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<document id="5A00FF4D6A4831EA10AA1D068E684540" ID-DOI="10.5281/zenodo.4509152" ID-ISSN="2345-7600" ID-Zenodo-Dep="4509152" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612548437016" checkinUser="carolina" docAuthor="ski, P. Jałoszy" docDate="2009" docId="03FEC075FF80C162FF3BFE49FF34FD51" docLanguage="en" docName="RafflesBZool.57.2.297-303.pdf" docOrigin="Raffles Bulletin of Zoology 57 (2)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Horaeomorphus fakfakensis ski 2009, new species" docType="treatment" docVersion="5" lastPageNumber="302" masterDocId="FFC7B80DFF84C167FFE6FFB7FFCAFFA4" masterDocTitle="Four New Species Of Horaeomorphus Schaufuss From The Oriental Region (Coleoptera: Staphylinidae, Scydmaeninae)" masterLastPageNumber="303" masterPageNumber="297" pageNumber="301" updateTime="1698867913614" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="0E94B55921333FEE5A55796B63E718CA">Four New Species Of Horaeomorphus Schaufuss From The Oriental Region (Coleoptera: Staphylinidae, Scydmaeninae)</mods:title>
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<mods:namePart id="58C4F8CA86E170C47C987A735182F994">ski, P. Jałoszy</mods:namePart>
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<mods:title id="369B05026B93D7C078F000FEDC461D86">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="CCC9AFD88E68CF342C21D46506CBF6BA">2009</mods:date>
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<mods:number id="C8D54462C404A7A7C43A1E21BBB778E1">2009-08-31</mods:number>
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<subSubSection id="C34D22E8FF80C163FF3BFE49FD67FDB2" box="[221,685,510,534]" pageId="4" pageNumber="301" type="nomenclature">
<paragraph id="8BE87163FF80C163FF3BFE49FD67FDB2" blockId="4.[221,685,510,565]" box="[221,685,510,534]" pageId="4" pageNumber="301">
<heading id="D0A0C60FFF80C163FF3BFE49FD67FDB2" bold="true" box="[221,685,510,534]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="301" reason="2">
<emphasis id="B923AD71FF80C163FF3BFE49FD67FDB2" bold="true" box="[221,685,510,534]" pageId="4" pageNumber="301">
<taxonomicName id="4C570AE0FF80C163FF3BFE49FDD2FDB2" ID-CoL="3MNJY" authority="ski, 2009" authorityName="ski" authorityYear="2009" box="[221,536,510,534]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="301" phylum="Arthropoda" rank="species" species="fakfakensis" status="sp. nov.">
<emphasis id="B923AD71FF80C163FF3BFE49FDD2FDB2" bold="true" box="[221,536,510,534]" italics="true" pageId="4" pageNumber="301">Horaeomorphus fakfakensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A210100AFF80C163FDCEFE49FD67FDB2" box="[552,685,510,534]" pageId="4" pageNumber="301" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34D22E8FF80C163FE88FDA9FDD6FD91" box="[366,540,542,565]" pageId="4" pageNumber="301" type="description">
<paragraph id="8BE87163FF80C163FE88FDA9FDD6FD91" blockId="4.[221,685,510,565]" box="[366,540,542,565]" pageId="4" pageNumber="301">
(
<figureCitation id="136C6DE6FF80C163FE90FDA9FE04FD91" box="[374,462,542,565]" captionStart="Fig" captionStartId="2.[134,169,1165,1186]" captionTargetBox="[134,1466,358,1140]" captionTargetId="figure-89@2.[130,1472,338,1154]" captionTargetPageId="2" captionText="Fig. 1. Dorsal habitus of holotype males: a, Horaeomorphus imitator, new species; b, Horaeomorphus minor, new species; c, Horaeomorphus fakfakensis, new species; d, Horaeomorphus samosirensis, new species (d). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4509154" httpUri="https://zenodo.org/record/4509154/files/figure.png" pageId="4" pageNumber="301">Figs. 1c</figureCitation>
,
<figureCitation id="136C6DE6FF80C163FE3BFDA9FDD9FD91" box="[477,531,542,565]" captionStart="Fig" captionStartId="3.[113,148,2061,2082]" captionTargetBox="[113,1440,1364,2029]" captionTargetId="figure-522@3.[80,1459,1346,2048]" captionTargetPageId="3" captionText="Fig. 3. Aedeagus in ventral (a, c) and lateral (b, d) views: a, b, Horaeomorphus fakfakensis, new species, c, d, Horaeomorphus samosirensis, new species. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4509158" httpUri="https://zenodo.org/record/4509158/files/figure.png" pageId="4" pageNumber="301">3a, b</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34D22E8FF80C163FF63FDEBFF14FD42" pageId="4" pageNumber="301" type="materials_examined">
<paragraph id="8BE87163FF80C163FF63FDEBFF14FD42" blockId="4.[133,772,604,742]" pageId="4" pageNumber="301">
Material examined.
<materialsCitation id="3B3F7B3EFF80C163FE9AFDEBFF14FD42" collectingDate="1996-08-13" collectionCode="IRIAN, JAYA, SMNS" country="Iceland" location="Male" pageId="4" pageNumber="301" specimenCount="2" typeStatus="holotype">
<emphasis id="B923AD71FF80C163FE9AFDEBFE22FDD5" bold="true" box="[380,488,604,625]" pageId="4" pageNumber="301">
<typeStatus id="54ECCFC1FF80C163FE9AFDEBFE29FDD5" box="[380,483,604,625]" pageId="4" pageNumber="301" type="holotype">Holotype</typeStatus>
.
</emphasis>
<location id="8E8827B8FF80C163FE15FDEAFDE0FDD6" LSID="urn:lsid:plazi:treatment:03FEC075FF80C162FF3BFE49FF34FD51:8E8827B8FF80C163FE15FDEAFDE0FDD6" box="[499,554,605,626]" country="Iceland" name="Male" pageId="4" pageNumber="301">Male</location>
, two labels “
<collectionCode id="ED46E9A6FF80C163FD5DFDEAFF09FD2A" pageId="4" pageNumber="301">IRIAN JAYA</collectionCode>
:Fakfak \ Kokas
<collectingCountry id="F34031F3FF80C163FE80FDCEFE5FFD2A" box="[358,405,633,654]" name="Iceland" pageId="4" pageNumber="301">Insel</collectingCountry>
Ogar \
<date id="FFE957A3FF80C163FE04FDCEFDAAFD2A" box="[482,608,633,654]" pageId="4" pageNumber="301" value="1996-08-13">
<collectingDate id="EFADAE4BFF80C163FE04FDCEFDAAFD2A" box="[482,608,633,654]" pageId="4" pageNumber="301" value="1996-08-13">13.VIII.1996</collectingDate>
</date>
\ leg. SCHÜLE/ STÜBEN” [pale blue, printed], “
<taxonomicName id="4C570AE0FF80C163FE5FFD21FD40FD0F" box="[441,650,662,683]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="301" phylum="Arthropoda" rank="genus">
<emphasis id="B923AD71FF80C163FE5FFD21FD40FD0F" box="[441,650,662,683]" italics="true" pageId="4" pageNumber="301">HORAEOMORPHUS</emphasis>
</taxonomicName>
\
<taxonomicName id="4C570AE0FF80C163FD7FFD21FCCEFD0F" authorityName="ski" authorityYear="2009" box="[665,772,662,683]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="301" phylum="Arthropoda" rank="species" species="fakfakensis">
<emphasis id="B923AD71FF80C163FD7FFD21FCCEFD0F" box="[665,772,662,683]" italics="true" pageId="4" pageNumber="301">fakfakensis</emphasis>
</taxonomicName>
m. \
<typeStatus id="54ECCFC1FF80C163FF53FD03FE8EFD6D" box="[181,324,692,713]" pageId="4" pageNumber="301" type="holotype">HOLOTYPUS</typeStatus>
\ det. P. JAŁOSZY SKI, 08” [red, printed] (
<collectionCode id="ED46E9A6FF80C163FF6AFD66FF18FD42" box="[140,210,721,742]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="4" pageNumber="301">SMNS</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34D22E8FF80C163FF63FCB8FBE0F871" pageId="4" pageNumber="301" type="diagnosis">
<paragraph id="8BE87163FF80C163FF63FCB8FF26FBA2" blockId="4.[133,773,783,1030]" pageId="4" pageNumber="301">
<emphasis id="B923AD71FF80C163FF63FCB8FEDEFC83" bold="true" box="[133,276,783,807]" italics="true" pageId="4" pageNumber="301">Diagnosis. </emphasis>
This moderately large
<taxonomicName id="4C570AE0FF80C163FDC7FCB8FD1EFC82" box="[545,724,783,806]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="301" phylum="Arthropoda" rank="genus">
<emphasis id="B923AD71FF80C163FDC7FCB8FD1EFC82" box="[545,724,783,806]" italics="true" pageId="4" pageNumber="301">Horaeomorphus</emphasis>
</taxonomicName>
can be distinguished from all other species of the genus on the basis of the following set of characters: pronotum very broad, disc nearly circular, with relatively small oval median part covered with large, deep and dense punctures while remaining surface bears small and very shallow punctures; base of pronotum with five pits, without groove; legs nonmodified.
</paragraph>
<paragraph id="8BE87163FF80C163FF63FB98FB45FD11" blockId="4.[133,772,1071,1158]" lastBlockId="4.[833,1472,190,693]" pageId="4" pageNumber="301">
<emphasis id="B923AD71FF80C163FF63FB98FF2BFBE3" bold="true" box="[133,225,1071,1095]" italics="true" pageId="4" pageNumber="301">Male. </emphasis>
Body (
<figureCitation id="136C6DE6FF80C163FEDBFB98FE5BFBE2" box="[317,401,1071,1094]" captionStart="Fig" captionStartId="2.[134,169,1165,1186]" captionTargetBox="[134,1466,358,1140]" captionTargetId="figure-89@2.[130,1472,338,1154]" captionTargetPageId="2" captionText="Fig. 1. Dorsal habitus of holotype males: a, Horaeomorphus imitator, new species; b, Horaeomorphus minor, new species; c, Horaeomorphus fakfakensis, new species; d, Horaeomorphus samosirensis, new species (d). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4509154" httpUri="https://zenodo.org/record/4509154/files/figure.png" pageId="4" pageNumber="301">Fig. 1c</figureCitation>
) moderately large (length
<quantity id="4CAFDC86FF80C163FD35FB98FF78FBC2" metricMagnitude="-3" metricUnit="m" metricValue="2.08" pageId="4" pageNumber="301" unit="mm" value="2.08">2.08 mm</quantity>
), pronotum and elytra very broad, moderately strongly convex, dark brown, vestiture slightly lighter than cuticle. Head broadest at large, strongly convex eyes, length
<quantity id="4CAFDC86FF80C163FCA7FF69FC62FF51" box="[833,936,222,245]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="4" pageNumber="301" unit="mm" value="0.33">0.33 mm</quantity>
, width
<quantity id="4CAFDC86FF80C163FBE6FF69FBA3FF51" box="[1024,1129,222,245]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="4" pageNumber="301" unit="mm" value="0.43">0.43 mm</quantity>
; tempora slightly shorter than eye in dorsal view, weakly and regularly rounded up to posterior 2/3, where they are strongly bent toward occipital constriction; vertex weakly convex, with pair of small and shallow pits located near posterior margins of supraantennal tubercles; frons weakly convex; supraantennal tubercles very distinct, strongly raised, well delimited from frons but indistinctly delimited from vertex. Punctures on vertex and frons very fine but sharply marked, very sparse, unevenly distributed, separated by spaces 46× as long as puncture diameters; setae moderately long, sparse, suberect to erect. Antennae moderately slender, shorter than half length of body, length
<quantity id="4CAFDC86FF80C163FC3EFDE9FB8BFDD1" box="[984,1089,606,629]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="301" unit="mm" value="0.9">0.90 mm</quantity>
; relative lengths of antennomeres (shortest antennomere VII as 1): 1.59; 1.36; 1.36; 1.36; 1.36; 1.27; 1; 1.36; 1.36; 1.36; 2.27.
</paragraph>
<paragraph id="8BE87163FF80C163FCA7FD69FA7BFB31" blockId="4.[833,1472,734,1173]" pageId="4" pageNumber="301">
Pronotum very broad and nearly circular, broadest between middle and anterior third, length
<quantity id="4CAFDC86FF80C163FB7FFD49FB33FCB1" box="[1177,1273,766,789]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="4" pageNumber="301" unit="mm" value="0.68">0.68 mm</quantity>
, width at base
<quantity id="4CAFDC86FF80C163FA77FD49FCA7FC91" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="4" pageNumber="301" unit="mm" value="0.43">0.43 mm</quantity>
, maximum width
<quantity id="4CAFDC86FF80C163FBCBFCA9FB45FC91" box="[1069,1167,798,821]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="4" pageNumber="301" unit="mm" value="0.65">0.65 mm</quantity>
; anterior and lateral margins rounded together; subbasal constriction barely marked; hind angles obtuse and blunt; posterior margin nearly straight; posterior collar very short, delimited from disc by transverse dorsal row of five pits; median pit moderately large, deep, two lateral pairs composed of smaller and shallower pits, external ones are shifted anteriorly. Disc with median oval area densely covered with large and deep, oval or slightly irregular in shape punctures separated by spaces 12× as long as puncture diameters; remaining surface of disc with similarly dense, but much smaller and very shallow punctures; vestiture moderately dense and long, suberect.
</paragraph>
<paragraph id="8BE87163FF80C163FCA7FB09FB54F9D1" blockId="4.[833,1472,1214,1653]" pageId="4" pageNumber="301">
Elytra oval, very broad and relatively short, as convex as pronotum, broadest near anterior fourth, length
<quantity id="4CAFDC86FF80C163FAB0FB69FA71FB51" box="[1366,1467,1246,1269]" metricMagnitude="-3" metricUnit="m" metricValue="1.08" pageId="4" pageNumber="301" unit="mm" value="1.08">1.08 mm</quantity>
, width
<quantity id="4CAFDC86FF80C163FC6CFB49FC38FAB1" box="[906,1010,1278,1301]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="4" pageNumber="301" unit="mm" value="0.73">0.73 mm</quantity>
, EI 1.48. Base of elytra arcuate, so that humeri are projecting anteriorly; humeral calli moderately distinct, delimited from adsutural region by short and deep, broad basal impressions; basal foveae indiscernible; apices of elytra separately rounded. Punctures in anterior 2/3 of elytra much more distinct than those on sides of pronotum, but much smaller than those on median part of pronotal disc, separated by spaces about equal to puncture diameters; punctures become gradually smaller and shallower toward apices and lateral margins of elytra. Vestiture similar to that on pronotum but slightly longer and slightly more suberect. Metathoracic wings not studied.
</paragraph>
<paragraph id="8BE87163FF80C163FCA7F929FBDFF911" blockId="4.[833,1045,1694,1717]" box="[833,1045,1694,1717]" pageId="4" pageNumber="301">Legs non-modified.</paragraph>
<paragraph id="8BE87163FF80C163FCA7F969FB78F831" blockId="4.[833,1472,1758,1941]" pageId="4" pageNumber="301">
Aedeagus (
<figureCitation id="136C6DE6FF80C163FC5CF969FBE4F951" box="[954,1070,1758,1781]" captionStart="Fig" captionStartId="3.[113,148,2061,2082]" captionTargetBox="[113,1440,1364,2029]" captionTargetId="figure-522@3.[80,1459,1346,2048]" captionTargetPageId="3" captionText="Fig. 3. Aedeagus in ventral (a, c) and lateral (b, d) views: a, b, Horaeomorphus fakfakensis, new species, c, d, Horaeomorphus samosirensis, new species. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4509158" httpUri="https://zenodo.org/record/4509158/files/figure.png" pageId="4" pageNumber="301">Figs. 3a, b</figureCitation>
)
<quantity id="4CAFDC86FF80C163FBD9F969FB55F951" box="[1087,1183,1758,1781]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="301" unit="mm" value="0.3">0.30 mm</quantity>
in length; relatively stout; median lobe with subtrapezoidal apical part distinctly projected in middle; internal sac relatively simple and lightly sclerotized, with small central complex; parameres slender, with apices reaching apex of median lobe, each with three very short apical setae.
</paragraph>
<paragraph id="8BE87163FF80C163FCA7F809FBE0F871" blockId="4.[833,1066,1982,2006]" box="[833,1066,1982,2006]" pageId="4" pageNumber="301">
<emphasis id="B923AD71FF80C163FCA7F809FC7AF872" bold="true" box="[833,944,1982,2006]" italics="true" pageId="4" pageNumber="301">Female. </emphasis>
Unknown.
