381 lines
64 KiB
XML
381 lines
64 KiB
XML
<document id="04B6B556A132FA32ED0F6CF1ABB25908" ID-DOI="10.25674/so95iss1id311" ID-ISSN="2509-9523" ID-Zenodo-Dep="10724236" checkinTime="1709147239332" checkinUser="felipe" docAuthor="Frisch, Johannes & Narakusumo, Raden Pramesa" docDate="2022" docId="03B7C77AFFD3FFE7BC8FFE4D0EB60108" docLanguage="en" docName="SoilOrg.95.1.23-73.pdf" docOrigin="Soil Organisms 95 (1)" docSource="http://dx.doi.org/10.25674/so95iss1id311" docStyle="DocumentStyle:EA2ECAC87EC21C07AB805D5B9B4601F2:SoilOrg.2018-.journal_article" docStyleId="EA2ECAC87EC21C07AB805D5B9B4601F2" docStyleName="SoilOrg.2018-.journal_article" docTitle="Scopaeus heronifer Frisch 2022, spec. nov." docType="treatment" docVersion="2" lastPageNumber="42" masterDocId="FF8EBF02FFC0FFF4BC5BFFF80D1D040D" masterDocTitle="Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species" masterLastPageNumber="73" masterPageNumber="23" pageNumber="42" updateTime="1709147743548" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="D85DDDEC1EC0B7EE4B67EA4F0A269885">
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<mods:title id="A87D1B22E3332658356F3830397BF9DE">Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species</mods:title>
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<mods:name id="948A696A02F9E1AA7E752D0A28C6F0F9" type="personal">
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<mods:roleTerm id="0E6B6F2AE25BE190BE22A2B4EFA9845B">Author</mods:roleTerm>
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<mods:namePart id="449A6C38DB2DD9A691EA021F12AF21B8">Frisch, Johannes</mods:namePart>
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<mods:name id="FCA277F6B982AD0647FF5CCEF7C60084" type="personal">
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<mods:roleTerm id="74A454A0F08A83049C4F8BB717F669AD">Author</mods:roleTerm>
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<mods:namePart id="EB07635B65EC65A9A47149CC90980C0D">Narakusumo, Raden Pramesa</mods:namePart>
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<mods:affiliation id="B738FEEA3C3A2EE0587AD53267EEBCA2">MUseUm ZoologicUm Bogoriense, Research Center for Biosystematics and EvolUtion, National Research and Innovation Agency (BRIN), Jl. Raya Jakarta-Bogor KM 46, Cibinong, West Java, 16911, Indonesia</mods:affiliation>
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<mods:title id="34C025B7C3358B51C42749FC0E8488FD">Soil Organisms</mods:title>
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<mods:part id="8ADD7BDEE17324657CB306DB4984B6F0">
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<mods:date id="62515F1626694BE6DDA6DC77DC1DBA57">2022</mods:date>
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<mods:number id="92AA39E5D0C0F71AC59C55F3178E3FB3">2022-04-01</mods:number>
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<mods:number id="5294C18C8F5879E002774C15E26B2D20">95</mods:number>
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<mods:url id="CD2044407F9847FD8C87F3CCC540AAE3">http://dx.doi.org/10.25674/so95iss1id311</mods:url>
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<mods:classification id="B290243B6215419CBCF21F6843BF5B3B">journal article</mods:classification>
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<mods:identifier id="7CC4AA330ACA57A5D4F91C3A038A6904" type="DOI">10.25674/so95iss1id311</mods:identifier>
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<treatment id="03B7C77AFFD3FFE7BC8FFE4D0EB60108" ID-DOI="http://doi.org/10.5281/zenodo.10724300" ID-Zenodo-Dep="10724300" LSID="urn:lsid:plazi:treatment:03B7C77AFFD3FFE7BC8FFE4D0EB60108" httpUri="http://treatment.plazi.org/id/03B7C77AFFD3FFE7BC8FFE4D0EB60108" lastPageNumber="42" pageId="19" pageNumber="42">
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<subSubSection id="C30425E7FFD3FFE7BC8FFE4D0F8605DD" box="[212,667,436,464]" pageId="19" pageNumber="42" type="nomenclature">
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<paragraph id="8BA1766CFFD3FFE7BC8FFE4D0F8605DD" blockId="19.[212,667,436,497]" box="[212,667,436,464]" pageId="19" pageNumber="42">
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<heading id="D0E9C100FFD3FFE7BC8FFE4D0F8605DD" box="[212,667,436,464]" fontSize="11" level="2" pageId="19" pageNumber="42" reason="5">
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<taxonomicName id="4C1E0DEFFFD3FFE7BC8FFE4D0F0405C2" authority="Frisch" authorityName="Frisch" authorityYear="2022" box="[212,537,436,463]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="heronifer" status="sp. nov.">
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<emphasis id="B96AAA7EFFD3FFE7BC8FFE4D0F0405C2" bold="true" box="[212,537,436,463]" pageId="19" pageNumber="42">
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<emphasis id="B96AAA7EFFD3FFE7BC8FFE4D0CDD05C2" bold="true" box="[212,448,436,463]" italics="true" pageId="19" pageNumber="42">Scopaeus heronifer</emphasis>
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Frisch
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</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2591705FFD3FFE7BE7CFE4E0F8605DD" box="[551,667,438,464]" pageId="19" pageNumber="42" rank="species">spec. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30425E7FFD3FFE7BC8FFE2E0F2E05FC" box="[212,563,470,497]" pageId="19" pageNumber="42" type="description">
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<paragraph id="8BA1766CFFD3FFE7BC8FFE2E0F2E05FC" blockId="19.[212,667,436,497]" box="[212,563,470,497]" pageId="19" pageNumber="42">
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(
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<figureCitation id="13256AE9FFD3FFE7BC87FE2E0C2905FD" box="[220,308,470,496]" captionStart="Figures 14–16" captionStartId="9.[189,262,1930,1952]" captionTargetBox="[262,1384,910,1880]" captionTargetId="figure-57@9.[189,1436,894,1911]" captionTargetPageId="9" captionText="Figures 14–16. Habitus of Scopaeus heronifer, paratype, Jawa Barat: Mt Salak (14); S. diversilobatus, paratype, Jawa Barat, Sukabumi: Djampang Kulon (15); S. cuspilobatus, paratype, Jawa Barat, Sukabumi: Sirnarasa (16)." figureDoi="http://doi.org/10.5281/zenodo.10724246" httpUri="https://zenodo.org/record/10724246/files/figure.png" pageId="19" pageNumber="42">Figs 14</figureCitation>
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,
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<figureCitation id="13256AE9FFD3FFE7BD1BFE2E0C4005FD" box="[320,349,470,496]" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="19" pageNumber="42">41</figureCitation>
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,
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<figureCitation id="13256AE9FFD3FFE7BD32FE2E0CA605FD" box="[361,443,470,496]" captionStart="Figures 86–91" captionStartId="27.[189,262,1863,1885]" captionTargetBox="[261,1365,582,1806]" captionTargetId="figure-268@27.[189,1436,563,1844]" captionTargetPageId="27" captionText="Figures 86–91. Apex of aedeagus in lateral (86, 89), ventral (87, 90), dorsal view (88, 91) of Scopaeus heronifer, holotype, Banten, Lebak: Majasari (86– 88); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (89–91). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: distoventral incision of dextral apical lobe (I); sinistral, apical tooth of flagellum (II); distal emargination of dorsal lobe (III)." figureDoi="http://doi.org/10.5281/zenodo.10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="19" pageNumber="42">86– 88</figureCitation>
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,
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<figureCitation id="13256AE9FFD3FFE7BD9CFE2E0CE905FD" box="[455,500,470,496]" captionStart="Figures 119–133" captionStartId="38.[151,224,1796,1818]" captionTargetBox="[224,1369,297,1686]" captionTargetId="figure-201@38.[151,1398,236,1777]" captionTargetPageId="38" captionText="Figures 119–133. Abdominal sternite VII (126) and abdominal sternite VIII (119–125, 127– 133) of ♂ of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (119); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (120); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (121); S. jacobsoni, Jawa Barat: Mt Salak (122); S. mixtus, Malaysia, Sabah: Lokan (123); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (124); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (125); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (126, 127); S. spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (128); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi:Cidahu (129); S. pulcher, holotype, Jawa Barat, Sukabumi:Cikaniki (130); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (131); S. heronifer, holotype, Banten, Lebak: Majasari (132); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (133)." figureDoi="http://doi.org/10.5281/zenodo.10724304" httpUri="https://zenodo.org/record/10724304/files/figure.png" pageId="19" pageNumber="42">132</figureCitation>
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,
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<figureCitation id="13256AE9FFD3FFE7BE5BFE2E0F3605FC" box="[512,555,470,497]" captionStart="Figures 163–172" captionStartId="42.[151,224,1839,1861]" captionTargetBox="[225,1322,230,1817]" captionTargetId="figure-168@42.[157,1398,210,1820]" captionTargetPageId="42" captionText="Figures 163–172. Sperm pump (163–172) and bursa (165, 167, 170, 172) of ♀ of Scopaeus grandis, paratype, Jawa Barat, Sukabumi: Cidahu (163); S.heronifer, paratype, Jawa Barat, Sukabumi:Sirnarasa (164); S. diversilobatus, paratype, Jawa Barat, Sukabumi, Djampang Kulon (165, with lateral gonocoxal plates IX folded laterally; 166); S. cuspilobatus, paratype, Jawa Barat, Sukabumi: Sirnarasa (167, with lateral gonocoxal plates IX, 168); S. velifer, paratype, Banten, Lebak: Citorek Kidul (169); S. elegantulus, Jawa Barat, Sukabumi: Djampang Kulon (170, with lateral gonocoxal plates IX, 171), S. uncinatus, paratype, Banten, Lebak: Citorek Kidul (172). Long scale bar: 163, 164, 166, 168, 169, 171, 172; short scale bar: 165, 167, 170." figureDoi="http://doi.org/10.5281/zenodo.10724315" httpUri="https://zenodo.org/record/10724315/files/figure.png" pageId="19" pageNumber="42">164</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C30425E7FFD3FFE7BC8FFDE20CF507B1" pageId="19" pageNumber="42" type="materials_examined">
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<paragraph id="8BA1766CFFD3FFE7BC8FFDE20CF507B1" blockId="19.[189,800,537,1976]" pageId="19" pageNumber="42">
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<emphasis id="B96AAA7EFFD3FFE7BC8FFDE20C81063C" bold="true" box="[212,412,537,562]" pageId="19" pageNumber="42">Type specimens:</emphasis>
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<materialsCitation id="3B767C31FFD3FFE7BDF0FDE10F66067E" collectingDate="2016-05-26" collectionCode="MZB" collectorName="Frisch" country="Indonesia" elevation="460" latitude="-6.6255555" location="Majasari" longLatPrecision="21" longitude="106.398056" municipality="Lebak" pageId="19" pageNumber="42" specimenCount="1" specimenCount-male="1" stateProvince="Banten" typeStatus="holotype">
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<collectingCountry id="F30936FCFFD3FFE7BDF0FDE10F07063C" box="[427,538,537,561]" name="Indonesia" pageId="19" pageNumber="42">Indonesia</collectingCountry>
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:
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<typeStatus id="54A5C8CEFFD3FFE7BE76FDE10F8E063C" box="[557,659,537,561]" pageId="19" pageNumber="42" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D18BDE5FFD3FFE7BEF9FDE20FAB063C" box="[674,694,538,561]" count="1" pageId="19" pageNumber="42" type="male">♂</specimenCount>
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,
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||
<collectingRegion id="49DAB88EFFD3FFE7BE91FDE20E06063C" box="[714,795,538,561]" country="Indonesia" name="Banten" pageId="19" pageNumber="42">Banten</collectingRegion>
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,
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<collectingMunicipality id="6BC5EC16FFD3FFE7BCE6FDC20C1B065F" box="[189,262,570,594]" pageId="19" pageNumber="42">Lebak</collectingMunicipality>
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: NE
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<location id="8EC120B7FFD3FFE7BD3BFDC20CDC065F" LSID="urn:lsid:plazi:treatment:03B7C77AFFD3FFE7BC8FFE4D0EB60108:8EC120B7FFD3FFE7BD3BFDC20CDC065F" box="[352,449,570,594]" country="Indonesia" latitude="-6.6255555" longLatPrecision="21" longitude="106.398056" municipality="Lebak" name="Majasari" pageId="19" pageNumber="42" stateProvince="Banten">Majasari</location>
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(
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<geoCoordinate id="EE2A10ABFFD3FFE7BDB9FDC20F78065F" box="[482,613,570,594]" degrees="06" direction="south" minutes="37" orientation="latitude" pageId="19" pageNumber="42" precision="15" seconds="32" value="-6.6255555">06°37’32’’S</geoCoordinate>
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,
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<geoCoordinate id="EE2A10ABFFD3FFE7BED9FDC20E0E065F" box="[642,787,570,594]" degrees="106" direction="east" minutes="23" orientation="longitude" pageId="19" pageNumber="42" precision="15" seconds="53" value="106.398056">106°23’53’’E</geoCoordinate>
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),
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<quantity id="4CE6DB89FFD3FFE7BCE6FDA30C10067E" box="[189,269,603,627]" metricMagnitude="2" metricUnit="m" metricValue="4.6" pageId="19" pageNumber="42" unit="m" value="460.0">
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<elevation id="0033915FFFD3FFE7BCE6FDA30C10067E" box="[189,269,603,627]" metricMagnitude="2" metricUnit="m" metricValue="4.6" pageId="19" pageNumber="42" unit="m" value="460.0">460 m</elevation>
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</quantity>
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,
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<date id="FFA050ACFFD3FFE7BD7AFDA30C91067E" box="[289,396,603,627]" pageId="19" pageNumber="42" value="2016-05-26">
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||
<collectingDate id="EFE4A944FFD3FFE7BD7AFDA30C91067E" box="[289,396,603,627]" pageId="19" pageNumber="42" value="2016-05-26">26.5.2016</collectingDate>
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</date>
|
||
, leg.
|
||
<collectorName id="26EB13BAFFD3FFE7BD8CFDA30F06067E" box="[471,539,603,627]" pageId="19" pageNumber="42">Frisch</collectorName>
|
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(
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||
<collectionCode id="ED0FEEA9FFD3FFE7BE68FDA30F6E067E" box="[563,627,603,627]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="19" pageNumber="42" type="Museum">MZB</collectionCode>
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)
|
||
</materialsCitation>
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||
.
|
||
<materialsCitation id="3B767C31FFD3FFE7BED5FDA40CDC0699" country="Indonesia" location="Jawa Barat" pageId="19" pageNumber="42" specimenCount="5" stateProvince="Jawa Barat" typeStatus="paratype">
|
||
<typeStatus id="54A5C8CEFFD3FFE7BED5FDA40FE60679" box="[654,763,604,628]" pageId="19" pageNumber="42" type="paratype">Paratypes</typeStatus>
|
||
(
|
||
<specimenCount id="9D18BDE5FFD3FFE7BF48FDA30C290699" count="5" pageId="19" pageNumber="42" type="generic">5 specimens</specimenCount>
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||
):
|
||
<collectingRegion id="49DAB88EFFD3FFE7BD1EFD850CDC0699" box="[325,449,637,660]" country="Indonesia" name="Jawa Barat" pageId="19" pageNumber="42">Jawa Barat</collectingRegion>
|
||
</materialsCitation>
|
||
:
|
||
<materialsCitation id="3B767C31FFD3FFE7BD91FD840C1006DB" collectingDate="2015-09-13" collectionCode="MZB" collectorName="Frisch" country="Indonesia" elevation="640" latitude="-6.665278" location="Mt Salak" longLatPrecision="21" longitude="106.76" municipality="Bogor" pageId="19" pageNumber="42" specimenCount="1" specimenCount-female="1" stateProvince="Jawa Barat" typeStatus="paratype">
|
||
<specimenCount id="9D18BDE5FFD3FFE7BD91FD840CEC0699" box="[458,497,636,660]" count="1" pageId="19" pageNumber="42" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BC5EC16FFD3FFE7BDA6FD850F5F0698" box="[509,578,637,661]" pageId="19" pageNumber="42">Bogor</collectingMunicipality>
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||
, NO-slope
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||
<location id="8EC120B7FFD3FFE7BEE6FD850E020699" LSID="urn:lsid:plazi:treatment:03B7C77AFFD3FFE7BC8FFE4D0EB60108:8EC120B7FFD3FFE7BEE6FD850E020699" box="[701,799,636,660]" country="Indonesia" latitude="-6.665278" longLatPrecision="21" longitude="106.76" municipality="Bogor" name="Mt Salak" pageId="19" pageNumber="42" stateProvince="Jawa Barat">Mt Salak</location>
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(
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<geoCoordinate id="EE2A10ABFFD3FFE7BC9EFD650C5506B8" box="[197,328,669,693]" degrees="06" direction="south" minutes="39" orientation="latitude" pageId="19" pageNumber="42" precision="15" seconds="55" value="-6.665278">06°39’55’’S</geoCoordinate>
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,
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<geoCoordinate id="EE2A10ABFFD3FFE7BD0BFD650CFD06B8" box="[336,480,669,693]" degrees="106" direction="east" minutes="45" orientation="longitude" pageId="19" pageNumber="42" precision="15" seconds="36" value="106.76">106°45’36’’E</geoCoordinate>
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||
),
|
||
<quantity id="4CE6DB89FFD3FFE7BDAAFD650F2B06B9" box="[497,566,669,693]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="19" pageNumber="42" unit="m" value="640.0">
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<elevation id="0033915FFFD3FFE7BDAAFD650F2B06B9" box="[497,566,669,693]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="19" pageNumber="42" unit="m" value="640.0">640 m</elevation>
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</quantity>
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,
|
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<date id="FFA050ACFFD3FFE7BE65FD650FBA06B8" box="[574,679,669,693]" pageId="19" pageNumber="42" value="2015-09-13">
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<collectingDate id="EFE4A944FFD3FFE7BE65FD650FBA06B8" box="[574,679,669,693]" pageId="19" pageNumber="42" value="2015-09-13">13.9.2015</collectingDate>
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</date>
|
||
, leg.
|
||
<collectorName id="26EB13BAFFD3FFE7BE80FD650E0206B8" box="[731,799,669,693]" pageId="19" pageNumber="42">Frisch</collectorName>
|
||
(
|
||
<collectionCode id="ED0FEEA9FFD3FFE7BC9EFD460C1806DB" box="[197,261,702,726]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="19" pageNumber="42" type="Museum">MZB</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B767C31FFD3FFE7BD44FD460F430715" collectingDate="2015-09-19" collectionCode="MZB" collectorName="Frisch" country="Indonesia" county="Sukabumi" elevation="870" latitude="-6.755833" location="Cisalimas River" longLatPrecision="21" longitude="106.56056" municipality="Cisalimas" pageId="19" pageNumber="42" specimenCount="1" specimenCount-female="1" stateProvince="Jawa Barat" typeStatus="paratype">
|
||
<specimenCount id="9D18BDE5FFD3FFE7BD44FD460C5006D8" box="[287,333,702,726]" count="1" pageId="19" pageNumber="42" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectingCounty id="62C00EE0FFD3FFE7BD3AFD460CC906DB" box="[353,468,702,726]" pageId="19" pageNumber="42">Sukabumi</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BC5EC16FFD3FFE7BDBDFD460F4A06DB" box="[486,599,702,726]" pageId="19" pageNumber="42">Cisalimas</collectingMunicipality>
|
||
:
|
||
<location id="8EC120B7FFD3FFE7BE33FD460E0206DB" LSID="urn:lsid:plazi:treatment:03B7C77AFFD3FFE7BC8FFE4D0EB60108:8EC120B7FFD3FFE7BE33FD460E0206DB" box="[616,799,702,726]" country="Indonesia" county="Sukabumi" latitude="-6.755833" longLatPrecision="21" longitude="106.56056" municipality="Cisalimas" name="Cisalimas River" pageId="19" pageNumber="42" stateProvince="Jawa Barat">Cisalimas River</location>
|
||
(E-slope
|
||
<location id="8EC120B7FFD3FFE7BD76FD270CD906FA" LSID="urn:lsid:plazi:treatment:03B7C77AFFD3FFE7BC8FFE4D0EB60108:8EC120B7FFD3FFE7BD76FD270CD906FA" box="[301,452,735,759]" country="Indonesia" county="Sukabumi" latitude="-6.755833" longLatPrecision="21" longitude="106.56056" municipality="Cisalimas" name="Mt Halimun" pageId="19" pageNumber="42" stateProvince="Jawa Barat">Mt Halimun</location>
|
||
), (
|
||
<geoCoordinate id="EE2A10ABFFD3FFE7BDB1FD270F7606FA" box="[490,619,735,759]" degrees="06" direction="south" minutes="45" orientation="latitude" pageId="19" pageNumber="42" precision="15" seconds="21" value="-6.755833">06°45’21’’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2A10ABFFD3FFE7BED9FD270E0E06FA" box="[642,787,735,759]" degrees="106" direction="east" minutes="33" orientation="longitude" pageId="19" pageNumber="42" precision="15" seconds="38" value="106.56056">106°33’38’’E</geoCoordinate>
|
||
),
|
||
<quantity id="4CE6DB89FFD3FFE7BCE6FCF80C1A071A" box="[189,263,768,792]" metricMagnitude="2" metricUnit="m" metricValue="8.7" pageId="19" pageNumber="42" unit="m" value="870.0">
|
||
<elevation id="0033915FFFD3FFE7BCE6FCF80C1A071A" box="[189,263,768,792]" metricMagnitude="2" metricUnit="m" metricValue="8.7" pageId="19" pageNumber="42" unit="m" value="870.0">870 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFA050ACFFD3FFE7BD4DFD070C63071A" box="[278,382,767,791]" pageId="19" pageNumber="42" value="2015-09-19">
|
||
<collectingDate id="EFE4A944FFD3FFE7BD4DFD070C63071A" box="[278,382,767,791]" pageId="19" pageNumber="42" value="2015-09-19">19.9.2015</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26EB13BAFFD3FFE7BDE4FCF80F1E0715" box="[447,515,768,792]" pageId="19" pageNumber="42">Frisch</collectorName>
|
||
(
|
||
<collectionCode id="ED0FEEA9FFD3FFE7BE4DFCF80F4B0715" box="[534,598,768,792]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="19" pageNumber="42" type="Museum">MZB</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B767C31FFD3FFE7BE36FCF80E060754" collectingDate="2015-09-20" collectionCode="MFNB" collectorName="Frisch" country="Indonesia" county="Sukabumi" elevation="680" latitude="-6.860833" location="Cisarua" longLatPrecision="21" longitude="106.513336" municipality="Sirnarasa" pageId="19" pageNumber="42" specimenCount="1" specimenCount-male="1" stateProvince="Jawa Barat" typeStatus="paratype">
|
||
<specimenCount id="9D18BDE5FFD3FFE7BE36FCF80F85071A" box="[621,664,768,792]" count="1" pageId="19" pageNumber="42" type="male">1 ♂</specimenCount>
|
||
,
|
||
<collectingCounty id="62C00EE0FFD3FFE7BEF3FCF80E060715" box="[680,795,768,792]" pageId="19" pageNumber="42">Sukabumi</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BC5EC16FFD3FFE7BCE6FCD80C340735" box="[189,297,800,824]" pageId="19" pageNumber="42">Sirnarasa</collectingMunicipality>
|
||
,
|
||
<location id="8EC120B7FFD3FFE7BD1DFCD80CBD0735" LSID="urn:lsid:plazi:treatment:03B7C77AFFD3FFE7BC8FFE4D0EB60108:8EC120B7FFD3FFE7BD1DFCD80CBD0735" box="[326,416,800,824]" country="Indonesia" county="Sukabumi" latitude="-6.860833" longLatPrecision="21" longitude="106.513336" municipality="Sirnarasa" name="Cisarua" pageId="19" pageNumber="42" stateProvince="Jawa Barat">Cisarua</location>
|
||
:
|
||
<location id="8EC120B7FFD3FFE7BDE6FCD80F650735" LSID="urn:lsid:plazi:treatment:03B7C77AFFD3FFE7BC8FFE4D0EB60108:8EC120B7FFD3FFE7BDE6FCD80F650735" box="[445,632,800,824]" country="Indonesia" county="Sukabumi" latitude="-6.860833" longLatPrecision="21" longitude="106.513336" municipality="Sirnarasa" name="Ciawitali River" pageId="19" pageNumber="42" stateProvince="Jawa Barat">Ciawitali River</location>
|
||
(
|
||
<geoCoordinate id="EE2A10ABFFD3FFE7BEC1FCD80E060735" box="[666,795,800,824]" degrees="06" direction="south" minutes="51" orientation="latitude" pageId="19" pageNumber="42" precision="15" seconds="39" value="-6.860833">06°51’39’’S</geoCoordinate>
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||
,
|
||
<geoCoordinate id="EE2A10ABFFD3FFE7BCE6FCB90C520754" box="[189,335,833,857]" degrees="106" direction="east" minutes="30" orientation="longitude" pageId="19" pageNumber="42" precision="15" seconds="48" value="106.513336">106°30’48’’E</geoCoordinate>
|
||
),
|
||
<quantity id="4CE6DB89FFD3FFE7BD3EFCB90CAD0754" box="[357,432,833,857]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="19" pageNumber="42" unit="m" value="680.0">
|
||
<elevation id="0033915FFFD3FFE7BD3EFCB90CAD0754" box="[357,432,833,857]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="19" pageNumber="42" unit="m" value="680.0">680 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFA050ACFFD3FFE7BDE4FCB90F340754" box="[447,553,833,857]" pageId="19" pageNumber="42" value="2015-09-20">
|
||
<collectingDate id="EFE4A944FFD3FFE7BDE4FCB90F340754" box="[447,553,833,857]" pageId="19" pageNumber="42" value="2015-09-20">20.9.2015</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26EB13BAFFD3FFE7BE32FCB90FB00754" box="[617,685,833,857]" pageId="19" pageNumber="42">Frisch</collectorName>
|
||
(
|
||
<collectionCode id="ED0FEEA9FFD3FFE7BE9BFCB90E0E0754" box="[704,787,833,857]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="19" pageNumber="42">MFNB</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B767C31FFD3FFE7BCE6FC9A0CF907B1" collectingDate="2015-09-21" collectionCode="MFNB, MZB" collectorName="Frisch" country="Indonesia" county="Sukabumi" elevation="670" latitude="-6.8588886" location="Cimaja River" longLatPrecision="21" longitude="106.51833" municipality="Sirnarasa" pageId="19" pageNumber="42" specimenCount="2" specimenCount-female="2" stateProvince="Jawa Barat" typeStatus="paratype">
|
||
<specimenCount id="9D18BDE5FFD3FFE7BCE6FC9A0DF40777" box="[189,233,866,890]" count="2" pageId="19" pageNumber="42" type="female">2 ♀</specimenCount>
|
||
,
|
||
<collectingCounty id="62C00EE0FFD3FFE7BCA0FC9A0C730777" box="[251,366,866,890]" pageId="19" pageNumber="42">Sukabumi</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BC5EC16FFD3FFE7BD26FC9A0CF70777" box="[381,490,866,890]" pageId="19" pageNumber="42">Sirnarasa</collectingMunicipality>
|
||
:
|
||
<location id="8EC120B7FFD3FFE7BDA2FC9A0F8D0777" LSID="urn:lsid:plazi:treatment:03B7C77AFFD3FFE7BC8FFE4D0EB60108:8EC120B7FFD3FFE7BDA2FC9A0F8D0777" box="[505,656,866,890]" country="Indonesia" county="Sukabumi" latitude="-6.8588886" longLatPrecision="21" longitude="106.51833" municipality="Sirnarasa" name="Cimaja River" pageId="19" pageNumber="42" stateProvince="Jawa Barat">Cimaja River</location>
|
||
(S-slope
|
||
<location id="8EC120B7FFD3FFE7BF5BFC9B0C3C0796" LSID="urn:lsid:plazi:treatment:03B7C77AFFD3FFE7BC8FFE4D0EB60108:8EC120B7FFD3FFE7BF5BFC9B0C3C0796" country="Indonesia" county="Sukabumi" latitude="-6.8588886" longLatPrecision="21" longitude="106.51833" municipality="Sirnarasa" name="Mt Halimun" pageId="19" pageNumber="42" stateProvince="Jawa Barat">Mt Halimun</location>
|
||
), (
|
||
<geoCoordinate id="EE2A10ABFFD3FFE7BD61FC7B0CA60796" box="[314,443,899,923]" degrees="06" direction="south" minutes="51" orientation="latitude" pageId="19" pageNumber="42" precision="15" seconds="32" value="-6.8588886">06°51’32’’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2A10ABFFD3FFE7BD9FFC7B0F490796" box="[452,596,899,923]" degrees="106" direction="east" minutes="31" orientation="longitude" pageId="19" pageNumber="42" precision="15" seconds="06" value="106.51833">106°31’06’’E</geoCoordinate>
|
||
),
|
||
<quantity id="4CE6DB89FFD3FFE7BE3DFC7B0FB70796" box="[614,682,899,923]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="19" pageNumber="42" unit="m" value="670.0">
|
||
<elevation id="0033915FFFD3FFE7BE3DFC7B0FB70796" box="[614,682,899,923]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="19" pageNumber="42" unit="m" value="670.0">670 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFA050ACFFD3FFE7BEE8FC7B0E060796" box="[691,795,899,923]" pageId="19" pageNumber="42" value="2015-09-21">
|
||
<collectingDate id="EFE4A944FFD3FFE7BEE8FC7B0E060796" box="[691,795,899,923]" pageId="19" pageNumber="42" value="2015-09-21">21.9.2015</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26EB13BAFFD3FFE7BCB7FC5C0C2D07B1" box="[236,304,932,956]" pageId="19" pageNumber="42">Frisch</collectorName>
|
||
(
|
||
<collectionCode id="ED0FEEA9FFD3FFE7BD64FC5C0C8F07B1" box="[319,402,932,956]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="19" pageNumber="42">MFNB</collectionCode>
|
||
,
|
||
<collectionCode id="ED0FEEA9FFD3FFE7BDC5FC5C0CC007B1" box="[414,477,932,956]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="19" pageNumber="42" type="Museum">MZB</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30425E7FFD3FFE7BC8FFC3D0EB007F3" pageId="19" pageNumber="42" type="description">
|
||
<paragraph id="8BA1766CFFD3FFE7BC8FFC3D0C8E0109" blockId="19.[189,800,537,1976]" pageId="19" pageNumber="42">
|
||
<emphasis id="B96AAA7EFFD3FFE7BC8FFC3D0C7F07D0" bold="true" box="[212,354,965,989]" pageId="19" pageNumber="42">Description:</emphasis>
|
||
Habitus and coloring as in
|
||
<figureCitation id="13256AE9FFD3FFE7BECBFC3D0FC107D0" box="[656,732,965,989]" captionStart="Figures 14–16" captionStartId="9.[189,262,1930,1952]" captionTargetBox="[262,1384,910,1880]" captionTargetId="figure-57@9.[189,1436,894,1911]" captionTargetPageId="9" captionText="Figures 14–16. Habitus of Scopaeus heronifer, paratype, Jawa Barat: Mt Salak (14); S. diversilobatus, paratype, Jawa Barat, Sukabumi: Djampang Kulon (15); S. cuspilobatus, paratype, Jawa Barat, Sukabumi: Sirnarasa (16)." figureDoi="http://doi.org/10.5281/zenodo.10724246" httpUri="https://zenodo.org/record/10724246/files/figure.png" pageId="19" pageNumber="42">Fig. 14</figureCitation>
|
||
. Head subquadrate. Penultimate antennal segment elongate. Mesotibia slender. Body surface with fine, dense, setose punctation, without microreticulation, shiny. Pubescence of body surface short, decumbent, without conspicuous macrosetae. Body medium brown with head and elytra somewhat darker than pronotum; antennae light brown with darker median segments; maxillary palpi and legs yellow-brown. Total body length
|
||
<quantity id="4CE6DB89FFD3FFE7BE72FB340FAE00EE" box="[553,691,1228,1252]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.6" metricValueMin="2.9" pageId="19" pageNumber="42" unit="mm" value="3.25" valueMax="3.6" valueMin="2.9">2.9 – 3.6 mm</quantity>
|
||
; forebody length
|
||
<quantity id="4CE6DB89FFD3FFE7BD53FB150C920109" box="[264,399,1261,1285]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="19" pageNumber="42" unit="mm" value="1.75" valueMax="1.9" valueMin="1.6">1.6 – 1.9 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA1766CFFD3FFE7BC8FFAF6096605A3" blockId="19.[189,800,537,1976]" lastBlockId="19.[824,1437,241,1285]" pageId="19" pageNumber="42">
|
||
Male: Abdominal sternite VII with unmodified posterior margin. Abdominal sternite VIII with short, convex emargination in posterior seventh (
|
||
<figureCitation id="13256AE9FFD3FFE7BEE9FAB70E090165" box="[690,788,1359,1384]" captionStart="Figures 119–133" captionStartId="38.[151,224,1796,1818]" captionTargetBox="[224,1369,297,1686]" captionTargetId="figure-201@38.[151,1398,236,1777]" captionTargetPageId="38" captionText="Figures 119–133. Abdominal sternite VII (126) and abdominal sternite VIII (119–125, 127– 133) of ♂ of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (119); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (120); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (121); S. jacobsoni, Jawa Barat: Mt Salak (122); S. mixtus, Malaysia, Sabah: Lokan (123); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (124); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (125); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (126, 127); S. spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (128); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi:Cidahu (129); S. pulcher, holotype, Jawa Barat, Sukabumi:Cikaniki (130); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (131); S. heronifer, holotype, Banten, Lebak: Majasari (132); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (133)." figureDoi="http://doi.org/10.5281/zenodo.10724304" httpUri="https://zenodo.org/record/10724304/files/figure.png" pageId="19" pageNumber="42">Fig. 132</figureCitation>
|
||
). Aedeagus (
|
||
<figureCitation id="13256AE9FFD3FFE7BD65FA880C8A0185" box="[318,407,1392,1416]" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="19" pageNumber="42">Figs 41</figureCitation>
|
||
,
|
||
<figureCitation id="13256AE9FFD3FFE7BDF3FA880CEE0185" box="[424,499,1392,1416]" captionStart="Figures 86–91" captionStartId="27.[189,262,1863,1885]" captionTargetBox="[261,1365,582,1806]" captionTargetId="figure-268@27.[189,1436,563,1844]" captionTargetPageId="27" captionText="Figures 86–91. Apex of aedeagus in lateral (86, 89), ventral (87, 90), dorsal view (88, 91) of Scopaeus heronifer, holotype, Banten, Lebak: Majasari (86– 88); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (89–91). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: distoventral incision of dextral apical lobe (I); sinistral, apical tooth of flagellum (II); distal emargination of dorsal lobe (III)." figureDoi="http://doi.org/10.5281/zenodo.10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="19" pageNumber="42">86 – 88</figureCitation>
|
||
) about
|
||
<quantity id="4CE6DB89FFD3FFE7BE09FA880FB90185" box="[594,676,1392,1416]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="19" pageNumber="42" unit="mm" value="0.7">0.7 mm</quantity>
|
||
long with distal lobes asymmetrical and notably bent dextrad; dextral apical lobe slender in lateral view with distoventral incision (
|
||
<figureCitation id="13256AE9FFD3FFE7BDE7FA2B0F0901E6" box="[444,532,1491,1515]" captionStart="Figures 86–91" captionStartId="27.[189,262,1863,1885]" captionTargetBox="[261,1365,582,1806]" captionTargetId="figure-268@27.[189,1436,563,1844]" captionTargetPageId="27" captionText="Figures 86–91. Apex of aedeagus in lateral (86, 89), ventral (87, 90), dorsal view (88, 91) of Scopaeus heronifer, holotype, Banten, Lebak: Majasari (86– 88); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (89–91). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: distoventral incision of dextral apical lobe (I); sinistral, apical tooth of flagellum (II); distal emargination of dorsal lobe (III)." figureDoi="http://doi.org/10.5281/zenodo.10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="19" pageNumber="42">Fig. 86</figureCitation>
|
||
: arrow I) and tapered towards dorsad curved end (
|
||
<figureCitation id="13256AE9FFD3FFE7BDADFA0C0F540201" box="[502,585,1524,1548]" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="19" pageNumber="42">Figs 41</figureCitation>
|
||
,
|
||
<figureCitation id="13256AE9FFD3FFE7BE0FFA0C0F6F0201" box="[596,626,1524,1548]" captionStart="Figures 86–91" captionStartId="27.[189,262,1863,1885]" captionTargetBox="[261,1365,582,1806]" captionTargetId="figure-268@27.[189,1436,563,1844]" captionTargetPageId="27" captionText="Figures 86–91. Apex of aedeagus in lateral (86, 89), ventral (87, 90), dorsal view (88, 91) of Scopaeus heronifer, holotype, Banten, Lebak: Majasari (86– 88); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (89–91). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: distoventral incision of dextral apical lobe (I); sinistral, apical tooth of flagellum (II); distal emargination of dorsal lobe (III)." figureDoi="http://doi.org/10.5281/zenodo.10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="19" pageNumber="42">86</figureCitation>
|
||
), in ventral and dorsal view convex laterally with retracted, narrow end (
|
||
<figureCitation id="13256AE9FFD3FFE7BC9EF9CE0C5A0243" box="[197,327,1590,1614]" captionStart="Figures 86–91" captionStartId="27.[189,262,1863,1885]" captionTargetBox="[261,1365,582,1806]" captionTargetId="figure-268@27.[189,1436,563,1844]" captionTargetPageId="27" captionText="Figures 86–91. Apex of aedeagus in lateral (86, 89), ventral (87, 90), dorsal view (88, 91) of Scopaeus heronifer, holotype, Banten, Lebak: Majasari (86– 88); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (89–91). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: distoventral incision of dextral apical lobe (I); sinistral, apical tooth of flagellum (II); distal emargination of dorsal lobe (III)." figureDoi="http://doi.org/10.5281/zenodo.10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="19" pageNumber="42">Figs 87, 88</figureCitation>
|
||
); sinistral apical lobe weakly sclerotized, evenly curved towards subacute apex (
|
||
<figureCitation id="13256AE9FFD3FFE7BED1F9AE0E0F0262" box="[650,786,1622,1647]" captionStart="Figures 86–91" captionStartId="27.[189,262,1863,1885]" captionTargetBox="[261,1365,582,1806]" captionTargetId="figure-268@27.[189,1436,563,1844]" captionTargetPageId="27" captionText="Figures 86–91. Apex of aedeagus in lateral (86, 89), ventral (87, 90), dorsal view (88, 91) of Scopaeus heronifer, holotype, Banten, Lebak: Majasari (86– 88); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (89–91). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: distoventral incision of dextral apical lobe (I); sinistral, apical tooth of flagellum (II); distal emargination of dorsal lobe (III)." figureDoi="http://doi.org/10.5281/zenodo.10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="19" pageNumber="42">Figs 87, 88</figureCitation>
|
||
); deep distomedian emargination (
|
||
<figureCitation id="13256AE9FFD3FFE7BE64F98F0F850282" box="[575,664,1655,1679]" captionStart="Figures 86–91" captionStartId="27.[189,262,1863,1885]" captionTargetBox="[261,1365,582,1806]" captionTargetId="figure-268@27.[189,1436,563,1844]" captionTargetPageId="27" captionText="Figures 86–91. Apex of aedeagus in lateral (86, 89), ventral (87, 90), dorsal view (88, 91) of Scopaeus heronifer, holotype, Banten, Lebak: Majasari (86– 88); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (89–91). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: distoventral incision of dextral apical lobe (I); sinistral, apical tooth of flagellum (II); distal emargination of dorsal lobe (III)." figureDoi="http://doi.org/10.5281/zenodo.10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="19" pageNumber="42">Fig. 88</figureCitation>
|
||
: arrow
|
||
<collectionCode id="ED0FEEA9FFD3FFE7BEA2F98F0E0A0282" box="[761,791,1655,1679]" pageId="19" pageNumber="42">III</collectionCode>
|
||
) dividing dorsal lobe in short, convex, dextral lobe and long, slender, dextrad curved, sinistral lobe angled bent towards truncate apex (
|
||
<figureCitation id="13256AE9FFD3FFE7BD98F9220F5D02FF" box="[451,576,1754,1778]" captionStart="Figures 86–91" captionStartId="27.[189,262,1863,1885]" captionTargetBox="[261,1365,582,1806]" captionTargetId="figure-268@27.[189,1436,563,1844]" captionTargetPageId="27" captionText="Figures 86–91. Apex of aedeagus in lateral (86, 89), ventral (87, 90), dorsal view (88, 91) of Scopaeus heronifer, holotype, Banten, Lebak: Majasari (86– 88); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (89–91). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: distoventral incision of dextral apical lobe (I); sinistral, apical tooth of flagellum (II); distal emargination of dorsal lobe (III)." figureDoi="http://doi.org/10.5281/zenodo.10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="19" pageNumber="42">Figs 87, 88</figureCitation>
|
||
); flagellum strongly projecting from apical lobes ventrally, evenly curved dorsad and projecting from apex of dextral apical lobe (
|
||
<figureCitation id="13256AE9FFD3FFE7BC9EF8C50C020358" box="[197,287,1853,1877]" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="19" pageNumber="42">Figs 41</figureCitation>
|
||
,
|
||
<figureCitation id="13256AE9FFD3FFE7BD69F8C50C4D0358" box="[306,336,1853,1877]" captionStart="Figures 86–91" captionStartId="27.[189,262,1863,1885]" captionTargetBox="[261,1365,582,1806]" captionTargetId="figure-268@27.[189,1436,563,1844]" captionTargetPageId="27" captionText="Figures 86–91. Apex of aedeagus in lateral (86, 89), ventral (87, 90), dorsal view (88, 91) of Scopaeus heronifer, holotype, Banten, Lebak: Majasari (86– 88); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (89–91). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: distoventral incision of dextral apical lobe (I); sinistral, apical tooth of flagellum (II); distal emargination of dorsal lobe (III)." figureDoi="http://doi.org/10.5281/zenodo.10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="19" pageNumber="42">86</figureCitation>
|
||
), with strongly sclerotized (darker), in dorsal view acute-angled dextrad bent end, at point of bend with tooth pointing sinistrad (
|
||
<figureCitation id="13256AE9FFD3FFE7BE0AF8860FB9039B" box="[593,676,1918,1942]" captionStart="Figures 86–91" captionStartId="27.[189,262,1863,1885]" captionTargetBox="[261,1365,582,1806]" captionTargetId="figure-268@27.[189,1436,563,1844]" captionTargetPageId="27" captionText="Figures 86–91. Apex of aedeagus in lateral (86, 89), ventral (87, 90), dorsal view (88, 91) of Scopaeus heronifer, holotype, Banten, Lebak: Majasari (86– 88); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (89–91). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: distoventral incision of dextral apical lobe (I); sinistral, apical tooth of flagellum (II); distal emargination of dorsal lobe (III)." figureDoi="http://doi.org/10.5281/zenodo.10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="19" pageNumber="42">Fig. 87</figureCitation>
|
||
: arrow
|
||
<collectionCode id="ED0FEEA9FFD3FFE7BEA7F8860E0C039B" box="[764,785,1918,1942]" pageId="19" pageNumber="42">II</collectionCode>
|
||
); ventral lobe very short, triangularly projecting distad in lateral view (
|
||
<figureCitation id="13256AE9FFD3FFE7BFB8FF0909290507" box="[995,1076,241,266]" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="19" pageNumber="42">Figs 41</figureCitation>
|
||
,
|
||
<figureCitation id="13256AE9FFD3FFE7B865FF0909410504" box="[1086,1116,241,265]" captionStart="Figures 86–91" captionStartId="27.[189,262,1863,1885]" captionTargetBox="[261,1365,582,1806]" captionTargetId="figure-268@27.[189,1436,563,1844]" captionTargetPageId="27" captionText="Figures 86–91. Apex of aedeagus in lateral (86, 89), ventral (87, 90), dorsal view (88, 91) of Scopaeus heronifer, holotype, Banten, Lebak: Majasari (86– 88); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (89–91). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: distoventral incision of dextral apical lobe (I); sinistral, apical tooth of flagellum (II); distal emargination of dorsal lobe (III)." figureDoi="http://doi.org/10.5281/zenodo.10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="19" pageNumber="42">86</figureCitation>
|
||
), with apical emargination in ventral view (
|
||
<figureCitation id="13256AE9FFD3FFE7BF8BFEEA093C0527" box="[976,1057,274,298]" captionStart="Figures 86–91" captionStartId="27.[189,262,1863,1885]" captionTargetBox="[261,1365,582,1806]" captionTargetId="figure-268@27.[189,1436,563,1844]" captionTargetPageId="27" captionText="Figures 86–91. Apex of aedeagus in lateral (86, 89), ventral (87, 90), dorsal view (88, 91) of Scopaeus heronifer, holotype, Banten, Lebak: Majasari (86– 88); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (89–91). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: distoventral incision of dextral apical lobe (I); sinistral, apical tooth of flagellum (II); distal emargination of dorsal lobe (III)." figureDoi="http://doi.org/10.5281/zenodo.10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="19" pageNumber="42">Fig. 87</figureCitation>
|
||
); lateral lobes reduced, marked by lateroventral group of short setae (
|
||
<figureCitation id="13256AE9FFD3FFE7B8ECFECB082B0546" box="[1207,1334,307,331]" captionStart="Figures 86–91" captionStartId="27.[189,262,1863,1885]" captionTargetBox="[261,1365,582,1806]" captionTargetId="figure-268@27.[189,1436,563,1844]" captionTargetPageId="27" captionText="Figures 86–91. Apex of aedeagus in lateral (86, 89), ventral (87, 90), dorsal view (88, 91) of Scopaeus heronifer, holotype, Banten, Lebak: Majasari (86– 88); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (89–91). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: distoventral incision of dextral apical lobe (I); sinistral, apical tooth of flagellum (II); distal emargination of dorsal lobe (III)." figureDoi="http://doi.org/10.5281/zenodo.10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="19" pageNumber="42">Figs 86, 87</figureCitation>
|
||
); median foramen elongate with acute proximal end, slightly shifted dextrad, with transverse ridge narrow and bent posteriad in middle (
|
||
<figureCitation id="13256AE9FFD3FFE7B845FE6E097305A3" box="[1054,1134,406,430]" captionStart="Figures 86–91" captionStartId="27.[189,262,1863,1885]" captionTargetBox="[261,1365,582,1806]" captionTargetId="figure-268@27.[189,1436,563,1844]" captionTargetPageId="27" captionText="Figures 86–91. Apex of aedeagus in lateral (86, 89), ventral (87, 90), dorsal view (88, 91) of Scopaeus heronifer, holotype, Banten, Lebak: Majasari (86– 88); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (89–91). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: distoventral incision of dextral apical lobe (I); sinistral, apical tooth of flagellum (II); distal emargination of dorsal lobe (III)." figureDoi="http://doi.org/10.5281/zenodo.10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="19" pageNumber="42">Fig. 87</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA1766CFFD3FFE7BF14FE4F095405FD" blockId="19.[824,1437,241,1285]" pageId="19" pageNumber="42">
|
||
Female: Sperm pump stout with circular chamber (
|
||
<figureCitation id="13256AE9FFD3FFE7B929FE4F0E7F05FD" captionStart="Figures 163–172" captionStartId="42.[151,224,1839,1861]" captionTargetBox="[225,1322,230,1817]" captionTargetId="figure-168@42.[157,1398,210,1820]" captionTargetPageId="42" captionText="Figures 163–172. Sperm pump (163–172) and bursa (165, 167, 170, 172) of ♀ of Scopaeus grandis, paratype, Jawa Barat, Sukabumi: Cidahu (163); S.heronifer, paratype, Jawa Barat, Sukabumi:Sirnarasa (164); S. diversilobatus, paratype, Jawa Barat, Sukabumi, Djampang Kulon (165, with lateral gonocoxal plates IX folded laterally; 166); S. cuspilobatus, paratype, Jawa Barat, Sukabumi: Sirnarasa (167, with lateral gonocoxal plates IX, 168); S. velifer, paratype, Banten, Lebak: Citorek Kidul (169); S. elegantulus, Jawa Barat, Sukabumi: Djampang Kulon (170, with lateral gonocoxal plates IX, 171), S. uncinatus, paratype, Banten, Lebak: Citorek Kidul (172). Long scale bar: 163, 164, 166, 168, 169, 171, 172; short scale bar: 165, 167, 170." figureDoi="http://doi.org/10.5281/zenodo.10724315" httpUri="https://zenodo.org/record/10724315/files/figure.png" pageId="19" pageNumber="42">Fig. 164</figureCitation>
|
||
); bursa membranous.
|
||
</paragraph>
|
||
<paragraph id="8BA1766CFFD3FFE7BF14FE010EB007F3" blockId="19.[824,1437,241,1285]" pageId="19" pageNumber="42">
|
||
<emphasis id="B96AAA7EFFD3FFE7BF14FE010ED0061C" bold="true" box="[847,973,505,529]" pageId="19" pageNumber="42">Phylogeny:</emphasis>
|
||
<taxonomicName id="4C1E0DEFFFD3FFE7BF85FE0109A7061D" authorityName="Frisch" authorityYear="2022" box="[990,1210,504,528]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="heronifer">
|
||
<emphasis id="B96AAA7EFFD3FFE7BF85FE0109A7061D" box="[990,1210,504,528]" italics="true" pageId="19" pageNumber="42">Scopaeus heronifer</emphasis>
|
||
</taxonomicName>
|
||
belongs to the
|
||
<taxonomicName id="4C1E0DEFFFD3FFE7B9D3FE010EAE063C" authorityName="Fauvel" authorityYear="1905" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="longicornis">
|
||
<emphasis id="B96AAA7EFFD3FFE7B9D3FE010EAE063C" italics="true" pageId="19" pageNumber="42">S. longicornis</emphasis>
|
||
</taxonomicName>
|
||
complex (
|
||
<bibRefCitation id="EF8F0B9DFFD3FFE7B86DFDE1081C063F" author="Frisch, J." box="[1078,1281,537,562]" pageId="19" pageNumber="42" pagination="195 - 218" refId="ref31602" refString="Frisch, J. (2007): A review of the Scopaeus gracilis species group (Coleoptera, Staphylinidae, Paederinae), with description of new species from Sardinia, southern Africa and Middle East. - Deutsche Entomologische Zeitschrift 54 (2): 195 - 218." type="journal article" year="2007">Frisch 2007: 204</bibRefCitation>
|
||
, 205) of the
|
||
<taxonomicName id="4C1E0DEFFFD3FFE7BF63FDC30EB7065F" authorityName="Oke" authorityYear="1933" box="[824,938,570,594]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B96AAA7EFFD3FFE7BF63FDC30EB7065F" box="[824,938,570,594]" italics="true" pageId="19" pageNumber="42">S. gracilis</emphasis>
|
||
</taxonomicName>
|
||
species group (
|
||
<bibRefCitation id="EF8F0B9DFFD3FFE7B804FDC20848065F" author="Frisch, J. & D. Burckhardt & V. Wolters" box="[1119,1365,570,594]" pageId="19" pageNumber="42" pagination="27 - 53" refId="ref31920" refString="Frisch, J., D. Burckhardt & V. Wolters (2002): Rove beetles of the subtribe Scopaeina Mulsant & Rey (Coleoptera: Staphylinidae) in the West Palaearctic: Phylogeny, biogeography and species catalogue. - Organisms, Diversity & Evolution 2: 27 - 53." type="journal article" year="2002">Frisch et al. 2002: 39</bibRefCitation>
|
||
). This subgroup comprises two phylogeographical clades, the South African and Madagascan
|
||
<taxonomicName id="4C1E0DEFFFD3FFE7B8FCFD84085F0699" authorityName="Fauvel" authorityYear="1905" box="[1191,1346,636,660]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="longicornis">
|
||
<emphasis id="B96AAA7EFFD3FFE7B8FCFD84085F0699" box="[1191,1346,636,660]" italics="true" pageId="19" pageNumber="42">S. longicornis</emphasis>
|
||
</taxonomicName>
|
||
lineage and the Middle Eastern
|
||
<taxonomicName id="4C1E0DEFFFD3FFE7B805FD6509FF06B8" authorityName="Frisch" authorityYear="2007" box="[1118,1250,669,693]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="asirensis">
|
||
<emphasis id="B96AAA7EFFD3FFE7B805FD6509FF06B8" box="[1118,1250,669,693]" italics="true" pageId="19" pageNumber="42">S. asirensis</emphasis>
|
||
</taxonomicName>
|
||
lineage (
|
||
<bibRefCitation id="EF8F0B9DFFD3FFE7B90CFD650EF806DB" author="Frisch, J." pageId="19" pageNumber="42" pagination="195 - 218" refId="ref31602" refString="Frisch, J. (2007): A review of the Scopaeus gracilis species group (Coleoptera, Staphylinidae, Paederinae), with description of new species from Sardinia, southern Africa and Middle East. - Deutsche Entomologische Zeitschrift 54 (2): 195 - 218." type="journal article" year="2007">Frisch 2007: 213 – 215</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF8F0B9DFFD3FFE7BFA0FD46092F06DB" author="Frisch, J." box="[1019,1074,702,726]" pageId="19" pageNumber="42" pagination="61 - 70" refId="ref31734" refString="Frisch, J. (2015): A new species of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) from Socotra Island, with distributional and phylogeographical notes on related species. - Soil Organisms 87 (1): 61 - 70." type="journal article" year="2015">2015</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C1E0DEFFFD3FFE7B816FD46083506DB" authorityName="Frisch" authorityYear="2022" box="[1101,1320,702,726]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="heronifer">
|
||
<emphasis id="B96AAA7EFFD3FFE7B816FD46083506DB" box="[1101,1320,702,726]" italics="true" pageId="19" pageNumber="42">Scopaeus heronifer</emphasis>
|
||
</taxonomicName>
|
||
, the first known Oriental species of the
|
||
<taxonomicName id="4C1E0DEFFFD3FFE7B8D4FD27083506FA" authorityName="Fauvel" authorityYear="1905" box="[1167,1320,735,759]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="longicornis">
|
||
<emphasis id="B96AAA7EFFD3FFE7B8D4FD27083506FA" box="[1167,1320,735,759]" italics="true" pageId="19" pageNumber="42">S. longicornis</emphasis>
|
||
</taxonomicName>
|
||
subgroup, represents another phylogenetic lineage. It lacks the characteristic, horn-shaped, sinistral extension of the base of the flagellum of the Afrotropical and Palaearctic species (
|
||
<bibRefCitation id="EF8F0B9DFFD3FFE7BFCEFC9A09060777" author="Frisch, J." box="[917,1051,866,890]" pageId="19" pageNumber="42" pagination="195 - 218" refId="ref31602" refString="Frisch, J. (2007): A review of the Scopaeus gracilis species group (Coleoptera, Staphylinidae, Paederinae), with description of new species from Sardinia, southern Africa and Middle East. - Deutsche Entomologische Zeitschrift 54 (2): 195 - 218." type="journal article" year="2007">Frisch 2007</bibRefCitation>
|
||
: e.g.
|
||
<figureCitation id="13256AE9FFD3FFE7B80FFC9A09B90777" box="[1108,1188,866,890]" captionStart="Figures 23–26" captionStartId="14.[151,224,1930,1952]" captionTargetBox="[263,1352,435,1899]" captionTargetId="figure-131@14.[151,1398,422,1911]" captionTargetPageId="14" captionText="Figures 23–26. Habitus of Scopaeus cuspidatus, paratype, Jawa Barat: Mt Salak (23); S. sulawesianus, paratype, Sulawesi Tengah: Tambarana (24); S. niger, paralectotype, Singapore (25); S. borneensis, holotype, Kalimantan Barat: Pontianak (26)." figureDoi="http://doi.org/10.5281/zenodo.10724252" httpUri="https://zenodo.org/record/10724252/files/figure.png" pageId="19" pageNumber="42">Fig. 26</figureCitation>
|
||
). The sinistral tooth at the sharp apical bend of the flagellum (arrow in
|
||
<figureCitation id="13256AE9FFD3FFE7B913FC7B08880796" box="[1352,1429,899,923]" captionStart="Figures 86–91" captionStartId="27.[189,262,1863,1885]" captionTargetBox="[261,1365,582,1806]" captionTargetId="figure-268@27.[189,1436,563,1844]" captionTargetPageId="27" captionText="Figures 86–91. Apex of aedeagus in lateral (86, 89), ventral (87, 90), dorsal view (88, 91) of Scopaeus heronifer, holotype, Banten, Lebak: Majasari (86– 88); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (89–91). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: distoventral incision of dextral apical lobe (I); sinistral, apical tooth of flagellum (II); distal emargination of dorsal lobe (III)." figureDoi="http://doi.org/10.5281/zenodo.10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="19" pageNumber="42">Fig. 87</figureCitation>
|
||
) is probably homologous to the sinistral enlargement of the flagellum of the
|
||
<taxonomicName id="4C1E0DEFFFD3FFE7B855FC3D099407D0" authorityName="Frisch" authorityYear="2007" box="[1038,1161,965,989]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="asirensis">
|
||
<emphasis id="B96AAA7EFFD3FFE7B855FC3D099407D0" box="[1038,1161,965,989]" italics="true" pageId="19" pageNumber="42">S. asirensis</emphasis>
|
||
</taxonomicName>
|
||
lineage (
|
||
<bibRefCitation id="EF8F0B9DFFD3FFE7B8B7FC3D087007D0" author="Frisch, J." box="[1260,1389,965,989]" pageId="19" pageNumber="42" pagination="61 - 70" refId="ref31734" refString="Frisch, J. (2015): A new species of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) from Socotra Island, with distributional and phylogeographical notes on related species. - Soil Organisms 87 (1): 61 - 70." type="journal article" year="2015">Frisch 2015</bibRefCitation>
|
||
: e.g.
|
||
<figureCitation id="13256AE9FFD3FFE7BF63FC1E0EBC07F3" box="[824,929,998,1022]" captionStart="Figures 8–10" captionStartId="5.[189,262,1930,1952]" captionTargetBox="[237,1406,902,1874]" captionTargetId="figure-357@5.[189,1436,889,1911]" captionTargetPageId="5" captionText="Figures 8–10. Habitus of Scopaeus tortuosiflagellatus, paratype, Jawa Barat, Sukabumi: Cisalimas (8); S. posoanus, paratype, Sulawesi Tengah: Lake Poso (9); S. spiraliflagellatus, paratype, Sulawesi Tengah: Lake Poso (10)." figureDoi="http://doi.org/10.5281/zenodo.10724240" httpUri="https://zenodo.org/record/10724240/files/figure.png" pageId="19" pageNumber="42">Figs 9, 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30425E7FFD3FFE7BF14FBFE09DF006D" pageId="19" pageNumber="42" type="distribution">
|
||
<paragraph id="8BA1766CFFD3FFE7BF14FBFE09DF006D" blockId="19.[824,1437,241,1285]" pageId="19" pageNumber="42">
|
||
<emphasis id="B96AAA7EFFD3FFE7BF14FBFE0EF80013" bold="true" box="[847,997,1030,1054]" pageId="19" pageNumber="42">Distribution:</emphasis>
|
||
<taxonomicName id="4C1E0DEFFFD3FFE7BFAFFBFF09D20012" authorityName="Frisch" authorityYear="2022" box="[1012,1231,1031,1055]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="heronifer">
|
||
<emphasis id="B96AAA7EFFD3FFE7BFAFFBFF09D20012" box="[1012,1231,1031,1055]" italics="true" pageId="19" pageNumber="42">Scopaeus heronifer</emphasis>
|
||
</taxonomicName>
|
||
was collected at the edge of Mount Halimun-Salak National Park in the provinces of
|
||
<collectingRegion id="49DAB88EFFD3FFE7BF9FFBB1090D006D" box="[964,1040,1097,1120]" country="Indonesia" name="Banten" pageId="19" pageNumber="42">Banten</collectingRegion>
|
||
and
|
||
<collectingRegion id="49DAB88EFFD3FFE7B81EFBB109A3006D" box="[1093,1214,1097,1120]" country="Indonesia" name="Jawa Barat" pageId="19" pageNumber="42">Jawa Barat</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30425E7FFD3FFE7BF14FB910EB60108" pageId="19" pageNumber="42" type="etymology">
|
||
<paragraph id="8BA1766CFFD3FFE7BF14FB910EB60108" blockId="19.[824,1437,241,1285]" pageId="19" pageNumber="42">
|
||
<emphasis id="B96AAA7EFFD3FFE7BF14FB910EC8008C" bold="true" box="[847,981,1129,1153]" pageId="19" pageNumber="42">Etymology:</emphasis>
|
||
The epithet
|
||
<taxonomicName id="4C1E0DEFFFD3FFE7B834FB9109CE008C" authorityName="Frisch" authorityYear="2022" box="[1135,1235,1129,1153]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="heronifer">
|
||
<emphasis id="B96AAA7EFFD3FFE7B834FB9109CE008C" box="[1135,1235,1129,1153]" italics="true" pageId="19" pageNumber="42">heronifer</emphasis>
|
||
</taxonomicName>
|
||
[adjective, Latin, composed of the noun
|
||
<emphasis id="B96AAA7EFFD3FFE7B86DFB72096800AF" box="[1078,1141,1162,1186]" italics="true" pageId="19" pageNumber="42">heron</emphasis>
|
||
(heron) and the verb
|
||
<emphasis id="B96AAA7EFFD3FFE7B933FB72088000AF" box="[1384,1437,1162,1186]" italics="true" pageId="19" pageNumber="42">ferre</emphasis>
|
||
(to carry, to bring)] refers to the apex of the flagellum of the aedeagus (
|
||
<figureCitation id="13256AE9FFD3FFE7BF8FFB34093E00E9" box="[980,1059,1228,1252]" captionStart="Figures 92–97" captionStartId="28.[151,224,1886,1908]" captionTargetBox="[223,1329,734,1834]" captionTargetId="figure-235@28.[151,1398,685,1867]" captionTargetPageId="28" captionText="Figures 92–97. Apex of aedeagus in lateral (92, 95), ventral (93, 96), dorsal view (94, 97) of Scopaeus cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (92–94); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (95– 97). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral tooth of dextral apical lobe (I), lateral ridges of median foramen (II, III)." figureDoi="http://doi.org/10.5281/zenodo.10724282" httpUri="https://zenodo.org/record/10724282/files/figure.png" pageId="19" pageNumber="42">Fig. 94</figureCitation>
|
||
), which is reminescent of the head of a heron.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |