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<document id="0346E090886E9DCE4248B3BD35882D55" ID-DOI="10.11646/zootaxa.5541.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14240870" ID-ZooBank="86A7CD79-F5A2-4AA6-A6AA-01C0CB64B29C" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1732833600772" checkinUser="plazi" docAuthor="Karanovic, Tomislav" docDate="2024" docId="03E8C55F9C08FFDCFF0B923CFB3F674F" docLanguage="en" docName="zootaxa.5541.2.1.pdf" docOrigin="Zootaxa 5541 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5541.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Diacyclops leijsi Karanovic 2024, sp. nov." docType="treatment" docVersion="4" lastPageNumber="111" masterDocId="FFD1BD279C0AFFD6FF9C9466FFE06572" masterDocTitle="Six new species of Diacyclops (Copepoda: Cyclopoida) from Australia" masterLastPageNumber="143" masterPageNumber="101" pageNumber="103" updateTime="1733245968451" updateUser="jonas" zenodo-license-document="CLOSED">
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<mods:title id="CA9761EB7CFE5E8F015BDB50407B296F">Six new species of Diacyclops (Copepoda: Cyclopoida) from Australia</mods:title>
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<mods:namePart id="693D2F62D57DBF4EAB65EB81ADD70706">Karanovic, Tomislav</mods:namePart>
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<mods:title id="97BF97F1DDDC023A0B19BC36C79D14C5">Zootaxa</mods:title>
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<mods:date id="4A40DFE46DB3C011EF5DDA6615976507">2024</mods:date>
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<mods:number id="F086FCFEEF170C8F0114519AA11ED977">2024-11-26</mods:number>
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<treatment id="03E8C55F9C08FFDCFF0B923CFB3F674F" ID-DOI="http://doi.org/10.5281/zenodo.14248341" ID-Zenodo-Dep="14248341" LSID="urn:lsid:plazi:treatment:03E8C55F9C08FFDCFF0B923CFB3F674F" httpUri="http://treatment.plazi.org/id/03E8C55F9C08FFDCFF0B923CFB3F674F" lastPageId="10" lastPageNumber="111" pageId="2" pageNumber="103">
<subSubSection id="C35B27C29C08FFD4FF0B923CFE206307" box="[151,448,1626,1653]" pageId="2" pageNumber="103" type="nomenclature">
<paragraph id="8BFE74499C08FFD4FF0B923CFE206307" blockId="2.[151,974,1626,1725]" box="[151,448,1626,1653]" pageId="2" pageNumber="103">
<heading id="D0B6C3259C08FFD4FF0B923CFE206307" bold="true" box="[151,448,1626,1653]" fontSize="11" level="1" pageId="2" pageNumber="103" reason="1">
<emphasis id="B935A85B9C08FFD4FF0B923CFE206307" bold="true" box="[151,448,1626,1653]" pageId="2" pageNumber="103">
<taxonomicName id="4C410FCA9C08FFD4FF0B923CFEBB6307" authority="Karanovic, 2024" authorityName="Karanovic" authorityYear="2024" box="[151,347,1626,1653]" class="Copepoda" family="Cyclopidae" genus="Diacyclops" kingdom="Animalia" order="Cyclopoida" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="leijsi" status="sp. nov.">
<emphasis id="B935A85B9C08FFD4FF0B923CFEBB6307" bold="true" box="[151,347,1626,1653]" italics="true" pageId="2" pageNumber="103">Diacyclops leijsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20615209C08FFD4FEFF923DFE206307" box="[355,448,1627,1653]" pageId="2" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35B27C29C08FFD4FF0B9219FC2E63CF" pageId="2" pageNumber="103" type="description">
<paragraph id="8BFE74499C08FFD4FF0B9219FEA263EB" blockId="2.[151,974,1626,1725]" box="[151,322,1663,1689]" pageId="2" pageNumber="103">
(
<figureCitation id="137A68CC9C08FFD4FF039219FEE363EB" box="[159,259,1663,1689]" captionStart="FIGURE 1" captionStartId="3.[151,250,1059,1083]" captionTargetBox="[155,1436,181,1036]" captionTargetId="figure-15@3.[151,1436,181,1036]" captionTargetPageId="3" captionText="FIGURE 1. Habitus in dorsal view of holotype females of all six new species: A, Diacyclops leijsi sp. nov.; B, D. eberhardi sp. nov.; C, D. ballaballaensis sp. nov.; D, D. hancocki sp. nov.; E, D. tomlinsonae sp. nov.; F, D. huntervalleyensis sp. nov. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14240872" httpUri="https://zenodo.org/record/14240872/files/figure.png" pageId="2" pageNumber="103">Figs. 1A</figureCitation>
,
<figureCitation id="137A68CC9C08FFD4FE939219FEDB63EB" box="[271,315,1663,1689]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStartId-0="4.[151,250,1943,1967]" captionStartId-1="5.[151,250,1621,1645]" captionStartId-2="6.[151,250,1943,1967]" captionStartId-3="7.[151,250,1828,1852]" captionStartId-4="8.[151,250,1544,1568]" captionStartId-5="9.[151,250,1887,1911]" captionTargetBox-0="[174,1414,181,1914]" captionTargetBox-1="[151,1435,181,1594]" captionTargetBox-2="[171,1411,181,1918]" captionTargetBox-3="[159,1436,181,1797]" captionTargetBox-4="[151,1436,181,1520]" captionTargetBox-5="[151,1436,525,1864]" captionTargetId-0="figure-27@4.[174,1414,181,1919]" captionTargetId-1="figure-15@5.[151,1435,181,1597]" captionTargetId-2="figure-27@6.[171,1415,181,1919]" captionTargetId-3="figure-15@7.[151,1436,181,1804]" captionTargetId-4="figure-27@8.[151,1436,181,1520]" captionTargetId-5="figure-201@9.[151,1436,525,1864]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionText-0="FIGURE 2. Diacyclops leijsi sp. nov., line drawings, holotype female: A, urosome, ventral view; B, antennula. Scale bar 50 μm." captionText-1="FIGURE 3. Diacyclops leijsi sp. nov., line drawings, holotype female:A, antenna; B, labrum; C, mandibula; D, cutting edge of mandibula; E. maxillula; F, praecoxal arthrite of maxillula; G, maxilla; H, maxilliped. Scale bar 50 μm." captionText-2="FIGURE 4. Diacyclops leijsi sp. nov., line drawings, holotype female: A, first swimming leg; B, second swimming leg; C, endopod of third swimming leg; D, third exopodal segment of third swimming leg; E, fourth swimming leg; F, fifth leg; G, sixth leg. Scale bar 50 μm." captionText-3="FIGURE 5. Diacyclops leijsi sp. nov., line drawings, paratype male 1: A, urosome, ventral; B, antennula; C, third endopodal segment of fourth swimming leg; D, sixth leg. Scale bar 50 μm." captionText-4="FIGURE 6. Diacyclops leijsi sp. nov., scanning electron micrographs, AD, paratype female 1, ventral; E, paratype female 2, lateral; F, paratype male 2, lateral: A, habitus; B, caudal rami; C, mouth appendages; D, fourth swimming leg; E, distal part of antennula; F, habitus." captionText-5="FIGURE 7. Diacyclops leijsi sp. nov., scanning electron micrographs, A &amp; B, paratype male 2, lateral; CF, paratype male 3, dorsal:A, left antennula; B, detail of right antennula; C, free prosomites; D, first two urosomites; E, third and fourth urosomites; F, last two urosomites." figureDoi-0="http://doi.org/10.5281/zenodo.14240874" figureDoi-1="http://doi.org/10.5281/zenodo.14240876" figureDoi-2="http://doi.org/10.5281/zenodo.14240878" figureDoi-3="http://doi.org/10.5281/zenodo.14240880" figureDoi-4="http://doi.org/10.5281/zenodo.14240882" figureDoi-5="http://doi.org/10.5281/zenodo.14240884" httpUri-0="https://zenodo.org/record/14240874/files/figure.png" httpUri-1="https://zenodo.org/record/14240876/files/figure.png" httpUri-2="https://zenodo.org/record/14240878/files/figure.png" httpUri-3="https://zenodo.org/record/14240880/files/figure.png" httpUri-4="https://zenodo.org/record/14240882/files/figure.png" httpUri-5="https://zenodo.org/record/14240884/files/figure.png" pageId="2" pageNumber="103">27</figureCitation>
)
</paragraph>
<paragraph id="8BFE74499C08FFD4FF0B92C4FC2E63CF" blockId="2.[151,974,1626,1725]" box="[151,974,1698,1725]" pageId="2" pageNumber="103">
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urn:lsid:zoobank.org:act:
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</paragraph>
</subSubSection>
<subSubSection id="C35B27C29C08FFD4FF0B928DFCB462CB" pageId="2" pageNumber="103" type="materials_examined">
<paragraph id="8BFE74499C08FFD4FF0B928DFDFD625B" blockId="2.[151,1436,1770,1977]" pageId="2" pageNumber="103">
<emphasis id="B935A85B9C08FFD4FF0B928DFED56276" bold="true" box="[151,309,1770,1797]" pageId="2" pageNumber="103">Type locality.</emphasis>
<collectingRegion id="4985BAAB9C08FFD4FEA7928DFE0B6277" box="[315,491,1771,1797]" country="Australia" name="South Australia" pageId="2" pageNumber="103">
South
<collectingCountry id="F35634D99C08FFD4FE1D928DFE0B6277" box="[385,491,1771,1797]" name="Australia" pageId="2" pageNumber="103">Australia</collectingCountry>
</collectingRegion>
,
<collectingMunicipality id="6B9AEE339C08FFD4FE68928DFC856277" box="[500,869,1771,1797]" latitude="-29.453112" longLatPrecision="48" longitude="136.858" name="Wabma Kadarbu Mound Springs" pageId="2" pageNumber="103" stateProvince="South Australia">Wabma Kadarbu Mound Springs</collectingMunicipality>
, sample no. TC17049,
<geoCoordinate id="EE75128E9C08FFD4FBFA928CFB156277" box="[1126,1269,1770,1797]" degrees="29.453111" direction="south" orientation="latitude" pageId="2" pageNumber="103" precision="1" value="-29.453112">29.453111°S</geoCoordinate>
<geoCoordinate id="EE75128E9C08FFD4FB60928CFA966277" box="[1276,1398,1770,1797]" degrees="136.858" direction="east" orientation="longitude" pageId="2" pageNumber="103" precision="55" value="136.858">136.858°E</geoCoordinate>
,
<date id="FFFF52899C08FFD4FA1C928CFEC2625B" pageId="2" pageNumber="103" value="2003-03-15">
<collectingDate id="EFBBAB619C08FFD4FA1C928CFEC2625B" pageId="2" pageNumber="103" value="2003-03-15">15 March 2003</collectingDate>
</date>
, collected by
<collectorName id="26B4119F9C08FFD4FE209369FDF9625B" box="[444,537,1807,1833]" pageId="2" pageNumber="103">R. Leijs</collectorName>
.
</paragraph>
<paragraph id="8BFE74499C08FFD4FF5B9355FCBF623F" blockId="2.[151,1436,1770,1977]" box="[199,863,1843,1869]" pageId="2" pageNumber="103">
<materialsCitation id="3B297E149C08FFD4FF5B9355FCBF623F" box="[199,863,1843,1869]" collectingDate="2003-03-15" collectorName="R. Leijs" latitude="-29.453112" location="Wabma Kadarbu Mound Springs" longLatPrecision="48" longitude="136.858" municipality="Wabma Kadarbu Mound Springs" pageId="2" pageNumber="103" specimenCount="2" specimenCount-adult="1" specimenCount-female="1" stateProvince="South Australia" typeStatus="holotype">
<emphasis id="B935A85B9C08FFD4FF5B9355FEDB623F" bold="true" box="[199,315,1843,1869]" pageId="2" pageNumber="103">
<typeStatus id="54FACAEB9C08FFD4FF5B9355FED6623F" box="[199,310,1843,1869]" pageId="2" pageNumber="103" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="9D47BFC09C08FFD4FEDD9355FE61623F" box="[321,385,1843,1869]" count="1" pageId="2" pageNumber="103" type="adult">Adult</specimenCount>
<specimenCount id="9D47BFC09C08FFD4FE1B9355FE33623F" box="[391,467,1843,1869]" count="1" pageId="2" pageNumber="103" type="female">female</specimenCount>
dissected on one microscope slide.
</materialsCitation>
</paragraph>
<paragraph id="8BFE74499C08FFD4FF5B9331FCB462CB" blockId="2.[151,1436,1770,1977]" pageId="2" pageNumber="103">
<materialsCitation id="3B297E149C08FFD4FF5B9331FB106203" box="[199,1264,1879,1905]" collectingDate="2003-03-15" collectorName="R. Leijs" latitude="-29.453112" location="Wabma Kadarbu Mound Springs" longLatPrecision="48" longitude="136.858" municipality="Wabma Kadarbu Mound Springs" pageId="2" pageNumber="103" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="South Australia" typeStatus="paratype">
<emphasis id="B935A85B9C08FFD4FF5B9331FEA66203" bold="true" box="[199,326,1879,1905]" pageId="2" pageNumber="103">
<typeStatus id="54FACAEB9C08FFD4FF5B9331FEA16203" box="[199,321,1879,1905]" pageId="2" pageNumber="103" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9D47BFC09C08FFD4FECC9331FE5E6203" box="[336,446,1879,1905]" count="1" pageId="2" pageNumber="103" type="male">One male</specimenCount>
and
<specimenCount id="9D47BFC09C08FFD4FE669331FDBD6203" box="[506,605,1879,1905]" count="1" pageId="2" pageNumber="103" type="female">1 female</specimenCount>
from type locality dissected on 1 microscope slide each;
</materialsCitation>
<materialsCitation id="3B297E149C08FFD4FB669331FAFB62E7" collectingDate="2003-03-15" collectorName="R. Leijs" latitude="-29.453112" location="Wabma Kadarbu Mound Springs" longLatPrecision="48" longitude="136.858" municipality="Wabma Kadarbu Mound Springs" pageId="2" pageNumber="103" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="South Australia">
<specimenCount id="9D47BFC09C08FFD4FB669331FAB26203" box="[1274,1362,1879,1905]" count="2" pageId="2" pageNumber="103" type="male">2 males</specimenCount>
and
<specimenCount id="9D47BFC09C08FFD4FA129331FF0E62E7" count="3" pageId="2" pageNumber="103" type="female">3 females</specimenCount>
from type locality on 1 SEM stub (row no. 6), together with 5 other species described here;
</materialsCitation>
<materialsCitation id="3B297E149C08FFD4FAB9931DFCAF62CB" collectingDate="2003-03-15" collectorName="R. Leijs" latitude="-29.453112" location="Wabma Kadarbu Mound Springs" longLatPrecision="48" longitude="136.858" municipality="Wabma Kadarbu Mound Springs" pageId="2" pageNumber="103" specimenCount="11" specimenCount-female="7" specimenCount-male="4" stateProvince="South Australia">
<specimenCount id="9D47BFC09C08FFD4FAB9931DFA6062E7" box="[1317,1408,1915,1941]" count="4" pageId="2" pageNumber="103" type="male">4 males</specimenCount>
,
<specimenCount id="9D47BFC09C08FFD4FA12931DFF1162CB" count="7" pageId="2" pageNumber="103" type="female">7 females</specimenCount>
, and 2 copepodids from type locality in 1 alcohol vial
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF3E24C19C09FFD5FF0B9045FEB161F6" ID-DOI="http://doi.org/10.5281/zenodo.14240872" ID-Zenodo-Dep="14240872" httpUri="https://zenodo.org/record/14240872/files/figure.png" pageId="3" pageNumber="104" startId="3.[151,250,1059,1083]" targetBox="[155,1436,181,1036]" targetPageId="3" targetType="figure">
<paragraph id="8BFE74499C09FFD5FF0B9045FEB161F6" blockId="3.[151,1437,1059,1156]" pageId="3" pageNumber="104">
<emphasis id="B935A85B9C09FFD5FF0B9045FEF4614E" bold="true" box="[151,276,1059,1084]" pageId="3" pageNumber="104">FIGURE 1.</emphasis>
Habitus in dorsal view of holotype females of all six new species: A,
<taxonomicName id="4C410FCA9C09FFD5FC749042FB73614E" authorityName="Karanovic" authorityYear="2024" box="[1000,1171,1060,1084]" class="Copepoda" family="Cyclopidae" genus="Diacyclops" kingdom="Animalia" order="Cyclopoida" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="leijsi" status="sp. nov.">
<emphasis id="B935A85B9C09FFD5FC749042FB73614E" box="[1000,1171,1060,1084]" italics="true" pageId="3" pageNumber="104">Diacyclops leijsi</emphasis>
</taxonomicName>
<emphasis id="B935A85B9C09FFD5FB069042FB0C614E" bold="true" box="[1178,1260,1060,1084]" pageId="3" pageNumber="104">
<taxonomicNameLabel id="A20615209C09FFD5FB069042FB0C614E" box="[1178,1260,1060,1084]" pageId="3" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; B,
<taxonomicName id="4C410FCA9C09FFD5FA869042FA7D614E" authorityName="Karanovic" authorityYear="2024" box="[1306,1437,1060,1084]" class="Copepoda" family="Cyclopidae" genus="Diacyclops" kingdom="Animalia" order="Cyclopoida" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="eberhardi" status="sp. nov.">
<emphasis id="B935A85B9C09FFD5FA869042FA7D614E" box="[1306,1437,1060,1084]" italics="true" pageId="3" pageNumber="104">D. eberhardi</emphasis>
</taxonomicName>
<emphasis id="B935A85B9C09FFD5FF0B902EFF086112" bold="true" box="[151,232,1096,1120]" pageId="3" pageNumber="104">
<taxonomicNameLabel id="A20615209C09FFD5FF0B902EFF086112" box="[151,232,1096,1120]" pageId="3" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; C,
