treatments-xml/data/F1/6B/FB/F16BFB33FFFBFFA2FF6AFC09FF1AFA5D.xml
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<document id="C297B58659513F28A54B6BF5934F1A5A" ID-CLB-Dataset="37321" ID-DOI="10.11646/zootaxa.4209.1.1" ID-GBIF-Dataset="2de5b7bf-3975-458a-b200-07ec69f4e013" ID-ISSN="1175-5326" ID-Zenodo-Dep="272646" ID-ZooBank="849BAB5C-464A-4B4A-A586-5742411EDC01" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484407904318" checkinUser="plazi" docAuthor="Ng, Peter K. L. &amp; Castro, Peter" docDate="2016" docId="F16BFB33FFFBFFA2FF6AFC09FF1AFA5D" docLanguage="en" docName="zootaxa.4209.1.1.pdf.imf" docOrigin="Zootaxa 4209 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Megaesthesius sagedae Rathbun 1909" docType="treatment" docVersion="8" lastPageNumber="90" masterDocId="0D52834BFFA3FFFBFFFDFFD1FFCDFFB9" masterDocTitle="Revision of the family Chasmocarcinidae Serène, 1964 (Crustacea, Brachyura, Goneplacoidea)" masterLastPageNumber="182" masterPageNumber="1" pageNumber="89" updateTime="1698435566492" updateUser="ExternalLinkService">
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<mods:title id="8085A81FE963732CE24698E902CC2FA3">Revision of the family Chasmocarcinidae Serène, 1964 (Crustacea, Brachyura, Goneplacoidea)</mods:title>
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<mods:namePart id="FB0E6398CE97A21A37A9BC512D30EB1A">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="F926F28F6C1B5C9654A722100284D09F">Castro, Peter</mods:namePart>
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<treatment id="F16BFB33FFFBFFA2FF6AFC09FF1AFA5D" ID-DOI="http://doi.org/10.5281/zenodo.5617227" ID-GBIF-Taxon="127419537" ID-Zenodo-Dep="5617227" LSID="urn:lsid:plazi:treatment:F16BFB33FFFBFFA2FF6AFC09FF1AFA5D" httpUri="http://treatment.plazi.org/id/F16BFB33FFFBFFA2FF6AFC09FF1AFA5D" lastPageId="89" lastPageNumber="90" pageId="88" pageNumber="89">
<subSubSection id="31D819AEFFFBFFA3FF6AFC09FC47FBAA" pageId="88" pageNumber="89" type="nomenclature">
<paragraph id="797D4A25FFFBFFA3FF6AFC09FDA2FC4B" blockId="88.[151,906,984,1043]" box="[151,623,984,1010]" pageId="88" pageNumber="89">
<heading id="2235FD49FFFBFFA3FF6AFC09FDA2FC4B" bold="true" box="[151,623,984,1010]" fontSize="11" level="1" pageId="88" pageNumber="89" reason="1">
<taxonomicName id="BEC231A6FFFBFFA3FF6AFC09FDA2FC4B" ID-CoL="3YTMZ" authority="Rathbun, 1909" authorityName="Rathbun" authorityYear="1909" box="[151,623,984,1010]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sagedae">
<emphasis id="4BB69637FFFBFFA3FF6AFC09FDA2FC4B" bold="true" box="[151,623,984,1010]" pageId="88" pageNumber="89">
<emphasis id="4BB69637FFFBFFA3FF6AFC09FE7DFC4B" bold="true" box="[151,432,984,1010]" italics="true" pageId="88" pageNumber="89">Megaesthesius sagedae</emphasis>
<bibRefCitation id="1D5337D4FFFBFFA3FE4BFC09FDA2FC4B" author="Rathbun" box="[438,623,984,1010]" pageId="88" pageNumber="89" refString="Rathbun, M. J. (1909) New crabs from the Gulf of Siam. Proceedings of the Biological Society of Washington, 22, 107 - 114." type="journal article" year="1909">Rathbun, 1909</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="797D4A25FFFBFFA3FF6AFC2BFC47FBAA" blockId="88.[151,906,984,1043]" box="[151,906,1018,1043]" pageId="88" pageNumber="89">
(
<figureCitation id="E1F956A0FFFBFFA3FF62FC2BFF37FBAA" box="[159,250,1018,1043]" captionStart="FIGURE 19" captionStartId="92.[151,250,1665,1687]" captionTargetBox="[203,1371,195,1618]" captionTargetId="figure@92.[203,1371,195,1632]" captionTargetPageId="92" captionText="FIGURE 19. Species of Megaesthesius and Alainthesius n. gen. A, B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); C F, Megaesthesius yokoyai (after Takeda &amp; Miyake 1969 a: figs. 3 a, 4 f, 4 a, 4 e, respectively); G L, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472) (after Crosnier 1975: fig. 1 a, 1 k, 1 j, 1 b, 1 c, 1 i, respectively); M R, Megaesthesius westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: figs. 1 A, B, 2 A, D, E, 3 E); S, W, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); T, X, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); U, Y, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); V, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012). A, overall habitus; B, C, G, M, dorsal view of carapace; E, left P 4; F, I, frontal view of cephalothorax; H, O, frontal and anterolateral margin of carapace; N, ventral view of cephalothorax; J, right P 3; K, buccal cavity and posterior margin of epistome; Q, right P 4; R, right P 5; D, L, P, male pleon; S V, posterior margin of epistome; W Y, left thoracic sternites 7 and 8, “ supplementary plate ”, penis tube and penis. Abbreviations: cx 4 = coxa of P 4; cx 5 = coxa of P 5; p = penis; pt = calcified penis tube; sp = “ supplementary plate ”; st 7 = thoracic sternite 7; st 8 = thoracic sternite 8." httpUri="https://zenodo.org/record/272665/files/figure.png" pageId="88" pageNumber="89">Figs. 19</figureCitation>
A, B, S, W; 27B; 34A; 48C; 55C; 62B; 82AC; 88F; 92B)
</paragraph>
</subSubSection>
<subSubSection id="31D819AEFFFBFFA3FF6AFB93FBC1FB0D" pageId="88" pageNumber="89" type="reference_group">
<paragraph id="797D4A25FFFBFFA3FF6AFB93FA50FBE1" blockId="88.[151,1437,1090,1205]" box="[151,1437,1090,1112]" pageId="88" pageNumber="89">
<treatmentCitationGroup id="59D26D0BFFFBFFA3FF6AFB93FA50FBE1" box="[151,1437,1090,1112]" pageId="88" pageNumber="89">
<taxonomicName id="BEC231A6FFFBFFA3FF6AFB93FD96FBE1" ID-CoL="3YTMZ" authority="Rathbun, 1909: 112" authorityName="Rathbun" authorityPageNumber="112" authorityYear="1909" box="[151,603,1090,1112]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sagedae">
<emphasis id="4BB69637FFFBFFA3FF6AFB93FE4AFBE1" box="[151,391,1090,1112]" italics="true" pageId="88" pageNumber="89">Megaesthesius sagedae</emphasis>
<treatmentCitation id="F8636C34FFFBFFA3FE72FB93FD96FBE1" author="Rathbun" box="[399,603,1090,1112]" page="112" pageId="88" pageNumber="89" year="1909">
<bibRefCitation id="1D5337D4FFFBFFA3FE72FB93FDE7FBE1" author="Rathbun" box="[399,554,1090,1112]" pageId="88" pageNumber="89" refString="Rathbun, M. J. (1909) New crabs from the Gulf of Siam. Proceedings of the Biological Society of Washington, 22, 107 - 114." type="journal article" year="1909">Rathbun, 1909</bibRefCitation>
: 112
</treatmentCitation>
</taxonomicName>
[type locality: Singapore]; 1910: 344, figs. 30, 31, pl. 2, fig. 5.—
<bibRefCitation id="1D5337D4FFFBFFA3FAF8FB93FA55FBE1" author="Serene" box="[1285,1432,1090,1112]" pageId="88" pageNumber="89" refString="Serene, R. (1964 b) Redescription du genre Megaesthesius Rathbun et definition des Chasmocarcininae, nouvelle sous-famille des Goneplacidae (Decapoda Brachyura). Crustaceana, 7 (3), 175 - 187." type="journal article" year="1964" yearSuffix="b">Serène 1964b</bibRefCitation>
:
</treatmentCitationGroup>
</paragraph>
<paragraph id="797D4A25FFFBFFA3FF3AFBB0FBC1FB0D" blockId="88.[151,1437,1090,1205]" pageId="88" pageNumber="89">
176, figs. 16 [South China Sea]; 1968: 92 [in list].—
<bibRefCitation id="1D5337D4FFFBFFA3FD22FBB0FC4DFBCE" author="Zarenkov" box="[735,896,1121,1143]" pageId="88" pageNumber="89" refString="Zarenkov, N. A. (1972) [New data on Indo-Pacific crabs (Fam. Goneplacidae, Pinnotheridae, Parthenopidae, Dorippidae) and the problem of seasonal reproduction of decapod animals in the Bay of Tonkin]. In: [The Complex Investigations of Ocean Nature], part 3, Moscow State University, Moscow, pp. 229 - 253. [In Russian]" type="book chapter" year="1972">Zarenkov 1972</bibRefCitation>
: 241, fig. 6-1 [Vietnam].—Ng
<emphasis id="4BB69637FFFBFFA3FB4BFBB3FB24FBCE" box="[1206,1257,1121,1143]" italics="true" pageId="88" pageNumber="89">et al.</emphasis>
2008: 76 [in list].?
<taxonomicName id="BEC231A6FFFBFFA3FF55FB51FE5AFB2F" ID-CoL="3YTMZ" box="[168,407,1152,1174]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sagedae">
<emphasis id="4BB69637FFFBFFA3FF55FB51FE5AFB2F" box="[168,407,1152,1174]" italics="true" pageId="88" pageNumber="89">Megaesthesius sagedae</emphasis>
</taxonomicName>
<bibRefCitation id="1D5337D4FFFBFFA3FE4CFB51FD5FFB2F" author="Serene" box="[433,658,1152,1174]" pageId="88" pageNumber="89" refString="Serene, R. &amp; Vadon, C. (1981) Crustaces decapodes: brachyoures. Liste preliminaire, description de formes nouvelles et remarques taxonomiques, In: Resultats des campagnes MUSORSTOM I. Philippines (18 - 28 mars 1976), vol. 1. Memoires ORSTOM (Paris), 91, 117 - 140." type="book chapter" year="1981">Serène &amp; Vadon 1981</bibRefCitation>
: 118, 119, 123 [Philippines].—Ng &amp; Davie 2002: 378 [Western Thailand?] Not
<taxonomicName id="BEC231A6FFFBFFA3FF3EFB4FFE7CFB0D" ID-CoL="3YTMZ" box="[195,433,1182,1204]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sagedae">
<emphasis id="4BB69637FFFBFFA3FF3EFB4FFE7CFB0D" box="[195,433,1182,1204]" italics="true" pageId="88" pageNumber="89">Megaesthesius sagedae</emphasis>
</taxonomicName>
—Davie 2013: 475, figs. 3JN, 4. [=
<taxonomicName id="BEC231A6FFFBFFA3FCD6FB4FFBCEFB0C" ID-CoL="666GD" box="[811,1027,1182,1205]" class="Malacostraca" family="Chasmocarcinidae" genus="Alainthesius" kingdom="Animalia" order="Decapoda" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="signatus">
<emphasis id="4BB69637FFFBFFA3FCD6FB4FFBCEFB0C" box="[811,1027,1182,1205]" italics="true" pageId="88" pageNumber="89">Alainthesius signatus</emphasis>
</taxonomicName>
]
</paragraph>
</subSubSection>
<subSubSection id="31D819AEFFFBFFA3FF6AFB30FE20FA17" pageId="88" pageNumber="89" type="materials_examined">
<paragraph id="797D4A25FFFBFFA3FF6AFB30FE56FAA4" blockId="88.[151,1437,1249,2030]" pageId="88" pageNumber="89">
<materialsCitation id="C9AA4078FFFBFFA3FF6AFB30FE56FAA4" ID-GBIF-Occurrence="1424726212" collectingDate="1899" collectingDateMax="1900" collectingDateMin="1899" collectionCode="ZMUC" country="Singapore" elevation="5" location="Mortensen Expedition" pageId="88" pageNumber="89" specimenCount="2" typeStatus="holotype">
<emphasis id="4BB69637FFFBFFA3FF6AFB30FEF0FB43" bold="true" box="[151,317,1249,1274]" pageId="88" pageNumber="89">
<typeStatus id="A679F487FFFBFFA3FF6AFB30FF1FFB43" box="[151,210,1249,1274]" pageId="88" pageNumber="89">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="A679F487FFFBFFA3FEB6FB30FE7EFB43" box="[331,435,1249,1274]" pageId="88" pageNumber="89" type="holotype">Holotype</typeStatus>
male (2.5 ×
<quantity id="BE3AE7C0FFFBFFA3FDBCFB33FD55FB43" box="[577,664,1250,1274]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="88" pageNumber="89" unit="mm" value="2.8">2.8 mm</quantity>
) (
<collectionCode id="1FD3D2E0FFFBFFA3FD53FB33FCCFFB43" box="[686,770,1250,1274]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="88" pageNumber="89">ZMUC</collectionCode>
) [no pleon or gonopods],
<collectingCountry id="01D50AB5FFFBFFA3FBDEFB30FB55FB43" box="[1059,1176,1249,1274]" name="Singapore" pageId="88" pageNumber="89">Singapore</collectingCountry>
,
<location id="7C1D1CFEFFFBFFA3FB5EFB33FA51FB43" LSID="urn:lsid:plazi:treatment:F16BFB33FFFBFFA2FF6AFC09FF1AFA5D:7C1D1CFEFFFBFFA3FB5EFB33FA51FB43" box="[1187,1436,1249,1274]" country="Singapore" name="Mortensen Expedition" pageId="88" pageNumber="89">Mortensen Expedition</location>
<collectingDate id="1D38950DFFFBFFA3FF6AFAD7FED5FAA7" box="[151,280,1286,1310]" pageId="88" pageNumber="89" value="1899" valueMax="1900">
<date id="0D7C6CE5FFFBFFA3FF6AFAD7FED5FAA7" box="[151,280,1286,1310]" pageId="88" pageNumber="89" value="1899" valueMax="1900">18991900</date>
</collectingDate>
,
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<quantity id="BE3AE7C0FFFBFFA3FEDEFAD4FE5AFAA4" box="[291,407,1285,1310]" metricMagnitude="0" metricUnit="m" metricValue="4.55" metricValueMax="5.5" metricValueMin="3.6" pageId="88" pageNumber="89" unit="m" value="4.55" valueMax="5.5" valueMin="3.6">3.65.5 m</quantity>
</elevation>
.
</materialsCitation>
</paragraph>
<paragraph id="797D4A25FFFBFFA3FF3AFAF8FD10FADF" blockId="88.[151,1437,1249,2030]" pageId="88" pageNumber="89">
<emphasis id="4BB69637FFFBFFA3FF3AFAF8FE34FAFB" bold="true" box="[199,505,1321,1346]" pageId="88" pageNumber="89">Other material examined</emphasis>
.
<emphasis id="4BB69637FFFBFFA3FDF7FAFBFDA8FAF8" box="[522,613,1322,1345]" italics="true" pageId="88" pageNumber="89">Vietnam</emphasis>
.
