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<document id="1BD4352F7C8DAD90A9D7C209A3CE704D" ID-CLB-Dataset="37321" ID-DOI="10.11646/zootaxa.4209.1.1" ID-GBIF-Dataset="2de5b7bf-3975-458a-b200-07ec69f4e013" ID-ISSN="1175-5326" ID-Zenodo-Dep="272646" ID-ZooBank="849BAB5C-464A-4B4A-A586-5742411EDC01" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484407904318" checkinUser="plazi" docAuthor="Ng, Peter K. L. &amp; Castro, Peter" docDate="2016" docId="F16BFB33FF3EFF65FF6AF995FB9CFE31" docLanguage="en" docName="zootaxa.4209.1.1.pdf.imf" docOrigin="Zootaxa 4209 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Australocarcinus palauensis Davie &amp; Guinot 1996" docType="treatment" docVersion="8" lastPageNumber="159" masterDocId="0D52834BFFA3FFFBFFFDFFD1FFCDFFB9" masterDocTitle="Revision of the family Chasmocarcinidae Serène, 1964 (Crustacea, Brachyura, Goneplacoidea)" masterLastPageNumber="182" masterPageNumber="1" pageNumber="158" updateTime="1698435566492" updateUser="ExternalLinkService">
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<mods:title id="914B657BB0BF31BDBA9FC659CBD10304">Revision of the family Chasmocarcinidae Serène, 1964 (Crustacea, Brachyura, Goneplacoidea)</mods:title>
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<treatment id="F16BFB33FF3EFF65FF6AF995FB9CFE31" ID-DOI="http://doi.org/10.5281/zenodo.5617285" ID-GBIF-Taxon="127419614" ID-Zenodo-Dep="5617285" LSID="urn:lsid:plazi:treatment:F16BFB33FF3EFF65FF6AF995FB9CFE31" httpUri="http://treatment.plazi.org/id/F16BFB33FF3EFF65FF6AF995FB9CFE31" lastPageId="158" lastPageNumber="159" pageId="157" pageNumber="158">
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<paragraph id="797D4A25FF3EFF66FF6AF995FCCAF9E7" blockId="157.[151,775,1604,1663]" box="[151,775,1604,1630]" pageId="157" pageNumber="158">
<heading id="2235FD49FF3EFF66FF6AF995FCCAF9E7" bold="true" box="[151,775,1604,1630]" fontSize="11" level="1" pageId="157" pageNumber="158" reason="1">
<taxonomicName id="BEC231A6FF3EFF66FF6AF995FCCAF9E7" ID-CoL="5W9PP" authority="Davie &amp; Guinot, 1996" authorityName="Davie &amp; Guinot" authorityYear="1996" box="[151,775,1604,1630]" class="Malacostraca" family="Chasmocarcinidae" genus="Australocarcinus" kingdom="Animalia" order="Decapoda" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="palauensis">
<emphasis id="4BB69637FF3EFF66FF6AF995FCCAF9E7" bold="true" box="[151,775,1604,1630]" pageId="157" pageNumber="158">
<emphasis id="4BB69637FF3EFF66FF6AF995FE22F9E7" bold="true" box="[151,495,1604,1630]" italics="true" pageId="157" pageNumber="158">Australocarcinus palauensis</emphasis>
<bibRefCitation id="1D5337D4FF3EFF66FE0BF995FCCAF9E7" author="Davie" box="[502,775,1604,1630]" pageId="157" pageNumber="158" refString="Davie, P. J. F. &amp; Guinot, D. (1996) Two new freshwater species in Australocarcinus Davie, with remarks on Trogloplacinae Guinot and Goneplacidae Macleay [sic] (Crustacea: Decapoda: Brachyura). Memoirs of the Queensland Museum, 39, 277 - 287." type="journal article" year="1996">Davie &amp; Guinot, 1996</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="797D4A25FF3EFF66FF6AF9B7FE25F9C6" blockId="157.[151,775,1604,1663]" box="[151,488,1638,1663]" pageId="157" pageNumber="158">
(
