171 lines
16 KiB
XML
171 lines
16 KiB
XML
<document id="F2B003C49DDE6CC78A41F63C4C555194" ID-DOI="10.3897/zookeys.1230.136906" ID-ZooBank="0FBF41B0-1D35-4A31-9653-B1F475FBDF80" ID-publisher-id="136906" URI-arpha="3B6146E9-7C33-583C-A633-97ED3C18A10A" URI-zoobank="http://zoobank.org/0FBF41B0-1D35-4A31-9653-B1F475FBDF80" article-type="research-article" checkinTime="1741198404937" checkinUser="pensoft" docAuthor="Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J. & Liew, Thor-Seng" docDate="2025" docId="DA698A0517A658D987281C376E2F7279" docLanguage="en" docName="ZooKeys 1230: 99-154" docOrigin="ZooKeys 1230" docSource="https://zookeys.pensoft.net/article/136906/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Atopos tourannensis BOR" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="154" masterDocId="3B6146E97C33583CA63397ED3C18A10A" masterDocTitle="From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia" masterLastPageNumber="154" masterPageNumber="99" pageNumber="99" updateTime="1741199410933" updateUser="ExternalLinkService">
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<mods:titleInfo id="40365C6A7C22F17D68E9A6056503E550">
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<mods:title id="26776D08F31F28265C7AE6046F84B15C">From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia</mods:title>
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<mods:name id="ED8FA7128AB8C20E50B145844FDD3F58" type="personal">
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<mods:roleTerm id="E6DE15CA04CBE6D918474697C928935C">Author</mods:roleTerm>
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<mods:namePart id="D7A4A7D96F5EAA0F5950734D1EF0E727">Goh, Soo-Mun</mods:namePart>
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<mods:affiliation id="A83B698709D94B4260B0E1970DC6C2B9">Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu, Malaysia</mods:affiliation>
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</mods:name>
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<mods:name id="B951A7FA9655B77EB26FB29BB849F5BE" type="personal">
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<mods:roleTerm id="AA3B375A33EC286E5119E5C469B7CE0B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="330955139469090E409008B04387747E">Jasrul, Dulipat</mods:namePart>
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<mods:affiliation id="936D20CCB241FFBF4F783D6BAA324E6F">Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu, Malaysia</mods:affiliation>
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</mods:name>
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<mods:name id="70C487F120C669EB90276C4390B8AAC1" type="personal">
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<mods:roleTerm id="5842C53F81333E08EC80EC2D7E9F81FE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="106A2C4421CD4EFAE17E19D175B34EE7">Lee, Mei-Yi</mods:namePart>
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<mods:nameIdentifier id="9F041EFD712F8986B1C1ABF0C62F0069" type="ORCID">https://orcid.org/0009-0005-8200-3508</mods:nameIdentifier>
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<mods:affiliation id="053E13EC03AD190C672CB30AFA3620FE">School of Biological Science, Universiti Sains Malaysia, Penang, Malaysia</mods:affiliation>
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</mods:name>
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<mods:name id="EABFAE77BD382FC89CEC7E6EA549F287" type="personal">
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<mods:roleTerm id="CEF612EA5B3FFCB51DA60FF415FAD962">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9C83666F552CC2F2F32514852B9E6864">Vermeulen, Jaap J.</mods:namePart>
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<mods:affiliation id="A89A0FCFBAB3B6C2E243F744939B1AD8">jk. artandscience, Lauwerbes 8, 2318 AT, Leiden, Netherlands</mods:affiliation>
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</mods:name>
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<mods:namePart id="5EDF11D30918C31C11D3D557F1A55599">Liew, Thor-Seng</mods:namePart>
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<mods:nameIdentifier id="5198D643B5335C00052FEC534C91E8F3" type="ORCID">0000-0002-9437-5924</mods:nameIdentifier>
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<mods:affiliation id="355603C2A60CA95BE2CA6C68D3FB269D">Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu, Malaysia</mods:affiliation>
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<mods:typeOfResource id="C0828C4C0C5FEE94403E27EAB0FA9301">text</mods:typeOfResource>
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<mods:titleInfo id="3939EEEEA4324E818157AFDA88FF32D7">
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<mods:title id="24915073DAAB27F3DDB0018B78996D94">ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="FD01B3F9ECEFBEE9AC1BC2A57DEB7231">
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<mods:date id="D0C1DD13979F7330210591922CE6C076">2025</mods:date>
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<mods:detail id="38CC75174EA03C6C4955182DD0E1CB8E" type="pubDate">
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<mods:number id="F2802B2B44B90218A3D78B3B54A2023C">2025-03-05</mods:number>
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<mods:detail id="74D38545184FA9CD28E6B645C40C2879" type="volume">
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<mods:number id="407751CA1EE17F69C4E9B201FB6105BC">1230</mods:number>
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<mods:start id="C70CF8B489F8B14EE8EDBEEAF258D3E0">99</mods:start>
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<mods:end id="CF3E3A838070EA2C05B2FB72F5B02CF8">154</mods:end>
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<mods:classification id="1F2CB6B8544E9EB5EC302A105761DBBC">journal article</mods:classification>
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<mods:identifier id="4C473222B14AC7E0ABE22F07FFDA494A" type="DOI">10.3897/zookeys.1230.136906</mods:identifier>
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<mods:identifier id="2610F6BF0CC9FC82A192B22134F193B9" type="ZooBank">0FBF41B0-1D35-4A31-9653-B1F475FBDF80</mods:identifier>
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</mods:mods>
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<treatment id="DA698A0517A658D987281C376E2F7279" ID-DOI="http://doi.org/10.5281/zenodo.14976216" ID-Zenodo-Dep="14976216" ID-arpha="DA698A05-17A6-58D9-8728-1C376E2F7279" LSID="urn:lsid:plazi:treatment:DA698A0517A658D987281C376E2F7279" httpUri="http://treatment.plazi.org/id/DA698A0517A658D987281C376E2F7279">
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<subSubSection id="D8AAB58E91475FE12672241999C2BF28" type="nomenclature">
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<paragraph id="1BD7296B05F25F8D0255B4629278BE92">
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<taxonomicName id="C41313CBEB935A0C4765C105A4F1B7C2" ID-CoL="7Q8TP" authority="(Souleyet, 1852)" authorityName="BOR" baseAuthorityName="Souleyet" baseAuthorityYear="1852" class="Gastropoda" family="Rathouisiidae" genus="Atopos" kingdom="Animalia" order="Systellommatophora" phylum="Mollusca" rank="species" species="tourannensis">
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<emphasis id="9AC018CB493C08C4046008CC3D9376B5" italics="true">Atopos tourannensis</emphasis>
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(
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<bibRefCitation id="690B02E68D283EAC69B2E6932E13821B" author="Souleyet LFA" journalOrPublisher="Arthus Bertrand, Paris" refId="B43" refString="Souleyet LFA (1852) Tome Deuxieme. Mollusques. In: Eydoux F, Souleyet LFA (1841 - 1852). Voyage autour du monde execute pendant les annees 1836 et 1837 sur la corvette La Bonite. Zoologie. Arthus Bertrand, Paris, 664 pp. https://www.biodiversitylibrary.org/page/28264738" title="Tome Deuxième. Mollusques. In: Eydoux F, Souleyet LFA (1841-1852). Voyage autour du monde exécuté pendant les années 1836 et 1837 sur la corvette La Bonite. Zoologie." url="https://www.biodiversitylibrary.org/page/28264738" year="1852">Souleyet, 1852</bibRefCitation>
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)
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="7430CF3650C7A8969E0FE128F92D1F03" type="description">
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<paragraph id="CE5073040C4236E8C28BB257AD7783B6">
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<figureCitation id="85ACF2DDABB2EA8F578561A927ADCA7D" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. A Atopos tourannensis (Souleyet, 1852) BOR / MOL 15227 B Atopos punctata Collinge, 1902 BOR / MOL 15228 C Laevicaulis alte (A. Férussac, 1822) BOR / MOL 15773. Scale bar: 10 mm." figureDoi="10.3897/zookeys.1230.136906.figure18" httpUri="https://binary.pensoft.net/fig/1270453">Figs 18 A</figureCitation>
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,
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<figureCitation id="15711BCBC6E0F6068A31F8B2BCA7C8EE" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. A Pseudoplecta bijuga (Stoliczka, 1873) B Atopos tourannensis (Souleyet, 1852) C Atopos punctata Collinge, 1902 D Laevicaulis alte (A. Férussac, 1822) E Meghimatium pictum (Stoliczka, 1873), photo credit Xin Wei Ooi." figureDoi="10.3897/zookeys.1230.136906.figure22" httpUri="https://binary.pensoft.net/fig/1270457">22 B</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="SECID0E1RDK" type="material">
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<paragraph id="B29D2A250C74E74BB275DA36767C0678">
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<heading id="C742095BD8E0A3147DC407E201FF6F24" reason="title">Material examined.</heading>
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</paragraph>
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<paragraph id="75EA404FD1BFA0C42EA7DA0012DDA97F">
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<materialsCitation id="8B3BF74FB33C535B227ADCEBF3DD1F62" collectingDate="2022-10-22" collectionCode="T" collectorName="S. Liew & J. Dulipat" country="Malaysia" county="Hill" elevation="180" latitude="5.418" location="Entrance" longLatPrecision="55" longitude="100.2159" municipality="Teluk Bahang-Balik Pulau" specimenCount="1" stateProvince="Pulau Pinang">
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<collectingCountry id="67007EB9B03F4AA641D1763A32A57527" name="Malaysia">Malaysia</collectingCountry>
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•
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<collectingRegion id="E2D0FAD40748123ACDDCEB2AE6D3D4A0" country="Malaysia" name="Pulau Pinang">Penang</collectingRegion>
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<collectingCounty id="F7FD6E2E5F6CCBC77E29E5ED737C8F27">Hill</collectingCounty>
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,
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<collectingMunicipality id="7B347BEE65BDDE5609C4F97C57BDF5C1">Teluk Bahang-Balik Pulau</collectingMunicipality>
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,
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<location id="5CC3A8A1AA1EC2540FED910493546A7D" LSID="urn:lsid:plazi:treatment:DA698A0517A658D987281C376E2F7279:5CC3A8A1AA1EC2540FED910493546A7D" country="Malaysia" county="Hill" latitude="5.418" longLatPrecision="55" longitude="100.2159" municipality="Teluk Bahang-Balik Pulau" name="Entrance" stateProvince="Pulau Pinang">Entrance</location>
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of a durian farm, near
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<location id="55C6BACC4C340C820EBD45146D8DFD3C" LSID="urn:lsid:plazi:treatment:DA698A0517A658D987281C376E2F7279:55C6BACC4C340C820EBD45146D8DFD3C" country="Malaysia" county="Hill" latitude="5.418" longLatPrecision="55" longitude="100.2159" municipality="Teluk Bahang-Balik Pulau" name="Boulder Valley" stateProvince="Pulau Pinang">Boulder Valley</location>
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,
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<location id="BDF324D09409A5911CF42D0EF39104D0" LSID="urn:lsid:plazi:treatment:DA698A0517A658D987281C376E2F7279:BDF324D09409A5911CF42D0EF39104D0" country="Malaysia" county="Hill" latitude="5.418" longLatPrecision="55" longitude="100.2159" municipality="Teluk Bahang-Balik Pulau" name="Old" stateProvince="Pulau Pinang">Old</location>
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rubber tree, Plot. 17;
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<named-content content-type="dwc:verbatimCoordinates" id="NCID0EDSDK" specific-use="{"type":"Point","coordinates":[100.215900,5.418000]}">
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<geoCoordinate id="FB01AB85F71E766BC4F8C881C780C543" degrees="5.418" direction="north" orientation="latitude" precision="55" value="5.418">5.418 ° N</geoCoordinate>
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,
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<geoCoordinate id="4212F38B8C7C24ED205820ED118431E8" degrees="100.2159" direction="east" orientation="longitude" precision="5" value="100.2159">100.2159 ° E</geoCoordinate>
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</named-content>
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;
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<elevation id="A54D6569160586A9541684FF0139E9C9" metricMagnitude="2" metricUnit="m" metricValue="1.8" unit="m" value="180.0">
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<quantity id="3325A7D967B0EDC427B46A6449B971FF" metricMagnitude="2" metricUnit="m" metricValue="1.8" unit="m" value="180.0">180 m</quantity>
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a. s. l.
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</elevation>
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;
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<date id="918A242B10101C0384B873499C42095C" value="2022-10-22">
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<collectingDate id="469F4F5405CA036CC863C591DD782192" value="2022-10-22">22 Oct. 2022</collectingDate>
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</date>
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;
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<collectionCode id="C9E2BCCB08BA0ECFC87E5DA646443874" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
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.
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<collectorName id="587E178C413A5716ABB8960819C35A44">S. Liew</collectorName>
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,
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<collectorName id="2444929A2533C079C72790E73FEC7B0D">J. Dulipat</collectorName>
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leg.; BOR / MOL 15227
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</materialsCitation>
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.
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</paragraph>
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<caption id="882B2989C9D7CF0046BDBC2C4E198CA8" ID-DOI="10.3897/zookeys.1230.136906.figure18" ID-arpha="B59C1C31-9E25-5361-BA41-9ED93FD9FE99" httpUri="https://binary.pensoft.net/fig/1270453" startId="F18">
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<paragraph id="BFA58362FDAA6301E1C38834E5586895">
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<label id="56AB888DC88654733548E001F65C2689">Figure 18.</label>
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</paragraph>
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<paragraph id="573A3B920E34854D4DA9D2AF415E5378">
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<emphasis id="E5800555913FE0447C1C99E28D1C8EFB" bold="true">A</emphasis>
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<taxonomicName id="7DCF73F321AAFF346985BA5A4ECCB2E2" authority="(Souleyet, 1852) BOR" authorityName="BOR" baseAuthorityName="Souleyet" baseAuthorityYear="1852" class="Gastropoda" family="Rathouisiidae" genus="Atopos" kingdom="Animalia" order="Systellommatophora" phylum="Mollusca" rank="species" species="tourannensis">
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<emphasis id="684AB81642E285127C6F80AF11967299" italics="true">Atopos tourannensis</emphasis>
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(
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<bibRefCitation id="7AA34F772AC07799CB2CA8E246371C34" author="Souleyet LFA" journalOrPublisher="Arthus Bertrand, Paris" refId="B43" refString="Souleyet LFA (1852) Tome Deuxieme. Mollusques. In: Eydoux F, Souleyet LFA (1841 - 1852). Voyage autour du monde execute pendant les annees 1836 et 1837 sur la corvette La Bonite. Zoologie. Arthus Bertrand, Paris, 664 pp. https://www.biodiversitylibrary.org/page/28264738" title="Tome Deuxième. Mollusques. In: Eydoux F, Souleyet LFA (1841-1852). Voyage autour du monde exécuté pendant les années 1836 et 1837 sur la corvette La Bonite. Zoologie." url="https://www.biodiversitylibrary.org/page/28264738" year="1852">Souleyet, 1852</bibRefCitation>
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) BOR
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</taxonomicName>
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/ MOL 15227
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<emphasis id="D81FDB7E6CD576D85A05A8E62E62C83D" bold="true">B</emphasis>
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<taxonomicName id="79540A4A4F3CE2664F29FC0BEFD9853B" authority="Collinge, 1902 BOR" authorityName="Collinge" authorityYear="1902" class="Gastropoda" family="Rathouisiidae" genus="Atopos" kingdom="Animalia" order="Systellommatophora" phylum="Mollusca" rank="species" species="punctata">
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<emphasis id="091DC811F7E7A84247DDA0496A3AB67C" italics="true">Atopos punctata</emphasis>
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<bibRefCitation id="2E177FBA7AC5E24E1EE98E3D5E3D57C9" author="Collinge WE" journalOrPublisher="The Journal of Malacology" pagination="71-95" refId="B13" refString="Collinge WE (1902) On the non-operculate land and freshwater Molluscs collected by the members of the " Skeat-Expedition " in the Malay Peninsula, 1899 - 1900. The Journal of Malacology 9: 71-95. https://www.biodiversitylibrary.org/item/89648#page/253/mode/1up" title="On the non-operculate land and freshwater Molluscs collected by the members of the “ Skeat-Expedition ” in the Malay Peninsula, 1899-1900." url="https://www.biodiversitylibrary.org/item/89648#page/253/mode/1up" volume="9" year="1902">Collinge, 1902</bibRefCitation>
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BOR
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</taxonomicName>
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/ MOL 15228
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<emphasis id="E9AC1DB5CD83A4D832744C9B02D57FAD" bold="true">C</emphasis>
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<taxonomicName id="592060C24E8478050AA93E0C4AFF7A5E" authority="(A. Ferussac, 1822) BOR" authorityName="BOR" baseAuthorityName="A. Ferussac" baseAuthorityYear="1822" class="Gastropoda" family="Veronicellidae" genus="Laevicaulis" kingdom="Animalia" order="Systellommatophora" phylum="Mollusca" rank="species" species="alte">
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<emphasis id="C025EBC3546DCE23C9A398C99C19E022" italics="true">Laevicaulis alte</emphasis>
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(A. Férussac, 1822) BOR
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</taxonomicName>
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/ MOL 15773. Scale bar: 10 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="SECID0E3TDK" type="remarks">
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<paragraph id="B1BF65D3244C6511400489FD77539A5F">
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<heading id="A6480F7CA4EB5FFA467A38DC2F260C85" reason="title">Remarks.</heading>
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</paragraph>
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<paragraph id="1AA8FE8D384FA610256E37CAABB39030">
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This is a new record for
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<collectingRegion id="2E352FE94BEEEEE3635B1A0BCB8F4306" country="Malaysia" name="Pulau Pinang">Penang</collectingRegion>
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Hill. This species differs from
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<taxonomicName id="AA3152E0498EC0FFECA2886DB4F92BDE" authorityName="Collinge" authorityYear="1902" class="Gastropoda" family="Rathouisiidae" genus="Atopos" kingdom="Animalia" order="Systellommatophora" phylum="Mollusca" rank="species" species="punctata">
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<emphasis id="7C162F59EF4C6B76F95E6EDF26AA759C" italics="true">Atopos punctata</emphasis>
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</taxonomicName>
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by having a black mantle with white areas at both ends (head and tail) of the mantle.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |