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<mods:title id="69B0D5EAF4EF9A96BE19826F7D1050F7">The Early Miocene Bats (Chiroptera, Mammalia) From The Karstic Sites Of Erkertshofen And Petersbuch 2 (Southern Germany)</mods:title>
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<mods:namePart id="7E7C2FC6F8845AD267EC60F11BAB646E">Rosina, Valentina V.</mods:namePart>
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<treatment id="03FF8F57F33AFFA3D0A66286FC3F1DE8" LSID="urn:lsid:plazi:treatment:03FF8F57F33AFFA3D0A66286FC3F1DE8" httpUri="http://treatment.plazi.org/id/03FF8F57F33AFFA3D0A66286FC3F1DE8" lastPageNumber="428" pageId="16" pageNumber="428">
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<paragraph id="8BE93E41F33AFFA3D0A66286FAFC188B" blockId="16.[933,1391,486,511]" box="[933,1391,486,511]" pageId="16" pageNumber="428">
<heading id="D0A1892DF33AFFA3D0A66286FAFC188B" box="[933,1391,486,511]" centered="true" fontSize="8" level="2" pageId="16" pageNumber="428" reason="2">
<taxonomicName id="4C5645C2F33AFFA3D0A66286FAFC188B" authority="ZIEGLER, 1993" authorityName="ZIEGLER" authorityYear="1993" box="[933,1391,486,511]" class="Mammalia" family="Vespertilionidae" genus="Eptesicus" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="428" phylum="Chordata" rank="species" species="aurelianensis">
<emphasis id="B922E253F33AFFA3D0A66286FAFC188B" bold="true" box="[933,1391,486,511]" pageId="16" pageNumber="428">
<emphasis id="B922E253F33AFFA3D0A66286FB9D188A" bold="true" box="[933,1038,487,511]" italics="true" pageId="16" pageNumber="428">Eptesicus</emphasis>
cf.
<emphasis id="B922E253F33AFFA3D7346287FB58188B" bold="true" box="[1079,1227,486,510]" italics="true" pageId="16" pageNumber="428">aurelianensis</emphasis>
<bibRefCitation id="EFC743B0F33AFFA3D7D16286FABE1888" author="Ziegler, R." box="[1234,1325,487,511]" pageId="16" pageNumber="428" pagination="119 - 154" refId="ref24078" refString="Ziegler, R. (1993): Die Chiroptera (Mammalia) aus dem Untermiozan von Wintershof-West bei Eichstatt (Bayern). - Mitteilungen der Bayerischen Staatssammlung fur Palaontologie und historische Geologie, 33: 119 - 154." type="journal article" year="1993">ZIEGLER</bibRefCitation>
, 1993
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<paragraph id="8BE93E41F33AFFA3D74F6166FB5B1B6A" blockId="16.[1100,1224,519,543]" box="[1100,1224,519,543]" pageId="16" pageNumber="428">
<figureCitation id="136D22C4F33AFFA3D74F6166FB5B1B6A" box="[1100,1224,519,543]" captionStart="Text-fig" captionStartId="17.[125,206,1840,1861]" captionTargetBox="[125,1440,151,1821]" captionTargetId="figure-2@17.[125,1440,146,1827]" captionTargetPageId="17" captionText="Text-fig. 8. a, o – Miostrellus petersbuchensis, Petersbuch 28: a – NMA P28/0335, right maxillary bone with P4–M3, holotype, ventral view; o – NMA P28/0449, right mnd with C inf., p4–m3, occlusal view; b–f, k, n – M. cf. noctuloides: b – BSP 1962 XIX 4198, right M3, Erkertshofen 1, occlusal view; c – BSP 1962 XIX 4195, left M1, Erkertshofen 1, occlusal view; d – PCMRCh4, left C sup., Petersbuch 2, lingual (d1) and occlusal (d2) views; e – BSP 1974 XIV 1204, left C sup., Erkertshofen 2, occlusal (e1) and lingual (e2) views; f – PCMRCh23, right P4, Petersbuch 2, occlusal view; k – BSP 1974 XIV 1200, right C inf., Erkertshofen 2, occlusal view; n – BSP 1962 XIX 4192, right mnd with m1–2; Erkertshofen 1, lateral (n1) and occlusal (n2) views; g – Eptesicus cf. aurelianensis, BSP 1962 XIX 4197, right M2, Erkertshofen 1, occlusal view; h, i – cf. Miostrellus sp., Erkertshofen 2: h – BSP 1974 XIV 1203, left m1, occlusal view; i – BSP 1974 XIV 1207, right P4; j, l, m – Miostrellus cf. petersbuchensis, Petersbuch 2, occlusal views: j – PCMRCh5, left m1; l – PCMRCh7, left p4; m – PCMRCh6, right mnd with m1." figureDoi="http://doi.org/10.5281/zenodo.5383159" httpUri="https://zenodo.org/record/5383159/files/figure.png" pageId="16" pageNumber="428">
Text-fig.
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<paragraph id="8BE93E41F33AFFA3D0726157FAB21B1B" blockId="16.[843,1482,566,686]" pageId="16" pageNumber="428">M a t e r i a l a n d m e a s u r e m e n t s. Erkertshofen 1: BSP 1962 XIX 4197, right M2, 1.45 × 1.90.</paragraph>
<paragraph id="8BE93E41F33AFFA3D0726117FC1B1BDB" blockId="16.[843,1482,566,686]" pageId="16" pageNumber="428">Petersbuch 2: BSP 1980 XXII 5366, right M2, ≈1.45 × ≈1.90</paragraph>
<paragraph id="8BE93E41F33AFFA3D07261A4FC3F1DE8" blockId="16.[843,1482,709,1181]" pageId="16" pageNumber="428">
D e s c r i p t i o n a n d c o m p a r i s o n. The M2 crowns from Erkertshofen 1 and Petersbuch 2 show a welldeveloped cingulum and paralophs (
<figureCitation id="136D22C4F33AFFA3D7ED6064FAE91A68" box="[1262,1402,773,797]" captionStart="Text-fig" captionStartId="17.[125,206,1840,1861]" captionTargetBox="[125,1440,151,1821]" captionTargetId="figure-2@17.[125,1440,146,1827]" captionTargetPageId="17" captionText="Text-fig. 8. a, o – Miostrellus petersbuchensis, Petersbuch 28: a – NMA P28/0335, right maxillary bone with P4–M3, holotype, ventral view; o – NMA P28/0449, right mnd with C inf., p4–m3, occlusal view; b–f, k, n – M. cf. noctuloides: b – BSP 1962 XIX 4198, right M3, Erkertshofen 1, occlusal view; c – BSP 1962 XIX 4195, left M1, Erkertshofen 1, occlusal view; d – PCMRCh4, left C sup., Petersbuch 2, lingual (d1) and occlusal (d2) views; e – BSP 1974 XIV 1204, left C sup., Erkertshofen 2, occlusal (e1) and lingual (e2) views; f – PCMRCh23, right P4, Petersbuch 2, occlusal view; k – BSP 1974 XIV 1200, right C inf., Erkertshofen 2, occlusal view; n – BSP 1962 XIX 4192, right mnd with m1–2; Erkertshofen 1, lateral (n1) and occlusal (n2) views; g – Eptesicus cf. aurelianensis, BSP 1962 XIX 4197, right M2, Erkertshofen 1, occlusal view; h, i – cf. Miostrellus sp., Erkertshofen 2: h – BSP 1974 XIV 1203, left m1, occlusal view; i – BSP 1974 XIV 1207, right P4; j, l, m – Miostrellus cf. petersbuchensis, Petersbuch 2, occlusal views: j – PCMRCh5, left m1; l – PCMRCh7, left p4; m – PCMRCh6, right mnd with m1." figureDoi="http://doi.org/10.5281/zenodo.5383159" httpUri="https://zenodo.org/record/5383159/files/figure.png" pageId="16" pageNumber="428">
Text-fig.
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<figureCitation id="136D22C4F33AFFA3D67A6064FA171A68" box="[1401,1412,773,797]" captionStart="Text-fig" captionStartId="2.[147,228,2019,2040]" captionTargetBox="[147,1462,810,2000]" captionTargetId="figure-371@2.[146,806,809,1451]" captionTargetPageId="2" captionText="Text-fig. 1. Megaderma franconica, Petersbuch 2: a – BPS 1976 XXII 4147, right mxl with P4–M3, ventral view; b – BSP 1976 XXII 4112, left mnd with m2–3, lateral view; c – BSP 1976 XXII 4124, left mnd with p2–m1, (c1) lateral view, (c2) occlusal view; d – BSP 1976 XXII 4110, right mnd with p4–m3, (d1) occlusal view, (d2) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5383143" httpUri="https://zenodo.org/record/5383143/files/figure.png" pageId="16" pageNumber="428">1</figureCitation>
). The absence of a large hypocone suggests that the tooth belongs to a vespertilionid bat. Both molars have para- and metalophs but lack the paraconules. The trigon basins are closed, the hypocones are weakly developed (
<figureCitation id="136D22C4F33AFFA3D7C060E4FAD51AE8" box="[1219,1350,901,925]" captionStart="Text-fig" captionStartId="17.[125,206,1840,1861]" captionTargetBox="[125,1440,151,1821]" captionTargetId="figure-2@17.[125,1440,146,1827]" captionTargetPageId="17" captionText="Text-fig. 8. a, o – Miostrellus petersbuchensis, Petersbuch 28: a – NMA P28/0335, right maxillary bone with P4–M3, holotype, ventral view; o – NMA P28/0449, right mnd with C inf., p4–m3, occlusal view; b–f, k, n – M. cf. noctuloides: b – BSP 1962 XIX 4198, right M3, Erkertshofen 1, occlusal view; c – BSP 1962 XIX 4195, left M1, Erkertshofen 1, occlusal view; d – PCMRCh4, left C sup., Petersbuch 2, lingual (d1) and occlusal (d2) views; e – BSP 1974 XIV 1204, left C sup., Erkertshofen 2, occlusal (e1) and lingual (e2) views; f – PCMRCh23, right P4, Petersbuch 2, occlusal view; k – BSP 1974 XIV 1200, right C inf., Erkertshofen 2, occlusal view; n – BSP 1962 XIX 4192, right mnd with m1–2; Erkertshofen 1, lateral (n1) and occlusal (n2) views; g – Eptesicus cf. aurelianensis, BSP 1962 XIX 4197, right M2, Erkertshofen 1, occlusal view; h, i – cf. Miostrellus sp., Erkertshofen 2: h – BSP 1974 XIV 1203, left m1, occlusal view; i – BSP 1974 XIV 1207, right P4; j, l, m – Miostrellus cf. petersbuchensis, Petersbuch 2, occlusal views: j – PCMRCh5, left m1; l – PCMRCh7, left p4; m – PCMRCh6, right mnd with m1." figureDoi="http://doi.org/10.5281/zenodo.5383159" httpUri="https://zenodo.org/record/5383159/files/figure.png" pageId="16" pageNumber="428">
Text-fig.
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). The upper molars from Erkertshofen 1 and Petersbuch 2 share these features with
<taxonomicName id="4C5645C2F33AFFA3D0E760A4FBCE1AA8" authorityName="RACHL" authorityYear="1983" box="[996,1117,965,989]" class="Mammalia" family="Vespertilionidae" genus="Miostrellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="434" phylum="Chordata" rank="genus" sensu="stricto">
<emphasis id="B922E253F33AFFA3D0E760A4FBCE1AA8" box="[996,1117,965,989]" italics="true" pageId="16" pageNumber="428">Miostrellus</emphasis>
</taxonomicName>
or
<taxonomicName id="4C5645C2F33AFFA3D78A60A7FB7C1AA8" authorityName="RAFINESQUE" authorityYear="1820" box="[1161,1263,966,989]" class="Mammalia" family="Vespertilionidae" genus="Eptesicus" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="433" phylum="Chordata" rank="genus">
<emphasis id="B922E253F33AFFA3D78A60A7FB7C1AA8" box="[1161,1263,966,989]" italics="true" pageId="16" pageNumber="428">Eptesicus</emphasis>
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. Nevertheless, they are larger than all
<taxonomicName id="4C5645C2F33AFFA3D71D6084FB7C1A88" box="[1054,1263,997,1021]" class="Mammalia" family="Vespertilionidae" genus="Miostrellus" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="428" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B922E253F33AFFA3D71D6084FB041A88" box="[1054,1175,997,1021]" italics="true" pageId="16" pageNumber="428">Miostrellus</emphasis>
species
</taxonomicName>
(
<tableCitation id="C6D40BFAF33AFFA3D6016084FAD91A88" box="[1282,1354,997,1021]" captionStart="Table 8" captionStartId="18.[102,158,641,662]" captionText="Table 8. Comparison of different Miostrellus species and small-sized Eptesicus species from the Miocene of Central Europe (in mm, the size difference between maximum and minimum values, the arithmetic mean is in brackets; after Baudelot 1972, Rachl 1983, Ziegler 1993, 1994, 2000, 2003, Horáček 2001, Rosina and Rummel 2012)." pageId="16" pageNumber="428">Tab. 8</tableCitation>
) and, thus, more similar in size to
<taxonomicName id="4C5645C2F33AFFA3D74B6767FB651D68" authorityName="ZIEGLER" authorityYear="1993" box="[1096,1270,1029,1053]" class="Mammalia" family="Vespertilionidae" genus="Eptesicus" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="428" phylum="Chordata" rank="species" species="aurelianensis">
<emphasis id="B922E253F33AFFA3D74B6767FB651D68" box="[1096,1270,1029,1053]" italics="true" pageId="16" pageNumber="428">E. aurelianensis</emphasis>
</taxonomicName>
(compare with e.g. specimen SMNS 45744 H1;
<bibRefCitation id="EFC743B0F33AFFA3D77E6744FAAF1D48" author="Ziegler, R." box="[1149,1340,1061,1085]" pageId="16" pageNumber="428" pagination="97 - 116" refId="ref24119" refString="Ziegler, R. (1994): Die Chiroptera (Mammalia) aus dem Unteromiozan von Stubersheim 3 (Baden-Wurttemberg). - Munchner Geowissenschaftliche Abhandlun- gen, A, 26: 97 - 116." type="journal article" year="1994">Ziegler 1994: 113</bibRefCitation>
, pl. 5, fig. 6). However, they differ from
<taxonomicName id="4C5645C2F33AFFA3D76D6727FA8F1D28" authorityName="ZIEGLER" authorityYear="1993" box="[1134,1308,1093,1117]" class="Mammalia" family="Vespertilionidae" genus="Eptesicus" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="428" phylum="Chordata" rank="species" species="aurelianensis">
<emphasis id="B922E253F33AFFA3D76D6727FA8F1D28" box="[1134,1308,1093,1117]" italics="true" pageId="16" pageNumber="428">E. aurelianensis</emphasis>
</taxonomicName>
in having some undulated metaloph and a less developed hypocone (Textfig.
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2).
</paragraph>
</subSubSection>
</treatment>
</document>
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