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<document id="CB7AE098C5480524709F717E9C9AAD46" ID-DOI="10.11646/phytotaxa.363.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13703414" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725581356715" checkinUser="felipe" docAuthor="Negaresh, Kazem &amp; Rahiminejad, Mohammad Reza" docDate="2018" docId="03E887F4DF4DFF9319D43C80F327FB2A" docLanguage="en" docName="phytotaxa.363.1.1.pdf" docOrigin="Phytotaxa 363 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.363.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.4:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="4" docTitle="Centaurea regia var. longispinosa Negaresh 2018, var. nov." docType="treatment" docVersion="2" lastPageNumber="450" masterDocId="FFD1FF8CDF12FFCC195C3E3CF14EFF83" masterDocTitle="A revision of Centaurea sect. Cynaroides (Asteraceae, Cardueae-Centaureinae)" masterLastPageNumber="450" masterPageNumber="448" pageNumber="448" updateTime="1725593506943" updateUser="ExternalLinkService">
<mods:mods id="BC4DF004FCA6861F18C3E3F41BC46A3F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F1E6DF5A14AD38B28A6770BA2BBFA899">
<mods:title id="1AD50CD257DC005ACB515319F18FB464">A revision of Centaurea sect. Cynaroides (Asteraceae, Cardueae-Centaureinae)</mods:title>
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<mods:name id="67B70BFDFE0E7D29D1E6093D4F0915AB" type="personal">
<mods:role id="DD8F0463E56E0795E813857D48A311B3">
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<mods:namePart id="9AF197EFB2A51B903565B3E9652BD9BB">Negaresh, Kazem</mods:namePart>
<mods:affiliation id="0B5D1BB1A2C21B4ADF3C42037A5D3097">Department of Horticulture, Faculty of Agriculture, Khuzestan Agricultural Sciences and Natural Resources University, Mollasani, Ahvaz, Iran. (* Corresponding author e-mail: negaresh @ ramin. ac. ir)</mods:affiliation>
<mods:nameIdentifier id="BBEDDA8B67B68CDC343C4225EDF8E1F9" type="email">negaresh@ramin.ac.ir</mods:nameIdentifier>
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<mods:roleTerm id="2E21AF9F6EF831E43E8D960C32CAACE3">Author</mods:roleTerm>
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<mods:namePart id="C6AD7A297EC69D88107261FBBBC5FD88">Rahiminejad, Mohammad Reza</mods:namePart>
<mods:affiliation id="0D21AB01B4743031009794E007DB6FC5">Department of Horticulture, Faculty of Agriculture, Khuzestan Agricultural Sciences and Natural Resources University, Mollasani, Ahvaz, Iran. (* Corresponding author e-mail: negaresh @ ramin. ac. ir) &amp; Department of Biology, University of Isfahan, Isfahan 81746 - 73441, Iran. &amp; Department of Horticulture, Faculty of Agriculture, Khuzestan Agricultural Sciences and Natural Resources University, Mollasani, Ahvaz, Iran. (* Corresponding author e-mail: negaresh @ ramin. ac. ir)</mods:affiliation>
<mods:nameIdentifier id="E4650A10B8EB954F914FA3708300D6AC" type="email">negaresh@ramin.ac.ir</mods:nameIdentifier>
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<mods:title id="42C22256E51AED29CA427AEA07334C25">Phytotaxa</mods:title>
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<mods:part id="E4E38E0CB6471FAEF8F2C9D6CFF92FD8">
<mods:date id="5505A85912250BD3CBD2D8053AD84B07">2018</mods:date>
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<mods:number id="2D5AEABCE9DD5767B80F3C39B4692ABD">2018-08-02</mods:number>
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<mods:number id="C207FFEF0B8AB292EA03BFA01840B3EB">363</mods:number>
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<mods:classification id="E2E240240DD5E6792525C2D9E7C3593C">journal article</mods:classification>
<mods:identifier id="9DD1D7676DF225E2C93AD81426D78213" type="DOI">10.11646/phytotaxa.363.1.1</mods:identifier>
<mods:identifier id="6F10466E76B101391EBF565DE8BA5426" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="C46DED0EF9D251327B4AAFD8CCEF16A5" type="Zenodo-Dep">13703414</mods:identifier>
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<treatment id="03E887F4DF4DFF9319D43C80F327FB2A" ID-DOI="http://doi.org/10.5281/zenodo.13703796" ID-Zenodo-Dep="13703796" LSID="urn:lsid:plazi:treatment:03E887F4DF4DFF9319D43C80F327FB2A" httpUri="http://treatment.plazi.org/id/03E887F4DF4DFF9319D43C80F327FB2A" pageId="95">
<subSubSection id="C35B6569DF4DFF9319D43C80F2DEFD56" box="[136,912,698,725]" pageId="95" type="nomenclature">
<paragraph id="8BFE36E2DF4DFF9319D43C80F2DEFD56" blockId="95.[136,1451,698,867]" box="[136,912,698,725]" pageId="95">
<emphasis id="B935EAF0DF4DFF9319D43C80F1F9FD56" bold="true" box="[136,183,700,725]" pageId="95">28c.</emphasis>
<taxonomicName id="4C414D61DF4DFF9319E23C86F3F9FD56" ID-CoL="7LGQG" authority="Negaresh" authorityName="Negaresh" authorityYear="2018" box="[190,695,698,725]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="95" phylum="Tracheophyta" rank="variety" species="regia" status="var. nov." variety="longispinosa">
<emphasis id="B935EAF0DF4DFF9319E23C86F034FD57" bold="true" box="[190,378,698,724]" italics="true" pageId="95">Centaurea regia</emphasis>
var.
<emphasis id="B935EAF0DF4DFF9318ED3C86F30DFD56" bold="true" box="[433,579,698,725]" italics="true" pageId="95">longispinosa</emphasis>
Negaresh
</taxonomicName>
,
<emphasis id="B935EAF0DF4DFF931B9E3C80F269FD56" bold="true" box="[706,807,700,725]" pageId="95">
<taxonomicNameLabel id="A206578BDF4DFF931B9E3C80F269FD56" box="[706,807,700,725]" pageId="95" rank="variety">var. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="137A2A67DF4DFF931A6A3C80F2C6FD56" box="[822,904,700,725]" captionStart="FIGURE 65" captionStartId="96.[136,229,1887,1908]" captionTargetBox="[151,1435,190,1862]" captionTargetId="figure-17@96.[151,1436,190,1862]" captionTargetPageId="96" captionText="FIGURE 65. Centaurea regia Boissier var. longispinosa Negaresh (Negaresh &amp; Kamalnejad 112, herb. Negaresh).A. close up of capitule. B. inflorescence. C. habit. D. median cauline leaves. Scale A = 2 cm, B = 10 cm, C and D = 8 cm." figureDoi="http://doi.org/10.5281/zenodo.13703590" httpUri="https://zenodo.org/record/13703590/files/figure.png" pageId="95">Fig. 65</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35B6569DF4DFF9319D43CDEF5B8FC2E" pageId="95" type="description">
<paragraph id="8BFE36E2DF4DFF9319D43CDEF00EFC98" blockId="95.[136,1451,698,867]" pageId="95">
<emphasis id="B935EAF0DF4DFF9319D43CDEF00EFC98" italics="true" pageId="95">
<taxonomicName id="4C414D61DF4DFF9319D43CDEF34FFD74" box="[136,513,738,759]" class="Magnoliopsida" family="Asteraceae" genus="Centaureae" kingdom="Plantae" order="Asterales" pageId="95" phylum="Tracheophyta" rank="variety" species="regiae" variety="cynarocephalae">Centaureae regiae var. cynarocephalae</taxonomicName>
<taxonomicName id="4C414D61DF4DFF931B543CDEF309FD74" box="[520,583,738,759]" class="Magnoliopsida" family="Asteraceae" genus="Cynaroides" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asterales" pageId="95" phylum="Tracheophyta" rank="species" species="affinis">affinis</taxonomicName>
, a qua spinis terminalibus appendicum mediorum (17)
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(nec (3)48(13) mm) longis differt.
</emphasis>
</paragraph>
<paragraph id="8BFE36E2DF4DFF9319D43D16F229FCE0" blockId="95.[136,1451,698,867]" pageId="95">
<materialsCitation id="3B293CBFDF4DFF9319D43D16F22DFCE0" collectingDate="2014-05-30" collectionCode="FUMH, RUH" collectorName="Negaresh &amp; Kamalnejad" country="Iran" county="Type" elevation="1388" location="Kamyaran" municipality="Ravansar" pageId="95" specimenCount="2" stateProvince="Kermanshah" typeStatus="holotype">
<collectingCounty id="629F4E6EDF4DFF9319D43D16F1F5FCBC" box="[136,187,810,831]" pageId="95">Type</collectingCounty>
:—
<collectingCountry id="F3567672DF4DFF93198B3D16F05AFCBC" box="[215,276,810,831]" name="Iran" pageId="95">IRAN</collectingCountry>
. Prov.
<collectingRegion id="4985F800DF4DFF93180F3D16F09EFCBC" box="[339,464,810,831]" country="Iran" name="Kermanshah" pageId="95">Kermanshah</collectingRegion>
:
<collectingMunicipality id="6B9AAC98DF4DFF9318853D16F37CFCBC" box="[473,562,810,831]" pageId="95">Ravansar</collectingMunicipality>
to
<location id="8E9E6039DF4DFF931B133D16F3FBFCBC" LSID="urn:lsid:plazi:treatment:03E887F4DF4DFF9319D43C80F327FB2A:8E9E6039DF4DFF931B133D16F3FBFCBC" box="[591,693,810,831]" country="Iran" county="Type" municipality="Ravansar" name="Kamyaran" pageId="95" stateProvince="Kermanshah">Kamyaran</location>
, 13 after
<location id="8E9E6039DF4DFF931A453D16F23AFCBC" LSID="urn:lsid:plazi:treatment:03E887F4DF4DFF9319D43C80F327FB2A:8E9E6039DF4DFF931A453D16F23AFCBC" box="[793,884,810,831]" country="Iran" county="Type" municipality="Ravansar" name="Ravansar" pageId="95" stateProvince="Kermanshah">Ravansar</location>
,
<quantity id="4CB99B07DF4DFF931A213D16F54DFCBC" box="[893,1027,810,831]" metricMagnitude="3" metricUnit="m" metricValue="1.3875" metricValueMax="1.4" metricValueMin="1.375" pageId="95" unit="m" value="1387.5" valueMax="1400.0" valueMin="1375.0">
<elevation id="006CD1D1DF4DFF931A213D16F54DFCBC" box="[893,1027,810,831]" metricMagnitude="3" metricUnit="m" metricValue="1.3875" metricValueMax="1.4" metricValueMin="1.375" pageId="95" unit="m" value="1387.5" valueMax="1400.0" valueMin="1375.0">13751400 m</elevation>
</quantity>
,
<date id="FFFF1022DF4DFF931D503D16F5C3FCBC" box="[1036,1165,810,831]" pageId="95" value="2014-05-30">
<collectingDate id="EFBBE9CADF4DFF931D503D16F5C3FCBC" box="[1036,1165,810,831]" pageId="95" value="2014-05-30">30 May 2014</collectingDate>
</date>
,
<emphasis id="B935EAF0DF4DFF931DCA3D16F4E5FCBC" box="[1174,1451,810,831]" italics="true" pageId="95">
<collectorName id="26B45334DF4DFF931DCA3D16F5BFFCBC" box="[1174,1265,810,831]" pageId="95">Negaresh</collectorName>
&amp;
<collectorName id="26B45334DF4DFF931C523D16F4CDFCBC" box="[1294,1411,810,831]" pageId="95">Kamalnejad</collectorName>
112
</emphasis>
(
<typeStatus id="54FA8840DF4DFF9319983D72F054FCE0" box="[196,282,846,867]" pageId="95" type="holotype">holotype</typeStatus>
herb. Negaresh!,
<typeStatus id="54FA8840DF4DFF9318993D72F35AFCE0" box="[453,532,846,867]" pageId="95" type="isotype">isotypes</typeStatus>
<collectionCode id="ED50AE27DF4DFF931B453D72F328FCE0" box="[537,614,846,867]" country="Iran" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15384" lsid="urn:lsid:biocol.org:col:15384" name="Ferdowsi University" pageId="95" type="Herbarium">FUMH</collectionCode>
!, herb. Negaresh.!,
<collectionCode id="ED50AE27DF4DFF931A423D72F217FCE0" box="[798,857,846,867]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15206" lsid="urn:lsid:biocol.org:col:15206" name="Rhodes University" pageId="95" type="Herbarium">RUH</collectionCode>
!)
</materialsCitation>
.
</paragraph>
<paragraph id="8BFE36E2DF4DFF9319D43DA8F5B8FC2E" blockId="95.[136,1452,916,1193]" box="[136,1270,916,941]" pageId="95">
Terminal spine of median appenadges (17)
<quantity id="4CB99B07DF4DFF931B2D3DA8F3A3FC2E" box="[625,749,916,941]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="95" unit="mm" value="22.5" valueMax="25.0" valueMin="20.0">2025 mm</quantity>
long; appendages brown; cilia
<quantity id="4CB99B07DF4DFF931D173DA8F5F8FC2E" box="[1099,1206,916,941]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="10.0" metricValueMin="3.0" pageId="95" unit="mm" value="6.5" valueMax="10.0" valueMin="3.0">310 mm</quantity>
long.
</paragraph>
</subSubSection>
<subSubSection id="C35B6569DF4DFF9319E13D84F327FB2A" pageId="95" type="etymology">
<paragraph id="8BFE36E2DF4DFF9319E13D84F553FC52" blockId="95.[136,1452,916,1193]" box="[189,1053,952,977]" pageId="95">
<emphasis id="B935EAF0DF4DFF9319E13D84F006FC52" bold="true" box="[189,328,952,977]" pageId="95">Etymology:</emphasis>
—The specific epithet longispinosa means terminal spine long.
</paragraph>
<paragraph id="8BFE36E2DF4DFF9319E13DE0F345FBBE" blockId="95.[136,1452,916,1193]" pageId="95">
<emphasis id="B935EAF0DF4DFF9319E13DE0F3CBFC76" bold="true" box="[189,645,988,1013]" pageId="95">Taxonomic and distribution remarks:</emphasis>
<taxonomicName id="4C414D61DF4DFF931BFD3DE0F56BFC76" authorityName="Negaresh" authorityYear="2018" box="[673,1061,988,1013]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="95" phylum="Tracheophyta" rank="variety" species="regia" variety="longispinosa">
<emphasis id="B935EAF0DF4DFF931BFD3DE0F214FC76" box="[673,858,988,1013]" italics="true" pageId="95">Centaurea regia</emphasis>
<materialsCitation id="3B293CBFDF4DFF931A3F3DE0F56BFC76" box="[867,1061,988,1013]" collectionCode="W" country="Iran" location="Kermanshah Province" pageId="95" specimenCount="1" stateProvince="Kermanshah">
var.
<emphasis id="B935EAF0DF4DFF931ACA3DE0F56BFC76" box="[918,1061,988,1013]" italics="true" pageId="95">longispinosa</emphasis>
</materialsCitation>
</taxonomicName>
is a rare endemic of
<collectingRegion id="4985F800DF4DFF931C423DE0F1A0FB9A" country="Iran" name="Kermanshah" pageId="95">Kermanshah Province</collectingRegion>
, W
<collectingCountry id="F3567672DF4DFF9318433A3CF002FB9A" box="[287,332,1024,1049]" name="Iran" pageId="95">Iran</collectingCountry>
(
<figureCitation id="137A2A67DF4DFF9318023A3CF0FBFB9A" box="[350,437,1024,1049]" captionStart="FIGURE 60" captionStartId="89.[136,229,1022,1043]" captionTargetBox="[154,1419,228,968]" captionTargetId="figure-14@89.[151,1436,190,997]" captionTargetPageId="89" captionText="FIGURE 60. Distribution of Centaurea regia var. longispinosa (circles) and C. regia var. purpurea (squares)." figureDoi="http://doi.org/10.5281/zenodo.13703572" httpUri="https://zenodo.org/record/13703572/files/figure.png" pageId="95">Fig. 60</figureCitation>
). It is distinguished from the other varieties by a longer spine (17)
<quantity id="4CB99B07DF4DFF931D983A3CF40DFB9A" box="[1220,1347,1024,1049]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="95" unit="mm" value="22.5" valueMax="25.0" valueMin="20.0">2025 mm</quantity>
and also somewhat longer cilia (
<quantity id="4CB99B07DF4DFF9318D33A18F0B1FBBE" box="[399,511,1060,1085]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="10.0" metricValueMin="3.0" pageId="95" unit="mm" value="6.5" valueMax="10.0" valueMin="3.0">310 mm</quantity>
).
</paragraph>
<paragraph id="8BFE36E2DF4DFF9319E13A74F25FFB06" blockId="95.[136,1452,916,1193]" pageId="95">
<materialsCitation id="3B293CBFDF4DFF9319E13A74F243FB06" collectingDate="2017-06-16" collectionCode="RUH" country="Iran" elevation="1370" latitude="34.717888" location="Sardem village" longLatPrecision="1" longitude="46.67739" municipality="Ravansar" pageId="95" specimenCount="1" stateProvince="Kermanshah" typeStatus="paratype">
<emphasis id="B935EAF0DF4DFF9319E13A74F064FBE2" bold="true" box="[189,298,1096,1121]" pageId="95">
<typeStatus id="54FA8840DF4DFF9319E13A74F064FBE2" box="[189,298,1096,1121]" pageId="95" type="paratype">Paratype</typeStatus>
</emphasis>
:—
<collectingCountry id="F3567672DF4DFF9318103A74F0D8FBE2" box="[332,406,1096,1121]" name="Iran" pageId="95">IRAN</collectingCountry>
. Prov.
<collectingRegion id="4985F800DF4DFF9318B83A74F338FBE2" box="[484,630,1096,1121]" country="Iran" name="Kermanshah" pageId="95">Kermanshah</collectingRegion>
:
<collectingMunicipality id="6B9AAC98DF4DFF931BDD3A74F3A7FBE2" box="[641,745,1096,1121]" pageId="95">Ravansar</collectingMunicipality>
toward
<location id="8E9E6039DF4DFF931A143A74F2BAFBE2" LSID="urn:lsid:plazi:treatment:03E887F4DF4DFF9319D43C80F327FB2A:8E9E6039DF4DFF931A143A74F2BAFBE2" box="[840,1012,1096,1121]" country="Iran" latitude="34.717888" longLatPrecision="1" longitude="46.67739" municipality="Ravansar" name="Sardem village" pageId="95" stateProvince="Kermanshah">Sardem village</location>
,
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,
<geoCoordinate id="EE755025DF4DFF931DE93A74F428FBE2" box="[1205,1382,1096,1121]" degrees="46" direction="east" minutes="40" orientation="longitude" pageId="95" precision="1" seconds="38.6" value="46.67739">46° 40 38.6″ E</geoCoordinate>
,
<quantity id="4CB99B07DF4DFF931C2F3A74F1EEFB06" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" pageId="95" unit="m" value="1370.0">
<elevation id="006CD1D1DF4DFF931C2F3A74F1EEFB06" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" pageId="95" unit="m" value="1370.0">1370 m</elevation>
</quantity>
,
<date id="FFFF1022DF4DFF9319F03A50F00AFB06" box="[172,324,1132,1157]" pageId="95" value="2017-06-16">
<collectingDate id="EFBBE9CADF4DFF9319F03A50F00AFB06" box="[172,324,1132,1157]" pageId="95" value="2017-06-16">16 June 2017</collectingDate>
</date>
,
<emphasis id="B935EAF0DF4DFF9318133A50F0A4FB06" box="[335,490,1132,1157]" italics="true" pageId="95">Negaresh 180</emphasis>
(herb. Negaresh!,
<collectionCode id="ED50AE27DF4DFF931BE03A50F24FFB06" box="[700,769,1132,1157]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15206" lsid="urn:lsid:biocol.org:col:15206" name="Rhodes University" pageId="95" type="Herbarium">RUH</collectionCode>
!)
</materialsCitation>
.
</paragraph>
<paragraph id="8BFE36E2DF4DFF9319E13AACF327FB2A" blockId="95.[136,1452,916,1193]" box="[189,617,1168,1193]" pageId="95">
<emphasis id="B935EAF0DF4DFF9319E13AACF099FB2A" bold="true" box="[189,471,1168,1193]" pageId="95">Chromosome numbers:</emphasis>
—Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>