<taxonomicName id="4C410FCA9C09FFD5FE89902EFE306112" authorityName="Karanovic" authorityYear="2024" box="[277,464,1096,1120]" class="Copepoda" family="Cyclopidae" genus="Diacyclops" kingdom="Animalia" order="Cyclopoida" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="ballaballaensis" status="sp. nov.">
<emphasis id="B935A85B9C09FFD5FE89902EFE306112" box="[277,464,1096,1120]" italics="true" pageId="3" pageNumber="104">D. ballaballaensis</emphasis>
</taxonomicName>
<emphasis id="B935A85B9C09FFD5FE4B902EFDC86112" bold="true" box="[471,552,1096,1120]" pageId="3" pageNumber="104">
<taxonomicNameLabel id="A20615209C09FFD5FE4B902EFDC86112" box="[471,552,1096,1120]" pageId="3" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; D,
<taxonomicName id="4C410FCA9C09FFD5FDCA902EFD316112" authorityName="Karanovic" authorityYear="2024" box="[598,721,1096,1120]" class="Copepoda" family="Cyclopidae" genus="Diacyclops" kingdom="Animalia" order="Cyclopoida" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="hancocki" status="sp. nov.">
<emphasis id="B935A85B9C09FFD5FDCA902EFD316112" box="[598,721,1096,1120]" italics="true" pageId="3" pageNumber="104">D. hancocki</emphasis>
</taxonomicName>
<emphasis id="B935A85B9C09FFD5FD45902EFCCA6112" bold="true" box="[729,810,1096,1120]" pageId="3" pageNumber="104">
<taxonomicNameLabel id="A20615209C09FFD5FD45902EFCCA6112" box="[729,810,1096,1120]" pageId="3" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; E,
<taxonomicName id="4C410FCA9C09FFD5FCC9902EFC116112" authorityName="Karanovic" authorityYear="2024" box="[853,1009,1096,1120]" class="Copepoda" family="Cyclopidae" genus="Diacyclops" kingdom="Animalia" order="Cyclopoida" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="tomlinsonae" status="sp. nov.">
<emphasis id="B935A85B9C09FFD5FCC9902EFC116112" box="[853,1009,1096,1120]" italics="true" pageId="3" pageNumber="104">D. tomlinsonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20615209C09FFD5FC64902EFBAB612D" box="[1016,1099,1096,1120]" pageId="3" pageNumber="104" rank="species">
<emphasis id="B935A85B9C09FFD5FC64902EFBA46112" bold="true" box="[1016,1092,1096,1120]" pageId="3" pageNumber="104">sp. nov</emphasis>
.
</taxonomicNameLabel>
; F,
<taxonomicName id="4C410FCA9C09FFD5FBEE902EFAA36112" authorityName="Karanovic" authorityYear="2024" box="[1138,1347,1096,1120]" class="Copepoda" family="Cyclopidae" genus="Diacyclops" kingdom="Animalia" order="Cyclopoida" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="huntervalleyensis" status="sp. nov.">
<emphasis id="B935A85B9C09FFD5FBEE902EFAA36112" box="[1138,1347,1096,1120]" italics="true" pageId="3" pageNumber="104">D. huntervalleyensis</emphasis>
</taxonomicName>
<emphasis id="B935A85B9C09FFD5FAD7902EFA7C6112" bold="true" box="[1355,1436,1096,1120]" pageId="3" pageNumber="104">
<taxonomicNameLabel id="A20615209C09FFD5FAD7902EFA7C6112" box="[1355,1436,1096,1120]" pageId="3" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Scale bar 100 µm.
</paragraph>
</caption>
<subSubSection id="C35B27C29C09FFD5FF5B90B0FB2C6066" pageId="3" pageNumber="104" type="etymology">
<paragraph id="8BFE74499C09FFD5FF5B90B0FB2C6066" blockId="3.[151,1437,1238,2020]" pageId="3" pageNumber="104">
<emphasis id="B935A85B9C09FFD5FF5B90B0FEAE6182" bold="true" box="[199,334,1238,1264]" pageId="3" pageNumber="104">Etymology.</emphasis>
The species is named after Dr. Remko Leijs, South Australian Museum, Adelaide, who collected this species and entrusted it to me for identification. The name is a noun in the genitive singular.
</paragraph>
</subSubSection>
<subSubSection id="C35B27C29C09FFDCFF5B9178FB3F674F" lastPageId="10" lastPageNumber="111" pageId="3" pageNumber="104" type="diagnosis">
<paragraph id="8BFE74499C09FFDFFF5B9178FD8A6487" blockId="3.[151,1437,1238,2020]" lastBlockId="9.[151,1437,151,501]" lastPageId="9" lastPageNumber="110" pageId="3" pageNumber="104">
<emphasis id="B935A85B9C09FFD5FF5B9178FEA2604A" bold="true" box="[199,322,1310,1336]" pageId="3" pageNumber="104">Diagnosis.</emphasis>
Female. Body length, excluding caudal setae, from 440 to 485 µm. Habitus (
<figureCitation id="137A68CC9C09FFD5FB3C9178FAE4604A" box="[1184,1284,1310,1336]" captionStart="FIGURE 1" captionStartId="3.[151,250,1059,1083]" captionTargetBox="[155,1436,181,1036]" captionTargetId="figure-15@3.[151,1436,181,1036]" captionTargetPageId="3" captionText="FIGURE 1. Habitus in dorsal view of holotype females of all six new species: A, Diacyclops leijsi sp. nov.; B, D. eberhardi sp. nov.; C, D. ballaballaensis sp. nov.; D, D. hancocki sp. nov.; E, D. tomlinsonae sp. nov.; F, D. huntervalleyensis sp. nov. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14240872" httpUri="https://zenodo.org/record/14240872/files/figure.png" pageId="3" pageNumber="104">Figs. 1A</figureCitation>
,
<figureCitation id="137A68CC9C09FFD5FA8C9178FAD3604A" box="[1296,1331,1310,1336]" captionStart="FIGURE 6" captionStartId="8.[151,250,1544,1568]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure-27@8.[151,1436,181,1520]" captionTargetPageId="8" captionText="FIGURE 6. Diacyclops leijsi sp. nov., scanning electron micrographs, AD, paratype female 1, ventral; E, paratype female 2, lateral; F, paratype male 2, lateral: A, habitus; B, caudal rami; C, mouth appendages; D, fourth swimming leg; E, distal part of antennula; F, habitus." figureDoi="http://doi.org/10.5281/zenodo.14240882" httpUri="https://zenodo.org/record/14240882/files/figure.png" pageId="3" pageNumber="104">6A</figureCitation>
) spindle-shaped, nearly 2.8 times as long as wide, with prosome/urosome ratio of 1.25, and posterior end of cephalothorax about 2.5 times as wide as genital double-somite in dorsal view. Integument on all somites thin and wrinkled, with dense bacterial cover, spinules only on anal somite and caudal rami, cuticular pores on all somites, and sensilla on all but penultimate somite. Hyaline fringes of prosomites smooth, while those of genital double-somite and 2 subsequent urosomites coarsely serrated (
<figureCitation id="137A68CC9C09FFD5FDFA91B4FD29609E" box="[614,713,1490,1516]" captionStart="FIGURE 1" captionStartId="3.[151,250,1059,1083]" captionTargetBox="[155,1436,181,1036]" captionTargetId="figure-15@3.[151,1436,181,1036]" captionTargetPageId="3" captionText="FIGURE 1. Habitus in dorsal view of holotype females of all six new species: A, Diacyclops leijsi sp. nov.; B, D. eberhardi sp. nov.; C, D. ballaballaensis sp. nov.; D, D. hancocki sp. nov.; E, D. tomlinsonae sp. nov.; F, D. huntervalleyensis sp. nov. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14240872" httpUri="https://zenodo.org/record/14240872/files/figure.png" pageId="3" pageNumber="104">Figs. 1A</figureCitation>
,
<figureCitation id="137A68CC9C09FFD5FD4891B4FD18609E" box="[724,760,1490,1516]" captionStart="FIGURE 6" captionStartId="8.[151,250,1544,1568]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure-27@8.[151,1436,181,1520]" captionTargetPageId="8" captionText="FIGURE 6. Diacyclops leijsi sp. nov., scanning electron micrographs, AD, paratype female 1, ventral; E, paratype female 2, lateral; F, paratype male 2, lateral: A, habitus; B, caudal rami; C, mouth appendages; D, fourth swimming leg; E, distal part of antennula; F, habitus." figureDoi="http://doi.org/10.5281/zenodo.14240882" httpUri="https://zenodo.org/record/14240882/files/figure.png" pageId="3" pageNumber="104">6B</figureCitation>
). Genital double-somite (
<figureCitation id="137A68CC9C09FFD5FB8E91B4FB8A609E" box="[1042,1130,1490,1516]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[174,1414,181,1914]" captionTargetId="figure-27@4.[174,1414,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Diacyclops leijsi sp. nov., line drawings, holotype female: A, urosome, ventral view; B, antennula. Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.14240874" httpUri="https://zenodo.org/record/14240874/files/figure.png" pageId="3" pageNumber="104">Fig. 2A</figureCitation>
) slightly wider than long in ventral view, widest at first fifth of its length and gradually tapering posteriorly, widest part about 1.5 times as wide as posterior margin; seminal receptacle large and shaped like a mature mushroom; copulatory pore relatively large; copulatory duct short, narrow, and well-sclerotized. Anal somite (
<figureCitation id="137A68CC9C09FFD5FCF19258FC24632A" box="[877,964,1598,1624]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[174,1414,181,1914]" captionTargetId="figure-27@4.[174,1414,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Diacyclops leijsi sp. nov., line drawings, holotype female: A, urosome, ventral view; B, antennula. Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.14240874" httpUri="https://zenodo.org/record/14240874/files/figure.png" pageId="3" pageNumber="104">Fig. 2A</figureCitation>
) with small spinules along ventral margin. Caudal rami (
<figureCitation id="137A68CC9C09FFD5FEAC9204FE69630E" box="[304,393,1634,1660]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[174,1414,181,1914]" captionTargetId="figure-27@4.[174,1414,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Diacyclops leijsi sp. nov., line drawings, holotype female: A, urosome, ventral view; B, antennula. Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.14240874" httpUri="https://zenodo.org/record/14240874/files/figure.png" pageId="3" pageNumber="104">Fig. 2A</figureCitation>
) slender, narrowly spaced, nearly 5 times as long as wide and 3 times as long as anal somite; principal terminal setae without breaking planes, inner one about 1.2 times as long as entire urosome and 1.6 times as long as outer one; dorsal seta about as long as caudal ramus, 1.5 times as long as innermost terminal seta, and twice as long as outermost terminal seta. Antennula (
<figureCitation id="137A68CC9C09FFD5FD0192A8FD1E639A" box="[669,766,1742,1768]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[174,1414,181,1914]" captionTargetId="figure-27@4.[174,1414,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Diacyclops leijsi sp. nov., line drawings, holotype female: A, urosome, ventral view; B, antennula. Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.14240874" httpUri="https://zenodo.org/record/14240874/files/figure.png" pageId="3" pageNumber="104">Figs. 2B</figureCitation>
,
<figureCitation id="137A68CC9C09FFD5FC9592A8FCC9639A" box="[777,809,1742,1768]" captionStart="FIGURE 6" captionStartId="8.[151,250,1544,1568]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure-27@8.[151,1436,181,1520]" captionTargetPageId="8" captionText="FIGURE 6. Diacyclops leijsi sp. nov., scanning electron micrographs, AD, paratype female 1, ventral; E, paratype female 2, lateral; F, paratype male 2, lateral: A, habitus; B, caudal rami; C, mouth appendages; D, fourth swimming leg; E, distal part of antennula; F, habitus." figureDoi="http://doi.org/10.5281/zenodo.14240882" httpUri="https://zenodo.org/record/14240882/files/figure.png" pageId="3" pageNumber="104">6E</figureCitation>
) 17-segmented, slightly longer than cephalothorax, with single aesthetasc on twelfth segment and setae formula 8.4.2.6.3.2.2.1.1.0.1.1.0.1.2.2.8; ultimate segment about 2.4 times as long as wide. Antenna (
<figureCitation id="137A68CC9C09FFD5FD9C9370FDB86242" box="[512,600,1814,1840]" captionStart="FIGURE 3" captionStartId="5.[151,250,1621,1645]" captionTargetBox="[151,1435,181,1594]" captionTargetId="figure-15@5.[151,1435,181,1597]" captionTargetPageId="5" captionText="FIGURE 3. Diacyclops leijsi sp. nov., line drawings, holotype female:A, antenna; B, labrum; C, mandibula; D, cutting edge of mandibula; E. maxillula; F, praecoxal arthrite of maxillula; G, maxilla; H, maxilliped. Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.14240876" httpUri="https://zenodo.org/record/14240876/files/figure.png" pageId="3" pageNumber="104">Fig. 3A</figureCitation>
) 5-segmented, without exopodal seta, with setae formula 0.2.1.8.7; second and fifth segment of similar length. Labrum (
<figureCitation id="137A68CC9C09FFD5FD13935CFD046226" box="[655,740,1850,1876]" captionStart="FIGURE 3" captionStartId="5.[151,250,1621,1645]" captionTargetBox="[151,1435,181,1594]" captionTargetId="figure-15@5.[151,1435,181,1597]" captionTargetPageId="5" captionText="FIGURE 3. Diacyclops leijsi sp. nov., line drawings, holotype female:A, antenna; B, labrum; C, mandibula; D, cutting edge of mandibula; E. maxillula; F, praecoxal arthrite of maxillula; G, maxilla; H, maxilliped. Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.14240876" httpUri="https://zenodo.org/record/14240876/files/figure.png" pageId="3" pageNumber="104">Fig. 3B</figureCitation>
) trapezoidal, with 2 diagonal rows of 10 slender spinules each on anterior surface; cutting edge nearly straight, with 14 sharp teeth between pointed lateral corners. Mandibula (
<figureCitation id="137A68CC9C09FFD5FF0393E4FEFC62EE" box="[159,284,1922,1948]" captionStart="FIGURE 3" captionStartId="5.[151,250,1621,1645]" captionTargetBox="[151,1435,181,1594]" captionTargetId="figure-15@5.[151,1435,181,1597]" captionTargetPageId="5" captionText="FIGURE 3. Diacyclops leijsi sp. nov., line drawings, holotype female:A, antenna; B, labrum; C, mandibula; D, cutting edge of mandibula; E. maxillula; F, praecoxal arthrite of maxillula; G, maxilla; H, maxilliped. Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.14240876" httpUri="https://zenodo.org/record/14240876/files/figure.png" pageId="3" pageNumber="104">Fig. 3C, D</figureCitation>
) slender, with 2 long and 1 short seta on 1-segmented palp; cutting edge with 4 slender spinules on anterior surface, 8 apical teeth, and dorsalmost unipinnate seta; ventralmost tooth strongest and quadricuspidate, second, fourth, and seventh teeth from ventral side bicuspidate, all other teeth unicuspidate. Maxillula (
<figureCitation id="137A68CC9C09FFD5FA9B93ACFA876296" box="[1287,1383,1994,2020]" captionStart="FIGURE 3" captionStartId="5.[151,250,1621,1645]" captionTargetBox="[151,1435,181,1594]" captionTargetId="figure-15@5.[151,1435,181,1597]" captionTargetPageId="5" captionText="FIGURE 3. Diacyclops leijsi sp. nov., line drawings, holotype female:A, antenna; B, labrum; C, mandibula; D, cutting edge of mandibula; E. maxillula; F, praecoxal arthrite of maxillula; G, maxilla; H, maxilliped. Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.14240876" httpUri="https://zenodo.org/record/14240876/files/figure.png" pageId="3" pageNumber="104">Figs. 3E</figureCitation>
,
<figureCitation id="137A68CC9C09FFD5FAEE93ACFA736296" box="[1394,1427,1994,2020]" captionStart="FIGURE 6" captionStartId="8.[151,250,1544,1568]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure-27@8.[151,1436,181,1520]" captionTargetPageId="8" captionText="FIGURE 6. Diacyclops leijsi sp. nov., scanning electron micrographs, AD, paratype female 1, ventral; E, paratype female 2, lateral; F, paratype male 2, lateral: A, habitus; B, caudal rami; C, mouth appendages; D, fourth swimming leg; E, distal part of antennula; F, habitus." figureDoi="http://doi.org/10.5281/zenodo.14240882" httpUri="https://zenodo.org/record/14240882/files/figure.png" pageId="3" pageNumber="104">6C</figureCitation>
) composed of praecoxa and 2-segmented palp, unornamented; praecoxal arthrite bearing 4 strong distal spines and 6 medial elements; palp with slender exopodal seta, 3 slender setae on endopod, and 1 strong and 2 slender setae on inner margin of coxobasis. Maxilla (
<figureCitation id="137A68CC9C02FFDEFDCC9364FD53626E" box="[592,691,1794,1820]" captionStart="FIGURE 3" captionStartId="5.[151,250,1621,1645]" captionTargetBox="[151,1435,181,1594]" captionTargetId="figure-15@5.[151,1435,181,1597]" captionTargetPageId="5" captionText="FIGURE 3. Diacyclops leijsi sp. nov., line drawings, holotype female:A, antenna; B, labrum; C, mandibula; D, cutting edge of mandibula; E. maxillula; F, praecoxal arthrite of maxillula; G, maxilla; H, maxilliped. Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.14240876" httpUri="https://zenodo.org/record/14240876/files/figure.png" pageId="8" pageNumber="109">Figs. 3G</figureCitation>
,
<figureCitation id="137A68CC9C02FFDEFD229364FD01626E" box="[702,737,1794,1820]" captionStart="FIGURE 6" captionStartId="8.[151,250,1544,1568]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure-27@8.[151,1436,181,1520]" captionTargetPageId="8" captionText="FIGURE 6. Diacyclops leijsi sp. nov., scanning electron micrographs, AD, paratype female 1, ventral; E, paratype female 2, lateral; F, paratype male 2, lateral: A, habitus; B, caudal rami; C, mouth appendages; D, fourth swimming leg; E, distal part of antennula; F, habitus." figureDoi="http://doi.org/10.5281/zenodo.14240882" httpUri="https://zenodo.org/record/14240882/files/figure.png" pageId="8" pageNumber="109">6C</figureCitation>
) 4-segmented, with 2 slender setae on praecoxa, 1 slender seta on proximal endite of coxa, 1 strong and 1 slender seta on highly mobile distal endite of coxa, basis expanded into robust claw with 1 strong and 1 very slender seta, and endopod with 3 strong and 2 slender setae. Maxilliped (
<figureCitation id="137A68CC9C02FFDEFAF8932CFF5B62FA" captionStart="FIGURE 3" captionStartId="5.[151,250,1621,1645]" captionTargetBox="[151,1435,181,1594]" captionTargetId="figure-15@5.[151,1435,181,1597]" captionTargetPageId="5" captionText="FIGURE 3. Diacyclops leijsi sp. nov., line drawings, holotype female:A, antenna; B, labrum; C, mandibula; D, cutting edge of mandibula; E. maxillula; F, praecoxal arthrite of maxillula; G, maxilla; H, maxilliped. Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.14240876" httpUri="https://zenodo.org/record/14240876/files/figure.png" pageId="8" pageNumber="109">Figs. 3H</figureCitation>
,
<figureCitation id="137A68CC9C02FFDEFF569308FF0C62FA" box="[202,236,1902,1928]" captionStart="FIGURE 6" captionStartId="8.[151,250,1544,1568]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure-27@8.[151,1436,181,1520]" captionTargetPageId="8" captionText="FIGURE 6. Diacyclops leijsi sp. nov., scanning electron micrographs, AD, paratype female 1, ventral; E, paratype female 2, lateral; F, paratype male 2, lateral: A, habitus; B, caudal rami; C, mouth appendages; D, fourth swimming leg; E, distal part of antennula; F, habitus." figureDoi="http://doi.org/10.5281/zenodo.14240882" httpUri="https://zenodo.org/record/14240882/files/figure.png" pageId="8" pageNumber="109">6C</figureCitation>
) 4-segmented, with long spinules on second and third segment and setae formula 3.2.1.3. All swimming legs (
<figureCitation id="137A68CC9C02FFDEFF4893F4FE5A62DF" box="[212,442,1938,1965]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[171,1411,181,1918]" captionTargetId="figure-27@6.[171,1415,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Diacyclops leijsi sp. nov., line drawings, holotype female: A, first swimming leg; B, second swimming leg; C, endopod of third swimming leg; D, third exopodal segment of third swimming leg; E, fourth swimming leg; F, fifth leg; G, sixth leg. Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.14240878" httpUri="https://zenodo.org/record/14240878/files/figure.png" pageId="8" pageNumber="109">Figs. 4A, B, C, D, E</figureCitation>
,
<figureCitation id="137A68CC9C02FFDEFE5A93F4FE0962DE" box="[454,489,1938,1964]" captionStart="FIGURE 6" captionStartId="8.[151,250,1544,1568]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure-27@8.[151,1436,181,1520]" captionTargetPageId="8" captionText="FIGURE 6. Diacyclops leijsi sp. nov., scanning electron micrographs, AD, paratype female 1, ventral; E, paratype female 2, lateral; F, paratype male 2, lateral: A, habitus; B, caudal rami; C, mouth appendages; D, fourth swimming leg; E, distal part of antennula; F, habitus." figureDoi="http://doi.org/10.5281/zenodo.14240882" httpUri="https://zenodo.org/record/14240882/files/figure.png" pageId="8" pageNumber="109">6D</figureCitation>
) large and wide, with smooth intercoxal sclerite with slightly concave distal margin, each leg composed of minute and triangular praecoxa, large and rectangular coxa, short basis, 3-segmented exopod and 3-segmented endopod; basis of first leg with long outer seta and long inner spine, latter slightly longer that first 2 endopodal segments combined; basis of second to fourth legs with shorter outer seta and inner distal corner pointed; all first and second exopodal segments, all first endopodal segments, and second endopodal segment of first and second legs with single inner seta; second endopodal segment of third and fourth legs with 2 inner setae; third exopodal segments spine formula 2.3.3.3 and setae formula 4.4.4.4; third endopodal segments of first to third leg with 3 inner setae, 1 apical seta, 1 apical spine, and 1 outer seta; third endopodal segment of fourth leg nearly twice as long as wide, with 2 inner setae, 2 apical spines, and 1 outer seta; its outer spine nearly 1.3 times as long as segment or inner spine. Fifth leg (
<figureCitation id="137A68CC9C03FFDFFDA69509FD7F64FB" box="[570,671,367,393]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[174,1414,181,1914]" captionTargetId="figure-27@4.[174,1414,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Diacyclops leijsi sp. nov., line drawings, holotype female: A, urosome, ventral view; B, antennula. Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.14240874" httpUri="https://zenodo.org/record/14240874/files/figure.png" pageId="9" pageNumber="110">Figs. 2A</figureCitation>
,
<figureCitation id="137A68CC9C03FFDFFD319509FD2C64FB" box="[685,716,367,393]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[171,1411,181,1918]" captionTargetId="figure-27@6.[171,1415,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Diacyclops leijsi sp. nov., line drawings, holotype female: A, first swimming leg; B, second swimming leg; C, endopod of third swimming leg; D, third exopodal segment of third swimming leg; E, fourth swimming leg; F, fifth leg; G, sixth leg. Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.14240878" httpUri="https://zenodo.org/record/14240878/files/figure.png" pageId="9" pageNumber="110">4F</figureCitation>
) small and slender, 2-segmented, with single outer seta on first segment and apical seta and subapical inner spine on second segment; second segment about 2.2 times as long as wide and only slightly longer than its spine. Sixth leg (
<figureCitation id="137A68CC9C03FFDFFD6D95D1FCA764A3" box="[753,839,439,465]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[171,1411,181,1918]" captionTargetId="figure-27@6.[171,1415,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Diacyclops leijsi sp. nov., line drawings, holotype female: A, first swimming leg; B, second swimming leg; C, endopod of third swimming leg; D, third exopodal segment of third swimming leg; E, fourth swimming leg; F, fifth leg; G, sixth leg. Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.14240878" httpUri="https://zenodo.org/record/14240878/files/figure.png" pageId="9" pageNumber="110">Fig. 4G</figureCitation>
) small semicircular plate, with long seta and 2 minute spines; seta almost as long as plate width.
</paragraph>
<caption id="DF3E24C19C0EFFD2FF0B93F1FF2062A6" ID-DOI="http://doi.org/10.5281/zenodo.14240874" ID-Zenodo-Dep="14240874" httpUri="https://zenodo.org/record/14240874/files/figure.png" pageId="4" pageNumber="105" startId="4.[151,250,1943,1967]" targetBox="[174,1414,181,1914]" targetPageId="4" targetType="figure">
<paragraph id="8BFE74499C0EFFD2FF0B93F1FF2062A6" blockId="4.[151,1436,1943,2004]" pageId="4" pageNumber="105">
<emphasis id="B935A85B9C0EFFD2FF0B93F1FEF462DD" bold="true" box="[151,276,1943,1967]" pageId="4" pageNumber="105">FIGURE 2.</emphasis>
<taxonomicName id="4C410FCA9C0EFFD2FE8093FEFE2762C2" authorityName="Karanovic" authorityYear="2024" box="[284,455,1944,1968]" class="Copepoda" family="Cyclopidae" genus="Diacyclops" kingdom="Animalia" order="Cyclopoida" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="leijsi" status="sp. nov.">
<emphasis id="B935A85B9C0EFFD2FE8093FEFE2762C2" box="[284,455,1944,1968]" italics="true" pageId="4" pageNumber="105">Diacyclops leijsi</emphasis>
</taxonomicName>
<emphasis id="B935A85B9C0EFFD2FE5293FEFDC062C2" bold="true" box="[462,544,1944,1968]" pageId="4" pageNumber="105">
<taxonomicNameLabel id="A20615209C0EFFD2FE5293FEFDC062C2" box="[462,544,1944,1968]" pageId="4" pageNumber="105" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, line drawings, holotype female: A, urosome, ventral view; B, antennula. Scale bar 50 μm.
</paragraph>
</caption>
<caption id="DF3E24C19C0FFFD3FF0B9233FB4E63E0" ID-DOI="http://doi.org/10.5281/zenodo.14240876" ID-Zenodo-Dep="14240876" httpUri="https://zenodo.org/record/14240876/files/figure.png" pageId="5" pageNumber="106" startId="5.[151,250,1621,1645]" targetBox="[151,1435,181,1594]" targetPageId="5" targetType="figure">
<paragraph id="8BFE74499C0FFFD3FF0B9233FB4E63E0" blockId="5.[151,1436,1621,1682]" pageId="5" pageNumber="106">
<emphasis id="B935A85B9C0FFFD3FF0B9233FEF2631F" bold="true" box="[151,274,1621,1645]" pageId="5" pageNumber="106">FIGURE 3.</emphasis>
<taxonomicName id="4C410FCA9C0FFFD3FE849233FE21631F" authorityName="Karanovic" authorityYear="2024" box="[280,449,1621,1645]" class="Copepoda" family="Cyclopidae" genus="Diacyclops" kingdom="Animalia" order="Cyclopoida" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="leijsi" status="sp. nov.">
<emphasis id="B935A85B9C0FFFD3FE849233FE21631F" box="[280,449,1621,1645]" italics="true" pageId="5" pageNumber="106">Diacyclops leijsi</emphasis>
</taxonomicName>
<emphasis id="B935A85B9C0FFFD3FE5B9233FDF7631F" bold="true" box="[455,535,1621,1645]" pageId="5" pageNumber="106">
<taxonomicNameLabel id="A20615209C0FFFD3FE5B9233FDF7631F" box="[455,535,1621,1645]" pageId="5" pageNumber="106" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, line drawings, holotype female: A, antenna; B, labrum; C, mandibula; D, cutting edge of mandibula; E. maxillula; F, praecoxal arthrite of maxillula; G, maxilla; H, maxilliped. Scale bar 50 μm.
</paragraph>
</caption>
<caption id="DF3E24C19C0CFFD0FF0B93F1FE8F628A" ID-DOI="http://doi.org/10.5281/zenodo.14240878" ID-Zenodo-Dep="14240878" httpUri="https://zenodo.org/record/14240878/files/figure.png" pageId="6" pageNumber="107" startId="6.[151,250,1943,1967]" targetBox="[171,1411,181,1918]" targetPageId="6" targetType="figure">
<paragraph id="8BFE74499C0CFFD0FF0B93F1FE8F628A" blockId="6.[151,1436,1943,2040]" pageId="6" pageNumber="107">
<emphasis id="B935A85B9C0CFFD0FF0B93F1FEF562DD" bold="true" box="[151,277,1943,1967]" pageId="6" pageNumber="107">FIGURE 4.</emphasis>
<taxonomicName id="4C410FCA9C0CFFD0FE8193FEFE2962C2" authorityName="Karanovic" authorityYear="2024" box="[285,457,1944,1968]" class="Copepoda" family="Cyclopidae" genus="Diacyclops" kingdom="Animalia" order="Cyclopoida" pageId="6" pageNumber="107" phylum="Arthropoda" rank="species" species="leijsi" status="sp. nov.">
<emphasis id="B935A85B9C0CFFD0FE8193FEFE2962C2" box="[285,457,1944,1968]" italics="true" pageId="6" pageNumber="107">Diacyclops leijsi</emphasis>
</taxonomicName>
<emphasis id="B935A85B9C0CFFD0FE4D93FEFDC462C2" bold="true" box="[465,548,1944,1968]" pageId="6" pageNumber="107">
<taxonomicNameLabel id="A20615209C0CFFD0FE4D93FEFDC462C2" box="[465,548,1944,1968]" pageId="6" pageNumber="107" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, line drawings, holotype female: A, first swimming leg; B, second swimming leg; C, endopod of third swimming leg; D, third exopodal segment of third swimming leg; E, fourth swimming leg; F, fifth leg; G, sixth leg. Scale bar 50 μm.
</paragraph>
</caption>
<caption id="DF3E24C19C0DFFD1FF0B9342FCFD6213" ID-DOI="http://doi.org/10.5281/zenodo.14240880" ID-Zenodo-Dep="14240880" httpUri="https://zenodo.org/record/14240880/files/figure.png" pageId="7" pageNumber="108" startId="7.[151,250,1828,1852]" targetBox="[159,1436,181,1797]" targetPageId="7" targetType="figure">
<paragraph id="8BFE74499C0DFFD1FF0B9342FCFD6213" blockId="7.[151,1437,1828,1889]" pageId="7" pageNumber="108">
<emphasis id="B935A85B9C0DFFD1FF0B9342FEF4624E" bold="true" box="[151,276,1828,1852]" pageId="7" pageNumber="108">FIGURE 5.</emphasis>
<taxonomicName id="4C410FCA9C0DFFD1FE879342FE25624E" authorityName="Karanovic" authorityYear="2024" box="[283,453,1828,1852]" class="Copepoda" family="Cyclopidae" genus="Diacyclops" kingdom="Animalia" order="Cyclopoida" pageId="7" pageNumber="108" phylum="Arthropoda" rank="species" species="leijsi" status="sp. nov.">
<emphasis id="B935A85B9C0DFFD1FE879342FE25624E" box="[283,453,1828,1852]" italics="true" pageId="7" pageNumber="108">Diacyclops leijsi</emphasis>
</taxonomicName>
<emphasis id="B935A85B9C0DFFD1FE519342FDFE624E" bold="true" box="[461,542,1828,1852]" pageId="7" pageNumber="108">
<taxonomicNameLabel id="A20615209C0DFFD1FE519342FDFE624E" box="[461,542,1828,1852]" pageId="7" pageNumber="108" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, line drawings, paratype male 1: A, urosome, ventral; B, antennula; C, third endopodal segment of fourth swimming leg; D, sixth leg. Scale bar 50 μm.
</paragraph>
</caption>
<caption id="DF3E24C19C02FFDEFF0B926EFE90631A" ID-DOI="http://doi.org/10.5281/zenodo.14240882" ID-Zenodo-Dep="14240882" httpUri="https://zenodo.org/record/14240882/files/figure.png" pageId="8" pageNumber="109" startId="8.[151,250,1544,1568]" targetBox="[151,1436,181,1520]" targetPageId="8" targetType="figure">
<paragraph id="8BFE74499C02FFDEFF0B926EFE90631A" blockId="8.[151,1437,1544,1640]" pageId="8" pageNumber="109">
<emphasis id="B935A85B9C02FFDEFF0B926EFEF36352" bold="true" box="[151,275,1544,1568]" pageId="8" pageNumber="109">FIGURE 6.</emphasis>
<taxonomicName id="4C410FCA9C02FFDEFE86926EFE246352" authorityName="Karanovic" authorityYear="2024" box="[282,452,1544,1568]" class="Copepoda" family="Cyclopidae" genus="Diacyclops" kingdom="Animalia" order="Cyclopoida" pageId="8" pageNumber="109" phylum="Arthropoda" rank="species" species="leijsi" status="sp. nov.">
<emphasis id="B935A85B9C02FFDEFE86926EFE246352" box="[282,452,1544,1568]" italics="true" pageId="8" pageNumber="109">Diacyclops leijsi</emphasis>
</taxonomicName>
<emphasis id="B935A85B9C02FFDEFE57926EFDFC6352" bold="true" box="[459,540,1544,1568]" pageId="8" pageNumber="109">
<taxonomicNameLabel id="A20615209C02FFDEFE57926EFDFC6352" box="[459,540,1544,1568]" pageId="8" pageNumber="109" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, scanning electron micrographs, AD, paratype female 1, ventral; E, paratype female 2, lateral; F, paratype male 2, lateral: A, habitus; B, caudal rami; C, mouth appendages; D, fourth swimming leg; E, distal part of antennula; F, habitus.
</paragraph>
</caption>
<caption id="DF3E24C19C03FFDFFF0B9339FE9C62CD" ID-DOI="http://doi.org/10.5281/zenodo.14240884" ID-Zenodo-Dep="14240884" httpUri="https://zenodo.org/record/14240884/files/figure.png" pageId="9" pageNumber="110" startId="9.[151,250,1887,1911]" targetBox="[151,1436,525,1864]" targetPageId="9" targetType="figure">
<paragraph id="8BFE74499C03FFDFFF0B9339FE9C62CD" blockId="9.[151,1436,1887,1983]" pageId="9" pageNumber="110">
<emphasis id="B935A85B9C03FFDFFF0B9339FEF46205" bold="true" box="[151,276,1887,1911]" pageId="9" pageNumber="110">FIGURE 7.</emphasis>
<taxonomicName id="4C410FCA9C03FFDFFE879339FE246205" authorityName="Karanovic" authorityYear="2024" box="[283,452,1887,1911]" class="Copepoda" family="Cyclopidae" genus="Diacyclops" kingdom="Animalia" order="Cyclopoida" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="leijsi" status="sp. nov.">
<emphasis id="B935A85B9C03FFDFFE879339FE246205" box="[283,452,1887,1911]" italics="true" pageId="9" pageNumber="110">Diacyclops leijsi</emphasis>
</taxonomicName>
<emphasis id="B935A85B9C03FFDFFE509339FDFD6205" bold="true" box="[460,541,1887,1911]" pageId="9" pageNumber="110">
<taxonomicNameLabel id="A20615209C03FFDFFE509339FDFD6205" box="[460,541,1887,1911]" pageId="9" pageNumber="110" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, scanning electron micrographs, A &amp; B, paratype male 2, lateral; CF, paratype male 3, dorsal: A, left antennula; B, detail of right antennula; C, free prosomites; D, first two urosomites; E, third and fourth urosomites; F, last two urosomites.
</paragraph>
</caption>
<paragraph id="8BFE74499C00FFDCFF5B94F1FB3F674F" blockId="10.[151,1437,150,573]" pageId="10" pageNumber="111">
Male. Body length from 435 to 465 µm. Habitus (
<figureCitation id="137A68CC9C00FFDCFC9694F1FC8065C3" box="[778,864,151,177]" captionStart="FIGURE 6" captionStartId="8.[151,250,1544,1568]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure-27@8.[151,1436,181,1520]" captionTargetPageId="8" captionText="FIGURE 6. Diacyclops leijsi sp. nov., scanning electron micrographs, AD, paratype female 1, ventral; E, paratype female 2, lateral; F, paratype male 2, lateral: A, habitus; B, caudal rami; C, mouth appendages; D, fourth swimming leg; E, distal part of antennula; F, habitus." figureDoi="http://doi.org/10.5281/zenodo.14240882" httpUri="https://zenodo.org/record/14240882/files/figure.png" pageId="10" pageNumber="111">Fig. 6F</figureCitation>
) and urosome (
<figureCitation id="137A68CC9C00FFDCFB8994F1FB8E65C3" box="[1045,1134,150,177]" captionStart="FIGURE 5" captionStartId="7.[151,250,1828,1852]" captionTargetBox="[159,1436,181,1797]" captionTargetId="figure-15@7.[151,1436,181,1804]" captionTargetPageId="7" captionText="FIGURE 5. Diacyclops leijsi sp. nov., line drawings, paratype male 1: A, urosome, ventral; B, antennula; C, third endopodal segment of fourth swimming leg; D, sixth leg. Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.14240880" httpUri="https://zenodo.org/record/14240880/files/figure.png" pageId="10" pageNumber="111">Fig. 5A</figureCitation>
) slightly slenderer than in female; free genital somite (
<figureCitation id="137A68CC9C00FFDCFE5794DDFDCD65A7" box="[459,557,186,213]" captionStart="FIGURE 5" captionStartId="7.[151,250,1828,1852]" captionTargetBox="[159,1436,181,1797]" captionTargetId="figure-15@7.[151,1436,181,1804]" captionTargetPageId="7" captionText="FIGURE 5. Diacyclops leijsi sp. nov., line drawings, paratype male 1: A, urosome, ventral; B, antennula; C, third endopodal segment of fourth swimming leg; D, sixth leg. Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.14240880" httpUri="https://zenodo.org/record/14240880/files/figure.png" pageId="10" pageNumber="111">Figs. 5A</figureCitation>
,
<figureCitation id="137A68CC9C00FFDCFDAB94DDFDBB65A7" box="[567,603,187,213]" captionStart="FIGURE 7" captionStartId="9.[151,250,1887,1911]" captionTargetBox="[151,1436,525,1864]" captionTargetId="figure-201@9.[151,1436,525,1864]" captionTargetPageId="9" captionText="FIGURE 7. Diacyclops leijsi sp. nov., scanning electron micrographs, A &amp; B, paratype male 2, lateral; CF, paratype male 3, dorsal:A, left antennula; B, detail of right antennula; C, free prosomites; D, first two urosomites; E, third and fourth urosomites; F, last two urosomites." figureDoi="http://doi.org/10.5281/zenodo.14240884" httpUri="https://zenodo.org/record/14240884/files/figure.png" pageId="10" pageNumber="111">7D</figureCitation>
) more than 1.5 times as wide as subsequent urosomite, with relatively small ovoid spermatophores. Ornamentation of cephalothorax (
<figureCitation id="137A68CC9C00FFDCFCBC94B9FC96658B" box="[800,886,223,249]" captionStart="FIGURE 6" captionStartId="8.[151,250,1544,1568]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure-27@8.[151,1436,181,1520]" captionTargetPageId="8" captionText="FIGURE 6. Diacyclops leijsi sp. nov., scanning electron micrographs, AD, paratype female 1, ventral; E, paratype female 2, lateral; F, paratype male 2, lateral: A, habitus; B, caudal rami; C, mouth appendages; D, fourth swimming leg; E, distal part of antennula; F, habitus." figureDoi="http://doi.org/10.5281/zenodo.14240882" httpUri="https://zenodo.org/record/14240882/files/figure.png" pageId="10" pageNumber="111">Fig. 6F</figureCitation>
), free prosomites (
<figureCitation id="137A68CC9C00FFDCFBD794B9FB4B658B" box="[1099,1195,223,249]" captionStart="FIGURE 6" captionStartId="8.[151,250,1544,1568]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure-27@8.[151,1436,181,1520]" captionTargetPageId="8" captionText="FIGURE 6. Diacyclops leijsi sp. nov., scanning electron micrographs, AD, paratype female 1, ventral; E, paratype female 2, lateral; F, paratype male 2, lateral: A, habitus; B, caudal rami; C, mouth appendages; D, fourth swimming leg; E, distal part of antennula; F, habitus." figureDoi="http://doi.org/10.5281/zenodo.14240882" httpUri="https://zenodo.org/record/14240882/files/figure.png" pageId="10" pageNumber="111">Figs. 6F</figureCitation>
,
<figureCitation id="137A68CC9C00FFDCFB2B94B9FB3B658B" box="[1207,1243,223,249]" captionStart="FIGURE 7" captionStartId="9.[151,250,1887,1911]" captionTargetBox="[151,1436,525,1864]" captionTargetId="figure-201@9.[151,1436,525,1864]" captionTargetPageId="9" captionText="FIGURE 7. Diacyclops leijsi sp. nov., scanning electron micrographs, A &amp; B, paratype male 2, lateral; CF, paratype male 3, dorsal:A, left antennula; B, detail of right antennula; C, free prosomites; D, first two urosomites; E, third and fourth urosomites; F, last two urosomites." figureDoi="http://doi.org/10.5281/zenodo.14240884" httpUri="https://zenodo.org/record/14240884/files/figure.png" pageId="10" pageNumber="111">7C</figureCitation>
), and urosomites (
<figureCitation id="137A68CC9C00FFDCFF039565FEDD646F" box="[159,317,259,285]" captionStart="FIGURE 7" captionStartId="9.[151,250,1887,1911]" captionTargetBox="[151,1436,525,1864]" captionTargetId="figure-201@9.[151,1436,525,1864]" captionTargetPageId="9" captionText="FIGURE 7. Diacyclops leijsi sp. nov., scanning electron micrographs, A &amp; B, paratype male 2, lateral; CF, paratype male 3, dorsal:A, left antennula; B, detail of right antennula; C, free prosomites; D, first two urosomites; E, third and fourth urosomites; F, last two urosomites." figureDoi="http://doi.org/10.5281/zenodo.14240884" httpUri="https://zenodo.org/record/14240884/files/figure.png" pageId="10" pageNumber="111">Fig. 7D, E, F</figureCitation>
) as in female. Caudal rami (
<figureCitation id="137A68CC9C00FFDCFD169565FD11646F" box="[650,753,258,285]" captionStart="FIGURE 5" captionStartId="7.[151,250,1828,1852]" captionTargetBox="[159,1436,181,1797]" captionTargetId="figure-15@7.[151,1436,181,1804]" captionTargetPageId="7" captionText="FIGURE 5. Diacyclops leijsi sp. nov., line drawings, paratype male 1: A, urosome, ventral; B, antennula; C, third endopodal segment of fourth swimming leg; D, sixth leg. Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.14240880" httpUri="https://zenodo.org/record/14240880/files/figure.png" pageId="10" pageNumber="111">Figs. 5A</figureCitation>
,
<figureCitation id="137A68CC9C00FFDCFC9C9565FCFE646F" box="[768,798,259,285]" captionStart="FIGURE 6" captionStartId="8.[151,250,1544,1568]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure-27@8.[151,1436,181,1520]" captionTargetPageId="8" captionText="FIGURE 6. Diacyclops leijsi sp. nov., scanning electron micrographs, AD, paratype female 1, ventral; E, paratype female 2, lateral; F, paratype male 2, lateral: A, habitus; B, caudal rami; C, mouth appendages; D, fourth swimming leg; E, distal part of antennula; F, habitus." figureDoi="http://doi.org/10.5281/zenodo.14240882" httpUri="https://zenodo.org/record/14240882/files/figure.png" pageId="10" pageNumber="111">6F</figureCitation>
) slightly slenderer than in female, with slightly shorter dorsal and innermost terminal setae. Antennula (
<figureCitation id="137A68CC9C00FFDCFD549541FCCD6433" box="[712,813,294,321]" captionStart="FIGURE 5" captionStartId="7.[151,250,1828,1852]" captionTargetBox="[159,1436,181,1797]" captionTargetId="figure-15@7.[151,1436,181,1804]" captionTargetPageId="7" captionText="FIGURE 5. Diacyclops leijsi sp. nov., line drawings, paratype male 1: A, urosome, ventral; B, antennula; C, third endopodal segment of fourth swimming leg; D, sixth leg. Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.14240880" httpUri="https://zenodo.org/record/14240880/files/figure.png" pageId="10" pageNumber="111">Figs. 5B</figureCitation>
,
<figureCitation id="137A68CC9C00FFDCFCA19541FC636433" box="[829,899,295,321]" captionStart="FIGURE 7" captionStartId="9.[151,250,1887,1911]" captionTargetBox="[151,1436,525,1864]" captionTargetId="figure-201@9.[151,1436,525,1864]" captionTargetPageId="9" captionText="FIGURE 7. Diacyclops leijsi sp. nov., scanning electron micrographs, A &amp; B, paratype male 2, lateral; CF, paratype male 3, dorsal:A, left antennula; B, detail of right antennula; C, free prosomites; D, first two urosomites; E, third and fourth urosomites; F, last two urosomites." figureDoi="http://doi.org/10.5281/zenodo.14240884" httpUri="https://zenodo.org/record/14240884/files/figure.png" pageId="10" pageNumber="111">7A, B</figureCitation>
) slender, strongly prehensile and digeniculate, 16-segmented, with anvil-shaped cuticular ridges on anterior margin of fourteenth and fifteenth segments (distal geniculation); 3 aesthetascs present on first segment, while fourth, ninth, and thirteenth segments carry 1 aesthetasc each; setae formula 8.4.2.2.2.2.1.1.2.2.2.2.1.2.1.13; several setae minute and slender, 1 on each ninth, eleventh, and twelfth segment spiniform, and 2 pairs of setae on ultimate segment fused basally; penultimate segment about 2.5 times as long as wide. Antenna, labrum, mandibula, maxillula, maxilla, maxilliped, all swimming legs (
<figureCitation id="137A68CC9C00FFDCFAA995BDFA6E6487" box="[1333,1422,474,501]" captionStart="FIGURE 5" captionStartId="7.[151,250,1828,1852]" captionTargetBox="[159,1436,181,1797]" captionTargetId="figure-15@7.[151,1436,181,1804]" captionTargetPageId="7" captionText="FIGURE 5. Diacyclops leijsi sp. nov., line drawings, paratype male 1: A, urosome, ventral; B, antennula; C, third endopodal segment of fourth swimming leg; D, sixth leg. Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.14240880" httpUri="https://zenodo.org/record/14240880/files/figure.png" pageId="10" pageNumber="111">Fig. 5C</figureCitation>
), and fifth leg (
<figureCitation id="137A68CC9C00FFDCFEAD9599FE69676B" box="[305,393,510,537]" captionStart="FIGURE 5" captionStartId="7.[151,250,1828,1852]" captionTargetBox="[159,1436,181,1797]" captionTargetId="figure-15@7.[151,1436,181,1804]" captionTargetPageId="7" captionText="FIGURE 5. Diacyclops leijsi sp. nov., line drawings, paratype male 1: A, urosome, ventral; B, antennula; C, third endopodal segment of fourth swimming leg; D, sixth leg. Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.14240880" httpUri="https://zenodo.org/record/14240880/files/figure.png" pageId="10" pageNumber="111">Fig. 5A</figureCitation>
) as in female. Sixth leg (
<figureCitation id="137A68CC9C00FFDCFD3A9599FCC1676B" box="[678,801,510,537]" captionStart="FIGURE 5" captionStartId="7.[151,250,1828,1852]" captionTargetBox="[159,1436,181,1797]" captionTargetId="figure-15@7.[151,1436,181,1804]" captionTargetPageId="7" captionText="FIGURE 5. Diacyclops leijsi sp. nov., line drawings, paratype male 1: A, urosome, ventral; B, antennula; C, third endopodal segment of fourth swimming leg; D, sixth leg. Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.14240880" httpUri="https://zenodo.org/record/14240880/files/figure.png" pageId="10" pageNumber="111">Fig. 5A, D</figureCitation>
) longer than in female, with better developed innermost spine and with central slender seta; outermost seta about 2.7 times as long as spine or central seta.
</paragraph>
</subSubSection>
</treatment>
</document>