<materialsCitation id="C9AA4078FFFBFFA3FD8BFAFBFD10FADF" ID-GBIF-Occurrence="1424726208" collectingDate="1963-04-22" collectorName="Orlik, N. &amp; Zarenkov" country="Vietnam" location="Gulf" pageId="88" pageNumber="89" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Gulf">
<specimenCount id="6FC481ACFFFBFFA3FD8BFAFBFD0BFAFB" box="[630,710,1321,1346]" pageId="88" pageNumber="89" type="male">1 male</specimenCount>
,
<specimenCount id="6FC481ACFFFBFFA3FD28FAFBFCF5FAFB" box="[725,824,1321,1346]" pageId="88" pageNumber="89" type="female">1 female</specimenCount>
(MNHN-IU-2013-9473 [= MNHN-B10215),
<collectingRegion id="BB0684C7FFFBFFA3FAB5FAF8FAB6FAFB" box="[1352,1403,1321,1346]" country="Papua New Guinea" name="Gulf" pageId="88" pageNumber="89">Gulf</collectingRegion>
of
<collectingCountry id="01D50AB5FFFBFFA3FF6AFA9CFF24FADF" box="[151,233,1357,1382]" name="Vietnam" pageId="88" pageNumber="89">Tonkin</collectingCountry>
, trawl,
<collectorName id="D4372FF3FFFBFFA3FEC6FA9CFDDFFADF" box="[315,530,1357,1382]" pageId="88" pageNumber="89">
<emphasis id="4BB69637FFFBFFA3FEC6FA9CFEBBFADF" box="[315,374,1357,1382]" italics="true" pageId="88" pageNumber="89">Orlik</emphasis>
, N. Zarenkov
</collectorName>
coll.,
<collectingDate id="1D38950DFFFBFFA3FDA5FA9CFD14FADF" box="[600,729,1357,1382]" pageId="88" pageNumber="89" value="1963-04-22">
<date id="0D7C6CE5FFFBFFA3FDA5FA9CFD14FADF" box="[600,729,1357,1382]" pageId="88" pageNumber="89" value="1963-04-22">22.04.1963</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
<paragraph id="797D4A25FFFBFFA3FF3AFAA0FE20FA17" blockId="88.[151,1437,1249,2030]" pageId="88" pageNumber="89">
<emphasis id="4BB69637FFFBFFA3FF3AFAA0FEE2FA33" box="[199,303,1393,1418]" italics="true" pageId="88" pageNumber="89">Malaysia</emphasis>
.
<materialsCitation id="C9AA4078FFFBFFA3FEC2FAA3FE20FA17" ID-GBIF-Occurrence="1424726196" collectionCode="ZRC" collectorName="Bintulu &amp; T. S. Leong" country="Malaysia" location="Sarawak" pageId="88" pageNumber="89" specimenCode="ZRC 2015.303" specimenCount="1" specimenCount-male="1">
<specimenCount id="6FC481ACFFFBFFA3FEC2FAA3FE41FA33" box="[319,396,1393,1418]" pageId="88" pageNumber="89" type="male">1 male</specimenCount>
2.3 ×
<quantity id="BE3AE7C0FFFBFFA3FE27FAA3FDFFFA30" box="[474,562,1394,1418]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="88" pageNumber="89" unit="mm" value="2.8">2.8 mm</quantity>
(
<specimenCode id="2964E25EFFFBFFA3FDBEFAA3FD3CFA33" box="[579,753,1393,1418]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="88" pageNumber="89">
<collectionCode id="1FD3D2E0FFFBFFA3FDBEFAA3FDB6FA33" box="[579,635,1394,1418]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="88" pageNumber="89">ZRC</collectionCode>
2015.303
</specimenCode>
), stn F28 oilfield, offshore
<collectorName id="D4372FF3FFFBFFA3FBD6FAA0FB4FFA33" box="[1067,1154,1393,1418]" pageId="88" pageNumber="89">Bintulu</collectorName>
, northern
<collectingCountry id="01D50AB5FFFBFFA3FB0AFAA0FA97FA33" box="[1271,1370,1393,1418]" name="Malaysia" pageId="88" pageNumber="89">Sarawak</collectingCountry>
, deep water,
<collectorName id="D4372FF3FFFBFFA3FF1FFA47FE90FA17" box="[226,349,1430,1454]" pageId="88" pageNumber="89">T.S. Leong</collectorName>
coll., 2000s.
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</paragraph>
</subSubSection>
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<paragraph id="797D4A25FFFBFFA3FF3AFA6BFB3FF913" blockId="88.[151,1437,1249,2030]" pageId="88" pageNumber="89">
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<collectingCountry id="01D50AB5FFFBFFA3FF3AFA6BFEF4FA68" box="[199,313,1466,1489]" name="Singapore" pageId="88" pageNumber="89">
<emphasis id="4BB69637FFFBFFA3FF3AFA6BFEF4FA68" box="[199,313,1466,1489]" italics="true" pageId="88" pageNumber="89">Singapore</emphasis>
</collectingCountry>
.
<specimenCount id="6FC481ACFFFBFFA3FEBAFA6BFE5FFA6B" box="[327,402,1465,1490]" pageId="88" pageNumber="89" type="male">1 male</specimenCount>
(2.4 ×
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)
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, 1 female (cl
<quantity id="BE3AE7C0FFFBFFA3FD23FA6BFCFBFA68" box="[734,822,1466,1490]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="88" pageNumber="89" unit="mm" value="2.9">2.9 mm</quantity>
, carapace damaged) (ZRC 1995.883), Pulau Semakau, station 2, grab 2, Reef Ecology Study Team,
<date id="0D7C6CE5FFFBFFA3FD6FFA0CFCDAFA4F" box="[658,791,1501,1526]" pageId="88" pageNumber="89" value="1995-08-06">08.06.1995</date>
.—1 male (2.5 × 3.0 mm) (ZRC 1995.885), Reef Ecology Study Team, Pulau Semakau, station 2, grab 4,
<date id="0D7C6CE5FFFBFFA3FD59F9D0FCE4F9A3" box="[676,809,1537,1562]" pageId="88" pageNumber="89" value="1995-08-06">08.06.1995</date>
.—
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<specimenCount id="6FC481ACFFFBFFA3FCB7F9D0FC42F9A3" box="[842,911,1537,1562]" pageId="88" pageNumber="89" type="male">1 male</specimenCount>
(2.2 ×
<quantity id="BE3AE7C0FFFBFFA3FC22F9D0FBFBF9A3" box="[991,1078,1537,1562]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="88" pageNumber="89" unit="mm" value="2.5">2.5 mm</quantity>
) (
<specimenCode id="2964E25EFFFBFFA3FBB6F9D3FB35F9A3" box="[1099,1272,1537,1562]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="88" pageNumber="89">
<collectionCode id="1FD3D2E0FFFBFFA3FBB6F9D3FB4EF9A3" box="[1099,1155,1538,1562]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="88" pageNumber="89">ZRC</collectionCode>
2015.264
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), station 5515- ORI-010, northeast of
<collectorName id="D4372FF3FFFBFFA3FE6AF9F7FDC5F984" box="[407,520,1574,1598]" pageId="88" pageNumber="89">
<location id="7C1D1CFEFFFBFFA3FE6AF9F7FE23F987" LSID="urn:lsid:plazi:treatment:F16BFB33FFFBFFA2FF6AFC09FF1AFA5D:7C1D1CFEFFFBFFA3FE6AF9F7FE23F987" box="[407,494,1574,1598]" country="Singapore" latitude="1.2595667" name="Sentosa" pageId="88" pageNumber="89">Sentosa</location>
I.
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,
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</materialsCitation>
01°15.831' N to 103°51.675'E103°51.524'E, rectangular dredge,
<quantity id="BE3AE7C0FFFBFFA3FF6AF99BFF31F9D8" box="[151,252,1610,1634]" metricMagnitude="1" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="88" pageNumber="89" unit="m" value="18.5" valueMax="19.0" valueMin="18.0">1819 m</quantity>
, mud, Comprehensive Marine Biodiversity Survey,
<date id="0D7C6CE5FFFBFFA3FCB9F99BFC04F9DB" box="[836,969,1610,1634]" pageId="88" pageNumber="89" value="2013-03-21">21.03.2013</date>
.—1 male (2.6 ×
<quantity id="BE3AE7C0FFFBFFA3FB7EF99BFB10F9DB" box="[1155,1245,1610,1634]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="88" pageNumber="89" unit="mm" value="2.9">2.9 mm</quantity>
), 1 female (3.0 ×
<quantity id="BE3AE7C0FFFBFFA3FF6AF9BCFF23F93F" box="[151,238,1645,1670]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="88" pageNumber="89" unit="mm" value="3.6">3.6 mm</quantity>
) (ZRC 2013.1804), Pulau Semakau, station PS25T2, Reef Ecology Study Team coll., 1990s; 1 female (3.3 × 4.0 mm), 1 female (3.3 ×
<quantity id="BE3AE7C0FFFBFFA3FE30F943FDE9F913" box="[461,548,1682,1706]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="88" pageNumber="89" unit="mm" value="3.7">3.7 mm</quantity>
) (QM), Pulau Semakau, Reef Ecology Study Team coll., 1990s.
</paragraph>
</subSubSection>
<subSubSection id="31D819AEFFFBFFA2FF3AF964FC5DFD49" lastPageId="89" lastPageNumber="90" pageId="88" pageNumber="89" type="diagnosis">
<paragraph id="797D4A25FFFBFFA3FF3AF964FA50F857" blockId="88.[151,1437,1249,2030]" pageId="88" pageNumber="89">
<emphasis id="4BB69637FFFBFFA3FF3AF964FEF7F977" bold="true" box="[199,314,1717,1742]" pageId="88" pageNumber="89">Diagnosis</emphasis>
. Small size. Carapace (
<figureCitation id="E1F956A0FFFBFFA3FDB0F964FD6DF977" box="[589,672,1717,1742]" captionStart="FIGURE 19" captionStartId="92.[151,250,1665,1687]" captionTargetBox="[203,1371,195,1618]" captionTargetId="figure@92.[203,1371,195,1632]" captionTargetPageId="92" captionText="FIGURE 19. Species of Megaesthesius and Alainthesius n. gen. A, B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); C F, Megaesthesius yokoyai (after Takeda &amp; Miyake 1969 a: figs. 3 a, 4 f, 4 a, 4 e, respectively); G L, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472) (after Crosnier 1975: fig. 1 a, 1 k, 1 j, 1 b, 1 c, 1 i, respectively); M R, Megaesthesius westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: figs. 1 A, B, 2 A, D, E, 3 E); S, W, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); T, X, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); U, Y, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); V, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012). A, overall habitus; B, C, G, M, dorsal view of carapace; E, left P 4; F, I, frontal view of cephalothorax; H, O, frontal and anterolateral margin of carapace; N, ventral view of cephalothorax; J, right P 3; K, buccal cavity and posterior margin of epistome; Q, right P 4; R, right P 5; D, L, P, male pleon; S V, posterior margin of epistome; W Y, left thoracic sternites 7 and 8, “ supplementary plate ”, penis tube and penis. Abbreviations: cx 4 = coxa of P 4; cx 5 = coxa of P 5; p = penis; pt = calcified penis tube; sp = “ supplementary plate ”; st 7 = thoracic sternite 7; st 8 = thoracic sternite 8." httpUri="https://zenodo.org/record/272665/files/figure.png" pageId="88" pageNumber="89">Fig. 19</figureCitation>
A, B;
<bibRefCitation id="1D5337D4FFFBFFA3FD17F967FC47F977" author="Serene" box="[746,906,1717,1742]" pageId="88" pageNumber="89" refString="Serene, R. (1964 b) Redescription du genre Megaesthesius Rathbun et definition des Chasmocarcininae, nouvelle sous-famille des Goneplacidae (Decapoda Brachyura). Crustaceana, 7 (3), 175 - 187." type="journal article" year="1964" yearSuffix="b">Serène 1964b</bibRefCitation>
: fig. 1) subquadrate, 1.11.2 wider than long; front bilobed, with well-defined median cleft; anterolateral margins straight, with numerous small, sharp teeth, spines; conspicuous granules through most of carapace, larger along posterolateral and anterior portions (
<figureCitation id="E1F956A0FFFBFFA3FACDF92CFA4CF8AF" box="[1328,1409,1789,1814]" captionStart="FIGURE 19" captionStartId="92.[151,250,1665,1687]" captionTargetBox="[203,1371,195,1618]" captionTargetId="figure@92.[203,1371,195,1632]" captionTargetPageId="92" captionText="FIGURE 19. Species of Megaesthesius and Alainthesius n. gen. A, B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); C F, Megaesthesius yokoyai (after Takeda &amp; Miyake 1969 a: figs. 3 a, 4 f, 4 a, 4 e, respectively); G L, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472) (after Crosnier 1975: fig. 1 a, 1 k, 1 j, 1 b, 1 c, 1 i, respectively); M R, Megaesthesius westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: figs. 1 A, B, 2 A, D, E, 3 E); S, W, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); T, X, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); U, Y, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); V, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012). A, overall habitus; B, C, G, M, dorsal view of carapace; E, left P 4; F, I, frontal view of cephalothorax; H, O, frontal and anterolateral margin of carapace; N, ventral view of cephalothorax; J, right P 3; K, buccal cavity and posterior margin of epistome; Q, right P 4; R, right P 5; D, L, P, male pleon; S V, posterior margin of epistome; W Y, left thoracic sternites 7 and 8, “ supplementary plate ”, penis tube and penis. Abbreviations: cx 4 = coxa of P 4; cx 5 = coxa of P 5; p = penis; pt = calcified penis tube; sp = “ supplementary plate ”; st 7 = thoracic sternite 7; st 8 = thoracic sternite 8." httpUri="https://zenodo.org/record/272665/files/figure.png" pageId="88" pageNumber="89">Fig. 19</figureCitation>
A, B). Orbits long. Epistome (
<figureCitation id="E1F956A0FFFBFFA3FE2EF8F0FDF9F883" box="[467,564,1825,1850]" captionStart="FIGURE 19" captionStartId="92.[151,250,1665,1687]" captionTargetBox="[203,1371,195,1618]" captionTargetId="figure@92.[203,1371,195,1632]" captionTargetPageId="92" captionText="FIGURE 19. Species of Megaesthesius and Alainthesius n. gen. A, B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); C F, Megaesthesius yokoyai (after Takeda &amp; Miyake 1969 a: figs. 3 a, 4 f, 4 a, 4 e, respectively); G L, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472) (after Crosnier 1975: fig. 1 a, 1 k, 1 j, 1 b, 1 c, 1 i, respectively); M R, Megaesthesius westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: figs. 1 A, B, 2 A, D, E, 3 E); S, W, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); T, X, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); U, Y, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); V, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012). A, overall habitus; B, C, G, M, dorsal view of carapace; E, left P 4; F, I, frontal view of cephalothorax; H, O, frontal and anterolateral margin of carapace; N, ventral view of cephalothorax; J, right P 3; K, buccal cavity and posterior margin of epistome; Q, right P 4; R, right P 5; D, L, P, male pleon; S V, posterior margin of epistome; W Y, left thoracic sternites 7 and 8, “ supplementary plate ”, penis tube and penis. Abbreviations: cx 4 = coxa of P 4; cx 5 = coxa of P 5; p = penis; pt = calcified penis tube; sp = “ supplementary plate ”; st 7 = thoracic sternite 7; st 8 = thoracic sternite 8." httpUri="https://zenodo.org/record/272665/files/figure.png" pageId="88" pageNumber="89">Figs. 19</figureCitation>
S; 27B) compressed, posterior margin with broad median lobe with fissure, lateral margins short, straight. Eye peduncle conspicuously long, immobile, cornea long (
<figureCitation id="E1F956A0FFFBFFA3FB7AF894FB16F8E7" box="[1159,1243,1861,1886]" captionStart="FIGURE 27" captionStartId="100.[151,250,1447,1469]" captionTargetBox="[151,1436,193,1426]" captionTargetId="figure@100.[151,1436,193,1426]" captionTargetPageId="100" captionText="FIGURE 27. Frontal view of species of Megaesthesius and Microtopsis. A, Microtopsis takedai, Japan: paratype female 3.6 × 2.7 mm (ZRC 2012.0146); B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); C, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); D, Alainthesius signatus n. sp., Madagascar: paratype female 2.7 × 3.0 mm (MNHN-IU- 2013 - 9471); E, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); F, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012)." httpUri="https://zenodo.org/record/272673/files/figure.png" pageId="88" pageNumber="89">Fig. 27</figureCitation>
B;
<bibRefCitation id="1D5337D4FFFBFFA3FB07F897FA54F8E7" author="Serene" box="[1274,1433,1861,1886]" pageId="88" pageNumber="89" refString="Serene, R. (1964 b) Redescription du genre Megaesthesius Rathbun et definition des Chasmocarcininae, nouvelle sous-famille des Goneplacidae (Decapoda Brachyura). Crustaceana, 7 (3), 175 - 187." type="journal article" year="1964" yearSuffix="b">Serène 1964b</bibRefCitation>
: fig. 2), pigmentation reduced. Antennule (
<figureCitation id="E1F956A0FFFBFFA3FD87F8B8FD1AF83B" box="[634,727,1897,1922]" captionStart="FIGURE 19" captionStartId="92.[151,250,1665,1687]" captionTargetBox="[203,1371,195,1618]" captionTargetId="figure@92.[203,1371,195,1632]" captionTargetPageId="92" captionText="FIGURE 19. Species of Megaesthesius and Alainthesius n. gen. A, B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); C F, Megaesthesius yokoyai (after Takeda &amp; Miyake 1969 a: figs. 3 a, 4 f, 4 a, 4 e, respectively); G L, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472) (after Crosnier 1975: fig. 1 a, 1 k, 1 j, 1 b, 1 c, 1 i, respectively); M R, Megaesthesius westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: figs. 1 A, B, 2 A, D, E, 3 E); S, W, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); T, X, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); U, Y, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); V, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012). A, overall habitus; B, C, G, M, dorsal view of carapace; E, left P 4; F, I, frontal view of cephalothorax; H, O, frontal and anterolateral margin of carapace; N, ventral view of cephalothorax; J, right P 3; K, buccal cavity and posterior margin of epistome; Q, right P 4; R, right P 5; D, L, P, male pleon; S V, posterior margin of epistome; W Y, left thoracic sternites 7 and 8, “ supplementary plate ”, penis tube and penis. Abbreviations: cx 4 = coxa of P 4; cx 5 = coxa of P 5; p = penis; pt = calcified penis tube; sp = “ supplementary plate ”; st 7 = thoracic sternite 7; st 8 = thoracic sternite 8." httpUri="https://zenodo.org/record/272665/files/figure.png" pageId="88" pageNumber="89">Figs. 19</figureCitation>
A, B; 27B) greatly enlarged (particularly in males), cannot be folded. Third maxillipeds (
<figureCitation id="E1F956A0FFFBFFA3FE24F85CFDFEF81F" box="[473,563,1933,1958]" captionStart="FIGURE 34" captionStartId="107.[151,250,1457,1479]" captionTargetBox="[188,1388,200,1429]" captionTargetId="figure@107.[151,1436,193,1456]" captionTargetPageId="107" captionText="FIGURE 34. Third maxilliped of species of Megaesthesius and Alainthesius n. gen. A, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); B, Megaesthesius yokoyai, Japan: male 3.5 × 3.8 mm (after Takeda &amp; Miyake 1969: fig. 4 c); C, Megaethesius westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: fig. 2 C); D, Megaesthesius migmus n. sp., Philippines: paratype female 4.6 × 5.2 mm (ZRC 2015.243); E, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (after Crosnier 1975: fig. 1 e); F, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073). Scales = 0.5 mm." httpUri="https://zenodo.org/record/272680/files/figure.png" pageId="88" pageNumber="89">Fig. 34</figureCitation>
A;
<bibRefCitation id="1D5337D4FFFBFFA3FDA6F85CFCDCF81F" author="Zarenkov" box="[603,785,1933,1958]" pageId="88" pageNumber="89" refString="Zarenkov, N. A. (1972) [New data on Indo-Pacific crabs (Fam. Goneplacidae, Pinnotheridae, Parthenopidae, Dorippidae) and the problem of seasonal reproduction of decapod animals in the Bay of Tonkin]. In: [The Complex Investigations of Ocean Nature], part 3, Moscow State University, Moscow, pp. 229 - 253. [In Russian]" type="book chapter" year="1972">Zarenkov 1972</bibRefCitation>
: 6-
<date id="0D7C6CE5FFFBFFA3FCC7F85CFCA8F81F" box="[826,869,1933,1958]" pageId="88" pageNumber="89">1III</date>
) nearly fill buccal cavern when closed; merus subtriangular, outer margin convex, no defined anteroexternal angle; ischium subquadrate, about same length than merus; outer margins of ischium, merus with small spines and teeth. Chelipeds (
<figureCitation id="E1F956A0FFFBFFA3FBE9F804FBA2F857" box="[1044,1135,2005,2030]" captionStart="FIGURE 19" captionStartId="92.[151,250,1665,1687]" captionTargetBox="[203,1371,195,1618]" captionTargetId="figure@92.[203,1371,195,1632]" captionTargetPageId="92" captionText="FIGURE 19. Species of Megaesthesius and Alainthesius n. gen. A, B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); C F, Megaesthesius yokoyai (after Takeda &amp; Miyake 1969 a: figs. 3 a, 4 f, 4 a, 4 e, respectively); G L, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472) (after Crosnier 1975: fig. 1 a, 1 k, 1 j, 1 b, 1 c, 1 i, respectively); M R, Megaesthesius westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: figs. 1 A, B, 2 A, D, E, 3 E); S, W, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); T, X, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); U, Y, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); V, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012). A, overall habitus; B, C, G, M, dorsal view of carapace; E, left P 4; F, I, frontal view of cephalothorax; H, O, frontal and anterolateral margin of carapace; N, ventral view of cephalothorax; J, right P 3; K, buccal cavity and posterior margin of epistome; Q, right P 4; R, right P 5; D, L, P, male pleon; S V, posterior margin of epistome; W Y, left thoracic sternites 7 and 8, “ supplementary plate ”, penis tube and penis. Abbreviations: cx 4 = coxa of P 4; cx 5 = coxa of P 5; p = penis; pt = calcified penis tube; sp = “ supplementary plate ”; st 7 = thoracic sternite 7; st 8 = thoracic sternite 8." httpUri="https://zenodo.org/record/272665/files/figure.png" pageId="88" pageNumber="89">Figs. 19</figureCitation>
A; 48C;
<bibRefCitation id="1D5337D4FFFBFFA3FB32F807FAA1F857" author="Serene" box="[1231,1388,2005,2030]" pageId="88" pageNumber="89" refString="Serene, R. (1964 b) Redescription du genre Megaesthesius Rathbun et definition des Chasmocarcininae, nouvelle sous-famille des Goneplacidae (Decapoda Brachyura). Crustaceana, 7 (3), 175 - 187." type="journal article" year="1964" yearSuffix="b">Serène 1964b</bibRefCitation>
: fig.
