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<mods:title id="4F8B2261927B6B8D63B861187B757EA0">From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia</mods:title>
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<mods:namePart id="4AEF647479CD226B487AA1CC4AA64275">Goh, Soo-Mun</mods:namePart>
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<taxonomicName id="14D8BAA73A8BEB725D68C6ED0B3F65F9" authority="Stoliczka, 1873" authorityName="Stoliczka" authorityYear="1873" class="Gastropoda" family="Helicarionidae" genus="Helicarion" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="permolle">
<emphasis id="0AA80FF90958892B2C7B7C60D82ADE4B" italics="true">Helicarion permolle</emphasis>
<bibRefCitation id="0CE064AF3B8F9CFE0DBB1669A1778748" author="Stoliczka F" journalOrPublisher="Journal of the Asiatic Society of Bengal" pagination="11-38" refId="B48" refString="Stoliczka F (1873) On the land shells of Penang Island, with descriptions of the animals and anatomical notes; part second, Helicacea. Journal of the Asiatic Society of Bengal 42: 11-38. https://www.biodiversitylibrary.org/page/35546163#page/25/mode/1up" title="On the land shells of Penang Island, with descriptions of the animals and anatomical notes; part second, Helicacea." url="https://www.biodiversitylibrary.org/page/35546163#page/25/mode/1up" volume="42" year="1873">Stoliczka, 1873</bibRefCitation>
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<figureCitation id="3DAA6A814E185E1E2E042974B9BAEFEA" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A Tanychlamys Penang sp. 2 BOR / MOL 15759 B Tanychlamys stephoides (Stoliczka, 1873) BOR / MOL 15164 C Tanychlamys tersa (Issel, 1874) BOR / MOL 15533 D Helicarion permolle Stoliczka, 1873 BOR / MOL 15741 E Bradybaena similaris (A. Férussac, 1821) BOR / MOL 15772 F Trichochloritis penangensis (Stoliczka, 1873) BOR / MOL 15291. Scale bars: 1 mm (A C); 10 mm (D F)." figureDoi="10.3897/zookeys.1230.136906.figure11" httpUri="https://binary.pensoft.net/fig/1270446">Figs 11 D</figureCitation>
,
<figureCitation id="7E665BD142068833C29D8D3B5AACC2CE" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. A Tanychlamys Penang sp. 2 B Tanychlamys tersa (Issel, 1874) C Helicarion permolle Stoliczka, 1873 D Bradybaena similaris (A. Férussac, 1821) E Vitrinopsis sp. F Vitrinopsis nucleata (Stoliczka, 1873)." figureDoi="10.3897/zookeys.1230.136906.figure21" httpUri="https://binary.pensoft.net/fig/1270456">21 C</figureCitation>
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<heading id="A9ABFD87271B8AADE82AF293C1489B38" reason="title">Material examined.</heading>
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<elevation id="010C22FDD9946F7E8C01443B01172DBB" metricMagnitude="1" metricUnit="m" metricValue="8.5" unit="m" value="85.0">
<quantity id="162BE2BFAAA3E15D1D5199855799064C" metricMagnitude="1" metricUnit="m" metricValue="8.5" unit="m" value="85.0">85 m</quantity>
a. s. l.
</elevation>
;
<date id="44354AF1D85D6CD5B1333C4C880AC5AC" value="2022-09-04">
<collectingDate id="9B2F012A2D0741446B64275A16345891" value="2022-09-04">4 Sept. 2022</collectingDate>
</date>
;
<collectionCode id="4FE1501D0C1CF6DEC057CF6C9499F9EE" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
.
<collectorName id="67A4FD6EDE5128505831CB020622CCBC">S. Liew</collectorName>
,
<collectorName id="E129E146443C9C0464CEC0C0EA6EE33B">J. Dulipat</collectorName>
,
<collectorName id="CC05549A423EA28F3D4D2F573656935C">S. M. Goh</collectorName>
leg.; BOR / MOL 15121
</materialsCitation>
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,
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from
<location id="86CF7D27A7752163592369D36982A3E0" LSID="urn:lsid:plazi:treatment:BA505C825B745084AAF275A2C6081E18:86CF7D27A7752163592369D36982A3E0" country="Malaysia" county="Hill" latitude="5.4205" longLatPrecision="7" longitude="100.2724" municipality="Trail" name="Viaduct" stateProvince="Pulau Pinang">Viaduct</location>
to
<location id="DBA9EFF5E79AE76726C6962C25582C41" LSID="urn:lsid:plazi:treatment:BA505C825B745084AAF275A2C6081E18:DBA9EFF5E79AE76726C6962C25582C41" country="Malaysia" county="Hill" latitude="5.4205" longLatPrecision="7" longitude="100.2724" municipality="Trail" name="Claremont" stateProvince="Pulau Pinang">Claremont</location>
, after shelter,
<location id="512F180433C6155A324166F214B2092B" LSID="urn:lsid:plazi:treatment:BA505C825B745084AAF275A2C6081E18:512F180433C6155A324166F214B2092B" country="Malaysia" county="Hill" latitude="5.4205" longLatPrecision="7" longitude="100.2724" municipality="Trail" name="Plot." stateProvince="Pulau Pinang">Plot.</location>
27;
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<geoCoordinate id="8D4CAB3377B900C8BA9D7EB0BA0A83D3" degrees="5.4205" direction="north" orientation="latitude" precision="5" value="5.4205">5.4205 ° N</geoCoordinate>
,
<geoCoordinate id="9B7C78D99468409C28F3FAAB3A17B9AE" degrees="100.2724" direction="east" orientation="longitude" precision="5" value="100.2724">100.2724 ° E</geoCoordinate>
</named-content>
;
<elevation id="FAA2057DB8C2EA08115D159C061366F8" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">
<quantity id="4F705486A9B95E77C9323B8805B1A6E9" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">500 m</quantity>
a. s. l.
</elevation>
;
<date id="B1FD792030F90F117E0F2F92A5753584" value="2022-10-26">
<collectingDate id="7719BB20C6CDC6639FEED50B6DBCBA59" value="2022-10-26">26 Oct. 2022</collectingDate>
</date>
;
<collectionCode id="D96F9F527691E55C0D8B75FE80AE6A29" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
.
<collectorName id="9D674E804C5B97B09200E7BE6104E708">S. Liew</collectorName>
,
<collectorName id="CCEA1EF49715FBF5E27A69B00F124E2F">J. Dulipat</collectorName>
leg.; BOR / MOL 15235
</materialsCitation>
<materialsCitation id="59B429B16B97ADD9E9B01500CD68667E" collectingDate="2023-02-02" collectionCode="T" collectorName="S. Liew &amp; M. Y. Lee &amp; A. Yusni" country="Malaysia" county="Hill" elevation="75" latitude="5.44" location="Plot." longLatPrecision="7" longitude="100.2866" municipality="Botanical" specimenCount="1" stateProvince="Pulau Pinang">
<collectingRegion id="B17944C6C434A1239E0D6E380844CF2F" country="Malaysia" name="Pulau Pinang">Penang</collectingRegion>
<collectingCounty id="6C1370EFA178E81368E88A5E551F106E">Hill</collectingCounty>
,
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garden,
<location id="36F06BE01A4ECDCAE8CE4D57CB0B12DE" LSID="urn:lsid:plazi:treatment:BA505C825B745084AAF275A2C6081E18:36F06BE01A4ECDCAE8CE4D57CB0B12DE" country="Malaysia" county="Hill" latitude="5.44" longLatPrecision="7" longitude="100.2866" municipality="Botanical" name="Plot." stateProvince="Pulau Pinang">Plot.</location>
29;
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<geoCoordinate id="517221B834368853A2901DCAFF0F1A30" degrees="5.4400" direction="north" orientation="latitude" precision="5" value="5.44">5.4400 ° N</geoCoordinate>
,
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</named-content>
;
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<quantity id="B8CE966687AF5350A7B327138DC7F17B" metricMagnitude="1" metricUnit="m" metricValue="7.5" unit="m" value="75.0">75 m</quantity>
a. s. l.
</elevation>
;
<date id="C59FF5E9F991BFAD0E6344B6B56B8039" value="2023-02-02">
<collectingDate id="96FCE0C986B99D43B9DE2C1715EA8157" value="2023-02-02">2 Feb. 2023</collectingDate>
</date>
;
<collectionCode id="348292BE83281176EC65E860F8221D1A" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
.
<collectorName id="4FAA0A14CCF22589370B7A0F92899E61">S. Liew</collectorName>
,
<collectorName id="7E3C60ED8BD75515B7F0695D02E28CE8">M. Y. Lee</collectorName>
,
<collectorName id="D2E22EE124DEAF95CECF2AF2D8783FBF">A. Yusni</collectorName>
leg.; BOR / MOL 15742
</materialsCitation>
<materialsCitation id="AF9ECC1D009CD9C6A8B9AD3237711BF1" collectingDate="2023-02-02" collectionCode="T" collectorName="S. Liew &amp; M. Y. Lee &amp; A. Yusni" country="Malaysia" county="Hill" elevation="75" latitude="5.44" location="Plot." longLatPrecision="7" longitude="100.2866" municipality="Botanical" specimenCount="1" stateProvince="Pulau Pinang">same data as for preceding; BOR / MOL 15768</materialsCitation>
<materialsCitation id="EB2CA56F68318C13884AF91EBFFA3070" collectingDate="2023-02-04" collectionCode="T" collectorName="Hill &amp; By Path H &amp; Plot. &amp; S. Liew &amp; A. Yusni" country="Malaysia" elevation="610" latitude="5.421" location="Penang" longLatPrecision="7" longitude="100.2701" specimenCount="1" stateProvince="Pulau Pinang">
<collectingRegion id="B8EE5E4CF9B045EC03761742E0F764B8" country="Malaysia" name="Pulau Pinang">Penang</collectingRegion>
<collectorName id="A02C4B94E6A427B5A97446E3065194DA">Hill</collectorName>
,
<collectorName id="0EEEA288A2F172337E29236560CB056F">By Path H</collectorName>
,
<collectorName id="A4DD63C83C1F486354F159CC8F7A69E9">Plot.</collectorName>
33;
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<geoCoordinate id="5D7E6FB9AE0DCC383D97ED71462B3727" degrees="5.4210" direction="north" orientation="latitude" precision="5" value="5.421">5.4210 ° N</geoCoordinate>
,
<geoCoordinate id="99638D33A213D96510EB7A0B7F81910E" degrees="100.2701" direction="east" orientation="longitude" precision="5" value="100.2701">100.2701 ° E</geoCoordinate>
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;
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<quantity id="E62609AEF1B78FDAF553794106847F0C" metricMagnitude="2" metricUnit="m" metricValue="6.1" unit="m" value="610.0">610 m</quantity>
a. s. l.
</elevation>
;
<date id="D076747B4D4A456871E0038EB3AA834A" value="2023-02-04">
<collectingDate id="E2FD87DA93BBAEBC75E6782E8B9A82B7" value="2023-02-04">4 Feb. 2023</collectingDate>
</date>
;
<collectionCode id="F7C5C711DABED1EB7CE15C6D04637594" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
.
<collectorName id="A035D95F3109A2F5EC835AEABADFE3E7">S. Liew</collectorName>
,
<collectorName id="CE4F2F1C5C73A32E93CE32685DB703B2">A. Yusni</collectorName>
leg.; BOR / MOL 15741
</materialsCitation>
<materialsCitation id="BCC003CD6FED3157568ED2D4E65E759E" collectingDate="2022-09-04" collectionCode="T" collectorName="Hill &amp; Path B &amp; Random &amp; S. Liew &amp; J. Dulipat" country="Malaysia" elevation="730" latitude="5.4216" location="Penang" longLatPrecision="7" longitude="100.2674" specimenCount="1" stateProvince="Pulau Pinang">
<collectingRegion id="0723BECDE841BDA5CF5122872DCA5BAF" country="Malaysia" name="Pulau Pinang">Penang</collectingRegion>
<collectorName id="48ED0C3836C75CEFD7A298ADDA366FEC">Hill</collectorName>
,
<collectorName id="DB30FE09AE71475C150FAA4F55E2E3D5">Path B</collectorName>
,
<collectorName id="C1CBC82273C0CB7AA850F4DCB9D4F9CE">Random</collectorName>
1. PathB;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E1XAK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[100.267400,5.421600]}">
<geoCoordinate id="587E29EDBFFCB9DDC76759165FAAF825" degrees="5.4216" direction="north" orientation="latitude" precision="5" value="5.4216">5.4216 ° N</geoCoordinate>
,
<geoCoordinate id="1170C1475123E306A73A78C4BB9EB4E3" degrees="100.2674" direction="east" orientation="longitude" precision="5" value="100.2674">100.2674 ° E</geoCoordinate>
</named-content>
;
<elevation id="977C13C515BF675AD1B5E9C8C497F184" metricMagnitude="2" metricUnit="m" metricValue="7.3" unit="m" value="730.0">
<quantity id="E54A10D5A01AA5A745C1DD07DB2E05FC" metricMagnitude="2" metricUnit="m" metricValue="7.3" unit="m" value="730.0">730 m</quantity>
a. s. l.
</elevation>
;
<date id="5DD4AD4D69469DE31BDAF59C14A4D750" value="2022-09-04">
<collectingDate id="A6FFC55DEE221F14B9A8810781A020C2" value="2022-09-04">4 Sept. 2022</collectingDate>
</date>
;
<collectionCode id="6A26741B77D32E3BD9628F38BD806AAB" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
.
<collectorName id="15D81EC5C1E31268FF9A516C22C0FD66">S. Liew</collectorName>
,
<collectorName id="7D8036EB43CC39B4F20DBB0265CDE1E4">J. Dulipat</collectorName>
leg.; BOR / MOL 15109
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6XAK" type="remarks">
<paragraph id="B01F509E5C0C6A714E5F2CECCDCEABCF">
<heading id="D079626235C0EACC9B0A521BDC1F2E0C" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="72572A75AB618792DED56484F7128BA5">
This species was previously described by
<bibRefCitation id="92A9429F4569D642B973EA222FA52C26" author="Stoliczka F" firstAuthor="Stoliczka" journalOrPublisher="Journal of the Asiatic Society of Bengal" pagination="11-38" refId="B48" refString="Stoliczka F (1873) On the land shells of Penang Island, with descriptions of the animals and anatomical notes; part second, Helicacea. Journal of the Asiatic Society of Bengal 42: 1138. https://www.biodiversitylibrary.org/page/35546163#page/25/mode/1up" title="On the land shells of Penang Island, with descriptions of the animals and anatomical notes; part second, Helicacea." url="https://www.biodiversitylibrary.org/page/35546163#page/25/mode/1up" volume="42" year="1873">Stoliczka (1873)</bibRefCitation>
, and it is one of the most common semi-slugs from
<collectingRegion id="BF71CA152F277A0B5A9217FDD9D107F4" country="Malaysia" name="Pulau Pinang">Penang</collectingRegion>
Hill. It was listed as taxon inquirendum on
<bibRefCitation id="C48E8A48A58A8EC869B3EF2E1CFBE62E" DOI="10.14284/448" refId="B30" refString="MolluscaBase (2024) MolluscaBase. https://doi.org/10.14284/448 [accessed 22 September 2024]" year="2024">MolluscaBase (2024</bibRefCitation>
) [
<ext-link id="70B7BEC0F0F8D3295F7EC8869FD19ED6" ext-link-type="uri" xlink_href="https://molluscabase.org/aphia.php?p=taxdetails&amp;id=1337679" xlink_type="simple">https://molluscabase.org/aphia.php?p=taxdetails&amp;id=1337679</ext-link>
]. We suggest that this is a valid species as the specimens fit the description in
<bibRefCitation id="40D87D4272B3154145778B110E7A1570" author="Stoliczka F" firstAuthor="Stoliczka" journalOrPublisher="Journal of the Asiatic Society of Bengal" pagination="11-38" refId="B48" refString="Stoliczka F (1873) On the land shells of Penang Island, with descriptions of the animals and anatomical notes; part second, Helicacea. Journal of the Asiatic Society of Bengal 42: 1138. https://www.biodiversitylibrary.org/page/35546163#page/25/mode/1up" title="On the land shells of Penang Island, with descriptions of the animals and anatomical notes; part second, Helicacea." url="https://www.biodiversitylibrary.org/page/35546163#page/25/mode/1up" volume="42" year="1873">Stoliczka (1873)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>