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<document id="16FE4702EC6F8342EE74D7E083B53631" ID-CLB-Dataset="27747" ID-DOI="10.11646/zootaxa.4532.3.1" ID-GBIF-Dataset="92005c51-bac6-4193-acc5-5a6b56c0b5cd" ID-ISSN="1175-5326" ID-Zenodo-Dep="2615331" ID-ZooBank="2929A344-3A07-4FB6-96C9-C1B7497BF6CF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553788124377" checkinUser="plazi" docAuthor="Wang, Wendy Y., Yong, Gordon W. J. &amp; Jaitrong, Weeyawat" docDate="2018" docId="03E887EE9748FFDDFF35E5EFFD9FFCDC" docLanguage="en" docName="zootaxa.4532.3.1.pdf" docOrigin="Zootaxa 4532 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhopalomastix javana Wheeler, W. M. 1929" docType="treatment" docVersion="8" lastPageNumber="316" masterDocId="FFD1FF969741FFD2FFA2E144FF96FFC8" masterDocTitle="The ant genus Rhopalomastix (Hymenoptera: Formicidae: Myrmicinae) in Southeast Asia, with descriptions of four new species from Singapore based on morphology and DNA barcoding" masterLastPageNumber="340" masterPageNumber="301" pageNumber="310" updateTime="1698715070841" updateUser="ExternalLinkService">
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<mods:title id="FD3F77027DF3D1182056C3F28796EBDB">The ant genus Rhopalomastix (Hymenoptera: Formicidae: Myrmicinae) in Southeast Asia, with descriptions of four new species from Singapore based on morphology and DNA barcoding</mods:title>
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<mods:namePart id="57FF82844630DE123AAE4A576F69CF9D">Wang, Wendy Y.</mods:namePart>
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<mods:namePart id="58C6E36B871B0BC8B0310C2BF0E2CB95">Yong, Gordon W. J.</mods:namePart>
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<mods:namePart id="6AEA9664E581D0885CC4EC654B1AC9F4">Jaitrong, Weeyawat</mods:namePart>
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<treatment id="03E887EE9748FFDDFF35E5EFFD9FFCDC" ID-DOI="http://doi.org/10.5281/zenodo.5979187" ID-GBIF-Taxon="154521343" ID-Zenodo-Dep="5979187" LSID="urn:lsid:plazi:treatment:03E887EE9748FFDDFF35E5EFFD9FFCDC" httpUri="http://treatment.plazi.org/id/03E887EE9748FFDDFF35E5EFFD9FFCDC" lastPageId="15" lastPageNumber="316" pageId="9" pageNumber="310">
<subSubSection id="C35B65739748FFDBFF35E5EFFC9AFB0E" box="[151,780,1195,1222]" pageId="9" pageNumber="310" type="nomenclature">
<paragraph id="8BFE36F89748FFDBFF35E5EFFC9AFB0E" blockId="9.[151,780,1195,1255]" box="[151,780,1195,1222]" pageId="9" pageNumber="310">
<heading id="D0B681949748FFDBFF35E5EFFC9AFB0E" bold="true" box="[151,780,1195,1222]" fontSize="11" level="1" pageId="9" pageNumber="310" reason="1">
<emphasis id="B935EAEA9748FFDBFF35E5EFFC9AFB0E" bold="true" box="[151,780,1195,1222]" pageId="9" pageNumber="310">
<taxonomicName id="4C414D7B9748FFDBFF35E5EFFD27FB0E" authority="Wheeler, W. M., 1929" authorityName="Wheeler, W. M." authorityYear="1929" box="[151,689,1195,1222]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="B935EAEA9748FFDBFF35E5EFFE31FB0D" bold="true" box="[151,423,1195,1221]" italics="true" pageId="9" pageNumber="310">Rhopalomastix javana</emphasis>
<bibRefCitation id="EFD04B099748FFDBFE0FE5E8FD27FB0E" author="Wheeler, W. M." box="[429,689,1196,1222]" pageId="9" pageNumber="310" pagination="95 - 101" refId="ref25770" refString="Wheeler, W. M. (1929) The ant genus Rhopalomastix. Psyche, Cambridge, 36, 95 - 101. https: // doi. org / 10.1155 / 1929 / 60827" type="journal article" year="1929">Wheeler, W.M., 1929</bibRefCitation>
</taxonomicName>
stat. n.
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C35B65739748FFDBFF35E58AFEB6FB2F" box="[151,288,1230,1255]" pageId="9" pageNumber="310" type="description">
<paragraph id="8BFE36F89748FFDBFF35E58AFEB6FB2F" blockId="9.[151,780,1195,1255]" box="[151,288,1230,1255]" pageId="9" pageNumber="310">
(
<figureCitation id="137A2A7D9748FFDBFF3DE58AFE8EFB2F" box="[159,280,1230,1255]" captionStart-0="FIGURES 811" captionStart-1="FIGURES 1215" captionStart-2="FIGURES 1619" captionStart-3="FIGURES 2023" captionStartId-0="10.[151,264,1704,1727]" captionStartId-1="11.[151,264,1903,1926]" captionStartId-2="12.[151,264,1167,1190]" captionStartId-3="14.[151,264,1983,2006]" captionTargetBox-0="[151,1436,625,1684]" captionTargetBox-1="[151,1436,833,1892]" captionTargetBox-2="[151,1436,193,1146]" captionTargetBox-3="[226,1360,1088,1975]" captionTargetId-0="figure@10.[151,1436,625,1684]" captionTargetId-1="figure@11.[151,1436,833,1892]" captionTargetId-2="figure@12.[151,1436,193,1146]" captionTargetId-3="figure@14.[226,1360,1088,1975]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionTargetPageId-3="14" captionText-0="FIGURES 811. Lectotype worker of R. javana stat. n. 8, habitus in profile; 9, head in full-face view; 10, closeup of head in profile view; 11, body in dorsal view." captionText-1="FIGURES 1215. Non-type worker of R. javana stat. n. from Singapore, Mandai Track, ZRC_HYM_0000576; 12, habitus in profile; 13, head in full face view; 14, closeup of head in profile view; 15, body in dorsal view." captionText-2="FIGURES 1619. Non-type queen of R. javana from Singapore, Bukit Timah Nature Reserve, ZRC_HYM_0001732; 16, habitus in profile; 17, head in full face view; 18, body in dorsal view; 19, closeup of mesosoma in dorsal view." captionText-3="FIGURES 2023. Non-type male of R. javana from Singapore, Mandai track, ZRC_HYM_0000576; 20, habitus in profile; 21, head in full face view; 22, body in dorsal view; 23, closeup of mesosoma in dorsal view." httpUri-0="https://zenodo.org/record/2615341/files/figure.png" httpUri-1="https://zenodo.org/record/2615345/files/figure.png" httpUri-2="https://zenodo.org/record/2615347/files/figure.png" httpUri-3="https://zenodo.org/record/2615349/files/figure.png" pageId="9" pageNumber="310">Figs. 823</figureCitation>
)
</paragraph>
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<subSubSection id="C35B65739748FFDBFF35E452FB6DFA72" pageId="9" pageNumber="310" type="reference_group">
<paragraph id="8BFE36F89748FFDBFF35E452FCB1FAE4" blockId="9.[151,807,1302,1325]" box="[151,807,1302,1325]" pageId="9" pageNumber="310">
<treatmentCitationGroup id="AB5111D69748FFDBFF35E452FCB1FAE4" box="[151,807,1302,1325]" pageId="9" pageNumber="310">
<taxonomicName id="4C414D7B9748FFDBFF35E452FCB5FAE4" ID-CoL="5KX5Y" authority="Wheeler, W. M., 1929: 96" authorityName="Wheeler, W. M." authorityPageNumber="96" authorityYear="1929" box="[151,803,1302,1325]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="subSpecies" species="rothneyi" subSpecies="javana">
<emphasis id="B935EAEA9748FFDBFF35E452FE1EFAE4" box="[151,392,1302,1324]" italics="true" pageId="9" pageNumber="310">Rhopalomastix rothneyi</emphasis>
subsp.
<emphasis id="B935EAEA9748FFDBFE77E45CFD8CFAE5" box="[469,538,1304,1325]" italics="true" pageId="9" pageNumber="310">javana</emphasis>
<treatmentCitation id="0AE010E99748FFDBFD82E452FCB5FAE4" author="Wheeler, W. M." box="[544,803,1302,1324]" page="96" pageId="9" pageNumber="310" year="1929">
<bibRefCitation id="EFD04B099748FFDBFD82E452FD6BFAE4" author="Wheeler, W. M." box="[544,765,1302,1324]" pageId="9" pageNumber="310" pagination="95 - 101" refId="ref25770" refString="Wheeler, W. M. (1929) The ant genus Rhopalomastix. Psyche, Cambridge, 36, 95 - 101. https: // doi. org / 10.1155 / 1929 / 60827" type="journal article" year="1929">Wheeler, W.M., 1929</bibRefCitation>
: 96
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFE36F89748FFDBFF35E41DFB6DFA72" blockId="9.[151,1437,1369,2006]" pageId="9" pageNumber="310">
<emphasis id="B935EAEA9748FFDBFF35E41DFF75FABA" bold="true" box="[151,227,1369,1394]" pageId="9" pageNumber="310">Types.</emphasis>
<taxonomicName id="4C414D7B9748FFDBFF54E41DFCC1FAB9" ID-CoL="5KX5Y" authority=": Numerous" authorityName="Numerous" box="[246,855,1369,1394]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="subSpecies" species="rothneyi" subSpecies="javana">
<emphasis id="B935EAEA9748FFDBFF54E41DFD98FAB9" box="[246,526,1369,1393]" italics="true" pageId="9" pageNumber="310">Rhopalomastix rothneyi</emphasis>
subsp.
<emphasis id="B935EAEA9748FFDBFDD9E41EFD51FAB9" box="[635,711,1370,1393]" italics="true" pageId="9" pageNumber="310">javana</emphasis>
: Numerous
</taxonomicName>
workers, four females and three males from
<collectingCountry id="F35676689748FFDBFF35E43AFEA4FA5E" box="[151,306,1406,1430]" name="Indonesia" pageId="9" pageNumber="310">INDONESIA</collectingCountry>
, Eastern
<collectingRegion id="4985F81A9748FFDBFE11E43AFE7EFA5D" box="[435,488,1406,1429]" country="Indonesia" name="Jawa Timur" pageId="9" pageNumber="310">Java</collectingRegion>
, Besoeki, Bondowoso. Three
<typeStatus id="54FA885A9748FFDBFCCCE43AFC53FA5E" box="[878,965,1406,1430]" pageId="9" pageNumber="310" type="syntype">syntype</typeStatus>
workers on 1 pin (USNM, cotype: MCZ.6.9.20783/ SNM.595.31) were examined. One worker (top on pin) is selected as the
<typeStatus id="54FA885A9748FFDBFB2EE4E5FB61FA72" box="[1164,1271,1441,1466]" pageId="9" pageNumber="310" type="lectotype">lectotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C35B65739748FFDBFF65E481FCDFF95A" pageId="9" pageNumber="310" type="diagnosis">
<paragraph id="8BFE36F89748FFDBFF65E481FCDFF95A" blockId="9.[151,1437,1369,2006]" pageId="9" pageNumber="310">
<emphasis id="B935EAEA9748FFDBFF65E481FE3DFA16" bold="true" box="[199,427,1477,1502]" pageId="9" pageNumber="310">Diagnosis. Worker.</emphasis>
Workers monomorphic with little size variation (
<typeStatus id="54FA885A9748FFDBFC7EE481FBD3FA16" box="[988,1093,1477,1502]" pageId="9" pageNumber="310" type="lectotype">lectotype</typeStatus>
and
<typeStatus id="54FA885A9748FFDBFBDCE481FAB2FA16" box="[1150,1316,1477,1502]" pageId="9" pageNumber="310" type="paralectotype">paralectotypes</typeStatus>
: HL 0.48 0.52, HW 0.450.48). Head in full face view subrectangular, longer than wide, posterior margin with a shallow median concavity. In profile, clypeus broadly and weakly convex; entire dorsum finely striated; posterior half of head striate-reticulate and weakly shining; ventral surface of head largely superficially reticulate; outer surface of femora reticulate with shining interspaces; petiole bell-shaped, anterior face weakly concave. Head, mesosoma dorsum and gaster dark brown, rest of body yellowish brown.
</paragraph>
</subSubSection>
<subSubSection id="C35B65739748FFDFFF65E7D9FCE5FB20" lastPageId="13" lastPageNumber="314" pageId="9" pageNumber="310" type="description">
<paragraph id="8BFE36F89748FFDBFF65E7D9FB96F8A2" blockId="9.[151,1437,1369,2006]" pageId="9" pageNumber="310">
<emphasis id="B935EAEA9748FFDBFF65E7D9FE8AF97E" bold="true" box="[199,284,1693,1718]" pageId="9" pageNumber="310">Queen.</emphasis>
Small-medium size (non-types: HL 0.480.49, HW 0.440.45), head slightly less rectangular than that of worker. In profile view, clypeus very weakly convex, almost flat; posterior half and ventral surface of head largely smooth and shiny, vertex feebly striated; outer surface of femora and gaster superficially reticulate; dorsum of mesosoma finely striate and shiny; anterior pronotal margin roundly convex; anterior face of petiole evenly concave, petiole distinctly narrower apically. Body colour generally uniformly dark brown; head, dorsum of mesosoma and gaster slightly darker brown, tibia and tarsus yellowish brown.
</paragraph>
<paragraph id="8BFE36F89748FFD8FF65E631FC99FF30" blockId="9.[151,1437,1369,2006]" lastBlockId="10.[151,1436,151,573]" lastPageId="10" lastPageNumber="311" pageId="9" pageNumber="310">
<emphasis id="B935EAEA9748FFDBFF65E631FE9DF846" bold="true" box="[199,267,1909,1934]" pageId="9" pageNumber="310">Male.</emphasis>
Small size (non-types: HL 0.390.41, HW 0.380.4). Head almost entirely spherical in full face view, posterior margin broad and round; eye relatively smaller than other species, maximum diameter of eye only slightly exceeding midline of head; clypeus in profile roundly convex, strongly projecting form dorsum of head, rugulose with punctured interspaces and weakly shining; petiole short and sub-trapezoidal, apex smooth and blunt; entire head substriate-reticulate with smooth and shining interspaces, rest of body superficially reticulate; short standing hairs on entire dorsum including petiole and postpetiole.
</paragraph>
<paragraph id="8BFE36F8974BFFD8FF65E040FAC4FE88" blockId="10.[151,1436,151,573]" pageId="10" pageNumber="311">
<emphasis id="B935EAEA974BFFD8FF65E040FE49FED5" bold="true" box="[199,479,260,285]" pageId="10" pageNumber="311">Worker measurements.</emphasis>
<typeStatus id="54FA885A974BFFD8FE45E041FDC1FED5" box="[487,599,261,285]" pageId="10" pageNumber="311" type="lectotype">Lectotype</typeStatus>
and
<typeStatus id="54FA885A974BFFD8FD2DE040FCA7FED5" box="[655,817,260,285]" pageId="10" pageNumber="311" type="paralectotype">paralectotypes</typeStatus>
(n = 3): TL 1.831.98; HL 0.480.52; HW 0.450.48; SL 0.180.20; ML 0.480.50; PronW 0.280.32; EL 0.08; EW 0.040.06; CI 9294; REL 1718; SI 3942.
</paragraph>
<paragraph id="8BFE36F8974BFFD8FF65E008FC41FE40" blockId="10.[151,1436,151,573]" pageId="10" pageNumber="311">Nine non-types (n = 9): TL 1.692.02; HL 0.460.48; HW 0.410.44; SL 0.170.19; ML 0.500.54; PronW 0.280.32; EL 0.080.1; EW 0.040.06; CI 8993; REL 1823; SI 4044.</paragraph>
<paragraph id="8BFE36F8974BFFD8FF65E0D0FE92FE3D" blockId="10.[151,1436,151,573]" pageId="10" pageNumber="311">
<emphasis id="B935EAEA974BFFD8FF65E0D0FE44FE65" bold="true" box="[199,466,404,429]" pageId="10" pageNumber="311">Queen measurements.</emphasis>
Three non-types (n = 3; 1 alate and 2 dealate): TL 2.212.43; HL 0.480.49; HW 0.440.45; SL 0.180.20; ML 0.700.72; MsW 0.360.40; EL 0.150.16; EW 0.090.10; CI 9294; REL 3336; SI 4045.
</paragraph>
<paragraph id="8BFE36F8974BFFD8FF65E0BBFBFCFDF5" blockId="10.[151,1436,151,573]" pageId="10" pageNumber="311">
<emphasis id="B935EAEA974BFFD8FF65E0BBFE57FDD0" bold="true" box="[199,449,511,536]" pageId="10" pageNumber="311">Male measurements.</emphasis>
Two non-types (n = 2): TL 1.981.99; HL 0.390.41; HW 0.380.40; SL 0.06; ML 0.660.70; MsW 0.400.44; EL 0.180.20; EW 0.14; CI 9798; REL 4553; SI 1516.
</paragraph>
<caption id="DF3E6670974BFFD8FF35E7ECFD84F916" httpUri="https://zenodo.org/record/2615341/files/figure.png" pageId="10" pageNumber="311" startId="10.[151,264,1704,1727]" targetBox="[151,1436,625,1684]" targetPageId="10">
<paragraph id="8BFE36F8974BFFD8FF35E7ECFD84F916" blockId="10.[151,1436,1704,1758]" pageId="10" pageNumber="311">
<emphasis id="B935EAEA974BFFD8FF35E7ECFED0F976" bold="true" box="[151,326,1704,1727]" pageId="10" pageNumber="311">FIGURES 811.</emphasis>
Lectotype worker of
<taxonomicName id="4C414D7B974BFFD8FD86E7EEFD10F977" box="[548,646,1706,1727]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="B935EAEA974BFFD8FD86E7EEFD10F977" box="[548,646,1706,1727]" italics="true" pageId="10" pageNumber="311">R. javana</emphasis>
</taxonomicName>
<emphasis id="B935EAEA974BFFD8FD2EE7EDFD40F976" bold="true" box="[652,726,1705,1726]" pageId="10" pageNumber="311">stat. n.</emphasis>
8, habitus in profile; 9, head in full-face view; 10, closeup of head in profile view; 11, body in dorsal view.
</paragraph>
</caption>
<paragraph id="8BFE36F8974BFFD9FF65E641FB40FE88" blockId="10.[151,1437,1797,2038]" lastBlockId="11.[151,1436,151,789]" lastPageId="11" lastPageNumber="312" pageId="10" pageNumber="311">
<emphasis id="B935EAEA974BFFD8FF65E641FE4FF8D6" bold="true" box="[199,473,1797,1822]" pageId="10" pageNumber="311">Redescription. Worker</emphasis>
(
<typeStatus id="54FA885A974BFFD8FE4BE641FDC2F8D6" box="[489,596,1797,1822]" pageId="10" pageNumber="311" type="lectotype">lectotype</typeStatus>
,
<typeStatus id="54FA885A974BFFD8FDC2E641FC94F8D6" box="[608,770,1797,1822]" pageId="10" pageNumber="311" type="paralectotype">paralectotypes</typeStatus>
and non-types).Workers monomorphic with little variation in size. Head in full face view subrectangular, longer than wide, posterior margin with a shallow median concavity, posterolateral corners roundly convex, lateral margins of head broadly convex and weakly converging anteriorlyanterior clypeal margin less wide than posterior margin of head. Eye with 1116 ommatidia. In profile view, outline of clypeus evenly and weakly convex, projecting slightly forward from dorsal margin of head. Mesosoma in profile box-shaped, subcylindrical, dorsal outline weakly convex and almost straight, propodeal declivity steep and shallowly concave. In dorsal view, mesosoma subrectangular, anterior pronotal margin broadly convex, sometimes angulate at its median point, humeral angles rounded, posterior propodeal margin weakly concave. In profile, petiole inclined posteriorly, slightly longer than high, apex distinctly narrower than base, anterior margin broadly concave; petiolar node bell-shaped, apex rounded and smooth, posterior face nearly straight and almost as steep as anterior face; petiole higher and longer than postpetiole, dorsal margin of postpetiole weakly convex and nearly flat. In dorsal view, petiole suboval, wider than long; postpetiole globular and larger than petiole.
</paragraph>
<paragraph id="8BFE36F8974AFFD9FF65E009FBA5FDD0" blockId="11.[151,1436,151,789]" pageId="11" pageNumber="312">Dorsum and lateral surface of head finely striated and shining, posterolateral corners also feebly reticulate, interspaces smooth and shining; posterior-most portion of vertex of head just above occiput in posterodorsal view largely unsculptured, smooth and shiny. Ventral surface of posterior half of head mostly superficially reticulate with smooth and shining interspaces. Mandible mostly smooth and shining, area close to masticatory margin punctate. Lateral surface of mesosoma striate-reticulate and shining; dorsum of mesosoma striated. Petiole, postpetiole and gaster superficially reticulate with smooth and shining interspaces.</paragraph>
<paragraph id="8BFE36F8974AFFD9FF65E360FCE2FCDD" blockId="11.[151,1436,151,789]" pageId="11" pageNumber="312">Dorsal and ventral surfaces of head with abundant short standing and suberect hairs interspersed by sparse longer erect hairs, gena with abundant short appressed hairs. Mesosoma dorsum with sparse, scattered short standing hairs and sparse longer erect hairs close to dorsolateral margin. Anterior rising face of petiole with uniformly short appressed hairs, posterior declivity covered with short erect and subdecumbent hairs; apex of petiolar node with short erect hairs and 12 pairs of much longer erect hairs. Postpetiole with abundant short erect and suberect hairs, and 1 pair of long erect hairs. Gaster very pilose, all surfaces covered with abundant short suberect and erect hairs, interspersed by sparse longer erect hairs.</paragraph>
<caption id="DF3E6670974AFFD9FF35E62BFBCEF86C" httpUri="https://zenodo.org/record/2615345/files/figure.png" pageId="11" pageNumber="312" startId="11.[151,264,1903,1926]" targetBox="[151,1436,833,1892]" targetPageId="11">
<paragraph id="8BFE36F8974AFFD9FF35E62BFBCEF86C" blockId="11.[151,1436,1903,1956]" pageId="11" pageNumber="312">
<emphasis id="B935EAEA974AFFD9FF35E62BFEC2F84C" bold="true" box="[151,340,1903,1926]" pageId="11" pageNumber="312">FIGURES 1215.</emphasis>
Non-type worker of
<taxonomicName id="4C414D7B974AFFD9FD88E635FD1DF84E" box="[554,651,1905,1926]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="B935EAEA974AFFD9FD88E635FD1DF84E" box="[554,651,1905,1926]" italics="true" pageId="11" pageNumber="312">R. javana</emphasis>
</taxonomicName>
<emphasis id="B935EAEA974AFFD9FD30E634FD4CF84D" bold="true" box="[658,730,1904,1925]" pageId="11" pageNumber="312">stat. n.</emphasis>
from Singapore, Mandai Track, ZRC_HYM_0000576; 12, habitus in profile; 13, head in full face view; 14, closeup of head in profile view; 15, body in dorsal view.
</paragraph>
</caption>
<caption id="DF3E6670974DFFDEFF35E5CBFB61FB0C" httpUri="https://zenodo.org/record/2615347/files/figure.png" pageId="12" pageNumber="313" startId="12.[151,264,1167,1190]" targetBox="[151,1436,193,1146]" targetPageId="12">
<paragraph id="8BFE36F8974DFFDEFF35E5CBFB61FB0C" blockId="12.[151,1435,1167,1220]" pageId="12" pageNumber="313">
<emphasis id="B935EAEA974DFFDEFF35E5CBFEC1FB6C" bold="true" box="[151,343,1167,1190]" pageId="12" pageNumber="313">FIGURES 1619.</emphasis>
Non-type queen of
<taxonomicName id="4C414D7B974DFFDEFD8DE5D5FD02FB6E" box="[559,660,1169,1190]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="B935EAEA974DFFDEFD8DE5D5FD02FB6E" box="[559,660,1169,1190]" italics="true" pageId="12" pageNumber="313">R. javana</emphasis>
</taxonomicName>
from Singapore, Bukit Timah Nature Reserve, ZRC_HYM_0001732; 16, habitus in profile; 17, head in full face view; 18, body in dorsal view; 19, closeup of mesosoma in dorsal view.
</paragraph>
</caption>
<paragraph id="8BFE36F8974DFFDEFF65E5B4FDABFAE5" blockId="12.[151,1437,1264,2010]" pageId="12" pageNumber="313">Head, dorsum of mesosoma and gaster generally darker brown in colour, rest of body more yellowish brown, tips of antennae and legs pale in tone.</paragraph>
<paragraph id="8BFE36F8974DFFDEFF65E47CFE0EF8C9" blockId="12.[151,1437,1264,2010]" pageId="12" pageNumber="313">
<emphasis id="B935EAEA974DFFDEFF65E47CFE83FA99" bold="true" box="[199,277,1336,1361]" pageId="12" pageNumber="313">Queen</emphasis>
(non-types). Small-medium size. Head in full face view subrectangular, slightly less rectangular than that of worker, longer than wide; lateral margins almost entirely straight and parallel to each other, posterolateral corners smooth and rounded, posterior margin nearly straight with a shallow median concavity. Eye large (&gt;100 ommatidia); posterior margin of eye located slightly below midline of head in full face view. In profile, clypeus very weakly convex, almost entirely flat. Scape short, not exceeding midline of head when lain back. In profile, mesosoma sub-cylindrical, more elongate than that of worker, dorsal outline weakly and broadly convex, propodeal junction rounded and smooth, posterior face of propodeum steep and almost vertical, shallowly concave. In dorsal view, mesosoma sub-cylindrical, anterior pronotal margin roundly convex, humeral angles distinctly obtuse though rounded and smooth, mesoscutum wider than long. In profile view, petiole inclined posteriorly, longer than high, anterior margin weakly concave; petiolar node bell-shaped, apex roundly convex, posterior face steeper than anterior face; dorsal margin of postpetiole weakly convex, not as long as postpetiole, dorsal margin weakly convex. In dorsal view, petiole suboval, less than twice as wide as long; postpetiole globular, much larger and wider than petiole.
</paragraph>
<paragraph id="8BFE36F8974DFFDFFF65E649FE91FF1D" blockId="12.[151,1437,1264,2010]" lastBlockId="13.[151,1437,151,2013]" lastPageId="13" lastPageNumber="314" pageId="12" pageNumber="313">Posterior half of head mostly smooth and shining, feeble striations on frons and venter, anterior half of head finely striated and shiny, clypeus striated but with a median spot that is smooth and shining. Mandible mostly smooth and shining, area next to masticatory margin slightly striate-punctate Lateral surface of mesosoma weakly substriate-reticulate, interspaces smooth and shining. Dorsum of mesosoma mostly striated and shining, median section of pronotum superficially reticulate with smooth and shining interspaces. Lateral surfaces of petiole and postpetiole reticulate, interspaces smooth and shining; apices of petiole and postpetiole feebly and superficially reticulate, mostly smooth and shining. Gaster largely smooth and shining, partly superficially and weakly reticulate.</paragraph>
<paragraph id="8BFE36F8974CFFDFFF65E1A4FDFDFE65" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="314">Dorsum and ventral surface of head covered with abundant short erect and suberect hairs, with sparse longer erect hairs. Dorsum of mesosoma with sparse short standing hairs and sparser long erect hairs, mainly along dorsolateral margin. Anterior face of petiole with short appressed and/or decumbent hairs, posterior face with multiple short erect hairs, apex with short erect hairs, 12 pairs of longer erect hairs; postpetiole with multiple short standing hairs and a pair of longer erect hairs. Gaster pilose, with abundant short appressed hairs, scattered short standing hairs and sparse long erect hairs.</paragraph>
<paragraph id="8BFE36F8974CFFDFFF65E0F3FD17FE3D" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="314">Entire body generally uniform dark brown; head, dorsum of mesosoma and gaster slightly darker brown; tibia and tarsus paler and more yellowish brown.</paragraph>
<paragraph id="8BFE36F8974CFFDFFF65E0BBFE22FC48" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="314">
<emphasis id="B935EAEA974CFFDFFF65E0BBFE92FDD0" bold="true" box="[199,260,511,536]" pageId="13" pageNumber="314">Male</emphasis>
(non-types). Small size. Head in full face view almost entirely spherical, posterior margin broad and round; eye relatively smaller as compared to other species, maximum diameter of eye only slightly exceeding midline of head, outline of eye exceeding lateral margin of head by a little. In profile, head broad-ovate, lateral corners smooth and rounded, ventral and posterior margins forming a continuous curved outline; clypeus roundly convex and strongly projecting from dorsum of head. In profile, mesosoma sub-oblong, dorsal margin almost completely straight, propodeal junction roundly convex, posterior face of propodeum steep and almost entirely vertical. In dorsal view, mesosoma obovate, notauli absent. In profile view, petiole inclined posteriorly, short and subtrapezoidal, longer than high, anterior margin weakly concave, apex smooth and blunt, weakly convex and sloping downwards gently, posterior margin very short; anteroventral extension of subpetiolar process shallow and subtriangular, longer at base than apex. Dorsal outline of postpetiole weakly convex, differentiated from gaster by a weak but distinct cinctus.
</paragraph>
<paragraph id="8BFE36F8974CFFDFFF65E2C8FE8DFBFD" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="314">Entire head mostly substriate-reticulate, with smooth and shining interspaces and scattered punctures; clypeus rugulose with punctured interspaces and weakly shining. Lateral surface of mesosoma weakly striate-reticulate, interspaces smooth and shining; dorsum largely striated and shining, propodeum superficially reticulate with smooth and shiny interspaces. Petiole, postpetiole and gaster superficially reticulate with smooth and shiny interspaces.</paragraph>
<paragraph id="8BFE36F8974CFFDFFF65E504FC44FB68" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="314">Dorsum of head with abundant short standing hairs, lateral and ventral surfaces with numerous short decumbent and appressed hairs. Dorsal surfaces of mesosoma, petiole, postpetiole and gaster mostly with abundant short standing hairs, propodeum with visibly much sparser standing hairs.</paragraph>
<paragraph id="8BFE36F8974CFFDFFF65E5E8FCE5FB20" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="314">Generally dark greyish-brown; head, dorsum of mesosoma and gaster darker blackish-brown; funiculi of antennae light brown; joints of legs, tarsus pale brownish-yellow.</paragraph>
</subSubSection>
<subSubSection id="C35B6573974CFFDFFF65E5B0FDB4FA55" pageId="13" pageNumber="314" type="distribution">
<paragraph id="8BFE36F8974CFFDFFF65E5B0FC41FAC5" blockId="13.[151,1437,151,2013]" box="[199,983,1268,1293]" pageId="13" pageNumber="314">
<emphasis id="B935EAEA974CFFDFFF65E5B0FEF6FAC5" bold="true" box="[199,352,1268,1293]" pageId="13" pageNumber="314">Distribution.</emphasis>
<collectingCountry id="F3567668974CFFDFFEC5E5B0FE42FAC5" box="[359,468,1268,1293]" name="Indonesia" pageId="13" pageNumber="314">Indonesia</collectingCountry>
(eastern
<collectingRegion id="4985F81A974CFFDFFD99E5B0FDE7FAC4" box="[571,625,1268,1292]" country="Indonesia" name="Jawa Timur" pageId="13" pageNumber="314">Java</collectingRegion>
),
<collectingCountry id="F3567668974CFFDFFD26E5B0FD6FFAC5" box="[644,761,1268,1293]" name="Singapore" pageId="13" pageNumber="314">Singapore</collectingCountry>
,
<collectingCountry id="F3567668974CFFDFFCA1E5B0FCF1FAC5" box="[771,871,1268,1293]" name="Thailand" pageId="13" pageNumber="314">Thailand</collectingCountry>
(
<figureCitation id="137A2A7D974CFFDFFCD4E5B0FC5CFAC5" box="[886,970,1268,1293]" captionStart="FIGURE 76" captionStartId="36.[151,250,1737,1759]" captionTargetBox="[151,1436,543,1684]" captionTargetId="figure@36.[151,1436,543,1684]" captionTargetPageId="36" captionText="FIGURE 76. Distribution of R. javana and R. johorensis in Southeast Asia." httpUri="https://zenodo.org/record/2615385/files/figure.png" pageId="13" pageNumber="314">Fig. 76</figureCitation>
).
</paragraph>
<paragraph id="8BFE36F8974CFFDFFF65E453FDB4FA55" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="314">
<emphasis id="B935EAEA974CFFDFFF65E453FEBCFAF8" bold="true" box="[199,298,1303,1328]" pageId="13" pageNumber="314">Habitat.</emphasis>
Non-type specimens from
<collectingCountry id="F3567668974CFFDFFDD9E453FD7AFAF8" box="[635,748,1303,1328]" name="Singapore" pageId="13" pageNumber="314">Singapore</collectingCountry>
were collected from bark of common native trees, e.g.
<taxonomicName id="4C414D7B974CFFDFFF35E478FD31FA9D" authority="(Blume) Hook." authorityName="Hook." baseAuthorityName="Blume" box="[151,679,1340,1365]" class="Magnoliopsida" family="Anacardiaceae" genus="Campnosperma" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="13" pageNumber="314" phylum="Tracheophyta" rank="species" species="auriculatum">
<emphasis id="B935EAEA974CFFDFFF35E478FE77FA9C" box="[151,481,1340,1364]" italics="true" pageId="13" pageNumber="314">Campnosperma auriculatum</emphasis>
(Blume) Hook.
</taxonomicName>
f., in secondary forests and nature reserves which used to be abandoned plantations; specimens from
<collectingCountry id="F3567668974CFFDFFDFEE41BFD56FAB0" box="[604,704,1375,1400]" name="Thailand" pageId="13" pageNumber="314">Thailand</collectingCountry>
were collected from bark of mango trees. Nests do not appear to occupy large expanses of tree bark.
</paragraph>
</subSubSection>
<subSubSection id="C35B6573974CFFDDFF65E4E3FE5AFED5" lastPageId="15" lastPageNumber="316" pageId="13" pageNumber="314" type="discussion">
<paragraph id="8BFE36F8974CFFDFFF65E4E3FAC8F928" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="314">
<emphasis id="B935EAEA974CFFDFFF65E4E3FEADFA08" bold="true" box="[199,315,1447,1472]" pageId="13" pageNumber="314">Remarks.</emphasis>
We compared the queen specimens of
<taxonomicName id="4C414D7B974CFFDFFCA4E4EDFCE3FA77" box="[774,885,1448,1472]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="B935EAEA974CFFDFFCA4E4EDFCE3FA77" box="[774,885,1448,1472]" italics="true" pageId="13" pageNumber="314">R. javana</emphasis>
</taxonomicName>
from
<collectingCountry id="F3567668974CFFDFFC62E4E3FBA4FA08" box="[960,1074,1447,1472]" name="Singapore" pageId="13" pageNumber="314">Singapore</collectingCountry>
with the
<typeStatus id="54FA885A974CFFDFFB07E4E3FA91FA08" box="[1189,1287,1447,1472]" pageId="13" pageNumber="314" type="holotype">holotype</typeStatus>
queen (head missing) of
<taxonomicName id="4C414D7B974CFFDFFEBCE488FDBFFA2C" authorityName="Forel" authorityYear="1900" box="[286,553,1484,1508]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="rothneyi">
<emphasis id="B935EAEA974CFFDFFEBCE488FDBFFA2C" box="[286,553,1484,1508]" italics="true" pageId="13" pageNumber="314">Rhopalomastix rothneyi</emphasis>
</taxonomicName>
from
<collectingCountry id="F3567668974CFFDFFDCDE488FD3AFA2D" box="[623,684,1484,1509]" name="India" pageId="13" pageNumber="314">India</collectingCountry>
. We found the
<collectingCountry id="F3567668974CFFDFFCF4E488FC5EFA2D" box="[854,968,1484,1509]" name="Singapore" pageId="13" pageNumber="314">Singapore</collectingCountry>
queen specimens of
<taxonomicName id="4C414D7B974CFFDFFB17E489FAB7FA2C" box="[1205,1313,1485,1508]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="B935EAEA974CFFDFFB17E489FAB7FA2C" box="[1205,1313,1485,1508]" italics="true" pageId="13" pageNumber="314">R. javana</emphasis>
</taxonomicName>
(SQJV) to be quite different from the
<typeStatus id="54FA885A974CFFDFFE65E4ABFDBFF9C0" box="[455,553,1519,1544]" pageId="13" pageNumber="314" type="holotype">holotype</typeStatus>
queen of
<taxonomicName id="4C414D7B974CFFDFFD3BE4B5FC83F9C0" authorityName="Forel" authorityYear="1900" box="[665,789,1520,1544]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="rothneyi">
<emphasis id="B935EAEA974CFFDFFD3BE4B5FC83F9C0" box="[665,789,1520,1544]" italics="true" pageId="13" pageNumber="314">R. rothneyi</emphasis>
</taxonomicName>
(QR): 1) in SQJV, pronotal length in dorsal view is more than half of propodeal length; in QR, the pronotal length is less than half of propodeal length; 2) mesoscutellar disc of SQJV less than twice as wide as long; in QR, mesoscutellar disc more than twice as wide as long; 3) SQJV propodeum in dorsal view more than twice as wide as long, with broadly concave posterior margin; in QR, propodeum less than twice as wide as long, with roundly convex posterior margin; 4) anterior margin of petiole of SQJV broadly concave, posterior margin steep and nearly flat; in QR, anterior margin not distinctly concave, as steep and flat as posterior margin; 5) SQJV is generally smaller (MsW
<quantity id="4CB99B1D974CFFDFFC11E783FBD6F928" box="[947,1088,1735,1760]" metricMagnitude="-3" metricUnit="m" metricValue="9.652" metricValueMax="10.16" metricValueMin="9.144" pageId="13" pageNumber="314" unit="in" value="0.38" valueMax="0.4" valueMin="0.36">0.360.40 in</quantity>
SQJV, MsW
<quantity id="4CB99B1D974CFFDFFB7BE78CFAB1F928" box="[1241,1319,1735,1760]" metricMagnitude="-2" metricUnit="m" metricValue="1.0668" pageId="13" pageNumber="314" unit="in" value="0.42">0.42 in</quantity>
QR)
</paragraph>
<paragraph id="8BFE36F8974CFFDCFF65E7A8FBECFF1D" blockId="13.[151,1437,151,2013]" lastBlockId="14.[151,1437,151,1040]" lastPageId="14" lastPageNumber="315" pageId="13" pageNumber="314">
Three workers, two males on two pins each with typus labels (MHNG) and
<specimenCount id="9D47FD71974CFFDFFBD4E7A8FB6FF8CD" box="[1142,1273,1772,1797]" pageId="13" pageNumber="314" type="worker">23 workers</specimenCount>
on pins with cotypus labels of
<taxonomicName id="4C414D7B974CFFDFFECFE655FE7EF8E0" authorityName="Forel" authorityYear="1900" box="[365,488,1808,1832]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="rothneyi">
<emphasis id="B935EAEA974CFFDFFECFE655FE7EF8E0" box="[365,488,1808,1832]" italics="true" pageId="13" pageNumber="314">R. rothneyi</emphasis>
</taxonomicName>
(MHNG) from Peradineya [sic],
<collectingCountry id="F3567668974CFFDFFCFFE64BFC5CF8E0" box="[861,970,1807,1832]" name="Sri Lanka" pageId="13" pageNumber="314">Sri Lanka</collectingCountry>
were also examined. Although labeled as such, they are not type specimens of
<taxonomicName id="4C414D7B974CFFDFFD99E671FD2EF884" authorityName="Forel" authorityYear="1900" box="[571,696,1844,1868]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="rothneyi">
<emphasis id="B935EAEA974CFFDFFD99E671FD2EF884" box="[571,696,1844,1868]" italics="true" pageId="13" pageNumber="314">R. rothneyi</emphasis>
</taxonomicName>
. We compared workers of the Sri Lankan specimens (SLW) with the
<typeStatus id="54FA885A974CFFDFFF62E613FEB0F8B8" box="[192,294,1879,1904]" pageId="13" pageNumber="314" type="lectotype">lectotype</typeStatus>
and
<typeStatus id="54FA885A974CFFDFFEFCE613FD96F8B8" box="[350,512,1879,1904]" pageId="13" pageNumber="314" type="paralectotype">paralectotypes</typeStatus>
of
<taxonomicName id="4C414D7B974CFFDFFD87E61DFD06F8A7" box="[549,656,1880,1904]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="B935EAEA974CFFDFFD87E61DFD06F8A7" box="[549,656,1880,1904]" italics="true" pageId="13" pageNumber="314">R. javana</emphasis>
</taxonomicName>
from Java (JVW). We found the following differences between SLW and JVW: 1) for SLW, straight-line distance of eye to mandibular insertion in profile view ± maximum diameter of eye; in JVW, distance of eye to mandibular insertion&gt; maximum diameter of eye; 2) in JVW, mesosoma relatively short in proportion to head length (HL/ML 8991); in SLW, mesosoma slightly more elongate (HL/ML 8387); 3) in JVW, slope of the propodeal declivity is more deeply concave, with a distinctly concave posterior propodeal margin in dorsal view; for SLW, slope of the propodeal declivity is steep and mostly flat.
</paragraph>
<paragraph id="8BFE36F8974FFFDCFF65E19BFE35FE65" blockId="14.[151,1437,151,1040]" pageId="14" pageNumber="315">
The male of
<taxonomicName id="4C414D7B974FFFDCFEF1E1A5FE2BFF3F" box="[339,445,224,248]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="B935EAEA974FFFDCFEF1E1A5FE2BFF3F" box="[339,445,224,248]" italics="true" pageId="14" pageNumber="315">R. javana</emphasis>
</taxonomicName>
from
<collectingCountry id="F3567668974FFFDCFDA3E19BFDE4FF30" box="[513,626,223,248]" name="Singapore" pageId="14" pageNumber="315">Singapore</collectingCountry>
(SMJV) differs from the Sri Lankan male (SLM) by the following traits: 1) SMJV has a more orbicular head, broadly and evenly rounded posteriorly in full face view; SLM has a more longitudinally oval head, with posterior margin more elongate; 2) SMJV is smaller (SMJV HL 0.390.41, HW 0.380.40, MsW 0.40.44; SLM HL 0.44, HW 0.400.42, MsW 0.480.5); 3) mesoscutum of SMJV in profile view roughly as high as pronotum; in SLM, mesoscutum distinctly much higher than pronotum, with a roundly convex anterior margin.
</paragraph>
<paragraph id="8BFE36F8974FFFDCFF65E0F3FEB9FDA8" blockId="14.[151,1437,151,1040]" pageId="14" pageNumber="315">
Given the character differences between
<taxonomicName id="4C414D7B974FFFDCFD3EE0FDFC8FFE18" authorityName="Forel" authorityYear="1900" box="[668,793,440,464]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="rothneyi">
<emphasis id="B935EAEA974FFFDCFD3EE0FDFC8FFE18" box="[668,793,440,464]" italics="true" pageId="14" pageNumber="315">R. rothneyi</emphasis>
</taxonomicName>
and both the types and the
<collectingCountry id="F3567668974FFFDCFBFFE0F3FB59FE18" box="[1117,1231,439,464]" name="Singapore" pageId="14" pageNumber="315">Singapore</collectingCountry>
populations of
<taxonomicName id="4C414D7B974FFFDCFA26E0FDFF75FE3C" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="B935EAEA974FFFDCFA26E0FDFF75FE3C" italics="true" pageId="14" pageNumber="315">R. javana</emphasis>
</taxonomicName>
, especially those between the
<typeStatus id="54FA885A974FFFDCFDE9E098FD3BFE3D" box="[587,685,476,501]" pageId="14" pageNumber="315" type="holotype">holotype</typeStatus>
queen of
<taxonomicName id="4C414D7B974FFFDCFC8AE099FC31FE3C" authorityName="Forel" authorityYear="1900" box="[808,935,476,500]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="rothneyi">
<emphasis id="B935EAEA974FFFDCFC8AE099FC31FE3C" box="[808,935,476,500]" italics="true" pageId="14" pageNumber="315">R. rothneyi</emphasis>
</taxonomicName>
and SQJV, we thereby choose to raise the subspecies
<taxonomicName id="4C414D7B974FFFDCFEBAE344FEF2FDDF" box="[280,356,512,535]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="B935EAEA974FFFDCFEBAE344FEF2FDDF" box="[280,356,512,535]" italics="true" pageId="14" pageNumber="315">javana</emphasis>
</taxonomicName>
to species status. There remains the strong possibility of future revision of
<taxonomicName id="4C414D7B974FFFDCFB6BE345FAD0FDD0" authorityName="Forel" authorityYear="1900" box="[1225,1350,512,536]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="rothneyi">
<emphasis id="B935EAEA974FFFDCFB6BE345FAD0FDD0" box="[1225,1350,512,536]" italics="true" pageId="14" pageNumber="315">R. rothneyi</emphasis>
</taxonomicName>
and its previous subspecies, when fresh material from across the species range is available for DNA/molecular investigation.
</paragraph>
<paragraph id="8BFE36F8974FFFDCFF65E328FD77FBD8" blockId="14.[151,1437,151,1040]" pageId="14" pageNumber="315">
The
<typeStatus id="54FA885A974FFFDCFF5EE328FEF5FD4D" box="[252,355,620,645]" pageId="14" pageNumber="315" type="lectotype">lectotype</typeStatus>
and
<typeStatus id="54FA885A974FFFDCFE3FE328FDA9FD4D" box="[413,575,620,645]" pageId="14" pageNumber="315" type="paralectotype">paralectotypes</typeStatus>
of
<taxonomicName id="4C414D7B974FFFDCFDCBE329FD4EFD4C" box="[617,728,621,644]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="B935EAEA974FFFDCFDCBE329FD4EFD4C" box="[617,728,621,644]" italics="true" pageId="14" pageNumber="315">R. javana</emphasis>
</taxonomicName>
(JVW) are morphologically similar to the
<collectingCountry id="F3567668974FFFDCFB67E328FAA1FD4D" box="[1221,1335,620,645]" name="Singapore" pageId="14" pageNumber="315">Singapore</collectingCountry>
(SWJV) and Thai (TWJV) specimens, except for a few minor differences: 1) JVW is overall uniformly yellowish brown with pale yellow antennal tips, but SWJV and TWJV are in general not uniform in colour, with head and mesosoma usually darker brown than rest of body, antennal tips and legs paler yellowish brown; 2) eye size of JVW relatively smaller in proportion to head width compared to SWJV and TWJV (JVW: REL 1718; SWJV, TWJV: REL 18 23). On the other hand, the heads of JVW and TWJV are generally larger than that of SWJV (JVW, TWJV: HL 0.480.52, HW 0.450.48; SWJV: HL 0.460.48; HW 0.410.44). These minor differences may simply be morphological variation or polyphenism among geographically distant allopatric populations. For now, given the lack of DNA/molecular evidence and the minor morphological differences, we here choose to treat the
<collectingCountry id="F3567668974FFFDCFA89E2C8FA0BFC6D" box="[1323,1437,908,933]" name="Singapore" pageId="14" pageNumber="315">Singapore</collectingCountry>
and Thai populations as
<emphasis id="B935EAEA974FFFDCFE0BE2F5FD8DFC0F" box="[425,539,944,968]" italics="true" pageId="14" pageNumber="315">
<taxonomicName id="4C414D7B974FFFDCFE0BE2F5FD81FC0F" box="[425,535,944,968]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="javana">R. javana</taxonomicName>
,
</emphasis>
until fresh specimens of
<taxonomicName id="4C414D7B974FFFDCFC94E2F5FC34FC0F" box="[822,930,944,968]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="B935EAEA974FFFDCFC94E2F5FC34FC0F" box="[822,930,944,968]" italics="true" pageId="14" pageNumber="315">R. javana</emphasis>
</taxonomicName>
can be obtained from its type locality (Java), and more compelling DNA information from different genetic markers (nuclear and mitochondrial) is available for comparison with the
<collectingCountry id="F3567668974FFFDCFE21E2B3FE62FBD8" box="[387,500,1015,1040]" name="Singapore" pageId="14" pageNumber="315">Singapore</collectingCountry>
and Thai specimens.
</paragraph>
<caption id="DF3E6670974FFFDCFF35E6FBFB80F83C" httpUri="https://zenodo.org/record/2615349/files/figure.png" pageId="14" pageNumber="315" startId="14.[151,264,1983,2006]" targetBox="[226,1360,1088,1975]" targetPageId="14">
<paragraph id="8BFE36F8974FFFDCFF35E6FBFB80F83C" blockId="14.[151,1435,1983,2036]" pageId="14" pageNumber="315">
<emphasis id="B935EAEA974FFFDCFF35E6FBFEC5F81D" bold="true" box="[151,339,1983,2006]" pageId="14" pageNumber="315">FIGURES 2023.</emphasis>
Non-type male of
<taxonomicName id="4C414D7B974FFFDCFDB2E685FDE7F81E" box="[528,625,1985,2006]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="B935EAEA974FFFDCFDB2E685FDE7F81E" box="[528,625,1985,2006]" italics="true" pageId="14" pageNumber="315">R. javana</emphasis>
</taxonomicName>
from Singapore, Mandai track, ZRC_HYM_0000576; 20, habitus in profile; 21, head in full face view; 22, body in dorsal view; 23, closeup of mesosoma in dorsal view.
</paragraph>
</caption>
<paragraph id="8BFE36F8974EFFDDFF65E1D3FE5AFED5" blockId="15.[151,1437,151,788]" pageId="15" pageNumber="316">
Workers of
<taxonomicName id="4C414D7B974EFFDDFEECE1DDFE2AFF67" box="[334,444,152,176]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="B935EAEA974EFFDDFEECE1DDFE2AFF67" box="[334,444,152,176]" italics="true" pageId="15" pageNumber="316">R. javana</emphasis>
</taxonomicName>
are similar to small workers of
<taxonomicName id="4C414D7B974EFFDDFC8EE1DDFC57FF78" box="[812,961,152,176]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="johorensis">
<emphasis id="B935EAEA974EFFDDFC8EE1DDFC57FF78" box="[812,961,152,176]" italics="true" pageId="15" pageNumber="316">R. johorensis</emphasis>
</taxonomicName>
, but may be differentiated from the latter based on characters indicated in couplet (5) of the species key. Workers of
<taxonomicName id="4C414D7B974EFFDDFBABE1F9FBEEFF1C" box="[1033,1144,189,212]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="B935EAEA974EFFDDFBABE1F9FBEEFF1C" box="[1033,1144,189,212]" italics="true" pageId="15" pageNumber="316">R. javana</emphasis>
</taxonomicName>
are also similar to small workers of
<taxonomicName id="4C414D7B974EFFDDFEBFE1A5FE02FF30" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2018" box="[285,404,224,248]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="tenebra">
<emphasis id="B935EAEA974EFFDDFEBFE1A5FE02FF30" box="[285,404,224,248]" italics="true" pageId="15" pageNumber="316">R. tenebra</emphasis>
</taxonomicName>
, but can be distinguished from the latter based on characters as explained in “Remarks” under the latter description.
</paragraph>
</subSubSection>
<subSubSection id="C35B6573974EFFDDFF65E063FD9FFCDC" pageId="15" pageNumber="316" type="materials_examined">
<paragraph id="8BFE36F8974EFFDDFF65E063FA02FDF4" blockId="15.[151,1437,151,788]" pageId="15" pageNumber="316">
<emphasis id="B935EAEA974EFFDDFF65E063FDA6FE88" bold="true" box="[199,560,295,320]" pageId="15" pageNumber="316">Non-type material examined.</emphasis>
<materialsCitation id="3B293CA5974EFFDDFD9FE06CFDE8FEAD" ID-GBIF-Occurrence="2118818653" collectingDate="2017-01-04" collectorName="G. W. Yong" country="Singapore" latitude="1.40337" location="Mandai Track" longLatPrecision="1" longitude="103.77824" pageId="15" pageNumber="316" specimenCount="20" specimenCount-male="1" specimenCount-queen="1" specimenCount-worker="18">
<collectingCountry id="F3567668974EFFDDFD9FE06CFD49FE88" box="[573,735,296,320]" name="Singapore" pageId="15" pageNumber="316">SINGAPORE</collectingCountry>
:
<specimenCount id="9D47FD71974EFFDDFD4DE06CFCE0FE88" box="[751,886,295,320]" pageId="15" pageNumber="316" type="worker">18 workers</specimenCount>
,
<specimenCount id="9D47FD71974EFFDDFC25E06CFC7FFEF7" box="[903,1001,296,320]" pageId="15" pageNumber="316" type="queen">1 queen</specimenCount>
,
<specimenCount id="9D47FD71974EFFDDFC58E06CFBD9FE88" box="[1018,1103,295,320]" pageId="15" pageNumber="316" type="male">1 male</specimenCount>
,
<location id="8E9E6023974EFFDDFBC2E063FA92FE88" LSID="urn:lsid:plazi:treatment:03E887EE9748FFDDFF35E5EFFD9FFCDC:8E9E6023974EFFDDFBC2E063FA92FE88" box="[1120,1284,295,320]" country="Singapore" latitude="1.40337" longLatPrecision="1" longitude="103.77824" name="Mandai Track" pageId="15" pageNumber="316">Mandai Track</location>
(
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,
<geoCoordinate id="EE75503F974EFFDDFF35E008FEA2FEAC" box="[151,308,332,356]" direction="east" orientation="longitude" pageId="15" pageNumber="316" precision="1" value="103.77824">103.77824° E</geoCoordinate>
),
<date id="FFFF1038974EFFDDFEEBE008FE50FEAC" box="[329,454,332,356]" pageId="15" pageNumber="316" value="2017-01-04">
<collectingDate id="EFBBE9D0974EFFDDFEEBE008FE50FEAC" box="[329,454,332,356]" pageId="15" pageNumber="316" value="2017-01-04">4 Jan 2017</collectingDate>
</date>
,
<collectorName id="26B4532E974EFFDDFE70E008FDDDFEAD" box="[466,587,332,357]" pageId="15" pageNumber="316">G.W. Yong</collectorName>
leg.
</materialsCitation>
,
<materialsCitation id="3B293CA5974EFFDDFD28E008FC70FE64" ID-GBIF-Occurrence="2118815106" collectingDate="2017-06-21" collectionCode="ZRC, THNHM" collectorName="C. Peeters" country="Singapore" latitude="1.35449" location="Bukit Timah Nature Reserve" longLatPrecision="1" longitude="103.78211" pageId="15" pageNumber="316" specimenCode="HYM000576" specimenCount="12" specimenCount-worker="12">
colony no. GY-SG17-RhoN,
<collectionCode id="ED50AE3D974EFFDDFC73E008FB9EFEAD" box="[977,1032,332,357]" country="Singapore" name="Zoological Reference Collection, National University of Singapore" pageId="15" pageNumber="316">ZRC</collectionCode>
_
<specimenCode id="DBE79E83974EFFDDFBB4E008FB3DFEAD" box="[1046,1195,332,357]" collectionCode="ZRC" country="Singapore" name="Zoological Reference Collection, National University of Singapore" pageId="15" pageNumber="316">HYM000576</specimenCode>
(
<collectionCode id="ED50AE3D974EFFDDFB18E008FAB3FEAC" box="[1210,1317,332,356]" pageId="15" pageNumber="316">THNHM</collectionCode>
,
<collectionCode id="ED50AE3D974EFFDDFA93E008FAFAFEAC" box="[1329,1388,332,356]" country="Singapore" name="Zoological Reference Collection, National University of Singapore" pageId="15" pageNumber="316">ZRC</collectionCode>
);
<specimenCount id="9D47FD71974EFFDDFA22E008FF62FE40" pageId="15" pageNumber="316" type="worker">12 workers</specimenCount>
, 2 queens,
<location id="8E9E6023974EFFDDFE20E02BFD40FE4F" LSID="urn:lsid:plazi:treatment:03E887EE9748FFDDFF35E5EFFD9FFCDC:8E9E6023974EFFDDFE20E02BFD40FE4F" box="[386,726,367,392]" country="Singapore" latitude="1.35449" longLatPrecision="1" longitude="103.78211" name="Bukit Timah Nature Reserve" pageId="15" pageNumber="316">Bukit Timah Nature Reserve</location>
(
<geoCoordinate id="EE75503F974EFFDDFD49E02BFCFCFE40" box="[747,874,367,392]" direction="north" orientation="latitude" pageId="15" pageNumber="316" precision="1" value="1.35449">1.35449°N</geoCoordinate>
,
<geoCoordinate id="EE75503F974EFFDDFCD9E034FB87FE40" box="[891,1041,368,392]" direction="east" orientation="longitude" pageId="15" pageNumber="316" precision="1" value="103.78211">103.78211°E</geoCoordinate>
), nest in bark of
<taxonomicName id="4C414D7B974EFFDDFB49E034FEB6FE64" class="Magnoliopsida" family="Anacardiaceae" genus="Campnosperma" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="15" pageNumber="316" phylum="Tracheophyta" rank="species" species="auriculatum">
<emphasis id="B935EAEA974EFFDDFB49E034FEB6FE64" italics="true" pageId="15" pageNumber="316">Campnosperma auriculatum</emphasis>
</taxonomicName>
,
<date id="FFFF1038974EFFDDFE8CE0D0FE2CFE64" box="[302,442,404,428]" pageId="15" pageNumber="316" value="2017-06-21">
<collectingDate id="EFBBE9D0974EFFDDFE8CE0D0FE2CFE64" box="[302,442,404,428]" pageId="15" pageNumber="316" value="2017-06-21">21 Jun 2017</collectingDate>
</date>
,
<collectorName id="26B4532E974EFFDDFE67E0D0FDA0FE64" box="[453,566,404,428]" pageId="15" pageNumber="316">C. Peeters</collectorName>
leg., colony no. WW-SG17-015,
<collectionCode id="ED50AE3D974EFFDDFC12E0D0FC70FE64" box="[944,998,404,428]" country="Singapore" name="Zoological Reference Collection, National University of Singapore" pageId="15" pageNumber="316">ZRC</collectionCode>
</materialsCitation>
_HYM_0001732 (ZRC).
<materialsCitation id="3B293CA5974EFFDDFAA6E0D0FD00FE3D" ID-GBIF-Occurrence="2118916486" collectingDate="2018-03-11" collectorName="W. Thailand &amp; W. Jaitrong" country="Thailand" county="Prov." elevation="280" latitude="14.33389" location="Ban Tha Sao" longLatPrecision="538" longitude="98.98" municipality="Sai Yok Dist." pageId="15" pageNumber="316" specimenCount="1" stateProvince="Kanchanaburi">
<collectingCountry id="F3567668974EFFDDFAA6E0D0FA0FFE64" box="[1284,1433,404,428]" name="Thailand" pageId="15" pageNumber="316">THAILAND</collectingCountry>
:
<collectorName id="26B4532E974EFFDDFF35E0FCFEB4FE18" box="[151,290,439,464]" pageId="15" pageNumber="316">W. Thailand</collectorName>
,
<collectingRegion id="4985F81A974EFFDDFE8FE0F3FE5CFE18" box="[301,458,439,464]" country="Thailand" name="Kanchanaburi" pageId="15" pageNumber="316">Kanchanaburi</collectingRegion>
<collectingCounty id="629F4E74974EFFDDFE73E0FCFD99FE07" box="[465,527,440,463]" pageId="15" pageNumber="316">Prov.</collectingCounty>
,
<collectingMunicipality id="6B9AAC82974EFFDDFDBBE0F3FD26FE18" box="[537,688,439,464]" pageId="15" pageNumber="316">Sai Yok Dist.</collectingMunicipality>
,
<location id="8E9E6023974EFFDDFD19E0FCFCD8FE18" LSID="urn:lsid:plazi:treatment:03E887EE9748FFDDFF35E5EFFD9FFCDC:8E9E6023974EFFDDFD19E0FCFCD8FE18" box="[699,846,439,464]" country="Thailand" county="Prov." latitude="14.33389" longLatPrecision="538" longitude="98.98" municipality="Sai Yok Dist." name="Ban Tha Sao" pageId="15" pageNumber="316" stateProvince="Kanchanaburi">Ban Tha Sao</location>
,
<elevation id="006CD1CB974EFFDDFCFAE0FCFC4CFE18" box="[856,986,439,464]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="15" pageNumber="316" unit="m" value="280.0">
<quantity id="4CB99B1D974EFFDDFCFAE0FCFC09FE07" box="[856,927,440,464]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="15" pageNumber="316" unit="m" value="280.0">280 m</quantity>
a.s.l.
</elevation>
(
<geoCoordinate id="EE75503F974EFFDDFC4AE0FCFBEFFE07" box="[1000,1145,440,464]" direction="north" orientation="latitude" pageId="15" pageNumber="316" precision="1" value="14.33389">14.33389° N</geoCoordinate>
,
<geoCoordinate id="EE75503F974EFFDDFB26E0FCFB70FE18" box="[1156,1254,440,464]" direction="east" orientation="longitude" pageId="15" pageNumber="316" precision="555" value="98.98">98.98° E</geoCoordinate>
), collected from a mango tree,
<date id="FFFF1038974EFFDDFE9EE098FE44FE3C" box="[316,466,476,500]" pageId="15" pageNumber="316" value="2018-03-11">
<collectingDate id="EFBBE9D0974EFFDDFE9EE098FE44FE3C" box="[316,466,476,500]" pageId="15" pageNumber="316" value="2018-03-11">11 Mar 2018</collectingDate>
</date>
,
<collectorName id="26B4532E974EFFDDFE42E099FDF7FE3D" box="[480,609,476,501]" pageId="15" pageNumber="316">W. Jaitrong</collectorName>
leg.
</materialsCitation>
,
<materialsCitation id="3B293CA5974EFFDDFD06E098FA06FDF4" ID-GBIF-Occurrence="2118820490" collectingDate="18-03-18" collectingDateMax="2018-03-18" collectingDateMin="18-03-18" collectionCode="THNHM, ZRC" collectorName="W. Jaitrong" country="Thailand" county="Central Thailand" elevation="89" latitude="14.67056" location="Phu Kae Botanical Garden" longLatPrecision="1" longitude="100.885" municipality="Prov." pageId="15" pageNumber="316" specimenCount="1" stateProvince="Sara Buri">
colony no.WJT
<date id="FFFF1038974EFFDDFCF7E098FC3BFE3C" box="[853,941,476,500]" pageId="15" pageNumber="316" value="18-11-03">
<collectingDate id="EFBBE9D0974EFFDDFCF7E098FC3BFE3C" box="[853,941,476,500]" pageId="15" pageNumber="316" value="18-11-03">110318</collectingDate>
</date>
-1 (
<collectionCode id="ED50AE3D974EFFDDFC6CE098FBAFFE3C" box="[974,1081,476,500]" pageId="15" pageNumber="316">THNHM</collectionCode>
,
<collectionCode id="ED50AE3D974EFFDDFBE4E098FB17FE3C" box="[1094,1153,476,500]" country="Singapore" name="Zoological Reference Collection, National University of Singapore" pageId="15" pageNumber="316">ZRC</collectionCode>
); same locality, date and collector, colony no.WJT
<date id="FFFF1038974EFFDDFE10E344FD91FDD0" box="[434,519,512,536]" pageId="15" pageNumber="316" value="18-11-03">
<collectingDate id="EFBBE9D0974EFFDDFE10E344FD91FDD0" box="[434,519,512,536]" pageId="15" pageNumber="316" value="18-11-03">110318</collectingDate>
</date>
-2 (
<collectionCode id="ED50AE3D974EFFDDFD89E344FD00FDD0" box="[555,662,512,536]" pageId="15" pageNumber="316">THNHM</collectionCode>
,
<collectionCode id="ED50AE3D974EFFDDFD00E344FD4BFDD0" box="[674,733,512,536]" country="Singapore" name="Zoological Reference Collection, National University of Singapore" pageId="15" pageNumber="316">ZRC</collectionCode>
);
<collectingCounty id="629F4E74974EFFDDFD52E0BBFC26FDD0" box="[752,944,511,536]" pageId="15" pageNumber="316">Central Thailand</collectingCounty>
,
<collectingRegion id="4985F81A974EFFDDFC19E0BBFB8CFDD0" box="[955,1050,511,536]" country="Thailand" name="Sara Buri" pageId="15" pageNumber="316">Saraburi</collectingRegion>
<collectingMunicipality id="6B9AAC82974EFFDDFB83E344FBC9FDDF" box="[1057,1119,512,535]" pageId="15" pageNumber="316">Prov.</collectingMunicipality>
,
<location id="8E9E6023974EFFDDFBCBE0BBFA0EFDD0" LSID="urn:lsid:plazi:treatment:03E887EE9748FFDDFF35E5EFFD9FFCDC:8E9E6023974EFFDDFBCBE0BBFA0EFDD0" box="[1129,1432,511,536]" country="Thailand" county="Central Thailand" latitude="14.67056" longLatPrecision="1" longitude="100.885" municipality="Prov." name="Phu Kae Botanical Garden" pageId="15" pageNumber="316" stateProvince="Sara Buri">Phu Kae Botanical Garden</location>
,
<elevation id="006CD1CB974EFFDDFF35E360FE96FDF5" box="[151,256,548,573]" metricMagnitude="1" metricUnit="m" metricValue="8.9" pageId="15" pageNumber="316" unit="m" value="89.0">
<quantity id="4CB99B1D974EFFDDFF35E360FF46FDF4" box="[151,208,548,572]" metricMagnitude="1" metricUnit="m" metricValue="8.9" pageId="15" pageNumber="316" unit="m" value="89.0">89 m</quantity>
a.s.l.
</elevation>
(
<geoCoordinate id="EE75503F974EFFDDFEAAE360FE0AFDF5" box="[264,412,548,573]" direction="north" orientation="latitude" pageId="15" pageNumber="316" precision="1" value="14.67056">14.67056°N</geoCoordinate>
,
<geoCoordinate id="EE75503F974EFFDDFE0AE360FDD6FDF5" box="[424,576,548,573]" direction="east" orientation="longitude" pageId="15" pageNumber="316" precision="1" value="100.885">100.88500°E</geoCoordinate>
),
<date id="FFFF1038974EFFDDFDF1E360FD70FDF4" box="[595,742,548,572]" pageId="15" pageNumber="316" value="2018-03-18">
<collectingDate id="EFBBE9D0974EFFDDFDF1E360FD70FDF4" box="[595,742,548,572]" pageId="15" pageNumber="316" value="2018-03-18">18 Mar 2018</collectingDate>
</date>
,
<collectorName id="26B4532E974EFFDDFD53E361FCE7FDF5" box="[753,881,548,573]" pageId="15" pageNumber="316">W. Jaitrong</collectorName>
leg., colony no.WJT
<date id="FFFF1038974EFFDDFBFFE360FB25FDF4" box="[1117,1203,548,572]" pageId="15" pageNumber="316" value="18-03-18">
<collectingDate id="EFBBE9D0974EFFDDFBFFE360FB25FDF4" box="[1117,1203,548,572]" pageId="15" pageNumber="316" value="18-03-18">180318</collectingDate>
</date>
-2 (
<collectionCode id="ED50AE3D974EFFDDFB74E360FAD7FDF4" box="[1238,1345,548,572]" pageId="15" pageNumber="316">THNHM</collectionCode>
,
<collectionCode id="ED50AE3D974EFFDDFAEEE360FA1EFDF4" box="[1356,1416,548,572]" country="Singapore" name="Zoological Reference Collection, National University of Singapore" pageId="15" pageNumber="316">ZRC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BFE36F8974EFFDDFF65E303FD9FFCDC" blockId="15.[151,1437,151,788]" pageId="15" pageNumber="316">
<emphasis id="B935EAEA974EFFDDFF65E303FE07FDA8" bold="true" box="[199,401,583,608]" pageId="15" pageNumber="316">DNA Barcoding.</emphasis>
<collectingCountry id="F3567668974EFFDDFE39E30CFDABFDA8" box="[411,573,584,608]" name="Singapore" pageId="15" pageNumber="316">SINGAPORE</collectingCountry>
:
<specimenCount id="9D47FD71974EFFDDFDE9E303FD5DFDA8" box="[587,715,583,608]" pageId="15" pageNumber="316" type="worker">16 workers</specimenCount>
were successfully barcoded, 8 each from a colony in Mandai (ZRC_HYM_0000576) and another colony in BTNR (ZRC_HYM_0001732).
<collectingCountry id="F3567668974EFFDDFB93E328FB50FD4C" box="[1073,1222,620,644]" name="Thailand" pageId="15" pageNumber="316">THAILAND</collectingCountry>
:
<specimenCount id="9D47FD71974EFFDDFB74E328FACFFD4D" box="[1238,1369,620,645]" pageId="15" pageNumber="316" type="worker">14 workers</specimenCount>
were successfully barcoded, 5 each from 2 colonies in West Thailand (WJT
<date id="FFFF1038974EFFDDFC6DE3D4FBBEFD60" box="[975,1064,656,680]" pageId="15" pageNumber="316" value="18-11-03">110318</date>
-1, WJT
<date id="FFFF1038974EFFDDFB23E3D4FB41FD60" box="[1153,1239,656,680]" pageId="15" pageNumber="316" value="18-11-03">110318</date>
-2), and 4 from a colony in Central Thailand (WJT
<date id="FFFF1038974EFFDDFD8CE3F0FD10FD04" box="[558,646,692,716]" pageId="15" pageNumber="316" value="18-03-18">180318</date>
-2). Based on the combined cluster dendrogram (
<figureCitation id="137A2A7D974EFFDDFB78E3F0FAB5FD04" box="[1242,1315,692,717]" captionStart="FIGURE 1" captionStartId="3.[151,250,1919,1941]" captionTargetBox="[229,1342,421,1774]" captionTargetId="figure@3.[150,1436,338,1867]" captionTargetPageId="3" captionText="FIGURE 1. Cluster dendrogram of COI (313 bp) barcodes of Rhopalomastix from Singapore and Thailand. Objective clustering of barcodes was performed based on uncorrected p-distances using the best close match criteria; the number at each node represents the percentage divergence or pairwise distance threshold at which clusters split or lump together. Each cluster identified as a species is highlighted in one colour per respective species. Sequences of Thai Rhopalomastix are outlined in red." httpUri="https://zenodo.org/record/2615333/files/figure.png" pageId="15" pageNumber="316">Fig. 1</figureCitation>
), barcode divergences from other species are as follows:
<taxonomicName id="4C414D7B974EFFDDFD03E39DFC8BFD38" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2018" box="[673,797,728,752]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="murphyi">
<emphasis id="B935EAEA974EFFDDFD03E39DFC8BFD38" box="[673,797,728,752]" italics="true" pageId="15" pageNumber="316">R. murphyi</emphasis>
</taxonomicName>
, 19.2%,
<taxonomicName id="4C414D7B974EFFDDFC23E39DFBABFD38" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2018" box="[897,1085,728,752]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="glabricephala">
<emphasis id="B935EAEA974EFFDDFC23E39DFBABFD38" box="[897,1085,728,752]" italics="true" pageId="15" pageNumber="316">R. glabricephala</emphasis>
</taxonomicName>
, 17%;
<taxonomicName id="4C414D7B974EFFDDFB2FE39DFAB6FD38" box="[1165,1312,728,752]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="johorensis">
<emphasis id="B935EAEA974EFFDDFB2FE39DFAB6FD38" box="[1165,1312,728,752]" italics="true" pageId="15" pageNumber="316">R. johorensis</emphasis>
</taxonomicName>
, 13.8%;
<taxonomicName id="4C414D7B974EFFDDFA26E39DFF76FCDC" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2018" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="striata">
<emphasis id="B935EAEA974EFFDDFA26E39DFF76FCDC" italics="true" pageId="15" pageNumber="316">R. striata</emphasis>
</taxonomicName>
, 13.8%;
<taxonomicName id="4C414D7B974EFFDDFEE7E3B9FE2FFCDC" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2018" box="[325,441,764,788]" class="Insecta" family="Formicidae" genus="Rhopalomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="tenebra">
<emphasis id="B935EAEA974EFFDDFEE7E3B9FE2FFCDC" box="[325,441,764,788]" italics="true" pageId="15" pageNumber="316">R. tenebra</emphasis>
</taxonomicName>
, 7.1%.
</paragraph>
</subSubSection>
</treatment>
</document>