316 lines
40 KiB
XML
316 lines
40 KiB
XML
<document id="8C6BCD876B477616E1DF5781DA6BE184" ID-CLB-Dataset="279445" ID-DOI="10.11646/zootaxa.5365.1.1" ID-GBIF-Dataset="23a3b51c-63b7-4937-bdba-384f28db6be5" ID-ISSN="1175-5334" ID-Zenodo-Dep="10146969" ID-ZooBank="1DC72714-D0E8-49D8-821D-03C6B2A7AE80" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1700197580075" checkinUser="plazi" docAuthor="Colloff, Matthew J." docDate="2023" docId="03A2C77C466AFFC0C79CB6D514F6D960" docLanguage="en" docName="zootaxa.5365.1.1.pdf" docOrigin="Zootaxa 5365 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Eutegaeidae J. Balogh 1965" docType="treatment" docVersion="5" lastPageNumber="13" masterDocId="FF9BBF044661FFCCC70BB31F1724DE65" masterDocTitle="The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families" masterLastPageNumber="93" masterPageNumber="1" pageNumber="12" updateTime="1701119607223" updateUser="ExternalLinkService">
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<mods:title id="C9270568804B12B142CEF9A719A87525">The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families</mods:title>
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<mods:namePart id="DE1411BE96F549DE4FA782B87B717F52">Colloff, Matthew J.</mods:namePart>
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<mods:typeOfResource id="27B1359729F6EB64FD7B25750693BB09">text</mods:typeOfResource>
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<mods:title id="BA3CB3E199F83FECA1E3AD8A1BBCC671">Zootaxa</mods:title>
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<mods:part id="7C60B91B79F3F476191AAAC1A4048D99">
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<mods:date id="E186799CF75940068643E20BAEE0F07F">2023</mods:date>
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<mods:number id="23ED0685A597CB3489B4FB8A3321893B">5365</mods:number>
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<mods:identifier id="40F21DFD76865AE56ED495A2CEE40FF3" type="CLB-Dataset">279445</mods:identifier>
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<mods:identifier id="931467E5E2878A125A8DDA0888548A03" type="DOI">10.11646/zootaxa.5365.1.1</mods:identifier>
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</mods:mods>
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<treatment id="03A2C77C466AFFC0C79CB6D514F6D960" ID-GBIF-Taxon="219262774" LSID="urn:lsid:plazi:treatment:03A2C77C466AFFC0C79CB6D514F6D960" httpUri="http://treatment.plazi.org/id/03A2C77C466AFFC0C79CB6D514F6D960" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
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<subSubSection id="C31125E1466AFFC7C79CB6D516D3DB81" box="[151,503,1482,1509]" pageId="11" pageNumber="12" type="nomenclature">
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<paragraph id="8BB4766A466AFFC7C79CB6D516D3DB81" blockId="11.[151,503,1482,1509]" box="[151,503,1482,1509]" pageId="11" pageNumber="12">
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<heading id="D0FCC106466AFFC7C79CB6D516D3DB81" bold="true" box="[151,503,1482,1509]" fontSize="11" level="2" pageId="11" pageNumber="12" reason="3">
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<taxonomicName id="4C0B0DE9466AFFC7C79CB6D516D3DB81" ID-CoL="9ZW" authority="J. Balogh, 1965" authorityName="J. Balogh" authorityYear="1965" box="[151,503,1482,1509]" class="Arachnida" family="Eutegaeidae" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="12" phylum="Arthropoda" rank="family">
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<emphasis id="B97FAA78466AFFC7C79CB6D516D3DB81" bold="true" box="[151,503,1482,1509]" pageId="11" pageNumber="12">
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Eutegaeidae J.
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<bibRefCitation id="EF9A0B9B466AFFC7C65EB6D416D3DB81" author="Balogh, J." box="[341,503,1482,1509]" pageId="11" pageNumber="12" pagination="5 - 99" refId="ref64020" refString="Balogh, J. (1965) A synopsis of the world oribatid (Acari) genera. Acta Zoologica Academiae Scientiarum Hungaricae, 11, 5 - 99." type="journal article" year="1965">Balogh, 1965</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31125E1466AFFC7C79CB51016DBD842" box="[151,511,1551,1575]" pageId="11" pageNumber="12" type="reference_group">
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<paragraph id="8BB4766A466AFFC7C79CB51016DBD842" blockId="11.[151,602,1551,1606]" box="[151,511,1551,1575]" pageId="11" pageNumber="12">
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<treatmentCitationGroup id="AB1B5144466AFFC7C79CB51016DBD842" box="[151,511,1551,1575]" pageId="11" pageNumber="12">
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<treatmentCitation id="0AAA507B466AFFC7C79CB51016DED842" author="Balogh, J." box="[151,506,1551,1575]" page="59" pageId="11" pageNumber="12" year="1965">
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<taxonomicName id="4C0B0DE9466AFFC7C79CB51016DED842" ID-CoL="9ZW" authority="J. Balogh, 1965, p. 59" authorityName="J. Balogh" authorityPageNumber="59" authorityYear="1965" box="[151,506,1551,1575]" class="Arachnida" family="Eutegaeidae" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="12" phylum="Arthropoda" rank="family">
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Eutegaeidae J.
