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<mods:title id="B297C5037FE293FBBD486DF476FD1492">Revision of the family Chasmocarcinidae Serène, 1964 (Crustacea, Brachyura, Goneplacoidea)</mods:title>
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<treatment id="F16BFB33FFBDFFE4FF6AFB04FC32F89E" ID-DOI="http://doi.org/10.5281/zenodo.5617086" ID-GBIF-Taxon="127419567" ID-Zenodo-Dep="5617086" LSID="urn:lsid:plazi:treatment:F16BFB33FFBDFFE4FF6AFB04FC32F89E" httpUri="http://treatment.plazi.org/id/F16BFB33FFBDFFE4FF6AFB04FC32F89E" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
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<paragraph id="797D4A25FFBDFFE5FF6AFB04FD0FFB56" blockId="30.[151,706,1237,1263]" box="[151,706,1237,1263]" pageId="30" pageNumber="31">
<heading id="2235FD49FFBDFFE5FF6AFB04FD0FFB56" bold="true" box="[151,706,1237,1263]" fontSize="11" level="1" pageId="30" pageNumber="31" reason="1">
<emphasis id="4BB69637FFBDFFE5FF6AFB04FD0FFB56" bold="true" box="[151,706,1237,1263]" pageId="30" pageNumber="31">
Genus
<taxonomicName id="BEC231A6FFBDFFE5FF13FB04FD0FFB56" ID-CoL="7NVQX" authority="Alcock &amp; Anderson, 1899" authorityName="Alcock &amp; Anderson" authorityYear="1899" box="[238,706,1237,1263]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="4BB69637FFBDFFE5FF13FB04FEB7FB56" bold="true" box="[238,378,1237,1263]" italics="true" pageId="30" pageNumber="31">Camatopsis</emphasis>
<bibRefCitation id="1D5337D4FFBDFFE5FE7CFB04FD0FFB56" author="Alcock" box="[385,706,1237,1263]" pageId="30" pageNumber="31" refString="Alcock, A. &amp; Anderson, A. R. S. (1899) An account of the deep-sea Crustacea dredged during the surveying-season of 1987 - 98. Natural history notes from H. M. Royal Indian Marine Survey Ship ' Investigator, ' Commander T. H. Heming, R. N., commanding, Series III, No. 2. Annals and Magazine of Natural History, ser. 7 (3), 1 - 27, 278 - 292." type="journal article" year="1899">Alcock &amp; Anderson, 1899</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
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<treatmentCitationGroup id="59D26D0BFFBDFFE5FF6AFACDFB28FAD6" pageId="30" pageNumber="31">
<taxonomicName id="BEC231A6FFBDFFE5FF6AFACDFD80FA8B" ID-CoL="7NVQX" authority="Alcock &amp; Anderson, 1899: 13" authorityName="Alcock &amp; Anderson" authorityPageNumber="13" authorityYear="1899" box="[151,589,1308,1330]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="4BB69637FFBDFFE5FF6AFACDFEC2FA88" box="[151,271,1308,1329]" italics="true" pageId="30" pageNumber="31">Camatopsis</emphasis>
<treatmentCitation id="F8636C34FFBDFFE5FEEAFACDFD80FA8B" author="Alcock" box="[279,589,1308,1330]" page="13" pageId="30" pageNumber="31" year="1899">
<bibRefCitation id="1D5337D4FFBDFFE5FEEAFACDFDE6FA8B" author="Alcock" box="[279,555,1308,1330]" pageId="30" pageNumber="31" refString="Alcock, A. &amp; Anderson, A. R. S. (1899) An account of the deep-sea Crustacea dredged during the surveying-season of 1987 - 98. Natural history notes from H. M. Royal Indian Marine Survey Ship ' Investigator, ' Commander T. H. Heming, R. N., commanding, Series III, No. 2. Annals and Magazine of Natural History, ser. 7 (3), 1 - 27, 278 - 292." type="journal article" year="1899">Alcock &amp; Anderson, 1899</bibRefCitation>
: 13
</treatmentCitation>
</taxonomicName>
.—Alcock 1899: 75; 1900: 318 [in key], 328.—
<treatmentCitation id="F8636C34FFBDFFE5FBBDFACDFB23FA8B" author="Tesch" box="[1088,1262,1308,1330]" page="202" pageId="30" pageNumber="31" year="1918">
<bibRefCitation id="1D5337D4FFBDFFE5FBBDFACDFB71FA8B" author="Tesch" box="[1088,1212,1308,1330]" pageId="30" pageNumber="31" refString="Tesch J. J. (1918) Goneplacidae and Pinnotheridae. The Decapoda Brachyura of the Siboga Expedition, 2. Siboga-Expeditie 39 (c 1), livr. 84, 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch 1918</bibRefCitation>
: 202
</treatmentCitation>
[in key], 234.— Balss 1957: 1658.—
<treatmentCitation id="F8636C34FFBDFFE5FE69FAEBFD9FFAE9" author="Serene" box="[404,594,1338,1360]" page="268" pageId="30" pageNumber="31" year="1964">
<bibRefCitation id="1D5337D4FFBDFFE5FE69FAEBFDEEFAE9" author="Serene" box="[404,547,1338,1360]" pageId="30" pageNumber="31" refString="Serene, R. (1964 a) Goneplacidae et Pinnotheridae recoltes par le Dr. Mortensen. Papers from Dr. Th. Mortensen's Pacific Expedition 1914 - 1916, part 80. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening I KObenhavn, 126, 181 - 282, pls. 16 - 24." type="journal article" year="1964" yearSuffix="a">Serène 1964a</bibRefCitation>
: 268
</treatmentCitation>
.—
<treatmentCitation id="F8636C34FFBDFFE5FD8CFAEBFCDAFAE9" author="Sakai" box="[625,791,1338,1360]" page="542" pageId="30" pageNumber="31" year="1976">
<bibRefCitation id="1D5337D4FFBDFFE5FD8CFAEBFD2AFAE9" author="Sakai" box="[625,743,1338,1360]" pageId="30" pageNumber="31" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Kodansha, Tokyo, vol. 1 [English text], xxix + 773 pp., figs. 1 - 379, maps 1 - 3; vol. 2 [Japanese text], 461 pp., figs. 1, 2; vol. 3 [plates], 16 pp. + pls. 1 - 251." type="book chapter" year="1976">Sakai 1976</bibRefCitation>
: 542
</treatmentCitation>
[in key], 552.—
<treatmentCitation id="F8636C34FFBDFFE5FC40FAEBFB03FAE9" author="Hsueh" box="[957,1230,1338,1360]" page="113" pageId="30" pageNumber="31" year="2002">
<bibRefCitation id="1D5337D4FFBDFFE5FC40FAEBFB6DFAE9" author="Hsueh" box="[957,1184,1338,1360]" pageId="30" pageNumber="31" refString="Hsueh, P. - W. &amp; Huang, J. - F. (2002) Crabs of the family Goneplacidae (Decapoda, Brachyura) from Taiwan. Crustaceana, 75 (2), 111 - 136. http: // dx. doi. org / 10.1163 / 15685400252902344" type="journal article" year="2002">Hsueh &amp; Huang 2002</bibRefCitation>
: 113
</treatmentCitation>
[in key].—Ng
<emphasis id="4BB69637FFBDFFE5FA94FAEAFA50FAE9" box="[1385,1437,1338,1360]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
2008: 76 [in list].—
<treatmentCitation id="F8636C34FFBDFFE5FE73FA88FD47FAD6" author="De" box="[398,650,1369,1391]" page="32" pageId="30" pageNumber="31" year="2009">
<bibRefCitation id="1D5337D4FFBDFFE5FE73FA88FDAAFAD6" author="De" box="[398,615,1369,1391]" pageId="30" pageNumber="31" refString="De Grave, S., Pentcheff, N. D., Ahyong, S. T., Chan, T. - Y., Crandall, K. A., Dworschak, P. C., Felder, D. L., Feldmann, R. M., Fransen, C. H. J. M., Goulding, L. Y. D., Lemaitre, R., Low, M. E. Y., Martin, J. W., Ng, P. K. L., Schweitzer, C. E., Tan, S. H., Tshudy, D. &amp; Wetzer, R. (2009) A classification of living and fossil genera of decapod crustaceans. Raffles Bulletin of Zoology, suppl. 21, 1 - 109." type="journal article" year="2009">
De Grave
<emphasis id="4BB69637FFBDFFE5FE05FA8BFDE6FAD6" box="[504,555,1369,1391]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
2009
</bibRefCitation>
: 32
</treatmentCitation>
[in list].—Castro
<emphasis id="4BB69637FFBDFFE5FCBFFA8BFCBBFAD6" box="[834,886,1369,1391]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
2010: 41.—
<treatmentCitation id="F8636C34FFBDFFE5FC0CFA88FB2CFAD6" author="Komai" box="[1009,1249,1369,1391]" page="144" pageId="30" pageNumber="31" year="2012">
<bibRefCitation id="1D5337D4FFBDFFE5FC0CFA88FB62FAD6" author="Komai" box="[1009,1199,1369,1391]" pageId="30" pageNumber="31" refString="Komai, T., Ng, P. K. L. &amp; Yamada, Y. (2012) A new genus and new species of chasmocarcinid crab (Decapoda, Brachyura, Goneplacoidea) from shallow waters in Japan. Studies on Eumalacostraca: a Homage to Masatsune Takeda. Crustaceana Monographs, 17, 137 - 154." type="journal article" year="2012">
Komai
<emphasis id="4BB69637FFBDFFE5FBC2FA8BFBBEFAD6" box="[1087,1139,1369,1391]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
2012
</bibRefCitation>
: 144
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="31D819AEFFBDFFE5FF6AFA4DFE3BFA44" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="797D4A25FFBDFFE5FF6AFA4DFE10FA6E" blockId="30.[151,1045,1436,1677]" pageId="30" pageNumber="31">
<emphasis id="4BB69637FFBDFFE5FF6AFA4DFEE6FA0C" bold="true" box="[151,299,1436,1461]" pageId="30" pageNumber="31">Type species</emphasis>
.
<taxonomicName id="BEC231A6FFBDFFE5FEC4FA4CFCF0FA0D" authority="Alcock &amp; Anderson, 1899" authorityName="Alcock &amp; Anderson" authorityYear="1899" box="[313,829,1436,1461]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="rubida">
<emphasis id="4BB69637FFBDFFE5FEC4FA4CFDC2FA0C" box="[313,527,1436,1461]" italics="true" pageId="30" pageNumber="31">Camatopsis rubida</emphasis>
<bibRefCitation id="1D5337D4FFBDFFE5FDEBFA4DFCF0FA0D" author="Alcock" box="[534,829,1436,1461]" pageId="30" pageNumber="31" refString="Alcock, A. &amp; Anderson, A. R. S. (1899) An account of the deep-sea Crustacea dredged during the surveying-season of 1987 - 98. Natural history notes from H. M. Royal Indian Marine Survey Ship ' Investigator, ' Commander T. H. Heming, R. N., commanding, Series III, No. 2. Annals and Magazine of Natural History, ser. 7 (3), 1 - 27, 278 - 292." type="journal article" year="1899">Alcock &amp; Anderson, 1899</bibRefCitation>
</taxonomicName>
(gender feminine).
<emphasis id="4BB69637FFBDFFE5FF3AFA6EFE18FA61" bold="true" box="[199,469,1471,1496]" pageId="30" pageNumber="31">Other species included</emphasis>
:
</paragraph>
<paragraph id="797D4A25FFBDFFE5FF3AFA34FE3BFA44" blockId="30.[151,1045,1436,1677]" box="[199,502,1508,1533]" pageId="30" pageNumber="31">
<taxonomicName id="BEC231A6FFBDFFE5FF3AFA34FE7CFA45" box="[199,433,1508,1532]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="africana" status="sp. nov.">
<emphasis id="4BB69637FFBDFFE5FF3AFA34FE7CFA45" box="[199,433,1508,1532]" italics="true" pageId="30" pageNumber="31">Camatopsis africana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50852B4CFFBDFFE5FE45FA35FE3BFA44" box="[440,502,1508,1533]" pageId="30" pageNumber="31" rank="species">
<emphasis id="4BB69637FFBDFFE5FE45FA35FE3BFA44" bold="true" box="[440,502,1508,1533]" pageId="30" pageNumber="31">n. sp.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="31D819AEFFBDFFE5FF3AF9D9FE13F934" pageId="30" pageNumber="31" type="description">
<paragraph id="797D4A25FFBDFFE5FF3AF9D9FDD9F999" blockId="30.[151,1045,1436,1677]" box="[199,532,1543,1568]" pageId="30" pageNumber="31">
<taxonomicName id="BEC231A6FFBDFFE5FF3AF9D9FE02F999" box="[199,463,1543,1568]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="leptomerus" status="sp. nov.">
<emphasis id="4BB69637FFBDFFE5FF3AF9D9FE02F999" box="[199,463,1543,1568]" italics="true" pageId="30" pageNumber="31">Camatopsis leptomerus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50852B4CFFBDFFE5FE28F9D6FDD9F999" box="[469,532,1543,1568]" pageId="30" pageNumber="31" rank="species">
<emphasis id="4BB69637FFBDFFE5FE28F9D6FDD9F999" bold="true" box="[469,532,1543,1568]" pageId="30" pageNumber="31">n. sp.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="797D4A25FFBDFFE5FF3AF9FCFE16F9FC" blockId="30.[151,1045,1436,1677]" box="[199,475,1580,1605]" pageId="30" pageNumber="31">
<taxonomicName id="BEC231A6FFBDFFE5FF3AF9FCFE58F9FD" box="[199,405,1581,1604]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="minor" status="sp. nov.">
<emphasis id="4BB69637FFBDFFE5FF3AF9FCFE58F9FD" box="[199,405,1581,1604]" italics="true" pageId="30" pageNumber="31">Camatopsis minor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50852B4CFFBDFFE5FE61F9FDFE16F9FC" box="[412,475,1580,1605]" pageId="30" pageNumber="31" rank="species">
<emphasis id="4BB69637FFBDFFE5FE61F9FDFE16F9FC" bold="true" box="[412,475,1580,1605]" pageId="30" pageNumber="31">n. sp.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="797D4A25FFBDFFE5FF3AF981FE1FF9D1" blockId="30.[151,1045,1436,1677]" box="[199,466,1615,1640]" pageId="30" pageNumber="31">
<taxonomicName id="BEC231A6FFBDFFE5FF3AF981FE41F9D1" box="[199,396,1615,1640]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="thula" status="sp. nov.">
<emphasis id="4BB69637FFBDFFE5FF3AF981FE41F9D1" box="[199,396,1615,1640]" italics="true" pageId="30" pageNumber="31">Camatopsis thula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50852B4CFFBDFFE5FE6EF99EFE1FF9D1" box="[403,466,1615,1640]" pageId="30" pageNumber="31" rank="species">
<emphasis id="4BB69637FFBDFFE5FE6EF99EFE1FF9D1" bold="true" box="[403,466,1615,1640]" pageId="30" pageNumber="31">n. sp.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="797D4A25FFBDFFE5FF3AF9A4FE13F934" blockId="30.[151,1045,1436,1677]" box="[199,478,1652,1677]" pageId="30" pageNumber="31">
<taxonomicName id="BEC231A6FFBDFFE5FF3AF9A4FE55F934" box="[199,408,1652,1677]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="valida" status="sp. nov.">
<emphasis id="4BB69637FFBDFFE5FF3AF9A4FE55F934" box="[199,408,1652,1677]" italics="true" pageId="30" pageNumber="31">Camatopsis valida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50852B4CFFBDFFE5FE62F9A5FE13F934" box="[415,478,1652,1677]" pageId="30" pageNumber="31" rank="species">
<emphasis id="4BB69637FFBDFFE5FE62F9A5FE13F934" bold="true" box="[415,478,1652,1677]" pageId="30" pageNumber="31">n. sp.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="31D819AEFFBDFFE4FF3AF96DFA41FE4C" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="diagnosis">
<paragraph id="797D4A25FFBDFFE4FF3AF96DFA41FE4C" blockId="30.[151,1436,1724,2037]" lastBlockId="31.[151,1437,151,1832]" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
<emphasis id="4BB69637FFBDFFE5FF3AF96DFEF7F96C" bold="true" box="[199,314,1724,1749]" pageId="30" pageNumber="31">Diagnosis</emphasis>
. Carapace subtrapezoidal, high; front bilobed, with shallow or deep median cleft; anterolateral margins arcuate, minutely granular, without distinct lobes or teeth. Epistome slightly depressed; semicircular median lobe with deep median fissure, semicircular lateral margins without visible fissures. Eye peduncle filling orbit, short, slightly mobile; cornea reduced, with reduced pigmentation. Third maxillipeds almost entirely fill buccal cavern when closed; merus subcircular, anteroexternal angle produced to different degrees; ischium subquadrate or slightly elongated, about same length as merus. Chelipeds subequal in length, slightly dissimilar in females, heteromorphic in males; cutting margins blade-like, lined with short setae, at least distal part of pollex with sharp spines that overlap dactylus when closed, most proximal teeth on cutting margins of major chela of males enlarged. Inner margin of cheliped carpus smooth or with tooth (
<taxonomicName id="BEC231A6FFBDFFE5FC1CF80CFBA8F84D" box="[993,1125,2012,2036]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="4BB69637FFBDFFE5FC1CF80CFBA8F84D" box="[993,1125,2012,2036]" italics="true" pageId="30" pageNumber="31">C. africana</emphasis>
</taxonomicName>
males). Ventral surface of cheliped merus without teeth or conspicuous tubercles. Meri of ambulatory legs with microscopic granules, unarmed; long setae along margins of propodi, dactyli. P5 dactylus upcurved. Fused thoracic sternites 1, 2 triangular, short; fused thoracic sternites 3, 4 relatively broad. Male pleon lateral margins of somite 6 convex, of fused somites 3–5 nearly straight; telson proportionally short. Sterno-pleonal cavity of male deep, press-button for pleonal holding as small, short tubercle just posterior to thoracic sternal suture 4/5 at edge of sterno-pleonal cavity. Male thoracic sternite 8 short, quadrate; “supplementary plate” short, longer at rounded outer margin; structures tightly appressed with most of penis concealed. G1 stout, distal segment slightly curved, distal segment curved inwards or upwards, with short to long spinules. G2 short, about 3/4 G1 length, straight, slender, with long basal segment, conspicuously short distal segment. Female pleon with lateral margins of somites strongly convex; telson proportionally short. Sterno-pleonal cavity of female moderately deep, vulvae close together or widely separated.
</paragraph>
</subSubSection>
<subSubSection id="31D819AEFFBCFFE4FF3AFE2EFB1FF8BC" pageId="31" pageNumber="32" type="discussion">
<paragraph id="797D4A25FFBCFFE4FF3AFE2EFBF2FD3C" blockId="31.[151,1437,151,1832]" pageId="31" pageNumber="32">
<emphasis id="4BB69637FFBCFFE4FF3AFE2EFEF9FDA1" bold="true" box="[199,308,511,536]" pageId="31" pageNumber="32">Remarks</emphasis>
. Long regarded as monotypic, “
<taxonomicName id="BEC231A6FFBCFFE4FD60FDD1FCB9FDA1" box="[669,884,511,536]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="rubida">
<emphasis id="4BB69637FFBCFFE4FD60FDD1FCB9FDA1" box="[669,884,511,536]" italics="true" pageId="31" pageNumber="32">Camatopsis rubida</emphasis>
</taxonomicName>
,” which
<bibRefCitation id="1D5337D4FFBCFFE4FC24FE2EFAC0FDA1" author="Alcock" box="[985,1293,511,536]" pageId="31" pageNumber="32" refString="Alcock, A. &amp; Anderson, A. R. S. (1899) An account of the deep-sea Crustacea dredged during the surveying-season of 1987 - 98. Natural history notes from H. M. Royal Indian Marine Survey Ship ' Investigator, ' Commander T. H. Heming, R. N., commanding, Series III, No. 2. Annals and Magazine of Natural History, ser. 7 (3), 1 - 27, 278 - 292." type="journal article" year="1899">Alcock &amp; Anderson (1899)</bibRefCitation>
described in
<taxonomicName id="BEC231A6FFBCFFE4FF6AFDF5FED7FD84" box="[151,282,548,573]" class="Malacostraca" family="Pilumnidae" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subFamily" subFamily="Rhizopinae">Rhizopinae</taxonomicName>
, has been reported from many locations in the Indo-West Pacific region. Comparisons of a large series of specimens from the range shows that there are consistent differences in the proportions of the ambulatory legs, form of the male pleon and G1 structures, and six species can in fact be recognised.
</paragraph>
<paragraph id="797D4A25FFBCFFE4FF3AFD5EFE06FC54" blockId="31.[151,1437,151,1832]" pageId="31" pageNumber="32">
The genus is closest to
<taxonomicName id="BEC231A6FFBCFFE4FE37FD40FD82FD11" box="[458,591,657,680]" class="Malacostraca" family="Chasmocarcinidae" genus="Microtopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="4BB69637FFBCFFE4FE37FD40FD82FD11" box="[458,591,657,680]" italics="true" pageId="31" pageNumber="32">Microtopsis</emphasis>
</taxonomicName>
in the form of the carapace and chelipeds.
<bibRefCitation id="1D5337D4FFBCFFE4FBD3FD5EFAC7FD11" box="[1070,1290,655,680]" pageId="31" pageNumber="32" refString="Komai, T., Ng, P. K. L. &amp; Yamada, Y. (2012) A new genus and new species of chasmocarcinid crab (Decapoda, Brachyura, Goneplacoidea) from shallow waters in Japan. Studies on Eumalacostraca: a Homage to Masatsune Takeda. Crustaceana Monographs, 17, 137 - 154." type="journal article">
Komai
<emphasis id="4BB69637FFBCFFE4FB7CFD40FB7FFD11" box="[1153,1202,655,680]" italics="true" pageId="31" pageNumber="32">et al</emphasis>
. (2012)
</bibRefCitation>
listed a suite of characters that separate
<taxonomicName id="BEC231A6FFBCFFE4FE37FD64FD83FD75" box="[458,590,693,716]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="4BB69637FFBCFFE4FE37FD64FD83FD75" box="[458,590,693,716]" italics="true" pageId="31" pageNumber="32">Camatopsis</emphasis>
</taxonomicName>
from
<taxonomicName id="BEC231A6FFBCFFE4FD6AFD64FCD1FD75" box="[663,796,693,716]" class="Malacostraca" family="Chasmocarcinidae" genus="Microtopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="4BB69637FFBCFFE4FD6AFD64FCD1FD75" box="[663,796,693,716]" italics="true" pageId="31" pageNumber="32">Microtopsis</emphasis>
</taxonomicName>
. The third maxilliped merus of
<taxonomicName id="BEC231A6FFBCFFE4FB73FD64FADEFD75" box="[1166,1299,693,716]" class="Malacostraca" family="Chasmocarcinidae" genus="Microtopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="4BB69637FFBCFFE4FB73FD64FADEFD75" box="[1166,1299,693,716]" italics="true" pageId="31" pageNumber="32">Microtopsis</emphasis>
</taxonomicName>
is subovate without any anteroexternal expansion (
<figureCitation id="E1F956A0FFBCFFE4FDB0FD06FD53FD49" box="[589,670,727,752]" captionStart="FIGURE 80" captionStartId="153.[151,250,1480,1502]" captionTargetBox="[166,1396,204,1436]" captionTargetId="figure@153.[151,1436,193,1459]" captionTargetPageId="153" captionText="FIGURE 80. Third maxilliped and male pleon of species of Microtopsis. A, B, M. takedai, Japan: holotype male 2.4 × 3.1 mm (after Komai et al. 2012: figs. 3 A, 2 H, respectively); C, D, F, M. teschi n. sp., Indonesia: holotype male 1.5 × 2.1 mm (NNM- ZMA); E, M. teschi n. sp., Indonesia: paratype 1.5 × 1.9 mm (NNM-ZMA). A, left third maxilliped; B, F, male pleon; C, right third maxilliped (denuded, exopod not drawn); D, left third maxilliped (denuded, exopod not drawn); E, right third maxilliped (denuded). Scales = 0.5 mm." httpUri="https://zenodo.org/record/272726/files/figure.png" pageId="31" pageNumber="32">Fig. 80</figureCitation>
A) (merus more quadrate with the anteroexternal angle auriculiform in various degrees in
<taxonomicName id="BEC231A6FFBCFFE4FE79FD2CFDC5FCAD" box="[388,520,765,788]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="4BB69637FFBCFFE4FE79FD2CFDC5FCAD" box="[388,520,765,788]" italics="true" pageId="31" pageNumber="32">Camatopsis</emphasis>
</taxonomicName>
, e.g.,
<figureCitation id="E1F956A0FFBCFFE4FDB6FD2DFD56FCAD" box="[587,667,764,789]" captionStart="FIGURE 30" captionStartId="103.[151,250,1353,1375]" captionTargetBox="[151,1436,193,1332]" captionTargetId="figure@103.[151,1436,193,1332]" captionTargetPageId="103" captionText="FIGURE 30. Third maxilliped of species of Camatopsis and Chasmocarcinops. A, Camatopsis rubida, Australia: male 8.3 × 8.5 mm (NMV J 54591); B, Camatopsis africana n. sp., Madagascar: holotype male 5.7 × 6.9 mm (MNHN-IU- 2010 - 1574); C, Camatopsis leptomerus n. sp., Philippines: holotype male 10.5 × 11.3 mm (NMCR); D, Camatopsis thula n. sp., Taiwan: holotype male 8.9 × 9.3 mm (ZRC 1999.0662); E, Chasmocarcinops gelasimoides, Andaman Sea: male 8.8 × 9.5 mm (PMBC 16849); F, Chasmocarcinops gelasimoides, Singapore: male 9.5 × 10.3 mm (ZRC 2000.1341)." httpUri="https://zenodo.org/record/272676/files/figure.png" pageId="31" pageNumber="32">Fig. 30</figureCitation>
A); the cutting margins of the fingers of the
<taxonomicName id="BEC231A6FFBCFFE4FB7FFD2DFB08FCAC" box="[1154,1221,764,789]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="minor">minor</taxonomicName>
chelae have comblike setae and no sharp teeth (
<figureCitation id="E1F956A0FFBCFFE4FE1AFCCEFDF7FC81" box="[487,570,799,824]" captionStart="FIGURE 48" captionStartId="121.[151,250,1288,1310]" captionTargetBox="[151,1436,217,1267]" captionTargetId="figure@121.[151,1436,193,1267]" captionTargetPageId="121" captionText="FIGURE 48. Chelae of species of Megaesthesius, Alainthesius n. gen. and Microtopsis. A, B, Microtopsis takedai, Japan: holotype male 2.4 × 3.1 mm (after Komai et al. 2012: fig. 3 B, D); C, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); D, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); E, Megaesthesius yokoyai, Japan: male (after Takeda &amp; Miyake 1969: fig. 4 d); F, Alainthesius signatus n. sp., Madagascar: holotype male 3.5 × 3.8 mm (after Crosnier 1975: fig. 1 d); G, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN- IU- 2013 - 9073); H, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012)." httpUri="https://zenodo.org/record/272694/files/figure.png" pageId="31" pageNumber="32">Fig. 48</figureCitation>
B) (with one or two sharp teeth in
<taxonomicName id="BEC231A6FFBCFFE4FC43FCF1FB8FFC8E" box="[958,1090,800,823]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="4BB69637FFBCFFE4FC43FCF1FB8FFC8E" box="[958,1090,800,823]" italics="true" pageId="31" pageNumber="32">Camatopsis</emphasis>
</taxonomicName>
, e.g.,
<figureCitation id="E1F956A0FFBCFFE4FB75FCCEFB16FC81" box="[1160,1243,799,824]" captionStart="FIGURE 39" captionStartId="112.[151,250,1909,1931]" captionTargetBox="[151,1436,409,1887]" captionTargetId="figure@112.[151,1436,409,1887]" captionTargetPageId="112" captionText="FIGURE 39. Chelae of species of Camatopsis. A, B, C. leptomerus n. sp., Philippines: holotype male 10.5 × 11.3 mm (NMCR); C, D, C. leptomerus n. sp., Philippines: paratype female 3.4 × 4.0 mm (ZRC 2015.159); E, F, C. thula n. sp., Taiwan: holotype male 8.9 × 9.3 mm (ZRC 1999.0662); G, H, C. minor n. sp., Indonesia: holotype male 3.6 × 4.3 mm (NNM-ZMA); I, J, C. valida n. sp., Fiji: holotype male 3.6 × 4.9 mm (MNHN-IU- 2013 - 9067)." httpUri="https://zenodo.org/record/272685/files/figure.png" pageId="31" pageNumber="32">Fig. 39</figureCitation>
B); male thoracic sternites 7 and 8 are proportionately broader (
<figureCitation id="E1F956A0FFBCFFE4FD39FC95FCD6FCE4" box="[708,795,836,861]" captionStart="FIGURE 62" captionStartId="135.[151,250,926,948]" captionTargetBox="[151,1436,303,904]" captionTargetId="figure@135.[151,1436,303,904]" captionTargetPageId="135" captionText="FIGURE 62. Thoracic sternites 7 and 8 and male pleon of species of Microtopsis and Megaesthesius. A, Microtopsis takedai, Japan: paratype male, carapace damaged (ZRC 2012.0146); B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); C, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); D, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073)." httpUri="https://zenodo.org/record/272708/files/figure.png" pageId="31" pageNumber="32">Fig. 62</figureCitation>
A) than in
<taxonomicName id="BEC231A6FFBCFFE4FC5BFC94FBE7FCE5" box="[934,1066,837,860]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="4BB69637FFBCFFE4FC5BFC94FBE7FCE5" box="[934,1066,837,860]" italics="true" pageId="31" pageNumber="32">Camatopsis</emphasis>
</taxonomicName>
(
<figureCitation id="E1F956A0FFBCFFE4FBBDFC95FB54FCE4" box="[1088,1177,836,861]" captionStart="FIGURE 58" captionStartId="131.[151,250,1830,1852]" captionTargetBox="[164,1408,930,1807]" captionTargetId="figure@131.[164,1408,930,1807]" captionTargetPageId="131" captionText="FIGURE 58. Thoracic sternites 7 and 8 and male pleon of species of Camatopsis and Chasmocarcinops. A, Camatopsis rubida, Australia: male 8.3 × 8.5 mm (NMV J 54591); B, Camatopsis africana n. sp., Madagascar: holotype male 5.7 × 6.9 mm (MNHN-IU- 2010 - 1574); C, Camatopsis leptomerus n. sp., Philippines: holotype male 10.5 × 11.3 mm (NMCR); D, Camatopsis thula n. sp., Taiwan: holotype male 8.9 × 9.3 mm (ZRC 1999.0662); E, Camatopsis valida n. sp., Fiji: paratype female 3.5 × 4.3 mm (MNHN-IU- 2013 - 9467); F, Chasmocarcinops gelasimoides, Singapore: male 9.5 × 10.3 mm (ZRC 2000.1341)." httpUri="https://zenodo.org/record/272704/files/figure.png" pageId="31" pageNumber="32">Fig. 58</figureCitation>
A); the male pleon is proportionally shorter (
<figureCitation id="E1F956A0FFBCFFE4FE66FCB6FE20FC39" box="[411,493,871,896]" captionStart="FIGURE 80" captionStartId="153.[151,250,1480,1502]" captionTargetBox="[166,1396,204,1436]" captionTargetId="figure@153.[151,1436,193,1459]" captionTargetPageId="153" captionText="FIGURE 80. Third maxilliped and male pleon of species of Microtopsis. A, B, M. takedai, Japan: holotype male 2.4 × 3.1 mm (after Komai et al. 2012: figs. 3 A, 2 H, respectively); C, D, F, M. teschi n. sp., Indonesia: holotype male 1.5 × 2.1 mm (NNM- ZMA); E, M. teschi n. sp., Indonesia: paratype 1.5 × 1.9 mm (NNM-ZMA). A, left third maxilliped; B, F, male pleon; C, right third maxilliped (denuded, exopod not drawn); D, left third maxilliped (denuded, exopod not drawn); E, right third maxilliped (denuded). Scales = 0.5 mm." httpUri="https://zenodo.org/record/272726/files/figure.png" pageId="31" pageNumber="32">Fig. 80</figureCitation>
B) than in
<taxonomicName id="BEC231A6FFBCFFE4FD9EFCB9FD2AFCC6" box="[611,743,872,895]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="4BB69637FFBCFFE4FD9EFCB9FD2AFCC6" box="[611,743,872,895]" italics="true" pageId="31" pageNumber="32">Camatopsis</emphasis>
</taxonomicName>
(e.g., 51A); the G1 is short, stout, distinctly twisted medially, with the distal part broad and shortened (
<figureCitation id="E1F956A0FFBCFFE4FD81FC5DFD15FC1D" box="[636,728,908,933]" captionStart="FIGURE 81" captionStartId="154.[151,250,1949,1971]" captionTargetBox="[218,1333,202,1887]" captionTargetId="figure@154.[207,1366,195,1912]" captionTargetPageId="154" captionText="FIGURE 81. Gonopods of species of Microtopsis. A ‒ F, M. takedai, Japan: holotype male 2.4 × 3.1 mm (after Komai et al. 2012: fig. 4); G ‒ N, M. teschi n. sp., Indonesia: holotype male 1.5 × 2.1 mm (NNM-ZMA). A ‒ C, G ‒ I, left G 1; D, E, J ‒ L, distal part of left G 1; F, M, left G 2; N, distal part of left G 2. Scales: G ‒ I, M = 0.1 mm; J ‒ L = 0.05 mm; N = 0.01 mm." httpUri="https://zenodo.org/record/272727/files/figure.png" pageId="31" pageNumber="32">Figs. 81</figureCitation>
A‒E) (slender and elongated, with the distal part tapering in
<taxonomicName id="BEC231A6FFBCFFE4FF6AFC61FED6FC7E" box="[151,283,944,967]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="4BB69637FFBCFFE4FF6AFC61FED6FC7E" box="[151,283,944,967]" italics="true" pageId="31" pageNumber="32">Camatopsis</emphasis>
</taxonomicName>
; e.g.,
<figureCitation id="E1F956A0FFBCFFE4FE9CFC7EFE7CFC71" box="[353,433,943,968]" captionStart="FIGURE 67" captionStartId="140.[151,250,1502,1524]" captionTargetBox="[168,1426,200,1445]" captionTargetId="figure@140.[151,1436,195,1480]" captionTargetPageId="140" captionText="FIGURE 67. Gonopods of species of Camatopsis. A ‒ C, C. rubida, Andaman Sea: male 4.2 × 4.5 mm (ZMUC CRU 20539); D ‒ F, C. rubida, Australia: male 8.3 × 8.5 mm (NMV J 54591); G ‒ I, C. africana n. sp., Mozambique Channel: holotype male 6.8 × 7.7 mm (MNHN-IU- 2010 - 1500); J ‒ L, C. leptomerus n. sp., Philippines: holotype male 10.5 × 11.3 mm (NMCR); M ‒ O, C. thula n. sp., Taiwan: holotype male 8.9 × 9.3 mm (ZRC 1999.0662). A, B, D, E, G, H, J, K, M, N, ventral view of left G 1; F, dorsal view of left G 1; C, I, L, O, left G 2. Scales: A, C, D, G, I, J, L, M, O = 0.5 mm; B, E, F, H, K, N = 0.1 mm." httpUri="https://zenodo.org/record/272713/files/figure.png" pageId="31" pageNumber="32">Fig. 67</figureCitation>
A, B); and the G2 is as long as the G1 (Fig. e.g., 81F) (G2 shorter, about 3/4 G1 length in
<taxonomicName id="BEC231A6FFBCFFE4FF6AFC04FED6FC55" box="[151,283,981,1004]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="4BB69637FFBCFFE4FF6AFC04FED6FC55" box="[151,283,981,1004]" italics="true" pageId="31" pageNumber="32">Camatopsis</emphasis>
</taxonomicName>
, e.g.
<figureCitation id="E1F956A0FFBCFFE4FEA5FC05FE66FC54" box="[344,427,980,1005]" captionStart="FIGURE 67" captionStartId="140.[151,250,1502,1524]" captionTargetBox="[168,1426,200,1445]" captionTargetId="figure@140.[151,1436,195,1480]" captionTargetPageId="140" captionText="FIGURE 67. Gonopods of species of Camatopsis. A ‒ C, C. rubida, Andaman Sea: male 4.2 × 4.5 mm (ZMUC CRU 20539); D ‒ F, C. rubida, Australia: male 8.3 × 8.5 mm (NMV J 54591); G ‒ I, C. africana n. sp., Mozambique Channel: holotype male 6.8 × 7.7 mm (MNHN-IU- 2010 - 1500); J ‒ L, C. leptomerus n. sp., Philippines: holotype male 10.5 × 11.3 mm (NMCR); M ‒ O, C. thula n. sp., Taiwan: holotype male 8.9 × 9.3 mm (ZRC 1999.0662). A, B, D, E, G, H, J, K, M, N, ventral view of left G 1; F, dorsal view of left G 1; C, I, L, O, left G 2. Scales: A, C, D, G, I, J, L, M, O = 0.5 mm; B, E, F, H, K, N = 0.1 mm." httpUri="https://zenodo.org/record/272713/files/figure.png" pageId="31" pageNumber="32">Fig. 67</figureCitation>
C).
</paragraph>
<paragraph id="797D4A25FFBCFFE4FF3AFC26FB1FF8BC" blockId="31.[151,1437,151,1832]" pageId="31" pageNumber="32">
Prior to this revision, adults of
<taxonomicName id="BEC231A6FFBCFFE4FDB5FC29FD01FBB6" box="[584,716,1016,1039]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="4BB69637FFBCFFE4FDB5FC29FD01FBB6" box="[584,716,1016,1039]" italics="true" pageId="31" pageNumber="32">Camatopsis</emphasis>
</taxonomicName>
species were believed to be large, with carapace widths of approximately
<quantity id="BE3AE7C0FFBCFFE4FEC2FBCDFE4DFB8D" box="[319,384,1052,1076]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="31" pageNumber="32" unit="mm" value="8.0">8 mm</quantity>
or more; with
<taxonomicName id="BEC231A6FFBCFFE4FDD8FBCCFD67FB8D" box="[549,682,1053,1076]" class="Malacostraca" family="Chasmocarcinidae" genus="Microtopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="4BB69637FFBCFFE4FDD8FBCCFD67FB8D" box="[549,682,1053,1076]" italics="true" pageId="31" pageNumber="32">Microtopsis</emphasis>
</taxonomicName>
maturing at carapace widths of only
<quantity id="BE3AE7C0FFBCFFE4FBBAFBCDFB45FB8D" box="[1095,1160,1052,1076]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="31" pageNumber="32" unit="mm" value="3.0">3 mm</quantity>
(see
<bibRefCitation id="1D5337D4FFBCFFE4FB3FFBCDFA5CFB8D" author="Komai" box="[1218,1425,1052,1077]" pageId="31" pageNumber="32" refString="Komai, T., Ng, P. K. L. &amp; Yamada, Y. (2012) A new genus and new species of chasmocarcinid crab (Decapoda, Brachyura, Goneplacoidea) from shallow waters in Japan. Studies on Eumalacostraca: a Homage to Masatsune Takeda. Crustaceana Monographs, 17, 137 - 154." type="journal article" year="2012">
Komai
<emphasis id="4BB69637FFBCFFE4FAE8FBCCFA8BFB8C" box="[1301,1350,1052,1077]" italics="true" pageId="31" pageNumber="32">et al</emphasis>
. 2012
</bibRefCitation>
). The discovery of
<taxonomicName id="BEC231A6FFBCFFE4FE98FB91FE00FBE1" box="[357,461,1088,1112]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="minor" status="sp. nov.">
<emphasis id="4BB69637FFBCFFE4FE98FB91FE00FBE1" box="[357,461,1088,1112]" italics="true" pageId="31" pageNumber="32">C. minor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50852B4CFFBCFFE4FE2AFBEEFDD4FBE1" box="[471,537,1087,1112]" pageId="31" pageNumber="32" rank="species">
<emphasis id="4BB69637FFBCFFE4FE2AFBEEFDD4FBE1" bold="true" box="[471,537,1087,1112]" pageId="31" pageNumber="32">n. sp.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="BEC231A6FFBCFFE4FDABFB91FD0CFBE1" box="[598,705,1087,1112]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="valida" status="sp. nov.">
<emphasis id="4BB69637FFBCFFE4FDABFB91FD0CFBE1" box="[598,705,1087,1112]" italics="true" pageId="31" pageNumber="32">C. valida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50852B4CFFBCFFE4FD36FBEEFCC0FBE1" box="[715,781,1087,1112]" pageId="31" pageNumber="32" rank="species">
<emphasis id="4BB69637FFBCFFE4FD36FBEEFCC0FBE1" bold="true" box="[715,781,1087,1112]" pageId="31" pageNumber="32">n. sp.</emphasis>
</taxonomicNameLabel>
as well as a good series of specimens of
<taxonomicName id="BEC231A6FFBCFFE4FB06FB91FA51FBE1" box="[1275,1436,1087,1112]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="leptomerus">
<emphasis id="4BB69637FFBCFFE4FB06FB91FA51FBE1" box="[1275,1436,1087,1112]" italics="true" pageId="31" pageNumber="32">C. leptomerus</emphasis>
</taxonomicName>
changes our perspective on these differences. The structure of the male pleon and G1 remains diagnostic, and easily distinguishes
<taxonomicName id="BEC231A6FFBCFFE4FE80FB58FDCFFB19" box="[381,514,1161,1184]" class="Malacostraca" family="Chasmocarcinidae" genus="Microtopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="4BB69637FFBCFFE4FE80FB58FDCFFB19" box="[381,514,1161,1184]" italics="true" pageId="31" pageNumber="32">Microtopsis</emphasis>
</taxonomicName>
from
<taxonomicName id="BEC231A6FFBCFFE4FDB7FB59FD03FB26" box="[586,718,1160,1183]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="4BB69637FFBCFFE4FDB7FB59FD03FB26" box="[586,718,1160,1183]" italics="true" pageId="31" pageNumber="32">Camatopsis</emphasis>
</taxonomicName>
. The structures of the third maxilliped and chela, on the other hand, appear to be size-dependent. Specimens of
<taxonomicName id="BEC231A6FFBCFFE4FD2EFB7CFCBEFB7C" box="[723,883,1196,1221]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="leptomerus">
<emphasis id="4BB69637FFBCFFE4FD2EFB7CFCBEFB7C" box="[723,883,1196,1221]" italics="true" pageId="31" pageNumber="32">C. leptomerus</emphasis>
</taxonomicName>
up to 7.0 mm in carapace width (e.g., CP2383, ZRC 2015.139) are typical, with the anteroexternal angle of the third maxilliped merus with auriculiform anteroexternal angle and the
<taxonomicName id="BEC231A6FFBCFFE4FE1DFB25FDEEFAB4" box="[480,547,1268,1293]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="minor">minor</taxonomicName>
chela possessing the two vertical teeth on the cutting margin of the propodus. Smaller specimens, especially those below 6.0 mm in carapace width have the third maxilliped merus varying from subovate to round but without an auriculiform anteroexternal angle (
<figureCitation id="E1F956A0FFBCFFE4FC51FAEDFC32FAEC" box="[940,1023,1340,1365]" captionStart="FIGURE 68" captionStartId="141.[151,250,1809,1831]" captionTargetBox="[164,1418,212,1765]" captionTargetId="figure@141.[151,1436,193,1788]" captionTargetPageId="141" captionText="FIGURE 68. Gonopods and third maxilliped of Camatopsis leptomerus. A, Philippines: paratype male 4.7 × 5.9 mm (ZRC 2015.120); B ‒ D, Indonesia: male 2.5 × 3.9 mm (NNM-ZMA 241605 e), E, Philippines: paratype female 3.4 × 4.0 mm (ZRC 2015.159); F, Philippines: paratype male 3.1 × 3.4 mm (ZRC 2015.120); G, Philippines: paratype female 4.8 × 5.8 mm (ZRC 2015.154); H, Philippines: male 4.7 × 5.9 mm (ZRC 2015.120). A, B, ventral view of left G 1; D, dorsal view of left G 1; C, ventral view of distal part of left G 1; E, right third maxilliped (denuded); F ‒ H, left third maxilliped (denuded). Scales: A, B, D = 0.25 mm; E ‒ H = 0.5 mm; C = 0.01 mm." httpUri="https://zenodo.org/record/272714/files/figure.png" pageId="31" pageNumber="32">Fig. 68</figureCitation>
E‒H), and the cutting margin of the
<taxonomicName id="BEC231A6FFBCFFE4FF6AFA8EFF17FAC1" box="[151,218,1375,1400]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="minor">minor</taxonomicName>
chela propodus is entire, without any vertical teeth. Surprisingly, the G1 structure even in the very small males of
<taxonomicName id="BEC231A6FFBCFFE4FF00FA54FE57FA24" box="[253,410,1412,1437]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="leptomerus">
<emphasis id="4BB69637FFBCFFE4FF00FA54FE57FA24" box="[253,410,1412,1437]" italics="true" pageId="31" pageNumber="32">C. leptomerus</emphasis>
</taxonomicName>
(e.g., two males 3.1 ×
<quantity id="BE3AE7C0FFBCFFE4FD64FA55FD3CFA25" box="[665,753,1412,1436]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="31" pageNumber="32" unit="mm" value="3.4">3.4 mm</quantity>
, 4.7 ×
<quantity id="BE3AE7C0FFBCFFE4FCC0FA55FC5BFA25" box="[829,918,1412,1437]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="31" pageNumber="32" unit="mm" value="5.9">5.9 mm</quantity>
; ZRC 2015.120) is distinct, resembling that of the adult (
<figureCitation id="E1F956A0FFBCFFE4FEF5FA76FE97FA79" box="[264,346,1447,1472]" captionStart="FIGURE 68" captionStartId="141.[151,250,1809,1831]" captionTargetBox="[164,1418,212,1765]" captionTargetId="figure@141.[151,1436,193,1788]" captionTargetPageId="141" captionText="FIGURE 68. Gonopods and third maxilliped of Camatopsis leptomerus. A, Philippines: paratype male 4.7 × 5.9 mm (ZRC 2015.120); B ‒ D, Indonesia: male 2.5 × 3.9 mm (NNM-ZMA 241605 e), E, Philippines: paratype female 3.4 × 4.0 mm (ZRC 2015.159); F, Philippines: paratype male 3.1 × 3.4 mm (ZRC 2015.120); G, Philippines: paratype female 4.8 × 5.8 mm (ZRC 2015.154); H, Philippines: male 4.7 × 5.9 mm (ZRC 2015.120). A, B, ventral view of left G 1; D, dorsal view of left G 1; C, ventral view of distal part of left G 1; E, right third maxilliped (denuded); F ‒ H, left third maxilliped (denuded). Scales: A, B, D = 0.25 mm; E ‒ H = 0.5 mm; C = 0.01 mm." httpUri="https://zenodo.org/record/272714/files/figure.png" pageId="31" pageNumber="32">Fig. 68</figureCitation>
A‒D). Although the G1s of these small males is relatively more slender, the distal part is twisted with the tip distinctly turned upwards (when detached and with the ventral surface facing the observer) (
<figureCitation id="E1F956A0FFBCFFE4FA99FA1DFF79F9B1" captionStart="FIGURE 68" captionStartId="141.[151,250,1809,1831]" captionTargetBox="[164,1418,212,1765]" captionTargetId="figure@141.[151,1436,193,1788]" captionTargetPageId="141" captionText="FIGURE 68. Gonopods and third maxilliped of Camatopsis leptomerus. A, Philippines: paratype male 4.7 × 5.9 mm (ZRC 2015.120); B ‒ D, Indonesia: male 2.5 × 3.9 mm (NNM-ZMA 241605 e), E, Philippines: paratype female 3.4 × 4.0 mm (ZRC 2015.159); F, Philippines: paratype male 3.1 × 3.4 mm (ZRC 2015.120); G, Philippines: paratype female 4.8 × 5.8 mm (ZRC 2015.154); H, Philippines: male 4.7 × 5.9 mm (ZRC 2015.120). A, B, ventral view of left G 1; D, dorsal view of left G 1; C, ventral view of distal part of left G 1; E, right third maxilliped (denuded); F ‒ H, left third maxilliped (denuded). Scales: A, B, D = 0.25 mm; E ‒ H = 0.5 mm; C = 0.01 mm." httpUri="https://zenodo.org/record/272714/files/figure.png" pageId="31" pageNumber="32">Figs. 68</figureCitation>
A‒D). Although the distal part of the G1 is gently twisted when the G1 is still attached on the pleon and when it is viewed at an angle, the distal third appears to be curving outwards, as depicted by
<bibRefCitation id="1D5337D4FFBCFFE4FB9EF9C5FA55F994" author="Tesch" box="[1123,1432,1556,1581]" pageId="31" pageNumber="32" refString="Tesch J. J. (1918) Goneplacidae and Pinnotheridae. The Decapoda Brachyura of the Siboga Expedition, 2. Siboga-Expeditie 39 (c 1), livr. 84, 149 - 295, pls. 7 - 18." type="journal article" year="1918" yearSuffix="i">
Tesch (1918: pl. 16 fig.
<date id="0D7C6CE5FFBCFFE4FA8BF9C5FA43F994" box="[1398,1422,1556,1581]" pageId="31" pageNumber="32">3i</date>
)
</bibRefCitation>
.
<taxonomicName id="BEC231A6FFBCFFE4FF6AF9E9FEA8F9E9" box="[151,357,1592,1616]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="minor" status="sp. nov.">
<emphasis id="4BB69637FFBCFFE4FF6AF9E9FEA8F9E9" box="[151,357,1592,1616]" italics="true" pageId="31" pageNumber="32">Camatopsis minor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50852B4CFFBCFFE4FE90F9E6FE61F9E9" box="[365,428,1591,1616]" pageId="31" pageNumber="32" rank="species">
<emphasis id="4BB69637FFBCFFE4FE90F9E6FE61F9E9" bold="true" box="[365,428,1591,1616]" pageId="31" pageNumber="32">n. sp.</emphasis>
</taxonomicNameLabel>
is also a small-size species like
<taxonomicName id="BEC231A6FFBCFFE4FCE4F9E8FC53F9E9" box="[793,926,1593,1616]" class="Malacostraca" family="Chasmocarcinidae" genus="Microtopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="4BB69637FFBCFFE4FCE4F9E8FC53F9E9" box="[793,926,1593,1616]" italics="true" pageId="31" pageNumber="32">Microtopsis</emphasis>
</taxonomicName>
, maturing at carapace widths of about
<quantity id="BE3AE7C0FFBCFFE4FAA9F9E9FA55F9F6" box="[1364,1432,1592,1616]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="31" pageNumber="32" unit="mm" value="4.0">4 mm</quantity>
. The merus of the third maxilliped has a subauriculiform anteroexternal angle but less well developed than adults of other
<taxonomicName id="BEC231A6FFBCFFE4FF2AF951FE96F92E" box="[215,347,1664,1687]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="4BB69637FFBCFFE4FF2AF951FE96F92E" box="[215,347,1664,1687]" italics="true" pageId="31" pageNumber="32">Camatopsis</emphasis>
</taxonomicName>
species. Although the larger male chela is slightly inflated, the cutting margin of the propodus of the smaller chela is entire, without the two vertical teeth. Its G1 structure resembles those of its congeners (
<figureCitation id="E1F956A0FFBCFFE4FA92F975FF79F959" captionStart="FIGURE 69" captionStartId="142.[151,250,1853,1875]" captionTargetBox="[162,1412,212,1817]" captionTargetId="figure@142.[151,1436,193,1831]" captionTargetPageId="142" captionText="FIGURE 69. Thoracic sternum, third maxilliped, pleon and gonopods of Camatopsis species. A, B, D ‒ F, C. minor n. sp., Indonesia: holotype male 3.6 × 4.3 mm (NNM-ZMA); C, C. minor n. sp., Indonesia: paratype male 3.4 × 4.0 mm (NNM- ZMA); G ‒ K, C. valida n. sp., Fiji: holotype male 3.6 × 4.9 mm (MNHN-IU- 2013 - 9067). A, anterior thoracic sternum; B, male pleon; C, D, H, I, ventral view of left G 1; E, J, dorsal view of left G 1; F, K, left G 2; G, left third maxilliped (setae not drawn). Scales: A, B, G = 0.5 mm; C – F = 0.25 mm; H – K = 0.1 mm." httpUri="https://zenodo.org/record/272715/files/figure.png" pageId="31" pageNumber="32">Fig. 69</figureCitation>
C‒E), and is not strongly twisted with the distal parts adorned with long setae (see discussion for
<taxonomicName id="BEC231A6FFBCFFE4FB08F919FA94F959" box="[1269,1369,1736,1760]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="4BB69637FFBCFFE4FB08F919FA94F959" box="[1269,1369,1736,1760]" italics="true" pageId="31" pageNumber="32">C. minor</emphasis>
</taxonomicName>
). The structure of the male pleon and position of the vulvae of
<taxonomicName id="BEC231A6FFBCFFE4FCEFF93CFCBBF8BD" box="[786,886,1773,1796]" class="Malacostraca" family="Chasmocarcinidae" genus="Camatopsis" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="4BB69637FFBCFFE4FCEFF93CFCBBF8BD" box="[786,886,1773,1796]" italics="true" pageId="31" pageNumber="32">C. minor</emphasis>
</taxonomicName>
are like those of its congeners.
</paragraph>
</subSubSection>
<subSubSection id="31D819AEFFBCFFE4FF3AF8DEFC32F89E" box="[199,1023,1807,1832]" pageId="31" pageNumber="32" type="distribution">
<paragraph id="797D4A25FFBCFFE4FF3AF8DEFC32F89E" blockId="31.[151,1437,151,1832]" box="[199,1023,1807,1832]" pageId="31" pageNumber="32">
<emphasis id="4BB69637FFBCFFE4FF3AF8DEFE94F891" bold="true" box="[199,345,1807,1832]" pageId="31" pageNumber="32">Distribution</emphasis>
. Indo-West Pacific region: Indian to western Pacific oceans.
</paragraph>
</subSubSection>
</treatment>
</document>
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