treatments-xml/data/03/B7/C7/03B7C77AFFDEFFEABCF5FF080834065F.xml
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<document id="EBF239CBF6DF8BC105D3DAD502570759" ID-DOI="10.25674/so95iss1id311" ID-ISSN="2509-9523" ID-Zenodo-Dep="10724236" checkinTime="1709147239332" checkinUser="felipe" docAuthor="Frisch, Johannes &amp; Narakusumo, Raden Pramesa" docDate="2022" docId="03B7C77AFFDEFFEABCF5FF080834065F" docLanguage="en" docName="SoilOrg.95.1.23-73.pdf" docOrigin="Soil Organisms 95 (1)" docSource="http://dx.doi.org/10.25674/so95iss1id311" docStyle="DocumentStyle:EA2ECAC87EC21C07AB805D5B9B4601F2:SoilOrg.2018-.journal_article" docStyleId="EA2ECAC87EC21C07AB805D5B9B4601F2" docStyleName="SoilOrg.2018-.journal_article" docTitle="Scopaeus velifer Frisch 2022, spec. nov." docType="treatment" docVersion="2" lastPageNumber="53" masterDocId="FF8EBF02FFC0FFF4BC5BFFF80D1D040D" masterDocTitle="Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species" masterLastPageNumber="73" masterPageNumber="23" pageNumber="53" updateTime="1709147743548" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="AC02CF523D7EFFCD13332922A2D10B24">Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species</mods:title>
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<mods:namePart id="592F5BA0A77BD86BE19261A0A1C7911A">Frisch, Johannes</mods:namePart>
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<mods:namePart id="E7730B1EE88571EFD5023AAB9D5CC7BF">Narakusumo, Raden Pramesa</mods:namePart>
<mods:affiliation id="D4E68A287BA704BF17B2014890B9F86F">MUseUm ZoologicUm Bogoriense, Research Center for Biosystematics and EvolUtion, National Research and Innovation Agency (BRIN), Jl. Raya Jakarta-Bogor KM 46, Cibinong, West Java, 16911, Indonesia</mods:affiliation>
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<mods:title id="EB8940D1C5908E2ACDEB94AEDFF23A81">Soil Organisms</mods:title>
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<mods:date id="60566DC8BF10FFE74E27FAC8761EE443">2022</mods:date>
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<treatment id="03B7C77AFFDEFFEABCF5FF080834065F" ID-DOI="http://doi.org/10.5281/zenodo.10724319" ID-Zenodo-Dep="10724319" LSID="urn:lsid:plazi:treatment:03B7C77AFFDEFFEABCF5FF080834065F" httpUri="http://treatment.plazi.org/id/03B7C77AFFDEFFEABCF5FF080834065F" lastPageNumber="53" pageId="30" pageNumber="53">
<subSubSection id="C30425E7FFDEFFEABCF5FF080F4D0507" box="[174,592,239,266]" pageId="30" pageNumber="53" type="nomenclature">
<paragraph id="8BA1766CFFDEFFEABCF5FF080F4D0507" blockId="30.[174,592,239,299]" box="[174,592,239,266]" pageId="30" pageNumber="53">
<heading id="D0E9C100FFDEFFEABCF5FF080F4D0507" box="[174,592,239,266]" fontSize="11" level="2" pageId="30" pageNumber="53" reason="5">
<taxonomicName id="4C1E0DEFFFDEFFEABCF5FF080CD30504" authority="Frisch" authorityName="Frisch" authorityYear="2022" box="[174,462,239,266]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="53" phylum="Arthropoda" rank="species" species="velifer" status="sp. nov.">
<emphasis id="B96AAA7EFFDEFFEABCF5FF080CD30504" bold="true" box="[174,462,239,266]" pageId="30" pageNumber="53">
<emphasis id="B96AAA7EFFDEFFEABCF5FF080C680507" bold="true" box="[174,373,239,266]" italics="true" pageId="30" pageNumber="53">Scopaeus velifer</emphasis>
Frisch
</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2591705FFDEFFEABD87FF080F4D0507" box="[476,592,240,266]" pageId="30" pageNumber="53" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
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<subSubSection id="C30425E7FFDEFFEABCF5FEE80F040526" box="[174,537,272,299]" pageId="30" pageNumber="53" type="description">
<paragraph id="8BA1766CFFDEFFEABCF5FEE80F040526" blockId="30.[174,592,239,299]" box="[174,537,272,299]" pageId="30" pageNumber="53">
(
<figureCitation id="13256AE9FFDEFFEABCEDFEE80C100526" box="[182,269,272,299]" captionStart="Figures 1722" captionStartId="11.[189,262,1908,1930]" captionTargetBox="[279,1382,235,1877]" captionTargetId="figure-90@11.[206,1416,206,1889]" captionTargetPageId="11" captionText="Figures 1722. Habitus of Scopaeusvelifer, paratype,Banten, Lebak:Citorek Kidul (17); S. elegantulus, Jawa Barat, Sukabumi:Djampang Kulon (18); S. uncinatus, paratype, Jawa Barat, Sukabumi: Cidahu (19); S. bipenicillatus, paratype, Sulawesi Tengah, Sigi: Pagana (20); S. bipectenatus, paratype, Sulawesi Tengah: Lake Poso (21); S. crassipunctatus, paratype, Jawa Barat, Sukabumi: Sirnarasa (22)." figureDoi="http://doi.org/10.5281/zenodo.10724250" httpUri="https://zenodo.org/record/10724250/files/figure.png" pageId="30" pageNumber="53">Figs 17</figureCitation>
,
<figureCitation id="13256AE9FFDEFFEABD43FEE90C250526" box="[280,312,273,299]" captionStart="Figures 4249" captionStartId="17.[190,263,1886,1908]" captionTargetBox="[263,1341,889,1832]" captionTargetId="figure-379@17.[189,1436,851,1867]" captionTargetPageId="17" captionText="Figures 4249. Aedeagus in lateral view of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul (42); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (43); S. uncinatus, holotype,Banten, Lebak:Citorek Kidul (44); S. bipenicillatus, holotype,Sulawesi Tengah, Sigi: Pagana (45); S. bipectenatus, holotype, Sulawesi Tengah, Poso: Lake Poso (46); S. cuspidatus, holotype, Bali, Jembrana: Cekik (47); S. crassipunctatus, holotype, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (48); S. sulawesianus, holotype, Sulawesi Selatan: Engrekang (49)." figureDoi="http://doi.org/10.5281/zenodo.10724262" httpUri="https://zenodo.org/record/10724262/files/figure.png" pageId="30" pageNumber="53">42</figureCitation>
,
<figureCitation id="13256AE9FFDEFFEABD1FFEE90CB90526" box="[324,420,273,299]" captionStart="Figures 98103" captionStartId="29.[189,262,1884,1906]" captionTargetBox="[262,1360,393,1733]" captionTargetId="figure-128@29.[189,1436,334,1865]" captionTargetPageId="29" captionText="Figures 98103. Aedeagus of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul in lateral (98), ventral (99), dorsal view (100); apex of aedeagus of S. uncinatus, holotype, Banten, Lebak: Citorek Kidul in lateral (101), ventral (102), dorsal view (103). Abbreviations: al apical lobes, dl dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, tr transverse ridge of median foramen, v velum, vl ventral lobe. Arrows: small, ventral lobe (I); distal enlargement of lateral lobe (II); apicodorsal bridge of apical lobes (III)." figureDoi="http://doi.org/10.5281/zenodo.10724284" httpUri="https://zenodo.org/record/10724284/files/figure.png" pageId="30" pageNumber="53">98 100</figureCitation>
, 135, 169)
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFDEFFEABCF5FEAC0CE105C2" pageId="30" pageNumber="53" type="materials_examined">
<paragraph id="8BA1766CFFDEFFEABCF5FEAC0CE105C2" blockId="30.[151,764,340,1975]" pageId="30" pageNumber="53">
<emphasis id="B96AAA7EFFDEFFEABCF5FEAC0C680561" bold="true" box="[174,373,340,364]" pageId="30" pageNumber="53">Type specimens:</emphasis>
<materialsCitation id="3B767C31FFDEFFEABDDCFEAC0CC905A3" collectingDate="2016-05-23" collectionCode="MZB" collectorName="Frisch" country="Indonesia" elevation="870" latitude="-6.7475004" location="Citorek Kidul" longLatPrecision="21" longitude="106.320274" municipality="Lebak" pageId="30" pageNumber="53" specimenCount="1" specimenCount-male="1" stateProvince="Banten" typeStatus="holotype">
<collectingCountry id="F30936FCFFDEFFEABDDCFEAC0CE90561" box="[391,500,340,364]" name="Indonesia" pageId="30" pageNumber="53">Indonesia</collectingCountry>
:
<typeStatus id="54A5C8CEFFDEFFEABE53FEAC0F710561" box="[520,620,340,364]" pageId="30" pageNumber="53" type="holotype">Holotype</typeStatus>
<specimenCount id="9D18BDE5FFDEFFEABE25FEAD0F8F0561" box="[638,658,341,364]" count="1" pageId="30" pageNumber="53" type="male"></specimenCount>
,
<collectingRegion id="49DAB88EFFDEFFEABEF3FEAD0FEA0561" box="[680,759,341,364]" country="Indonesia" name="Banten" pageId="30" pageNumber="53">Banten</collectingRegion>
,
<collectingMunicipality id="6BC5EC16FFDEFFEABCCCFE8D0DC20580" box="[151,223,373,397]" pageId="30" pageNumber="53">Lebak</collectingMunicipality>
:
<location id="8EC120B7FFDEFFEABCB2FE8D0C600580" LSID="urn:lsid:plazi:treatment:03B7C77AFFDEFFEABCF5FF080834065F:8EC120B7FFDEFFEABCB2FE8D0C600580" box="[233,381,373,397]" country="Indonesia" latitude="-6.7475004" longLatPrecision="21" longitude="106.320274" municipality="Lebak" name="Citorek Kidul" pageId="30" pageNumber="53" stateProvince="Banten">Citorek Kidul</location>
(
<geoCoordinate id="EE2A10ABFFDEFFEABDD7FE8D0F140580" box="[396,521,373,397]" degrees="06" direction="south" minutes="44" orientation="latitude" pageId="30" pageNumber="53" precision="15" seconds="51" value="-6.7475004">06°4451S</geoCoordinate>
,
<geoCoordinate id="EE2A10ABFFDEFFEABE4FFE8D0F820580" box="[532,671,373,397]" degrees="106" direction="east" minutes="19" orientation="longitude" pageId="30" pageNumber="53" precision="15" seconds="13" value="106.320274">106°1913E</geoCoordinate>
),
<quantity id="4CE6DB89FFDEFFEABEE9FE8D0FEA0580" box="[690,759,373,397]" metricMagnitude="2" metricUnit="m" metricValue="8.7" pageId="30" pageNumber="53" unit="m" value="870.0">
<elevation id="0033915FFFDEFFEABEE9FE8D0FEA0580" box="[690,759,373,397]" metricMagnitude="2" metricUnit="m" metricValue="8.7" pageId="30" pageNumber="53" unit="m" value="870.0">870 m</elevation>
</quantity>
,
<date id="FFA050ACFFDEFFEABCCCFE6E0C1D05A3" box="[151,256,406,430]" pageId="30" pageNumber="53" value="2016-05-23">
<collectingDate id="EFE4A944FFDEFFEABCCCFE6E0C1D05A3" box="[151,256,406,430]" pageId="30" pageNumber="53" value="2016-05-23">23.5.2016</collectingDate>
</date>
, leg.
<collectorName id="26EB13BAFFDEFFEABD61FE6E0C6005A3" box="[314,381,406,430]" pageId="30" pageNumber="53">Frisch</collectorName>
(
<collectionCode id="ED0FEEA9FFDEFFEABDD6FE6E0CD105A3" box="[397,460,406,430]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="30" pageNumber="53" type="Museum">MZB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B767C31FFDEFFEABD84FE6E0CE505C2" collectingDate="2016-05-23" collectionCode="MFNB, MZB" collectorName="Frisch" country="Indonesia" elevation="870" latitude="-6.7475004" location="Citorek Kidul" longLatPrecision="21" longitude="106.320274" municipality="Lebak" pageId="30" pageNumber="53" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Banten" typeStatus="paratype">
<typeStatus id="54A5C8CEFFDEFFEABD84FE6E0F5005A3" box="[479,589,406,430]" pageId="30" pageNumber="53" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D18BDE5FFDEFFEABE03FE6E0F9C05A0" box="[600,641,406,430]" count="1" pageId="30" pageNumber="53" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D18BDE5FFDEFFEABED5FE6E0FAB05A0" box="[654,694,406,430]" count="4" pageId="30" pageNumber="53" type="female">4 ♀</specimenCount>
, same data as holotype (
<collectionCode id="ED0FEEA9FFDEFFEABD0DFE4F0CB505C2" box="[342,424,439,463]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="30" pageNumber="53">MFNB</collectionCode>
,
<collectionCode id="ED0FEEA9FFDEFFEABDE9FE4F0CED05C2" box="[434,496,439,463]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="30" pageNumber="53" type="Museum">MZB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFDEFFEABCF5FE2F09D10561" pageId="30" pageNumber="53" type="description">
<paragraph id="8BA1766CFFDEFFEABCF5FE2F0F7507D1" blockId="30.[151,764,340,1975]" pageId="30" pageNumber="53">
<emphasis id="B96AAA7EFFDEFFEABCF5FE2F0C2105E2" bold="true" box="[174,316,471,495]" pageId="30" pageNumber="53">Description:</emphasis>
Habitus and coloring as in
<figureCitation id="13256AE9FFDEFFEABE36FE200FA505FD" box="[621,696,472,496]" captionStart="Figures 1722" captionStartId="11.[189,262,1908,1930]" captionTargetBox="[279,1382,235,1877]" captionTargetId="figure-90@11.[206,1416,206,1889]" captionTargetPageId="11" captionText="Figures 1722. Habitus of Scopaeusvelifer, paratype,Banten, Lebak:Citorek Kidul (17); S. elegantulus, Jawa Barat, Sukabumi:Djampang Kulon (18); S. uncinatus, paratype, Jawa Barat, Sukabumi: Cidahu (19); S. bipenicillatus, paratype, Sulawesi Tengah, Sigi: Pagana (20); S. bipectenatus, paratype, Sulawesi Tengah: Lake Poso (21); S. crassipunctatus, paratype, Jawa Barat, Sukabumi: Sirnarasa (22)." figureDoi="http://doi.org/10.5281/zenodo.10724250" httpUri="https://zenodo.org/record/10724250/files/figure.png" pageId="30" pageNumber="53">Fig. 17</figureCitation>
. Head subquadrate. Penultimate antennal segment quadrate. Mesotibia moderately thickened. Forebody surface matt with dense, rugulose, setose punctation; abdomen somewhat shiny; microreticulation absent. Pubescence of body surface short, decumbent, without conspicuous macrosetae. Body dark brown with slight, reddish tinge except for yellow-brown tip of abdomen and appendages; antennae unicolorous yellow-brown, or with scapus medium brown and median segments, beginning with pedicellus, with black tinge, but gradually lighter towards yellow-brown terminal segments; maxillary palpi unicolorous yellow-brown or with darker penultimate segment; femora darker brown in variable extent, gradually lighter towards distal ends. Total body length
<quantity id="4CE6DB89FFDEFFEABCCCFC3D0C3D07D1" box="[151,288,965,989]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.4" metricValueMin="2.7" pageId="30" pageNumber="53" unit="mm" value="3.05" valueMax="3.4" valueMin="2.7">2.7 3.4 mm</quantity>
; forebody length
<quantity id="4CE6DB89FFDEFFEABD87FC3D0F7907D1" box="[476,612,965,989]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="30" pageNumber="53" unit="mm" value="1.7" valueMax="1.8" valueMin="1.6">1.6 1.8 mm</quantity>
.
</paragraph>
<paragraph id="8BA1766CFFDEFFEABCF5FC1E081E0527" blockId="30.[151,764,340,1975]" lastBlockId="30.[786,1400,241,594]" pageId="30" pageNumber="53">
Male: Abdominal sternite VII with unmodified posterior margin. Abdominal sternite VIII with two lateral, narrow, deep, triangular incisions occupying almost posterior half of sternite length; subbasal ridge of sternite angled projecting posteriad (Fig. 135). Aedeagus (
<figureCitation id="13256AE9FFDEFFEABD4EFB720C7000AF" box="[277,365,1162,1186]" captionStart="Figures 4249" captionStartId="17.[190,263,1886,1908]" captionTargetBox="[263,1341,889,1832]" captionTargetId="figure-379@17.[189,1436,851,1867]" captionTargetPageId="17" captionText="Figures 4249. Aedeagus in lateral view of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul (42); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (43); S. uncinatus, holotype,Banten, Lebak:Citorek Kidul (44); S. bipenicillatus, holotype,Sulawesi Tengah, Sigi: Pagana (45); S. bipectenatus, holotype, Sulawesi Tengah, Poso: Lake Poso (46); S. cuspidatus, holotype, Bali, Jembrana: Cekik (47); S. crassipunctatus, holotype, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (48); S. sulawesianus, holotype, Sulawesi Selatan: Engrekang (49)." figureDoi="http://doi.org/10.5281/zenodo.10724262" httpUri="https://zenodo.org/record/10724262/files/figure.png" pageId="30" pageNumber="53">Figs 42</figureCitation>
,
<figureCitation id="13256AE9FFDEFFEABD26FB720CCE00AF" box="[381,467,1162,1186]" captionStart="Figures 98103" captionStartId="29.[189,262,1884,1906]" captionTargetBox="[262,1360,393,1733]" captionTargetId="figure-128@29.[189,1436,334,1865]" captionTargetPageId="29" captionText="Figures 98103. Aedeagus of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul in lateral (98), ventral (99), dorsal view (100); apex of aedeagus of S. uncinatus, holotype, Banten, Lebak: Citorek Kidul in lateral (101), ventral (102), dorsal view (103). Abbreviations: al apical lobes, dl dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, tr transverse ridge of median foramen, v velum, vl ventral lobe. Arrows: small, ventral lobe (I); distal enlargement of lateral lobe (II); apicodorsal bridge of apical lobes (III)." figureDoi="http://doi.org/10.5281/zenodo.10724284" httpUri="https://zenodo.org/record/10724284/files/figure.png" pageId="30" pageNumber="53">98 100</figureCitation>
) about
<quantity id="4CE6DB89FFDEFFEABE76FB720F6300AF" box="[557,638,1162,1186]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="30" pageNumber="53" unit="mm" value="0.6">0.6 mm</quantity>
long, with conspicuous, unique, subcircular, lateral structure, here termed velum, made up of translucent membrane studded with numerous microsetae, stretched between aedeagus and extremely lengthened flagellum accompanied by outer, thin, sclerotized ring and two inner, yellow stripes (
<figureCitation id="13256AE9FFDEFFEABCC4FAB70C200165" box="[159,317,1359,1384]" captionStart="Figures 98103" captionStartId="29.[189,262,1884,1906]" captionTargetBox="[262,1360,393,1733]" captionTargetId="figure-128@29.[189,1436,334,1865]" captionTargetPageId="29" captionText="Figures 98103. Aedeagus of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul in lateral (98), ventral (99), dorsal view (100); apex of aedeagus of S. uncinatus, holotype, Banten, Lebak: Citorek Kidul in lateral (101), ventral (102), dorsal view (103). Abbreviations: al apical lobes, dl dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, tr transverse ridge of median foramen, v velum, vl ventral lobe. Arrows: small, ventral lobe (I); distal enlargement of lateral lobe (II); apicodorsal bridge of apical lobes (III)." figureDoi="http://doi.org/10.5281/zenodo.10724284" httpUri="https://zenodo.org/record/10724284/files/figure.png" pageId="30" pageNumber="53">Figs 99, 100</figureCitation>
); flagellum thin, filamentous, arising from strongly sclerotized median structure of aedeagus dextroproximad, running dorsally across phallobase and widely sinistrad of aedeagus, and projecting from apex of apical lobes ventrally, seemingly attached to velum sinistroproximally (
<figureCitation id="13256AE9FFDEFFEABDB3FA0C0F5F0201" box="[488,578,1524,1548]" captionStart="Figures 4249" captionStartId="17.[190,263,1886,1908]" captionTargetBox="[263,1341,889,1832]" captionTargetId="figure-379@17.[189,1436,851,1867]" captionTargetPageId="17" captionText="Figures 4249. Aedeagus in lateral view of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul (42); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (43); S. uncinatus, holotype,Banten, Lebak:Citorek Kidul (44); S. bipenicillatus, holotype,Sulawesi Tengah, Sigi: Pagana (45); S. bipectenatus, holotype, Sulawesi Tengah, Poso: Lake Poso (46); S. cuspidatus, holotype, Bali, Jembrana: Cekik (47); S. crassipunctatus, holotype, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (48); S. sulawesianus, holotype, Sulawesi Selatan: Engrekang (49)." figureDoi="http://doi.org/10.5281/zenodo.10724262" httpUri="https://zenodo.org/record/10724262/files/figure.png" pageId="30" pageNumber="53">Figs 42</figureCitation>
,
<figureCitation id="13256AE9FFDEFFEABE0EFA0C0FBC0201" box="[597,673,1524,1548]" captionStart="Figures 98103" captionStartId="29.[189,262,1884,1906]" captionTargetBox="[262,1360,393,1733]" captionTargetId="figure-128@29.[189,1436,334,1865]" captionTargetPageId="29" captionText="Figures 98103. Aedeagus of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul in lateral (98), ventral (99), dorsal view (100); apex of aedeagus of S. uncinatus, holotype, Banten, Lebak: Citorek Kidul in lateral (101), ventral (102), dorsal view (103). Abbreviations: al apical lobes, dl dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, tr transverse ridge of median foramen, v velum, vl ventral lobe. Arrows: small, ventral lobe (I); distal enlargement of lateral lobe (II); apicodorsal bridge of apical lobes (III)." figureDoi="http://doi.org/10.5281/zenodo.10724284" httpUri="https://zenodo.org/record/10724284/files/figure.png" pageId="30" pageNumber="53">98, 99</figureCitation>
); apical lobes strongly curved ventrad, tapered towards wide, truncate apex, with subacute, ventral extension (
<figureCitation id="13256AE9FFDEFFEABEFFF9CE0FEA0243" box="[676,759,1590,1614]" captionStart="Figures 4249" captionStartId="17.[190,263,1886,1908]" captionTargetBox="[263,1341,889,1832]" captionTargetId="figure-379@17.[189,1436,851,1867]" captionTargetPageId="17" captionText="Figures 4249. Aedeagus in lateral view of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul (42); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (43); S. uncinatus, holotype,Banten, Lebak:Citorek Kidul (44); S. bipenicillatus, holotype,Sulawesi Tengah, Sigi: Pagana (45); S. bipectenatus, holotype, Sulawesi Tengah, Poso: Lake Poso (46); S. cuspidatus, holotype, Bali, Jembrana: Cekik (47); S. crassipunctatus, holotype, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (48); S. sulawesianus, holotype, Sulawesi Selatan: Engrekang (49)." figureDoi="http://doi.org/10.5281/zenodo.10724262" httpUri="https://zenodo.org/record/10724262/files/figure.png" pageId="30" pageNumber="53">Figs 42</figureCitation>
,
<figureCitation id="13256AE9FFDEFFEABCCCF9AF0DAE0262" box="[151,179,1623,1647]" captionStart="Figures 98103" captionStartId="29.[189,262,1884,1906]" captionTargetBox="[262,1360,393,1733]" captionTargetId="figure-128@29.[189,1436,334,1865]" captionTargetPageId="29" captionText="Figures 98103. Aedeagus of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul in lateral (98), ventral (99), dorsal view (100); apex of aedeagus of S. uncinatus, holotype, Banten, Lebak: Citorek Kidul in lateral (101), ventral (102), dorsal view (103). Abbreviations: al apical lobes, dl dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, tr transverse ridge of median foramen, v velum, vl ventral lobe. Arrows: small, ventral lobe (I); distal enlargement of lateral lobe (II); apicodorsal bridge of apical lobes (III)." figureDoi="http://doi.org/10.5281/zenodo.10724284" httpUri="https://zenodo.org/record/10724284/files/figure.png" pageId="30" pageNumber="53">98</figureCitation>
), in proximal portion widened laterally with stout, lateroapicad pointing tooth, in distal portion convexly widened laterally and strongly tapered towards narrow, short, subtruncate apices (
<figureCitation id="13256AE9FFDEFFEABDE8F9410F2702DC" box="[435,570,1721,1745]" captionStart="Figures 98103" captionStartId="29.[189,262,1884,1906]" captionTargetBox="[262,1360,393,1733]" captionTargetId="figure-128@29.[189,1436,334,1865]" captionTargetPageId="29" captionText="Figures 98103. Aedeagus of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul in lateral (98), ventral (99), dorsal view (100); apex of aedeagus of S. uncinatus, holotype, Banten, Lebak: Citorek Kidul in lateral (101), ventral (102), dorsal view (103). Abbreviations: al apical lobes, dl dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, tr transverse ridge of median foramen, v velum, vl ventral lobe. Arrows: small, ventral lobe (I); distal enlargement of lateral lobe (II); apicodorsal bridge of apical lobes (III)." figureDoi="http://doi.org/10.5281/zenodo.10724284" httpUri="https://zenodo.org/record/10724284/files/figure.png" pageId="30" pageNumber="53">Figs 99, 100</figureCitation>
); dorsal lobe with membranous apex (
<figureCitation id="13256AE9FFDEFFEABD2DF9220CCA02FF" box="[374,471,1754,1778]" captionStart="Figures 98103" captionStartId="29.[189,262,1884,1906]" captionTargetBox="[262,1360,393,1733]" captionTargetId="figure-128@29.[189,1436,334,1865]" captionTargetPageId="29" captionText="Figures 98103. Aedeagus of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul in lateral (98), ventral (99), dorsal view (100); apex of aedeagus of S. uncinatus, holotype, Banten, Lebak: Citorek Kidul in lateral (101), ventral (102), dorsal view (103). Abbreviations: al apical lobes, dl dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, tr transverse ridge of median foramen, v velum, vl ventral lobe. Arrows: small, ventral lobe (I); distal enlargement of lateral lobe (II); apicodorsal bridge of apical lobes (III)." figureDoi="http://doi.org/10.5281/zenodo.10724284" httpUri="https://zenodo.org/record/10724284/files/figure.png" pageId="30" pageNumber="53">Fig. 100</figureCitation>
); lateral lobes marked by numerous, short, spacious setae (
<figureCitation id="13256AE9FFDEFFEABDA2F9030F5A031E" box="[505,583,1787,1811]" captionStart="Figures 98103" captionStartId="29.[189,262,1884,1906]" captionTargetBox="[262,1360,393,1733]" captionTargetId="figure-128@29.[189,1436,334,1865]" captionTargetPageId="29" captionText="Figures 98103. Aedeagus of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul in lateral (98), ventral (99), dorsal view (100); apex of aedeagus of S. uncinatus, holotype, Banten, Lebak: Citorek Kidul in lateral (101), ventral (102), dorsal view (103). Abbreviations: al apical lobes, dl dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, tr transverse ridge of median foramen, v velum, vl ventral lobe. Arrows: small, ventral lobe (I); distal enlargement of lateral lobe (II); apicodorsal bridge of apical lobes (III)." figureDoi="http://doi.org/10.5281/zenodo.10724284" httpUri="https://zenodo.org/record/10724284/files/figure.png" pageId="30" pageNumber="53">Fig. 99</figureCitation>
), slightly convex in ventral view (
<figureCitation id="13256AE9FFDEFFEABD17F8E40C810339" box="[332,412,1820,1844]" captionStart="Figures 98103" captionStartId="29.[189,262,1884,1906]" captionTargetBox="[262,1360,393,1733]" captionTargetId="figure-128@29.[189,1436,334,1865]" captionTargetPageId="29" captionText="Figures 98103. Aedeagus of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul in lateral (98), ventral (99), dorsal view (100); apex of aedeagus of S. uncinatus, holotype, Banten, Lebak: Citorek Kidul in lateral (101), ventral (102), dorsal view (103). Abbreviations: al apical lobes, dl dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, tr transverse ridge of median foramen, v velum, vl ventral lobe. Arrows: small, ventral lobe (I); distal enlargement of lateral lobe (II); apicodorsal bridge of apical lobes (III)." figureDoi="http://doi.org/10.5281/zenodo.10724284" httpUri="https://zenodo.org/record/10724284/files/figure.png" pageId="30" pageNumber="53">Fig. 99</figureCitation>
); ventral lobe slender, narrowed in middle, somewhat widened towards subacute apex and reaching apical lobes (
<figureCitation id="13256AE9FFDEFFEABD9CF8A50F02037B" box="[455,543,1885,1910]" captionStart="Figures 4249" captionStartId="17.[190,263,1886,1908]" captionTargetBox="[263,1341,889,1832]" captionTargetId="figure-379@17.[189,1436,851,1867]" captionTargetPageId="17" captionText="Figures 4249. Aedeagus in lateral view of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul (42); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (43); S. uncinatus, holotype,Banten, Lebak:Citorek Kidul (44); S. bipenicillatus, holotype,Sulawesi Tengah, Sigi: Pagana (45); S. bipectenatus, holotype, Sulawesi Tengah, Poso: Lake Poso (46); S. cuspidatus, holotype, Bali, Jembrana: Cekik (47); S. crassipunctatus, holotype, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (48); S. sulawesianus, holotype, Sulawesi Selatan: Engrekang (49)." figureDoi="http://doi.org/10.5281/zenodo.10724262" httpUri="https://zenodo.org/record/10724262/files/figure.png" pageId="30" pageNumber="53">Figs 42</figureCitation>
,
<figureCitation id="13256AE9FFDEFFEABE75F8A60F56037B" box="[558,587,1886,1910]" captionStart="Figures 98103" captionStartId="29.[189,262,1884,1906]" captionTargetBox="[262,1360,393,1733]" captionTargetId="figure-128@29.[189,1436,334,1865]" captionTargetPageId="29" captionText="Figures 98103. Aedeagus of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul in lateral (98), ventral (99), dorsal view (100); apex of aedeagus of S. uncinatus, holotype, Banten, Lebak: Citorek Kidul in lateral (101), ventral (102), dorsal view (103). Abbreviations: al apical lobes, dl dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, tr transverse ridge of median foramen, v velum, vl ventral lobe. Arrows: small, ventral lobe (I); distal enlargement of lateral lobe (II); apicodorsal bridge of apical lobes (III)." figureDoi="http://doi.org/10.5281/zenodo.10724284" httpUri="https://zenodo.org/record/10724284/files/figure.png" pageId="30" pageNumber="53">98</figureCitation>
); small, narrow lobe projecting ventrally distal of ventral lobe (
<figureCitation id="13256AE9FFDEFFEABEFCF8860FE5039B" box="[679,760,1918,1942]" captionStart="Figures 98103" captionStartId="29.[189,262,1884,1906]" captionTargetBox="[262,1360,393,1733]" captionTargetId="figure-128@29.[189,1436,334,1865]" captionTargetPageId="29" captionText="Figures 98103. Aedeagus of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul in lateral (98), ventral (99), dorsal view (100); apex of aedeagus of S. uncinatus, holotype, Banten, Lebak: Citorek Kidul in lateral (101), ventral (102), dorsal view (103). Abbreviations: al apical lobes, dl dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, tr transverse ridge of median foramen, v velum, vl ventral lobe. Arrows: small, ventral lobe (I); distal enlargement of lateral lobe (II); apicodorsal bridge of apical lobes (III)." figureDoi="http://doi.org/10.5281/zenodo.10724284" httpUri="https://zenodo.org/record/10724284/files/figure.png" pageId="30" pageNumber="53">Fig. 98</figureCitation>
: arrow I); median foramen conspicuously large and wide, longer than half of phallobase, limited distally by weakly sclerotized, straight transverse ridge (
<figureCitation id="13256AE9FFDEFFEAB8F3FEEA09EA0527" box="[1192,1271,274,298]" captionStart="Figures 98103" captionStartId="29.[189,262,1884,1906]" captionTargetBox="[262,1360,393,1733]" captionTargetId="figure-128@29.[189,1436,334,1865]" captionTargetPageId="29" captionText="Figures 98103. Aedeagus of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul in lateral (98), ventral (99), dorsal view (100); apex of aedeagus of S. uncinatus, holotype, Banten, Lebak: Citorek Kidul in lateral (101), ventral (102), dorsal view (103). Abbreviations: al apical lobes, dl dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, tr transverse ridge of median foramen, v velum, vl ventral lobe. Arrows: small, ventral lobe (I); distal enlargement of lateral lobe (II); apicodorsal bridge of apical lobes (III)." figureDoi="http://doi.org/10.5281/zenodo.10724284" httpUri="https://zenodo.org/record/10724284/files/figure.png" pageId="30" pageNumber="53">Fig. 99</figureCitation>
).
</paragraph>
<paragraph id="8BA1766CFFDEFFEABF72FECB09D10561" blockId="30.[786,1400,241,594]" pageId="30" pageNumber="53">
Female: Chamber segment of sperm pump without apophysis (
<figureCitation id="13256AE9FFDEFFEABFD0FEAC0EFB0561" box="[907,998,340,364]" captionStart="Figures 163172" captionStartId="42.[151,224,1839,1861]" captionTargetBox="[225,1322,230,1817]" captionTargetId="figure-168@42.[157,1398,210,1820]" captionTargetPageId="42" captionText="Figures 163172. Sperm pump (163172) and bursa (165, 167, 170, 172) of ♀ of Scopaeus grandis, paratype, Jawa Barat, Sukabumi: Cidahu (163); S.heronifer, paratype, Jawa Barat, Sukabumi:Sirnarasa (164); S. diversilobatus, paratype, Jawa Barat, Sukabumi, Djampang Kulon (165, with lateral gonocoxal plates IX folded laterally; 166); S. cuspilobatus, paratype, Jawa Barat, Sukabumi: Sirnarasa (167, with lateral gonocoxal plates IX, 168); S. velifer, paratype, Banten, Lebak: Citorek Kidul (169); S. elegantulus, Jawa Barat, Sukabumi: Djampang Kulon (170, with lateral gonocoxal plates IX, 171), S. uncinatus, paratype, Banten, Lebak: Citorek Kidul (172). Long scale bar: 163, 164, 166, 168, 169, 171, 172; short scale bar: 165, 167, 170." figureDoi="http://doi.org/10.5281/zenodo.10724315" httpUri="https://zenodo.org/record/10724315/files/figure.png" pageId="30" pageNumber="53">Fig. 169</figureCitation>
); bursa membranous.
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFDEFFEABF72FE8D0EEC05C2" pageId="30" pageNumber="53" type="distribution">
<paragraph id="8BA1766CFFDEFFEABF72FE8D0EEC05C2" blockId="30.[786,1400,241,594]" pageId="30" pageNumber="53">
<emphasis id="B96AAA7EFFDEFFEABF72FE8D0EA20580" bold="true" box="[809,959,373,397]" pageId="30" pageNumber="53">Distribution:</emphasis>
<taxonomicName id="4C1E0DEFFFDEFFEABF90FE8D099E0580" authorityName="Frisch" authorityYear="2022" box="[971,1155,373,397]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="53" phylum="Arthropoda" rank="species" species="velifer">
<emphasis id="B96AAA7EFFDEFFEABF90FE8D099E0580" box="[971,1155,373,397]" italics="true" pageId="30" pageNumber="53">Scopaeus velifer</emphasis>
</taxonomicName>
is known only from the
<typeStatus id="54A5C8CEFFDEFFEABF61FE6E0E7705A3" box="[826,874,406,430]" pageId="30" pageNumber="53">type</typeStatus>
locality at the western edge of Mount Halimun-Salak National Park.
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFDEFFEABF72FE200834065F" pageId="30" pageNumber="53" type="etymology">
<paragraph id="8BA1766CFFDEFFEABF72FE200834065F" blockId="30.[786,1400,241,594]" pageId="30" pageNumber="53">
<emphasis id="B96AAA7EFFDEFFEABF72FE200EB205FD" bold="true" box="[809,943,472,496]" pageId="30" pageNumber="53">Etymology:</emphasis>
The epithet
<taxonomicName id="4C1E0DEFFFDEFFEAB807FE2009BD05FD" authorityName="Frisch" authorityYear="2022" box="[1116,1184,472,496]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="53" phylum="Arthropoda" rank="species" species="velifer">
<emphasis id="B96AAA7EFFDEFFEAB807FE2009BD05FD" box="[1116,1184,472,496]" italics="true" pageId="30" pageNumber="53">velifer</emphasis>
</taxonomicName>
[adjective, Latin, composed of the noun
<emphasis id="B96AAA7EFFDEFFEAB851FE000957061D" box="[1034,1098,504,528]" italics="true" pageId="30" pageNumber="53">velum</emphasis>
(sail) and the verb
<emphasis id="B96AAA7EFFDEFFEAB945FE01084E061C" box="[1310,1363,505,529]" italics="true" pageId="30" pageNumber="53">ferre</emphasis>
(to carry, to bring)] addresses the peculiar, dorsolateral sail of the aedeagus of this species (
<figureCitation id="13256AE9FFDEFFEAB83CFDC209A7065F" box="[1127,1210,570,594]" captionStart="Figures 4249" captionStartId="17.[190,263,1886,1908]" captionTargetBox="[263,1341,889,1832]" captionTargetId="figure-379@17.[189,1436,851,1867]" captionTargetPageId="17" captionText="Figures 4249. Aedeagus in lateral view of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul (42); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (43); S. uncinatus, holotype,Banten, Lebak:Citorek Kidul (44); S. bipenicillatus, holotype,Sulawesi Tengah, Sigi: Pagana (45); S. bipectenatus, holotype, Sulawesi Tengah, Poso: Lake Poso (46); S. cuspidatus, holotype, Bali, Jembrana: Cekik (47); S. crassipunctatus, holotype, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (48); S. sulawesianus, holotype, Sulawesi Selatan: Engrekang (49)." figureDoi="http://doi.org/10.5281/zenodo.10724262" httpUri="https://zenodo.org/record/10724262/files/figure.png" pageId="30" pageNumber="53">Figs 42</figureCitation>
,
<figureCitation id="13256AE9FFDEFFEAB89EFDC20800065F" box="[1221,1309,570,594]" captionStart="Figures 98103" captionStartId="29.[189,262,1884,1906]" captionTargetBox="[262,1360,393,1733]" captionTargetId="figure-128@29.[189,1436,334,1865]" captionTargetPageId="29" captionText="Figures 98103. Aedeagus of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul in lateral (98), ventral (99), dorsal view (100); apex of aedeagus of S. uncinatus, holotype, Banten, Lebak: Citorek Kidul in lateral (101), ventral (102), dorsal view (103). Abbreviations: al apical lobes, dl dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, tr transverse ridge of median foramen, v velum, vl ventral lobe. Arrows: small, ventral lobe (I); distal enlargement of lateral lobe (II); apicodorsal bridge of apical lobes (III)." figureDoi="http://doi.org/10.5281/zenodo.10724284" httpUri="https://zenodo.org/record/10724284/files/figure.png" pageId="30" pageNumber="53">98 100</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>