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<bibRefCitation id="EF9A0B9B466AFFC7C63BB510169FD842" author="Balogh, J." box="[304,443,1551,1575]" pageId="11" pageNumber="12" pagination="5 - 99" refId="ref64020" refString="Balogh, J. (1965) A synopsis of the world oribatid (Acari) genera. Acta Zoologica Academiae Scientiarum Hungaricae, 11, 5 - 99." type="journal article" year="1965">Balogh, 1965</bibRefCitation>
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, p. 59
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31125E1466AFFC7C79CB531157ED823" box="[151,602,1582,1606]" pageId="11" pageNumber="12" type="materials_examined">
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<paragraph id="8BB4766A466AFFC7C79CB531157ED823" blockId="11.[151,602,1551,1606]" box="[151,602,1582,1606]" pageId="11" pageNumber="12">
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<typeStatus id="54B0C8C8466AFFC7C79CB53117EDD823" box="[151,201,1582,1606]" pageId="11" pageNumber="12">Type</typeStatus>
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genus:
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<taxonomicName id="4C0B0DE9466AFFC7C613B5301571D823" authority="Berlese, 1916, p. 62" authorityName="Berlese" authorityPageNumber="62" authorityYear="1916" box="[280,597,1582,1606]" class="Arachnida" family="Eutegaeidae" genus="Eutegaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="B97FAA78466AFFC7C613B53016A6D823" box="[280,386,1583,1606]" italics="true" pageId="11" pageNumber="12">Eutegaeus</emphasis>
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<bibRefCitation id="EF9A0B9B466AFFC7C683B5311532D823" author="Berlese, A." box="[392,534,1582,1606]" pageId="11" pageNumber="12" pagination="19 - 67" refId="ref64788" refString="Berlese, A. (1916) Centuria prima di Acari nuovi. Redia, 12, 19 - 67." type="journal article" year="1916">Berlese, 1916</bibRefCitation>
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, p. 62
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31125E1466AFFC0C79CB56B1675DF24" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="diagnosis">
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<paragraph id="8BB4766A466AFFC0C79CB56B1675DF24" blockId="11.[151,1437,1652,2044]" lastBlockId="12.[151,1437,151,1797]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
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<emphasis id="B97FAA78466AFFC7C79CB56B1636D8EB" bold="true" box="[151,274,1652,1678]" pageId="11" pageNumber="12">Diagnosis.</emphasis>
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the following diagnosis is modified from those of J.
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<bibRefCitation id="EF9A0B9B466AFFC7C45CB56B14DED8EA" author="Balogh, J." box="[855,1018,1652,1679]" pageId="11" pageNumber="12" pagination="5 - 99" refId="ref64020" refString="Balogh, J. (1965) A synopsis of the world oribatid (Acari) genera. Acta Zoologica Academiae Scientiarum Hungaricae, 11, 5 - 99." type="journal article" year="1965">Balogh (1965)</bibRefCitation>
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,
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<bibRefCitation id="EF9A0B9B466AFFC7C30EB56B1397D8EA" author="Woolley, T. A." box="[1029,1203,1652,1679]" pageId="11" pageNumber="12" pagination="381 - 388" refId="ref69684" refString="Woolley, T. A. (1965) Eutegaeidae, a new family of oribatid mites, with a description of a new species from New Zealand (Acarina: Oribatei). Acarologia, 7, 381 - 388." type="journal article" year="1965">Woolley (1965)</bibRefCitation>
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and
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<bibRefCitation id="EF9A0B9B466AFFC7C3ECB56A12BCD8EB" author="Luxton, M." box="[1255,1432,1652,1679]" pageId="11" pageNumber="12" pagination="71 - 91" refId="ref67435" refString="Luxton, M. (1988 a) A new mite superfamily (Acari: Cryptostigmata). Zoological Journal of the Linnean Society, 93, 71 - 91. https: // doi. org / 10.1111 / j. 1096 - 3642.1988. tb 01360. x" type="journal article" year="1988">Luxton (1988a)</bibRefCitation>
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. Rostrum acute, entire; lamellae very broad, covering lateral margins of prodorsum, with free cusps; lamellar seta emerging from excavation at apex of cusp, often between two teeth. Interlamellar setae positioned medial of bothridia; bothridium not fused with bases of lamellae, long, corniculate, opening anterolaterally, often with anterior condyle of enantiophysis H on posterior margin. Bothridial seta typically very long, setiform or bacilliform. Prodorsum waisted posterior of bothridium. Humeral processes prominent, ovoid, waisted basally, pointed apically, extending from anterolateral margin of notogaster to point at least halfway along length of lamella, but not reflexed ventrally. Notogaster circular to ovoid, convex, smooth, broader than long. With eight or nine pairs of notogastral setae, either all setae of the
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<emphasis id="B97FAA78466AFFC7C69BB48B16BCD9CB" box="[400,408,1940,1966]" italics="true" pageId="11" pageNumber="12">l</emphasis>
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and
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<emphasis id="B97FAA78466AFFC7C6DAB48B16FBD9CB" box="[465,479,1940,1966]" italics="true" pageId="11" pageNumber="12">h</emphasis>
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series in the centrodorsal position (
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<taxonomicName id="4C0B0DE9466AFFC7C472B48A1337D9CB" authorityName="Woolley" authorityYear="1965" box="[889,1043,1941,1966]" class="Arachnida" family="Eutegaeidae" genus="Neseutegaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="B97FAA78466AFFC7C472B48A1337D9CB" box="[889,1043,1941,1966]" italics="true" pageId="11" pageNumber="12">Neseutegaeus</emphasis>
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</taxonomicName>
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) or at least one pair in the
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<emphasis id="B97FAA78466AFFC7C247B48B1270D9CB" box="[1356,1364,1940,1966]" italics="true" pageId="11" pageNumber="12">l</emphasis>
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series in centrodorsal position (
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<taxonomicName id="4C0B0DE9466AFFC7C6BFB4A6150DD9B7" authorityName="Berlese" authorityYear="1916" box="[436,553,1977,2002]" class="Arachnida" family="Eutegaeidae" genus="Eutegaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="B97FAA78466AFFC7C6BFB4A6150DD9B7" box="[436,553,1977,2002]" italics="true" pageId="11" pageNumber="12">Eutegaeus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C0B0DE9466AFFC7C569B4A715D0D9B7" authorityName="Luxton" authorityYear="1988" box="[610,756,1976,2002]" class="Arachnida" family="Eutegaeidae" genus="Atalotegaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="B97FAA78466AFFC7C569B4A715D0D9B7" box="[610,756,1976,2002]" italics="true" pageId="11" pageNumber="12">Atalotegaeus</emphasis>
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</taxonomicName>
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); setae of
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<emphasis id="B97FAA78466AFFC7C465B4A71452D9B7" box="[878,886,1976,2002]" italics="true" pageId="11" pageNumber="12">l</emphasis>
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and
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<emphasis id="B97FAA78466AFFC7C4A4B4A71499D9B7" box="[943,957,1976,2002]" italics="true" pageId="11" pageNumber="12">h</emphasis>
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series (and occasionally
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<emphasis id="B97FAA78466AFFC7C3D7B4A613CED9B7" box="[1244,1258,1977,2002]" italics="true" pageId="11" pageNumber="12">p</emphasis>
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<subScript id="178F742F466AFFC7C3E1B4D513D6D9BD" attach="left" box="[1258,1266,1994,2008]" fontSize="6" pageId="11" pageNumber="12">1</subScript>
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) longer and of different morphology from those of
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<emphasis id="B97FAA78466AFFC7C527B4C2151ED993" box="[556,570,2013,2038]" italics="true" pageId="11" pageNumber="12">p</emphasis>
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series (or
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<emphasis id="B97FAA78466AFFC7C5A4B4C215E1D999" box="[687,709,2013,2044]" italics="true" pageId="11" pageNumber="12">
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p
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<subScript id="178F742F466AFFC7C5B6B4F115E1D999" attach="left" box="[701,709,2030,2044]" fontSize="6" pageId="11" pageNumber="12">2</subScript>
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</emphasis>
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||
and
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<emphasis id="B97FAA78466AFFC7C5F7B4C2142ED993" box="[764,778,2013,2038]" italics="true" pageId="11" pageNumber="12">p</emphasis>
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<subScript id="178F742F466AFFC7C401B4F11436D999" attach="left" box="[778,786,2030,2044]" fontSize="6" pageId="11" pageNumber="12">3</subScript>
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). Chelicerae chelate-dentate and of normal proportions or slim but not with needle-like modified digits. Pedotectum I (pd I) sub-rectangular in lateral outline; pd II triangular or rectangular; discidium pointed. Circumpedal carina present. Perigenital carinae and enantiophyses E4 absent. Five or six pairs of genital setae, three pairs of adanal setae. Anal plates lozenge-shaped. Pre-anal organ T-shaped. Nymphs with or without one or most gastronotic setae emerging from flat scales; lateral margin of lamellar region strongly incised.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31125E1466DFFC0C7CCB25414F6D960" pageId="12" pageNumber="13" type="discussion">
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<paragraph id="8BB4766A466DFFC0C7CCB25416C6DFB4" blockId="12.[151,1437,151,1797]" pageId="12" pageNumber="13">
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<emphasis id="B97FAA78466DFFC0C7CCB254161FDF00" bold="true" box="[199,315,331,357]" pageId="12" pageNumber="13">Remarks.</emphasis>
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Since the publication by J.
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<bibRefCitation id="EF9A0B9B466DFFC0C568B2541422DF00" author="Balogh, J." box="[611,774,330,357]" pageId="12" pageNumber="13" pagination="5 - 99" refId="ref64020" refString="Balogh, J. (1965) A synopsis of the world oribatid (Acari) genera. Acta Zoologica Academiae Scientiarum Hungaricae, 11, 5 - 99." type="journal article" year="1965">Balogh (1965)</bibRefCitation>
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, the
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<taxonomicName id="4C0B0DE9466DFFC0C43DB254149BDF00" authorityName="J. Balogh" authorityYear="1965" box="[822,959,331,357]" class="Arachnida" family="Eutegaeidae" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Eutegaeidae</taxonomicName>
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has contained between two and nine genera (
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<tableCitation id="C68943D1466DFFC0C794B27017D7DFEC" box="[159,243,367,393]" captionStart="TABLE 1" captionStartId="5.[145,237,449,475]" captionTargetPageId="5" captionText="TABLE 1. Classifications of families and genera of Eutegaeoidea and Cepheoidea. Luxton (1988a) covers Eutegaeoidea only. Norton and Behan-Pelletier (2009) did not include genera under the family group names." httpUri="http://table.plazi.org/id/DF7426E24664FFC9C79AB2DE1502DF9A" pageId="12" pageNumber="13" tableUuid="DF7426E24664FFC9C79AB2DE1502DF9A">Table 1</tableCitation>
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||
). The diagnosis above is based on a concept of the
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<taxonomicName id="4C0B0DE9466DFFC0C433B27014E5DFEC" authorityName="J. Balogh" authorityYear="1965" box="[824,961,367,393]" class="Arachnida" family="Eutegaeidae" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Eutegaeidae</taxonomicName>
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as containing only three genera (
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<tableCitation id="C68943D1466DFFC0C23DB27012AADFEC" box="[1334,1422,367,393]" captionStart="TABLE 2" captionStartId="10.[152,249,150,176]" captionTargetBox="[159,1385,189,1947]" captionTargetPageId="10" captionText="TABLE 2. A checklist of Eutegaeoidea J. Balogh, 1965 based upon the present work." httpUri="http://table.plazi.org/id/DF7426E2466BFFC6C793B389137EDED4" pageId="12" pageNumber="13" tableUuid="DF7426E2466BFFC6C793B389137EDED4">Table 2</tableCitation>
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||
):
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<taxonomicName id="4C0B0DE9466DFFC0C79CB28C1628DFC9" authorityName="Berlese" authorityYear="1916" box="[151,268,403,428]" class="Arachnida" family="Eutegaeidae" genus="Eutegaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
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<emphasis id="B97FAA78466DFFC0C79CB28C1628DFC9" box="[151,268,403,428]" italics="true" pageId="12" pageNumber="13">Eutegaeus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0B0DE9466DFFC0C610B28C1689DFC8" authorityName="Luxton" authorityYear="1988" box="[283,429,403,429]" class="Arachnida" family="Eutegaeidae" genus="Atalotegaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA78466DFFC0C610B28C1689DFC8" box="[283,429,403,429]" italics="true" pageId="12" pageNumber="13">Atalotegaeus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0B0DE9466DFFC0C6EEB28C155BDFC9" authorityName="Woolley" authorityYear="1965" box="[485,639,403,428]" class="Arachnida" family="Eutegaeidae" genus="Neseutegaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA78466DFFC0C6EEB28C155BDFC9" box="[485,639,403,428]" italics="true" pageId="12" pageNumber="13">Neseutegaeus</emphasis>
|
||
</taxonomicName>
|
||
, due to
|
||
<taxonomicName id="4C0B0DE9466DFFC0C5D6B28C1447DFC9" authorityName="Luxton" authorityYear="1988" box="[733,867,403,428]" class="Arachnida" family="Eutegaeidae" genus="Birotegaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA78466DFFC0C5D6B28C1447DFC9" box="[733,867,403,428]" italics="true" pageId="12" pageNumber="13">Birotegaeus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0B0DE9466DFFC0C490B28C1310DFC9" authorityName="Woolley" authorityYear="1965" box="[923,1076,403,428]" class="Arachnida" family="Eutegaeidae" genus="Pareutegaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA78466DFFC0C490B28C1310DFC9" box="[923,1076,403,428]" italics="true" pageId="12" pageNumber="13">Pareutegaeus</emphasis>
|
||
</taxonomicName>
|
||
being newly synonymised with
|
||
<taxonomicName id="4C0B0DE9466DFFC0C79CB2A81628DFB5" authorityName="Berlese" authorityYear="1916" box="[151,268,439,464]" class="Arachnida" family="Eutegaeidae" genus="Eutegaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA78466DFFC0C79CB2A81628DFB5" box="[151,268,439,464]" italics="true" pageId="12" pageNumber="13">Eutegaeus</emphasis>
|
||
</taxonomicName>
|
||
, as detailed below.
|
||
</paragraph>
|
||
<paragraph id="8BB4766A466DFFC0C7CCB2C41200DDE4" blockId="12.[151,1437,151,1797]" pageId="12" pageNumber="13">
|
||
The main character states that differentiate
|
||
<taxonomicName id="4C0B0DE9466DFFC0C5B8B2C41418DF90" authorityName="J. Balogh" authorityYear="1965" box="[691,828,475,501]" class="Arachnida" family="Eutegaeidae" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Eutegaeidae</taxonomicName>
|
||
from the other families of
|
||
<taxonomicName id="4C0B0DE9466DFFC0C37BB2C41223DF90" authorityName="Balogh" authorityYear="1965" box="[1136,1287,475,501]" class="Pilidiophora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heteronemertea" pageId="12" pageNumber="8" phylum="Nemertea" rank="superFamily" superFamily="Eutegaeoidea">Eutegaeoidea</taxonomicName>
|
||
are the long, pointed humeral processes that emerge from the anterolateral margin of the notogaster, directly posterior of the bothridia and have a narrow, waisted point of attachment to the notogaster; narrower than the anterior part of the humeral process and one or more of the setae of the
|
||
<emphasis id="B97FAA78466DFFC0C410B1581407DC04" box="[795,803,583,609]" italics="true" pageId="12" pageNumber="13">l</emphasis>
|
||
series in the centrodorsal position. These two character states are synapomorphies of
|
||
<taxonomicName id="4C0B0DE9466DFFC0C6FEB17415A6DCE0" authorityName="J. Balogh" authorityYear="1965" box="[501,642,619,645]" class="Arachnida" family="Eutegaeidae" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Eutegaeidae</taxonomicName>
|
||
. In
|
||
<taxonomicName id="4C0B0DE9466DFFC0C5B8B1741442DCE0" authority="Colloff, 2023" authorityName="Colloff" authorityYear="2023" box="[691,870,619,645]" family="Neoeutegaeidae" pageId="12" pageNumber="13" rank="family" status="fam. nov.">Neoeutegaeidae</taxonomicName>
|
||
<emphasis id="B97FAA78466DFFC0C479B17514C0DCE0" bold="true" box="[882,996,618,645]" pageId="12" pageNumber="13">
|
||
<taxonomicNameLabel id="A24C1703466DFFC0C479B17514C0DCE0" box="[882,996,618,645]" pageId="12" pageNumber="13" rank="family">fam. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(cf. below) the humeral process also emerges on the anterior notogaster but its base is broad and has a distinct bulge and incurve where it meets the notogaster. In
|
||
<taxonomicName id="4C0B0DE9466DFFC0C631B1AC16D8DCA8" authorityName="Luxton" authorityYear="1988" box="[314,508,691,717]" family="Compactozetidae" pageId="12" pageNumber="13" rank="family">Compactozetidae</taxonomicName>
|
||
the humeral process has its origin some distance posterior of that in
|
||
<taxonomicName id="4C0B0DE9466DFFC0C218B1AC12B8DCA8" authorityName="J. Balogh" authorityYear="1965" box="[1299,1436,691,717]" class="Arachnida" family="Eutegaeidae" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Eutegaeidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0B0DE9466DFFC0C7CCB1C8165EDC94" authority="Colloff, 2023" authorityName="Colloff" authorityYear="2023" box="[199,378,727,753]" family="Neoeutegaeidae" pageId="12" pageNumber="13" rank="family" status="fam. nov.">Neoeutegaeidae</taxonomicName>
|
||
<emphasis id="B97FAA78466DFFC0C68AB1C916CBDC94" bold="true" box="[385,495,726,753]" pageId="12" pageNumber="13">
|
||
<taxonomicNameLabel id="A24C1703466DFFC0C68AB1C916CBDC94" box="[385,495,726,753]" pageId="12" pageNumber="13" rank="family">fam. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, at the widest part of the notogaster. In
|
||
<taxonomicName id="4C0B0DE9466DFFC0C4A5B1C8131CDC94" authorityName="Luxton" authorityYear="1988" box="[942,1080,727,753]" class="Arachnida" family="Pterozetidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Pterozetidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0B0DE9466DFFC0C364B1C81201DC94" authority="Colloff, 2023" authorityName="Colloff" authorityYear="2023" box="[1135,1317,727,753]" family="Porrhotegaeidae" pageId="12" pageNumber="13" rank="family" status="fam. nov.">Porrhotegaeidae</taxonomicName>
|
||
<taxonomicNameLabel id="A24C1703466DFFC0C226B1C912B8DC94" box="[1325,1436,726,753]" pageId="12" pageNumber="13" rank="family">
|
||
<emphasis id="B97FAA78466DFFC0C226B1C912B2DC94" bold="true" box="[1325,1430,726,753]" pageId="12" pageNumber="13">fam. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
the humeral process also originates in this position and is pointed apically and broad basally; its lateral margin is straight and lacks any prominent constriction or bulge at its base. In
|
||
<taxonomicName id="4C0B0DE9466DFFC0C4BCB0001357DD5C" authority="Colloff, 2023" authorityName="Colloff" authorityYear="2023" box="[951,1139,799,825]" family="Bornebuschiidae" pageId="12" pageNumber="81" rank="family" status="fam. nov.">Bornebuschiidae</taxonomicName>
|
||
<emphasis id="B97FAA78466DFFC0C375B00113C6DD5C" bold="true" box="[1150,1250,798,825]" pageId="12" pageNumber="13">fam nov</emphasis>
|
||
. (cf. below) the humeral process also originates in this posterior position but it is rounded apically and does not extend anteriorly beyond the bothridium. In
|
||
<taxonomicName id="4C0B0DE9466DFFC0C6C8B078154DDDE4" authorityName="Mahunka" authorityYear="1986" box="[451,617,871,897]" class="Arachnida" family="Cerocepheidae" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Cerocepheidae</taxonomicName>
|
||
the humeral process is reduced to a short, triangular structure.
|
||
</paragraph>
|
||
<paragraph id="8BB4766A466DFFC0C7CCB0941261DA8C" blockId="12.[151,1437,151,1797]" pageId="12" pageNumber="13">
|
||
The lamellae of
|
||
<taxonomicName id="4C0B0DE9466DFFC0C68DB0941536DDC0" authorityName="J. Balogh" authorityYear="1965" box="[390,530,907,933]" class="Arachnida" family="Eutegaeidae" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Eutegaeidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0B0DE9466DFFC0C52BB09415E2DDC0" authorityName="Mahunka" authorityYear="1986" box="[544,710,907,933]" class="Arachnida" family="Cerocepheidae" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Cerocepheidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0B0DE9466DFFC0C408B0941492DDC0" authority="Colloff, 2023" authorityName="Colloff" authorityYear="2023" box="[771,950,907,933]" family="Neoeutegaeidae" pageId="12" pageNumber="13" rank="family" status="fam. nov.">Neoeutegaeidae</taxonomicName>
|
||
<emphasis id="B97FAA78466DFFC0C4B4B0951314DDC0" bold="true" box="[959,1072,906,933]" pageId="12" pageNumber="13">
|
||
<taxonomicNameLabel id="A24C1703466DFFC0C4B4B0951314DDC0" box="[959,1072,906,933]" pageId="12" pageNumber="13" rank="family">fam. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C0B0DE9466DFFC0C367B0941206DDC0" authority="Colloff, 2023" authorityName="Colloff" authorityYear="2023" box="[1132,1314,907,933]" family="Porrhotegaeidae" pageId="12" pageNumber="13" rank="family" status="fam. nov.">Porrhotegaeidae</taxonomicName>
|
||
<emphasis id="B97FAA78466DFFC0C227B09512B8DDC0" bold="true" box="[1324,1436,906,933]" pageId="12" pageNumber="13">
|
||
<taxonomicNameLabel id="A24C1703466DFFC0C227B09512B8DDC0" box="[1324,1436,906,933]" pageId="12" pageNumber="13" rank="family">fam. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
have free cusps and are broad, not apically convergent, with convex or straight lateral margins and the lamellar seta emerges from an excavation at the apex of the cusp, typically between two teeth, or the cusp is incurved and the lamellar seta emerge sub-apically (
|
||
<taxonomicName id="4C0B0DE9466DFFC0C55AB0E815C7DA74" authorityName="Luxton" authorityYear="1988" box="[593,739,1015,1041]" class="Arachnida" family="Eutegaeidae" genus="Atalotegaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA78466DFFC0C55AB0E815C7DA74" box="[593,739,1015,1041]" italics="true" pageId="12" pageNumber="13">Atalotegaeus</emphasis>
|
||
</taxonomicName>
|
||
). In
|
||
<taxonomicName id="4C0B0DE9466DFFC0C416B0E814FBDA74" authorityName="Luxton" authorityYear="1988" box="[797,991,1015,1041]" family="Compactozetidae" pageId="12" pageNumber="13" rank="family">Compactozetidae</taxonomicName>
|
||
the lamellae are fused with each other apically and curved ventrally and fused with the rostrum in
|
||
<taxonomicName id="4C0B0DE9466DFFC0C431B70414C0DA50" authorityName="Hammer" authorityYear="1966" box="[826,996,1051,1077]" class="Arachnida" family="Cepheusidae" genus="Compactozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA78466DFFC0C431B70414C0DA50" box="[826,996,1051,1077]" italics="true" pageId="12" pageNumber="13">Compactozetes</emphasis>
|
||
</taxonomicName>
|
||
or are slightly convergent apically and have a rounded or obliquely-angled free cusp (
|
||
<taxonomicName id="4C0B0DE9466DFFC0C5A0B7201419DA3C" authorityName="Aoki" authorityYear="1965" box="[683,829,1087,1113]" class="Arachnida" family="Cepheusidae" genus="Sadocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA78466DFFC0C5A0B7201419DA3C" box="[683,829,1087,1113]" italics="true" pageId="12" pageNumber="13">Sadocepheus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0B0DE9466DFFC0C446B72014FDDA3D" authorityName="J. Balogh & Mahunka" authorityYear="1969" box="[845,985,1087,1112]" class="Arachnida" family="Compactozetidae" genus="Hamotegeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA78466DFFC0C446B72014FDDA3D" box="[845,985,1087,1112]" italics="true" pageId="12" pageNumber="13">Hamotegeus</emphasis>
|
||
</taxonomicName>
|
||
). In
|
||
<taxonomicName id="4C0B0DE9466DFFC0C31AB72013BFDA3C" authorityName="Luxton" authorityYear="1988" box="[1041,1179,1087,1113]" class="Arachnida" family="Pterozetidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Pterozetidae</taxonomicName>
|
||
the lamellae are fused with each other apically but not folded ventrally. In
|
||
<taxonomicName id="4C0B0DE9466DFFC0C5D6B77C14BDDA18" authority="Colloff, 2023" authorityName="Colloff" authorityYear="2023" box="[733,921,1123,1149]" family="Bornebuschiidae" pageId="12" pageNumber="81" rank="family" status="fam. nov.">Bornebuschiidae</taxonomicName>
|
||
<emphasis id="B97FAA78466DFFC0C4ABB77D1322DA18" bold="true" box="[928,1030,1122,1149]" pageId="12" pageNumber="13">fam nov.</emphasis>
|
||
the lamellar seta is positioned some distance posterolaterally, at the base of the lamellar cusp, which is either curved and blunt (
|
||
<taxonomicName id="4C0B0DE9466DFFC0C394B798121FDAC4" authorityName="Hammer" authorityYear="1966" box="[1183,1339,1159,1185]" class="Arachnida" family="Cerocepheidae" genus="Bornebuschia" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA78466DFFC0C394B798121FDAC4" box="[1183,1339,1159,1185]" italics="true" pageId="12" pageNumber="13">Bornebuschia</emphasis>
|
||
</taxonomicName>
|
||
) or very long and pointed, extending well beyond the rostrum (
|
||
<taxonomicName id="4C0B0DE9466DFFC0C5F9B7B414ACDAA1" authorityName="Luxton" authorityYear="1988" box="[754,904,1195,1220]" class="Arachnida" family="Cerocepheidae" genus="Dicrotegaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA78466DFFC0C5F9B7B414ACDAA1" box="[754,904,1195,1220]" italics="true" pageId="12" pageNumber="13">Dicrotegaeus</emphasis>
|
||
</taxonomicName>
|
||
). In
|
||
<taxonomicName id="4C0B0DE9466DFFC0C4B7B7B41356DAA0" authority="Colloff, 2023" authorityName="Colloff" authorityYear="2023" box="[956,1138,1195,1221]" family="Porrhotegaeidae" pageId="12" pageNumber="13" rank="family" status="fam. nov.">Porrhotegaeidae</taxonomicName>
|
||
<taxonomicNameLabel id="A24C1703466DFFC0C373B7B513C2DAA0" box="[1144,1254,1194,1221]" pageId="12" pageNumber="13" rank="family">
|
||
<emphasis id="B97FAA78466DFFC0C373B7B513C4DAA0" bold="true" box="[1144,1248,1194,1221]" pageId="12" pageNumber="13">fam. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
the lamellae are similar to those of
|
||
<taxonomicName id="4C0B0DE9466DFFC0C662B7D016D1DA8C" authorityName="J. Balogh" authorityYear="1965" box="[361,501,1231,1257]" class="Arachnida" family="Eutegaeidae" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Eutegaeidae</taxonomicName>
|
||
, but the lamellar seta emerges from the ventral surface of the lamellar cusp.
|
||
</paragraph>
|
||
<paragraph id="8BB4766A466DFFC0C7CCB7EC14F6D960" blockId="12.[151,1437,151,1797]" pageId="12" pageNumber="13">
|
||
The margins of the coxisternum, comprising pedotecta I and II and the discidium, are relatively uniform in morphology in
|
||
<taxonomicName id="4C0B0DE9466DFFC0C647B60816F1DB54" authorityName="J. Balogh" authorityYear="1965" box="[332,469,1303,1329]" class="Arachnida" family="Eutegaeidae" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Eutegaeidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0B0DE9466DFFC0C51FB608159EDB54" authorityName="Mahunka" authorityYear="1986" box="[532,698,1303,1329]" class="Arachnida" family="Cerocepheidae" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Cerocepheidae</taxonomicName>
|
||
when viewed ventrally. The morphology of these structures in ventral aspect does not fully reveal their three-dimensional shape, which becomes more apparent in lateral aspect (compare
|
||
<figureCitation id="13306AEF466DFFC0C601B64016F3DB1C" box="[266,471,1375,1401]" captionStart="FIGURE 27" captionStartId="52.[151,250,1662,1686]" captionTargetBox="[232,1356,381,1639]" captionTargetId="figure-126@52.[232,1356,381,1639]" captionTargetPageId="52" captionText="FIGURE 27. Scanning electron micrographs of Neoeutegaeus sp. from O’Reilly’s Rainforest Retreat, Lamington National Park, Queensland (from Hunt et al., 1998b)." figureDoi="http://doi.org/10.5281/zenodo.10147023" httpUri="https://zenodo.org/record/10147023/files/figure.png" pageId="12" pageNumber="13">Figs. 27b and 27c</figureCitation>
|
||
) but, as mentioned in the methods section above, lateral aspects were not illustrated because of the difficulty in making temporary mounts of specimens in which the hysterosoma is broader than long. However, the ventral aspects of these structures (
|
||
<figureCitation id="13306AEF466DFFC0C409B6B81470DBA4" box="[770,852,1447,1473]" captionStart="FIGURE 44" captionStartId="85.[151,250,1671,1695]" captionTargetBox="[235,1352,957,1648]" captionTargetId="figure-403@85.[235,1352,957,1648]" captionTargetPageId="85" captionText="FIGURE 44. Distribution of type localities of species of Eutegaeoidea superimposed on a map of Gondwana prior to its break-up during the middle Jurassic (re-drawn from Veevers, 2000). Note that only genera present on more than one former Gondwanan land mass are included." figureDoi="http://doi.org/10.5281/zenodo.10147057" httpUri="https://zenodo.org/record/10147057/files/figure.png" pageId="12" pageNumber="13">Fig. 44</figureCitation>
|
||
) reveals useful similarities and differences between taxa. Pedoctectum I is sub-rectangular in all genera of
|
||
<taxonomicName id="4C0B0DE9466DFFC0C40CB6D414B4DB80" authorityName="J. Balogh" authorityYear="1965" box="[775,912,1483,1509]" class="Arachnida" family="Eutegaeidae" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Eutegaeidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0B0DE9466DFFC0C4C2B6D4134BDB80" authorityName="Mahunka" authorityYear="1986" box="[969,1135,1483,1509]" class="Arachnida" family="Cerocepheidae" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Cerocepheidae</taxonomicName>
|
||
and the anterior margin is transverse or oblique. Pedotectum II is pointed and triangular/rectangular in
|
||
<taxonomicName id="4C0B0DE9466DFFC0C4E9B6F01350D86C" authorityName="Luxton" authorityYear="1988" box="[994,1140,1519,1545]" class="Arachnida" family="Eutegaeidae" genus="Atalotegaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA78466DFFC0C4E9B6F01350D86C" box="[994,1140,1519,1545]" italics="true" pageId="12" pageNumber="13">Atalotegaeus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0B0DE9466DFFC0C3A3B6F0121DD86C" authorityName="TRAGARDH" authorityYear="1931" box="[1192,1337,1519,1545]" class="Arachnida" family="Cerocepheidae" genus="Cerocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA78466DFFC0C3A3B6F0121DD86C" box="[1192,1337,1519,1545]" italics="true" pageId="12" pageNumber="13">Cerocepheus</emphasis>
|
||
</taxonomicName>
|
||
, in those species of
|
||
<taxonomicName id="4C0B0DE9466DFFC0C605B50C16A7D849" authorityName="Berlese" authorityYear="1916" box="[270,387,1555,1580]" class="Arachnida" family="Eutegaeidae" genus="Eutegaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA78466DFFC0C605B50C16A7D849" box="[270,387,1555,1580]" italics="true" pageId="12" pageNumber="13">Eutegaeus</emphasis>
|
||
</taxonomicName>
|
||
for which the ventral surface has been described and in
|
||
<taxonomicName id="4C0B0DE9466DFFC0C4F4B50C13BDD849" authorityName="Woolley" authorityYear="1965" box="[1023,1177,1555,1580]" class="Arachnida" family="Eutegaeidae" genus="Neseutegaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97FAA78466DFFC0C4F4B50C13BDD849" box="[1023,1177,1555,1580]" italics="true" pageId="12" pageNumber="13">Neseutegaeus</emphasis>
|
||
</taxonomicName>
|
||
. The discidium is thin, triangular and pointed in
|
||
<taxonomicName id="4C0B0DE9466DFFC0C6B8B528157DD834" authorityName="Mahunka" authorityYear="1986" box="[435,601,1591,1617]" class="Arachnida" family="Cerocepheidae" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Cerocepheidae</taxonomicName>
|
||
and all species of
|
||
<taxonomicName id="4C0B0DE9466DFFC0C420B5281490D834" authorityName="J. Balogh" authorityYear="1965" box="[811,948,1591,1617]" class="Arachnida" family="Eutegaeidae" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Eutegaeidae</taxonomicName>
|
||
for which the structure has been described. In
|
||
<taxonomicName id="4C0B0DE9466DFFC0C7B2B5441648D810" authority="Colloff, 2023" authorityName="Colloff" authorityYear="2023" box="[185,364,1627,1653]" family="Neoeutegaeidae" pageId="12" pageNumber="13" rank="family" status="fam. nov.">Neoeutegaeidae</taxonomicName>
|
||
<taxonomicNameLabel id="A24C1703466DFFC0C67EB54516CCD810" box="[373,488,1626,1653]" pageId="12" pageNumber="13" rank="family">
|
||
<emphasis id="B97FAA78466DFFC0C67EB54516C5D810" bold="true" box="[373,481,1626,1653]" pageId="12" pageNumber="13">fam. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
pd I is sub-rectangular, pd II is massive and either trapezoid or lobe-like and the discidium is lobe-like or sub-circular. In
|
||
<taxonomicName id="4C0B0DE9466DFFC0C56EB560140ED8FC" authorityName="Luxton" authorityYear="1988" box="[613,810,1663,1689]" family="Compactozetidae" pageId="12" pageNumber="13" rank="family">Compactozetidae</taxonomicName>
|
||
, pd I has a prominent anterior spine, pd II is very large and lobe-like or rectangular and the discidium is lobe-like or sub-triangular. In
|
||
<taxonomicName id="4C0B0DE9466DFFC0C301B5BC13B0D8D8" authorityName="Luxton" authorityYear="1988" box="[1034,1172,1699,1725]" class="Arachnida" family="Pterozetidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Pterozetidae</taxonomicName>
|
||
pd I is sub-rectangular, pd II is rectangular and the discidium is rectangular or trapezoid. In
|
||
<taxonomicName id="4C0B0DE9466DFFC0C49FB5D8136ED884" authority="Colloff, 2023" authorityName="Colloff" authorityYear="2023" box="[916,1098,1735,1761]" family="Porrhotegaeidae" pageId="12" pageNumber="13" rank="family" status="fam. nov.">Porrhotegaeidae</taxonomicName>
|
||
<emphasis id="B97FAA78466DFFC0C35AB5D9139BD884" bold="true" box="[1105,1215,1734,1761]" pageId="12" pageNumber="13">
|
||
<taxonomicNameLabel id="A24C1703466DFFC0C35AB5D9139BD884" box="[1105,1215,1734,1761]" pageId="12" pageNumber="13" rank="family">fam. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
pd I and II are sub-rectangular, pd I lacks a spine and the discidium is triangular and pointed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |