250 lines
32 KiB
XML
250 lines
32 KiB
XML
<document id="CC5DA5B36D2EDA731BB30B0473FC7AF6" ID-CLB-Dataset="37321" ID-DOI="10.11646/zootaxa.4209.1.1" ID-GBIF-Dataset="2de5b7bf-3975-458a-b200-07ec69f4e013" ID-ISSN="1175-5326" ID-Zenodo-Dep="272646" ID-ZooBank="849BAB5C-464A-4B4A-A586-5742411EDC01" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484407904318" checkinUser="plazi" docAuthor="Ng, Peter K. L. & Castro, Peter" docDate="2016" docId="F16BFB33FFF4FFA3FF6AFDE3FBA5FE76" docLanguage="en" docName="zootaxa.4209.1.1.pdf.imf" docOrigin="Zootaxa 4209 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Megaesthesius Rathbun 1909" docType="treatment" docVersion="8" lastPageNumber="89" masterDocId="0D52834BFFA3FFFBFFFDFFD1FFCDFFB9" masterDocTitle="Revision of the family Chasmocarcinidae Serène, 1964 (Crustacea, Brachyura, Goneplacoidea)" masterLastPageNumber="182" masterPageNumber="1" pageNumber="88" updateTime="1698435566492" updateUser="ExternalLinkService">
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<mods:title id="BBCD57D8582FCC6D9A26E81ABCEA3589">Revision of the family Chasmocarcinidae Serène, 1964 (Crustacea, Brachyura, Goneplacoidea)</mods:title>
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<mods:namePart id="87765CA99B593D9D2EBD83397DE03587">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="6E11B9A16E081C5F4C2B5315172AC97A">Castro, Peter</mods:namePart>
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<mods:title id="9EB11E23684E20CA907967D9EB0C19AA">Zootaxa</mods:title>
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<mods:date id="D5807CE93CBB4D9900317E832049D6CB">2016</mods:date>
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<paragraph id="797D4A25FFF4FFACFF6AFDE3FD92FDF5" blockId="87.[151,607,562,588]" box="[151,607,562,588]" pageId="87" pageNumber="88">
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<heading id="2235FD49FFF4FFACFF6AFDE3FD92FDF5" bold="true" box="[151,607,562,588]" fontSize="11" level="1" pageId="87" pageNumber="88" reason="1">
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<emphasis id="4BB69637FFF4FFACFF6AFDE3FD92FDF5" bold="true" box="[151,607,562,588]" pageId="87" pageNumber="88">
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Genus
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<taxonomicName id="BEC231A6FFF4FFACFF13FDE3FD92FDF5" ID-CoL="7PNNB" authority="Rathbun, 1909" authorityName="Rathbun" authorityYear="1909" box="[238,607,562,588]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
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<emphasis id="4BB69637FFF4FFACFF13FDE3FE52FDF5" bold="true" box="[238,415,562,588]" italics="true" pageId="87" pageNumber="88">Megaesthesius</emphasis>
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<bibRefCitation id="1D5337D4FFF4FFACFE5AFDE3FD92FDF5" author="Rathbun" box="[423,607,562,588]" pageId="87" pageNumber="88" refString="Rathbun, M. J. (1909) New crabs from the Gulf of Siam. Proceedings of the Biological Society of Washington, 22, 107 - 114." type="journal article" year="1909">Rathbun, 1909</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="31D819AEFFF4FFACFF6AFDA8FDEAFD75" pageId="87" pageNumber="88" type="reference_group">
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<paragraph id="797D4A25FFF4FFACFF6AFDA8FDEAFD75" blockId="87.[151,1437,633,717]" pageId="87" pageNumber="88">
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<treatmentCitationGroup id="59D26D0BFFF4FFACFF6AFDA8FDEAFD75" pageId="87" pageNumber="88">
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<taxonomicName id="BEC231A6FFF4FFACFF6AFDA8FDC5FD36" ID-CoL="7PNNB" authority="Rathbun, 1909: 112" authorityName="Rathbun" authorityPageNumber="112" authorityYear="1909" box="[151,520,633,655]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
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<emphasis id="4BB69637FFF4FFACFF6AFDA8FEE1FD36" box="[151,300,633,655]" italics="true" pageId="87" pageNumber="88">Megaesthesius</emphasis>
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<treatmentCitation id="F8636C34FFF4FFACFECBFDA8FDC5FD36" author="Rathbun" box="[310,520,633,655]" page="112" pageId="87" pageNumber="88" year="1909">
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<bibRefCitation id="1D5337D4FFF4FFACFECBFDA8FE1EFD36" author="Rathbun" box="[310,467,633,655]" pageId="87" pageNumber="88" refString="Rathbun, M. J. (1909) New crabs from the Gulf of Siam. Proceedings of the Biological Society of Washington, 22, 107 - 114." type="journal article" year="1909">Rathbun, 1909</bibRefCitation>
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: 112
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</treatmentCitation>
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; 1910: 344.—
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<treatmentCitation id="F8636C34FFF4FFACFD63FDA8FCA5FD36" author="Serene" box="[670,872,633,655]" page="175" pageId="87" pageNumber="88" year="1964">
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<bibRefCitation id="1D5337D4FFF4FFACFD63FDA8FCFFFD36" author="Serene" box="[670,818,633,655]" pageId="87" pageNumber="88" refString="Serene, R. (1964 b) Redescription du genre Megaesthesius Rathbun et definition des Chasmocarcininae, nouvelle sous-famille des Goneplacidae (Decapoda Brachyura). Crustaceana, 7 (3), 175 - 187." type="journal article" year="1964" yearSuffix="b">Serène 1964b</bibRefCitation>
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: 175
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</treatmentCitation>
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; 1968: 92 (in list).—
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<treatmentCitation id="F8636C34FFF4FFACFBB5FDA8FAEEFD36" author="Zarenkov" box="[1096,1315,633,655]" page="241" pageId="87" pageNumber="88" year="1972">
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<bibRefCitation id="1D5337D4FFF4FFACFBB5FDA8FB3CFD36" author="Zarenkov" box="[1096,1265,633,655]" pageId="87" pageNumber="88" refString="Zarenkov, N. A. (1972) [New data on Indo-Pacific crabs (Fam. Goneplacidae, Pinnotheridae, Parthenopidae, Dorippidae) and the problem of seasonal reproduction of decapod animals in the Bay of Tonkin]. In: [The Complex Investigations of Ocean Nature], part 3, Moscow State University, Moscow, pp. 229 - 253. [In Russian]" type="book chapter" year="1972">Zarenkov, 1972</bibRefCitation>
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||
: 241
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||
</treatmentCitation>
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.—Crosnier 1975: 110.—
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<treatmentCitation id="F8636C34FFF4FFACFEB4FD49FE22FD17" author="Sakai" box="[329,495,664,686]" page="542" pageId="87" pageNumber="88" year="1976">
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<bibRefCitation id="1D5337D4FFF4FFACFEB4FD49FE72FD17" author="Sakai" box="[329,447,664,686]" pageId="87" pageNumber="88" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Kodansha, Tokyo, vol. 1 [English text], xxix + 773 pp., figs. 1 - 379, maps 1 - 3; vol. 2 [Japanese text], 461 pp., figs. 1, 2; vol. 3 [plates], 16 pp. + pls. 1 - 251." type="book chapter" year="1976">Sakai 1976</bibRefCitation>
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: 542
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||
</treatmentCitation>
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||
[in key], 552.—Ng
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<emphasis id="4BB69637FFF4FFACFD47FD48FD20FD17" box="[698,749,664,686]" italics="true" pageId="87" pageNumber="88">et al.</emphasis>
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2008: 76 [in list].—
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||
<treatmentCitation id="F8636C34FFF4FFACFC44FD49FB7FFD17" author="De" box="[953,1202,664,686]" page="33" pageId="87" pageNumber="88" year="2009">
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<bibRefCitation id="1D5337D4FFF4FFACFC44FD49FB5DFD17" author="De" box="[953,1168,664,686]" pageId="87" pageNumber="88" refString="De Grave, S., Pentcheff, N. D., Ahyong, S. T., Chan, T. - Y., Crandall, K. A., Dworschak, P. C., Felder, D. L., Feldmann, R. M., Fransen, C. H. J. M., Goulding, L. Y. D., Lemaitre, R., Low, M. E. Y., Martin, J. W., Ng, P. K. L., Schweitzer, C. E., Tan, S. H., Tshudy, D. & Wetzer, R. (2009) A classification of living and fossil genera of decapod crustaceans. Raffles Bulletin of Zoology, suppl. 21, 1 - 109." type="journal article" year="2009">
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De Grave
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||
<emphasis id="4BB69637FFF4FFACFBDFFD48FB98FD17" box="[1058,1109,664,686]" italics="true" pageId="87" pageNumber="88">et al.</emphasis>
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2009
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</bibRefCitation>
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||
: 33
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||
</treatmentCitation>
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||
[in list].—Castro
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||
<emphasis id="4BB69637FFF4FFACFA97FD48FA50FD17" box="[1386,1437,664,686]" italics="true" pageId="87" pageNumber="88">et al.</emphasis>
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2010: 41.—
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<treatmentCitation id="F8636C34FFF4FFACFEC0FD67FDEEFD75" author="Davie" box="[317,547,694,717]" page="43" pageId="87" pageNumber="88" year="2015">
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<bibRefCitation id="1D5337D4FFF4FFACFEC0FD67FE33FD75" author="Davie" box="[317,510,694,717]" pageId="87" pageNumber="88" refString="Davie, P. J. F., Guinot, D. & Ng, P. K. L. (2015 a) Anatomy and functional morphology of Brachyura. In: Castro, P., Davie, P. J. F., Guinot, D., Schram, F. & Von Vaupel Klein, C. (Eds.), Treatise on Zoology - Anatomy, Taxonomy, Biology - The Crustacea, complementary to the volumes translated from the French of the Traite de Zoologie, 9 (C) (I), Decapoda: Brachyura (Part 1), Brill, Leiden, pp. 11 - 163. http: // dx. doi. org / 10.1163 / 9789004190832 _ 004" type="journal article" year="2015" yearSuffix="a">
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||
Davie
|
||
<emphasis id="4BB69637FFF4FFACFE7EFD69FE7BFD75" box="[387,438,694,717]" italics="true" pageId="87" pageNumber="88">et al.</emphasis>
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||
2015a
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||
</bibRefCitation>
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||
: 43
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="31D819AEFFF4FFACFF6AFD28FDDDFCE3" pageId="87" pageNumber="88" type="materials_examined">
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<paragraph id="797D4A25FFF4FFACFF6AFD28FE10FC8C" blockId="87.[151,970,761,930]" pageId="87" pageNumber="88">
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<emphasis id="4BB69637FFF4FFACFF6AFD28FEE6FCAB" bold="true" box="[151,299,761,786]" pageId="87" pageNumber="88">Type species</emphasis>
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.
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||
<taxonomicName id="BEC231A6FFF4FFACFEC4FD28FD21FCAB" authority="Rathbun, 1909" authorityName="Rathbun" authorityYear="1909" box="[313,748,761,786]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="sagedae">
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||
<emphasis id="4BB69637FFF4FFACFEC4FD28FD8DFCAB" box="[313,576,761,786]" italics="true" pageId="87" pageNumber="88">Megaesthesius sagedae</emphasis>
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<bibRefCitation id="1D5337D4FFF4FFACFDBAFD28FD21FCAB" author="Rathbun" box="[583,748,761,786]" pageId="87" pageNumber="88" refString="Rathbun, M. J. (1909) New crabs from the Gulf of Siam. Proceedings of the Biological Society of Washington, 22, 107 - 114." type="journal article" year="1909">Rathbun, 1909</bibRefCitation>
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</taxonomicName>
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||
(gender masculine)
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||
<emphasis id="4BB69637FFF4FFACFF3AFCCCFE18FC8F" bold="true" box="[199,469,797,822]" pageId="87" pageNumber="88">Other species included</emphasis>
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:
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</paragraph>
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<paragraph id="797D4A25FFF4FFACFF3AFC90FDDDFCE3" blockId="87.[151,970,761,930]" box="[199,528,833,858]" pageId="87" pageNumber="88">
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<taxonomicName id="BEC231A6FFF4FFACFF3AFC90FE07FCE0" box="[199,458,833,858]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="migmus" status="sp. nov.">
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<emphasis id="4BB69637FFF4FFACFF3AFC90FE07FCE0" box="[199,458,833,858]" italics="true" pageId="87" pageNumber="88">Megaesthesius migmus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="50852B4CFFF4FFACFE2CFC90FDDDFCE3" box="[465,528,833,858]" pageId="87" pageNumber="88" rank="species">
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<emphasis id="4BB69637FFF4FFACFE2CFC90FDDDFCE3" bold="true" box="[465,528,833,858]" pageId="87" pageNumber="88">n. sp.</emphasis>
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</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="31D819AEFFF4FFACFF3AFCB4FDA6FCC7" box="[199,619,869,894]" pageId="87" pageNumber="88" type="description">
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<paragraph id="797D4A25FFF4FFACFF3AFCB4FDA6FCC7" blockId="87.[151,970,761,930]" box="[199,619,869,894]" pageId="87" pageNumber="88">
|
||
<taxonomicName id="BEC231A6FFF4FFACFF3AFCB4FDA6FCC7" authority="Davie, 2013" authorityName="Davie" authorityYear="2013" box="[199,619,869,894]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="westralia">
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||
<emphasis id="4BB69637FFF4FFACFF3AFCB4FE11FCC7" box="[199,476,869,894]" italics="true" pageId="87" pageNumber="88">Megaesthesius westralia</emphasis>
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||
Davie, 2013
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
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<subSubSection id="31D819AEFFF4FFACFF3AFC58FD98FC1B" box="[199,597,905,930]" pageId="87" pageNumber="88" type="reference_group">
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<paragraph id="797D4A25FFF4FFACFF3AFC58FD98FC1B" blockId="87.[151,970,761,930]" box="[199,597,905,930]" pageId="87" pageNumber="88">
|
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<taxonomicName id="BEC231A6FFF4FFACFF3AFC58FD98FC1B" ID-CoL="3YTN3" authority="Sakai, 1939" authorityName="Sakai" authorityYear="1939" box="[199,597,905,930]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="yokoyai">
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||
<emphasis id="4BB69637FFF4FFACFF3AFC58FE07FC1B" box="[199,458,905,930]" italics="true" pageId="87" pageNumber="88">Megaesthesius yokoyai</emphasis>
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||
<bibRefCitation id="1D5337D4FFF4FFACFE2CFC58FD98FC1B" author="Sakai" box="[465,597,905,930]" pageId="87" pageNumber="88" refString="Sakai, T. (1939) Studies on the crabs of Japan, IV. Brachygnatha, Brachyrhyncha. 741 + 23 pp., figs. 1 - 129, pls. 42 - 111." type="book chapter" year="1939">Sakai, 1939</bibRefCitation>
|
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</taxonomicName>
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="31D819AEFFF4FFACFF6AFC00FCBCF92F" pageId="87" pageNumber="88" type="diagnosis">
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<paragraph id="797D4A25FFF4FFACFF6AFC00FCBCF92F" blockId="87.[151,1437,977,2010]" pageId="87" pageNumber="88">
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||
<emphasis id="4BB69637FFF4FFACFF6AFC00FEC7FC53" bold="true" box="[151,266,977,1002]" pageId="87" pageNumber="88">Diagnosis</emphasis>
|
||
. Small size, carapace subquadrate, high, strongly arcuate; front bilobed, with deep or shallow median cleft; anterolateral margins straight, armed with small teeth often arranged in 4 or 5 lobes; dorsal surface with conspicuous round granules. Epistome compressed, posterior margin with broad median lobe with fissure. Eye peduncle conspicuously long, immobile, cornea long, reduced, pigmentation reduced. Antennule greatly enlarged (particularly in males), cannot be folded. Third maxillipeds nearly fill buccal cavern when closed; merus subtriangular, outer margin convex, no defined anteroexternal angle; ischium subquadrate, about same length or longer than merus. Chelipeds subequal, nearly identical in both sexes; cutting margins of both chelipeds with broad, shallow teeth in both sexes; fingers of
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<taxonomicName id="BEC231A6FFF4FFACFD40FB1CFCCDFB5F" box="[701,768,1229,1254]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="minor">minor</taxonomicName>
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chela relatively broad, proportionally short, prominently flattened laterally, cutting margins blade-like. Inner margin of major cheliped carpus with small tooth. Ventral surface of cheliped merus smooth or with teeth. Ambulatory legs with minute teeth on anterior or posterior or both margins of most articles. Fused thoracic sternites 1, 2 broadly triangular, short; fused sternites 3, 4 relatively broad. Male pleon with lateral margins of somite 6, fused somites 3–5 slightly convex; postero-lateral regions slightly swollen; telson proportionally short or long. Sterno-pleonal cavity of male deep. Press-button for pleonal holding as tubercle at middle portion of thoracic somite 5 near edge of sterno-pleonal cavity. Male thoracic sternite 8 proportionally long, rectangular; “supplementary plate” conspicuously narrow, only reaching median portion of exposed thorax, short, slightly longer at rounded outer margin. Outer (ventral) surface of penis calcified, resembling plate between “supplementary plate” (unknown in
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<taxonomicName id="BEC231A6FFF4FFACFC98F9C3FC38F993" box="[869,1013,1553,1578]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="westralia">
|
||
<emphasis id="4BB69637FFF4FFACFC98F9C3FC38F993" box="[869,1013,1553,1578]" italics="true" pageId="87" pageNumber="88">M. westralia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="BEC231A6FFF4FFACFBFBF9C3FB48F993" box="[1030,1157,1553,1578]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="yokoyai">
|
||
<emphasis id="4BB69637FFF4FFACFBFBF9C3FB48F993" box="[1030,1157,1553,1578]" italics="true" pageId="87" pageNumber="88">M. yokoyai</emphasis>
|
||
</taxonomicName>
|
||
), sternite 8. G1 slender, distal segment straight with spinules. G2 straight, slender, distal segment directed inwards, nearly as long as G1. Somites of female pleon with gently convex margins; telson proportionally short. Sterno-pleonal cavity of female deep, vulvae on outer margins of cavity between sutures 5/6, 6/7.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="31D819AEFFF4FFA3FF3AF970FCE2FE14" lastPageId="88" lastPageNumber="89" pageId="87" pageNumber="88" type="discussion">
|
||
<paragraph id="797D4A25FFF4FFACFF3AF970FE6BF863" blockId="87.[151,1437,977,2010]" pageId="87" pageNumber="88">
|
||
<emphasis id="4BB69637FFF4FFACFF3AF970FEF9F903" bold="true" box="[199,308,1697,1722]" pageId="87" pageNumber="88">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="BEC231A6FFF4FFACFEB8F970FD6CF903" authority="Rathbun, 1909" authorityName="Rathbun" authorityYear="1909" box="[325,673,1697,1722]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4BB69637FFF4FFACFEB8F970FE27F903" box="[325,490,1697,1722]" italics="true" pageId="87" pageNumber="88">Megaesthesius</emphasis>
|
||
<bibRefCitation id="1D5337D4FFF4FFACFE08F970FD6CF903" author="Rathbun" box="[501,673,1697,1722]" pageId="87" pageNumber="88" refString="Rathbun, M. J. (1909) New crabs from the Gulf of Siam. Proceedings of the Biological Society of Washington, 22, 107 - 114." type="journal article" year="1909">Rathbun, 1909</bibRefCitation>
|
||
</taxonomicName>
|
||
, is unusual in many respects, even within the
|
||
<taxonomicName id="BEC231A6FFF4FFACFB3BF970FA54F903" box="[1222,1433,1697,1722]" class="Malacostraca" family="Chasmocarcinidae" kingdom="Animalia" order="Decapoda" pageId="87" pageNumber="88" phylum="Arthropoda" rank="family">Chasmocarcinidae</taxonomicName>
|
||
. Most outstanding, a character that has been discussed at length by
|
||
<bibRefCitation id="1D5337D4FFF4FFACFC55F917FB95F967" author="Serene" box="[936,1112,1733,1758]" pageId="87" pageNumber="88" refString="Serene, R. (1964 b) Redescription du genre Megaesthesius Rathbun et definition des Chasmocarcininae, nouvelle sous-famille des Goneplacidae (Decapoda Brachyura). Crustaceana, 7 (3), 175 - 187." type="journal article" year="1964" yearSuffix="b">Serène (1964b)</bibRefCitation>
|
||
and Crosnier (1975), is the disproportionately large and exposed antennular peduncle that is densely covered with sensory setae (aesthetascs). The eye peduncle is also large and long, but is fused to the orbit with the cornea being barely discernible. It is probably for these reasons that
|
||
<bibRefCitation id="1D5337D4FFF4FFACFE00F8E0FD10F8F3" author="Stevcic" box="[509,733,1841,1866]" pageId="87" pageNumber="88" refString="Stevcic, Z. (2005) The reclassification of brachyuran crabs (Crustacea: Decapoda: Brachyura). Natura Croatica (Fauna Croatica), 14, supplement 1, 1 - 159." type="journal article" year="2005">Števčić (2005: 108)</bibRefCitation>
|
||
established a new subfamily,
|
||
<taxonomicName id="BEC231A6FFF4FFACFBC8F8E0FB3EF8F3" box="[1077,1267,1841,1866]" class="Malacostraca" family="Chasmocarcinidae" kingdom="Animalia" order="Decapoda" pageId="87" pageNumber="88" phylum="Arthropoda" rank="subFamily" subFamily="Megaesthesiinae">Megaesthesiinae</taxonomicName>
|
||
, for the genus, although he did not explain why. We have also observed that the penis is covered by a calcified plate that lies between the “supplementary plate” and thoracic sternite 8 (
|
||
<figureCitation id="E1F956A0FFF4FFACFCCDF8A8FCB2F82B" box="[816,895,1913,1938]" captionStart="FIGURE 19" captionStartId="92.[151,250,1665,1687]" captionTargetBox="[203,1371,195,1618]" captionTargetId="figure@92.[203,1371,195,1632]" captionTargetPageId="92" captionText="FIGURE 19. Species of Megaesthesius and Alainthesius n. gen. A, B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); C ‒ F, Megaesthesius yokoyai (after Takeda & Miyake 1969 a: figs. 3 a, 4 f, 4 a, 4 e, respectively); G ‒ L, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472) (after Crosnier 1975: fig. 1 a, 1 k, 1 j, 1 b, 1 c, 1 i, respectively); M ‒ R, Megaesthesius westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: figs. 1 A, B, 2 A, D, E, 3 E); S, W, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); T, X, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); U, Y, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); V, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012). A, overall habitus; B, C, G, M, dorsal view of carapace; E, left P 4; F, I, frontal view of cephalothorax; H, O, frontal and anterolateral margin of carapace; N, ventral view of cephalothorax; J, right P 3; K, buccal cavity and posterior margin of epistome; Q, right P 4; R, right P 5; D, L, P, male pleon; S ‒ V, posterior margin of epistome; W ‒ Y, left thoracic sternites 7 and 8, “ supplementary plate ”, penis tube and penis. Abbreviations: cx 4 = coxa of P 4; cx 5 = coxa of P 5; p = penis; pt = calcified penis tube; sp = “ supplementary plate ”; st 7 = thoracic sternite 7; st 8 = thoracic sternite 8." httpUri="https://zenodo.org/record/272665/files/figure.png" pageId="87" pageNumber="88">Fig. 19</figureCitation>
|
||
W–Y), a character unique to
|
||
<taxonomicName id="BEC231A6FFF4FFACFB3BF8A8FA54F82B" box="[1222,1433,1913,1938]" class="Malacostraca" family="Chasmocarcinidae" kingdom="Animalia" order="Decapoda" pageId="87" pageNumber="88" phylum="Arthropoda" rank="family">Chasmocarcinidae</taxonomicName>
|
||
. On the basis of these characters, it seems reasonable that
|
||
<taxonomicName id="BEC231A6FFF4FFACFCE8F84CFC77F80F" box="[789,954,1949,1974]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4BB69637FFF4FFACFCE8F84CFC77F80F" box="[789,954,1949,1974]" italics="true" pageId="87" pageNumber="88">Megaesthesius</emphasis>
|
||
</taxonomicName>
|
||
should be placed in its own subfamily (see also Ng
|
||
<emphasis id="4BB69637FFF4FFACFF09F813FEE8F863" box="[244,293,1985,2010]" italics="true" pageId="87" pageNumber="88">et al</emphasis>
|
||
. 2008: 76).
|
||
</paragraph>
|
||
<paragraph id="797D4A25FFFBFFA3FF3AFF46FCE2FE14" blockId="88.[151,1436,151,464]" pageId="88" pageNumber="89">
|
||
<materialsCitation id="C9AA4078FFFBFFA3FF3AFF46FCE6FE14" ID-GBIF-Occurrence="1424726206" country="Madagascar" location="western Pacific" pageId="88" pageNumber="89" specimenCount="1">
|
||
Crosnier’s (1975) record of “
|
||
<taxonomicName id="BEC231A6FFFBFFA3FDE2FF46FCE6FF09" box="[543,811,151,176]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sagedae">
|
||
<emphasis id="4BB69637FFFBFFA3FDE2FF46FCE6FF09" box="[543,811,151,176]" italics="true" pageId="88" pageNumber="89">Megaesthesius sagedae</emphasis>
|
||
</taxonomicName>
|
||
” from
|
||
<collectingCountry id="01D50AB5FFFBFFA3FC78FF46FBC1FF09" box="[901,1036,151,176]" name="Madagascar" pageId="88" pageNumber="89">Madagascar</collectingCountry>
|
||
requires discussion. Although the carapace of the
|
||
<collectingCountry id="01D50AB5FFFBFFA3FEB8FF6DFE01FF6C" box="[325,460,188,213]" name="Madagascar" pageId="88" pageNumber="89">Madagascar</collectingCountry>
|
||
material superficially resembles
|
||
<taxonomicName id="BEC231A6FFFBFFA3FCC3FF6CFC72FF6C" box="[830,959,188,213]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sagedae">
|
||
<emphasis id="4BB69637FFFBFFA3FCC3FF6CFC72FF6C" box="[830,959,188,213]" italics="true" pageId="88" pageNumber="89">M. sagedae</emphasis>
|
||
</taxonomicName>
|
||
, it is clearly not
|
||
<taxonomicName id="BEC231A6FFFBFFA3FB8AFF6CFB35FF6C" box="[1143,1272,188,213]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sagedae">
|
||
<emphasis id="4BB69637FFFBFFA3FB8AFF6CFB35FF6C" box="[1143,1272,188,213]" italics="true" pageId="88" pageNumber="89">M. sagedae</emphasis>
|
||
</taxonomicName>
|
||
as is known at present (see below) and must be recognised as a new genus and new species. More significantly, its G2 is short, with the basal segment short and the distal segment spatuliform with only with a relatively short elongation, and as such, is only about 2/3 length of the G1 (
|
||
<figureCitation id="E1F956A0FFFBFFA3FD8EFEF6FD0BFEF9" box="[627,710,295,320]" captionStart="FIGURE 83" captionStartId="156.[151,250,1910,1932]" captionTargetBox="[264,1306,207,1869]" captionTargetId="figure@156.[248,1336,193,1910]" captionTargetPageId="156" captionText="FIGURE 83. Gonopods of species of Alainthesius n. gen. A ‒ C, A. signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472) (after Crosnier 1975: fig. 1 f ‒ h); D ‒ H, A. bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073). A, ventral view of right G 1; D, ventral view of left G 1; B, dorsal view of right G 1; E, G, dorsal view of distal part of left G 1; F, ventral view of distal part of left G 1; C, right G 2; H, left G 2. Scales = 0.1 mm." httpUri="https://zenodo.org/record/272729/files/figure.png" pageId="88" pageNumber="89">Fig. 83</figureCitation>
|
||
C) (G2 basal and distal segments markedly long, exceeding G1 length in other
|
||
<taxonomicName id="BEC231A6FFFBFFA3FEB0FE9DFE3FFEDC" box="[333,498,332,357]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="88" pageNumber="89" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4BB69637FFFBFFA3FEB0FE9DFE3FFEDC" box="[333,498,332,357]" italics="true" pageId="88" pageNumber="89">Megaesthesius</emphasis>
|
||
</taxonomicName>
|
||
species,
|
||
<figureCitation id="E1F956A0FFFBFFA3FD9DFE9DFD78FEDD" box="[608,693,332,357]" captionStart="FIGURE 82" captionStartId="155.[151,250,1823,1845]" captionTargetBox="[177,1417,213,1802]" captionTargetId="figure@155.[151,1436,193,1802]" captionTargetPageId="155" captionText="FIGURE 82. Gonopods of species of Megaesthesius. A ‒ C, M. sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); D ‒ F, M. migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); G ‒ I, M. yokoyai, Japan: male 3.5 × 3.8 mm (after Takeda & Miyake 1969: fig. 4 h, i); J ‒ M, M. westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: fig. 3 A ‒ D). A, D, G, H, J, K, ventral view of left G 1; B, E, I, K, L, dorsal view of left G 1; C, F, M, left G 2. Scales = 0.1 mm." httpUri="https://zenodo.org/record/272728/files/figure.png" pageId="88" pageNumber="89">Fig. 82</figureCitation>
|
||
C, F, M). It makes more sense to separate the new
|
||
<collectingCountry id="01D50AB5FFFBFFA3FAE8FE9DFA51FEDC" box="[1301,1436,332,357]" name="Madagascar" pageId="88" pageNumber="89">Madagascar</collectingCountry>
|
||
species into a new genus, here named
|
||
<taxonomicName id="BEC231A6FFFBFFA3FD9FFEBEFD20FE31" box="[610,749,367,392]" class="Malacostraca" family="Chasmocarcinidae" genus="Alainthesius" kingdom="Animalia" order="Decapoda" pageId="88" pageNumber="89" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="4BB69637FFFBFFA3FD9FFEBEFD20FE31" box="[610,749,367,392]" italics="true" pageId="88" pageNumber="89">Alainthesius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="50852B4CFFFBFFA3FD05FEBEFC87FE31" box="[760,842,367,392]" pageId="88" pageNumber="89" rank="genus">
|
||
<emphasis id="4BB69637FFFBFFA3FD05FEBEFC87FE31" bold="true" box="[760,842,367,392]" pageId="88" pageNumber="89">n. gen.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(see below). A second species of
|
||
<taxonomicName id="BEC231A6FFFBFFA3FB1AFEBEFABFFE31" box="[1255,1394,367,392]" class="Malacostraca" family="Chasmocarcinidae" genus="Alainthesius" kingdom="Animalia" order="Decapoda" pageId="88" pageNumber="157" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="4BB69637FFFBFFA3FB1AFEBEFABFFE31" box="[1255,1394,367,392]" italics="true" pageId="88" pageNumber="89">Alainthesius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="BEC231A6FFFBFFA3FA79FEA1FECCFE14" class="Malacostraca" family="Chasmocarcinidae" genus="Alainthesius" kingdom="Animalia" order="Decapoda" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="bertrandi" status="sp. nov.">
|
||
<emphasis id="4BB69637FFFBFFA3FA79FEA1FECCFE14" italics="true" pageId="88" pageNumber="89">A. bertrandi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="50852B4CFFFBFFA3FEFAFE45FE8BFE14" box="[263,326,404,429]" pageId="88" pageNumber="89" rank="species">
|
||
<emphasis id="4BB69637FFFBFFA3FEFAFE45FE8BFE14" bold="true" box="[263,326,404,429]" pageId="88" pageNumber="89">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, is also described from the
|
||
<location id="7C1D1CFEFFFBFFA3FD86FE44FCE6FE14" LSID="urn:lsid:plazi:treatment:F16BFB33FFF4FFA3FF6AFDE3FBA5FE76:7C1D1CFEFFFBFFA3FD86FE44FCE6FE14" box="[635,811,404,429]" country="Madagascar" name="western Pacific" pageId="88" pageNumber="89">western Pacific</location>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="31D819AEFFFBFFA3FF3AFE66FBA5FE76" box="[199,1128,439,464]" pageId="88" pageNumber="89" type="distribution">
|
||
<paragraph id="797D4A25FFFBFFA3FF3AFE66FBA5FE76" blockId="88.[151,1436,151,464]" box="[199,1128,439,464]" pageId="88" pageNumber="89">
|
||
<emphasis id="4BB69637FFFBFFA3FF3AFE66FE94FE69" bold="true" box="[199,345,439,464]" pageId="88" pageNumber="89">Distribution</emphasis>
|
||
. Indo-West Pacific region: eastern Indian and western Pacific oceans.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |