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<document id="C51DC27694CFC2D8E98505EB7175C2B3" ID-DOI="10.11646/zootaxa.5414.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10681263" ID-ZooBank="3B1DFADA-C20E-473B-A5E9-843548B54AD3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1708377054888" checkinUser="plazi" docAuthor="Paredes-Munguía, Williams, Brescovit, Antonio D. &amp; Teixeira, Renato A." docDate="2024" docId="03E887D2BA0BFE2AFDDBFADF60C5F9C8" docLanguage="en" docName="zootaxa.5414.1.1.pdf" docOrigin="Zootaxa 5414 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5414.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.18:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="18" docTitle="Arctosa jibarosa Paredes-Munguía &amp; Brescovit &amp; Teixeira 2024" docType="key" docVersion="1" lastPageNumber="7" masterDocId="FFD1FFAABA0DFE2CFD4CFF8E6559FFD2" masterDocTitle="Revision of Neotropical wolf spider genus Arctosa C. L. Koch, 1847 (Araneae: Lycosidae), with description of seven new species" masterLastPageNumber="83" masterPageNumber="1" pageNumber="7" updateTime="1708377875290" updateUser="ExternalLinkService">
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<mods:title id="97A92C4FD81B99C348C7C56A12A24B75">Revision of Neotropical wolf spider genus Arctosa C. L. Koch, 1847 (Araneae: Lycosidae), with description of seven new species</mods:title>
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<mods:namePart id="AFCD30BFE6D07F4C5E7D98D999E5F166">Paredes-Munguía, Williams</mods:namePart>
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<mods:affiliation id="FA6563C43A4461FD6EC508E13DBA0765">Pontifícia Universidade Católica do Rio Grande do Sul, Escola de Ciências da Saúde e da Vida, Museu de Ciências e Tecnologia, Laboratório de Aracnologia, Porto Alegre, Brazil. &amp; Museo de Historia Natural de la Universidad Nacional Mayor de San Marcos, Laboratorio de Entomología, Av. Arenales 1256, Lima 14, Peru. &amp; williams. paredes @ acad. pucrs. br; https: // orcid. org / 0000 - 0002 - 3320 - 4611</mods:affiliation>
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<mods:namePart id="3397C32FB3E0F1D65F5D576F7A664932">Brescovit, Antonio D.</mods:namePart>
<mods:nameIdentifier id="D25D10A3D5DDC5C705E6C227A2215E95" type="ORCID">0000-0002-1511-5324</mods:nameIdentifier>
<mods:affiliation id="C58BFBACE7976D0B1423AD2C8C22A8D4">Laboratório de Coleções Zoológicas, Av. Vital Brasil, 1500, Butantã, 05503 - 900, São Paulo, SP, Brazil. &amp; antonio. brescovit @ butantan. gov. br; https: // orcid. org / 0000 - 0002 - 1511 - 5324</mods:affiliation>
<mods:nameIdentifier id="694DC0B92513C5B1D3A84A8D2F218E4D" type="email">antonio.brescovit@butantan.gov.br</mods:nameIdentifier>
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<mods:namePart id="5B2EC311D337F39DF12E34FEB43141A0">Teixeira, Renato A.</mods:namePart>
<mods:nameIdentifier id="82E98E03C4C854AB87A7F647B5C010A7" type="ORCID">0000-0002-1756-9821</mods:nameIdentifier>
<mods:affiliation id="17FD447364583DCFFCC2FA2BAFD6DE57">Pontifícia Universidade Católica do Rio Grande do Sul, Escola de Ciências da Saúde e da Vida, Museu de Ciências e Tecnologia, Laboratório de Aracnologia, Porto Alegre, Brazil. &amp; Pontifícia Universidade Católica do Rio Grande do Sul, Escola de Ciências da Saúde e da Vida, Museu de Ciências e Tecnologia, Laboratório de Aracnologia, Porto Alegre, Brazil. &amp; renatoaug. tx @ gmail. com; https: // orcid. org / 0000 - 0002 - 1756 - 9821</mods:affiliation>
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<mods:title id="0EFAC214D2EA07AA2896D884B4947826">Zootaxa</mods:title>
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<mods:date id="479723AF8C33BEB44939740941ED5533">2024</mods:date>
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<treatment id="03E887D2BA0BFE2AFDDBFADF60C5F9C8" LSID="urn:lsid:plazi:treatment:03E887D2BA0BFE2AFDDBFADF60C5F9C8" httpUri="http://treatment.plazi.org/id/03E887D2BA0BFE2AFDDBFADF60C5F9C8" lastPageNumber="7" pageId="6" pageNumber="7">
<subSubSection id="C35B654FBA0BFE2AFDDBFADF64D2FAB9" box="[151,395,1361,1387]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8BFE36C4BA0BFE2AFDDBFADF64D2FAB9" blockId="6.[151,1436,1361,1562]" box="[151,395,1361,1387]" pageId="6" pageNumber="7">
<heading id="D0B681A8BA0BFE2AFDDBFADF64D2FAB9" box="[151,395,1361,1387]" fontSize="11" level="2" pageId="6" pageNumber="7" reason="5">
<taxonomicName id="4C414D47BA0BFE2AFDDBFADF65AAFAB9" authorityName="Paredes-Munguía &amp; Brescovit &amp; Teixeira" authorityYear="2024" box="[151,243,1361,1387]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="jibarosa">
<emphasis id="B935EAD6BA0BFE2AFDDBFADF65AAFAB9" box="[151,243,1361,1387]" italics="true" pageId="6" pageNumber="7">jibarosa</emphasis>
</taxonomicName>
group, males
</heading>
</paragraph>
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<subSubSection id="C35B654FBA0BFE2AFDDBFAFC60C5F9C8" pageId="6" pageNumber="7" type="key">
<key id="FFD061F5BA0BFE2AFDDBFAFC60C5F9C8" pageId="6" pageNumber="7">
<keyStep id="30B52E61BA0BFE2AFDDBFAFC60C5FA76" pageId="6" pageNumber="7">
<paragraph id="8BFE36C4BA0BFE2AFDDBFAFC60C5FA55" blockId="6.[151,1436,1361,1562]" box="[151,1436,1393,1416]" pageId="6" pageNumber="7">
<keyLead id="30B095F1BA0BFE2AFDDBFAFC60C5FA55" box="[151,1436,1393,1416]" pageId="6" pageNumber="7">
1 Depression of Median apophysis (DMA) shallow (
<figureCitation id="137A2A41BA0BFE2AFF85FAFC6643FA5A" box="[713,794,1394,1416]" captionStart="FIGURE 20" captionStartId="25.[151,250,1482,1506]" captionTargetBox="[151,1436,489,1459]" captionTargetId="figure-180@25.[151,1436,489,1459]" captionTargetPageId="25" captionText="FIGURE 20. Arctosa jibarosa sp. nov. Male habitus (MUSM-ENT 504111) a, d, g–h (a, dorsal; d, ventral; g, lateral; h, frontal). Epigynum (MUSM-ENT 504111) b, e (b, dorsal; e, ventral). Female habitus paratype (MUSM-ENT 505269) c, f, i (c, dorsal; f, ventral; i, lateral). Scale bars: a, d, g, 1 mm, b, e, 0.1 mm, h, 200 µm, c, f, i, 1 mm. SS stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.10681305" httpUri="https://zenodo.org/record/10681305/files/figure.png" pageId="6" pageNumber="7">Fig. 20f</figureCitation>
)…............................. ...
<emphasis id="B935EAD6BA0BFE2AF9EBFAFC60C5FA55" bold="true" box="[1191,1436,1393,1415]" pageId="6" pageNumber="7">
<taxonomicName id="4C414D47BA0BFE2AF9EBFAFC6010FA55" authority="Paredes-Munguía &amp; Brescovit &amp; Teixeira, 2024" authorityName="Paredes-Munguía &amp; Brescovit &amp; Teixeira" authorityYear="2024" box="[1191,1353,1393,1415]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="jibarosa" status="sp. nov.">
<emphasis id="B935EAD6BA0BFE2AF9EBFAFC6010FA55" bold="true" box="[1191,1353,1393,1415]" italics="true" pageId="6" pageNumber="7">Arctosa jibarosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20657ADBA0BFE2AF81CFAFC60C5FA55" box="[1360,1436,1394,1415]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BFE36C4BA0BFE2AFDDBFA1E60C5FA76" blockId="6.[151,1436,1361,1562]" box="[151,1436,1423,1445]" pageId="6" pageNumber="7">
<keyLead id="30B095F1BA0BFE2AFDDBFA1E60C5FA76" box="[151,1436,1423,1445]" pageId="6" pageNumber="7">
- Depression of median apophysis (DMA) deep (
<figureCitation id="137A2A41BA0BFE2AFFE5FA0167B6FA77" box="[681,751,1423,1445]" captionStart="FIGURE 7" captionStartId="13.[151,250,1214,1238]" captionTargetBox="[151,1436,181,1191]" captionTargetId="figure-21@13.[151,1436,181,1191]" captionTargetPageId="13" captionText="FIGURE 7. Arctosa conflicta sp. nov. Male habitus (ICN-Ar 12814) a–c (a, dorsal; b, ventral; c, lateral), pedipalpal bulb d–g (d, ventral; e, prolateral; f, anterior; g, posterior). Scale bars: a–g, 200 µm. DMA depression of MA, E embolus, Pa posterior arm, Pp Palea protuberance, Ra retrolateral arm, SD seminal duct, ST subtegulum, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681277" httpUri="https://zenodo.org/record/10681277/files/figure.png" pageId="6" pageNumber="7">Fig. 7d</figureCitation>
).......................................................
<emphasis id="B935EAD6BA0BFE2AF8DCFA0160C5FA76" bold="true" box="[1424,1436,1423,1444]" pageId="6" pageNumber="7">2</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B52E61BA0BFE2AFDDBFA2360C5FA0D" pageId="6" pageNumber="7">
<paragraph id="8BFE36C4BA0BFE2AFDDBFA2360C5FA10" blockId="6.[151,1436,1361,1562]" box="[151,1436,1452,1474]" pageId="6" pageNumber="7">
<keyLead id="30B095F1BA0BFE2AFDDBFA2360C5FA10" box="[151,1436,1452,1474]" pageId="6" pageNumber="7">
2 Retrolateral arm of median apophysis (Ra) bulging and pronounced (
<figureCitation id="137A2A41BA0BFE2AFE34FA2266E5FA10" box="[888,956,1452,1474]" captionStart="FIGURE 6" captionStartId="12.[151,250,1952,1976]" captionTargetBox="[158,1432,1444,1921]" captionTargetId="figure-83@12.[151,1436,1437,1928]" captionTargetPageId="12" captionText="FIGURE 6. Arctosa conflicta sp. nov. Male bulb (ICN-Ar 12814) (a, ventral; b, retrolateral; c, apical. Scale bars: a–c, 200 µm. E embolus, Pa Posterior arm, Pp Palea protuberance, Ra Retrolateral arm, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681275" httpUri="https://zenodo.org/record/10681275/files/figure.png" pageId="6" pageNumber="7">Fig. 6a</figureCitation>
)................. ..
<emphasis id="B935EAD6BA0BFE2AF9EDFA2260C5FA10" bold="true" box="[1185,1436,1452,1474]" pageId="6" pageNumber="7">
<taxonomicName id="4C414D47BA0BFE2AF9EDFA226013FA10" authority="Paredes-Munguía &amp; Brescovit &amp; Teixeira, 2024" authorityName="Paredes-Munguía &amp; Brescovit &amp; Teixeira" authorityYear="2024" box="[1185,1354,1452,1474]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="conflicta" status="sp. nov.">
<emphasis id="B935EAD6BA0BFE2AF9EDFA226013FA10" bold="true" box="[1185,1354,1452,1474]" italics="true" pageId="6" pageNumber="7">Arctosa conflicta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20657ADBA0BFE2AF81CFA2360C5FA10" box="[1360,1436,1453,1474]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BFE36C4BA0BFE2AFDDBFA4460C5FA0D" blockId="6.[151,1436,1361,1562]" box="[151,1436,1482,1504]" pageId="6" pageNumber="7">
<keyLead id="30B095F1BA0BFE2AFDDBFA4460C5FA0D" box="[151,1436,1482,1504]" pageId="6" pageNumber="7">
- Retrolateral arm of median apophysis (Ra) with an acute tip (
<figureCitation id="137A2A41BA0BFE2AFE62FA446626FA0D" box="[814,895,1482,1504]" captionStart="FIGURE 40" captionStartId="47.[151,250,1581,1605]" captionTargetBox="[185,1402,181,1557]" captionTargetId="figure-21@47.[185,1402,181,1557]" captionTargetPageId="47" captionText="FIGURE 40. Arctosa tenella (Keyserling, 1877) comb. nov. Male habitus (ICN-Ar 2577) a–c (a, dorsal; b, lateral; c, ventral). Pedipalp, expanded d–e (d, prolateral; e, anterior). Scale bars: a–c 1 mm, d–e 150 µm. BH basal hematodocha, DH distal hematodocha, MA median apophysis, Pa posterior arm, Ra retrolateral arm, ST subtegulum, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681347" httpUri="https://zenodo.org/record/10681347/files/figure.png" pageId="6" pageNumber="7">Fig. 40c</figureCitation>
)...........................................
<emphasis id="B935EAD6BA0BFE2AF8DCFA4460C5FA0D" bold="true" box="[1424,1436,1482,1503]" pageId="6" pageNumber="7">3</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B52E61BA0BFE2AFDDBFA6960C5F9C8" pageId="6" pageNumber="7">
<paragraph id="8BFE36C4BA0BFE2AFDDBFA6960C5FA2E" blockId="6.[151,1436,1361,1562]" box="[151,1436,1511,1533]" pageId="6" pageNumber="7">
<keyLead id="30B095F1BA0BFE2AFDDBFA6960C5FA2E" box="[151,1436,1511,1533]" pageId="6" pageNumber="7">
3 Terminal apophysis (TA) curved (
<figureCitation id="137A2A41BA0BFE2AFF6AFA696722FA2E" box="[550,635,1511,1533]" captionStart="FIGURE 41" captionStartId="48.[151,250,1414,1438]" captionTargetBox="[159,1428,497,1383]" captionTargetId="figure-160@48.[151,1436,489,1391]" captionTargetPageId="48" captionText="FIGURE 41. Arctosa tenella (Keyserling, 1877) comb. nov. Epigynum (ICN-Ar 2577) (a, ventral; b, dorsal; c, posterolateral; d, posterior). Scale bars: a–d, 0.25 mm. CO copulatory opening, HS head of spermatheca, FD fertilization duct, MS median septum, VC vulval chamber." figureDoi="http://doi.org/10.5281/zenodo.10681349" httpUri="https://zenodo.org/record/10681349/files/figure.png" pageId="6" pageNumber="7">Figs 41e</figureCitation>
,
<figureCitation id="137A2A41BA0BFE2AFFC9FA6967F0FA2E" box="[645,681,1511,1532]" captionStart="FIGURE 43" captionStartId="49.[151,250,1942,1966]" captionTargetBox="[259,1329,1143,1919]" captionTargetId="figure-76@49.[259,1329,1143,1919]" captionTargetPageId="49" captionText="FIGURE 43. Arctosa tenella (Keyserling, 1877) comb. nov. Male palpal bulb SEM (MUSM-ENT 508928) a–c (a, ventral; b, conductor detail; c, MA details). Scale bars: a, 150 µm, b–c, 50 µm. C conductor, E embolus, Pa posterior arm, Ra retrolateral arm, ST subtegulum, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681353" httpUri="https://zenodo.org/record/10681353/files/figure.png" pageId="6" pageNumber="7">43a</figureCitation>
)............
<emphasis id="B935EAD6BA0BFE2AFE0FFA6960C5FA2E" bold="true" box="[835,1436,1511,1533]" pageId="6" pageNumber="7">
<taxonomicName id="4C414D47BA0BFE2AFE0FFA696070FA2F" authority="(F. O. Pickard-Cambridge, 1902)" authorityName="Paredes-Munguía &amp; Brescovit &amp; Teixeira" authorityYear="2024" baseAuthorityName="F. O. Pickard-Cambridge" baseAuthorityYear="1902" box="[835,1321,1511,1533]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tenella" status="comb. nov.">
<emphasis id="B935EAD6BA0BFE2AFE0FFA696681FA2F" bold="true" box="[835,984,1511,1533]" italics="true" pageId="6" pageNumber="7">Arctosa tenella</emphasis>
(F.O.
<bibRefCitation id="EFD04B35BA0BFE2AF95BFA696078FA2F" author="Pickard-Cambridge, F. O." box="[1047,1313,1511,1533]" pageId="6" pageNumber="7" refId="ref51737" refString="Pickard-Cambridge, F. O. (1902) Arachnida - Araneida and Opiliones. In: Godman, F. D. &amp; Salvin, O. (Eds.), Biologia Centrali- Americana. Vol. 2. Published for the editors by R. H. Porter, London, pp. 312 - 424." type="book" year="1902">Pickard-Cambridge, 1902</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A20657ADBA0BFE2AF862FA6960C5FA2E" box="[1326,1436,1511,1532]" pageId="6" pageNumber="7" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BFE36C4BA0BFE2AFDDBF98B60C5F9C8" blockId="6.[151,1436,1361,1562]" box="[151,1436,1540,1562]" pageId="6" pageNumber="7">
<keyLead id="30B095F1BA0BFE2AFDDBF98B60C5F9C8" box="[151,1436,1540,1562]" pageId="6" pageNumber="7">
- Terminal apophysis (TA) straight (
<figureCitation id="137A2A41BA0BFE2AFF61F98A67DBF9C8" box="[557,642,1540,1562]" captionStart="FIGURE 34" captionStartId="41.[151,250,1745,1769]" captionTargetBox="[162,1425,680,1711]" captionTargetId="figure-256@41.[151,1436,669,1722]" captionTargetPageId="41" captionText="FIGURE 34. Arctosa tenebrosa (Keyserling, 1877) comb. nov. Male. Pedipalp (ICN-Ar 1378) a–c (a, ventral; b, retrolateral; c, anterior). Scale bars: a–c, 0.25 mm. CMS cymbium macrosetae, Pa posterior arm, Pp palea protuberance, Ra retrolateral arm, T tegulum, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681333" httpUri="https://zenodo.org/record/10681333/files/figure.png" pageId="6" pageNumber="7">Figs 34a</figureCitation>
,
<figureCitation id="137A2A41BA0BFE2AFFC0F98A67EBF9C8" box="[652,690,1540,1562]" captionStart="FIGURE 35" captionStartId="42.[151,250,1851,1875]" captionTargetBox="[151,1436,181,1827]" captionTargetId="figure-25@42.[151,1436,181,1827]" captionTargetPageId="42" captionText="FIGURE 35. Arctosa tenebrosa (Keyserling, 1877) comb. nov. Male. Pedipalp (ICN-Ar 1378) a–c, i (a, ventral; b, retrolateral; c, prolateral; i, dorsal setae). Palpal bulb d–h (d, ventral; e, retrolateral; f, prolateral; g, posterior; h, anterior). Scale bars: a–c, 0.2 mm, d–i, 100 µm. C Conductor, DMA depression of median apophysis, E embolus, Pa posterior arm, ST subtegulum, T tegulum, TA terminal apophysis, TK tegular keel. * basal hump on median apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681335" httpUri="https://zenodo.org/record/10681335/files/figure.png" pageId="6" pageNumber="7">35d</figureCitation>
).....................
<emphasis id="B935EAD6BA0BFE2AFEFAF98A60C5F9C8" bold="true" box="[950,1436,1540,1562]" pageId="6" pageNumber="7">
<taxonomicName id="4C414D47BA0BFE2AFEFAF98A6070F9C8" authority="(Keyserling, 1877)" authorityName="Paredes-Munguía &amp; Brescovit &amp; Teixeira" authorityYear="2024" baseAuthorityName="Keyserling" baseAuthorityYear="1877" box="[950,1321,1540,1562]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tenebrosa" status="comb. nov.">
<emphasis id="B935EAD6BA0BFE2AFEFAF98A6131F9C8" bold="true" box="[950,1128,1540,1562]" italics="true" pageId="6" pageNumber="7">Arctosa tenebrosa</emphasis>
(
<bibRefCitation id="EFD04B35BA0BFE2AF938F98A6078F9C8" author="Keyserling, G. E." box="[1140,1313,1540,1562]" pageId="6" pageNumber="7" pagination="609 - 708" refId="ref50205" refString="Keyserling, G. E. (1877) Ueber amerikanische spinnenarten der unterordnung Citigradae. Verhandlungen Der Kaiserlich- Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 26, 609 - 708." type="journal article" year="1877">Keyserling, 1877</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A20657ADBA0BFE2AF862F98B60C5F9C8" box="[1326,1436,1541,1562]" pageId="6" pageNumber="7" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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</key>
</subSubSection>
</treatment>
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