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<document id="99F0911120AD5BA580300A5A2097C63B" ID-CLB-Dataset="37321" ID-DOI="10.11646/zootaxa.4209.1.1" ID-GBIF-Dataset="2de5b7bf-3975-458a-b200-07ec69f4e013" ID-ISSN="1175-5326" ID-Zenodo-Dep="272646" ID-ZooBank="849BAB5C-464A-4B4A-A586-5742411EDC01" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484407904318" checkinUser="plazi" docAuthor="Ng, Peter K. L. &amp; Castro, Peter" docDate="2016" docId="F16BFB33FFF7FFAEFF6AFB77FCD5F9C0" docLanguage="en" docName="zootaxa.4209.1.1.pdf.imf" docOrigin="Zootaxa 4209 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Microtopsis takedai Komai, Ng &amp; Yamada 2012" docType="treatment" docVersion="8" lastPageNumber="86" masterDocId="0D52834BFFA3FFFBFFFDFFD1FFCDFFB9" masterDocTitle="Revision of the family Chasmocarcinidae Serène, 1964 (Crustacea, Brachyura, Goneplacoidea)" masterLastPageNumber="182" masterPageNumber="1" pageNumber="85" updateTime="1698435566492" updateUser="ExternalLinkService">
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<mods:title id="0E8FCE25C00B2C3D01172A47D8B636A5">Revision of the family Chasmocarcinidae Serène, 1964 (Crustacea, Brachyura, Goneplacoidea)</mods:title>
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<mods:namePart id="089A4AF572F080AAB4F4515BFC96273C">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="7D1544D37185CD9B5F014FAAFC255B27">Castro, Peter</mods:namePart>
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<mods:title id="675C1D7A8FB3DA952F79484DB979AAAB">Zootaxa</mods:title>
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<mods:date id="0B88742D4EAB0EF342EC328AD1483E80">2016</mods:date>
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<mods:number id="D8934D24CE62485C0BFD659DFEDCD494">4209</mods:number>
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<mods:identifier id="1CE75339A3404831B51F73D7F24761EC" type="DOI">10.11646/zootaxa.4209.1.1</mods:identifier>
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<treatment id="F16BFB33FFF7FFAEFF6AFB77FCD5F9C0" ID-DOI="http://doi.org/10.5281/zenodo.5617207" ID-GBIF-Taxon="127419543" ID-Zenodo-Dep="5617207" LSID="urn:lsid:plazi:treatment:F16BFB33FFF7FFAEFF6AFB77FCD5F9C0" httpUri="http://treatment.plazi.org/id/F16BFB33FFF7FFAEFF6AFB77FCD5F9C0" lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85">
<subSubSection id="31D819AEFFF7FFAFFF6AFB77FCA7FB5B" pageId="84" pageNumber="85" type="nomenclature">
<paragraph id="797D4A25FFF7FFAFFF6AFB77FD24FB79" blockId="84.[151,874,1190,1250]" box="[151,745,1190,1216]" pageId="84" pageNumber="85">
<heading id="2235FD49FFF7FFAFFF6AFB77FD24FB79" bold="true" box="[151,745,1190,1216]" fontSize="11" level="1" pageId="84" pageNumber="85" reason="1">
<taxonomicName id="BEC231A6FFF7FFAFFF6AFB77FD24FB79" ID-CoL="438VQ" authority="Komai, Ng &amp; Yamada, 2012" authorityName="Komai, Ng &amp; Yamada" authorityYear="2012" box="[151,745,1190,1216]" class="Malacostraca" family="Chasmocarcinidae" genus="Microtopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="takedai">
<emphasis id="4BB69637FFF7FFAFFF6AFB77FD24FB79" bold="true" box="[151,745,1190,1216]" pageId="84" pageNumber="85">
<emphasis id="4BB69637FFF7FFAFFF6AFB77FE4EFB79" bold="true" box="[151,387,1190,1216]" italics="true" pageId="84" pageNumber="85">Microtopsis takedai</emphasis>
Komai, Ng &amp; Yamada, 2012
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="797D4A25FFF7FFAFFF6AFB18FCA7FB5B" blockId="84.[151,874,1190,1250]" box="[151,874,1225,1250]" pageId="84" pageNumber="85">
(
<figureCitation id="E1F956A0FFF7FFAFFF62FB18FF37FB5B" box="[159,250,1225,1250]" captionStart="FIGURE 18" captionStartId="85.[151,250,1325,1347]" captionTargetBox="[151,1436,517,1304]" captionTargetId="figure@85.[151,1436,517,1304]" captionTargetPageId="85" captionText="FIGURE 18. Overall dorsal view of species of Microtopsis. A, M. takedai, Japan: paratype female 3.6 × 2.7 mm (ZRC 2012.0146); B, M. teschi n. sp., Indonesia: holotype male 1.5 × 2.1 mm (NNM-ZMA); C, M. teschi n. sp., Indonesia: paratype male 1.5 × 1.9 mm (NNM-ZMA)." httpUri="https://zenodo.org/record/272664/files/figure.png" pageId="84" pageNumber="85">Figs. 18</figureCitation>
A; 27A; 48A, B; 55B; 62A; 80A, B; 81A–F; 88E; 92A)
</paragraph>
</subSubSection>
<subSubSection id="31D819AEFFF7FFAFFF6AFAC3FB49FA9E" box="[151,1156,1297,1319]" pageId="84" pageNumber="85" type="reference_group">
<paragraph id="797D4A25FFF7FFAFFF6AFAC3FB49FA9E" blockId="84.[151,1156,1297,1319]" box="[151,1156,1297,1319]" pageId="84" pageNumber="85">
<taxonomicName id="BEC231A6FFF7FFAFFF6AFAC3FD70FA9E" ID-CoL="438VQ" authority="Komai, Ng &amp; Yamada, 2012: 149" authorityName="Komai, Ng &amp; Yamada" authorityPageNumber="149" authorityYear="2012" box="[151,701,1297,1319]" class="Malacostraca" family="Chasmocarcinidae" genus="Microtopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="takedai">
<emphasis id="4BB69637FFF7FFAFFF6AFAC3FEADFA9E" box="[151,352,1297,1319]" italics="true" pageId="84" pageNumber="85">Microtopsis takedai</emphasis>
Komai, Ng &amp; Yamada, 2012: 149
</taxonomicName>
, figs. 1‒5 [type localilty: Japan, Ryukyu Is.].
</paragraph>
</subSubSection>
<subSubSection id="31D819AEFFF7FFAFFF6AFA85FCFCFA61" pageId="84" pageNumber="85" type="materials_examined">
<paragraph id="797D4A25FFF7FFAFFF6AFA85FACAFAD5" blockId="84.[151,1437,1364,2037]" box="[151,1287,1364,1389]" pageId="84" pageNumber="85">
<materialsCitation id="C9AA4078FFF7FFAFFF6AFA85FACAFAD5" ID-GBIF-Occurrence="1424726216" box="[151,1287,1364,1389]" country="Japan" location="Ryukyu Is" pageId="84" pageNumber="85" specimenCount="2" stateProvince="Okinawa" typeStatus="holotype">
<emphasis id="4BB69637FFF7FFAFFF6AFA85FEF0FAD4" bold="true" box="[151,317,1364,1389]" pageId="84" pageNumber="85">
<typeStatus id="A679F487FFF7FFAFFF6AFA85FF1FFAD4" box="[151,210,1364,1389]" pageId="84" pageNumber="85">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="A679F487FFF7FFAFFEB1FA85FE79FAD4" box="[332,436,1364,1389]" pageId="84" pageNumber="85" type="holotype">Holotype</typeStatus>
male (2.4 ×
<quantity id="BE3AE7C0FFF7FFAFFDBFFA85FD54FAD5" box="[578,665,1364,1388]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="84" pageNumber="85" unit="mm" value="3.1">3.1 mm</quantity>
) (CBM-ZC 10491),
<collectingCountry id="01D50AB5FFF7FFAFFC7CFA84FC09FAD5" box="[897,964,1365,1388]" name="Japan" pageId="84" pageNumber="85">Japan</collectingCountry>
,
<location id="7C1D1CFEFFF7FFAFFC32FA85FB89FAD5" LSID="urn:lsid:plazi:treatment:F16BFB33FFF7FFAEFF6AFB77FCD5F9C0:7C1D1CFEFFF7FFAFFC32FA85FB89FAD5" box="[975,1092,1364,1389]" country="Japan" name="Ryukyu Is" pageId="84" pageNumber="85" stateProvince="Okinawa">Ryukyu Is</location>
,
<collectingRegion id="BB0684C7FFF7FFAFFBADFA85FB75FAD4" box="[1104,1208,1364,1389]" country="Japan" name="Okinawa" pageId="84" pageNumber="85">Okinawa</collectingRegion>
, Nago.
</materialsCitation>
</paragraph>
<paragraph id="797D4A25FFF7FFAFFF6AFAA9FAC7FA29" blockId="84.[151,1437,1364,2037]" box="[151,1290,1399,1424]" pageId="84" pageNumber="85">
Paratype: female (2.8 ×
<quantity id="BE3AE7C0FFF7FFAFFE58FAA9FE30FA29" box="[421,509,1400,1424]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="84" pageNumber="85" unit="mm" value="3.4">3.4 mm</quantity>
) (CBM-ZC 10492), male and female (ZRC 2012.0146), type locality.
</paragraph>
<paragraph id="797D4A25FFF7FFAFFF3AFA4DFCFCFA61" blockId="84.[151,1437,1364,2037]" pageId="84" pageNumber="85">
<materialsCitation id="C9AA4078FFF7FFAFFF3AFA4DFCECFA0C" ID-GBIF-Occurrence="1424726223" box="[199,801,1436,1461]" country="Japan" location="Japan" pageId="84" pageNumber="85" specimenCount="1" specimenCount-male="1">
<emphasis id="4BB69637FFF7FFAFFF3AFA4DFE39FA0C" bold="true" box="[199,500,1436,1461]" pageId="84" pageNumber="85">Other material examined</emphasis>
.
<collectingCountry id="01D50AB5FFF7FFAFFDFFFA4CFD8BFA0D" box="[514,582,1437,1460]" name="Japan" pageId="84" pageNumber="85">
<emphasis id="4BB69637FFF7FFAFFDFFFA4CFD8BFA0D" box="[514,582,1437,1460]" italics="true" pageId="84" pageNumber="85">Japan</emphasis>
</collectingCountry>
.
<specimenCount id="6FC481ACFFF7FFAFFDA9FA4DFD52FA0C" box="[596,671,1436,1461]" pageId="84" pageNumber="85" type="male">1 male</specimenCount>
(damaged)
</materialsCitation>
,
<materialsCitation id="C9AA4078FFF7FFAFFCD1FA4DFCFCFA61" ID-GBIF-Occurrence="1424726202" collectionCode="ZRC" collectorName="Y. Yamada" country="Japan" elevation="13" location="Nago" municipality="Awa" pageId="84" pageNumber="85" specimenCode="ZRC 2012.0146" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa">
<specimenCount id="6FC481ACFFF7FFAFFCD1FA4DFC40FA0C" box="[812,909,1436,1461]" pageId="84" pageNumber="85" type="female">1 female</specimenCount>
(3.6 ×
<quantity id="BE3AE7C0FFF7FFAFFC23FA4DFBF8FA0D" box="[990,1077,1436,1460]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="84" pageNumber="85" unit="mm" value="2.7">2.7 mm</quantity>
) (
<specimenCode id="2964E25EFFF7FFAFFBB6FA4DFACBFA0C" box="[1099,1286,1436,1461]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="84" pageNumber="85">
<collectionCode id="1FD3D2E0FFF7FFAFFBB6FA4DFB4EFA0D" box="[1099,1155,1436,1460]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="84" pageNumber="85">ZRC</collectionCode>
2012.0146
</specimenCode>
),
<collectingMunicipality id="9919D05FFFF7FFAFFAE5FA4DFA80FA0D" box="[1304,1357,1436,1460]" pageId="84" pageNumber="85">Awa</collectingMunicipality>
,
<location id="7C1D1CFEFFF7FFAFFAA4FA4CFA55FA0C" LSID="urn:lsid:plazi:treatment:F16BFB33FFF7FFAEFF6AFB77FCD5F9C0:7C1D1CFEFFF7FFAFFAA4FA4CFA55FA0C" box="[1369,1432,1437,1461]" country="Japan" municipality="Awa" name="Nago" pageId="84" pageNumber="85" stateProvince="Okinawa">Nago</location>
,
<collectingRegion id="BB0684C7FFF7FFAFFF6AFA6EFF32FA61" box="[151,255,1471,1496]" country="Japan" name="Okinawa" pageId="84" pageNumber="85">Okinawa</collectingRegion>
,
<elevation id="F2EFAD16FFF7FFAFFEF7FA11FE88FA6E" box="[266,325,1472,1496]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="84" pageNumber="85" unit="m" value="13.0">
<quantity id="BE3AE7C0FFF7FFAFFEF7FA11FE88FA6E" box="[266,325,1472,1496]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="84" pageNumber="85" unit="m" value="13.0">13 m</quantity>
</elevation>
, scuba diving,
<collectorName id="D4372FF3FFF7FFAFFE10FA11FDAAFA61" box="[493,615,1471,1496]" pageId="84" pageNumber="85">Y. Yamada</collectorName>
coll., 0 2.01.2011.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="31D819AEFFF7FFAEFF3AFA35FD14FE69" lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85" type="diagnosis">
<paragraph id="797D4A25FFF7FFAEFF3AFA35FD14FE69" blockId="84.[151,1437,1364,2037]" lastBlockId="85.[151,1437,151,464]" lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85">
<emphasis id="4BB69637FFF7FFAFFF3AFA35FEF7FA44" bold="true" box="[199,314,1508,1533]" pageId="84" pageNumber="85">Diagnosis</emphasis>
. Carapace (
<figureCitation id="E1F956A0FFF7FFAFFE3FFA35FDD9FA45" box="[450,532,1508,1533]" captionStart="FIGURE 18" captionStartId="85.[151,250,1325,1347]" captionTargetBox="[151,1436,517,1304]" captionTargetId="figure@85.[151,1436,517,1304]" captionTargetPageId="85" captionText="FIGURE 18. Overall dorsal view of species of Microtopsis. A, M. takedai, Japan: paratype female 3.6 × 2.7 mm (ZRC 2012.0146); B, M. teschi n. sp., Indonesia: holotype male 1.5 × 2.1 mm (NNM-ZMA); C, M. teschi n. sp., Indonesia: paratype male 1.5 × 1.9 mm (NNM-ZMA)." httpUri="https://zenodo.org/record/272664/files/figure.png" pageId="84" pageNumber="85">Fig. 18</figureCitation>
A;
<bibRefCitation id="1D5337D4FFF7FFAFFDC5FA35FCC7FA45" author="Komai" box="[568,778,1508,1533]" pageId="84" pageNumber="85" refString="Komai, T., Ng, P. K. L. &amp; Yamada, Y. (2012) A new genus and new species of chasmocarcinid crab (Decapoda, Brachyura, Goneplacoidea) from shallow waters in Japan. Studies on Eumalacostraca: a Homage to Masatsune Takeda. Crustaceana Monographs, 17, 137 - 154." type="journal article" year="2012">
Komai
<emphasis id="4BB69637FFF7FFAFFD71FA34FD0BFA44" box="[652,710,1508,1533]" italics="true" pageId="84" pageNumber="85">et al.</emphasis>
2012
</bibRefCitation>
: figs. 1; 2A; 2B) subtrapezoidal, 1.2‒1.3 wider than long; front nearly straight; anterolateral margins arcuate, minutely dentate, without distinct lobes or teeth; dorsal surface almost smooth except for minute granules on lateral surfaces. Epistome (
<figureCitation id="E1F956A0FFF7FFAFFC19F9FDFBF5F9FD" box="[996,1080,1580,1605]" captionStart="FIGURE 27" captionStartId="100.[151,250,1447,1469]" captionTargetBox="[151,1436,193,1426]" captionTargetId="figure@100.[151,1436,193,1426]" captionTargetPageId="100" captionText="FIGURE 27. Frontal view of species of Megaesthesius and Microtopsis. A, Microtopsis takedai, Japan: paratype female 3.6 × 2.7 mm (ZRC 2012.0146); B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); C, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); D, Alainthesius signatus n. sp., Madagascar: paratype female 2.7 × 3.0 mm (MNHN-IU- 2013 - 9471); E, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); F, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012)." httpUri="https://zenodo.org/record/272673/files/figure.png" pageId="84" pageNumber="85">Fig. 27</figureCitation>
A;
<bibRefCitation id="1D5337D4FFF7FFAFFBA0F9FDFAF9F9FD" author="Komai" box="[1117,1332,1580,1605]" pageId="84" pageNumber="85" refString="Komai, T., Ng, P. K. L. &amp; Yamada, Y. (2012) A new genus and new species of chasmocarcinid crab (Decapoda, Brachyura, Goneplacoidea) from shallow waters in Japan. Studies on Eumalacostraca: a Homage to Masatsune Takeda. Crustaceana Monographs, 17, 137 - 154." type="journal article" year="2012">
Komai
<emphasis id="4BB69637FFF7FFAFFB4EF9FCFB23F9FC" box="[1203,1262,1580,1605]" italics="true" pageId="84" pageNumber="85">et al.</emphasis>
2012
</bibRefCitation>
: fig. 2C) slightly depressed; posterior margin with widely semicircular median lobe with median fissure, semicircular lateral margins without fissures. Eye peduncle (
<figureCitation id="E1F956A0FFF7FFAFFD9AF9A5FD76F935" box="[615,699,1652,1677]" captionStart="FIGURE 27" captionStartId="100.[151,250,1447,1469]" captionTargetBox="[151,1436,193,1426]" captionTargetId="figure@100.[151,1436,193,1426]" captionTargetPageId="100" captionText="FIGURE 27. Frontal view of species of Megaesthesius and Microtopsis. A, Microtopsis takedai, Japan: paratype female 3.6 × 2.7 mm (ZRC 2012.0146); B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); C, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); D, Alainthesius signatus n. sp., Madagascar: paratype female 2.7 × 3.0 mm (MNHN-IU- 2013 - 9471); E, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); F, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012)." httpUri="https://zenodo.org/record/272673/files/figure.png" pageId="84" pageNumber="85">Fig. 27</figureCitation>
A;
<bibRefCitation id="1D5337D4FFF7FFAFFD23F9A5FC7CF935" author="Komai" box="[734,945,1652,1677]" pageId="84" pageNumber="85" refString="Komai, T., Ng, P. K. L. &amp; Yamada, Y. (2012) A new genus and new species of chasmocarcinid crab (Decapoda, Brachyura, Goneplacoidea) from shallow waters in Japan. Studies on Eumalacostraca: a Homage to Masatsune Takeda. Crustaceana Monographs, 17, 137 - 154." type="journal article" year="2012">
Komai
<emphasis id="4BB69637FFF7FFAFFCCEF9A4FCA0F934" box="[819,877,1652,1677]" italics="true" pageId="84" pageNumber="85">et al.</emphasis>
2012
</bibRefCitation>
: fig. 2B) filling orbit, long, mobile; cornea reduced, pigmented. Third maxillipeds (
<figureCitation id="E1F956A0FFF7FFAFFD94F946FD72F909" box="[617,703,1687,1712]" captionStart="FIGURE 80" captionStartId="153.[151,250,1480,1502]" captionTargetBox="[166,1396,204,1436]" captionTargetId="figure@153.[151,1436,193,1459]" captionTargetPageId="153" captionText="FIGURE 80. Third maxilliped and male pleon of species of Microtopsis. A, B, M. takedai, Japan: holotype male 2.4 × 3.1 mm (after Komai et al. 2012: figs. 3 A, 2 H, respectively); C, D, F, M. teschi n. sp., Indonesia: holotype male 1.5 × 2.1 mm (NNM- ZMA); E, M. teschi n. sp., Indonesia: paratype 1.5 × 1.9 mm (NNM-ZMA). A, left third maxilliped; B, F, male pleon; C, right third maxilliped (denuded, exopod not drawn); D, left third maxilliped (denuded, exopod not drawn); E, right third maxilliped (denuded). Scales = 0.5 mm." httpUri="https://zenodo.org/record/272726/files/figure.png" pageId="84" pageNumber="85">Fig. 80</figureCitation>
A;
<bibRefCitation id="1D5337D4FFF7FFAFFD18F946FC0DF909" author="Komai" box="[741,960,1687,1712]" pageId="84" pageNumber="85" refString="Komai, T., Ng, P. K. L. &amp; Yamada, Y. (2012) A new genus and new species of chasmocarcinid crab (Decapoda, Brachyura, Goneplacoidea) from shallow waters in Japan. Studies on Eumalacostraca: a Homage to Masatsune Takeda. Crustaceana Monographs, 17, 137 - 154." type="journal article" year="2012">
Komai
<emphasis id="4BB69637FFF7FFAFFCC1F948FCB4F909" box="[828,889,1687,1712]" italics="true" pageId="84" pageNumber="85">et al.</emphasis>
2012
</bibRefCitation>
: fig. 3A) not filling buccal cavern when closed; merus subcircular, anteroexternal angle rounded, narrower than ischium; ischium quadrate, widened inner margin, about same length as merus (
<figureCitation id="E1F956A0FFF7FFAFFDC8F90EFD4BF941" box="[565,646,1759,1784]" captionStart="FIGURE 80" captionStartId="153.[151,250,1480,1502]" captionTargetBox="[166,1396,204,1436]" captionTargetId="figure@153.[151,1436,193,1459]" captionTargetPageId="153" captionText="FIGURE 80. Third maxilliped and male pleon of species of Microtopsis. A, B, M. takedai, Japan: holotype male 2.4 × 3.1 mm (after Komai et al. 2012: figs. 3 A, 2 H, respectively); C, D, F, M. teschi n. sp., Indonesia: holotype male 1.5 × 2.1 mm (NNM- ZMA); E, M. teschi n. sp., Indonesia: paratype 1.5 × 1.9 mm (NNM-ZMA). A, left third maxilliped; B, F, male pleon; C, right third maxilliped (denuded, exopod not drawn); D, left third maxilliped (denuded, exopod not drawn); E, right third maxilliped (denuded). Scales = 0.5 mm." httpUri="https://zenodo.org/record/272726/files/figure.png" pageId="84" pageNumber="85">Fig. 80</figureCitation>
A;
<bibRefCitation id="1D5337D4FFF7FFAFFD55F90EFCB9F941" author="Komai" box="[680,884,1759,1784]" pageId="84" pageNumber="85" refString="Komai, T., Ng, P. K. L. &amp; Yamada, Y. (2012) A new genus and new species of chasmocarcinid crab (Decapoda, Brachyura, Goneplacoidea) from shallow waters in Japan. Studies on Eumalacostraca: a Homage to Masatsune Takeda. Crustaceana Monographs, 17, 137 - 154." type="journal article" year="2012">
Komai
<emphasis id="4BB69637FFF7FFAFFD07F930FCFCF941" box="[762,817,1759,1784]" italics="true" pageId="84" pageNumber="85">et al.</emphasis>
2012
</bibRefCitation>
: 3A). Chelipeds (
<figureCitation id="E1F956A0FFF7FFAFFBCBF90EFB5CF941" box="[1078,1169,1759,1784]" captionStart="FIGURE 18" captionStartId="85.[151,250,1325,1347]" captionTargetBox="[151,1436,517,1304]" captionTargetId="figure@85.[151,1436,517,1304]" captionTargetPageId="85" captionText="FIGURE 18. Overall dorsal view of species of Microtopsis. A, M. takedai, Japan: paratype female 3.6 × 2.7 mm (ZRC 2012.0146); B, M. teschi n. sp., Indonesia: holotype male 1.5 × 2.1 mm (NNM-ZMA); C, M. teschi n. sp., Indonesia: paratype male 1.5 × 1.9 mm (NNM-ZMA)." httpUri="https://zenodo.org/record/272664/files/figure.png" pageId="84" pageNumber="85">Figs. 18</figureCitation>
A; 48A, B;
<bibRefCitation id="1D5337D4FFF7FFAFFAEFF90EFF19F8A5" author="Komai" pageId="84" pageNumber="85" refString="Komai, T., Ng, P. K. L. &amp; Yamada, Y. (2012) A new genus and new species of chasmocarcinid crab (Decapoda, Brachyura, Goneplacoidea) from shallow waters in Japan. Studies on Eumalacostraca: a Homage to Masatsune Takeda. Crustaceana Monographs, 17, 137 - 154." type="journal article" year="2012">
Komai
<emphasis id="4BB69637FFF7FFAFFA99F930FA51F941" box="[1380,1436,1759,1784]" italics="true" pageId="84" pageNumber="85">et al.</emphasis>
2012
</bibRefCitation>
: figs. 3B, D; 5A, B) subequal in length, slightly dissimilar in female, heteromorphic in males; cutting margins of both chelipeds of females,
<taxonomicName id="BEC231A6FFF7FFAFFE19F8F6FDEAF8F9" box="[484,551,1831,1856]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="minor">minor</taxonomicName>
in males with dense row of short setae (
<figureCitation id="E1F956A0FFF7FFAFFC10F8F6FB8CF8F9" box="[1005,1089,1831,1856]" captionStart="FIGURE 48" captionStartId="121.[151,250,1288,1310]" captionTargetBox="[151,1436,217,1267]" captionTargetId="figure@121.[151,1436,193,1267]" captionTargetPageId="121" captionText="FIGURE 48. Chelae of species of Megaesthesius, Alainthesius n. gen. and Microtopsis. A, B, Microtopsis takedai, Japan: holotype male 2.4 × 3.1 mm (after Komai et al. 2012: fig. 3 B, D); C, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); D, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); E, Megaesthesius yokoyai, Japan: male (after Takeda &amp; Miyake 1969: fig. 4 d); F, Alainthesius signatus n. sp., Madagascar: holotype male 3.5 × 3.8 mm (after Crosnier 1975: fig. 1 d); G, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN- IU- 2013 - 9073); H, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012)." httpUri="https://zenodo.org/record/272694/files/figure.png" pageId="84" pageNumber="85">Fig. 48</figureCitation>
B;
<bibRefCitation id="1D5337D4FFF7FFAFFB9DF8F6FAE2F8F9" author="Komai" box="[1120,1327,1831,1856]" pageId="84" pageNumber="85" refString="Komai, T., Ng, P. K. L. &amp; Yamada, Y. (2012) A new genus and new species of chasmocarcinid crab (Decapoda, Brachyura, Goneplacoidea) from shallow waters in Japan. Studies on Eumalacostraca: a Homage to Masatsune Takeda. Crustaceana Monographs, 17, 137 - 154." type="journal article" year="2012">
Komai
<emphasis id="4BB69637FFF7FFAFFB4EF8F8FB21F8F9" box="[1203,1260,1831,1856]" italics="true" pageId="84" pageNumber="85">et al.</emphasis>
2012
</bibRefCitation>
: figs. 3D, 5A, B), largest chela of male (
<figureCitation id="E1F956A0FFF7FFAFFE17F89DFDF0F8DD" box="[490,573,1868,1893]" captionStart="FIGURE 48" captionStartId="121.[151,250,1288,1310]" captionTargetBox="[151,1436,217,1267]" captionTargetId="figure@121.[151,1436,193,1267]" captionTargetPageId="121" captionText="FIGURE 48. Chelae of species of Megaesthesius, Alainthesius n. gen. and Microtopsis. A, B, Microtopsis takedai, Japan: holotype male 2.4 × 3.1 mm (after Komai et al. 2012: fig. 3 B, D); C, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); D, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); E, Megaesthesius yokoyai, Japan: male (after Takeda &amp; Miyake 1969: fig. 4 d); F, Alainthesius signatus n. sp., Madagascar: holotype male 3.5 × 3.8 mm (after Crosnier 1975: fig. 1 d); G, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN- IU- 2013 - 9073); H, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012)." httpUri="https://zenodo.org/record/272694/files/figure.png" pageId="84" pageNumber="85">Fig. 48</figureCitation>
A;
<bibRefCitation id="1D5337D4FFF7FFAFFDA3F89DFCE0F8DD" author="Komai" box="[606,813,1868,1893]" pageId="84" pageNumber="85" refString="Komai, T., Ng, P. K. L. &amp; Yamada, Y. (2012) A new genus and new species of chasmocarcinid crab (Decapoda, Brachyura, Goneplacoidea) from shallow waters in Japan. Studies on Eumalacostraca: a Homage to Masatsune Takeda. Crustaceana Monographs, 17, 137 - 154." type="journal article" year="2012">
Komai
<emphasis id="4BB69637FFF7FFAFFD4FF89CFD27F8DC" box="[690,746,1868,1893]" italics="true" pageId="84" pageNumber="85">et al.</emphasis>
2012
</bibRefCitation>
: 3B) high, with broad teeth on pollex, dentate dactylus. Inner margin of cheliped carpus (
<figureCitation id="E1F956A0FFF7FFAFFDF0F8BEFD93F831" box="[525,606,1903,1928]" captionStart="FIGURE 18" captionStartId="85.[151,250,1325,1347]" captionTargetBox="[151,1436,517,1304]" captionTargetId="figure@85.[151,1436,517,1304]" captionTargetPageId="85" captionText="FIGURE 18. Overall dorsal view of species of Microtopsis. A, M. takedai, Japan: paratype female 3.6 × 2.7 mm (ZRC 2012.0146); B, M. teschi n. sp., Indonesia: holotype male 1.5 × 2.1 mm (NNM-ZMA); C, M. teschi n. sp., Indonesia: paratype male 1.5 × 1.9 mm (NNM-ZMA)." httpUri="https://zenodo.org/record/272664/files/figure.png" pageId="84" pageNumber="85">Fig. 18</figureCitation>
A;
<bibRefCitation id="1D5337D4FFF7FFAFFD7FF8BEFC9CF831" author="Komai" box="[642,849,1903,1928]" pageId="84" pageNumber="85" refString="Komai, T., Ng, P. K. L. &amp; Yamada, Y. (2012) A new genus and new species of chasmocarcinid crab (Decapoda, Brachyura, Goneplacoidea) from shallow waters in Japan. Studies on Eumalacostraca: a Homage to Masatsune Takeda. Crustaceana Monographs, 17, 137 - 154." type="journal article" year="2012">
Komai
<emphasis id="4BB69637FFF7FFAFFD28F8A0FCC3F831" box="[725,782,1903,1928]" italics="true" pageId="84" pageNumber="85">et al.</emphasis>
2012
</bibRefCitation>
: 3C, E) with short tooth. Ventral surface of cheliped merus with short row of low microscopic tubercles on posterior outer margin in both sexes. Meri of ambulatory legs unarmed; P5 merus 0.5 cl in
<typeStatus id="A679F487FFF7FFAFFDE9F868FDB8F868" box="[532,629,1977,2001]" pageId="84" pageNumber="85" type="paratype">paratype</typeStatus>
female. Fused thoracic sternites 1, 2 (
<figureCitation id="E1F956A0FFF7FFAFFBD8F869FBB4F868" box="[1061,1145,1976,2001]" captionStart="FIGURE 55" captionStartId="128.[151,250,1514,1536]" captionTargetBox="[151,1436,179,1493]" captionTargetId="figure@128.[151,1436,179,1493]" captionTargetPageId="128" captionText="FIGURE 55. Anterior thoracic sternum and male pleon of species of Statommatia n. gen., Microtopsis and Megaesthesius. A, Statommatia granulosa n. sp., New Caledonia: holotype male 4.1 × 4.7 mm (MNHN-IU- 2013 - 9454); B, Microtopsis takedai, Japan: paratype male, carapace damaged (ZRC 2012.0146); C, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); D, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); E, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472); F, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073)." httpUri="https://zenodo.org/record/272701/files/figure.png" pageId="84" pageNumber="85">Fig. 55</figureCitation>
B;
<bibRefCitation id="1D5337D4FFF7FFAFFB64F869FAA7F869" author="Komai" box="[1177,1386,1976,2001]" pageId="84" pageNumber="85" refString="Komai, T., Ng, P. K. L. &amp; Yamada, Y. (2012) A new genus and new species of chasmocarcinid crab (Decapoda, Brachyura, Goneplacoidea) from shallow waters in Japan. Studies on Eumalacostraca: a Homage to Masatsune Takeda. Crustaceana Monographs, 17, 137 - 154." type="journal article" year="2012">
Komai
<emphasis id="4BB69637FFF7FFAFFB10F868FAEBF868" box="[1261,1318,1976,2001]" italics="true" pageId="84" pageNumber="85">et al.</emphasis>
2012
</bibRefCitation>
: fig. 2D) broadly triangular, proportionally long, short; fused sternites 3, 4 (
<figureCitation id="E1F956A0FFF7FFAFFC4CF80DFBC3F84D" box="[945,1038,2012,2037]" captionStart="FIGURE 88" captionStartId="161.[151,250,1770,1792]" captionTargetBox="[151,1436,193,1749]" captionTargetId="figure@161.[151,1436,193,1749]" captionTargetPageId="161" captionText="FIGURE 88. Female thoracic sternum and pleon of species of Statommatia n. gen., Tenagopelta n. gen., Microtopsis and Megaesthesius. A, Statommatia apta, Philippines: female 8.5 × 10.6 mm (ZRC 2015.238); B, Statommatia malagasy n. sp., Madagascar: paratype female 9.4 × 8.1 mm (ZRC 2015.215); C, Tenagopelta pacifica n. sp., Philippines: female 9.2 × 10.4 mm (ZRC 2015.187); D, Tenagopelta brachyphallus n. sp., Australia: female 6.9 × 7.6 mm (NMV J 61680); E, Microtopsis takedai, Japan: paratype female 3.6 × 2.7 mm (ZRC 2012.0146); F, Megaesthesius sagedae, Singapore: female 3.0 × 3.6 mm (ZRC 2013.1804); G, Megaesthesius migmus, Philippines: paratype female 3.9 × 4.5 mm (ZRC 2015.245); H, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012). Vulvae exposed in E." httpUri="https://zenodo.org/record/272734/files/figure.png" pageId="84" pageNumber="85">Figs. 88</figureCitation>
E; 92A;
<bibRefCitation id="1D5337D4FFF7FFAFFB91F80DFAF6F84D" author="Komai" box="[1132,1339,2012,2037]" pageId="84" pageNumber="85" refString="Komai, T., Ng, P. K. L. &amp; Yamada, Y. (2012) A new genus and new species of chasmocarcinid crab (Decapoda, Brachyura, Goneplacoidea) from shallow waters in Japan. Studies on Eumalacostraca: a Homage to Masatsune Takeda. Crustaceana Monographs, 17, 137 - 154." type="journal article" year="2012">
Komai
<emphasis id="4BB69637FFF7FFAFFB42F80CFB3AF84C" box="[1215,1271,2012,2037]" italics="true" pageId="84" pageNumber="85">et al.</emphasis>
2012
</bibRefCitation>
: fig. 2E) relatively broad. Male pleon (
<figureCitation id="E1F956A0FFF6FFAEFE18FF46FD8FFF09" box="[485,578,151,176]" captionStart="FIGURE 55" captionStartId="128.[151,250,1514,1536]" captionTargetBox="[151,1436,179,1493]" captionTargetId="figure@128.[151,1436,179,1493]" captionTargetPageId="128" captionText="FIGURE 55. Anterior thoracic sternum and male pleon of species of Statommatia n. gen., Microtopsis and Megaesthesius. A, Statommatia granulosa n. sp., New Caledonia: holotype male 4.1 × 4.7 mm (MNHN-IU- 2013 - 9454); B, Microtopsis takedai, Japan: paratype male, carapace damaged (ZRC 2012.0146); C, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); D, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); E, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472); F, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073)." httpUri="https://zenodo.org/record/272701/files/figure.png" pageId="85" pageNumber="86">Figs. 55</figureCitation>
B; 62A; 80B;
<bibRefCitation id="1D5337D4FFF6FFAEFD22FF46FC62FF09" author="Komai" box="[735,943,151,176]" pageId="85" pageNumber="86" refString="Komai, T., Ng, P. K. L. &amp; Yamada, Y. (2012) A new genus and new species of chasmocarcinid crab (Decapoda, Brachyura, Goneplacoidea) from shallow waters in Japan. Studies on Eumalacostraca: a Homage to Masatsune Takeda. Crustaceana Monographs, 17, 137 - 154." type="journal article" year="2012">
Komai
<emphasis id="4BB69637FFF6FFAEFCCEFF48FCA1FF09" box="[819,876,151,176]" italics="true" pageId="85" pageNumber="86">et al.</emphasis>
2012
</bibRefCitation>
: fig. 2D; 2H) with lateral margins of somite 6, fused somites 3‒5 convex; telson proportionally short; postero-lateral regions slightly swollen. Sterno-pleonal cavity deep. Press-button for pleonal holding (
<bibRefCitation id="1D5337D4FFF6FFAEFD4FFF0EFC41FF41" author="Komai" box="[690,908,223,248]" pageId="85" pageNumber="86" refString="Komai, T., Ng, P. K. L. &amp; Yamada, Y. (2012) A new genus and new species of chasmocarcinid crab (Decapoda, Brachyura, Goneplacoidea) from shallow waters in Japan. Studies on Eumalacostraca: a Homage to Masatsune Takeda. Crustaceana Monographs, 17, 137 - 154." type="journal article" year="2012">
Komai
<emphasis id="4BB69637FFF6FFAEFCF7FF30FC88FF41" box="[778,837,223,248]" italics="true" pageId="85" pageNumber="86">et al.</emphasis>
2012
</bibRefCitation>
: fig. 2G) as small, short tubercle posterior to thoracic sternal suture 4/5 near edge of sterno-pleonal cavity. G1 (
<figureCitation id="E1F956A0FFF6FFAEFC8AFED5FC0BFEA5" box="[887,966,260,285]" captionStart="FIGURE 81" captionStartId="154.[151,250,1949,1971]" captionTargetBox="[218,1333,202,1887]" captionTargetId="figure@154.[207,1366,195,1912]" captionTargetPageId="154" captionText="FIGURE 81. Gonopods of species of Microtopsis. A ‒ F, M. takedai, Japan: holotype male 2.4 × 3.1 mm (after Komai et al. 2012: fig. 4); G ‒ N, M. teschi n. sp., Indonesia: holotype male 1.5 × 2.1 mm (NNM-ZMA). A ‒ C, G ‒ I, left G 1; D, E, J ‒ L, distal part of left G 1; F, M, left G 2; N, distal part of left G 2. Scales: G ‒ I, M = 0.1 mm; J ‒ L = 0.05 mm; N = 0.01 mm." httpUri="https://zenodo.org/record/272727/files/figure.png" pageId="85" pageNumber="86">Fig. 81</figureCitation>
A‒E;
<bibRefCitation id="1D5337D4FFF6FFAEFBF4FED5FB1BFEA5" author="Komai" box="[1033,1238,260,285]" pageId="85" pageNumber="86" refString="Komai, T., Ng, P. K. L. &amp; Yamada, Y. (2012) A new genus and new species of chasmocarcinid crab (Decapoda, Brachyura, Goneplacoidea) from shallow waters in Japan. Studies on Eumalacostraca: a Homage to Masatsune Takeda. Crustaceana Monographs, 17, 137 - 154." type="journal article" year="2012">
Komai
<emphasis id="4BB69637FFF6FFAEFBA1FED4FB59FEA4" box="[1116,1172,260,285]" italics="true" pageId="85" pageNumber="86">et al.</emphasis>
2012
</bibRefCitation>
: fig. 4A‒E) stout, short, median part distinctly twisted, distal part slightly twisted, distal segment with short spinules. G2 (
<figureCitation id="E1F956A0FFF6FFAEFAD6FEF6FAB0FEF9" box="[1323,1405,295,320]" captionStart="FIGURE 81" captionStartId="154.[151,250,1949,1971]" captionTargetBox="[218,1333,202,1887]" captionTargetId="figure@154.[207,1366,195,1912]" captionTargetPageId="154" captionText="FIGURE 81. Gonopods of species of Microtopsis. A ‒ F, M. takedai, Japan: holotype male 2.4 × 3.1 mm (after Komai et al. 2012: fig. 4); G ‒ N, M. teschi n. sp., Indonesia: holotype male 1.5 × 2.1 mm (NNM-ZMA). A ‒ C, G ‒ I, left G 1; D, E, J ‒ L, distal part of left G 1; F, M, left G 2; N, distal part of left G 2. Scales: G ‒ I, M = 0.1 mm; J ‒ L = 0.05 mm; N = 0.01 mm." httpUri="https://zenodo.org/record/272727/files/figure.png" pageId="85" pageNumber="86">Fig. 81</figureCitation>
F);
<bibRefCitation id="1D5337D4FFF6FFAEFF6AFE9DFEBDFEDD" author="Komai" box="[151,368,332,357]" pageId="85" pageNumber="86" refString="Komai, T., Ng, P. K. L. &amp; Yamada, Y. (2012) A new genus and new species of chasmocarcinid crab (Decapoda, Brachyura, Goneplacoidea) from shallow waters in Japan. Studies on Eumalacostraca: a Homage to Masatsune Takeda. Crustaceana Monographs, 17, 137 - 154." type="journal article" year="2012">
Komai
<emphasis id="4BB69637FFF6FFAEFF13FE9CFEE7FEDC" box="[238,298,332,357]" italics="true" pageId="85" pageNumber="86">et al.</emphasis>
2012
</bibRefCitation>
: fig. 4F) slightly curved, slender, pointed distal segment, almost as long as G1. Somites of female pleon (
<figureCitation id="E1F956A0FFF6FFAEFEBDFEBEFE5BFE31" box="[320,406,367,392]" captionStart="FIGURE 88" captionStartId="161.[151,250,1770,1792]" captionTargetBox="[151,1436,193,1749]" captionTargetId="figure@161.[151,1436,193,1749]" captionTargetPageId="161" captionText="FIGURE 88. Female thoracic sternum and pleon of species of Statommatia n. gen., Tenagopelta n. gen., Microtopsis and Megaesthesius. A, Statommatia apta, Philippines: female 8.5 × 10.6 mm (ZRC 2015.238); B, Statommatia malagasy n. sp., Madagascar: paratype female 9.4 × 8.1 mm (ZRC 2015.215); C, Tenagopelta pacifica n. sp., Philippines: female 9.2 × 10.4 mm (ZRC 2015.187); D, Tenagopelta brachyphallus n. sp., Australia: female 6.9 × 7.6 mm (NMV J 61680); E, Microtopsis takedai, Japan: paratype female 3.6 × 2.7 mm (ZRC 2012.0146); F, Megaesthesius sagedae, Singapore: female 3.0 × 3.6 mm (ZRC 2013.1804); G, Megaesthesius migmus, Philippines: paratype female 3.9 × 4.5 mm (ZRC 2015.245); H, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012). Vulvae exposed in E." httpUri="https://zenodo.org/record/272734/files/figure.png" pageId="85" pageNumber="86">Fig. 88</figureCitation>
E;
<bibRefCitation id="1D5337D4FFF6FFAEFE44FEBEFD59FE31" author="Komai" box="[441,660,367,392]" pageId="85" pageNumber="86" refString="Komai, T., Ng, P. K. L. &amp; Yamada, Y. (2012) A new genus and new species of chasmocarcinid crab (Decapoda, Brachyura, Goneplacoidea) from shallow waters in Japan. Studies on Eumalacostraca: a Homage to Masatsune Takeda. Crustaceana Monographs, 17, 137 - 154." type="journal article" year="2012">
Komai
<emphasis id="4BB69637FFF6FFAEFDECFEA0FD80FE31" box="[529,589,367,392]" italics="true" pageId="85" pageNumber="86">et al.</emphasis>
2012
</bibRefCitation>
: fig. 5D) with convex lateral margins; telson proportionally short. Sterno-pleonal cavity of female (
<figureCitation id="E1F956A0FFF6FFAEFDFBFE45FD95FE15" box="[518,600,404,429]" captionStart="FIGURE 92" captionStartId="165.[151,250,934,956]" captionTargetBox="[151,1436,193,912]" captionTargetId="figure@165.[151,1436,193,912]" captionTargetPageId="165" captionText="FIGURE 92. Vulvae of species of Microtopsis and Megaesthesius. A, Microtopsis takedai, Japan: paratype female 3.6 × 2.7 mm (ZRC 2012.0146); B, Megaesthesius sagedae, Singapore: female carapace length 2.9 mm, sides damaged (ZRC 1995.883); C, Megaesthesius migmus n. sp., Philippines: paratype female 3.9 × 4.5 mm (ZRC 2015.245); D, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012)." httpUri="https://zenodo.org/record/272738/files/figure.png" pageId="85" pageNumber="86">Fig. 92</figureCitation>
A;
<bibRefCitation id="1D5337D4FFF6FFAEFD84FE45FC8AFE15" author="Komai" box="[633,839,404,429]" pageId="85" pageNumber="86" refString="Komai, T., Ng, P. K. L. &amp; Yamada, Y. (2012) A new genus and new species of chasmocarcinid crab (Decapoda, Brachyura, Goneplacoidea) from shallow waters in Japan. Studies on Eumalacostraca: a Homage to Masatsune Takeda. Crustaceana Monographs, 17, 137 - 154." type="journal article" year="2012">
Komai
<emphasis id="4BB69637FFF6FFAEFD31FE44FCC9FE14" box="[716,772,404,429]" italics="true" pageId="85" pageNumber="86">et al.</emphasis>
2012
</bibRefCitation>
: fig. 5C) moderately deep, vulvae far apart from each other on outer margins of cavity close to suture 5/6.
</paragraph>
</subSubSection>
<caption id="2DBD1AADFFF6FFAEFF6AFAFCFE3CFA38" httpUri="https://zenodo.org/record/272664/files/figure.png" pageId="85" pageNumber="86" targetBox="[151,1436,517,1304]" targetPageId="85">
<paragraph id="797D4A25FFF6FFAEFF6AFAFCFE3CFA38" blockId="85.[151,1436,1325,1409]" pageId="85" pageNumber="86">
<emphasis id="4BB69637FFF6FFAEFF6AFAFCFEE9FAFA" bold="true" box="[151,292,1325,1347]" pageId="85" pageNumber="86">FIGURE 18.</emphasis>
Overall dorsal view of species of
<taxonomicName id="BEC231A6FFF6FFAEFD66FAFEFCDEFAFD" box="[667,787,1327,1348]" class="Malacostraca" family="Chasmocarcinidae" genus="Microtopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="85" pageNumber="86" phylum="Arthropoda" rank="genus">
<emphasis id="4BB69637FFF6FFAEFD66FAFEFCDEFAFD" box="[667,787,1327,1348]" italics="true" pageId="85" pageNumber="86">Microtopsis</emphasis>
</taxonomicName>
. A,
<taxonomicName id="BEC231A6FFF6FFAEFCBBFAFEFC7AFAFA" box="[838,951,1325,1348]" class="Malacostraca" family="Chasmocarcinidae" genus="Microtopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="takedai">
<emphasis id="4BB69637FFF6FFAEFCBBFAFEFC7AFAFA" box="[838,951,1325,1348]" italics="true" pageId="85" pageNumber="86">M. takedai</emphasis>
</taxonomicName>
, Japan: paratype female 3.6 × 2.7 mm (ZRC 2012.0146); B,
<taxonomicName id="BEC231A6FFF6FFAEFECBFA9CFE5EFADB" box="[310,403,1356,1378]" class="Malacostraca" family="Chasmocarcinidae" genus="Microtopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="teschi" status="sp. nov.">
<emphasis id="4BB69637FFF6FFAEFECBFA9CFE5EFADB" box="[310,403,1356,1378]" italics="true" pageId="85" pageNumber="86">M. teschi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50852B4CFFF6FFAEFE64FA9CFE1FFADB" box="[409,466,1357,1378]" pageId="85" pageNumber="86" rank="species">
<emphasis id="4BB69637FFF6FFAEFE64FA9CFE1FFADB" bold="true" box="[409,466,1357,1378]" pageId="85" pageNumber="86">n. sp.</emphasis>
</taxonomicNameLabel>
, Indonesia: holotype male 1.5 × 2.1 mm (NNM-ZMA); C,
<taxonomicName id="BEC231A6FFF6FFAEFBD1FA9CFB45FADB" box="[1068,1160,1356,1378]" class="Malacostraca" family="Chasmocarcinidae" genus="Microtopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="teschi" status="sp. nov.">
<emphasis id="4BB69637FFF6FFAEFBD1FA9CFB45FADB" box="[1068,1160,1356,1378]" italics="true" pageId="85" pageNumber="86">M. teschi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50852B4CFFF6FFAEFB72FA9CFB05FADB" box="[1167,1224,1357,1378]" pageId="85" pageNumber="86" rank="species">
<emphasis id="4BB69637FFF6FFAEFB72FA9CFB05FADB" bold="true" box="[1167,1224,1357,1378]" pageId="85" pageNumber="86">n. sp.</emphasis>
</taxonomicNameLabel>
, Indonesia: paratype male 1.5 × 1.9 mm (NNM-ZMA).
</paragraph>
</caption>
<subSubSection id="31D819AEFFF6FFAEFF3AFA7CFCBCF9EF" pageId="85" pageNumber="86" type="discussion">
<paragraph id="797D4A25FFF6FFAEFF3AFA7CFCBCF9EF" blockId="85.[151,1437,1453,1658]" pageId="85" pageNumber="86">
<emphasis id="4BB69637FFF6FFAEFF3AFA7CFEF9FA7F" bold="true" box="[199,308,1453,1478]" pageId="85" pageNumber="86">Remarks</emphasis>
. The third maxilliped merus is incorrectly described and illustrated by
<bibRefCitation id="1D5337D4FFF6FFAEFBAAFA7CFAFDFA7F" author="Komai" box="[1111,1328,1453,1478]" pageId="85" pageNumber="86" refString="Komai, T., Ng, P. K. L. &amp; Yamada, Y. (2012) A new genus and new species of chasmocarcinid crab (Decapoda, Brachyura, Goneplacoidea) from shallow waters in Japan. Studies on Eumalacostraca: a Homage to Masatsune Takeda. Crustaceana Monographs, 17, 137 - 154." type="journal article" year="2012">
Komai
<emphasis id="4BB69637FFF6FFAEFB56FA7EFB11FA7F" box="[1195,1244,1453,1478]" italics="true" pageId="85" pageNumber="86">et al</emphasis>
. (2012
</bibRefCitation>
: fig. 3A). It is not subtriangular, but rather more subcircular, with the anterior margin even rather than distinctly produced. The “supplementary plate” (
<figureCitation id="E1F956A0FFF6FFAEFE2AFA24FDE6F9B7" box="[471,555,1525,1550]" captionStart="FIGURE 62" captionStartId="135.[151,250,926,948]" captionTargetBox="[151,1436,303,904]" captionTargetId="figure@135.[151,1436,303,904]" captionTargetPageId="135" captionText="FIGURE 62. Thoracic sternites 7 and 8 and male pleon of species of Microtopsis and Megaesthesius. A, Microtopsis takedai, Japan: paratype male, carapace damaged (ZRC 2012.0146); B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); C, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); D, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073)." httpUri="https://zenodo.org/record/272708/files/figure.png" pageId="85" pageNumber="86">Fig. 62</figureCitation>
A), which is about the same short length along thoracic sternite 8 and reaches episternite
<quantity id="BE3AE7C0FFF6FFAEFEE8F9CBFE8FF98B" box="[277,322,1561,1586]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="85" pageNumber="86" unit="in" value="7.0">7 in</quantity>
the
<typeStatus id="A679F487FFF6FFAEFE8BF9CBFE1AF98B" box="[374,471,1562,1586]" pageId="85" pageNumber="86" type="paratype">paratype</typeStatus>
male (ZRC 2012.0146), is not as narrow and not reaching the posterior edge of the pleon as depicted in the
<typeStatus id="A679F487FFF6FFAEFE5BF9ECFDC5F9EF" box="[422,520,1597,1622]" pageId="85" pageNumber="86" type="holotype">holotype</typeStatus>
by
<bibRefCitation id="1D5337D4FFF6FFAEFDCFF9ECFCC7F9EF" author="Komai" box="[562,778,1597,1622]" pageId="85" pageNumber="86" refString="Komai, T., Ng, P. K. L. &amp; Yamada, Y. (2012) A new genus and new species of chasmocarcinid crab (Decapoda, Brachyura, Goneplacoidea) from shallow waters in Japan. Studies on Eumalacostraca: a Homage to Masatsune Takeda. Crustaceana Monographs, 17, 137 - 154." type="journal article" year="2012">
Komai
<emphasis id="4BB69637FFF6FFAEFD78F9EEFD7BF9EF" box="[645,694,1597,1622]" italics="true" pageId="85" pageNumber="86">et al</emphasis>
. (2012
</bibRefCitation>
: fig. 3E).
</paragraph>
</subSubSection>
<subSubSection id="31D819AEFFF6FFAEFF3AF9B0FCD5F9C0" box="[199,792,1633,1658]" pageId="85" pageNumber="86" type="distribution">
<paragraph id="797D4A25FFF6FFAEFF3AF9B0FCD5F9C0" blockId="85.[151,1437,1453,1658]" box="[199,792,1633,1658]" pageId="85" pageNumber="86">
<emphasis id="4BB69637FFF6FFAEFF3AF9B0FE94F9C3" bold="true" box="[199,345,1633,1658]" pageId="85" pageNumber="86">Distribution</emphasis>
. Known only from
<collectingCountry id="01D50AB5FFF6FFAEFDCAF9B3FDB7F9C0" box="[567,634,1634,1657]" name="Japan" pageId="85" pageNumber="86">Japan</collectingCountry>
. Depth:
<specimenCount id="6FC481ACFFF6FFAEFD25F9B3FCD5F9C0" box="[728,792,1634,1658]" pageId="85" pageNumber="86" type="generic">
<quantity id="BE3AE7C0FFF6FFAEFD25F9B3FCD9F9C0" box="[728,788,1634,1658]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="85" pageNumber="86" unit="m" value="13.0">13 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
</treatment>
</document>
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