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<document id="BA070B9F4F015ACD6C09535E82AB3A78" ID-DOI="10.5852/ejt.2020.718.1101" ID-Zenodo-Dep="4069574" ID-ZooBank="F9E9A91E-488C-4DB1-9361-E788E9AC5BC1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1602021550335" checkinUser="valdenar" docAuthor="Huber, Bernhard A. &amp; Villarreal, Osvaldo" docDate="2020" docId="03E887ADFFE97A38FD2AFDE0FAA7FB61" docLanguage="en" docName="ejt-718_huber_villareal.pdf.imf" docOrigin="European Journal of Taxonomy 718" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Galapa Huber 2000" docType="treatment" docVersion="7" lastPageNumber="58" masterDocId="FFD1FFD5FFD07A01FF8BFF8FFA5BFFBA" masterDocTitle="On Venezuelan pholcid spiders (Araneae, Pholcidae)" masterLastPageNumber="317" masterPageNumber="1" pageNumber="58" updateTime="1698792674692" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="F8BB1598DD599A1066F26CDBFD48CCD3">On Venezuelan pholcid spiders (Araneae, Pholcidae)</mods:title>
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<mods:namePart id="EB67B5F876F19B4932BAB3D6BE8745C2">Huber, Bernhard A.</mods:namePart>
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<mods:affiliation id="889D3F9C91E9E992E132782435F464B2">Zoological Research Museum Alexander Koenig, Bonn, Germany.</mods:affiliation>
<mods:nameIdentifier id="5C7982CB8EDF0F0CFA1A02176B80BDED" type="email">b.huber@leibniz-zfmk.de</mods:nameIdentifier>
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<mods:namePart id="70E43AB40CD41B00940C796F74DBB63A">Villarreal, Osvaldo</mods:namePart>
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<mods:affiliation id="942C108071227640F25901B877151977">Museo del Instituto de Zoología Agrícola, Universidad Central de Venezuela, Maracay, Venezuela. &amp; Museu Nacional / UFRJ, Rio de Janeiro, Brazil.</mods:affiliation>
<mods:nameIdentifier id="DCD997E46D4BDA8133ABEC3F5636C674" type="email">osvaldovillarreal@gmail.com</mods:nameIdentifier>
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<mods:title id="C0A40DFB2A0E2C920686DCF45208E3B7">European Journal of Taxonomy</mods:title>
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<mods:date id="43F7D60E6E9BEB43DFE07C9CFC5167CE">2020</mods:date>
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<treatment id="03E887ADFFE97A38FD2AFDE0FAA7FB61" ID-DOI="http://doi.org/10.5281/zenodo.4343949" ID-Zenodo-Dep="4343949" LSID="urn:lsid:plazi:treatment:03E887ADFFE97A38FD2AFDE0FAA7FB61" httpUri="http://treatment.plazi.org/id/03E887ADFFE97A38FD2AFDE0FAA7FB61" lastPageNumber="58" pageId="57" pageNumber="58">
<subSubSection id="C35B6530FFE97A38FD2AFDE0FB58FD14" pageId="57" pageNumber="58" type="nomenclature">
<paragraph id="8BFE36BBFFE97A38FD2AFDE0F9C9FD33" blockId="57.[189,914,623,686]" box="[673,914,623,650]" pageId="57" pageNumber="58">
<heading id="D0B681D7FFE97A38FD2AFDE0F9C9FD33" box="[673,914,623,650]" centered="true" fontSize="11" level="2" pageId="57" pageNumber="58" reason="2">
<taxonomicName id="4C414D38FFE97A38FD2AFDE0F9C9FD33" ID-CoL="4LRF" authority="Huber, 2000" authorityName="Huber" authorityYear="2000" box="[673,914,623,650]" class="Arachnida" family="Pholcidae" genus="Galapa" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B935EAA9FFE97A38FD2AFDE0F8A2FD30" bold="true" box="[673,761,623,650]" italics="true" pageId="57" pageNumber="58">Galapa</emphasis>
<bibRefCitation id="EFD04B4AFFE97A38FC8AFDE0F9C9FD33" author="Huber B. A." box="[769,914,623,649]" pageId="57" pageNumber="58" pagination="1 - 348" refId="ref127973" refString="Huber B. A. 2000. New World pholcid spiders (Araneae: Pholcidae): a revision at generic level. Bulletin of the American Museum of Natural History 254: 1 - 348. https: // doi. org / 10.1206 / 0003 - 0090 (2000) 254 % 3 C 0001: NWPSAP % 3 E 2.0. CO; 2" type="journal article" year="2000">Huber, 2000</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BFE36BBFFE97A38FF36FD1BFB58FD14" blockId="57.[189,914,623,686]" box="[189,259,660,686]" pageId="57" pageNumber="58">
<heading id="D0B681D7FFE97A38FF36FD1BFB58FD14" bold="true" box="[189,259,660,686]" fontSize="11" level="3" pageId="57" pageNumber="58" reason="3">
<emphasis id="B935EAA9FFE97A38FF36FD1BFB58FD14" bold="true" box="[189,259,660,686]" pageId="57" pageNumber="58">Notes</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35B6530FFE97A38FF36FD4BFAA7FB61" pageId="57" pageNumber="58" type="description">
<paragraph id="8BFE36BBFFE97A38FF36FD4BF83CFBBF" blockId="57.[189,1399,708,1029]" pageId="57" pageNumber="58">
The genus
<taxonomicName id="4C414D38FFE97A38FECBFD4BFBC3FD64" authorityName="Huber" authorityYear="2000" box="[320,408,708,734]" class="Arachnida" family="Pholcidae" genus="Galapa" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B935EAA9FFE97A38FECBFD4BFBC3FD64" box="[320,408,708,734]" italics="true" pageId="57" pageNumber="58">Galapa</emphasis>
</taxonomicName>
was established for two species endemic to the
<collectingRegion id="4985F859FFE97A38FC5EFD4BFEF7FD64" box="[981,1196,708,734]" country="Ecuador" name="Galapagos" pageId="57" pageNumber="58">Galapagos Islands</collectingRegion>
(
<bibRefCitation id="EFD04B4AFFE97A38FB37FD4BFF0BFD64" author="Huber B. A." box="[1212,1360,708,734]" pageId="57" pageNumber="58" pagination="1 - 348" refId="ref127973" refString="Huber B. A. 2000. New World pholcid spiders (Araneae: Pholcidae): a revision at generic level. Bulletin of the American Museum of Natural History 254: 1 - 348. https: // doi. org / 10.1206 / 0003 - 0090 (2000) 254 % 3 C 0001: NWPSAP % 3 E 2.0. CO; 2" type="journal article" year="2000">Huber 2000</bibRefCitation>
). A third species was described in 2014, also from the
<collectingRegion id="4985F859FFE97A38FC8CFD66F987FCB9" box="[775,988,745,771]" country="Ecuador" name="Galapagos" pageId="57" pageNumber="58">Galapagos Islands</collectingRegion>
(
<bibRefCitation id="EFD04B4AFFE97A38FC60FD66FE29FCB9" author="Baert L." box="[1003,1138,745,771]" pageId="57" pageNumber="58" pagination="263 - 271" refId="ref126061" refString="Baert L. 2014. New spider species (Araneae) from the Galapagos Islands (Ecuador). Bulletin de la Societe royale belge d'Entomologie 149: 263 - 271." type="journal article" year="2014">Baert 2014</bibRefCitation>
). The new Venezuelan species below is assigned to
<taxonomicName id="4C414D38FFE97A38FDA8FC81F820FC92" authorityName="Huber" authorityYear="2000" box="[547,635,782,808]" class="Arachnida" family="Pholcidae" genus="Galapa" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B935EAA9FFE97A38FDA8FC81F820FC92" box="[547,635,782,808]" italics="true" pageId="57" pageNumber="58">Galapa</emphasis>
</taxonomicName>
because it shares two specific similarities: (1) unique (among
<taxonomicName id="4C414D38FFE97A38FF36FCBCFB6AFCF4" authorityName="Simon" authorityYear="1890" box="[189,305,819,846]" class="Arachnida" family="Pholcidae" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="subFamily" subFamily="Ninetinae">Ninetinae</taxonomicName>
) armature of the male chelicerae (processes on male cheliceral fangs but otherwise unmodified; compare
<bibRefCitation id="EFD04B4AFFE97A38FEA0FCD7FBE7FCC8" author="Huber B. A." box="[299,444,856,882]" pageId="57" pageNumber="58" pagination="1 - 348" refId="ref127973" refString="Huber B. A. 2000. New World pholcid spiders (Araneae: Pholcidae): a revision at generic level. Bulletin of the American Museum of Natural History 254: 1 - 348. https: // doi. org / 10.1206 / 0003 - 0090 (2000) 254 % 3 C 0001: NWPSAP % 3 E 2.0. CO; 2" type="journal article" year="2000">Huber 2000</bibRefCitation>
: fig. 384 and
<figureCitation id="137A2A3EFFE97A38FDEAFCD7F893FCC8" box="[609,712,856,882]" captionStart="Figs 178182" captionStartId="59.[189,241,1780,1806]" captionTargetBox="[192,1392,289,1696]" captionTargetId="figure-82@59.[189,1398,289,1718]" captionTargetPageId="59" captionText="Figs 178182. Galapa spiniphila Huber sp. nov.; from Falcón, near Cueva del Guano (type locality; ZFMK Ar 21860). 178179. Left male pedipalp, prolateral and retrolateral views. 180. Left procursus and genital bulb, dorsal view. 181. Male ocular area, clypeus, and chelicerae, frontal view (arrow: process on male cheliceral fang). 182. Cleared female genitalia, dorsal view. Abbreviations: b= genital bulb; p=procursus. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4069648" httpUri="https://zenodo.org/record/4069648/files/figure.png" pageId="57" pageNumber="58">Fig. 181</figureCitation>
); (2) dorsal (in the new species below slightly prolateral) process of the procursus (compare
<bibRefCitation id="EFD04B4AFFE97A38FDEAFCF3F8A9FC2D" author="Huber B. A." box="[609,754,892,919]" pageId="57" pageNumber="58" pagination="1 - 348" refId="ref127973" refString="Huber B. A. 2000. New World pholcid spiders (Araneae: Pholcidae): a revision at generic level. Bulletin of the American Museum of Natural History 254: 1 - 348. https: // doi. org / 10.1206 / 0003 - 0090 (2000) 254 % 3 C 0001: NWPSAP % 3 E 2.0. CO; 2" type="journal article" year="2000">Huber 2000</bibRefCitation>
: figs 383, 387 with
<figureCitation id="137A2A3EFFE97A38FC69FCF3FE11FC2D" box="[994,1098,892,919]" captionStart="Figs 178182" captionStartId="59.[189,241,1780,1806]" captionTargetBox="[192,1392,289,1696]" captionTargetId="figure-82@59.[189,1398,289,1718]" captionTargetPageId="59" captionText="Figs 178182. Galapa spiniphila Huber sp. nov.; from Falcón, near Cueva del Guano (type locality; ZFMK Ar 21860). 178179. Left male pedipalp, prolateral and retrolateral views. 180. Left procursus and genital bulb, dorsal view. 181. Male ocular area, clypeus, and chelicerae, frontal view (arrow: process on male cheliceral fang). 182. Cleared female genitalia, dorsal view. Abbreviations: b= genital bulb; p=procursus. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4069648" httpUri="https://zenodo.org/record/4069648/files/figure.png" pageId="57" pageNumber="58">Fig. 180</figureCitation>
). In addition, the general morphology of the proximal palpal segments is similar (slender femur, large tibia), and the genital bulb of the new species below strongly resembles the bulb of
<taxonomicName id="4C414D38FFE97A38FCE0FC48FEA5FC5A" authority="(Gertsch &amp; Peck, 1992)" baseAuthorityName="Gertsch &amp; Peck" baseAuthorityYear="1992" box="[875,1278,966,993]" class="Arachnida" family="Pholcidae" genus="Galapa" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="baerti">
<emphasis id="B935EAA9FFE97A38FCE0FC48F98CFC5A" box="[875,983,966,993]" italics="true" pageId="57" pageNumber="58">G. baerti</emphasis>
(
<bibRefCitation id="EFD04B4AFFE97A38FC62FC49FEADFC5A" author="Gertsch W. J. &amp; Peck S. B." box="[1001,1270,966,992]" pageId="57" pageNumber="58" pagination="1185 - 1199" refId="ref127074" refString="Gertsch W. J. &amp; Peck S. B. 1992. The pholcid spiders of the Galapagos Islands, Ecuador (Araneae: Pholcidae). Canadian Journal of Zoology 70: 1185 - 1199. https: // doi. org / 10.1139 / z 92 - 166" type="journal article" year="1992">Gertsch &amp; Peck, 1992</bibRefCitation>
)
</taxonomicName>
(compare
<bibRefCitation id="EFD04B4AFFE97A38FF36FC64FB16FBBF" author="Huber B. A." box="[189,333,1003,1029]" pageId="57" pageNumber="58" pagination="1 - 348" refId="ref127973" refString="Huber B. A. 2000. New World pholcid spiders (Araneae: Pholcidae): a revision at generic level. Bulletin of the American Museum of Natural History 254: 1 - 348. https: // doi. org / 10.1206 / 0003 - 0090 (2000) 254 % 3 C 0001: NWPSAP % 3 E 2.0. CO; 2" type="journal article" year="2000">Huber 2000</bibRefCitation>
: fig. 381 with
<figureCitation id="137A2A3EFFE97A38FE7FFC64F801FBBF" box="[500,602,1003,1029]" captionStart="Figs 178182" captionStartId="59.[189,241,1780,1806]" captionTargetBox="[192,1392,289,1696]" captionTargetId="figure-82@59.[189,1398,289,1718]" captionTargetPageId="59" captionText="Figs 178182. Galapa spiniphila Huber sp. nov.; from Falcón, near Cueva del Guano (type locality; ZFMK Ar 21860). 178179. Left male pedipalp, prolateral and retrolateral views. 180. Left procursus and genital bulb, dorsal view. 181. Male ocular area, clypeus, and chelicerae, frontal view (arrow: process on male cheliceral fang). 182. Cleared female genitalia, dorsal view. Abbreviations: b= genital bulb; p=procursus. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4069648" httpUri="https://zenodo.org/record/4069648/files/figure.png" pageId="57" pageNumber="58">Fig. 178</figureCitation>
).
</paragraph>
<paragraph id="8BFE36BBFFE97A38FF36FBA1FAA7FB61" blockId="57.[189,1399,1070,1243]" pageId="57" pageNumber="58">
This suggests that
<taxonomicName id="4C414D38FFE97A38FE1CFBA1FBB4FBF2" authorityName="Huber" authorityYear="2000" box="[407,495,1070,1096]" class="Arachnida" family="Pholcidae" genus="Galapa" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B935EAA9FFE97A38FE1CFBA1FBB4FBF2" box="[407,495,1070,1096]" italics="true" pageId="57" pageNumber="58">Galapa</emphasis>
</taxonomicName>
may in fact be a widespread genus. An undescribed species from
<collectingCountry id="F356762BFFE97A38FB7DFBA1FF2CFBF2" box="[1270,1399,1070,1096]" name="Costa Rica" pageId="57" pageNumber="58">Costa Rica</collectingCountry>
(
<collectingRegion id="4985F859FFE97A38FF4EFBDCFB0AFBD7" box="[197,337,1107,1133]" country="Costa Rica" name="Guanacaste" pageId="57" pageNumber="58">Guanacaste</collectingRegion>
, no precise locality data; deposited in ZFMK) may also belong in this group and thus support this view. It seems to lack a dorsal process on the procursus but shares the unique cheliceral morphology and is also otherwise very similar. A formal test of the monophyly of the genus does
<taxonomicName id="4C414D38FFE97A38FADAFB12FF2DFB0D" box="[1361,1398,1181,1207]" class="Not" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="57" pageNumber="241" phylum="Arthropoda" rank="class">not</taxonomicName>
exist.
</paragraph>
</subSubSection>
</treatment>
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