</paragraph>
<paragraph id="797D4A25FFFAFFA2FF6AFF49FC5DFD49" blockId="89.[151,1437,151,1508]" pageId="89" pageNumber="90">
1;
<bibRefCitation id="1D5337D4FFFAFFA2FF49FF46FEAEFF09" author="Zarenkov" box="[180,355,151,176]" pageId="89" pageNumber="90" refString="Zarenkov, N. A. (1972) [New data on Indo-Pacific crabs (Fam. Goneplacidae, Pinnotheridae, Parthenopidae, Dorippidae) and the problem of seasonal reproduction of decapod animals in the Bay of Tonkin]. In: [The Complex Investigations of Ocean Nature], part 3, Moscow State University, Moscow, pp. 229 - 253. [In Russian]" type="book chapter" year="1972">Zarenkov 1972</bibRefCitation>
: 6-
<date id="0D7C6CE5FFFAFFA2FE7AFF46FE50FF09" box="[391,413,151,176]" pageId="89" pageNumber="90">1I</date>
) nearly identical in both sexes; cutting margins of both chelae with broad, shallow teeth in both sexes; small chela laterally flattened, fingers short, prominently flattened laterally, cutting margins blade-like; ventral margin of palm dentiform; ventral surface of cheliped merus with smooth or with obtuse tubercles. Inner margin of cheliped carpus with small acute tooth. Ventral surface of cheliped merus smooth in both sexes. Proportionally long ambulatory legs (
<bibRefCitation id="1D5337D4FFFAFFA2FDC7FEF9FCD3FEF9" author="Serene" box="[570,798,295,320]" pageId="89" pageNumber="90" refString="Serene, R. (1964 b) Redescription du genre Megaesthesius Rathbun et definition des Chasmocarcininae, nouvelle sous-famille des Goneplacidae (Decapoda Brachyura). Crustaceana, 7 (3), 175 - 187." type="journal article" year="1964" yearSuffix="b">Serène 1964b: fig. 1</bibRefCitation>
), P5 merus not reaching front when folded. Minute teeth on anterior or posterior or both margins of most articles (
<bibRefCitation id="1D5337D4FFFAFFA2FCE4FE9DFC32FEDD" author="Serene" box="[793,1023,332,357]" pageId="89" pageNumber="90" refString="Serene, R. (1964 b) Redescription du genre Megaesthesius Rathbun et definition des Chasmocarcininae, nouvelle sous-famille des Goneplacidae (Decapoda Brachyura). Crustaceana, 7 (3), 175 - 187." type="journal article" year="1964" yearSuffix="b">Serène 1964b: fig. 1</bibRefCitation>
). Fused thoracic sternites 1, 2 (
<figureCitation id="E1F956A0FFFAFFA2FA99FE9DFF78FE31" captionStart="FIGURE 55" captionStartId="128.[151,250,1514,1536]" captionTargetBox="[151,1436,179,1493]" captionTargetId="figure@128.[151,1436,179,1493]" captionTargetPageId="128" captionText="FIGURE 55. Anterior thoracic sternum and male pleon of species of Statommatia n. gen., Microtopsis and Megaesthesius. A, Statommatia granulosa n. sp., New Caledonia: holotype male 4.1 × 4.7 mm (MNHN-IU- 2013 - 9454); B, Microtopsis takedai, Japan: paratype male, carapace damaged (ZRC 2012.0146); C, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); D, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); E, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472); F, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073)." httpUri="https://zenodo.org/record/272701/files/figure.png" pageId="89" pageNumber="90">Figs. 55</figureCitation>
C; 88F) broadly triangular, proportionally wide, short; fused sternites 3, 4 relatively broad. Male pleon (
<figureCitation id="E1F956A0FFFAFFA2FA99FEBEFF78FE14" captionStart="FIGURE 55" captionStartId="128.[151,250,1514,1536]" captionTargetBox="[151,1436,179,1493]" captionTargetId="figure@128.[151,1436,179,1493]" captionTargetPageId="128" captionText="FIGURE 55. Anterior thoracic sternum and male pleon of species of Statommatia n. gen., Microtopsis and Megaesthesius. A, Statommatia granulosa n. sp., New Caledonia: holotype male 4.1 × 4.7 mm (MNHN-IU- 2013 - 9454); B, Microtopsis takedai, Japan: paratype male, carapace damaged (ZRC 2012.0146); C, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); D, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); E, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472); F, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073)." httpUri="https://zenodo.org/record/272701/files/figure.png" pageId="89" pageNumber="90">Figs. 55</figureCitation>
C; 62B) with lateral margins of somite 6, fused somites 35 slightly convex; postero-lateral regions slightly swollen; telson proportionally long. Sterno-pleonal cavity of male deep, press-button for pleonal holding as tubercle at middle portion of thoracic somite 5 near edge of sterno-pleonal cavity. Male thoracic sternite 8 (
<figureCitation id="E1F956A0FFFAFFA2FA99FE0DFF7CFDA1" captionStart="FIGURE 19" captionStartId="92.[151,250,1665,1687]" captionTargetBox="[203,1371,195,1618]" captionTargetId="figure@92.[203,1371,195,1632]" captionTargetPageId="92" captionText="FIGURE 19. Species of Megaesthesius and Alainthesius n. gen. A, B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); C F, Megaesthesius yokoyai (after Takeda &amp; Miyake 1969 a: figs. 3 a, 4 f, 4 a, 4 e, respectively); G L, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472) (after Crosnier 1975: fig. 1 a, 1 k, 1 j, 1 b, 1 c, 1 i, respectively); M R, Megaesthesius westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: figs. 1 A, B, 2 A, D, E, 3 E); S, W, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); T, X, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); U, Y, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); V, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012). A, overall habitus; B, C, G, M, dorsal view of carapace; E, left P 4; F, I, frontal view of cephalothorax; H, O, frontal and anterolateral margin of carapace; N, ventral view of cephalothorax; J, right P 3; K, buccal cavity and posterior margin of epistome; Q, right P 4; R, right P 5; D, L, P, male pleon; S V, posterior margin of epistome; W Y, left thoracic sternites 7 and 8, “ supplementary plate ”, penis tube and penis. Abbreviations: cx 4 = coxa of P 4; cx 5 = coxa of P 5; p = penis; pt = calcified penis tube; sp = “ supplementary plate ”; st 7 = thoracic sternite 7; st 8 = thoracic sternite 8." httpUri="https://zenodo.org/record/272665/files/figure.png" pageId="89" pageNumber="90">Figs. 19</figureCitation>
W, 62B) proportionally long, rectangular; “supplementary plate” conspicuously narrow, only reaching median portion of exposed thorax, short, slightly longer at rounded outer margin. Outer (ventral) surface of penis calcified, resembling plate between “supplementary plate”, sternite 8 (
<figureCitation id="E1F956A0FFFAFFA2FCB3FD96FC6CFDD9" box="[846,929,583,608]" captionStart="FIGURE 19" captionStartId="92.[151,250,1665,1687]" captionTargetBox="[203,1371,195,1618]" captionTargetId="figure@92.[203,1371,195,1632]" captionTargetPageId="92" captionText="FIGURE 19. Species of Megaesthesius and Alainthesius n. gen. A, B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); C F, Megaesthesius yokoyai (after Takeda &amp; Miyake 1969 a: figs. 3 a, 4 f, 4 a, 4 e, respectively); G L, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472) (after Crosnier 1975: fig. 1 a, 1 k, 1 j, 1 b, 1 c, 1 i, respectively); M R, Megaesthesius westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: figs. 1 A, B, 2 A, D, E, 3 E); S, W, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); T, X, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); U, Y, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); V, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012). A, overall habitus; B, C, G, M, dorsal view of carapace; E, left P 4; F, I, frontal view of cephalothorax; H, O, frontal and anterolateral margin of carapace; N, ventral view of cephalothorax; J, right P 3; K, buccal cavity and posterior margin of epistome; Q, right P 4; R, right P 5; D, L, P, male pleon; S V, posterior margin of epistome; W Y, left thoracic sternites 7 and 8, “ supplementary plate ”, penis tube and penis. Abbreviations: cx 4 = coxa of P 4; cx 5 = coxa of P 5; p = penis; pt = calcified penis tube; sp = “ supplementary plate ”; st 7 = thoracic sternite 7; st 8 = thoracic sternite 8." httpUri="https://zenodo.org/record/272665/files/figure.png" pageId="89" pageNumber="90">Fig. 19</figureCitation>
W). G1 (
<figureCitation id="E1F956A0FFFAFFA2FBF7FD96FB93FDD9" box="[1034,1118,583,608]" captionStart="FIGURE 82" captionStartId="155.[151,250,1823,1845]" captionTargetBox="[177,1417,213,1802]" captionTargetId="figure@155.[151,1436,193,1802]" captionTargetPageId="155" captionText="FIGURE 82. Gonopods of species of Megaesthesius. A C, M. sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); D F, M. migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); G I, M. yokoyai, Japan: male 3.5 × 3.8 mm (after Takeda &amp; Miyake 1969: fig. 4 h, i); J M, M. westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: fig. 3 A D). A, D, G, H, J, K, ventral view of left G 1; B, E, I, K, L, dorsal view of left G 1; C, F, M, left G 2. Scales = 0.1 mm." httpUri="https://zenodo.org/record/272728/files/figure.png" pageId="89" pageNumber="90">Fig. 82</figureCitation>
A, B;
<bibRefCitation id="1D5337D4FFFAFFA2FB5BFD96FA9AFDD9" author="Zarenkov" box="[1190,1367,583,608]" pageId="89" pageNumber="90" refString="Zarenkov, N. A. (1972) [New data on Indo-Pacific crabs (Fam. Goneplacidae, Pinnotheridae, Parthenopidae, Dorippidae) and the problem of seasonal reproduction of decapod animals in the Bay of Tonkin]. In: [The Complex Investigations of Ocean Nature], part 3, Moscow State University, Moscow, pp. 229 - 253. [In Russian]" type="book chapter" year="1972">Zarenkov 1972</bibRefCitation>
: 6-
<date id="0D7C6CE5FFFAFFA2FA80FD96FA5EFDD9" box="[1405,1427,583,608]" pageId="89" pageNumber="90">1I</date>
) slender, distal segment straight with spinules, apex slanted. G2 (
<figureCitation id="E1F956A0FFFAFFA2FC94FDBDFC71FD3D" box="[873,956,620,645]" captionStart="FIGURE 82" captionStartId="155.[151,250,1823,1845]" captionTargetBox="[177,1417,213,1802]" captionTargetId="figure@155.[151,1436,193,1802]" captionTargetPageId="155" captionText="FIGURE 82. Gonopods of species of Megaesthesius. A C, M. sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); D F, M. migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); G I, M. yokoyai, Japan: male 3.5 × 3.8 mm (after Takeda &amp; Miyake 1969: fig. 4 h, i); J M, M. westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: fig. 3 A D). A, D, G, H, J, K, ventral view of left G 1; B, E, I, K, L, dorsal view of left G 1; C, F, M, left G 2. Scales = 0.1 mm." httpUri="https://zenodo.org/record/272728/files/figure.png" pageId="89" pageNumber="90">Fig. 82</figureCitation>
C;
<bibRefCitation id="1D5337D4FFFAFFA2FC21FDBDFB46FD3D" author="Zarenkov" box="[988,1163,620,645]" pageId="89" pageNumber="90" refString="Zarenkov, N. A. (1972) [New data on Indo-Pacific crabs (Fam. Goneplacidae, Pinnotheridae, Parthenopidae, Dorippidae) and the problem of seasonal reproduction of decapod animals in the Bay of Tonkin]. In: [The Complex Investigations of Ocean Nature], part 3, Moscow State University, Moscow, pp. 229 - 253. [In Russian]" type="book chapter" year="1972">Zarenkov 1972</bibRefCitation>
: 6-
<date id="0D7C6CE5FFFAFFA2FB53FDBDFB03FD3C" box="[1198,1230,620,645]" pageId="89" pageNumber="90">1II</date>
) nearly as long as G1, straight, slender, distal segment long, as long as basal segment, directed inward. Somites of female pleon with gently convex margins; telson proportionally short. Sterno-pleonal cavity of female (
<figureCitation id="E1F956A0FFFAFFA2FB94FD65FB72FD75" box="[1129,1215,692,717]" captionStart="FIGURE 92" captionStartId="165.[151,250,934,956]" captionTargetBox="[151,1436,193,912]" captionTargetId="figure@165.[151,1436,193,912]" captionTargetPageId="165" captionText="FIGURE 92. Vulvae of species of Microtopsis and Megaesthesius. A, Microtopsis takedai, Japan: paratype female 3.6 × 2.7 mm (ZRC 2012.0146); B, Megaesthesius sagedae, Singapore: female carapace length 2.9 mm, sides damaged (ZRC 1995.883); C, Megaesthesius migmus n. sp., Philippines: paratype female 3.9 × 4.5 mm (ZRC 2015.245); D, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012)." httpUri="https://zenodo.org/record/272738/files/figure.png" pageId="89" pageNumber="90">Fig. 92</figureCitation>
B) deep, vulvae far apart from each other on outer margins of cavity close to suture 5/6.
</paragraph>
</subSubSection>
<subSubSection id="31D819AEFFFAFFA2FF3AFD2DFD69FA24" pageId="89" pageNumber="90" type="discussion">
<paragraph id="797D4A25FFFAFFA2FF3AFD2DFB85FC1C" blockId="89.[151,1437,151,1508]" pageId="89" pageNumber="90">
<emphasis id="4BB69637FFFAFFA2FF3AFD2DFEF9FCAC" bold="true" box="[199,308,764,789]" pageId="89" pageNumber="90">Remarks</emphasis>
. The figure given in
<bibRefCitation id="1D5337D4FFFAFFA2FDDCFD2DFC24FCAC" author="Rathbun" box="[545,1001,764,789]" pageId="89" pageNumber="90" refString="Rathbun, M. J. (1910) The Danish Expedition to Siam 1899 - 1900. V. Brachyura. Konelige Danske Videnskabernes Selskat, Naturvidenskabelige Matematiske Afhandlinger, 5 (4), 301 - 367, pls. 1, 2, map." type="journal article" year="1910">Rathbun (1910: figs. 30, 31, pl. 2, fig. 5)</bibRefCitation>
of this species is relatively simple and the photograph of her only specimen, a male, is too small to discern any details.
<bibRefCitation id="1D5337D4FFFAFFA2FB92FCF1FA51FC81" author="Serene" box="[1135,1436,799,824]" pageId="89" pageNumber="90" refString="Serene, R. (1964 b) Redescription du genre Megaesthesius Rathbun et definition des Chasmocarcininae, nouvelle sous-famille des Goneplacidae (Decapoda Brachyura). Crustaceana, 7 (3), 175 - 187." type="journal article" year="1964" yearSuffix="b">Serène (1964b: figs. 16)</bibRefCitation>
examined the specimen and the carapace, chelipeds, and mouthparts in his figures agree with those of
<taxonomicName id="BEC231A6FFFAFFA2FF6AFCB6FE52FC39" box="[151,415,871,896]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="sagedae">
<emphasis id="4BB69637FFFAFFA2FF6AFCB6FE52FC39" box="[151,415,871,896]" italics="true" pageId="89" pageNumber="90">Megaesthesius sagedae</emphasis>
</taxonomicName>
examined here from the type locality in Singapore. Unfortunately, the present condition of the type specimen in ZMUC is very poor, and the pleon, G1, and G2 are all missing.
</paragraph>
<paragraph id="797D4A25FFFAFFA2FF3AFC61FCC8FAB4" blockId="89.[151,1437,151,1508]" pageId="89" pageNumber="90">
We are confident that Zarenkovs (1972: 241, fig. 6-1) record of this species from Vietnam is
<taxonomicName id="BEC231A6FFFAFFA2FB19FC60FAABFC71" box="[1252,1382,943,968]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="sagedae">
<emphasis id="4BB69637FFFAFFA2FB19FC60FAABFC71" box="[1252,1382,943,968]" italics="true" pageId="89" pageNumber="90">M. sagedae</emphasis>
</taxonomicName>
as it agrees very well with the specimens from Singapore.
<bibRefCitation id="1D5337D4FFFAFFA2FD09FC05FBF5FC55" author="Serene" box="[756,1080,980,1005]" pageId="89" pageNumber="90" refString="Serene, R. &amp; Vadon, C. (1981) Crustaces decapodes: brachyoures. Liste preliminaire, description de formes nouvelles et remarques taxonomiques, In: Resultats des campagnes MUSORSTOM I. Philippines (18 - 28 mars 1976), vol. 1. Memoires ORSTOM (Paris), 91, 117 - 140." type="book chapter" year="1981">Serène &amp; Vadon (1981: 123)</bibRefCitation>
reported on a female, 4.5 × 6.0 mm, collected by MUSORSTOM 2 from station CP15 from the entrance of Luzon and Lubang in the western Philippines (13°55N, 120°29'E,
<quantity id="BE3AE7C0FFFAFFA2FDF1FBCDFD43FB8D" box="[524,654,1052,1077]" metricMagnitude="2" metricUnit="m" metricValue="3.28" metricValueMax="3.3" metricValueMin="3.26" pageId="89" pageNumber="90" unit="m" value="328.0" valueMax="330.0" valueMin="326.0">326330 m</quantity>
,
<date id="0D7C6CE5FFFAFFA2FD67FBCDFCBBFB8D" box="[666,886,1052,1077]" pageId="89" pageNumber="90" value="1980-11-21">21 November 1980</date>
) but we have not been able to find the specimen in MNHN. Like many of the identified specimens from this expedition reported by these authors, the specimens could not be found (e.g., see Ng &amp;
<bibRefCitation id="1D5337D4FFFAFFA2FDCBFBB5FD7BFBC4" author="Chen" box="[566,694,1124,1149]" pageId="89" pageNumber="90" refString="Ng, P. K. L. &amp; Chen, H. - L. (2005) On two species of euxanthine crabs from the South China Sea, including a description of a new species of Crosnierius (Crustacea: Decapoda: Brachyura: Xanthidae). Proceedings of the Biological Society of Washington, 118 (2), 319 - 325." type="journal article" year="2005">Chen 2005</bibRefCitation>
; Richer de
<bibRefCitation id="1D5337D4FFFAFFA2FCC0FBB4FBD6FBC5" author="Forges" box="[829,1051,1124,1149]" pageId="89" pageNumber="90" refString="Richer De Forges, B. &amp; Ng, P. K. L. (2012) Griffinia takedai, a new species of deep sea majoid crab (Decapoda, Brachyura, Epialtidae) from the Philippines. In: Komatsu, H., Okuno, J. &amp; Fukuoka, K. (Eds.), Studies on Eumalacostraca: a homage to Takeda Masatsune. Crustaceana Monographs, 17, 275 - 284." type="journal article" year="2012">Forges &amp; Ng 2012</bibRefCitation>
). Its identity must be regarded as
<emphasis id="4BB69637FFFAFFA2FF6AFB58FEE9FB19" box="[151,292,1159,1184]" italics="true" pageId="89" pageNumber="90">incerta sedis</emphasis>
until it can be found.
<taxonomicName id="BEC231A6FFFAFFA2FDEAFB56FCD4FB19" box="[535,793,1159,1184]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="migmus" status="sp. nov.">
<emphasis id="4BB69637FFFAFFA2FDEAFB56FCD4FB19" box="[535,793,1159,1184]" italics="true" pageId="89" pageNumber="90">Megaesthesius migmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50852B4CFFFAFFA2FCDDFB56FC93FB19" box="[800,862,1159,1184]" pageId="89" pageNumber="90" rank="species">
<emphasis id="4BB69637FFFAFFA2FCDDFB56FC93FB19" bold="true" box="[800,862,1159,1184]" pageId="89" pageNumber="90">n. sp.</emphasis>
</taxonomicNameLabel>
has been recorded from deep waters in Philippines, and it is possible their record is this new species instead.
<taxonomicName id="BEC231A6FFFAFFA2FCEAFB7DFBD7FB7D" box="[791,1050,1196,1221]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="migmus" status="sp. nov.">
<emphasis id="4BB69637FFFAFFA2FCEAFB7DFBD7FB7D" box="[791,1050,1196,1221]" italics="true" pageId="89" pageNumber="90">Megaesthesius migmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50852B4CFFFAFFA2FBDCFB7DFBADFB7C" box="[1057,1120,1196,1221]" pageId="89" pageNumber="90" rank="species">
<emphasis id="4BB69637FFFAFFA2FBDCFB7DFBADFB7C" bold="true" box="[1057,1120,1196,1221]" pageId="89" pageNumber="90">n. sp.</emphasis>
</taxonomicNameLabel>
was found in the Bohol and Luzon Seas to the east. The record of “
<taxonomicName id="BEC231A6FFFAFFA2FDB5FB1EFC9DFB51" box="[584,848,1231,1256]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="sagedae">
<emphasis id="4BB69637FFFAFFA2FDB5FB1EFC9DFB51" box="[584,848,1231,1256]" italics="true" pageId="89" pageNumber="90">Megaesthesius sagedae</emphasis>
</taxonomicName>
” by Ng &amp; Davie (2002: 378, Western Thailand) will also need to re-examined to ascertain its actual identity.
</paragraph>
<paragraph id="797D4A25FFFAFFA2FF3AFAC6FEF2FAED" blockId="89.[151,1437,151,1508]" pageId="89" pageNumber="90">
The present specimens from Singapore were all collected from soft substrates in relatively shallow water, all less than
<quantity id="BE3AE7C0FFFAFFA2FF02FAEDFEF6FAED" box="[255,315,1340,1364]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="89" pageNumber="90" unit="m" value="20.0">20 m</quantity>
.
</paragraph>
<paragraph id="797D4A25FFFAFFA2FF3AFA8EFD69FA24" blockId="89.[151,1437,151,1508]" pageId="89" pageNumber="90">
The major differences between the four taxa are discussed under the
<emphasis id="4BB69637FFFAFFA2FC21FA8EFB84FAC1" bold="true" box="[988,1097,1375,1400]" pageId="89" pageNumber="90">Remarks</emphasis>
for
<taxonomicName id="BEC231A6FFFAFFA2FB81FAB0FB37FAC1" box="[1148,1274,1377,1400]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="migmus" status="sp. nov.">
<emphasis id="4BB69637FFFAFFA2FB81FAB0FB37FAC1" box="[1148,1274,1377,1400]" italics="true" pageId="89" pageNumber="90">M. migmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50852B4CFFFAFFA2FAFEFA8EFA8EFAC1" box="[1283,1347,1375,1400]" pageId="89" pageNumber="90" rank="species">
<emphasis id="4BB69637FFFAFFA2FAFEFA8EFA8EFAC1" bold="true" box="[1283,1347,1375,1400]" pageId="89" pageNumber="90">n. sp.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="BEC231A6FFFAFFA2FA80FAB0FF23FA24" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="yokoyai">
<emphasis id="4BB69637FFFAFFA2FA80FAB0FF23FA24" italics="true" pageId="89" pageNumber="90">M. yokoyai</emphasis>
</taxonomicName>
(see below and the key to the species).
</paragraph>
</subSubSection>
<subSubSection id="31D819AEFFFAFFA2FF3AFA76FF1AFA5D" pageId="89" pageNumber="90" type="distribution">
<paragraph id="797D4A25FFFAFFA2FF3AFA76FF1AFA5D" blockId="89.[151,1437,151,1508]" pageId="89" pageNumber="90">
<emphasis id="4BB69637FFFAFFA2FF3AFA76FE94FA79" bold="true" box="[199,345,1447,1472]" pageId="89" pageNumber="90">Distribution</emphasis>
.
<collectingRegion id="BB0684C7FFFAFFA2FE9AFA79FE0FFA06" box="[359,450,1448,1471]" country="Solomon Islands" name="Western" pageId="89" pageNumber="90">Western</collectingRegion>
Pacific Ocean (
<collectingCountry id="01D50AB5FFFAFFA2FD88FA76FD1BFA79" box="[629,726,1447,1472]" name="Vietnam" pageId="89" pageNumber="90">Vietnam</collectingCountry>
and
<collectingCountry id="01D50AB5FFFAFFA2FCF3FA76FC4EFA79" box="[782,899,1447,1472]" name="Singapore" pageId="89" pageNumber="90">Singapore</collectingCountry>
; questionably from Andaman Sea). Depth:
<quantity id="BE3AE7C0FFFAFFA2FA96FA76FF1EFA5D" metricMagnitude="1" metricUnit="m" metricValue="1.13" metricValueMax="1.9" metricValueMin="0.36" pageId="89" pageNumber="90" unit="m" value="11.3" valueMax="19.0" valueMin="3.6">
3.6
<specimenCount id="6FC481ACFFFAFFA2FF6AFA1DFF1EFA5D" box="[151,211,1484,1508]" pageId="89" pageNumber="90" type="generic">19 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>