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<document id="D61D7CC40AD13449662E9634A693FB70" ID-DOI="10.11646/zootaxa.5343.4.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="8334529" ID-ZooBank="FA99CD20-856C-4C53-AD8A-47E76400454D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1694430741822" checkinUser="plazi" docAuthor="Tennessen, K. J. &amp; Garrison, R. W." docDate="2023" docId="03FED774FFD28C27FF2F956539421505" docLanguage="en" docName="zootaxa.5343.4.7.pdf" docOrigin="Zootaxa 5343 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5343.4.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epipleoneura parmula Tennessen &amp; Garrison 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="400" masterDocId="FFC7AF0CFFD38C22FFB8970F3B54115C" masterDocTitle="Epipleoneura parmula sp. nov. (Odonata: Coenagrionidae: Protoneurinae) from Amazonian Ecuador" masterLastPageNumber="400" masterPageNumber="395" pageNumber="396" updateTime="1694453375681" updateUser="valdenar" zenodo-license-document="CLOSED">
<mods:mods id="5332E8882AC0A16820C9A73CEB833471" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E538AD0451C05661AAB7D70290C9C479">
<mods:title id="CE597A9DEECBFBD6FE0D079D3AE07D9C">Epipleoneura parmula sp. nov. (Odonata: Coenagrionidae: Protoneurinae) from Amazonian Ecuador</mods:title>
</mods:titleInfo>
<mods:name id="20B3CE526812524D02A67FFC930B2A22" type="personal">
<mods:role id="5B215210DF44CE04FD013426B3FB9D16">
<mods:roleTerm id="750E880E80210B3D39CF8C9591F4A049">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FAFCB958F28DDB7EB990B5F6025CC669">Tennessen, K. J.</mods:namePart>
<mods:affiliation id="9ABA3405A8C17A99F0E09ADCD9D79A0F">Research Associate, Florida State Collection of Arthropods, Gainesville, Florida, USA</mods:affiliation>
</mods:name>
<mods:name id="A39841BF8951D064EB3E893F93641CDF" type="personal">
<mods:role id="8BF2E39CE428827F3F752DD6161F0C2A">
<mods:roleTerm id="82E84C6FB8A8FF954DFFFF9A7A8CEAA3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7231470512929BE432F26E1223A76119">Garrison, R. W.</mods:namePart>
<mods:nameIdentifier id="99403208AA724D0AAE6125221B0F2249" type="ORCID">0000-0001-8586-2723</mods:nameIdentifier>
<mods:affiliation id="183DFF6D0595915D073A81896302203A">California Department of Food &amp; Agriculture, Plant Pest Diagnostics Center, Sacramento, California, USA argiavivida @ gmail. com; https: // orcid. org / 0000 - 0001 - 8586 - 2723</mods:affiliation>
<mods:nameIdentifier id="5687B1AE08E6954A960BE14FDC138BB4" type="email">argiavivida@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="2DBC25355125B7853639D3BC6FB9C8C5">text</mods:typeOfResource>
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<mods:title id="20189386F21A03D56899F3FD59B74328">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D6E087DC0BCA2471273688A8FC447F58">
<mods:date id="2FED39A79BD24D26715802DF03C19332">2023</mods:date>
<mods:detail id="DCDC4DB28BD33548629C7EEE7472C05E" type="pubDate">
<mods:number id="8DB4E47DE5ACA239485E8D432A90B290">2023-09-11</mods:number>
</mods:detail>
<mods:detail id="C93E1B60284F7B423A670D28ABDA01FD" type="volume">
<mods:number id="46DF53284F028C50DDBB725DA2BEAA0A">5343</mods:number>
</mods:detail>
<mods:detail id="EF81CD1CAE34DEC714ABE3B80CC34D90" type="issue">
<mods:number id="20409BA3601F7603E8759109F1292C62">4</mods:number>
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<mods:start id="34CC0A7A30385AAEA153470DD43B0EFA">395</mods:start>
<mods:end id="B4A2F81588AF92CAE16DF4C77A2DEB72">400</mods:end>
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<mods:location id="FF6A91FF9E9C7B0E05FC7411FFB375DD">
<mods:url id="F1121B3EA267B82C2962D3A5FFE3766D">http://dx.doi.org/10.11646/zootaxa.5343.4.7</mods:url>
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<mods:classification id="BCD6B2017416A7FB616979668FD8542B">journal article</mods:classification>
<mods:identifier id="B192D000CFD06E99EEE2315C7935B5CF" type="DOI">10.11646/zootaxa.5343.4.7</mods:identifier>
<mods:identifier id="52A20109E180F44716ACEF8AF92921EA" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C75C4328D324A1B58BC03D37061D4637" type="Zenodo-Dep">8334529</mods:identifier>
<mods:identifier id="283EC0211093DEC4EB0E0A747977E83E" type="ZooBank">FA99CD20-856C-4C53-AD8A-47E76400454D</mods:identifier>
</mods:mods>
<treatment id="03FED774FFD28C27FF2F956539421505" LSID="urn:lsid:plazi:treatment:03FED774FFD28C27FF2F956539421505" httpUri="http://treatment.plazi.org/id/03FED774FFD28C27FF2F956539421505" lastPageId="5" lastPageNumber="400" pageId="1" pageNumber="396">
<subSubSection id="C34D35E9FFD28C23FF2F9565386213D9" box="[151,822,618,645]" pageId="1" pageNumber="396" type="nomenclature">
<paragraph id="8BE86662FFD28C23FF2F9565386213D9" blockId="1.[151,822,618,645]" box="[151,822,618,645]" pageId="1" pageNumber="396">
<heading id="D0A0D10EFFD28C23FF2F9565386213D9" bold="true" box="[151,822,618,645]" fontSize="11" level="1" pageId="1" pageNumber="396" reason="1">
<emphasis id="B923BA70FFD28C23FF2F9565386213D9" bold="true" box="[151,822,618,645]" pageId="1" pageNumber="396">
<taxonomicName id="4C571DE1FFD28C23FF2F9565399F13D8" authority="Tennessen &amp; Garrison" authorityName="Tennessen &amp; Garrison" authorityYear="2023" box="[151,715,618,645]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="396" phylum="Arthropoda" rank="species" species="parmula" status="sp. nov.">
<emphasis id="B923BA70FFD28C23FF2F95653AFE13D9" bold="true" box="[151,426,618,645]" italics="true" pageId="1" pageNumber="396">Epipleoneura parmula</emphasis>
Tennessen &amp; Garrison
</taxonomicName>
,
<taxonomicNameLabel id="A210070BFFD28C23FD609564386213D9" box="[728,822,619,645]" pageId="1" pageNumber="396" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34D35E9FFD28C23FF2F95A53E6D139D" box="[151,1337,682,706]" pageId="1" pageNumber="396" type="description">
<paragraph id="8BE86662FFD28C23FF2F95A53E6D139D" blockId="1.[151,1337,682,706]" box="[151,1337,682,706]" pageId="1" pageNumber="396">
<figureCitation id="136C7AE7FFD28C23FF2F95A53B88139E" box="[151,220,682,706]" captionStart="FIGURES 1–6" captionStartId="2.[152,270,1827,1851]" captionTargetBox="[155,1431,241,1800]" captionTargetId="figure-58@2.[151,1435,237,1804]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 2, 5, 6: Epipleoneura parmula ♁ paratype; Fig. 3 Epipleoneura ocuene ♁ paratype; 4 Epipeoneura pallida ♁. Fig. 1. Habitus; Figs. 2–4 head and thorax, dorsolateral (2) and lateral views; Fig. 5 wings; 6 genital ligula ectal (left) and lateral (right) views." figureDoi="http://doi.org/10.5281/zenodo.8334531" httpUri="https://zenodo.org/record/8334531/files/figure.png" pageId="1" pageNumber="396">Figs. 1</figureCitation>
, whole body scan; 2, head &amp; thorax; 5, wings; 6, genital ligula; 7, 8, caudal appendages; 11, distribution map.
</paragraph>
</subSubSection>
<subSubSection id="C34D35E9FFD28C26FF2F95FF3AF216A4" lastPageId="4" lastPageNumber="399" pageId="1" pageNumber="396" type="etymology">
<paragraph id="8BE86662FFD28C23FF2F95FF3A6C1272" blockId="1.[151,1437,752,2039]" pageId="1" pageNumber="396">
<emphasis id="B923BA70FFD28C23FF2F95FF3A4A1256" bold="true" box="[151,286,752,778]" pageId="1" pageNumber="396">Etymology.</emphasis>
Named
<taxonomicName id="4C571DE1FFD28C23FEC795FF3A8A1256" authorityName="Tennessen &amp; Garrison" authorityYear="2023" box="[383,478,752,778]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="396" phylum="Arthropoda" rank="species" species="parmula">
<emphasis id="B923BA70FFD28C23FEC795FF3A8A1256" box="[383,478,752,778]" italics="true" pageId="1" pageNumber="396">parmula</emphasis>
</taxonomicName>
, noun in apposition (from L. noun
<emphasis id="B923BA70FFD28C23FCD795FE38EC1256" box="[879,952,753,778]" italics="true" pageId="1" pageNumber="396">parma</emphasis>
: small shield) referring to the shape of the male epiproct.
</paragraph>
<paragraph id="8BE86662FFD28C23FF7F94373BBA12C6" blockId="1.[151,1437,752,2039]" pageId="1" pageNumber="396">
<emphasis id="B923BA70FFD28C23FF7F94373AD3120E" bold="true" box="[199,391,824,850]" pageId="1" pageNumber="396">Primary Types:</emphasis>
<taxonomicName id="4C571DE1FFD28C23FE2B943739C3120E" authorityName="Tennessen &amp; Garrison" authorityYear="2023" box="[403,663,824,850]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="396" phylum="Arthropoda" rank="species" species="parmula">
<emphasis id="B923BA70FFD28C23FE2B943739C3120E" box="[403,663,824,850]" italics="true" pageId="1" pageNumber="396">Epipleoneura parmula</emphasis>
</taxonomicName>
:
<materialsCitation id="3B3F6C3FFFD28C23FD1394373BBF12C6" collectingDate="1977-07-27" collectionCode="FSCA" collectorName="Dennis R. Paulson &amp; Susan Hills" country="Ecuador" elevation="247" latitude="-0.4652" location="Laguna Taracoa" longLatPrecision="7" longitude="-76.7638" municipality="Primavera" pageId="1" pageNumber="396" specimenCount="1" stateProvince="Orellana" typeStatus="holotype">
<typeStatus id="54ECD8C0FFD28C23FD1394373859120E" box="[683,781,824,850]" pageId="1" pageNumber="396">holotype</typeStatus>
♁,
<collectingCountry id="F34026F2FFD28C23FCFA9437389A120E" box="[834,974,824,850]" name="Ecuador" pageId="1" pageNumber="396">ECUADOR</collectingCountry>
:
<collectingRegion id="4993A880FFD28C23FC6594373FFA120E" box="[989,1198,824,850]" country="Ecuador" name="Orellana" pageId="1" pageNumber="396">Orellana Province</collectingRegion>
:
<collectingMunicipality id="6B8CFC18FFD28C23FB7A94373E6C120E" box="[1218,1336,824,850]" pageId="1" pageNumber="396">Primavera</collectingMunicipality>
,
<location id="8E8830B9FFD28C23FAF094363BBB122B" LSID="urn:lsid:plazi:treatment:03FED774FFD28C27FF2F956539421505:8E8830B9FFD28C23FAF094363BBB122B" country="Ecuador" latitude="-0.4652" longLatPrecision="7" longitude="-76.7638" municipality="Primavera" name="Laguna Taracoa" pageId="1" pageNumber="396" stateProvince="Orellana">Laguna Taracoa</location>
{
<geoCoordinate id="EE6300A5FFD28C23FEBB94533A27122B" box="[259,371,860,887]" degrees="0.4652" direction="south" orientation="latitude" pageId="1" pageNumber="396" precision="5" value="-0.4652">0.4652°S</geoCoordinate>
,
<geoCoordinate id="EE6300A5FFD28C23FE3B94533958122A" box="[387,524,860,886]" degrees="76.7638" direction="west" orientation="longitude" pageId="1" pageNumber="396" precision="5" value="-76.7638">76.7638°W</geoCoordinate>
},
<quantity id="4CAFCB87FFD28C23FD9C94533926122B" box="[548,626,860,887]" metricMagnitude="2" metricUnit="m" metricValue="2.4699999999999998" pageId="1" pageNumber="396" unit="m" value="247.0">
<elevation id="007A8151FFD28C23FD9C94533926122B" box="[548,626,860,887]" metricMagnitude="2" metricUnit="m" metricValue="2.4699999999999998" pageId="1" pageNumber="396" unit="m" value="247.0">247 m</elevation>
</quantity>
,
<date id="FFE940A2FFD28C23FD3B94533874122A" box="[643,800,860,886]" pageId="1" pageNumber="396" value="1977-07-27">
<collectingDate id="EFADB94AFFD28C23FD3B94533874122A" box="[643,800,860,886]" pageId="1" pageNumber="396" value="1977-07-27">27 July 1977</collectingDate>
</date>
,
<collectorName id="26A203B4FFD28C23FC8894533F5E122A" box="[816,1034,860,887]" pageId="1" pageNumber="396">Dennis R. Paulson</collectorName>
&amp;
<collectorName id="26A203B4FFD28C23FB8F94533FE8122A" box="[1079,1212,860,886]" pageId="1" pageNumber="396">Susan Hills</collectorName>
leg. [examined, in
<collectionCode id="ED46FEA7FFD28C23FF2F948F3BB712C6" box="[151,227,896,922]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="1" pageNumber="396" type="Museum">FSCA</collectionCode>
]
</materialsCitation>
.
</paragraph>
<paragraph id="8BE86662FFD28C23FF7F94AB38FD1416" blockId="1.[151,1437,752,2039]" pageId="1" pageNumber="396">
<emphasis id="B923BA70FFD28C23FF7F94AB3A9E12E2" bold="true" box="[199,458,932,958]" pageId="1" pageNumber="396">Specimens examined:</emphasis>
<materialsCitation id="3B3F6C3FFFD28C23FE6B94AB3A1C12BE" collectingDate="1977-07-13" collectingDateMax="2011-01-26" collectingDateMin="1977-07-13" collectionCode="CSCA, RWG, FSCA" country="Ecuador" county="Rio Payamino" elevation="290" latitude="-1.0743" location="Rio Payamino" longLatPrecision="7" longitude="-77.6521" municipality="Coca" pageId="1" pageNumber="396" specimenCount="1" stateProvince="Orellana" typeStatus="paratype">
23 ♁♁ (all
<typeStatus id="54ECD8C0FFD28C23FDE094AB399D12E2" box="[600,713,932,958]" pageId="1" pageNumber="396">paratypes</typeStatus>
): 6 ♁♁,
<collectingCountry id="F34026F2FFD28C23FC9594AB38ED12E2" box="[813,953,932,958]" name="Ecuador" pageId="1" pageNumber="396">ECUADOR</collectingCountry>
:
<collectingRegion id="4993A880FFD28C23FC7C94AB3FC512E2" box="[964,1169,932,958]" country="Ecuador" name="Orellana" pageId="1" pageNumber="396">Orellana Province</collectingRegion>
: same data as holotype [
<collectionCode id="ED46FEA7FFD28C23FF2694C73BB912BE" box="[158,237,968,994]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="1" pageNumber="396" type="Museum">CSCA</collectionCode>
,
<collectionCode id="ED46FEA7FFD28C23FF4F94C73A6912BE" box="[247,317,968,994]" pageId="1" pageNumber="396">RWG</collectionCode>
];
</materialsCitation>
<materialsCitation id="3B3F6C3FFFD28C23FEF694C73FBD12BE" box="[334,1257,968,996]" collectingDate="1977-07-13" collectingDateMax="2011-01-26" collectingDateMin="1977-07-13" collectionCode="CSCA, RWG, FSCA" country="Ecuador" county="Rio Payamino" elevation="290" latitude="-1.0743" location="Rio Payamino" longLatPrecision="7" longitude="-77.6521" municipality="Coca" pageId="1" pageNumber="396" specimenCount="1" stateProvince="Orellana" typeStatus="paratype">
5 ♁♁, same data but
<date id="FFE940A2FFD28C23FD8194C7399F12BE" box="[569,715,968,994]" pageId="1" pageNumber="396" value="1977-07-13">
<collectingDate id="EFADB94AFFD28C23FD8194C7399F12BE" box="[569,715,968,994]" pageId="1" pageNumber="396" value="1977-07-13">13 July 1977</collectingDate>
</date>
, S. M. &amp;
<collectorName id="26A203B4FFD28C23FC8C94C73F5212BE" box="[820,1030,968,995]" pageId="1" pageNumber="396">Kenneth W. Knopf</collectorName>
leg. [
<collectionCode id="ED46FEA7FFD28C23FBFA94C73FDA12BE" box="[1090,1166,968,994]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="1" pageNumber="396" type="Museum">FSCA</collectionCode>
,
<collectionCode id="ED46FEA7FFD28C23FB2094C73F8A12BE" box="[1176,1246,968,994]" pageId="1" pageNumber="396">RWG</collectionCode>
];
</materialsCitation>
<materialsCitation id="3B3F6C3FFFD28C23FB5794C73A2C1576" collectingDate="1977-07-13" collectingDateMax="2011-01-26" collectingDateMin="1977-07-13" collectionCode="CSCA, RWG, FSCA" country="Ecuador" county="Rio Payamino" elevation="290" latitude="-1.0743" location="Rio Payamino" longLatPrecision="7" longitude="-77.6521" municipality="Coca" pageId="1" pageNumber="396" specimenCount="1" stateProvince="Orellana" typeStatus="paratype">
9 ♁♁, pond 8.2 km west of
<collectingCounty id="62891EEEFFD28C23FEA394E33AEF155A" box="[283,443,1004,1030]" pageId="1" pageNumber="396">Río Payamino</collectingCounty>
at
<collectingMunicipality id="6B8CFC18FFD28C23FE6594E3394D155A" box="[477,537,1004,1030]" pageId="1" pageNumber="396">Coca</collectingMunicipality>
,
<collectorName id="26A203B4FFD28C23FD9D94E2392E155A" box="[549,634,1004,1031]" pageId="1" pageNumber="396">Hwy. E</collectorName>
20,
<geoCoordinate id="EE6300A5FFD28C23FD1B94E339A6155A" box="[675,754,1004,1030]" degrees="0.48" direction="south" orientation="latitude" pageId="1" pageNumber="396" precision="555" value="-0.48">0.48°S</geoCoordinate>
,
<geoCoordinate id="EE6300A5FFD28C23FD4594E33837155A" box="[765,867,1004,1030]" degrees="77.08" direction="west" orientation="longitude" pageId="1" pageNumber="396" precision="555" value="-77.08">77.08°W</geoCoordinate>
, 290 m,
<date id="FFE940A2FFD28C23FC7B94E33FCA155A" box="[963,1182,1004,1030]" pageId="1" pageNumber="396" value="1997-11-18">
<collectingDate id="EFADB94AFFD28C23FC7B94E33FCA155A" box="[963,1182,1004,1030]" pageId="1" pageNumber="396" value="1997-11-18">18 November 1997</collectingDate>
</date>
,
<collectorName id="26A203B4FFD28C23FB1194E33EC8155B" box="[1193,1436,1004,1031]" pageId="1" pageNumber="396">Kenneth J. Tennessen</collectorName>
leg. [
<collectionCode id="ED46FEA7FFD28C23FF77931F3A4F1576" box="[207,283,1040,1066]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="1" pageNumber="396" type="Museum">FSCA</collectionCode>
,
<collectionCode id="ED46FEA7FFD28C23FE9F931F3A391576" box="[295,365,1040,1066]" pageId="1" pageNumber="396">RWG</collectionCode>
];
</materialsCitation>
<materialsCitation id="3B3F6C3FFFD28C23FE38931F385F1512" collectingDate="1977-07-13" collectingDateMax="2011-01-26" collectingDateMin="1977-07-13" collectionCode="CSCA, RWG, FSCA" country="Ecuador" county="Rio Payamino" elevation="290" latitude="-1.0743" location="Rio Payamino" longLatPrecision="7" longitude="-77.6521" municipality="Coca" pageId="1" pageNumber="396" specimenCount="1" stateProvince="Orellana" typeStatus="paratype">
1 ♁, pond 11.2 km west of
<location id="8E8830B9FFD28C23FD02931F380F1576" LSID="urn:lsid:plazi:treatment:03FED774FFD28C27FF2F956539421505:8E8830B9FFD28C23FD02931F380F1576" box="[698,859,1040,1066]" country="Ecuador" county="Rio Payamino" latitude="-1.0743" longLatPrecision="7" longitude="-77.6521" municipality="Coca" name="Rio Payamino" pageId="1" pageNumber="396" stateProvince="Orellana">Río Payamino</location>
at Coca,
<collectorName id="26A203B4FFD28C23FC71931E3F4A1576" box="[969,1054,1040,1067]" pageId="1" pageNumber="396">Hwy. E</collectorName>
20,
<geoCoordinate id="EE6300A5FFD28C23FBF1931F3FE71576" box="[1097,1203,1040,1066]" degrees="0.4827" direction="south" orientation="latitude" pageId="1" pageNumber="396" precision="5" value="-0.4827">0.4827°S</geoCoordinate>
,
<geoCoordinate id="EE6300A5FFD28C23FB78931F3E151576" box="[1216,1345,1040,1066]" degrees="77.0997" direction="west" orientation="longitude" pageId="1" pageNumber="396" precision="5" value="-77.0997">77.0997°W</geoCoordinate>
, 300 m,
<date id="FFE940A2FFD28C23FF2F933B3A201512" box="[151,372,1076,1102]" pageId="1" pageNumber="396" value="1997-11-18">
<collectingDate id="EFADB94AFFD28C23FF2F933B3A201512" box="[151,372,1076,1102]" pageId="1" pageNumber="396" value="1997-11-18">18 November 1997</collectingDate>
</date>
,
<collectorName id="26A203B4FFD28C23FEC7933B39271513" box="[383,627,1076,1103]" pageId="1" pageNumber="396">Kenneth J. Tennessen</collectorName>
leg. [
<collectionCode id="ED46FEA7FFD28C23FD0A933B38551512" box="[690,769,1076,1102]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="1" pageNumber="396" type="Museum">FSCA</collectionCode>
];
</materialsCitation>
<materialsCitation id="3B3F6C3FFFD28C23FCAB933B382115CA" collectingDate="1977-07-13" collectingDateMax="2011-01-26" collectingDateMin="1977-07-13" collectionCode="CSCA, RWG, FSCA" country="Ecuador" county="Rio Payamino" elevation="290" latitude="-1.0743" location="Rio Payamino" longLatPrecision="7" longitude="-77.6521" municipality="Coca" pageId="1" pageNumber="396" specimenCount="1" stateProvince="Orellana" typeStatus="paratype">
1 ♁,
<location id="8E8830B9FFD28C23FCF5933B38BA1512" LSID="urn:lsid:plazi:treatment:03FED774FFD28C27FF2F956539421505:8E8830B9FFD28C23FCF5933B38BA1512" box="[845,1006,1076,1102]" country="Ecuador" county="Rio Payamino" latitude="-1.0743" longLatPrecision="7" longitude="-77.6521" municipality="Coca" name="Rio Shiripuno" pageId="1" pageNumber="396" stateProvince="Orellana">Río Shiripuno</location>
, Shiripuno Lodge, 26 km southeast of
<location id="8E8830B9FFD28C23FF2F93573A51152E" LSID="urn:lsid:plazi:treatment:03FED774FFD28C27FF2F956539421505:8E8830B9FFD28C23FF2F93573A51152E" box="[151,261,1112,1138]" country="Ecuador" county="Rio Payamino" latitude="-1.0743" longLatPrecision="7" longitude="-77.6521" municipality="Coca" name="Shiripuno" pageId="1" pageNumber="396" stateProvince="Orellana">Shiripuno</location>
bridge [image # 3463-68],
<geoCoordinate id="EE6300A5FFD28C23FDF8935739FE152E" box="[576,682,1112,1138]" degrees="1.1047" direction="south" orientation="latitude" pageId="1" pageNumber="396" precision="5" value="-1.1047">1.1047°S</geoCoordinate>
,
<geoCoordinate id="EE6300A5FFD28C23FD0F9357386C152E" box="[695,824,1112,1138]" degrees="76.7317" direction="west" orientation="longitude" pageId="1" pageNumber="396" precision="5" value="-76.7317">76.7317°W</geoCoordinate>
, 235 m,
<date id="FFE940A2FFD28C23FC2493573FD1152E" box="[924,1157,1112,1139]" pageId="1" pageNumber="396" value="2009-01-23" valueMax="2009-01-26" valueMin="2009-01-23">
<collectingDate id="EFADB94AFFD28C23FC2493573FD1152E" box="[924,1157,1112,1139]" pageId="1" pageNumber="396" value="2009-01-23" valueMax="2009-01-26" valueMin="2009-01-23">23–26 January 2009</collectingDate>
</date>
, David Wagner, William Haber, Elacio Tapia,
<collectorName id="26A203B4FFD28C23FE3093733AAB15CA" box="[392,511,1148,1174]" pageId="1" pageNumber="396">Jim Miller</collectorName>
&amp;
<collectorName id="26A203B4FFD28C23FD9E937339B415CB" box="[550,736,1148,1175]" pageId="1" pageNumber="396">Virginia Wagner</collectorName>
leg. [
<collectionCode id="ED46FEA7FFD28C23FC9A9373383F15CA" box="[802,875,1148,1174]" pageId="1" pageNumber="396">RWG</collectionCode>
];
</materialsCitation>
<materialsCitation id="3B3F6C3FFFD28C23FCC693723A9A145E" collectingDate="1977-07-13" collectingDateMax="2011-01-26" collectingDateMin="1977-07-13" collectionCode="CSCA, RWG, FSCA" country="Ecuador" county="Rio Payamino" elevation="290" latitude="-1.0743" location="Rio Payamino" longLatPrecision="7" longitude="-77.6521" municipality="Coca" pageId="1" pageNumber="396" specimenCount="1" stateProvince="Orellana" typeStatus="paratype">
<collectingRegion id="4993A880FFD28C23FCC693723F7C15CA" box="[894,1064,1148,1175]" country="Ecuador" name="Napo" pageId="1" pageNumber="396">Napo Province</collectingRegion>
: 1 ♁, Airport pond,
<location id="8E8830B9FFD28C23FAAD93723EC815CA" LSID="urn:lsid:plazi:treatment:03FED774FFD28C27FF2F956539421505:8E8830B9FFD28C23FAAD93723EC815CA" box="[1301,1436,1148,1175]" country="Ecuador" county="Rio Payamino" latitude="-1.0743" longLatPrecision="7" longitude="-77.6521" municipality="Coca" name="Jatun Sacha" pageId="1" pageNumber="396" stateProvince="Orellana">Jatun Sacha</location>
Road, road to airport at Zancudo,
<locationDeviation id="9A5A7F40FFD28C23FDAE93AF380515E6" box="[534,849,1184,1211]" location="Jatun Sacha" pageId="1" pageNumber="396">3.8 km ENE of Jatun Sacha</locationDeviation>
, shallow pond with water lilies formed by widening of tiny stream (waypoint 460),
<geoCoordinate id="EE6300A5FFD28C23FE4493CB39321582" box="[508,614,1220,1246]" degrees="1.0483" direction="south" orientation="latitude" pageId="1" pageNumber="396" precision="5" value="-1.0483">1.0483°S</geoCoordinate>
,
<geoCoordinate id="EE6300A5FFD28C23FDCB93CB39A11583" box="[627,757,1220,1247]" degrees="77.5861" direction="west" orientation="longitude" pageId="1" pageNumber="396" precision="5" value="-77.5861">77.5861°W</geoCoordinate>
, 382 m,
<date id="FFE940A2FFD28C23FCE193CB3F431582" box="[857,1047,1220,1247]" pageId="1" pageNumber="396" value="2011-01-26">
<collectingDate id="EFADB94AFFD28C23FCE193CB3F431582" box="[857,1047,1220,1247]" pageId="1" pageNumber="396" value="2011-01-26">26 January 2011</collectingDate>
</date>
, William Haber,
<collectorName id="26A203B4FFD28C23FB6293CB3E2A1582" box="[1242,1406,1220,1246]" pageId="1" pageNumber="396">Fred Morrison</collectorName>
&amp;
<collectorName id="26A203B4FFD28C23FF2F93E73A68145F" box="[151,316,1256,1283]" pageId="1" pageNumber="396">Ronald Vargas</collectorName>
leg. [
<collectionCode id="ED46FEA7FFD28C23FEC393E73A90145E" box="[379,452,1256,1282]" pageId="1" pageNumber="396">RWG</collectionCode>
];
</materialsCitation>
<materialsCitation id="3B3F6C3FFFD28C23FE6F93E738F11416" collectingDate="1977-07-13" collectingDateMax="2011-01-26" collectingDateMin="1977-07-13" collectionCode="CSCA, RWG, FSCA" country="Ecuador" county="Rio Payamino" elevation="290" latitude="-1.0743" location="Rio Payamino" longLatPrecision="7" longitude="-77.6521" municipality="Coca" pageId="1" pageNumber="396" specimenCount="1" stateProvince="Orellana" typeStatus="paratype">
1 ♁, road south to Pulini from main road to airport and
<collectorName id="26A203B4FFD28C23FBE993E63FED145F" box="[1105,1209,1257,1283]" pageId="1" pageNumber="396">La Punta</collectorName>
, 21 km southeast of
<location id="8E8830B9FFD28C23FF2F92023B84147B" LSID="urn:lsid:plazi:treatment:03FED774FFD28C27FF2F956539421505:8E8830B9FFD28C23FF2F92023B84147B" box="[151,208,1293,1319]" country="Ecuador" county="Rio Payamino" latitude="-1.0743" longLatPrecision="7" longitude="-77.6521" municipality="Coca" name="Tena" pageId="1" pageNumber="396" stateProvince="Orellana">Tena</location>
, 1 km south of road junction, large pond (W-460),
<geoCoordinate id="EE6300A5FFD28C23FC9F920338C5147A" box="[807,913,1292,1318]" degrees="1.0743" direction="south" orientation="latitude" pageId="1" pageNumber="396" precision="5" value="-1.0743">1.0743°S</geoCoordinate>
,
<geoCoordinate id="EE6300A5FFD28C23FC1892033F75147B" box="[928,1057,1292,1319]" degrees="77.6521" direction="west" orientation="longitude" pageId="1" pageNumber="396" precision="5" value="-77.6521">77.6521°W</geoCoordinate>
, 470 m,
<date id="FFE940A2FFD28C23FB3192033E1E147A" box="[1161,1354,1292,1319]" pageId="1" pageNumber="396" value="2011-01-22">
<collectingDate id="EFADB94AFFD28C23FB3192033E1E147A" box="[1161,1354,1292,1319]" pageId="1" pageNumber="396" value="2011-01-22">22 January 2011</collectingDate>
</date>
,
<collectorName id="26A203B4FFD28C23FAE092033BBB1417" pageId="1" pageNumber="396">David Wagner</collectorName>
,
<collectorName id="26A203B4FFD28C23FF42923F3AF61416" box="[250,418,1328,1354]" pageId="1" pageNumber="396">William Haber</collectorName>
,
<collectorName id="26A203B4FFD28C23FE15923F391B1416" box="[429,591,1328,1354]" pageId="1" pageNumber="396">Fred Morrison</collectorName>
&amp;
<collectorName id="26A203B4FFD28C23FDCB923F384C1417" box="[627,792,1328,1355]" pageId="1" pageNumber="396">Ronald Vargus</collectorName>
leg. [
<collectionCode id="ED46FEA7FFD28C23FCEE923F38CB1416" box="[854,927,1328,1354]" pageId="1" pageNumber="396">RWG</collectionCode>
]
</materialsCitation>
.
</paragraph>
<paragraph id="8BE86662FFD28C23FF7F925B39E214CE" blockId="1.[151,1437,752,2039]" pageId="1" pageNumber="396">
A lentic species, slender (
<figureCitation id="136C7AE7FFD28C23FE5F925B397C1432" box="[487,552,1364,1390]" captionStart="FIGURES 1–6" captionStartId="2.[152,270,1827,1851]" captionTargetBox="[155,1431,241,1800]" captionTargetId="figure-58@2.[151,1435,237,1804]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 2, 5, 6: Epipleoneura parmula ♁ paratype; Fig. 3 Epipleoneura ocuene ♁ paratype; 4 Epipeoneura pallida ♁. Fig. 1. Habitus; Figs. 2–4 head and thorax, dorsolateral (2) and lateral views; Fig. 5 wings; 6 genital ligula ectal (left) and lateral (right) views." figureDoi="http://doi.org/10.5281/zenodo.8334531" httpUri="https://zenodo.org/record/8334531/files/figure.png" pageId="1" pageNumber="396">Fig. 1</figureCitation>
), dorsum dark with metallic reflections, thorax and abdomen tan laterally, and male epiproct with two elongate fused branches.
</paragraph>
<paragraph id="8BE86662FFD28C23FF7F92933A8D14EA" blockId="1.[151,1437,752,2039]" box="[199,473,1436,1462]" pageId="1" pageNumber="396">
Description of
<typeStatus id="54ECD8C0FFD28C23FED792933A8114EA" box="[367,469,1436,1462]" pageId="1" pageNumber="396">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BE86662FFD28C23FF7F92CF38DD17D2" blockId="1.[151,1437,752,2039]" pageId="1" pageNumber="396">
Head. Epicranium matte black with a dark metallic transverse crossbar between antennal bases; labium and maxilla tan; sides of mandibles yellow tan with medial brown spot; genae yellow tan with an upper brown triangular mark lateral to antefrons; labrum golden tan with small dark brown basomedial spot and seven stout setae on ectal surface and anterior margin; anteclypeus tan; postclypeus dark brown distally, tan basally; frontoclypeal suture brown; antefrons tan; frons angulate; antenna mostly tan, segment 1 darker than remainder; rear of head black except border adjacent to compound eyes narrowly yellow (
<figureCitation id="136C7AE7FFD28C23FC8E917B382F17D2" box="[822,891,1652,1678]" captionStart="FIGURES 1–6" captionStartId="2.[152,270,1827,1851]" captionTargetBox="[155,1431,241,1800]" captionTargetId="figure-58@2.[151,1435,237,1804]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 2, 5, 6: Epipleoneura parmula ♁ paratype; Fig. 3 Epipleoneura ocuene ♁ paratype; 4 Epipeoneura pallida ♁. Fig. 1. Habitus; Figs. 2–4 head and thorax, dorsolateral (2) and lateral views; Fig. 5 wings; 6 genital ligula ectal (left) and lateral (right) views." figureDoi="http://doi.org/10.5281/zenodo.8334531" httpUri="https://zenodo.org/record/8334531/files/figure.png" pageId="1" pageNumber="396">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8BE86662FFD28C20FF7F91973F7B1189" blockId="1.[151,1437,752,2039]" lastBlockId="2.[151,1436,151,213]" lastPageId="2" lastPageNumber="397" pageId="1" pageNumber="396">
Thorax. Prothorax black dorsally with a slight metallic luster, tan laterally (as in
<figureCitation id="136C7AE7FFD28C23FB8391973F2E17EE" box="[1083,1146,1688,1714]" captionStart="FIGURES 1–6" captionStartId="2.[152,270,1827,1851]" captionTargetBox="[155,1431,241,1800]" captionTargetId="figure-58@2.[151,1435,237,1804]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 2, 5, 6: Epipleoneura parmula ♁ paratype; Fig. 3 Epipleoneura ocuene ♁ paratype; 4 Epipeoneura pallida ♁. Fig. 1. Habitus; Figs. 2–4 head and thorax, dorsolateral (2) and lateral views; Fig. 5 wings; 6 genital ligula ectal (left) and lateral (right) views." figureDoi="http://doi.org/10.5281/zenodo.8334531" httpUri="https://zenodo.org/record/8334531/files/figure.png" pageId="1" pageNumber="396">Fig. 1</figureCitation>
), posterior margin of hind lobe broadly convex, tan, the tan border extended medially; propleuron mostly brown. Mesepisternum black with a slight metallic luster, no pale antehumeral stripe; mesepimeron yellow tan with an upper brown stripe isolating an upper narrow yellow-tan stripe (below mesopleural, or humeral, suture), the brown stripe beginning as a light brown mark on mesinfraepisternum, continuing light brown on anterior 1/8 of mesepimeron then continuing and widening as a dark metallic stripe to antealar carina, then extending posteroventrally and ending above metapleural suture; metapleural suture and metapleural fossa brown; metepisternum yellow tan except posteriorly where dark brown mesepimeral stripe runs along antealar carina; metepimeron yellow tan; pterothoracic venter light tan; coxae and trochanters yellow tan; femora light yellow except for wide, diffuse light brown ring in basal third, a narrow light brown ring in posterior third, and apex dark; tibiae, tarsi and claws yellow with darkened apices, spines dark. Wings hyaline, venation black to dark brown; MP ending 1/3 of a cell distal to vein descending from subnodus; 13 Px in Fw,
<quantity id="4CAFCB87FFD18C20FF7F97B43A541189" box="[199,256,187,213]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="2" pageNumber="397" unit="in" value="10.0">10 in</quantity>
left Hw,
<quantity id="4CAFCB87FFD18C20FED097B43AF41189" box="[360,416,187,213]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="2" pageNumber="397" unit="in" value="11.0">11 in</quantity>
right HW; Pt light brown, overlying 1.0 cell (as in
<figureCitation id="136C7AE7FFD18C20FC6597B43F751189" box="[989,1057,186,213]" captionStart="FIGURES 1–6" captionStartId="2.[152,270,1827,1851]" captionTargetBox="[155,1431,241,1800]" captionTargetId="figure-58@2.[151,1435,237,1804]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 2, 5, 6: Epipleoneura parmula ♁ paratype; Fig. 3 Epipleoneura ocuene ♁ paratype; 4 Epipeoneura pallida ♁. Fig. 1. Habitus; Figs. 2–4 head and thorax, dorsolateral (2) and lateral views; Fig. 5 wings; 6 genital ligula ectal (left) and lateral (right) views." figureDoi="http://doi.org/10.5281/zenodo.8334531" httpUri="https://zenodo.org/record/8334531/files/figure.png" pageId="2" pageNumber="397">Fig. 5</figureCitation>
).
</paragraph>
<caption id="DF2836EAFFD18C20FF20902C3A3216DF" ID-DOI="http://doi.org/10.5281/zenodo.8334531" ID-Zenodo-Dep="8334531" httpUri="https://zenodo.org/record/8334531/files/figure.png" pageId="2" pageNumber="397" startId="2.[152,270,1827,1851]" targetBox="[155,1431,241,1800]" targetPageId="2" targetType="figure">
<paragraph id="8BE86662FFD18C20FF20902C3A3216DF" blockId="2.[151,1437,1827,1923]" pageId="2" pageNumber="397">
<emphasis id="B923BA70FFD18C20FF20902C3A151667" bold="true" box="[152,321,1827,1851]" pageId="2" pageNumber="397">FIGURES 1–6.</emphasis>
1, 2, 5, 6:
<taxonomicName id="4C571DE1FFD18C20FE16902C39C11667" authorityName="Tennessen &amp; Garrison" authorityYear="2023" box="[430,661,1827,1851]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="397" phylum="Arthropoda" rank="species" species="parmula">
<emphasis id="B923BA70FFD18C20FE16902C39C11667" box="[430,661,1827,1851]" italics="true" pageId="2" pageNumber="397">Epipleoneura parmula</emphasis>
</taxonomicName>
♁ paratype; Fig. 3
<taxonomicName id="4C571DE1FFD18C20FCE3902C3F671667" authorityName="De Marmels" authorityYear="1989" box="[859,1075,1827,1851]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="397" phylum="Arthropoda" rank="species" species="ocuene">
<emphasis id="B923BA70FFD18C20FCE3902C3F671667" box="[859,1075,1827,1851]" italics="true" pageId="2" pageNumber="397">Epipleoneura ocuene</emphasis>
</taxonomicName>
♁ paratype; 4
<taxonomicName id="4C571DE1FFD18C20FB73902B3EC81667" box="[1227,1436,1827,1851]" class="Insecta" family="Coenagrionidae" genus="Epipeoneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="397" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B923BA70FFD18C20FB73902B3EC81667" box="[1227,1436,1827,1851]" italics="true" pageId="2" pageNumber="397">Epipeoneura pallida</emphasis>
</taxonomicName>
♁. Fig. 1. Habitus; Figs. 2–4 head and thorax, dorsolateral (2) and lateral views; Fig. 5 wings; 6 genital ligula ectal (left) and lateral (right) views.
</paragraph>
</caption>
<caption id="DF2836EAFFD08C21FF20913F3A7A17CC" ID-DOI="http://doi.org/10.5281/zenodo.8334533" ID-Zenodo-Dep="8334533" httpUri="https://zenodo.org/record/8334533/files/figure.png" pageId="3" pageNumber="398" startId="3.[152,270,1584,1608]" targetBox="[155,1431,185,1557]" targetPageId="3" targetType="figure">
<paragraph id="8BE86662FFD08C21FF20913F3A7A17CC" blockId="3.[151,1437,1584,1680]" pageId="3" pageNumber="398">
<emphasis id="B923BA70FFD08C21FF20913F3A1A1714" bold="true" box="[152,334,1584,1608]" pageId="3" pageNumber="398">FIGURES 7–10.</emphasis>
Figs. 7, 8:
<taxonomicName id="4C571DE1FFD08C21FE78913F39F21714" authorityName="Tennessen &amp; Garrison" authorityYear="2023" box="[448,678,1584,1608]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="398" phylum="Arthropoda" rank="species" species="parmula">
<emphasis id="B923BA70FFD08C21FE78913F39F21714" box="[448,678,1584,1608]" italics="true" pageId="3" pageNumber="398">Epipleoneura parmula</emphasis>
</taxonomicName>
♁ holotype; Fig. 9
<taxonomicName id="4C571DE1FFD08C21FCD6913F3F131714" authorityName="De Marmels" authorityYear="1989" box="[878,1095,1584,1608]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="398" phylum="Arthropoda" rank="species" species="ocuene">
<emphasis id="B923BA70FFD08C21FCD6913F3F131714" box="[878,1095,1584,1608]" italics="true" pageId="3" pageNumber="398">Epipleoneura ocuene</emphasis>
</taxonomicName>
♁ paratype; Fig. 10
<taxonomicName id="4C571DE1FFD08C21FAA1913E3B8B1730" class="Insecta" family="Coenagrionidae" genus="Epipeoneura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="398" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B923BA70FFD08C21FAA1913E3B8B1730" italics="true" pageId="3" pageNumber="398">Epipeoneura pallida</emphasis>
</taxonomicName>
♁. Fig. 7 caudal appendages, a (lateral), b. (lateroposterior) and c (posterior) views; Figs. 8–10 caudal appendages, posterior view.
</paragraph>
</caption>
<paragraph id="8BE86662FFD08C26FF7F91EC3AAB1041" blockId="3.[151,1437,1763,2041]" lastBlockId="4.[151,1436,151,285]" lastPageId="4" lastPageNumber="399" pageId="3" pageNumber="398">
Abdomen. Predominantly brown dorsally (as in
<figureCitation id="136C7AE7FFD08C21FD4991EC386717A1" box="[753,819,1763,1789]" captionStart="FIGURES 1–6" captionStartId="2.[152,270,1827,1851]" captionTargetBox="[155,1431,241,1800]" captionTargetId="figure-58@2.[151,1435,237,1804]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 2, 5, 6: Epipleoneura parmula ♁ paratype; Fig. 3 Epipleoneura ocuene ♁ paratype; 4 Epipeoneura pallida ♁. Fig. 1. Habitus; Figs. 2–4 head and thorax, dorsolateral (2) and lateral views; Fig. 5 wings; 6 genital ligula ectal (left) and lateral (right) views." figureDoi="http://doi.org/10.5281/zenodo.8334531" httpUri="https://zenodo.org/record/8334531/files/figure.png" pageId="3" pageNumber="398">Fig. 1</figureCitation>
); S1 and S2 tan laterally and ventrally; S3–7 with an extremely small anterior tan band, tan ventrally, brown laterally except light brown in posterior third of each segment; S8 largely tan laterally, S9 dark brown laterally, S10 dark brown. Anal appendages short, cercus and paraproct about ½ length of S10; cercus in lateral view with a dorsoposteriorly directed dorsal branch bearing an apical hook and a lobe-like ventral branch (
<figureCitation id="136C7AE7FFD08C21FE0C907C395216D1" box="[436,518,1907,1933]" captionStart="FIGURES 7–10" captionStartId="3.[152,270,1584,1608]" captionTargetBox="[155,1431,185,1557]" captionTargetId="figure-19@3.[151,1435,181,1561]" captionTargetPageId="3" captionText="FIGURES 7–10. Figs. 7, 8: Epipleoneura parmula ♁ holotype; Fig. 9 Epipleoneura ocuene ♁ paratype; Fig. 10 Epipeoneura pallida ♁. Fig. 7 caudal appendages, a (lateral), b. (lateroposterior) and c (posterior) views; Figs. 8–10 caudal appendages, posterior view." figureDoi="http://doi.org/10.5281/zenodo.8334533" httpUri="https://zenodo.org/record/8334533/files/figure.png" pageId="3" pageNumber="398">Fig. 7a</figureCitation>
), in posterior view with a reduced, rounded medial process bearing a small black tooth (
<figureCitation id="136C7AE7FFD08C21FF5990983A6716ED" box="[225,307,1943,1969]" captionStart="FIGURES 7–10" captionStartId="3.[152,270,1584,1608]" captionTargetBox="[155,1431,185,1557]" captionTargetId="figure-19@3.[151,1435,181,1561]" captionTargetPageId="3" captionText="FIGURES 7–10. Figs. 7, 8: Epipleoneura parmula ♁ holotype; Fig. 9 Epipleoneura ocuene ♁ paratype; Fig. 10 Epipeoneura pallida ♁. Fig. 7 caudal appendages, a (lateral), b. (lateroposterior) and c (posterior) views; Figs. 8–10 caudal appendages, posterior view." figureDoi="http://doi.org/10.5281/zenodo.8334533" httpUri="https://zenodo.org/record/8334533/files/figure.png" pageId="3" pageNumber="398">Fig. 7c</figureCitation>
); epiproct oriented vertically, in lateral view only partly visible (
<figureCitation id="136C7AE7FFD08C21FBB390983F0916ED" box="[1035,1117,1943,1969]" captionStart="FIGURES 7–10" captionStartId="3.[152,270,1584,1608]" captionTargetBox="[155,1431,185,1557]" captionTargetId="figure-19@3.[151,1435,181,1561]" captionTargetPageId="3" captionText="FIGURES 7–10. Figs. 7, 8: Epipleoneura parmula ♁ holotype; Fig. 9 Epipleoneura ocuene ♁ paratype; Fig. 10 Epipeoneura pallida ♁. Fig. 7 caudal appendages, a (lateral), b. (lateroposterior) and c (posterior) views; Figs. 8–10 caudal appendages, posterior view." figureDoi="http://doi.org/10.5281/zenodo.8334533" httpUri="https://zenodo.org/record/8334533/files/figure.png" pageId="3" pageNumber="398">Fig. 7a</figureCitation>
), in posterior view with two fused, elongate parallel lobes, the tips angled medially and produced as nipple-like lobes (
<figureCitation id="136C7AE7FFD08C21FB3090B43E4F1689" box="[1160,1307,1979,2005]" captionStart="FIGURES 7–10" captionStartId="3.[152,270,1584,1608]" captionTargetBox="[155,1431,185,1557]" captionTargetId="figure-19@3.[151,1435,181,1561]" captionTargetPageId="3" captionText="FIGURES 7–10. Figs. 7, 8: Epipleoneura parmula ♁ holotype; Fig. 9 Epipleoneura ocuene ♁ paratype; Fig. 10 Epipeoneura pallida ♁. Fig. 7 caudal appendages, a (lateral), b. (lateroposterior) and c (posterior) views; Figs. 8–10 caudal appendages, posterior view." figureDoi="http://doi.org/10.5281/zenodo.8334533" httpUri="https://zenodo.org/record/8334533/files/figure.png" pageId="3" pageNumber="398">Figs. 7b, c, 8</figureCitation>
); paraproct in lateral view about same length as cercus, apex conical and acute (
<figureCitation id="136C7AE7FFD08C21FC1090D038AF16A5" box="[936,1019,2015,2041]" captionStart="FIGURES 7–10" captionStartId="3.[152,270,1584,1608]" captionTargetBox="[155,1431,185,1557]" captionTargetId="figure-19@3.[151,1435,181,1561]" captionTargetPageId="3" captionText="FIGURES 7–10. Figs. 7, 8: Epipleoneura parmula ♁ holotype; Fig. 9 Epipleoneura ocuene ♁ paratype; Fig. 10 Epipeoneura pallida ♁. Fig. 7 caudal appendages, a (lateral), b. (lateroposterior) and c (posterior) views; Figs. 8–10 caudal appendages, posterior view." figureDoi="http://doi.org/10.5281/zenodo.8334533" httpUri="https://zenodo.org/record/8334533/files/figure.png" pageId="3" pageNumber="398">Fig. 7a</figureCitation>
). Genital ligula lacking internal fold but with a small, sharply pointed inner process (best seen in lateral view when ligula is relaxed and carefully flexed away from basal segment (
<figureCitation id="136C7AE7FFD78C26FE7A97B439511189" box="[450,517,187,213]" captionStart="FIGURES 1–6" captionStartId="2.[152,270,1827,1851]" captionTargetBox="[155,1431,241,1800]" captionTargetId="figure-58@2.[151,1435,237,1804]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 2, 5, 6: Epipleoneura parmula ♁ paratype; Fig. 3 Epipleoneura ocuene ♁ paratype; 4 Epipeoneura pallida ♁. Fig. 1. Habitus; Figs. 2–4 head and thorax, dorsolateral (2) and lateral views; Fig. 5 wings; 6 genital ligula ectal (left) and lateral (right) views." figureDoi="http://doi.org/10.5281/zenodo.8334531" httpUri="https://zenodo.org/record/8334531/files/figure.png" pageId="4" pageNumber="399">Fig. 6</figureCitation>
); flexure with a pair of posterolateral membranous pedunculate processes (
<figureCitation id="136C7AE7FFD78C26FAF197B43ED91189" box="[1353,1421,187,213]" captionStart="FIGURES 1–6" captionStartId="2.[152,270,1827,1851]" captionTargetBox="[155,1431,241,1800]" captionTargetId="figure-58@2.[151,1435,237,1804]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 2, 5, 6: Epipleoneura parmula ♁ paratype; Fig. 3 Epipleoneura ocuene ♁ paratype; 4 Epipeoneura pallida ♁. Fig. 1. Habitus; Figs. 2–4 head and thorax, dorsolateral (2) and lateral views; Fig. 5 wings; 6 genital ligula ectal (left) and lateral (right) views." figureDoi="http://doi.org/10.5281/zenodo.8334531" httpUri="https://zenodo.org/record/8334531/files/figure.png" pageId="4" pageNumber="399">Fig. 6</figureCitation>
); terminal segment in ectal view with lateral edges upcurved, apices expanded and rounded (
<figureCitation id="136C7AE7FFD78C26FB2797D03FB011A5" box="[1183,1252,223,249]" captionStart="FIGURES 1–6" captionStartId="2.[152,270,1827,1851]" captionTargetBox="[155,1431,241,1800]" captionTargetId="figure-58@2.[151,1435,237,1804]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 2, 5, 6: Epipleoneura parmula ♁ paratype; Fig. 3 Epipleoneura ocuene ♁ paratype; 4 Epipeoneura pallida ♁. Fig. 1. Habitus; Figs. 2–4 head and thorax, dorsolateral (2) and lateral views; Fig. 5 wings; 6 genital ligula ectal (left) and lateral (right) views." figureDoi="http://doi.org/10.5281/zenodo.8334531" httpUri="https://zenodo.org/record/8334531/files/figure.png" pageId="4" pageNumber="399">Fig. 6</figureCitation>
), in lateral view tapered to pointed apex (
<figureCitation id="136C7AE7FFD78C26FE15960C3AA51041" box="[429,497,259,285]" captionStart="FIGURES 1–6" captionStartId="2.[152,270,1827,1851]" captionTargetBox="[155,1431,241,1800]" captionTargetId="figure-58@2.[151,1435,237,1804]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 2, 5, 6: Epipleoneura parmula ♁ paratype; Fig. 3 Epipleoneura ocuene ♁ paratype; 4 Epipeoneura pallida ♁. Fig. 1. Habitus; Figs. 2–4 head and thorax, dorsolateral (2) and lateral views; Fig. 5 wings; 6 genital ligula ectal (left) and lateral (right) views." figureDoi="http://doi.org/10.5281/zenodo.8334531" httpUri="https://zenodo.org/record/8334531/files/figure.png" pageId="4" pageNumber="399">Fig. 6</figureCitation>
).
</paragraph>
<caption id="DF2836EAFFD78C26FF2091B03F37178B" ID-DOI="http://doi.org/10.5281/zenodo.8334535" ID-Zenodo-Dep="8334535" box="[152,1123,1727,1751]" httpUri="https://zenodo.org/record/8334535/files/figure.png" pageId="4" pageNumber="399" startId="4.[152,255,1727,1751]" targetBox="[157,1429,315,1698]" targetPageId="4" targetType="figure">
<paragraph id="8BE86662FFD78C26FF2091B03F37178B" blockId="4.[152,1123,1727,1751]" box="[152,1123,1727,1751]" pageId="4" pageNumber="399">
<emphasis id="B923BA70FFD78C26FF2091B03A70178B" bold="true" box="[152,292,1727,1751]" pageId="4" pageNumber="399">FIGURE 11.</emphasis>
Distribution of
<taxonomicName id="4C571DE1FFD78C26FE7E91B039F8178B" authorityName="Tennessen &amp; Garrison" authorityYear="2023" box="[454,684,1727,1751]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="399" phylum="Arthropoda" rank="species" species="parmula">
<emphasis id="B923BA70FFD78C26FE7E91B039F8178B" box="[454,684,1727,1751]" italics="true" pageId="4" pageNumber="399">Epipleoneura parmula</emphasis>
</taxonomicName>
within Napo and Orellana States, Ecuador.
</paragraph>
</caption>
<paragraph id="8BE86662FFD78C26FF7F902538861618" blockId="4.[151,1436,1833,2040]" box="[199,978,1833,1860]" pageId="4" pageNumber="399">
Measurements of
<typeStatus id="54ECD8C0FFD78C26FE2890263AA2161F" box="[400,502,1833,1859]" pageId="4" pageNumber="399">holotype</typeStatus>
. Hw 19.0, abdomen 29.0, total length 34.0
</paragraph>
<paragraph id="8BE86662FFD78C26FF7F90423A7116EC" blockId="4.[151,1436,1833,2040]" pageId="4" pageNumber="399">
Variation in
<typeStatus id="54ECD8C0FFD78C26FEE890413AE41634" box="[336,432,1870,1896]" pageId="4" pageNumber="399">paratype</typeStatus>
males. Some males with epicranium, dorsum of prothorax and mesepisterna dark metallic green as in
<figureCitation id="136C7AE7FFD78C26FEA2907E3A0B16D0" box="[282,351,1905,1932]" captionStart="FIGURES 1–6" captionStartId="2.[152,270,1827,1851]" captionTargetBox="[155,1431,241,1800]" captionTargetId="figure-58@2.[151,1435,237,1804]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 2, 5, 6: Epipleoneura parmula ♁ paratype; Fig. 3 Epipleoneura ocuene ♁ paratype; 4 Epipeoneura pallida ♁. Fig. 1. Habitus; Figs. 2–4 head and thorax, dorsolateral (2) and lateral views; Fig. 5 wings; 6 genital ligula ectal (left) and lateral (right) views." figureDoi="http://doi.org/10.5281/zenodo.8334531" httpUri="https://zenodo.org/record/8334531/files/figure.png" pageId="4" pageNumber="399">Fig. 2</figureCitation>
. Pterostigma surmounting 0.7–1.0 cells; postnodal crossveins: Fw 11–14 (usually 12), Hw 9–11 (usually 10).
</paragraph>
<paragraph id="8BE86662FFD78C26FF7F90B53AF216A4" blockId="4.[151,1436,1833,2040]" pageId="4" pageNumber="399">
Measurements of
<typeStatus id="54ECD8C0FFD78C26FE2190B539501688" box="[409,516,1978,2004]" pageId="4" pageNumber="399">paratypes</typeStatus>
(mean in parentheses). Hw 17.3–18.7 (17.9), abdomen 28.5–30.5 (29.2), total length 33.1–35.5 (34.0).
</paragraph>
</subSubSection>
<subSubSection id="C34D35E9FFD68C27FF7F9799392C13F5" pageId="5" pageNumber="400" type="diagnosis">
<paragraph id="8BE86662FFD68C27FF7F9799392C13F5" blockId="5.[151,1437,150,1113]" pageId="5" pageNumber="400">
<emphasis id="B923BA70FFD68C27FF7F97993A1011EC" bold="true" box="[199,324,150,176]" pageId="5" pageNumber="400">Diagnosis:</emphasis>
In
<bibRefCitation id="EFC61B93FFD68C27FED19798394211ED" author="Pessacq, P." box="[361,534,151,177]" pageId="5" pageNumber="400" pagination="201 - 234" refId="ref3039" refString="Pessacq, P. (2014) Synopsis of Epipleoneura (Zygoptera, Coenagrionidae, &quot; Protoneuridae &quot;), with emphasis on its Brazilian species. Zootaxa, 3872 (3), 201 - 234. https: // doi. org / 10.11646 / zootaxa. 3872.3.1" type="journal article" year="2014">Pessacq (2014)</bibRefCitation>
, the new species keys out to couplet
<quantity id="4CAFCB87FFD68C27FC0B979838B811ED" box="[947,1004,151,177]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" pageId="5" pageNumber="400" unit="in" value="23.0">23 in</quantity>
which
<taxonomicName id="4C571DE1FFD68C27FB8697983E5911ED" authority="Racenis" authorityName="Racenis" box="[1086,1293,151,177]" class="Insecta" family="Coenagrionidae" genus="Epipeoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B923BA70FFD68C27FB8697983FF811ED" box="[1086,1196,151,177]" italics="true" pageId="5" pageNumber="400">E. pallida</emphasis>
Rácenis
</taxonomicName>
(
<figureCitation id="136C7AE7FFD68C27FAA497983E3F11ED" box="[1308,1387,151,177]" captionStart="FIGURES 1–6" captionStartId="2.[152,270,1827,1851]" captionTargetBox="[155,1431,241,1800]" captionTargetId="figure-58@2.[151,1435,237,1804]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 2, 5, 6: Epipleoneura parmula ♁ paratype; Fig. 3 Epipleoneura ocuene ♁ paratype; 4 Epipeoneura pallida ♁. Fig. 1. Habitus; Figs. 2–4 head and thorax, dorsolateral (2) and lateral views; Fig. 5 wings; 6 genital ligula ectal (left) and lateral (right) views." figureDoi="http://doi.org/10.5281/zenodo.8334531" httpUri="https://zenodo.org/record/8334531/files/figure.png" pageId="5" pageNumber="400">Figs. 4</figureCitation>
,
<figureCitation id="136C7AE7FFD68C27FACF97983EC011ED" box="[1399,1428,151,177]" captionStart="FIGURES 7–10" captionStartId="3.[152,270,1584,1608]" captionTargetBox="[155,1431,185,1557]" captionTargetId="figure-19@3.[151,1435,181,1561]" captionTargetPageId="3" captionText="FIGURES 7–10. Figs. 7, 8: Epipleoneura parmula ♁ holotype; Fig. 9 Epipleoneura ocuene ♁ paratype; Fig. 10 Epipeoneura pallida ♁. Fig. 7 caudal appendages, a (lateral), b. (lateroposterior) and c (posterior) views; Figs. 8–10 caudal appendages, posterior view." figureDoi="http://doi.org/10.5281/zenodo.8334533" httpUri="https://zenodo.org/record/8334533/files/figure.png" pageId="5" pageNumber="400">10</figureCitation>
) and
<taxonomicName id="4C571DE1FFD68C27FF7E97B43A971189" authority="De Marmels" authorityName="De Marmels" authorityYear="1989" box="[198,451,187,213]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="ocuene">
<emphasis id="B923BA70FFD68C27FF7E97B43A671188" box="[198,307,187,212]" italics="true" pageId="5" pageNumber="400">E. ocuene</emphasis>
De Marmels
</taxonomicName>
(
<figureCitation id="136C7AE7FFD68C27FE6997B4394A1189" box="[465,542,187,213]" captionStart="FIGURES 1–6" captionStartId="2.[152,270,1827,1851]" captionTargetBox="[155,1431,241,1800]" captionTargetId="figure-58@2.[151,1435,237,1804]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 2, 5, 6: Epipleoneura parmula ♁ paratype; Fig. 3 Epipleoneura ocuene ♁ paratype; 4 Epipeoneura pallida ♁. Fig. 1. Habitus; Figs. 2–4 head and thorax, dorsolateral (2) and lateral views; Fig. 5 wings; 6 genital ligula ectal (left) and lateral (right) views." figureDoi="http://doi.org/10.5281/zenodo.8334531" httpUri="https://zenodo.org/record/8334531/files/figure.png" pageId="5" pageNumber="400">Figs. 3</figureCitation>
,
<figureCitation id="136C7AE7FFD68C27FD9297B4396D1189" box="[554,569,187,213]" captionStart="FIGURES 7–10" captionStartId="3.[152,270,1584,1608]" captionTargetBox="[155,1431,185,1557]" captionTargetId="figure-19@3.[151,1435,181,1561]" captionTargetPageId="3" captionText="FIGURES 7–10. Figs. 7, 8: Epipleoneura parmula ♁ holotype; Fig. 9 Epipleoneura ocuene ♁ paratype; Fig. 10 Epipeoneura pallida ♁. Fig. 7 caudal appendages, a (lateral), b. (lateroposterior) and c (posterior) views; Figs. 8–10 caudal appendages, posterior view." figureDoi="http://doi.org/10.5281/zenodo.8334533" httpUri="https://zenodo.org/record/8334533/files/figure.png" pageId="5" pageNumber="400">9</figureCitation>
) are separated. The dorsal branch of the cercus in those two species is directed dorsally compared to dorsoposteriorly in
<taxonomicName id="4C571DE1FFD68C27FDDE97D039B011A5" authorityName="Tennessen &amp; Garrison" authorityYear="2023" box="[614,740,223,249]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="parmula">
<emphasis id="B923BA70FFD68C27FDDE97D039B011A5" box="[614,740,223,249]" italics="true" pageId="5" pageNumber="400">E. parmula</emphasis>
</taxonomicName>
. Compared to
<taxonomicName id="4C571DE1FFD68C27FC3297D03F5D11A5" authorityName="Tennessen &amp; Garrison" authorityYear="2023" box="[906,1033,223,249]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="parmula">
<emphasis id="B923BA70FFD68C27FC3297D03F5D11A5" box="[906,1033,223,249]" italics="true" pageId="5" pageNumber="400">E. parmula</emphasis>
</taxonomicName>
, the epiproct is shorter in
<taxonomicName id="4C571DE1FFD68C27FA9697D03EC811A5" box="[1326,1436,223,249]" class="Insecta" family="Coenagrionidae" genus="Epipeoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B923BA70FFD68C27FA9697D03EC811A5" box="[1326,1436,223,249]" italics="true" pageId="5" pageNumber="400">E. pallida</emphasis>
</taxonomicName>
(
<figureCitation id="136C7AE7FFD68C27FF27960C3BA61041" box="[159,242,259,285]" captionStart="FIGURES 7–10" captionStartId="3.[152,270,1584,1608]" captionTargetBox="[155,1431,185,1557]" captionTargetId="figure-19@3.[151,1435,181,1561]" captionTargetPageId="3" captionText="FIGURES 7–10. Figs. 7, 8: Epipleoneura parmula ♁ holotype; Fig. 9 Epipleoneura ocuene ♁ paratype; Fig. 10 Epipeoneura pallida ♁. Fig. 7 caudal appendages, a (lateral), b. (lateroposterior) and c (posterior) views; Figs. 8–10 caudal appendages, posterior view." figureDoi="http://doi.org/10.5281/zenodo.8334533" httpUri="https://zenodo.org/record/8334533/files/figure.png" pageId="5" pageNumber="400">Fig. 10</figureCitation>
) and longer in
<taxonomicName id="4C571DE1FFD68C27FE1A960C39451040" authorityName="De Marmels" authorityYear="1989" box="[418,529,259,284]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="ocuene">
<emphasis id="B923BA70FFD68C27FE1A960C39451040" box="[418,529,259,284]" italics="true" pageId="5" pageNumber="400">E. ocuene</emphasis>
</taxonomicName>
(
<figureCitation id="136C7AE7FFD68C27FD9A960C393C1041" box="[546,616,259,285]" captionStart="FIGURES 7–10" captionStartId="3.[152,270,1584,1608]" captionTargetBox="[155,1431,185,1557]" captionTargetId="figure-19@3.[151,1435,181,1561]" captionTargetPageId="3" captionText="FIGURES 7–10. Figs. 7, 8: Epipleoneura parmula ♁ holotype; Fig. 9 Epipleoneura ocuene ♁ paratype; Fig. 10 Epipeoneura pallida ♁. Fig. 7 caudal appendages, a (lateral), b. (lateroposterior) and c (posterior) views; Figs. 8–10 caudal appendages, posterior view." figureDoi="http://doi.org/10.5281/zenodo.8334533" httpUri="https://zenodo.org/record/8334533/files/figure.png" pageId="5" pageNumber="400">Fig. 9</figureCitation>
). The gap between the two branches in both
<taxonomicName id="4C571DE1FFD68C27FBD1960C3F8C1040" authorityName="De Marmels" authorityYear="1989" box="[1129,1240,259,284]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="ocuene">
<emphasis id="B923BA70FFD68C27FBD1960C3F8C1040" box="[1129,1240,259,284]" italics="true" pageId="5" pageNumber="400">E. ocuene</emphasis>
</taxonomicName>
and
<taxonomicName id="4C571DE1FFD68C27FAA9960C3ED51041" box="[1297,1409,259,285]" class="Insecta" family="Coenagrionidae" genus="Epipeoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B923BA70FFD68C27FAA9960C3ED51041" box="[1297,1409,259,285]" italics="true" pageId="5" pageNumber="400">E. pallida</emphasis>
</taxonomicName>
is larger than in
<taxonomicName id="4C571DE1FFD68C27FE8B96283AE5101D" authorityName="Tennessen &amp; Garrison" authorityYear="2023" box="[307,433,295,321]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="parmula">
<emphasis id="B923BA70FFD68C27FE8B96283AE5101D" box="[307,433,295,321]" italics="true" pageId="5" pageNumber="400">E. parmula</emphasis>
</taxonomicName>
. Both
<taxonomicName id="4C571DE1FFD68C27FE449628392E101D" authorityName="Tennessen &amp; Garrison" authorityYear="2023" box="[508,634,295,321]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="parmula">
<emphasis id="B923BA70FFD68C27FE449628392E101D" box="[508,634,295,321]" italics="true" pageId="5" pageNumber="400">E. parmula</emphasis>
</taxonomicName>
(
<figureCitation id="136C7AE7FFD68C27FD31962839A7101D" box="[649,755,295,321]" captionStart="FIGURES 1–6" captionStartId="2.[152,270,1827,1851]" captionTargetBox="[155,1431,241,1800]" captionTargetId="figure-58@2.[151,1435,237,1804]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 2, 5, 6: Epipleoneura parmula ♁ paratype; Fig. 3 Epipleoneura ocuene ♁ paratype; 4 Epipeoneura pallida ♁. Fig. 1. Habitus; Figs. 2–4 head and thorax, dorsolateral (2) and lateral views; Fig. 5 wings; 6 genital ligula ectal (left) and lateral (right) views." figureDoi="http://doi.org/10.5281/zenodo.8334531" httpUri="https://zenodo.org/record/8334531/files/figure.png" pageId="5" pageNumber="400">Figs. 1, 2</figureCitation>
) and
<taxonomicName id="4C571DE1FFD68C27FC89962838CB101D" box="[817,927,295,321]" class="Insecta" family="Coenagrionidae" genus="Epipeoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B923BA70FFD68C27FC89962838CB101D" box="[817,927,295,321]" italics="true" pageId="5" pageNumber="400">E. pallida</emphasis>
</taxonomicName>
(
<figureCitation id="136C7AE7FFD68C27FC17962838A7101D" box="[943,1011,295,321]" captionStart="FIGURES 1–6" captionStartId="2.[152,270,1827,1851]" captionTargetBox="[155,1431,241,1800]" captionTargetId="figure-58@2.[151,1435,237,1804]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 2, 5, 6: Epipleoneura parmula ♁ paratype; Fig. 3 Epipleoneura ocuene ♁ paratype; 4 Epipeoneura pallida ♁. Fig. 1. Habitus; Figs. 2–4 head and thorax, dorsolateral (2) and lateral views; Fig. 5 wings; 6 genital ligula ectal (left) and lateral (right) views." figureDoi="http://doi.org/10.5281/zenodo.8334531" httpUri="https://zenodo.org/record/8334531/files/figure.png" pageId="5" pageNumber="400">Fig. 4</figureCitation>
, but see below) possess a pale narrow antehumeral stripe that is lacking in
<taxonomicName id="4C571DE1FFD68C27FD96964439C81038" authorityName="De Marmels" authorityYear="1989" box="[558,668,331,356]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="ocuene">
<emphasis id="B923BA70FFD68C27FD96964439C81038" box="[558,668,331,356]" italics="true" pageId="5" pageNumber="400">E. ocuene</emphasis>
</taxonomicName>
(
<figureCitation id="136C7AE7FFD68C27FD13964439A41039" box="[683,752,331,357]" captionStart="FIGURES 1–6" captionStartId="2.[152,270,1827,1851]" captionTargetBox="[155,1431,241,1800]" captionTargetId="figure-58@2.[151,1435,237,1804]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 2, 5, 6: Epipleoneura parmula ♁ paratype; Fig. 3 Epipleoneura ocuene ♁ paratype; 4 Epipeoneura pallida ♁. Fig. 1. Habitus; Figs. 2–4 head and thorax, dorsolateral (2) and lateral views; Fig. 5 wings; 6 genital ligula ectal (left) and lateral (right) views." figureDoi="http://doi.org/10.5281/zenodo.8334531" httpUri="https://zenodo.org/record/8334531/files/figure.png" pageId="5" pageNumber="400">Fig. 3</figureCitation>
). The original description of
<taxonomicName id="4C571DE1FFD68C27FB8E96443E021039" authority="(Racenis 1960)" baseAuthorityName="Racenis" baseAuthorityYear="1960" box="[1078,1366,331,357]" class="Insecta" family="Coenagrionidae" genus="Epipeoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B923BA70FFD68C27FB8E96443FF01039" box="[1078,1188,331,357]" italics="true" pageId="5" pageNumber="400">E. pallida</emphasis>
(
<bibRefCitation id="EFC61B93FFD68C27FB0B96443E191039" author="Racenis, J." box="[1203,1357,331,357]" pageId="5" pageNumber="400" pagination="25 - 42" refId="ref3174" refString="Racenis, J. (1960) Cuatro nuevas especies del genero Epipleoneura (Odonata: Protoneuridae). Acta Biologia Venezuelica, 3 (2), 25 - 42." type="journal article" year="1960">Rácenis 1960</bibRefCitation>
)
</taxonomicName>
based on the
<typeStatus id="54ECD8C0FFD68C27FF5E96603A1C10D5" box="[230,328,367,393]" pageId="5" pageNumber="400">holotype</typeStatus>
describes a largely pale thorax (“Cada mesepisternón, verde-metálico, color que ocupa más de la mitad de este esclerito; la parte posterior de los mesepisterna y todos los mesepimera, pardo-rojizos; en la parte dorsal de los mesepimera hay una mancha irregular, negruzca y con un brillo metálico; el resto del sintorax es de un color pardo-amarillento.”) with a metallic middorsal stripe confined to the middle half of each mesepisternum (hence the specific epithet
<taxonomicName id="4C571DE1FFD68C27FE7C96F039471345" baseAuthorityName="Racenis" baseAuthorityYear="1960" box="[452,531,511,537]" class="Insecta" family="Coenagrionidae" genus="Epipeoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B923BA70FFD68C27FE7C96F039471345" box="[452,531,511,537]" italics="true" pageId="5" pageNumber="400">pallida</emphasis>
</taxonomicName>
). Our small series of males from
<collectingCountry id="F34026F2FFD68C27FC3596F038941345" box="[909,960,511,537]" name="Peru" pageId="5" pageNumber="400">Peru</collectingCountry>
that we have identified as
<taxonomicName id="4C571DE1FFD68C27FB5596F03E081345" baseAuthorityName="Racenis" baseAuthorityYear="1960" box="[1261,1372,511,537]" class="Insecta" family="Coenagrionidae" genus="Epipeoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B923BA70FFD68C27FB5596F03E081345" box="[1261,1372,511,537]" italics="true" pageId="5" pageNumber="400">E. pallida</emphasis>
</taxonomicName>
has a more extensive dark thoracic pattern as shown in
<figureCitation id="136C7AE7FFD68C27FD79952C38511361" box="[705,773,547,573]" captionStart="FIGURES 1–6" captionStartId="2.[152,270,1827,1851]" captionTargetBox="[155,1431,241,1800]" captionTargetId="figure-58@2.[151,1435,237,1804]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, 2, 5, 6: Epipleoneura parmula ♁ paratype; Fig. 3 Epipleoneura ocuene ♁ paratype; 4 Epipeoneura pallida ♁. Fig. 1. Habitus; Figs. 2–4 head and thorax, dorsolateral (2) and lateral views; Fig. 5 wings; 6 genital ligula ectal (left) and lateral (right) views." figureDoi="http://doi.org/10.5281/zenodo.8334531" httpUri="https://zenodo.org/record/8334531/files/figure.png" pageId="5" pageNumber="400">Fig. 4</figureCitation>
. However, the male appendages of our Peruvian specimens match those figured by Rácenis for
<taxonomicName id="4C571DE1FFD68C27FD95954839C8133D" baseAuthorityName="Racenis" baseAuthorityYear="1960" box="[557,668,583,609]" class="Insecta" family="Coenagrionidae" genus="Epipeoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B923BA70FFD68C27FD95954839C8133D" box="[557,668,583,609]" italics="true" pageId="5" pageNumber="400">E. pallida</emphasis>
</taxonomicName>
in all essentials. The genital ligula of all three species is essentially the same though we have not observed the presence of the small, sharply pointed inner process in
<taxonomicName id="4C571DE1FFD68C27FB5895643E1A13D8" authorityName="De Marmels" authorityYear="1989" box="[1248,1358,619,644]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="ocuene">
<emphasis id="B923BA70FFD68C27FB5895643E1A13D8" box="[1248,1358,619,644]" italics="true" pageId="5" pageNumber="400">E. ocuene</emphasis>
</taxonomicName>
and
<taxonomicName id="4C571DE1FFD68C27FA3C95643BB213F5" baseAuthorityName="Racenis" baseAuthorityYear="1960" class="Insecta" family="Coenagrionidae" genus="Epipeoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B923BA70FFD68C27FA3C95643BB213F5" italics="true" pageId="5" pageNumber="400">E. pallida</emphasis>
</taxonomicName>
that is characteristic of
<taxonomicName id="4C571DE1FFD68C27FE4B9580392513F5" authorityName="Tennessen &amp; Garrison" authorityYear="2023" box="[499,625,655,681]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="parmula">
<emphasis id="B923BA70FFD68C27FE4B9580392513F5" box="[499,625,655,681]" italics="true" pageId="5" pageNumber="400">E. parmula</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34D35E9FFD68C27FF7F95BD39421505" pageId="5" pageNumber="400" type="discussion">
<paragraph id="8BE86662FFD68C27FF7F95BD3E761201" blockId="5.[151,1437,150,1113]" pageId="5" pageNumber="400">
<emphasis id="B923BA70FFD68C27FF7F95BD3A1B1390" bold="true" box="[199,335,690,716]" pageId="5" pageNumber="400">Discussion:</emphasis>
The presence of an inner process on the genital ligula of
<taxonomicName id="4C571DE1FFD68C27FC7F95BC3F111391" authorityName="Tennessen &amp; Garrison" authorityYear="2023" box="[967,1093,691,717]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="parmula">
<emphasis id="B923BA70FFD68C27FC7F95BC3F111391" box="[967,1093,691,717]" italics="true" pageId="5" pageNumber="400">E. parmula</emphasis>
</taxonomicName>
was unexpected as
<bibRefCitation id="EFC61B93FFD68C27FA9895BC3A5213AD" author="Garrison, R. W. &amp; von Ellenrieder, N. &amp; Louton, J. A." pageId="5" pageNumber="400" refId="ref2952" refString="Garrison, R. W., von Ellenrieder, N. &amp; Louton, J. A. (2010) Damselfly genera of the New World: an illustrated and annotated key to the Zygoptera. The Johns Hopkins University Press, Baltimore, Maryland, 383 pp." type="book" year="2010">
Garrison
<emphasis id="B923BA70FFD68C27FA3095BC3BF913AD" italics="true" pageId="5" pageNumber="400">et al</emphasis>
. (2010)
</bibRefCitation>
stated that
<taxonomicName id="4C571DE1FFD68C27FE3D95D8394913AD" authorityName="Williamson" authorityYear="1915" box="[389,541,727,753]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="genus">
<emphasis id="B923BA70FFD68C27FE3D95D8394913AD" box="[389,541,727,753]" italics="true" pageId="5" pageNumber="400">Epipleoneura</emphasis>
</taxonomicName>
lacked an inner process distal to the flexure, and Pessacq did not report an inner process in any of the 25 species he examined and illustrated. We examined 18 species for an inner process similar to that of
<taxonomicName id="4C571DE1FFD68C27FF5D94103A361265" authorityName="Tennessen &amp; Garrison" authorityYear="2023" box="[229,354,799,825]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="parmula">
<emphasis id="B923BA70FFD68C27FF5D94103A361265" box="[229,354,799,825]" italics="true" pageId="5" pageNumber="400">E. parmula</emphasis>
</taxonomicName>
and found it present only in
<taxonomicName id="4C571DE1FFD68C27FD18941038D31265" authority="(Selys)" baseAuthorityName="Selys" box="[672,903,799,825]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="B923BA70FFD68C27FD189410387A1265" box="[672,814,799,825]" italics="true" pageId="5" pageNumber="400">E. humeralis</emphasis>
(Selys)
</taxonomicName>
. In addition, reexamination of
<taxonomicName id="4C571DE1FFD68C27FB6494103EC91265" authority="Santos" authorityName="Santos" authorityYear="1964" box="[1244,1437,799,825]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="haroldi">
<emphasis id="B923BA70FFD68C27FB6494103E1F1265" box="[1244,1355,799,825]" italics="true" pageId="5" pageNumber="400">E. haroldi</emphasis>
Santos
</taxonomicName>
showed that it has a pair of hyaline, entally directed inner processes at the base of the terminal segment.
</paragraph>
<paragraph id="8BE86662FFD68C27FF7F946839421505" blockId="5.[151,1437,150,1113]" pageId="5" pageNumber="400">
The typical habitat for
<taxonomicName id="4C571DE1FFD68C27FE6A9468393E12DD" authorityName="Williamson" authorityYear="1915" box="[466,618,871,897]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="genus">
<emphasis id="B923BA70FFD68C27FE6A9468393E12DD" box="[466,618,871,897]" italics="true" pageId="5" pageNumber="400">Epipleoneura</emphasis>
</taxonomicName>
is slow to moderately flowing streams (
<bibRefCitation id="EFC61B93FFD68C27FB8694683FBC12DD" author="Lencioni, F. A. A." box="[1086,1256,870,897]" pageId="5" pageNumber="400" refId="ref3003" refString="Lencioni, F. A. A. (2005) Damselflies of Brazil. An illustrated identification guide. I. The non-Coenagrionidae families. All Print Editora, Sao Paulo, 324 pp." type="book" year="2005">Lencioni 2005</bibRefCitation>
;
<bibRefCitation id="EFC61B93FFD68C27FB4E94683B8712F9" author="Garrison, R. W. &amp; von Ellenrieder, N. &amp; Louton, J. A." pageId="5" pageNumber="400" refId="ref2952" refString="Garrison, R. W., von Ellenrieder, N. &amp; Louton, J. A. (2010) Damselfly genera of the New World: an illustrated and annotated key to the Zygoptera. The Johns Hopkins University Press, Baltimore, Maryland, 383 pp." type="book" year="2010">
Garrison
<emphasis id="B923BA70FFD68C27FADA94683EC112DD" box="[1378,1429,871,897]" italics="true" pageId="5" pageNumber="400">et al</emphasis>
. 2010
</bibRefCitation>
; pers. obs.). However, all adults of the new species were collected only at the margin of several ponds and a lake with nearby forest. Other
<taxonomicName id="4C571DE1FFD68C27FE4F94A039021295" authorityName="Fabricius" authorityYear="1793" box="[503,598,943,969]" class="Insecta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
observed at the various sites were:
<taxonomicName id="4C571DE1FFD68C27FC4B94A03F301295" box="[1011,1124,943,969]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B923BA70FFD68C27FC4B94A03F6D1294" box="[1011,1081,943,968]" italics="true" pageId="5" pageNumber="400">Lestes</emphasis>
sp.
</taxonomicName>
,
<taxonomicName id="4C571DE1FFD68C27FBCA94A03A4812B1" authority="Williamson" authorityName="Williamson" class="Insecta" family="Coenagrionidae" genus="Acanthallagma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="caeruleum">
<emphasis id="B923BA70FFD68C27FBCA94A03EC81295" box="[1138,1436,943,969]" italics="true" pageId="5" pageNumber="400">Acanthallagma caeruleum</emphasis>
Williamson
</taxonomicName>
,
<taxonomicName id="4C571DE1FFD68C27FE9394DC395712B1" authority="Hagen" authorityName="Hagen" box="[299,515,979,1005]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="pulla">
<emphasis id="B923BA70FFD68C27FE9394DC3AFB12B1" box="[299,431,979,1005]" italics="true" pageId="5" pageNumber="400">Argia pulla</emphasis>
Hagen
</taxonomicName>
<emphasis id="B923BA70FFD68C27FDB694DC397012B0" box="[526,548,979,1004]" italics="true" pageId="5" pageNumber="400">in</emphasis>
Selys,
<taxonomicName id="4C571DE1FFD68C27FDC694DC3F5412B1" authority="Williamson" authorityName="Williamson" authorityYear="1915" box="[638,1024,979,1005]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="lamina">
<emphasis id="B923BA70FFD68C27FDC694DC382412B1" box="[638,880,979,1005]" italics="true" pageId="5" pageNumber="400">Epipleoneura lamina</emphasis>
Williamson
</taxonomicName>
,
<taxonomicName id="4C571DE1FFD68C27FBB694DC3E3A12B1" authority="Daigle" authorityName="Daigle" authorityYear="2003" box="[1038,1390,979,1005]" class="Insecta" family="Coenagrionidae" genus="Metaleptobasis" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="minteri">
<emphasis id="B923BA70FFD68C27FBB694DC3E4212B0" box="[1038,1302,979,1005]" italics="true" pageId="5" pageNumber="400">Metaleptobasis minteri</emphasis>
Daigle
</taxonomicName>
,
<taxonomicName id="4C571DE1FFD68C27FAC694DC3AF4154D" authority="von Ellenrieder" authorityName="von Ellenrieder" authorityYear="2013" class="Insecta" family="Coenagrionidae" genus="Metaleptobasis" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="spatula">
<emphasis id="B923BA70FFD68C27FAC694DC3BBD154D" italics="true" pageId="5" pageNumber="400">M. spatula</emphasis>
von Ellenrieder
</taxonomicName>
,
<taxonomicName id="4C571DE1FFD68C27FE1394F83983154D" authority="(Martin)" baseAuthorityName="Martin" box="[427,727,1015,1041]" class="Insecta" family="Coenagrionidae" genus="Telebasis" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="griffinii">
<emphasis id="B923BA70FFD68C27FE1394F8393B154D" box="[427,623,1015,1041]" italics="true" pageId="5" pageNumber="400">Telebasis griffinii</emphasis>
(Martin)
</taxonomicName>
,
<taxonomicName id="4C571DE1FFD68C27FD5B94F83F70154D" authority="Selys" authorityName="Selys" authorityYear="1860" box="[739,1060,1015,1041]" class="Insecta" family="Protoneuridae" genus="Neoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="rubriventris">
<emphasis id="B923BA70FFD68C27FD5B94F8388A154D" box="[739,990,1015,1041]" italics="true" pageId="5" pageNumber="400">Neoneura rubriventris</emphasis>
Selys
</taxonomicName>
,
<taxonomicName id="4C571DE1FFD68C27FB9794F83ECC154D" authority="(Burmeister)" baseAuthorityName="Burmeister" box="[1071,1432,1015,1041]" class="Insecta" family="Libellulidae" genus="Diastatops" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="pullata">
<emphasis id="B923BA70FFD68C27FB9794F83FAB154D" box="[1071,1279,1015,1041]" italics="true" pageId="5" pageNumber="400">Diastatops pullata</emphasis>
(Burmeister)
</taxonomicName>
,
<taxonomicName id="4C571DE1FFD68C27FF2F9314391B1569" authority="(Burmeister)" baseAuthorityName="Burmeister" box="[151,591,1051,1077]" class="Insecta" family="Libellulidae" genus="Erythemis" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="haematogastra">
<emphasis id="B923BA70FFD68C27FF2F93143AE31569" box="[151,439,1051,1077]" italics="true" pageId="5" pageNumber="400">Erythemis haematogastra</emphasis>
(Burmeister)
</taxonomicName>
,
<taxonomicName id="4C571DE1FFD68C27FDE0931438DE1569" authority="(Perty)" baseAuthorityName="Perty" box="[600,906,1051,1077]" class="Insecta" family="Libellulidae" genus="Nephepeltia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="phryne">
<emphasis id="B923BA70FFD68C27FDE0931438671569" box="[600,819,1051,1077]" italics="true" pageId="5" pageNumber="400">Nephepeltia phryne</emphasis>
(Perty)
</taxonomicName>
,
<taxonomicName id="4C571DE1FFD68C27FC2C93143FBE1569" authority="Borror" authorityName="Borror" authorityYear="1931" box="[916,1258,1051,1077]" class="Insecta" family="Libellulidae" genus="Oligoclada" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="monosticha">
<emphasis id="B923BA70FFD68C27FC2C93143FCC1569" box="[916,1176,1051,1077]" italics="true" pageId="5" pageNumber="400">Oligoclada monosticha</emphasis>
Borror
</taxonomicName>
,
<taxonomicName id="4C571DE1FFD68C27FB4D93143BB11505" authority="(Perty)" baseAuthorityName="Perty" class="Insecta" family="Libellulidae" genus="Perithemis" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="lais">
<emphasis id="B923BA70FFD68C27FB4D93143EC81569" box="[1269,1436,1051,1077]" italics="true" pageId="5" pageNumber="400">Perithemis lais</emphasis>
(Perty)
</taxonomicName>
and
<taxonomicName id="4C571DE1FFD68C27FEA3933039421505" authority="Hoffman." authorityName="Hoffman." box="[283,534,1087,1113]" class="Insecta" family="Libellulidae" genus="Perithemis" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="parzefalli">
<emphasis id="B923BA70FFD68C27FEA393303AF61505" box="[283,418,1087,1113]" italics="true" pageId="5" pageNumber="400">P. parzefalli</emphasis>
Hoffman.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>
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