<figureCitation id="E1F956A0FF3EFF66FF62F9B7FF31F9C6" box="[159,252,1638,1663]" captionStart="FIGURE 93" captionStartId="166.[151,250,1487,1509]" captionTargetBox="[151,1436,193,1466]" captionTargetId="figure@166.[151,1436,193,1466]" captionTargetPageId="166" captionText="FIGURE 93. Overall dorsal view of species of Australocarcinus and Trogloplax. A, Australocarcinus riparius, Australia: male 8.8 × 10.2 mm (ZRC 2006.167); B, A. riparius, Australia: female 9.0 × 11.3 mm (ZRC 2006.167); C, Australocarcinus kanaka, New Caledonia: female 8.5 × 9.8 mm (ZRC 2009.111); D, Australocarcinus palauensis, Palau: holotype female (5.8 × 7.1 mm) (USNM 277853); E, F, Trogloplax joliveti, New Britain: holotype male 16.0 × 21.0 mm (MNHN-IU- 2014 - 10457) (after Guinot 1986: pl. 1 A, B); G, Trogloplax joliveti, New Britain: male 15.7 × 18.6 mm (ZRC 2015.294); H, Trogloplax joliveti, New Britain: female 14.1 × 16.8 mm (ZRC 2015.295)." httpUri="https://zenodo.org/record/272739/files/figure.png" pageId="157" pageNumber="158">Figs. 93</figureCitation>
D; 94C, D, I; 99E, F)
</paragraph>
</subSubSection>
<subSubSection id="31D819AEFF3EFF66FF6AF97FFA83F97D" box="[151,1358,1710,1733]" pageId="157" pageNumber="158" type="reference_group">
<paragraph id="797D4A25FF3EFF66FF6AF97FFA83F97D" blockId="157.[151,1358,1710,1733]" box="[151,1358,1710,1733]" pageId="157" pageNumber="158">
<treatmentCitationGroup id="59D26D0BFF3EFF66FF6AF97FFA83F97D" box="[151,1358,1710,1733]" pageId="157" pageNumber="158">
<taxonomicName id="BEC231A6FF3EFF66FF6AF97FFD1BF97D" ID-CoL="5W9PP" authority="Davie &amp; Guinot, 1996: 282" authorityName="Davie &amp; Guinot" authorityPageNumber="282" authorityYear="1996" box="[151,726,1710,1732]" class="Malacostraca" family="Chasmocarcinidae" genus="Australocarcinus" kingdom="Animalia" order="Decapoda" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="palauensis">
<emphasis id="4BB69637FF3EFF66FF6AF97FFE75F97D" box="[151,440,1710,1732]" italics="true" pageId="157" pageNumber="158">Australocarcinus palauensis</emphasis>
<treatmentCitation id="F8636C34FF3EFF66FE40F97FFD1BF97D" author="Davie" box="[445,726,1710,1732]" page="282" pageId="157" pageNumber="158" year="1996">
<bibRefCitation id="1D5337D4FF3EFF66FE40F97FFD6EF97D" author="Davie" box="[445,675,1710,1732]" pageId="157" pageNumber="158" refString="Davie, P. J. F. &amp; Guinot, D. (1996) Two new freshwater species in Australocarcinus Davie, with remarks on Trogloplacinae Guinot and Goneplacidae Macleay [sic] (Crustacea: Decapoda: Brachyura). Memoirs of the Queensland Museum, 39, 277 - 287." type="journal article" year="1996">Davie &amp; Guinot, 1996</bibRefCitation>
: 282
</treatmentCitation>
</taxonomicName>
, figs. 5, 8 [map] [Belau = Palau].—Ng
<emphasis id="4BB69637FF3EFF66FB98F961FB5FF97D" box="[1125,1170,1710,1733]" italics="true" pageId="157" pageNumber="158">et al</emphasis>
. 2008: 76 [in list].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="31D819AEFF3EFF66FF6AF920FD69F897" pageId="157" pageNumber="158" type="materials_examined">
<paragraph id="797D4A25FF3EFF66FF6AF920FD69F897" blockId="157.[151,1436,1777,2018]" pageId="157" pageNumber="158">
<materialsCitation id="C9AA4078FF3EFF66FF6AF920FD69F897" ID-GBIF-Occurrence="1424725831" collectionCode="USNM" collectorName="Belau &amp; Babelthuab I. &amp; D. Frey" country="Palau" location="Addeids River" pageId="157" pageNumber="158" specimenCode="USNM 277853" specimenCount="2" typeStatus="holotype">
<emphasis id="4BB69637FF3EFF66FF6AF920FEF3F8B3" bold="true" box="[151,318,1777,1802]" pageId="157" pageNumber="158">
<typeStatus id="A679F487FF3EFF66FF6AF920FF1FF8B3" box="[151,210,1777,1802]" pageId="157" pageNumber="158">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="A679F487FF3EFF66FEB1F920FE79F8B3" box="[332,436,1777,1802]" pageId="157" pageNumber="158" type="holotype">Holotype</typeStatus>
female (5.8 ×
<quantity id="BE3AE7C0FF3EFF66FDA7F923FD7FF8B3" box="[602,690,1778,1802]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="157" pageNumber="158" unit="mm" value="7.1">7.1 mm</quantity>
) (
<specimenCode id="2964E25EFF3EFF66FD34F923FCB6F8B3" box="[713,891,1777,1802]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="157" pageNumber="158">
<collectionCode id="1FD3D2E0FF3EFF66FD34F923FCD1F8B3" box="[713,796,1778,1802]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="157" pageNumber="158">USNM</collectionCode>
277853
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),
<collectorName id="D4372FF3FF3EFF66FC72F920FC1DF8B3" box="[911,976,1777,1802]" pageId="157" pageNumber="158">Belau</collectorName>
[=
<collectingCountry id="01D50AB5FF3EFF66FC05F920FBF1F8B3" box="[1016,1084,1777,1802]" name="Palau" pageId="157" pageNumber="158">Palau</collectingCountry>
],
<location id="7C1D1CFEFF3EFF66FBB3F920FB3FF8B3" LSID="urn:lsid:plazi:treatment:F16BFB33FF3EFF65FF6AF995FB9CFE31:7C1D1CFEFF3EFF66FBB3F920FB3FF8B3" box="[1102,1266,1777,1802]" country="Palau" name="Addeids River" pageId="157" pageNumber="158">Addeids River</location>
,
<collectorName id="D4372FF3FF3EFF66FB03F920FA5BF8B0" box="[1278,1430,1777,1802]" pageId="157" pageNumber="158">Babelthuab I.</collectorName>
, in small fast tributary, coll.
<collectorName id="D4372FF3FF3EFF66FE36F8C7FDEEF897" box="[459,547,1814,1838]" pageId="157" pageNumber="158">D. Frey</collectorName>
, 5.03.1946.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="31D819AEFF3EFF65FF3AF8E8FE0AFF41" lastPageId="158" lastPageNumber="159" pageId="157" pageNumber="158" type="diagnosis">
<paragraph id="797D4A25FF3EFF66FF3AF8E8FA51F85B" blockId="157.[151,1436,1777,2018]" pageId="157" pageNumber="158">
<emphasis id="4BB69637FF3EFF66FF3AF8E8FEF7F8EB" bold="true" box="[199,314,1849,1874]" pageId="157" pageNumber="158">Diagnosis</emphasis>
(modified from
<bibRefCitation id="1D5337D4FF3EFF66FDF8F8E8FCC3F8EB" author="Davie" box="[517,782,1849,1874]" pageId="157" pageNumber="158" refString="Davie, P. J. F. &amp; Guinot, D. (1996) Two new freshwater species in Australocarcinus Davie, with remarks on Trogloplacinae Guinot and Goneplacidae Macleay [sic] (Crustacea: Decapoda: Brachyura). Memoirs of the Queensland Museum, 39, 277 - 287." type="journal article" year="1996">Davie &amp; Guinot 1996</bibRefCitation>
). Carapace (
<figureCitation id="E1F956A0FF3EFF66FC58F8E8FC30F8EB" box="[933,1021,1849,1874]" captionStart="FIGURE 93" captionStartId="166.[151,250,1487,1509]" captionTargetBox="[151,1436,193,1466]" captionTargetId="figure@166.[151,1436,193,1466]" captionTargetPageId="166" captionText="FIGURE 93. Overall dorsal view of species of Australocarcinus and Trogloplax. A, Australocarcinus riparius, Australia: male 8.8 × 10.2 mm (ZRC 2006.167); B, A. riparius, Australia: female 9.0 × 11.3 mm (ZRC 2006.167); C, Australocarcinus kanaka, New Caledonia: female 8.5 × 9.8 mm (ZRC 2009.111); D, Australocarcinus palauensis, Palau: holotype female (5.8 × 7.1 mm) (USNM 277853); E, F, Trogloplax joliveti, New Britain: holotype male 16.0 × 21.0 mm (MNHN-IU- 2014 - 10457) (after Guinot 1986: pl. 1 A, B); G, Trogloplax joliveti, New Britain: male 15.7 × 18.6 mm (ZRC 2015.294); H, Trogloplax joliveti, New Britain: female 14.1 × 16.8 mm (ZRC 2015.295)." httpUri="https://zenodo.org/record/272739/files/figure.png" pageId="157" pageNumber="158">Fig. 93</figureCitation>
D;
<bibRefCitation id="1D5337D4FF3EFF66FBD9F8E8FAE0F8EB" author="Davie" box="[1060,1325,1849,1874]" pageId="157" pageNumber="158" refString="Davie, P. J. F. &amp; Guinot, D. (1996) Two new freshwater species in Australocarcinus Davie, with remarks on Trogloplacinae Guinot and Goneplacidae Macleay [sic] (Crustacea: Decapoda: Brachyura). Memoirs of the Queensland Museum, 39, 277 - 287." type="journal article" year="1996">Davie &amp; Guinot 1996</bibRefCitation>
: fig. 5A) subrectangular, 1.2 wider than long; front bilobed, with slight median cleft; dorsal surface smooth. Anterolateral margins arcuate, cristate, without teeth. Orbits short. Eye peduncle short, does not fill orbit, cornea enlarged. Third maxilliped (
<figureCitation id="E1F956A0FF3EFF66FEDEF874FEB4F807" box="[291,377,1957,1982]" captionStart="FIGURE 94" captionStartId="167.[151,250,1585,1607]" captionTargetBox="[151,1436,193,1564]" captionTargetId="figure@167.[151,1436,193,1564]" captionTargetPageId="167" captionText="FIGURE 94. Frontal view and third maxilliped of species of Australocarcinus and Trogloplax. A, Australocarcinus riparius, Australia: male 8.8 × 10.2 mm (ZRC 2006.167); B, Australocarcinus kanaka, New Caledonia: female 8.5 × 9.8 mm (ZRC 2009.111); C, D, Australocarcinus palauensis, Palau: holotype female (5.8 × 7.1 mm) (USNM 277853); E, Trogloplax joliveti, New Britain: male 15.7 × 18.6 mm (MNHN-B 20136); F, Trogloplax joliveti, New Britain: female 14.1 × 16.8 mm (ZRC 2015.295); G, Australocarcinus riparius, Australia: holotype male (after Davie 1988: fig. 1 C); H, Australocarcinus kanaka, New Caledonia: paratype female 9.5 × 11.1 mm (after Davie &amp; Guinot 1996: fig. 3 C); I, Australocarcinus palauensis, Palau: holotype female (5.8 × 7.1 mm) (USNM 277853) (after Davie &amp; Guinot 1996: fig. 5 B); J, Trogloplax joliveti, New Britain: holotype male 16.0 × 21.0 mm (MNHN-IU- 2014 - 10457) (after Guinot 1986: pl. 2 B)." httpUri="https://zenodo.org/record/272740/files/figure.png" pageId="157" pageNumber="158">Fig. 94</figureCitation>
D, I) merus quadrate, ischium quadrate, slightly longer than merus. Chelipeds subequal in length; cutting margins of both chelipeds with broad, shallow teeth larger in major chela (
<figureCitation id="E1F956A0FF3EFF66FB06F818FA9BF85B" box="[1275,1366,1993,2018]" captionStart="FIGURE 95" captionStartId="168.[151,250,1631,1653]" captionTargetBox="[151,1436,193,1609]" captionTargetId="figure@168.[151,1436,193,1609]" captionTargetPageId="168" captionText="FIGURE 95. Chelae of species of Australocarcinus and Trogloplax. A, B, Australocarcinus riparius, Australia: male 8.8 × 10.2 mm (ZRC 2006.167); C, D, Australocarcinus kanaka, New Caledonia: female 8.5 × 9.8 mm (ZRC 2009.111); E, F, Australocarcinus palauensis, Palau: holotype female (5.8 × 7.1 mm) (USNM 277853); G, H, Trogloplax joliveti, New Britain: male 15.7 × 18.6 mm (ZRC 2015.294)." httpUri="https://zenodo.org/record/272741/files/figure.png" pageId="157" pageNumber="158">Fig. 95</figureCitation>
E, F).
</paragraph>
<paragraph id="797D4A25FF3DFF65FF6AFF46FE0AFF41" blockId="158.[151,1436,151,392]" pageId="158" pageNumber="159">
Proportionally short ambulatory legs (
<figureCitation id="E1F956A0FF3DFF65FDC2FF46FD5CFF09" box="[575,657,151,176]" captionStart="FIGURE 93" captionStartId="166.[151,250,1487,1509]" captionTargetBox="[151,1436,193,1466]" captionTargetId="figure@166.[151,1436,193,1466]" captionTargetPageId="166" captionText="FIGURE 93. Overall dorsal view of species of Australocarcinus and Trogloplax. A, Australocarcinus riparius, Australia: male 8.8 × 10.2 mm (ZRC 2006.167); B, A. riparius, Australia: female 9.0 × 11.3 mm (ZRC 2006.167); C, Australocarcinus kanaka, New Caledonia: female 8.5 × 9.8 mm (ZRC 2009.111); D, Australocarcinus palauensis, Palau: holotype female (5.8 × 7.1 mm) (USNM 277853); E, F, Trogloplax joliveti, New Britain: holotype male 16.0 × 21.0 mm (MNHN-IU- 2014 - 10457) (after Guinot 1986: pl. 1 A, B); G, Trogloplax joliveti, New Britain: male 15.7 × 18.6 mm (ZRC 2015.294); H, Trogloplax joliveti, New Britain: female 14.1 × 16.8 mm (ZRC 2015.295)." httpUri="https://zenodo.org/record/272739/files/figure.png" pageId="158" pageNumber="159">Fig. 93</figureCitation>
D). Meri smooth. Somites of female pleon with slightly convex lateral margins; telson proportionally long (
<figureCitation id="E1F956A0FF3DFF65FDCDFF6DFD4FFF6D" box="[560,642,188,213]" captionStart="FIGURE 99" captionStartId="172.[151,250,1484,1506]" captionTargetBox="[151,1436,193,1462]" captionTargetId="figure@172.[151,1436,193,1462]" captionTargetPageId="172" captionText="FIGURE 99. Female pleon and vulvae of species of Australocarcinus and Trogloplax. A, B, Australocarcinus riparius, Australia: female 9.0 × 11.3 mm (ZRC 2006.167); C, D, Australocarcinus kanaka, New Caledonia: female 8.5 × 9.8 mm (ZRC 2009.111); E, F, Australocarcinus palauensis, Palau: holotype female (5.8 × 7.1 mm) (USNM 277853); G, H, Trogloplax joliveti, New Britain: female 14.1 × 16.8 mm (ZRC 2015.295). A, C, E, G, thoracic sternum with pleon; B, D, F, H, female sterno-pleonal cavity with vulvae." httpUri="https://zenodo.org/record/272745/files/figure.png" pageId="158" pageNumber="159">Fig. 99</figureCitation>
E). Sterno-pleonal cavity of female shallow, vulvae not fully developed (
<figureCitation id="E1F956A0FF3DFF65FF62FF0EFF3FFF41" box="[159,242,223,248]" captionStart="FIGURE 99" captionStartId="172.[151,250,1484,1506]" captionTargetBox="[151,1436,193,1462]" captionTargetId="figure@172.[151,1436,193,1462]" captionTargetPageId="172" captionText="FIGURE 99. Female pleon and vulvae of species of Australocarcinus and Trogloplax. A, B, Australocarcinus riparius, Australia: female 9.0 × 11.3 mm (ZRC 2006.167); C, D, Australocarcinus kanaka, New Caledonia: female 8.5 × 9.8 mm (ZRC 2009.111); E, F, Australocarcinus palauensis, Palau: holotype female (5.8 × 7.1 mm) (USNM 277853); G, H, Trogloplax joliveti, New Britain: female 14.1 × 16.8 mm (ZRC 2015.295). A, C, E, G, thoracic sternum with pleon; B, D, F, H, female sterno-pleonal cavity with vulvae." httpUri="https://zenodo.org/record/272745/files/figure.png" pageId="158" pageNumber="159">Fig. 99</figureCitation>
F). Male unknown.
</paragraph>
</subSubSection>
<subSubSection id="31D819AEFF3DFF65FF3AFED5FECAFEDC" pageId="158" pageNumber="159" type="discussion">
<paragraph id="797D4A25FF3DFF65FF3AFED5FECAFEDC" blockId="158.[151,1436,151,392]" pageId="158" pageNumber="159">
<emphasis id="4BB69637FF3DFF65FF3AFED5FEF9FEA4" bold="true" box="[199,308,260,285]" pageId="158" pageNumber="159">Remarks</emphasis>
. The species is known from a single young female specimen (pleon, pleopods and vulvae not mature,
<figureCitation id="E1F956A0FF3DFF65FF6AFEF6FF2AFEF9" box="[151,231,295,320]" captionStart="FIGURE 99" captionStartId="172.[151,250,1484,1506]" captionTargetBox="[151,1436,193,1462]" captionTargetId="figure@172.[151,1436,193,1462]" captionTargetPageId="172" captionText="FIGURE 99. Female pleon and vulvae of species of Australocarcinus and Trogloplax. A, B, Australocarcinus riparius, Australia: female 9.0 × 11.3 mm (ZRC 2006.167); C, D, Australocarcinus kanaka, New Caledonia: female 8.5 × 9.8 mm (ZRC 2009.111); E, F, Australocarcinus palauensis, Palau: holotype female (5.8 × 7.1 mm) (USNM 277853); G, H, Trogloplax joliveti, New Britain: female 14.1 × 16.8 mm (ZRC 2015.295). A, C, E, G, thoracic sternum with pleon; B, D, F, H, female sterno-pleonal cavity with vulvae." httpUri="https://zenodo.org/record/272745/files/figure.png" pageId="158" pageNumber="159">Fig. 99</figureCitation>
F) so the morphology of males, including the pleon, posterior part of thoracic sternum, G1, and G2, remains unknown.
</paragraph>
</subSubSection>
<subSubSection id="31D819AEFF3DFF65FF3AFEBEFB9CFE31" box="[199,1105,367,392]" pageId="158" pageNumber="159" type="distribution">
<paragraph id="797D4A25FF3DFF65FF3AFEBEFB9CFE31" blockId="158.[151,1436,151,392]" box="[199,1105,367,392]" pageId="158" pageNumber="159">
<emphasis id="4BB69637FF3DFF65FF3AFEBEFE94FE31" bold="true" box="[199,345,367,392]" pageId="158" pageNumber="159">Distribution</emphasis>
. Known only from
<collectingCountry id="01D50AB5FF3DFF65FDCAFEBEFDB5FE31" box="[567,632,367,392]" name="Palau" pageId="158" pageNumber="159">Belau</collectingCountry>
[=
<collectingCountry id="01D50AB5FF3DFF65FD62FEBEFD2DFE31" box="[671,736,367,392]" name="Palau" pageId="158" pageNumber="159">Palau</collectingCountry>
] (western Pacific Ocean); rivers.
</paragraph>
</subSubSection>
</treatment>
</document>