCC9C35C" bold="true" box="[720,775,1893,1919]" pageId="9" pageNumber="20">AB</emphasis>
. In vivo coloration in natural habitat.
<emphasis id="B9358608532BFFE4FB799907FAC1C35C" bold="true" box="[1238,1295,1893,1919]" pageId="9" pageNumber="20">CD</emphasis>
. In vivo coloration of collected specimens.
<emphasis id="B9358608532BFFE4FDD699EBFD64C380" bold="true" box="[633,682,1929,1955]" pageId="9" pageNumber="20">EJ</emphasis>
.
<taxonomicName id="4C412199532BFFE4FD1499EBFB47C380" authority="Lutken, 1859" authorityName="Lutken" authorityYear="1859" box="[699,1161,1928,1955]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="9" pageNumber="20" phylum="Echinodermata" rank="species" species="panamense">
<emphasis id="B9358608532BFFE4FD1499EBFC10C381" box="[699,990,1929,1955]" italics="true" pageId="9" pageNumber="20">Ophioderma panamense</emphasis>
<bibRefCitation id="EFD027EB532BFFE4FC4799EBFB47C380" author="Lutken C. F." box="[1000,1161,1928,1955]" pageId="9" pageNumber="20" pagination="75 - 169" refId="ref16788" refString="Lutken C. F. 1859. Additamenta ad Historiam Ophiuridarum. Beskrivelser af de fra Vestindien og Central Amerikas Vestkyst hidtil beckjendte Slangestjerner. Kongelige Danske Videnskabernes Selskab Biologiske Skrifter 2: 75 - 169." type="journal article" year="1859">Lütken, 1859</bibRefCitation>
</taxonomicName>
, holotype (NHMD- 107679, DD = 21.9 mm).
<emphasis id="B9358608532BFFE4FE5299CEFDDFC3E5" bold="true" box="[509,529,1964,1990]" pageId="9" pageNumber="20">E</emphasis>
. Dorsal disc.
<emphasis id="B9358608532BFFE4FD1499CEFD03C3E5" bold="true" box="[699,717,1964,1990]" pageId="9" pageNumber="20">F</emphasis>
. Ventral disc.
<emphasis id="B9358608532BFFE4FCD199CEFC5BC3E5" bold="true" box="[894,917,1964,1990]" pageId="9" pageNumber="20">G</emphasis>
. Mouth.
<emphasis id="B9358608532BFFE4FBA999CEFBD3C3E5" bold="true" box="[1030,1053,1964,1990]" pageId="9" pageNumber="20">H</emphasis>
. Dorsal arm.
<emphasis id="B9358608532BFFE4FB6999CEFB1FC3E5" bold="true" box="[1222,1233,1964,1990]" pageId="9" pageNumber="20">I</emphasis>
. Ventral arm.
<emphasis id="B9358608532BFFE4FF1299ADFF02C3CA" bold="true" box="[189,204,1999,2025]" pageId="9" pageNumber="20">J</emphasis>
. Dorsal view.
</paragraph>
</caption>
<caption id="DF3E0A925328FFE7FF129F6BFEBAC565" ID-Table-UUID="DF3E0A925328FFE7FF129F6BFEBAC565" httpUri="http://table.plazi.org/id/DF3E0A925328FFE7FF129F6BFEBAC565" pageId="10" pageNumber="21" targetBox="[219,1389,371,1330]" targetIsTable="true" targetPageId="10">
<paragraph id="8BFE5A1A5328FFE7FF129F6BFEBAC565" blockId="10.[189,1399,265,327]" pageId="10" pageNumber="21">
<emphasis id="B93586085328FFE7FF129F6BFEEBC500" bold="true" box="[189,293,265,291]" pageId="10" pageNumber="21">Table 1.</emphasis>
Comparison of morphological characters between some species of
<taxonomicName id="4C4121995328FFE7FBC39F6BFACEC500" authorityName="Muller &amp; Troschel" authorityYear="1840" box="[1132,1280,265,291]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="10" pageNumber="21" phylum="Echinodermata" rank="genus">
<emphasis id="B93586085328FFE7FBC39F6BFACEC500" box="[1132,1280,265,291]" italics="true" pageId="10" pageNumber="21">Ophioderma</emphasis>
</taxonomicName>
from the Eastern Pacific.
</paragraph>
</caption>
<paragraph id="8BFE5A1A5328FFE7FE699F11FA9DC113" pageId="10" pageNumber="21">
<table id="F941A8BA53280012FF749F11FAA3C111" box="[219,1389,371,1330]" gridcols="5" gridrows="10" pageId="10" pageNumber="21">
<tr id="3571585853280012FF749F11FAA3C5AA" box="[219,1389,371,393]" gridrow="0" pageId="10" pageNumber="21" rowspan-0="1">
<th id="76A0312453280012FE159F11FD59C5AA" box="[442,663,371,393]" gridcol="1" gridrow="0" pageId="10" pageNumber="21">
<emphasis id="B93586085328FFE7FE699F11FD47C5AB" bold="true" box="[454,649,371,393]" pageId="10" pageNumber="21">
<taxonomicName id="4C4121995328FFE7FE699F11FDF9C5AA" authority="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras, 2020" authorityName="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras" authorityYear="2020" box="[454,567,371,393]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="10" pageNumber="21" phylum="Echinodermata" rank="species" species="hendleri" status="sp. nov.">
<emphasis id="B93586085328FFE7FE699F11FDF9C5AA" bold="true" box="[454,567,371,393]" italics="true" pageId="10" pageNumber="21">O. hendleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2063B735328FFE7FD939F11FD47C5AB" box="[572,649,371,392]" pageId="10" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="76A0312453280012FD029F11FC48C5AA" box="[685,902,371,393]" gridcol="2" gridrow="0" pageId="10" pageNumber="21">
<taxonomicName id="4C4121995328FFE7FD7C9F11FCAEC5AA" authorityName="Lutken" authorityYear="1859" box="[723,864,371,393]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="10" pageNumber="21" phylum="Echinodermata" rank="species" species="panamense">
<emphasis id="B93586085328FFE7FD7C9F11FCAEC5AA" bold="true" box="[723,864,371,393]" italics="true" pageId="10" pageNumber="21">O. panamense</emphasis>
</taxonomicName>
</th>
<th id="76A0312453280012FC329F11FBB4C5AA" box="[925,1146,371,393]" gridcol="3" gridrow="0" pageId="10" pageNumber="21">
<taxonomicName id="4C4121995328FFE7FC199F11FBACC5AA" authorityName=", H. L. Clark" authorityYear="1917" box="[950,1122,371,393]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="10" pageNumber="21" phylum="Echinodermata" rank="species" species="pentacanthum">
<emphasis id="B93586085328FFE7FC199F11FBACC5AA" bold="true" box="[950,1122,371,393]" italics="true" pageId="10" pageNumber="21">O. pentacanthum</emphasis>
</taxonomicName>
</th>
<th id="76A0312453280012FB219F11FAA3C5AA" box="[1166,1389,371,393]" gridcol="4" gridrow="0" pageId="10" pageNumber="21">
<taxonomicName id="4C4121995328FFE7FB189F11FA8AC5AB" authorityName=", Lutken" authorityYear="1856" box="[1207,1348,371,392]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="10" pageNumber="21" phylum="Echinodermata" rank="species" species="variegatum">
<emphasis id="B93586085328FFE7FB189F11FA8AC5AB" bold="true" box="[1207,1348,371,392]" italics="true" pageId="10" pageNumber="21">O. variegatum</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="3571585853280012FF749FFCFAA3C597" box="[219,1389,414,436]" gridrow="1" pageId="10" pageNumber="21">
<th id="76A0312453280012FF749FFCFE5FC597" box="[219,401,414,436]" gridcol="0" gridrow="1" pageId="10" pageNumber="21">
<emphasis id="B93586085328FFE7FF419FFCFEB3C597" bold="true" box="[238,381,414,436]" pageId="10" pageNumber="21">Radial shields</emphasis>
</th>
<td id="76A0312453280012FE159FFCFD59C597" box="[442,663,414,436]" gridcol="1" gridrow="1" pageId="10" pageNumber="21">covered by granules</td>
<td id="76A0312453280012FD029FFCFC48C597" box="[685,902,414,436]" gridcol="2" gridrow="1" pageId="10" pageNumber="21">naked</td>
<td id="76A0312453280012FC329FFCFBB4C597" box="[925,1146,414,436]" gridcol="3" gridrow="1" pageId="10" pageNumber="21">covered by granules</td>
<td id="76A0312453280012FB219FFCFAA3C597" box="[1166,1389,414,436]" gridcol="4" gridrow="1" pageId="10" pageNumber="21">covered by granules</td>
</tr>
<tr id="3571585853280012FF749FAEFAA3C5DC" box="[219,1389,460,511]" gridrow="2" pageId="10" pageNumber="21">
<th id="76A0312453280012FF749FAEFE5FC5DC" box="[219,401,460,511]" gridcol="0" gridrow="2" pageId="10" pageNumber="21">
<emphasis id="B93586085328FFE7FF4E9FB8FE45C5CC" bold="true" box="[225,395,474,496]" pageId="10" pageNumber="21">Base of the arms</emphasis>
</th>
<td id="76A0312453280012FE159FAEFD59C5DC" box="[442,663,460,511]" gridcol="1" gridrow="2" pageId="10" pageNumber="21">covered by granules and scales</td>
<td id="76A0312453280012FD029FAEFC48C5DC" box="[685,902,460,511]" gridcol="2" gridrow="2" pageId="10" pageNumber="21">covered by granules and scales</td>
<td id="76A0312453280012FC329FAEFBB4C5DC" box="[925,1146,460,511]" gridcol="3" gridrow="2" pageId="10" pageNumber="21">covered by granules and scales</td>
<td id="76A0312453280012FB219FAEFAA3C5DC" box="[1166,1389,460,511]" gridcol="4" gridrow="2" pageId="10" pageNumber="21">covered by granules</td>
</tr>
<tr id="3571585853280012FF749C7AFAA3C64B" box="[219,1389,536,616]" gridrow="3" pageId="10" pageNumber="21">
<th id="76A0312453280012FF749C7AFE5FC64B" box="[219,401,536,616]" gridcol="0" gridrow="3" pageId="10" pageNumber="21">
<emphasis id="B93586085328FFE7FF549C57FEBCC668" bold="true" box="[251,370,565,587]" pageId="10" pageNumber="21">Arm-spines</emphasis>
</th>
<td id="76A0312453280012FE159C7AFD59C64B" box="[442,663,536,616]" gridcol="1" gridrow="3" pageId="10" pageNumber="21">up to 11; short and more or less robust</td>
<td id="76A0312453280012FD029C7AFC48C64B" box="[685,902,536,616]" gridcol="2" gridrow="3" pageId="10" pageNumber="21">up to 12; short and robust</td>
<td id="76A0312453280012FC329C7AFBB4C64B" box="[925,1146,536,616]" gridcol="3" gridrow="3" pageId="10" pageNumber="21">up to 6; large and thin, the ventral-most being much larger and robust</td>
<td id="76A0312453280012FB219C7AFAA3C64B" box="[1166,1389,536,616]" gridcol="4" gridrow="3" pageId="10" pageNumber="21">up to 9; very large and thin</td>
</tr>
<tr id="3571585853280012FF749CE3FAA3C697" box="[219,1389,641,692]" gridrow="4" pageId="10" pageNumber="21">
<th id="76A0312453280012FF749CE3FE5FC697" box="[219,401,641,692]" gridcol="0" gridrow="4" pageId="10" pageNumber="21">
<emphasis id="B93586085328FFE7FF449CF2FE4FC686" bold="true" box="[235,385,656,677]" pageId="10" pageNumber="21">Tentacle scales</emphasis>
</th>
<td id="76A0312453280012FE159CE3FD59C697" box="[442,663,641,692]" gridcol="1" gridrow="4" pageId="10" pageNumber="21">adradial slightly larger than abradial</td>
<td id="76A0312453280012FD029CE3FC48C697" box="[685,902,641,692]" gridcol="2" gridrow="4" pageId="10" pageNumber="21">adradial slightly larger than abradial</td>
<td id="76A0312453280012FC329CE3FBB4C697" box="[925,1146,641,692]" gridcol="3" gridrow="4" pageId="10" pageNumber="21">adradial much larger than abradial</td>
<td id="76A0312453280012FB219CE3FAA3C697" box="[1166,1389,641,692]" gridcol="4" gridrow="4" pageId="10" pageNumber="21">adradial slightly larger than abradial</td>
</tr>
<tr id="3571585853280012FF749CAEFAA3C751" box="[219,1389,716,882]" gridrow="5" pageId="10" pageNumber="21">
<th id="76A0312453280012FF749CAEFE5FC751" box="[219,401,716,882]" gridcol="0" gridrow="5" pageId="10" pageNumber="21">
<emphasis id="B93586085328FFE7FF579D76FEBDC709" bold="true" box="[248,371,788,810]" pageId="10" pageNumber="21">Oral shields</emphasis>
</th>
<td id="76A0312453280012FE159CAEFD59C751" box="[442,663,716,882]" gridcol="1" gridrow="5" pageId="10" pageNumber="21">broader than long, rounded triangular, with rounded obtuse proximal angle, straight distal edge, obtuse lateral angles</td>
<td id="76A0312453280012FD029CAEFC48C751" box="[685,902,716,882]" gridcol="2" gridrow="5" pageId="10" pageNumber="21">broader than long, rounded triangular, with rounded obtuse proximal angle, straight distal edge, obtuse lateral angles</td>
<td id="76A0312453280012FC329CAEFBB4C751" box="[925,1146,716,882]" gridcol="3" gridrow="5" pageId="10" pageNumber="21">broader than long, rounded triangular, with rounded obtuse proximal angle, straight distal edge, obtuse lateral angles</td>
<td id="76A0312453280012FB219CAEFAA3C751" box="[1166,1389,716,882]" gridcol="4" gridrow="5" pageId="10" pageNumber="21">longer than broad, oval, proximal margin rounded obtuse angled, distal edge convex, lateral margins straight</td>
</tr>
<tr id="3571585853280012FF749DE8FAA3C79E" box="[219,1389,906,957]" gridrow="6" pageId="10" pageNumber="21">
<th id="76A0312453280012FF749DE8FE5FC79E" box="[219,401,906,957]" gridcol="0" gridrow="6" pageId="10" pageNumber="21">
<emphasis id="B93586085328FFE7FF439DFAFEB1C78D" bold="true" box="[236,383,920,942]" pageId="10" pageNumber="21">Adoral shields</emphasis>
</th>
<td id="76A0312453280012FE159DE8FD59C79E" box="[442,663,906,957]" gridcol="1" gridrow="6" pageId="10" pageNumber="21">naked; equally long as broad</td>
<td id="76A0312453280012FD029DE8FC48C79E" box="[685,902,906,957]" gridcol="2" gridrow="6" pageId="10" pageNumber="21">covered by granules</td>
<td id="76A0312453280012FC329DE8FBB4C79E" box="[925,1146,906,957]" gridcol="3" gridrow="6" pageId="10" pageNumber="21">naked; longer than broad</td>
<td id="76A0312453280012FB219DE8FAA3C79E" box="[1166,1389,906,957]" gridcol="4" gridrow="6" pageId="10" pageNumber="21">naked; longer than broad</td>
</tr>
<tr id="3571585853280012FF749DB4FAA3C005" box="[219,1389,982,1062]" gridrow="7" pageId="10" pageNumber="21">
<th id="76A0312453280012FF749DB4FE5FC005" box="[219,401,982,1062]" gridcol="0" gridrow="7" pageId="10" pageNumber="21">
<emphasis id="B93586085328FFE7FF569D91FEBDC02A" bold="true" box="[249,371,1011,1033]" pageId="10" pageNumber="21">Genital slits</emphasis>
</th>
<td id="76A0312453280012FE159DB4FD59C005" box="[442,663,982,1062]" gridcol="1" gridrow="7" pageId="10" pageNumber="21">short and rounded; distally surrounded by granules and scales</td>
<td id="76A0312453280012FD029DB4FC48C005" box="[685,902,982,1062]" gridcol="2" gridrow="7" pageId="10" pageNumber="21">short and rounded; distally surrounded by granules and scales</td>
<td id="76A0312453280012FC329DB4FBB4C005" box="[925,1146,982,1062]" gridcol="3" gridrow="7" pageId="10" pageNumber="21">large and slender; distally surrounded by granules and scales</td>
<td id="76A0312453280012FB219DB4FAA3C005" box="[1166,1389,982,1062]" gridcol="4" gridrow="7" pageId="10" pageNumber="21">short and rounded; distally surrounded only by scales</td>
</tr>
<tr id="3571585853280012FF749A5DFAA3C121" box="[219,1389,1087,1282]" gridrow="8" pageId="10" pageNumber="21">
<th id="76A0312453280012FF749A5DFE5FC121" box="[219,401,1087,1282]" gridcol="0" gridrow="8" pageId="10" pageNumber="21">
<emphasis id="B93586085328FFE7FF749AF7FE5FC089" bold="true" box="[219,401,1173,1195]" pageId="10" pageNumber="21">
Coloration
<emphasis id="B93586085328FFE7FEE09AF7FE5FC089" bold="true" box="[335,401,1173,1194]" italics="true" pageId="10" pageNumber="21">in vivo</emphasis>
</emphasis>
</th>
<td id="76A0312453280012FE159A5DFD59C121" box="[442,663,1087,1282]" gridcol="1" gridrow="8" pageId="10" pageNumber="21">disc dark brown mottled with small and white spots, ventral margin beige; arms with transverse white bands</td>
<td id="76A0312453280012FD029A5DFC48C121" box="[685,902,1087,1282]" gridcol="2" gridrow="8" pageId="10" pageNumber="21">disc green, brown or grey mottled with red, orange or yellow, sometimes a central white spot; arms with transverse bands lighter and darker</td>
<td id="76A0312453280012FC329A5DFBB4C121" box="[925,1146,1087,1282]" gridcol="3" gridrow="8" pageId="10" pageNumber="21">disc orange with dark brown blotches; arms with transverse brown bands</td>
<td id="76A0312453280012FB219A5DFAA3C121" box="[1166,1389,1087,1282]" gridcol="4" gridrow="8" pageId="10" pageNumber="21">disc olive-green with dark pink or red; arms with transverse olivegreen and dark pinkred bands</td>
</tr>
<tr id="3571585853280012FF749B7BFAA3C111" box="[219,1389,1305,1330]" gridrow="9" pageId="10" pageNumber="21">
<th id="76A0312453280012FF749B7BFE5FC111" box="[219,401,1305,1330]" gridcol="0" gridrow="9" pageId="10" pageNumber="21">
<emphasis id="B93586085328FFE7FEB89B7BFE98C10D" bold="true" box="[279,342,1305,1326]" pageId="10" pageNumber="21">Depth</emphasis>
</th>
<td id="76A0312453280012FE159B7BFD59C111" box="[442,663,1305,1330]" gridcol="1" gridrow="9" pageId="10" pageNumber="21">intertidal 37 m</td>
<td id="76A0312453280012FD029B7BFC48C111" box="[685,902,1305,1330]" gridcol="2" gridrow="9" pageId="10" pageNumber="21">intertidal 20 m</td>
<td id="76A0312453280012FC329B7BFBB4C111" box="[925,1146,1305,1330]" gridcol="3" gridrow="9" pageId="10" pageNumber="21">102 183 m</td>
<td id="76A0312453280012FB219B7BFAA3C111" box="[1166,1389,1305,1330]" gridcol="4" gridrow="9" pageId="10" pageNumber="21">intertidal 110 m</td>
</tr>
</table>
</paragraph>
<paragraph id="8BFE5A1A5328FFE7FF129B07FD5FC3B1" blockId="10.[189,1399,1381,1938]" pageId="10" pageNumber="21">
Dorsal base of arms covered by granules and with approximately 11 oval scales, located laterally at first 3 dorsal arm plates (
<figureCitation id="137A469F5328FFE7FE439BEAFD86C181" box="[492,584,1416,1442]" captionStart="Fig" captionStartId="5.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@5.[189,1398,291,1855]" captionTargetPageId="5" captionText="Fig. 1. Ophioderma hendleri sp. nov., holotype (ICML-UNAM 18315, DD = 14.8 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Structures of a jaw (according to Hendler 2018). E. Basal dorsal arm. F. Dorsal arm. G. Ventral arm. H. Lateral arm plates and arm spines. I. Dorsal view. See Material and methods for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4326613" httpUri="https://zenodo.org/record/4326613/files/figure.png" pageId="10" pageNumber="21">Fig. 1E</figureCitation>
). Dorsal arm plates 3 × broader than long, rectangular, overlapping; proximal edge straight, distal edge slightly concave with rounded lateral edges (
<figureCitation id="137A469F5328FFE7FB2B9BC9FB11C1E5" box="[1156,1247,1451,1478]" captionStart="Fig" captionStartId="5.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@5.[189,1398,291,1855]" captionTargetPageId="5" captionText="Fig. 1. Ophioderma hendleri sp. nov., holotype (ICML-UNAM 18315, DD = 14.8 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Structures of a jaw (according to Hendler 2018). E. Basal dorsal arm. F. Dorsal arm. G. Ventral arm. H. Lateral arm plates and arm spines. I. Dorsal view. See Material and methods for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4326613" httpUri="https://zenodo.org/record/4326613/files/figure.png" pageId="10" pageNumber="21">Fig. 1F</figureCitation>
). Distalmost dorsal arm plates smaller, longer than broad, triangular. First ventral arm plate small, broader than long, with rounded edges; in contact with adoral shields. Second ventral arm plate quadrangular, as broad as long (
<figureCitation id="137A469F5328FFE7FE8A9874FE4DC213" box="[293,387,1558,1584]" captionStart="Fig" captionStartId="5.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@5.[189,1398,291,1855]" captionTargetPageId="5" captionText="Fig. 1. Ophioderma hendleri sp. nov., holotype (ICML-UNAM 18315, DD = 14.8 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Structures of a jaw (according to Hendler 2018). E. Basal dorsal arm. F. Dorsal arm. G. Ventral arm. H. Lateral arm plates and arm spines. I. Dorsal view. See Material and methods for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4326613" httpUri="https://zenodo.org/record/4326613/files/figure.png" pageId="10" pageNumber="21">Fig. 1C</figureCitation>
). Next ventral arm plates slightly longer than broad, quadrangular; proximal margin truncate and distal edge convex (
<figureCitation id="137A469F5328FFE7FDEE985BFD51C270" box="[577,671,1593,1619]" captionStart="Fig" captionStartId="5.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@5.[189,1398,291,1855]" captionTargetPageId="5" captionText="Fig. 1. Ophioderma hendleri sp. nov., holotype (ICML-UNAM 18315, DD = 14.8 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Structures of a jaw (according to Hendler 2018). E. Basal dorsal arm. F. Dorsal arm. G. Ventral arm. H. Lateral arm plates and arm spines. I. Dorsal view. See Material and methods for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4326613" httpUri="https://zenodo.org/record/4326613/files/figure.png" pageId="10" pageNumber="21">Fig. 1G</figureCitation>
). Distalmost ventral arm plates wider distally, with a tapering proximal angle. Paired rounded depressions between most proximal ventral arm plates (
<figureCitation id="137A469F5328FFE7FB6D983FFAD3C254" box="[1218,1309,1629,1655]" captionStart="Fig" captionStartId="5.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@5.[189,1398,291,1855]" captionTargetPageId="5" captionText="Fig. 1. Ophioderma hendleri sp. nov., holotype (ICML-UNAM 18315, DD = 14.8 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Structures of a jaw (according to Hendler 2018). E. Basal dorsal arm. F. Dorsal arm. G. Ventral arm. H. Lateral arm plates and arm spines. I. Dorsal view. See Material and methods for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4326613" httpUri="https://zenodo.org/record/4326613/files/figure.png" pageId="10" pageNumber="21">Fig. 1B</figureCitation>
). LAPs conspicuous, semicircular, broader than long (
<figureCitation id="137A469F5328FFE7FD4798E2FC86C2B9" box="[744,840,1664,1690]" captionStart="Fig" captionStartId="5.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@5.[189,1398,291,1855]" captionTargetPageId="5" captionText="Fig. 1. Ophioderma hendleri sp. nov., holotype (ICML-UNAM 18315, DD = 14.8 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Structures of a jaw (according to Hendler 2018). E. Basal dorsal arm. F. Dorsal arm. G. Ventral arm. H. Lateral arm plates and arm spines. I. Dorsal view. See Material and methods for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4326613" httpUri="https://zenodo.org/record/4326613/files/figure.png" pageId="10" pageNumber="21">Fig. 1H</figureCitation>
). LAP with up to 11 arm spines. First and 2
<superScript id="7C34F7525328FFE7FACA981DFAB8C2AE" attach="none" box="[1381,1398,1663,1677]" fontSize="6" pageId="10" pageNumber="21">nd</superScript>
LAP with 3 arm spines; 3
<superScript id="7C34F7525328FFE7FE5D98C1FDCEC292" attach="left" box="[498,512,1699,1713]" fontSize="6" pageId="10" pageNumber="21">rd</superScript>
with 4 arm spines; 4
<superScript id="7C34F7525328FFE7FCAF98C1FCC3C292" attach="left" box="[768,781,1699,1713]" fontSize="6" pageId="10" pageNumber="21">th</superScript>
with 5; 5
<superScript id="7C34F7525328FFE7FC2C98C1FC5EC292" attach="left" box="[899,912,1699,1713]" fontSize="6" pageId="10" pageNumber="21">th</superScript>
with 6; 6
<superScript id="7C34F7525328FFE7FBA998C1FBDDC292" attach="left" box="[1030,1043,1699,1713]" fontSize="6" pageId="10" pageNumber="21">th</superScript>
with 7; 7
<superScript id="7C34F7525328FFE7FB2698C1FB58C292" attach="left" box="[1161,1174,1699,1713]" fontSize="6" pageId="10" pageNumber="21">th</superScript>
with 8; 8
<superScript id="7C34F7525328FFE7FAA398C1FAD7C292" attach="left" box="[1292,1305,1699,1713]" fontSize="6" pageId="10" pageNumber="21">th</superScript>
with 9; 9
<superScript id="7C34F7525328FFE7FF6398A4FF26C2C2" attach="none" box="[204,232,1734,1761]" fontSize="6" pageId="10" pageNumber="21">th</superScript>
30
<superScript id="7C34F7525328FFE7FEAA98A4FEDCC2F7" attach="left" box="[261,274,1734,1748]" fontSize="6" pageId="10" pageNumber="21">th</superScript>
arm plates with 1011 arm spines; 35
<superScript id="7C34F7525328FFE7FD7898A4FD2AC2F7" attach="left" box="[727,740,1734,1748]" fontSize="6" pageId="10" pageNumber="21">th</superScript>
with 9 spines; 40
<superScript id="7C34F7525328FFE7FC1898A4FC1AC2C1" attach="none" box="[951,980,1734,1762]" fontSize="6" pageId="10" pageNumber="21">th</superScript>
50
<superScript id="7C34F7525328FFE7FC5E98A4FC30C2F7" attach="left" box="[1009,1022,1734,1748]" fontSize="6" pageId="10" pageNumber="21">th</superScript>
with 8; 60
<superScript id="7C34F7525328FFE7FBD098A4FB42C2F7" attach="left" box="[1151,1164,1734,1748]" fontSize="6" pageId="10" pageNumber="21">th</superScript>
with 7; 70
<superScript id="7C34F7525328FFE7FAA198A4FAD5C2F7" attach="left" box="[1294,1307,1734,1748]" fontSize="6" pageId="10" pageNumber="21">th</superScript>
with 6; 75
<superScript id="7C34F7525328FFE7FF759888FF29C2DB" attach="left" box="[218,231,1770,1784]" fontSize="6" pageId="10" pageNumber="21">th</superScript>
with 5; 80
<superScript id="7C34F7525328FFE7FEC69888FEB8C2DB" attach="left" box="[361,374,1770,1784]" fontSize="6" pageId="10" pageNumber="21">th</superScript>
with 4;&gt;80
<superScript id="7C34F7525328FFE7FDA89888FDDAC2DB" attach="left" box="[519,532,1770,1784]" fontSize="6" pageId="10" pageNumber="21">th</superScript>
with 3 arm spines. Arm spines with blunt tip; length approximately ½ of LAP. Dorsalmost arm spine shortest, blunt; ventralmost arm spine longest and more robust, as long as 1 arm segment, almost in contact with tentacle scale of succeeding joint (
<figureCitation id="137A469F5328FFE7FBB79953FBBBC36F" box="[1048,1141,1841,1868]" captionStart="Fig" captionStartId="5.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@5.[189,1398,291,1855]" captionTargetPageId="5" captionText="Fig. 1. Ophioderma hendleri sp. nov., holotype (ICML-UNAM 18315, DD = 14.8 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Structures of a jaw (according to Hendler 2018). E. Basal dorsal arm. F. Dorsal arm. G. Ventral arm. H. Lateral arm plates and arm spines. I. Dorsal view. See Material and methods for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4326613" httpUri="https://zenodo.org/record/4326613/files/figure.png" pageId="10" pageNumber="21">Fig. 1H</figureCitation>
). Two tentacle scales; adradial tentacle scale ovoid, approximately ½× length of ventral arm plate; abradial tentacle scale slightly shorter, subtriangular (
<figureCitation id="137A469F5328FFE7FD89991AFD4DC3B1" box="[550,643,1912,1938]" captionStart="Fig" captionStartId="5.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@5.[189,1398,291,1855]" captionTargetPageId="5" captionText="Fig. 1. Ophioderma hendleri sp. nov., holotype (ICML-UNAM 18315, DD = 14.8 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Structures of a jaw (according to Hendler 2018). E. Basal dorsal arm. F. Dorsal arm. G. Ventral arm. H. Lateral arm plates and arm spines. I. Dorsal view. See Material and methods for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4326613" httpUri="https://zenodo.org/record/4326613/files/figure.png" pageId="10" pageNumber="21">Fig. 1G</figureCitation>
).
</paragraph>
<paragraph id="8BFE5A1A5328FFE6FF1299C9FADFC5FD" blockId="10.[189,1398,1963,2025]" lastBlockId="11.[189,1399,265,479]" lastPageId="11" lastPageNumber="22" pageId="10" pageNumber="21">
General coloration dark brown (dry specimen) (
<figureCitation id="137A469F5328FFE7FD4099C9FCF1C3E6" box="[751,831,1963,1989]" captionStart="Fig" captionStartId="5.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@5.[189,1398,291,1855]" captionTargetPageId="5" captionText="Fig. 1. Ophioderma hendleri sp. nov., holotype (ICML-UNAM 18315, DD = 14.8 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Structures of a jaw (according to Hendler 2018). E. Basal dorsal arm. F. Dorsal arm. G. Ventral arm. H. Lateral arm plates and arm spines. I. Dorsal view. See Material and methods for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4326613" httpUri="https://zenodo.org/record/4326613/files/figure.png" pageId="10" pageNumber="21">Fig. 1I</figureCitation>
). Dorsal side: disc dark brown and mottled with small white spots; middle of margin on each interradius of disc bearing white spot (
<figureCitation id="137A469F5328FFE7FB6D99ACFAEDC3CB" box="[1218,1315,1998,2024]" captionStart="Fig" captionStartId="5.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@5.[189,1398,291,1855]" captionTargetPageId="5" captionText="Fig. 1. Ophioderma hendleri sp. nov., holotype (ICML-UNAM 18315, DD = 14.8 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Structures of a jaw (according to Hendler 2018). E. Basal dorsal arm. F. Dorsal arm. G. Ventral arm. H. Lateral arm plates and arm spines. I. Dorsal view. See Material and methods for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4326613" httpUri="https://zenodo.org/record/4326613/files/figure.png" pageId="10" pageNumber="21">Fig. 1A</figureCitation>
); arms dark brown with small black spots only observed microscopically and with every 56 arm plates with transverse white bands, located between distal part of ventral plate and proximal part of next arm plate (
<figureCitation id="137A469F5329FFE6FF699F36FEFDC54D" box="[198,307,340,366]" captionStart="Fig" captionStartId="5.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@5.[189,1398,291,1855]" captionTargetPageId="5" captionText="Fig. 1. Ophioderma hendleri sp. nov., holotype (ICML-UNAM 18315, DD = 14.8 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Structures of a jaw (according to Hendler 2018). E. Basal dorsal arm. F. Dorsal arm. G. Ventral arm. H. Lateral arm plates and arm spines. I. Dorsal view. See Material and methods for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4326613" httpUri="https://zenodo.org/record/4326613/files/figure.png" pageId="11" pageNumber="22">Fig. 1F, I</figureCitation>
). Ventral side: interradii, center and proximal part beige, margin dark brown with some beige marks (
<figureCitation id="137A469F5329FFE6FEBB9F1BFEA1C5B0" box="[276,367,377,403]" captionStart="Fig" captionStartId="5.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@5.[189,1398,291,1855]" captionTargetPageId="5" captionText="Fig. 1. Ophioderma hendleri sp. nov., holotype (ICML-UNAM 18315, DD = 14.8 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Structures of a jaw (according to Hendler 2018). E. Basal dorsal arm. F. Dorsal arm. G. Ventral arm. H. Lateral arm plates and arm spines. I. Dorsal view. See Material and methods for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4326613" httpUri="https://zenodo.org/record/4326613/files/figure.png" pageId="11" pageNumber="22">Fig. 1B</figureCitation>
). Jaw beige but oral shields can display brown color (
<figureCitation id="137A469F5329FFE6FC4B9F1BFBF1C5B0" box="[996,1087,377,403]" captionStart="Fig" captionStartId="5.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@5.[189,1398,291,1855]" captionTargetPageId="5" captionText="Fig. 1. Ophioderma hendleri sp. nov., holotype (ICML-UNAM 18315, DD = 14.8 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Structures of a jaw (according to Hendler 2018). E. Basal dorsal arm. F. Dorsal arm. G. Ventral arm. H. Lateral arm plates and arm spines. I. Dorsal view. See Material and methods for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4326613" httpUri="https://zenodo.org/record/4326613/files/figure.png" pageId="11" pageNumber="22">Fig. 1C</figureCitation>
). Ventral arm plates beige, but that on tip of arms dark brown with some beige transversal lines (
<figureCitation id="137A469F5329FFE6FC5B9FFDFB9CC59A" box="[1012,1106,415,441]" captionStart="Fig" captionStartId="5.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@5.[189,1398,291,1855]" captionTargetPageId="5" captionText="Fig. 1. Ophioderma hendleri sp. nov., holotype (ICML-UNAM 18315, DD = 14.8 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Structures of a jaw (according to Hendler 2018). E. Basal dorsal arm. F. Dorsal arm. G. Ventral arm. H. Lateral arm plates and arm spines. I. Dorsal view. See Material and methods for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4326613" httpUri="https://zenodo.org/record/4326613/files/figure.png" pageId="11" pageNumber="22">Fig. 1G</figureCitation>
). LAPs dark brown with some beige marks; arm spines dark brown and generally beige color on base and tip (
<figureCitation id="137A469F5329FFE6FB0A9FA6FACCC5FD" box="[1189,1282,452,478]" captionStart="Fig" captionStartId="5.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@5.[189,1398,291,1855]" captionTargetPageId="5" captionText="Fig. 1. Ophioderma hendleri sp. nov., holotype (ICML-UNAM 18315, DD = 14.8 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Structures of a jaw (according to Hendler 2018). E. Basal dorsal arm. F. Dorsal arm. G. Ventral arm. H. Lateral arm plates and arm spines. I. Dorsal view. See Material and methods for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4326613" httpUri="https://zenodo.org/record/4326613/files/figure.png" pageId="11" pageNumber="22">Fig. 1H</figureCitation>
).
</paragraph>
<paragraph id="8BFE5A1A5329FFE6FF459C6DFE30C60A" blockId="11.[189,1399,527,1527]" box="[234,510,527,553]" pageId="11" pageNumber="22">
<emphasis id="B93586085329FFE6FF459C6DFE30C60A" bold="true" box="[234,510,527,553]" pageId="11" pageNumber="22">Disarticulated ossicles</emphasis>
</paragraph>
<paragraph id="8BFE5A1A5329FFE6FF129C57FBC8C1D5" blockId="11.[189,1399,527,1527]" pageId="11" pageNumber="22">
Specimen analyzed:
<specimenCount id="9D4791935329FFE6FE199C57FDC0C66C" box="[438,526,565,591]" pageId="11" pageNumber="22" type="generic">1 spec.</specimenCount>
,
<typeStatus id="54FAE4B85329FFE6FDB29C57FD4CC66C" box="[541,642,565,591]" pageId="11" pageNumber="22" type="paratype">paratype</typeStatus>
ICML-UNAM 18323 (DD =
<quantity id="4CB9F7FF5329FFE6FC5E9C57FB84C66C" box="[1009,1098,565,592]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="22" unit="mm" value="15.0">15 mm</quantity>
, AL =
<quantity id="4CB9F7FF5329FFE6FB0C9C57FADAC66C" box="[1187,1300,565,591]" metricMagnitude="-2" metricUnit="m" metricValue="8.08" pageId="11" pageNumber="22" unit="mm" value="80.8">80.8 mm</quantity>
). Radial shield (external view) irregularly triangular, with convex proximal margin, incised abradial edge with projection, convex to straight distal edge and irregular adradial edge, series of 8 pores on median to proximal margin (which are covered/overlapped by disc scales and granules in intact animal) (
<figureCitation id="137A469F5329FFE6FAA29CC7FA9CC69C" box="[1293,1362,677,703]" captionStart="Fig" captionStartId="6.[189,232,1752,1778]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure-170@6.[294,1293,265,1714]" captionTargetPageId="6" captionText="Fig. 2. SEM images of skeletal elements of Ophioderma hendleri sp. nov., paratype (ICML-UNAM 18323, DD = 15 mm). AB. Radial shield (external view), deepest pores marked with arrows. C. Center of the distal portion of the radial shield, open meshed pores. D. Dorsal arm plate, with one spur. E. Ventral arm plate, with one spur. F. Lateral arm plate, external face, with two spurs and one condyle. G. Lateral arm plate, lateral face, with spine articulations. H. Lateral arm plate, internal face. I. Outer surface of the LAP (F) with finely meshed and polygonal knobs. J. Spine articulations with a weak sigmoidal fold and tilted lobes. K. Arm vertebra, distal face. L. Arm vertebra, proximal face. M. Tentacle scale. N. Arm spine. O. Dental plate fragments. P. Oral plate, abradial face. Q. Oral plate, adradial face." figureDoi="http://doi.org/10.5281/zenodo.4326617" httpUri="https://zenodo.org/record/4326617/files/figure.png" pageId="11" pageNumber="22">Fig. 2</figureCitation>
A B), most proximal ones deepest.
<collectingRegion id="498594F85329FFE6FD929CA9FD45C6C6" box="[573,651,715,741]" country="France" name="Centre" pageId="11" pageNumber="22">Center</collectingRegion>
of distal portion of radial shield covered in intact animal. Outer surface of stereom is open mesh of pores and small knobs (
<figureCitation id="137A469F5329FFE6FC2B9C92FC2CC729" box="[900,994,752,778]" captionStart="Fig" captionStartId="6.[189,232,1752,1778]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure-170@6.[294,1293,265,1714]" captionTargetPageId="6" captionText="Fig. 2. SEM images of skeletal elements of Ophioderma hendleri sp. nov., paratype (ICML-UNAM 18323, DD = 15 mm). AB. Radial shield (external view), deepest pores marked with arrows. C. Center of the distal portion of the radial shield, open meshed pores. D. Dorsal arm plate, with one spur. E. Ventral arm plate, with one spur. F. Lateral arm plate, external face, with two spurs and one condyle. G. Lateral arm plate, lateral face, with spine articulations. H. Lateral arm plate, internal face. I. Outer surface of the LAP (F) with finely meshed and polygonal knobs. J. Spine articulations with a weak sigmoidal fold and tilted lobes. K. Arm vertebra, distal face. L. Arm vertebra, proximal face. M. Tentacle scale. N. Arm spine. O. Dental plate fragments. P. Oral plate, abradial face. Q. Oral plate, adradial face." figureDoi="http://doi.org/10.5281/zenodo.4326617" httpUri="https://zenodo.org/record/4326617/files/figure.png" pageId="11" pageNumber="22">Fig. 2C</figureCitation>
). Dorsal arm plate slightly arched and somewhat rectangular, 3× as wide as long; proximal margin convex and distal margin straight, 1 spur on proximal portion of external surface (
<figureCitation id="137A469F5329FFE6FD7B9D59FCFDC776" box="[724,819,827,853]" captionStart="Fig" captionStartId="6.[189,232,1752,1778]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure-170@6.[294,1293,265,1714]" captionTargetPageId="6" captionText="Fig. 2. SEM images of skeletal elements of Ophioderma hendleri sp. nov., paratype (ICML-UNAM 18323, DD = 15 mm). AB. Radial shield (external view), deepest pores marked with arrows. C. Center of the distal portion of the radial shield, open meshed pores. D. Dorsal arm plate, with one spur. E. Ventral arm plate, with one spur. F. Lateral arm plate, external face, with two spurs and one condyle. G. Lateral arm plate, lateral face, with spine articulations. H. Lateral arm plate, internal face. I. Outer surface of the LAP (F) with finely meshed and polygonal knobs. J. Spine articulations with a weak sigmoidal fold and tilted lobes. K. Arm vertebra, distal face. L. Arm vertebra, proximal face. M. Tentacle scale. N. Arm spine. O. Dental plate fragments. P. Oral plate, abradial face. Q. Oral plate, adradial face." figureDoi="http://doi.org/10.5281/zenodo.4326617" httpUri="https://zenodo.org/record/4326617/files/figure.png" pageId="11" pageNumber="22">Fig. 2D</figureCitation>
). Ventral arm plate as long as wide, quadrangular with proximal side truncated and pointed spur, lateral sides concave forming border of tentacle pore (
<figureCitation id="137A469F5329FFE6FF699DE4FEEFC783" box="[198,289,902,928]" captionStart="Fig" captionStartId="6.[189,232,1752,1778]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure-170@6.[294,1293,265,1714]" captionTargetPageId="6" captionText="Fig. 2. SEM images of skeletal elements of Ophioderma hendleri sp. nov., paratype (ICML-UNAM 18323, DD = 15 mm). AB. Radial shield (external view), deepest pores marked with arrows. C. Center of the distal portion of the radial shield, open meshed pores. D. Dorsal arm plate, with one spur. E. Ventral arm plate, with one spur. F. Lateral arm plate, external face, with two spurs and one condyle. G. Lateral arm plate, lateral face, with spine articulations. H. Lateral arm plate, internal face. I. Outer surface of the LAP (F) with finely meshed and polygonal knobs. J. Spine articulations with a weak sigmoidal fold and tilted lobes. K. Arm vertebra, distal face. L. Arm vertebra, proximal face. M. Tentacle scale. N. Arm spine. O. Dental plate fragments. P. Oral plate, abradial face. Q. Oral plate, adradial face." figureDoi="http://doi.org/10.5281/zenodo.4326617" httpUri="https://zenodo.org/record/4326617/files/figure.png" pageId="11" pageNumber="22">Fig. 2E</figureCitation>
). LAP D-shaped, 2× as high as wide, with 8 spine articulations sunken in notches of distal edge (
<figureCitation id="137A469F5329FFE6FF699DC9FEC3C7E6" box="[198,269,939,965]" captionStart="Fig" captionStartId="6.[189,232,1752,1778]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure-170@6.[294,1293,265,1714]" captionTargetPageId="6" captionText="Fig. 2. SEM images of skeletal elements of Ophioderma hendleri sp. nov., paratype (ICML-UNAM 18323, DD = 15 mm). AB. Radial shield (external view), deepest pores marked with arrows. C. Center of the distal portion of the radial shield, open meshed pores. D. Dorsal arm plate, with one spur. E. Ventral arm plate, with one spur. F. Lateral arm plate, external face, with two spurs and one condyle. G. Lateral arm plate, lateral face, with spine articulations. H. Lateral arm plate, internal face. I. Outer surface of the LAP (F) with finely meshed and polygonal knobs. J. Spine articulations with a weak sigmoidal fold and tilted lobes. K. Arm vertebra, distal face. L. Arm vertebra, proximal face. M. Tentacle scale. N. Arm spine. O. Dental plate fragments. P. Oral plate, abradial face. Q. Oral plate, adradial face." figureDoi="http://doi.org/10.5281/zenodo.4326617" httpUri="https://zenodo.org/record/4326617/files/figure.png" pageId="11" pageNumber="22">Fig. 2</figureCitation>
FG). Ventral portion of LAP projecting ventro-proximalwards and ventro-distal tip projecting ventralwards (
<figureCitation id="137A469F5329FFE6FEC99DB2FE74C7C8" box="[358,442,976,1003]" captionStart="Fig" captionStartId="6.[189,232,1752,1778]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure-170@6.[294,1293,265,1714]" captionTargetPageId="6" captionText="Fig. 2. SEM images of skeletal elements of Ophioderma hendleri sp. nov., paratype (ICML-UNAM 18323, DD = 15 mm). AB. Radial shield (external view), deepest pores marked with arrows. C. Center of the distal portion of the radial shield, open meshed pores. D. Dorsal arm plate, with one spur. E. Ventral arm plate, with one spur. F. Lateral arm plate, external face, with two spurs and one condyle. G. Lateral arm plate, lateral face, with spine articulations. H. Lateral arm plate, internal face. I. Outer surface of the LAP (F) with finely meshed and polygonal knobs. J. Spine articulations with a weak sigmoidal fold and tilted lobes. K. Arm vertebra, distal face. L. Arm vertebra, proximal face. M. Tentacle scale. N. Arm spine. O. Dental plate fragments. P. Oral plate, abradial face. Q. Oral plate, adradial face." figureDoi="http://doi.org/10.5281/zenodo.4326617" httpUri="https://zenodo.org/record/4326617/files/figure.png" pageId="11" pageNumber="22">Fig 2F</figureCitation>
). Ventro-proximal margin with condyle (
<figureCitation id="137A469F5329FFE6FC0C9DB2FC30C7C8" box="[931,1022,976,1003]" captionStart="Fig" captionStartId="6.[189,232,1752,1778]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure-170@6.[294,1293,265,1714]" captionTargetPageId="6" captionText="Fig. 2. SEM images of skeletal elements of Ophioderma hendleri sp. nov., paratype (ICML-UNAM 18323, DD = 15 mm). AB. Radial shield (external view), deepest pores marked with arrows. C. Center of the distal portion of the radial shield, open meshed pores. D. Dorsal arm plate, with one spur. E. Ventral arm plate, with one spur. F. Lateral arm plate, external face, with two spurs and one condyle. G. Lateral arm plate, lateral face, with spine articulations. H. Lateral arm plate, internal face. I. Outer surface of the LAP (F) with finely meshed and polygonal knobs. J. Spine articulations with a weak sigmoidal fold and tilted lobes. K. Arm vertebra, distal face. L. Arm vertebra, proximal face. M. Tentacle scale. N. Arm spine. O. Dental plate fragments. P. Oral plate, abradial face. Q. Oral plate, adradial face." figureDoi="http://doi.org/10.5281/zenodo.4326617" httpUri="https://zenodo.org/record/4326617/files/figure.png" pageId="11" pageNumber="22">Fig. 2F</figureCitation>
). Proximal edge of LAP with 2 prominent and elongated spurs, protruding and modifying central-proximal edge (
<figureCitation id="137A469F5329FFE6FB309D94FB32C033" box="[1183,1276,1014,1040]" captionStart="Fig" captionStartId="6.[189,232,1752,1778]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure-170@6.[294,1293,265,1714]" captionTargetPageId="6" captionText="Fig. 2. SEM images of skeletal elements of Ophioderma hendleri sp. nov., paratype (ICML-UNAM 18323, DD = 15 mm). AB. Radial shield (external view), deepest pores marked with arrows. C. Center of the distal portion of the radial shield, open meshed pores. D. Dorsal arm plate, with one spur. E. Ventral arm plate, with one spur. F. Lateral arm plate, external face, with two spurs and one condyle. G. Lateral arm plate, lateral face, with spine articulations. H. Lateral arm plate, internal face. I. Outer surface of the LAP (F) with finely meshed and polygonal knobs. J. Spine articulations with a weak sigmoidal fold and tilted lobes. K. Arm vertebra, distal face. L. Arm vertebra, proximal face. M. Tentacle scale. N. Arm spine. O. Dental plate fragments. P. Oral plate, abradial face. Q. Oral plate, adradial face." figureDoi="http://doi.org/10.5281/zenodo.4326617" httpUri="https://zenodo.org/record/4326617/files/figure.png" pageId="11" pageNumber="22">Fig. 2F</figureCitation>
), between spurs and across remaining proximal margin discernible band of different stereom structure (
<figureCitation id="137A469F5329FFE6FAA19A79FAA7C016" box="[1294,1385,1051,1077]" captionStart="Fig" captionStartId="6.[189,232,1752,1778]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure-170@6.[294,1293,265,1714]" captionTargetPageId="6" captionText="Fig. 2. SEM images of skeletal elements of Ophioderma hendleri sp. nov., paratype (ICML-UNAM 18323, DD = 15 mm). AB. Radial shield (external view), deepest pores marked with arrows. C. Center of the distal portion of the radial shield, open meshed pores. D. Dorsal arm plate, with one spur. E. Ventral arm plate, with one spur. F. Lateral arm plate, external face, with two spurs and one condyle. G. Lateral arm plate, lateral face, with spine articulations. H. Lateral arm plate, internal face. I. Outer surface of the LAP (F) with finely meshed and polygonal knobs. J. Spine articulations with a weak sigmoidal fold and tilted lobes. K. Arm vertebra, distal face. L. Arm vertebra, proximal face. M. Tentacle scale. N. Arm spine. O. Dental plate fragments. P. Oral plate, abradial face. Q. Oral plate, adradial face." figureDoi="http://doi.org/10.5281/zenodo.4326617" httpUri="https://zenodo.org/record/4326617/files/figure.png" pageId="11" pageNumber="22">Fig. 2F</figureCitation>
). Outer surface finely meshed with relatively large polygonal knobs (
<figureCitation id="137A469F5329FFE6FC769A23FBE2C078" box="[985,1068,1089,1115]" captionStart="Fig" captionStartId="6.[189,232,1752,1778]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure-170@6.[294,1293,265,1714]" captionTargetPageId="6" captionText="Fig. 2. SEM images of skeletal elements of Ophioderma hendleri sp. nov., paratype (ICML-UNAM 18323, DD = 15 mm). AB. Radial shield (external view), deepest pores marked with arrows. C. Center of the distal portion of the radial shield, open meshed pores. D. Dorsal arm plate, with one spur. E. Ventral arm plate, with one spur. F. Lateral arm plate, external face, with two spurs and one condyle. G. Lateral arm plate, lateral face, with spine articulations. H. Lateral arm plate, internal face. I. Outer surface of the LAP (F) with finely meshed and polygonal knobs. J. Spine articulations with a weak sigmoidal fold and tilted lobes. K. Arm vertebra, distal face. L. Arm vertebra, proximal face. M. Tentacle scale. N. Arm spine. O. Dental plate fragments. P. Oral plate, abradial face. Q. Oral plate, adradial face." figureDoi="http://doi.org/10.5281/zenodo.4326617" httpUri="https://zenodo.org/record/4326617/files/figure.png" pageId="11" pageNumber="22">Fig. 2I</figureCitation>
). Inner surface of LAP with continuous ridge on proximal edge, and at ventro-distal margin 2 spurs matching those on external surface (
<figureCitation id="137A469F5329FFE6FE8E9AEEFEB0C085" box="[289,382,1164,1190]" captionStart="Fig" captionStartId="6.[189,232,1752,1778]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure-170@6.[294,1293,265,1714]" captionTargetPageId="6" captionText="Fig. 2. SEM images of skeletal elements of Ophioderma hendleri sp. nov., paratype (ICML-UNAM 18323, DD = 15 mm). AB. Radial shield (external view), deepest pores marked with arrows. C. Center of the distal portion of the radial shield, open meshed pores. D. Dorsal arm plate, with one spur. E. Ventral arm plate, with one spur. F. Lateral arm plate, external face, with two spurs and one condyle. G. Lateral arm plate, lateral face, with spine articulations. H. Lateral arm plate, internal face. I. Outer surface of the LAP (F) with finely meshed and polygonal knobs. J. Spine articulations with a weak sigmoidal fold and tilted lobes. K. Arm vertebra, distal face. L. Arm vertebra, proximal face. M. Tentacle scale. N. Arm spine. O. Dental plate fragments. P. Oral plate, abradial face. Q. Oral plate, adradial face." figureDoi="http://doi.org/10.5281/zenodo.4326617" httpUri="https://zenodo.org/record/4326617/files/figure.png" pageId="11" pageNumber="22">Fig. 2H</figureCitation>
), and 3 pores just below center. Spine articulations ventralwards increasing in size (
<figureCitation id="137A469F5329FFE6FAE89AEEFF2BC0E8" captionStart="Fig" captionStartId="6.[189,232,1752,1778]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure-170@6.[294,1293,265,1714]" captionTargetPageId="6" captionText="Fig. 2. SEM images of skeletal elements of Ophioderma hendleri sp. nov., paratype (ICML-UNAM 18323, DD = 15 mm). AB. Radial shield (external view), deepest pores marked with arrows. C. Center of the distal portion of the radial shield, open meshed pores. D. Dorsal arm plate, with one spur. E. Ventral arm plate, with one spur. F. Lateral arm plate, external face, with two spurs and one condyle. G. Lateral arm plate, lateral face, with spine articulations. H. Lateral arm plate, internal face. I. Outer surface of the LAP (F) with finely meshed and polygonal knobs. J. Spine articulations with a weak sigmoidal fold and tilted lobes. K. Arm vertebra, distal face. L. Arm vertebra, proximal face. M. Tentacle scale. N. Arm spine. O. Dental plate fragments. P. Oral plate, abradial face. Q. Oral plate, adradial face." figureDoi="http://doi.org/10.5281/zenodo.4326617" httpUri="https://zenodo.org/record/4326617/files/figure.png" pageId="11" pageNumber="22">Fig. 2G</figureCitation>
). Lobes with weak sigmoidal fold, tilted, curved, slightly joined on proximal margin by 3 or 4 knobs; ventral lobe smaller than dorsal lobe (
<figureCitation id="137A469F5329FFE6FDD19AB4FD1DC0D2" box="[638,723,1238,1265]" captionStart="Fig" captionStartId="6.[189,232,1752,1778]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure-170@6.[294,1293,265,1714]" captionTargetPageId="6" captionText="Fig. 2. SEM images of skeletal elements of Ophioderma hendleri sp. nov., paratype (ICML-UNAM 18323, DD = 15 mm). AB. Radial shield (external view), deepest pores marked with arrows. C. Center of the distal portion of the radial shield, open meshed pores. D. Dorsal arm plate, with one spur. E. Ventral arm plate, with one spur. F. Lateral arm plate, external face, with two spurs and one condyle. G. Lateral arm plate, lateral face, with spine articulations. H. Lateral arm plate, internal face. I. Outer surface of the LAP (F) with finely meshed and polygonal knobs. J. Spine articulations with a weak sigmoidal fold and tilted lobes. K. Arm vertebra, distal face. L. Arm vertebra, proximal face. M. Tentacle scale. N. Arm spine. O. Dental plate fragments. P. Oral plate, abradial face. Q. Oral plate, adradial face." figureDoi="http://doi.org/10.5281/zenodo.4326617" httpUri="https://zenodo.org/record/4326617/files/figure.png" pageId="11" pageNumber="22">Fig. 2J</figureCitation>
). Proximal vertebrae almost round, as wide as long, with large aboral muscle flange and smaller oral muscle flange (
<figureCitation id="137A469F5329FFE6FCDB9A9EFC1EC135" box="[884,976,1276,1302]" captionStart="Fig" captionStartId="6.[189,232,1752,1778]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure-170@6.[294,1293,265,1714]" captionTargetPageId="6" captionText="Fig. 2. SEM images of skeletal elements of Ophioderma hendleri sp. nov., paratype (ICML-UNAM 18323, DD = 15 mm). AB. Radial shield (external view), deepest pores marked with arrows. C. Center of the distal portion of the radial shield, open meshed pores. D. Dorsal arm plate, with one spur. E. Ventral arm plate, with one spur. F. Lateral arm plate, external face, with two spurs and one condyle. G. Lateral arm plate, lateral face, with spine articulations. H. Lateral arm plate, internal face. I. Outer surface of the LAP (F) with finely meshed and polygonal knobs. J. Spine articulations with a weak sigmoidal fold and tilted lobes. K. Arm vertebra, distal face. L. Arm vertebra, proximal face. M. Tentacle scale. N. Arm spine. O. Dental plate fragments. P. Oral plate, abradial face. Q. Oral plate, adradial face." figureDoi="http://doi.org/10.5281/zenodo.4326617" httpUri="https://zenodo.org/record/4326617/files/figure.png" pageId="11" pageNumber="22">Fig. 2L</figureCitation>
). Distal face of vertebrae with large muscle flanges, with typical zygospondylous articulation (
<figureCitation id="137A469F5329FFE6FCD69B43FC15C118" box="[889,987,1313,1339]" captionStart="Fig" captionStartId="6.[189,232,1752,1778]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure-170@6.[294,1293,265,1714]" captionTargetPageId="6" captionText="Fig. 2. SEM images of skeletal elements of Ophioderma hendleri sp. nov., paratype (ICML-UNAM 18323, DD = 15 mm). AB. Radial shield (external view), deepest pores marked with arrows. C. Center of the distal portion of the radial shield, open meshed pores. D. Dorsal arm plate, with one spur. E. Ventral arm plate, with one spur. F. Lateral arm plate, external face, with two spurs and one condyle. G. Lateral arm plate, lateral face, with spine articulations. H. Lateral arm plate, internal face. I. Outer surface of the LAP (F) with finely meshed and polygonal knobs. J. Spine articulations with a weak sigmoidal fold and tilted lobes. K. Arm vertebra, distal face. L. Arm vertebra, proximal face. M. Tentacle scale. N. Arm spine. O. Dental plate fragments. P. Oral plate, abradial face. Q. Oral plate, adradial face." figureDoi="http://doi.org/10.5281/zenodo.4326617" httpUri="https://zenodo.org/record/4326617/files/figure.png" pageId="11" pageNumber="22">Fig. 2K</figureCitation>
). Tentacle scale longer than wide, with scale-like surface (
<figureCitation id="137A469F5329FFE6FE749B25FD8FC142" box="[475,577,1351,1377]" captionStart="Fig" captionStartId="6.[189,232,1752,1778]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure-170@6.[294,1293,265,1714]" captionTargetPageId="6" captionText="Fig. 2. SEM images of skeletal elements of Ophioderma hendleri sp. nov., paratype (ICML-UNAM 18323, DD = 15 mm). AB. Radial shield (external view), deepest pores marked with arrows. C. Center of the distal portion of the radial shield, open meshed pores. D. Dorsal arm plate, with one spur. E. Ventral arm plate, with one spur. F. Lateral arm plate, external face, with two spurs and one condyle. G. Lateral arm plate, lateral face, with spine articulations. H. Lateral arm plate, internal face. I. Outer surface of the LAP (F) with finely meshed and polygonal knobs. J. Spine articulations with a weak sigmoidal fold and tilted lobes. K. Arm vertebra, distal face. L. Arm vertebra, proximal face. M. Tentacle scale. N. Arm spine. O. Dental plate fragments. P. Oral plate, abradial face. Q. Oral plate, adradial face." figureDoi="http://doi.org/10.5281/zenodo.4326617" httpUri="https://zenodo.org/record/4326617/files/figure.png" pageId="11" pageNumber="22">Fig. 2M</figureCitation>
). Spines with scale-like surface and blunt tip (
<figureCitation id="137A469F5329FFE6FBC09B25FB01C142" box="[1135,1231,1351,1377]" captionStart="Fig" captionStartId="6.[189,232,1752,1778]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure-170@6.[294,1293,265,1714]" captionTargetPageId="6" captionText="Fig. 2. SEM images of skeletal elements of Ophioderma hendleri sp. nov., paratype (ICML-UNAM 18323, DD = 15 mm). AB. Radial shield (external view), deepest pores marked with arrows. C. Center of the distal portion of the radial shield, open meshed pores. D. Dorsal arm plate, with one spur. E. Ventral arm plate, with one spur. F. Lateral arm plate, external face, with two spurs and one condyle. G. Lateral arm plate, lateral face, with spine articulations. H. Lateral arm plate, internal face. I. Outer surface of the LAP (F) with finely meshed and polygonal knobs. J. Spine articulations with a weak sigmoidal fold and tilted lobes. K. Arm vertebra, distal face. L. Arm vertebra, proximal face. M. Tentacle scale. N. Arm spine. O. Dental plate fragments. P. Oral plate, abradial face. Q. Oral plate, adradial face." figureDoi="http://doi.org/10.5281/zenodo.4326617" httpUri="https://zenodo.org/record/4326617/files/figure.png" pageId="11" pageNumber="22">Fig. 2N</figureCitation>
). Dental plate consists of several pieces, 1
<superScript id="7C34F7525329FFE6FDAA9B09FDDEC15A" attach="left" box="[517,528,1387,1401]" fontSize="6" pageId="11" pageNumber="22">st</superScript>
piece bears 2 TPas on small round socket and 1 wide tooth socket, while rest of pieces with single tooth socket per piece (
<quantity id="4CB9F7FF5329FFE6FCAB9BF3FCFDC188" box="[772,819,1425,1451]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="11" pageNumber="22" unit="in" value="4.0">4 in</quantity>
total), none of sockets penetrate plate (
<figureCitation id="137A469F5329FFE6FAA69BF3FAA4C188" box="[1289,1386,1425,1451]" captionStart="Fig" captionStartId="6.[189,232,1752,1778]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure-170@6.[294,1293,265,1714]" captionTargetPageId="6" captionText="Fig. 2. SEM images of skeletal elements of Ophioderma hendleri sp. nov., paratype (ICML-UNAM 18323, DD = 15 mm). AB. Radial shield (external view), deepest pores marked with arrows. C. Center of the distal portion of the radial shield, open meshed pores. D. Dorsal arm plate, with one spur. E. Ventral arm plate, with one spur. F. Lateral arm plate, external face, with two spurs and one condyle. G. Lateral arm plate, lateral face, with spine articulations. H. Lateral arm plate, internal face. I. Outer surface of the LAP (F) with finely meshed and polygonal knobs. J. Spine articulations with a weak sigmoidal fold and tilted lobes. K. Arm vertebra, distal face. L. Arm vertebra, proximal face. M. Tentacle scale. N. Arm spine. O. Dental plate fragments. P. Oral plate, abradial face. Q. Oral plate, adradial face." figureDoi="http://doi.org/10.5281/zenodo.4326617" httpUri="https://zenodo.org/record/4326617/files/figure.png" pageId="11" pageNumber="22">Fig. 2O</figureCitation>
). Oral plates as long as high, abradial face with muscle fossa highly triangular, almost covering whole surface (
<figureCitation id="137A469F5329FFE6FE8D9BBEFEB5C1D5" box="[290,379,1500,1526]" captionStart="Fig" captionStartId="6.[189,232,1752,1778]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure-170@6.[294,1293,265,1714]" captionTargetPageId="6" captionText="Fig. 2. SEM images of skeletal elements of Ophioderma hendleri sp. nov., paratype (ICML-UNAM 18323, DD = 15 mm). AB. Radial shield (external view), deepest pores marked with arrows. C. Center of the distal portion of the radial shield, open meshed pores. D. Dorsal arm plate, with one spur. E. Ventral arm plate, with one spur. F. Lateral arm plate, external face, with two spurs and one condyle. G. Lateral arm plate, lateral face, with spine articulations. H. Lateral arm plate, internal face. I. Outer surface of the LAP (F) with finely meshed and polygonal knobs. J. Spine articulations with a weak sigmoidal fold and tilted lobes. K. Arm vertebra, distal face. L. Arm vertebra, proximal face. M. Tentacle scale. N. Arm spine. O. Dental plate fragments. P. Oral plate, abradial face. Q. Oral plate, adradial face." figureDoi="http://doi.org/10.5281/zenodo.4326617" httpUri="https://zenodo.org/record/4326617/files/figure.png" pageId="11" pageNumber="22">Fig. 2P</figureCitation>
), adradial face in middle part with foot basin (
<figureCitation id="137A469F5329FFE6FC359BBEFC37C1D5" box="[922,1017,1500,1526]" captionStart="Fig" captionStartId="6.[189,232,1752,1778]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure-170@6.[294,1293,265,1714]" captionTargetPageId="6" captionText="Fig. 2. SEM images of skeletal elements of Ophioderma hendleri sp. nov., paratype (ICML-UNAM 18323, DD = 15 mm). AB. Radial shield (external view), deepest pores marked with arrows. C. Center of the distal portion of the radial shield, open meshed pores. D. Dorsal arm plate, with one spur. E. Ventral arm plate, with one spur. F. Lateral arm plate, external face, with two spurs and one condyle. G. Lateral arm plate, lateral face, with spine articulations. H. Lateral arm plate, internal face. I. Outer surface of the LAP (F) with finely meshed and polygonal knobs. J. Spine articulations with a weak sigmoidal fold and tilted lobes. K. Arm vertebra, distal face. L. Arm vertebra, proximal face. M. Tentacle scale. N. Arm spine. O. Dental plate fragments. P. Oral plate, abradial face. Q. Oral plate, adradial face." figureDoi="http://doi.org/10.5281/zenodo.4326617" httpUri="https://zenodo.org/record/4326617/files/figure.png" pageId="11" pageNumber="22">Fig. 2Q</figureCitation>
).
</paragraph>
<paragraph id="8BFE5A1A5329FFE6FF12984AFE7DC262" blockId="11.[189,435,1575,1602]" box="[189,435,1575,1602]" pageId="11" pageNumber="22">
<heading id="D0B6ED765329FFE6FF12984AFE7DC262" bold="true" box="[189,435,1575,1602]" fontSize="11" level="3" pageId="11" pageNumber="22" reason="3">
<emphasis id="B93586085329FFE6FF12984AFE7DC262" bold="true" box="[189,435,1575,1602]" pageId="11" pageNumber="22">
<typeStatus id="54FAE4B85329FFE6FF12984AFEE1C261" box="[189,303,1576,1602]" pageId="11" pageNumber="22" type="paratype">Paratype</typeStatus>
variations
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFE5A1A5329FFE6FF12983BFB06C3CB" blockId="11.[189,1399,1624,2025]" pageId="11" pageNumber="22">
Large specimens showed some differences in morphology compared to small ones (
<figureCitation id="137A469F5329FFE6FB3A983AFB11C251" box="[1173,1247,1624,1650]" captionStart="Fig" captionStartId="5.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@5.[189,1398,291,1855]" captionTargetPageId="5" captionText="Fig. 1. Ophioderma hendleri sp. nov., holotype (ICML-UNAM 18315, DD = 14.8 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Structures of a jaw (according to Hendler 2018). E. Basal dorsal arm. F. Dorsal arm. G. Ventral arm. H. Lateral arm plates and arm spines. I. Dorsal view. See Material and methods for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4326613" httpUri="https://zenodo.org/record/4326613/files/figure.png" pageId="11" pageNumber="22">Figs 1</figureCitation>
,
<figureCitation id="137A469F5329FFE6FB41983AFAD0C251" box="[1262,1310,1624,1650]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="7.[189,232,1929,1955]" captionStartId-1="8.[189,232,1928,1954]" captionTargetBox-0="[189,1398,327,1891]" captionTargetBox-1="[189,1398,325,1889]" captionTargetId-0="figure-11@7.[189,1398,327,1891]" captionTargetId-1="figure-17@8.[189,1398,325,1889]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Fig. 3. Ophioderma hendleri sp. nov., paratype (ICML-UNAM 10582, DD = 4.8 mm). A. Dorsal view. B. Basal dorsal arm. C. Dorsal arm. D. Ventral arm. E. Dorsal disc. F. Ventral disc. G. Interradii. H. Mouth." captionText-1="Fig. 4. Ophioderma hendleri sp. nov., paratype (ICML-UNAM 18319, DD = 2.2 mm). A. Dorsal view. B. Basal dorsal arm. C. Dorsal arm. D. Ventral arm. E. Dorsal disc. F. Ventral disc. G. Interradii. H. Mouth." figureDoi-0="http://doi.org/10.5281/zenodo.4326619" figureDoi-1="http://doi.org/10.5281/zenodo.4326621" httpUri-0="https://zenodo.org/record/4326619/files/figure.png" httpUri-1="https://zenodo.org/record/4326621/files/figure.png" pageId="11" pageNumber="22">34</figureCitation>
). Large individuals have granules and scales on the base of the arms (
<figureCitation id="137A469F5329FFE6FC0C981FFC30C2BB" box="[931,1022,1661,1688]" captionStart="Fig" captionStartId="5.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@5.[189,1398,291,1855]" captionTargetPageId="5" captionText="Fig. 1. Ophioderma hendleri sp. nov., holotype (ICML-UNAM 18315, DD = 14.8 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Structures of a jaw (according to Hendler 2018). E. Basal dorsal arm. F. Dorsal arm. G. Ventral arm. H. Lateral arm plates and arm spines. I. Dorsal view. See Material and methods for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4326613" httpUri="https://zenodo.org/record/4326613/files/figure.png" pageId="11" pageNumber="22">Fig. 1E</figureCitation>
), but in the smallest specimens (DD = ~
<quantity id="4CB9F7FF5329FFE6FE8998C1FE46C29D" box="[294,392,1699,1726]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="11" pageNumber="22" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">24 mm</quantity>
) the scales are covered by granules (
<figureCitation id="137A469F5329FFE6FC9D98C1FC40C29E" box="[818,910,1699,1725]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[189,1398,325,1889]" captionTargetId="figure-17@8.[189,1398,325,1889]" captionTargetPageId="8" captionText="Fig. 4. Ophioderma hendleri sp. nov., paratype (ICML-UNAM 18319, DD = 2.2 mm). A. Dorsal view. B. Basal dorsal arm. C. Dorsal arm. D. Ventral arm. E. Dorsal disc. F. Ventral disc. G. Interradii. H. Mouth." figureDoi="http://doi.org/10.5281/zenodo.4326621" httpUri="https://zenodo.org/record/4326621/files/figure.png" pageId="11" pageNumber="22">Fig. 4B</figureCitation>
). The scales begin to appear as the animal grows; therefore, in specimens with DD =
<quantity id="4CB9F7FF5329FFE6FD1498AAFCD3C2C0" box="[699,797,1736,1763]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="11" pageNumber="22" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">46 mm</quantity>
there are up to 3 elongated scales to either side of the base of the arms (
<figureCitation id="137A469F5329FFE6FE13988CFDD7C32B" box="[444,537,1774,1800]" captionStart="Fig" captionStartId="7.[189,232,1929,1955]" captionTargetBox="[189,1398,327,1891]" captionTargetId="figure-11@7.[189,1398,327,1891]" captionTargetPageId="7" captionText="Fig. 3. Ophioderma hendleri sp. nov., paratype (ICML-UNAM 10582, DD = 4.8 mm). A. Dorsal view. B. Basal dorsal arm. C. Dorsal arm. D. Ventral arm. E. Dorsal disc. F. Ventral disc. G. Interradii. H. Mouth." figureDoi="http://doi.org/10.5281/zenodo.4326619" httpUri="https://zenodo.org/record/4326619/files/figure.png" pageId="11" pageNumber="22">Fig. 3B</figureCitation>
), and with a DD =
<quantity id="4CB9F7FF5329FFE6FCAF988CFC8BC32B" box="[768,837,1774,1800]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="22" unit="mm" value="7.0">7 mm</quantity>
there are up to six scales; the number of scales in larger specimens reaches 11 (
<figureCitation id="137A469F5329FFE6FD979971FD5AC30E" box="[568,660,1811,1837]" captionStart="Fig" captionStartId="5.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@5.[189,1398,291,1855]" captionTargetPageId="5" captionText="Fig. 1. Ophioderma hendleri sp. nov., holotype (ICML-UNAM 18315, DD = 14.8 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Structures of a jaw (according to Hendler 2018). E. Basal dorsal arm. F. Dorsal arm. G. Ventral arm. H. Lateral arm plates and arm spines. I. Dorsal view. See Material and methods for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4326613" httpUri="https://zenodo.org/record/4326613/files/figure.png" pageId="11" pageNumber="22">Fig. 1E</figureCitation>
). Additionally, large individuals have numerous scales located on the distalmost part of the interradii (
<figureCitation id="137A469F5329FFE6FD39995BFD3FC370" box="[662,753,1849,1875]" captionStart="Fig" captionStartId="5.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@5.[189,1398,291,1855]" captionTargetPageId="5" captionText="Fig. 1. Ophioderma hendleri sp. nov., holotype (ICML-UNAM 18315, DD = 14.8 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Structures of a jaw (according to Hendler 2018). E. Basal dorsal arm. F. Dorsal arm. G. Ventral arm. H. Lateral arm plates and arm spines. I. Dorsal view. See Material and methods for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4326613" httpUri="https://zenodo.org/record/4326613/files/figure.png" pageId="11" pageNumber="22">Fig. 1B</figureCitation>
) but in the smallest specimens (DD =
<quantity id="4CB9F7FF5329FFE6FB6B995BFAE4C370" box="[1220,1322,1849,1875]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="11" pageNumber="22" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">36 mm</quantity>
) these scales are not evident, and instead, only granules are present (
<figureCitation id="137A469F5329FFE6FC0E993CFC25C35B" box="[929,1003,1886,1912]" captionStart="Fig" captionStartId="7.[189,232,1929,1955]" captionTargetBox="[189,1398,327,1891]" captionTargetId="figure-11@7.[189,1398,327,1891]" captionTargetPageId="7" captionText="Fig. 3. Ophioderma hendleri sp. nov., paratype (ICML-UNAM 10582, DD = 4.8 mm). A. Dorsal view. B. Basal dorsal arm. C. Dorsal arm. D. Ventral arm. E. Dorsal disc. F. Ventral disc. G. Interradii. H. Mouth." figureDoi="http://doi.org/10.5281/zenodo.4326619" httpUri="https://zenodo.org/record/4326619/files/figure.png" pageId="11" pageNumber="22">Figs 3</figureCitation>
FG, 4FG). In specimens larger than DD =
<quantity id="4CB9F7FF5329FFE6FEED99E6FE44C3BD" box="[322,394,1924,1950]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="22" unit="mm" value="7.0">7 mm</quantity>
, a row of elongated scales in the distal genital slits is evident. Finally, in the smallest specimens the adoral shields are completely covered by granules, which disappear as the animal grows, and individuals with DD
<emphasis id="B93586085329FFE6FE4999AFFE38C3CA" bold="true" box="[486,502,1997,2025]" pageId="11" pageNumber="22">=</emphasis>
<quantity id="4CB9F7FF5329FFE6FE5299ACFD8FC3CA" box="[509,577,1998,2025]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="22" unit="mm" value="5.0">5 mm</quantity>
tend to have totally or partially covered adoral shields.
</paragraph>
<paragraph id="8BFE5A1A532EFFE1FF129F6BFBECC5B7" blockId="12.[189,1399,265,405]" pageId="12" pageNumber="23">
Some specimens can have beige specks on the LAPs, the arm spines and the tentacle scales. Moreover, some specimens from the Gulf of California (
<collectingCountry id="F3561A8A532EFFE1FD609F4DFCE2C569" box="[719,812,303,330]" name="Mexico" pageId="12" pageNumber="23">Mexico</collectingCountry>
) have a large white spot in the center of the dorsal disc. It is important to note that in vivo coloration (
<figureCitation id="137A469F532EFFE1FC849F37FCBFC54C" box="[811,881,340,367]" captionStart="Fig" captionStartId="9.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@9.[189,1398,291,1855]" captionTargetPageId="9" captionText="Fig. 5. AD. Ophioderma hendleri sp. nov. AB. In vivo coloration in natural habitat. CD. In vivo coloration of collected specimens. EJ. Ophioderma panamense Lütken, 1859, holotype (NHMD- 107679, DD = 21.9 mm). E. Dorsal disc. F. Ventral disc. G. Mouth. H. Dorsal arm. I. Ventral arm. J. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4326625" httpUri="https://zenodo.org/record/4326625/files/figure.png" pageId="12" pageNumber="23">Fig. 5</figureCitation>
AD), generally conserved in all preserved specimens (
<figureCitation id="137A469F532EFFE1FEE89F18FE5FC5B6" box="[327,401,378,405]" captionStart="Fig" captionStartId="5.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@5.[189,1398,291,1855]" captionTargetPageId="5" captionText="Fig. 1. Ophioderma hendleri sp. nov., holotype (ICML-UNAM 18315, DD = 14.8 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Structures of a jaw (according to Hendler 2018). E. Basal dorsal arm. F. Dorsal arm. G. Ventral arm. H. Lateral arm plates and arm spines. I. Dorsal view. See Material and methods for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4326613" httpUri="https://zenodo.org/record/4326613/files/figure.png" pageId="12" pageNumber="23">Figs 1</figureCitation>
,
<figureCitation id="137A469F532EFFE1FE309F19FE01C5B6" box="[415,463,379,405]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="7.[189,232,1929,1955]" captionStartId-1="8.[189,232,1928,1954]" captionTargetBox-0="[189,1398,327,1891]" captionTargetBox-1="[189,1398,325,1889]" captionTargetId-0="figure-11@7.[189,1398,327,1891]" captionTargetId-1="figure-17@8.[189,1398,325,1889]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Fig. 3. Ophioderma hendleri sp. nov., paratype (ICML-UNAM 10582, DD = 4.8 mm). A. Dorsal view. B. Basal dorsal arm. C. Dorsal arm. D. Ventral arm. E. Dorsal disc. F. Ventral disc. G. Interradii. H. Mouth." captionText-1="Fig. 4. Ophioderma hendleri sp. nov., paratype (ICML-UNAM 18319, DD = 2.2 mm). A. Dorsal view. B. Basal dorsal arm. C. Dorsal arm. D. Ventral arm. E. Dorsal disc. F. Ventral disc. G. Interradii. H. Mouth." figureDoi-0="http://doi.org/10.5281/zenodo.4326619" figureDoi-1="http://doi.org/10.5281/zenodo.4326621" httpUri-0="https://zenodo.org/record/4326619/files/figure.png" httpUri-1="https://zenodo.org/record/4326621/files/figure.png" pageId="12" pageNumber="23">34</figureCitation>
), can be used as a distinctive character in the field.
</paragraph>
</subSubSection>
<subSubSection id="C35B0991532EFFE1FF129FDCFB0CC7D5" pageId="12" pageNumber="23" type="distribution">
<paragraph id="8BFE5A1A532EFFE1FF129FDCFE21C5FB" blockId="12.[189,495,446,473]" box="[189,495,446,473]" pageId="12" pageNumber="23">
<heading id="D0B6ED76532EFFE1FF129FDCFE21C5FB" bold="true" box="[189,495,446,473]" fontSize="11" level="3" pageId="12" pageNumber="23" reason="3">
<emphasis id="B9358608532EFFE1FF129FDCFE21C5FB" bold="true" box="[189,495,446,473]" pageId="12" pageNumber="23">Habitat and distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BFE5A1A532EFFE1FF129F92FB0CC7D5" blockId="12.[189,1399,496,1014]" pageId="12" pageNumber="23">
Widely distributed in the Eastern Tropical Pacific.
<taxonomicName id="4C412199532EFFE1FCBA9F92FBDDC629" authority="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras, 2020" authorityName="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras" authorityYear="2020" box="[789,1043,496,522]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="23" phylum="Echinodermata" rank="species" species="hendleri" status="sp. nov.">
<emphasis id="B9358608532EFFE1FCBA9F92FBDDC629" box="[789,1043,496,522]" italics="true" pageId="12" pageNumber="23">Ophioderma hendleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2063B73532EFFE1FBB49F92FBB9C629" box="[1051,1143,496,522]" pageId="12" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
has been collected in the Mexican Pacific from the Gulf of California (in the States of Sonora and Baja California Sur), Jalisco, Michoacán, Guerrero and Oaxaca as well as in Revillagigedo, Marías and Marietas Islands. In
<collectingCountry id="F3561A8A532EFFE1FA9B9C5EFF3DC658" name="Costa Rica" pageId="12" pageNumber="23">Costa Rica</collectingCountry>
it has been collected from Murciélago, Caño, and Cocos Islands; in
<collectingCountry id="F3561A8A532EFFE1FB829C00FB44C65F" box="[1069,1162,610,636]" name="Panama" pageId="12" pageNumber="23">Panama</collectingCountry>
from Pearl Islands; and finally, in
<collectingCountry id="F3561A8A532EFFE1FEDE9CE5FE2BC682" box="[369,485,647,673]" name="Colombia" pageId="12" pageNumber="23">Colombia</collectingCountry>
from the localities La Parguera and La Roñosa Rock (see: Supplementary material). The northernmost locality of its distribution is off San Francisquito Bay (Gulf of California,
<collectingCountry id="F3561A8A532EFFE1FF129CB1FED4C6CE" box="[189,282,723,749]" name="Mexico" pageId="12" pageNumber="23">Mexico</collectingCountry>
;
<geoCoordinate id="EE753CDD532EFFE1FE8A9CB1FEA5C6CE" box="[293,363,723,749]" degrees="28" direction="north" orientation="latitude" pageId="12" pageNumber="23" precision="55555" value="28.0">28° N</geoCoordinate>
) and the southernmost point is La Roñosa Rock,
<collectingCountry id="F3561A8A532EFFE1FC0D9CB1FBD8C6CE" box="[930,1046,723,749]" name="Colombia" pageId="12" pageNumber="23">Colombia</collectingCountry>
(
<geoCoordinate id="EE753CDD532EFFE1FB899CB1FB91C6CE" box="[1062,1119,723,749]" degrees="3" direction="north" orientation="latitude" pageId="12" pageNumber="23" precision="55555" value="3.0">3° N</geoCoordinate>
). Due to its distribution,
<taxonomicName id="4C412199532EFFE1FF129C9BFE88C730" authority="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras, 2020" authorityName="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras" authorityYear="2020" box="[189,326,761,787]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="23" phylum="Echinodermata" rank="species" species="hendleri" status="sp. nov.">
<emphasis id="B9358608532EFFE1FF129C9BFE88C730" box="[189,326,761,787]" italics="true" pageId="12" pageNumber="23">O. hendleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2063B73532EFFE1FEFF9C9BFE60C730" box="[336,430,761,787]" pageId="12" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
can be considered as a species with a Panamic affinity, suggesting that, similar to other ophiuroids (e.g.,
<taxonomicName id="4C412199532EFFE1FE549D7CFC6CC71A" authority="Lutken, 1859" authorityName="Lutken" authorityYear="1859" box="[507,930,798,825]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="23" phylum="Echinodermata" rank="species" species="aethiops">
<emphasis id="B9358608532EFFE1FE549D7CFD3DC71B" box="[507,755,798,824]" italics="true" pageId="12" pageNumber="23">Ophiocoma aethiops</emphasis>
<bibRefCitation id="EFD027EB532EFFE1FCAF9D7CFC6CC71A" author="Lutken C. F." box="[768,930,798,825]" pageId="12" pageNumber="23" pagination="75 - 169" refId="ref16788" refString="Lutken C. F. 1859. Additamenta ad Historiam Ophiuridarum. Beskrivelser af de fra Vestindien og Central Amerikas Vestkyst hidtil beckjendte Slangestjerner. Kongelige Danske Videnskabernes Selskab Biologiske Skrifter 2: 75 - 169." type="journal article" year="1859">Lütken, 1859</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C412199532EFFE1FC4B9D7CFEC3C77C" authority="(Lyman, 1860))" authorityName="" baseAuthorityName="Lyman" baseAuthorityYear="1860" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocomella" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="23" phylum="Echinodermata" rank="species" species="alexandri">
<emphasis id="B9358608532EFFE1FC4B9D7CFAC9C71B" box="[996,1287,798,824]" italics="true" pageId="12" pageNumber="23">Ophiocomella alexandri</emphasis>
(
<bibRefCitation id="EFD027EB532EFFE1FAB39D7DFF35C77C" author="Lyman T." pageId="12" pageNumber="23" pagination="252 - 262" refId="ref16936" refString="Lyman T. 1860. Descriptions of new Ophiuridae, belonging to the Smithsonian Institution and to the Museum of Comparative Zoology at Cambridge. Proceedings of the Boston Society of Natural History 7: 193 - 204, 252 - 262. https: // doi. org / 10.5962 / bhl. part. 4822" type="journal article" year="1860">Lyman, 1860</bibRefCitation>
))
</taxonomicName>
, its distribution may extend to northern
<collectingCountry id="F3561A8A532EFFE1FD579D27FCFFC77C" box="[760,817,837,863]" name="Peru" pageId="12" pageNumber="23">Peru</collectingCountry>
, where a transitional zone between the Panamic and the Peruvian Provinces is located (
<bibRefCitation id="EFD027EB532EFFE1FD3C9D08FBFAC7A7" author="Granja-Fernandez R. &amp; Hooker Y." box="[659,1076,874,900]" pageId="12" pageNumber="23" pagination="539 - 556" refId="ref15890" refString="Granja-Fernandez R. &amp; Hooker Y. 2020. Revisiting the diversity and distribution of the ophiuroids (Echinodermata: Ophiuroidea) from Peru. Zootaxa 4766 (4): 539 - 556. https: // doi. org / 10.11646 / zootaxa. 4766.4.2" type="journal article" year="2020">Granja-Fernández &amp; Hooker 2020</bibRefCitation>
), but more sampling effort is needed to confirm its presence or absence in this area. The species inhabits depths between
<quantity id="4CB9F7FF532EFFE1FA9B9DF2FF32C7F3" metricMagnitude="1" metricUnit="m" metricValue="1.8699999999999999" metricValueMax="3.7" metricValueMin="0.04" pageId="12" pageNumber="23" unit="m" value="18.7" valueMax="37.0" valueMin="0.4">
0.40
<specimenCount id="9D479193532EFFE1FF129DD4FF32C7F3" box="[189,252,950,976]" pageId="12" pageNumber="23" type="generic">37 m</specimenCount>
</quantity>
.
<taxonomicName id="4C412199532EFFE1FEA89DD4FDCAC7F3" authority="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras, 2020" authorityName="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras" authorityYear="2020" box="[263,516,950,976]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="23" phylum="Echinodermata" rank="species" species="hendleri" status="sp. nov.">
<emphasis id="B9358608532EFFE1FEA89DD4FDCAC7F3" box="[263,516,950,976]" italics="true" pageId="12" pageNumber="23">Ophioderma hendleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2063B73532EFFE1FDA49DD4FDA8C7F3" box="[523,614,950,976]" pageId="12" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
is mostly collected under rocks, burrowed in the sediment and it is common to find it associated with coral and rocky reefs (
<bibRefCitation id="EFD027EB532EFFE1FCF89DBEFB78C7D5" author="Granja-Fernandez R. &amp; Herrero-Perezrul M. D. &amp; Lopez-Perez R. A. &amp; Hernandez L. &amp; Rodriguez-Zaragoza F. A. &amp; Jones R. W. &amp; Pineda-Lopez R." box="[855,1206,988,1014]" pageId="12" pageNumber="23" pagination="101 - 145" refId="ref15937" refString="Granja-Fernandez R., Herrero-Perezrul M. D., Lopez-Perez R. A., Hernandez L., Rodriguez-Zaragoza F. A., Jones R. W. &amp; Pineda-Lopez R. 2014. Ophiuroidea (Echinodermata) from coral reefs in the Mexican Pacific. Zookeys 406: 101 - 145. https: // doi. org / 10.3897 / zookeys. 406.6306" type="journal article" year="2014">
Granja-Fernández
<emphasis id="B9358608532EFFE1FB9B9DBEFBA6C7D5" box="[1076,1128,988,1014]" italics="true" pageId="12" pageNumber="23">et al</emphasis>
. 2014
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35B0991532EFFE1FF129A42FEE3C3CB" pageId="12" pageNumber="23" type="discussion">
<paragraph id="8BFE5A1A532EFFE1FF129A42FEE1C019" blockId="12.[189,303,1056,1082]" box="[189,303,1056,1082]" pageId="12" pageNumber="23">
<heading id="D0B6ED76532EFFE1FF129A42FEE1C019" bold="true" box="[189,303,1056,1082]" fontSize="11" level="3" pageId="12" pageNumber="23" reason="3">
<emphasis id="B9358608532EFFE1FF129A42FEE1C019" bold="true" box="[189,303,1056,1082]" pageId="12" pageNumber="23">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BFE5A1A532EFFE1FF129A33FAA8C121" blockId="12.[189,1399,1105,1283]" pageId="12" pageNumber="23">
<taxonomicName id="4C412199532EFFE1FF129A33FE74C048" authority="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras, 2020" authorityName="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras" authorityYear="2020" box="[189,442,1105,1131]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="23" phylum="Echinodermata" rank="species" species="hendleri" status="sp. nov.">
<emphasis id="B9358608532EFFE1FF129A33FE74C048" box="[189,442,1105,1131]" italics="true" pageId="12" pageNumber="23">Ophioderma hendleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2063B73532EFFE1FE6F9A33FDD4C048" box="[448,538,1105,1131]" pageId="12" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
is one the most widely distributed ophiodermatids in the Eastern Tropical Pacific and it is of interest to note that although it has been collected since 1936, it has been misidentified in all the reviewed collections as
<taxonomicName id="4C412199532EFFE1FDE89AFFFD3CC095" authorityName=", Lutken" authorityYear="1856" box="[583,754,1181,1207]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="23" phylum="Echinodermata" rank="species" species="variegatum">
<emphasis id="B9358608532EFFE1FDE89AFFFD3CC095" box="[583,754,1181,1207]" italics="true" pageId="12" pageNumber="23">O. variegatum</emphasis>
</taxonomicName>
or
<taxonomicName id="4C412199532EFFE1FCB69AFFFC0CC095" authorityName="Lutken" authorityYear="1859" box="[793,962,1181,1207]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="23" phylum="Echinodermata" rank="species" species="panamense">
<emphasis id="B9358608532EFFE1FCB69AFFFC0CC095" box="[793,962,1181,1207]" italics="true" pageId="12" pageNumber="23">O. panamense</emphasis>
</taxonomicName>
(see Discussion). In previous works,
<bibRefCitation id="EFD027EB532EFFE1FF129AA0FDFFC0FE" author="Granja-Fernandez R. &amp; Herrero-Perezrul M. D. &amp; Lopez-Perez R. A. &amp; Hernandez L. &amp; Rodriguez-Zaragoza F. A. &amp; Jones R. W. &amp; Pineda-Lopez R." box="[189,561,1218,1245]" pageId="12" pageNumber="23" pagination="101 - 145" refId="ref15937" refString="Granja-Fernandez R., Herrero-Perezrul M. D., Lopez-Perez R. A., Hernandez L., Rodriguez-Zaragoza F. A., Jones R. W. &amp; Pineda-Lopez R. 2014. Ophiuroidea (Echinodermata) from coral reefs in the Mexican Pacific. Zookeys 406: 101 - 145. https: // doi. org / 10.3897 / zookeys. 406.6306" type="journal article" year="2014">
Granja-Fernández
<emphasis id="B9358608532EFFE1FE329AA1FE18C0FF" box="[413,470,1218,1244]" italics="true" pageId="12" pageNumber="23">et al</emphasis>
. (2014
</bibRefCitation>
,
<bibRefCitation id="EFD027EB532EFFE1FDEE9AA0FD42C0FE" author="Granja-Fernandez R. &amp; Herrero-Perezrul M. D. &amp; Lopez-Perez A. &amp; Hernandez-Morales A. &amp; Rangel-Solis P. D." box="[577,652,1218,1245]" pageId="12" pageNumber="23" pagination="37 - 47" refId="ref16005" refString="Granja-Fernandez R., Herrero-Perezrul M. D., Lopez-Perez A., Hernandez-Morales A. &amp; Rangel-Solis P. D. 2015 a. A literature review of the Ophiuroidea (Echinodermata) from the Pacific coast of Mexico. Revista de Biologia Tropical 63 (2): 37 - 47. https: // doi. org / 10.15517 / RBT. V 63 I 2.23127" type="journal article" year="2015">2015a</bibRefCitation>
,
<bibRefCitation id="EFD027EB532EFFE1FD329AA1FD17C0FE" author="Granja-Fernandez R. &amp; Rodriguez-Troncoso A. P. &amp; Herrero-Perezrul M. D. &amp; Sotelo-Casas R. C. &amp; Ortega J. R. &amp; Godinez-Dominguez E. &amp; Salazar-Silva P. &amp; Alarcon-Ortega L. C. &amp; Cazares-Salazar A. &amp; Cupul-Magana A. L." box="[669,729,1219,1245]" pageId="12" pageNumber="23" pagination="167 - 177" refId="ref16145" refString="Granja-Fernandez R., Rodriguez-Troncoso A. P., Herrero-Perezrul M. D., Sotelo-Casas R. C., Flores- Ortega J. R., Godinez-Dominguez E., Salazar-Silva P., Alarcon-Ortega L. C., Cazares-Salazar A. &amp; Cupul-Magana A. L. 2017. Ophiuroidea (Echinodermata) from the Central Mexican Pacific: an updated checklist including new distribution records. Marine Biodiversity 47: 167 - 177. https: // doi. org / 10.1007 / s 12526 - 016 - 0459 - 4" type="journal article" year="2017">2017</bibRefCitation>
) mentioned the presence of
<taxonomicName id="4C412199532EFFE1FB919AA0FB1CC0FF" authorityName="Muller &amp; Troschel" authorityYear="1840" box="[1086,1234,1218,1244]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="23" phylum="Echinodermata" rank="genus">
<emphasis id="B9358608532EFFE1FB919AA0FB1CC0FF" box="[1086,1234,1218,1244]" italics="true" pageId="12" pageNumber="23">Ophioderma</emphasis>
</taxonomicName>
sp. 1, which actually corresponds to
<taxonomicName id="4C412199532EFFE1FE7B9A8BFD94C121" authority="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras, 2020" authorityName="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras" authorityYear="2020" box="[468,602,1256,1283]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="23" phylum="Echinodermata" rank="species" species="hendleri" status="sp. nov.">
<emphasis id="B9358608532EFFE1FE7B9A8BFD94C121" box="[468,602,1256,1283]" italics="true" pageId="12" pageNumber="23">O. hendleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2063B73532EFFE1FDCE9A8BFD71C120" box="[609,703,1257,1283]" pageId="12" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
; therefore, these records are considered to be this species.
</paragraph>
<paragraph id="8BFE5A1A532EFFE1FF129B4FFD47C209" blockId="12.[189,1399,1324,1578]" pageId="12" pageNumber="23">
In our revision of the specimens, we did not observe any differences in the morphological characters from different geographical areas, suggesting that intraspecific variation in
<taxonomicName id="4C412199532EFFE1FB909B31FB09C14F" authority="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras, 2020" authorityName="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras" authorityYear="2020" box="[1087,1223,1362,1389]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="23" phylum="Echinodermata" rank="species" species="hendleri" status="sp. nov.">
<emphasis id="B9358608532EFFE1FB909B31FB09C14F" box="[1087,1223,1362,1389]" italics="true" pageId="12" pageNumber="23">O. hendleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2063B73532EFFE1FB609B31FAE2C14E" box="[1231,1324,1363,1389]" pageId="12" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
is not related to its distribution. On the other hand, it is very notable that this species in most cases presents the same color pattern (dorsal side dark brown, ventral side beige and arms with transversal lines;
<figureCitation id="137A469F532EFFE1FA8A9BFCFABEC19B" box="[1317,1392,1438,1464]" captionStart="Fig" captionStartId="5.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@5.[189,1398,291,1855]" captionTargetPageId="5" captionText="Fig. 1. Ophioderma hendleri sp. nov., holotype (ICML-UNAM 18315, DD = 14.8 mm). A. Dorsal disc. B. Ventral disc. C. Mouth. D. Structures of a jaw (according to Hendler 2018). E. Basal dorsal arm. F. Dorsal arm. G. Ventral arm. H. Lateral arm plates and arm spines. I. Dorsal view. See Material and methods for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4326613" httpUri="https://zenodo.org/record/4326613/files/figure.png" pageId="12" pageNumber="23">Figs 1</figureCitation>
,
<figureCitation id="137A469F532EFFE1FF129BA6FF22C1FC" box="[189,236,1476,1503]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="7.[189,232,1929,1955]" captionStartId-1="8.[189,232,1928,1954]" captionStartId-2="9.[189,232,1893,1919]" captionTargetBox-0="[189,1398,327,1891]" captionTargetBox-1="[189,1398,325,1889]" captionTargetBox-2="[189,1398,291,1855]" captionTargetId-0="figure-11@7.[189,1398,327,1891]" captionTargetId-1="figure-17@8.[189,1398,325,1889]" captionTargetId-2="figure-12@9.[189,1398,291,1855]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="Fig. 3. Ophioderma hendleri sp. nov., paratype (ICML-UNAM 10582, DD = 4.8 mm). A. Dorsal view. B. Basal dorsal arm. C. Dorsal arm. D. Ventral arm. E. Dorsal disc. F. Ventral disc. G. Interradii. H. Mouth." captionText-1="Fig. 4. Ophioderma hendleri sp. nov., paratype (ICML-UNAM 18319, DD = 2.2 mm). A. Dorsal view. B. Basal dorsal arm. C. Dorsal arm. D. Ventral arm. E. Dorsal disc. F. Ventral disc. G. Interradii. H. Mouth." captionText-2="Fig. 5. AD. Ophioderma hendleri sp. nov. AB. In vivo coloration in natural habitat. CD. In vivo coloration of collected specimens. EJ. Ophioderma panamense Lütken, 1859, holotype (NHMD- 107679, DD = 21.9 mm). E. Dorsal disc. F. Ventral disc. G. Mouth. H. Dorsal arm. I. Ventral arm. J. Dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.4326619" figureDoi-1="http://doi.org/10.5281/zenodo.4326621" figureDoi-2="http://doi.org/10.5281/zenodo.4326625" httpUri-0="https://zenodo.org/record/4326619/files/figure.png" httpUri-1="https://zenodo.org/record/4326621/files/figure.png" httpUri-2="https://zenodo.org/record/4326625/files/figure.png" pageId="12" pageNumber="23">35</figureCitation>
), with some exceptions (see
<typeStatus id="54FAE4B8532EFFE1FDEA9BA6FD62C1FD" box="[581,684,1476,1502]" pageId="12" pageNumber="23" type="paratype">Paratype</typeStatus>
variations). This is in contrast to other
<taxonomicName id="4C412199532EFFE1FB2A9BA6FAD7C1FD" authorityName="Muller &amp; Troschel" authorityYear="1840" box="[1157,1305,1476,1502]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="23" phylum="Echinodermata" rank="genus">
<emphasis id="B9358608532EFFE1FB2A9BA6FAD7C1FD" box="[1157,1305,1476,1502]" italics="true" pageId="12" pageNumber="23">Ophioderma</emphasis>
</taxonomicName>
species such as
<taxonomicName id="4C412199532EFFE1FEB89B88FE70C220" authorityName="Lutken" authorityYear="1859" box="[279,446,1514,1540]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="23" phylum="Echinodermata" rank="species" species="panamense">
<emphasis id="B9358608532EFFE1FEB89B88FE70C220" box="[279,446,1514,1540]" italics="true" pageId="12" pageNumber="23">O. panamense</emphasis>
</taxonomicName>
or
<taxonomicName id="4C412199532EFFE1FE4C9B88FD45C220" authorityName=", Lutken" authorityYear="1856" box="[483,651,1514,1540]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="23" phylum="Echinodermata" rank="species" species="variegatum">
<emphasis id="B9358608532EFFE1FE4C9B88FD45C220" box="[483,651,1514,1540]" italics="true" pageId="12" pageNumber="23">O. variegatum</emphasis>
</taxonomicName>
, which can display a great variety of colors (green, yellow, pink, etc.;
<bibRefCitation id="EFD027EB532EFFE1FF59986DFE5BC209" author="Nielsen E." box="[246,405,1551,1578]" pageId="12" pageNumber="23" pagination="241 - 346" refId="ref17315" refString="Nielsen E. 1932. Ophiurans from the Gulf of Panama, California and the Strait of Georgia. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening 91: 241 - 346." type="journal article" year="1932">Nielsen 1932</bibRefCitation>
;
<bibRefCitation id="EFD027EB532EFFE1FE0E986DFDB2C209" author="Ziesenhenne F. C." box="[417,636,1551,1578]" pageId="12" pageNumber="23" pagination="185 - 201" refId="ref18074" refString="Ziesenhenne F. C. 1955. A review of the genus Ophioderma Muller and Troschel. In: Allan Hancock Foundation for Scientific Research (eds) Essays in the Natural Sciences in Honor of Captain Allan Hancock: 185 - 201. University of Southern California Press, Los Angeles." type="book chapter" year="1955">Ziesenhenne 1955</bibRefCitation>
).
</paragraph>
<paragraph id="8BFE5A1A532EFFE1FF129831FEE3C3CB" blockId="12.[189,1399,1619,2024]" pageId="12" pageNumber="23">
This species is commonly found at the same localities as
<taxonomicName id="4C412199532EFFE1FCC29836FBD9C24E" authorityName="Lutken" authorityYear="1859" box="[877,1047,1620,1646]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="23" phylum="Echinodermata" rank="species" species="panamense">
<emphasis id="B9358608532EFFE1FCC29836FBD9C24E" box="[877,1047,1620,1646]" italics="true" pageId="12" pageNumber="23">O. panamense</emphasis>
</taxonomicName>
and other ophiuroids such as
<taxonomicName id="4C412199532EFFE1FF12981BFD90C2B7" authority="(Le Conte, 1851)" baseAuthorityName="Le Conte" baseAuthorityYear="1851" box="[189,606,1657,1684]" class="Ophiuroidea" family="Ophiactidae" genus="Ophiactis" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="23" phylum="Echinodermata" rank="species" species="simplex">
<emphasis id="B9358608532EFFE1FF12981BFE41C2B0" box="[189,399,1657,1683]" italics="true" pageId="12" pageNumber="23">Ophiactis simplex</emphasis>
(Le Conte, 1851)
</taxonomicName>
,
<taxonomicName id="4C412199532EFFE1FDC6981BFBBCC2B0" authority="(Muller &amp; Troshel, 1842)" baseAuthorityName="Muller &amp; Troshel" baseAuthorityYear="1842" box="[617,1138,1657,1683]" class="Ophiuroidea" family="Ophiactidae" genus="Ophiactis" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="23" phylum="Echinodermata" rank="species" species="savignyi">
<emphasis id="B9358608532EFFE1FDC6981BFC8DC2B0" box="[617,835,1657,1683]" italics="true" pageId="12" pageNumber="23">Ophiactis savignyi</emphasis>
(Müller &amp; Troshel, 1842)
</taxonomicName>
,
<taxonomicName id="4C412199532EFFE1FBD2981BFAA1C2B0" authorityName="Lutken" authorityYear="1859" box="[1149,1391,1657,1683]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocoma" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="23" phylum="Echinodermata" rank="species" species="aethiops">
<emphasis id="B9358608532EFFE1FBD2981BFAA1C2B0" box="[1149,1391,1657,1683]" italics="true" pageId="12" pageNumber="23">Ophiocoma aethiops</emphasis>
</taxonomicName>
,
<taxonomicName id="4C412199532EFFE1FF1298FDFE10C29A" authorityName="" baseAuthorityName="Lyman" baseAuthorityYear="1860" box="[189,478,1695,1721]" class="Ophiuroidea" family="Ophiocomidae" genus="Ophiocomella" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="23" phylum="Echinodermata" rank="species" species="alexandri">
<emphasis id="B9358608532EFFE1FF1298FDFE10C29A" box="[189,478,1695,1721]" italics="true" pageId="12" pageNumber="23">Ophiocomella alexandri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C412199532EFFE1FE5F98FDFC00C299" authority="(Le Conte, 1851)" baseAuthorityName="Le Conte" baseAuthorityYear="1851" box="[496,974,1695,1722]" class="Ophiuroidea" family="Ophionereididae" genus="Ophionereis" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="23" phylum="Echinodermata" rank="species" species="annulata">
<emphasis id="B9358608532EFFE1FE5F98FDFD3CC29A" box="[496,754,1695,1721]" italics="true" pageId="12" pageNumber="23">Ophionereis annulata</emphasis>
(Le Conte, 1851)
</taxonomicName>
,
<taxonomicName id="4C412199532EFFE1FC7298FDFAB9C29A" authority="Verrill, 1867" authorityName="Verrill" authorityYear="1867" box="[989,1399,1695,1721]" class="Ophiuroidea" family="Ophiotrichidae" genus="Ophiothela" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="23" phylum="Echinodermata" rank="species" species="mirabilis">
<emphasis id="B9358608532EFFE1FC7298FDFB1BC29A" box="[989,1237,1695,1721]" italics="true" pageId="12" pageNumber="23">Ophiothela mirabilis</emphasis>
<bibRefCitation id="EFD027EB532EFFE1FB7098FDFAB9C29A" author="Verrill A. E." box="[1247,1399,1695,1721]" pageId="12" pageNumber="23" pagination="251 - 322" refId="ref17963" refString="Verrill A. E. 1867. V. Notes on the Radiata in the Museum of Yale College, with descriptions of new genera and species. No. 2. Notes on the echinoderms of Panama and West Coast of America, with descriptions of new genera and species. Transactions of the Connecticut Academy of Arts and Sciences 1 (2): 251 - 322." type="journal article" year="1867">Verrill, 1867</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C412199532EFFE1FF5F98A7FC8BC2C3" authority="Le Conte, 1851" authorityName="Le Conte" authorityYear="1851" box="[240,837,1733,1760]" class="Ophiuroidea" family="Ophiotrichidae" genus="Ophiothrix" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="23" phylum="Echinodermata" rank="species" species="spiculata" subGenus="Ophiothrix">
<emphasis id="B9358608532EFFE1FF5F98A7FEBFC2FC" box="[240,369,1733,1759]" italics="true" pageId="12" pageNumber="23">Ophiothrix</emphasis>
(
<emphasis id="B9358608532EFFE1FE2C98A7FDCAC2FC" box="[387,516,1733,1759]" italics="true" pageId="12" pageNumber="23">Ophiothrix</emphasis>
)
<emphasis id="B9358608532EFFE1FDB998A7FD4CC2FC" box="[534,642,1733,1759]" italics="true" pageId="12" pageNumber="23">spiculata</emphasis>
Le Conte, 1851
</taxonomicName>
. It is one of the most conspicuous
<taxonomicName id="4C412199532EFFE1FB4D98A7FAB8C2FC" authorityName="Muller &amp; Troschel" authorityYear="1840" box="[1250,1398,1733,1759]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="23" phylum="Echinodermata" rank="genus">
<emphasis id="B9358608532EFFE1FB4D98A7FAB8C2FC" box="[1250,1398,1733,1759]" italics="true" pageId="12" pageNumber="23">Ophioderma</emphasis>
</taxonomicName>
species of the Eastern Tropical Pacific and it is always hiding under small and medium-sized rocks and in sediment of median size of rocky and coral reefs (
<figureCitation id="137A469F532EFFE1FC8C9973FCA9C308" box="[803,871,1808,1835]" captionStart="Fig" captionStartId="9.[189,232,1893,1919]" captionTargetBox="[189,1398,291,1855]" captionTargetId="figure-12@9.[189,1398,291,1855]" captionTargetPageId="9" captionText="Fig. 5. AD. Ophioderma hendleri sp. nov. AB. In vivo coloration in natural habitat. CD. In vivo coloration of collected specimens. EJ. Ophioderma panamense Lütken, 1859, holotype (NHMD- 107679, DD = 21.9 mm). E. Dorsal disc. F. Ventral disc. G. Mouth. H. Dorsal arm. I. Ventral arm. J. Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4326625" httpUri="https://zenodo.org/record/4326625/files/figure.png" pageId="12" pageNumber="23">Fig. 5</figureCitation>
AB). Its abundance has not been quantified, but using scuba diving, in an hour of sampling effort of all the species of ophiuroids at one locality, a total of 5
<specimenCount id="9D479193532EFFE1FE98993EFE06C355" box="[311,456,1884,1911]" pageId="12" pageNumber="23" type="generic">6 specimens</specimenCount>
were found, a smaller number compared to
<taxonomicName id="4C412199532EFFE1FC7B993FFBB3C355" authorityName="Lutken" authorityYear="1859" box="[980,1149,1885,1911]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="23" phylum="Echinodermata" rank="species" species="panamense">
<emphasis id="B9358608532EFFE1FC7B993FFC27C354" box="[980,1001,1885,1911]" italics="true" pageId="12" pageNumber="23">O</emphasis>
.
<emphasis id="B9358608532EFFE1FC56993FFBB3C355" box="[1017,1149,1885,1910]" italics="true" pageId="12" pageNumber="23">panamense</emphasis>
</taxonomicName>
(8
<specimenCount id="9D479193532EFFE1FB03993FFA9FC354" box="[1196,1361,1884,1911]" pageId="12" pageNumber="23" type="generic">12 specimens</specimenCount>
). It tends to be gregarious, and it is very common to find at least two specimens occupying the same space. Field notes denote that
<taxonomicName id="4C412199532EFFE1FE7399CAFDABC3E1" authority="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras, 2020" authorityName="Granja-Fernández &amp; Pineda-Enríquez &amp; Solís-Marín &amp; Laguarda-Figueras" authorityYear="2020" box="[476,613,1960,1986]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="23" phylum="Echinodermata" rank="species" species="hendleri" status="sp. nov.">
<emphasis id="B9358608532EFFE1FE7399CAFDABC3E1" box="[476,613,1960,1986]" italics="true" pageId="12" pageNumber="23">O. hendleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2063B73532EFFE1FDC099CBFD03C3E0" box="[623,717,1961,1987]" pageId="12" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
has been found in waters with a bottom temperature of 2729°C.
</paragraph>
</subSubSection>
</treatment>
</document>