</paragraph>
</subSubSection>
<caption id="DF2821EBFF80C163FF60F863FF78F7E4" ID-DOI="http://doi.org/10.5281/zenodo.4509160" ID-Zenodo-Dep="4509160" httpUri="https://zenodo.org/record/4509160/files/figure.png" pageId="4" pageNumber="301" startId="4.[134,171,2004,2025]" targetBox="[135,768,1281,1982]" targetPageId="4">
<paragraph id="8BE87163FF80C163FF60F863FF78F7E4" blockId="4.[134,772,2004,2112]" pageId="4" pageNumber="301">
Fig. 4. Hind trochanter and femur (a), and hind trochanter, femur and tibia (b).
<taxonomicName id="4C570AE0FF80C163FEBCF846FD9FF7A2" authority="ski, 2009" authorityName="ski" authorityYear="2009" box="[346,597,2033,2054]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="301" phylum="Arthropoda" rank="species" species="imitator" status="sp. nov.">
<emphasis id="B923AD71FF80C163FEBCF846FD9FF7A2" box="[346,597,2033,2054]" italics="true" pageId="4" pageNumber="301">Horaeomorphus imitator</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A210100AFF80C163FDB9F846FD13F7A2" box="[607,729,2033,2054]" pageId="4" pageNumber="301" rank="species">new species</taxonomicNameLabel>
(a),
<taxonomicName id="4C570AE0FF80C163FF60F7B9FE64F787" authority="ski, 2009" authorityName="ski" authorityYear="2009" box="[134,430,2062,2083]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="301" phylum="Arthropoda" rank="species" species="samosirensis" status="sp. nov.">
<emphasis id="B923AD71FF80C163FF60F7B9FE64F787" box="[134,430,2062,2083]" italics="true" pageId="4" pageNumber="301">Horaeomorphus samosirensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A210100AFF80C163FE5EF7B9FDFBF787" box="[440,561,2062,2083]" pageId="4" pageNumber="301" rank="species">new species</taxonomicNameLabel>
(b). Scale bar = 0.2 mm.
</paragraph>
</caption>
<subSubSection id="C34D22E8FF80C163FCA7F849FC2FF791" pageId="4" pageNumber="301" type="etymology">
<paragraph id="8BE87163FF80C163FCA7F849FC2FF791" blockId="4.[833,1471,2046,2101]" pageId="4" pageNumber="301">
<emphasis id="B923AD71FF80C163FCA7F849FC04F7B2" bold="true" box="[833,974,2046,2070]" italics="true" pageId="4" pageNumber="301">Etymology. </emphasis>
Locotypical, after the
<typeStatus id="54ECCFC1FF80C163FB5DF849FB23F7B1" box="[1211,1257,2046,2069]" pageId="4" pageNumber="301">type</typeStatus>
locality, Indonesian district Fakfak.
</paragraph>
</subSubSection>
<subSubSection id="C34D22E8FF81C162FF89FF09FD22FF71" box="[111,744,190,214]" pageId="5" pageNumber="302" type="distribution">
<paragraph id="8BE87163FF81C162FF89FF09FD22FF71" blockId="5.[111,744,190,214]" box="[111,744,190,214]" pageId="5" pageNumber="302">
<emphasis id="B923AD71FF81C162FF89FF09FEDAFF72" bold="true" box="[111,272,190,214]" italics="true" pageId="5" pageNumber="302">Distribution. </emphasis>
<collectingCountry id="F34031F3FF81C162FEFFFF09FE4BFF71" box="[281,385,190,213]" name="Indonesia" pageId="5" pageNumber="302">Indonesia</collectingCountry>
(New
<collectingCountry id="F34031F3FF81C162FE28FF09FDEBFF71" box="[462,545,190,213]" name="Guinea-Bissau" pageId="5" pageNumber="302">Guinea</collectingCountry>
): Irian Jaya Barat.
</paragraph>
</subSubSection>
<subSubSection id="C34D22E8FF81C162FF89FF49FF34FD51" pageId="5" pageNumber="302" type="discussion">
<paragraph id="8BE87163FF81C162FF89FF49FF34FD51" blockId="5.[111,750,254,757]" pageId="5" pageNumber="302">
<emphasis id="B923AD71FF81C162FF89FF49FF21FEB2" bold="true" box="[111,235,254,278]" italics="true" pageId="5" pageNumber="302">Remarks. </emphasis>
This species is most similar to
<taxonomicName id="4C570AE0FF81C162FDDBFF49FEC3FE91" authority="Franz, 1992" authorityName="Franz" authorityYear="1992" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="302" phylum="Arthropoda" rank="species" species="sarawakensis">
<emphasis id="B923AD71FF81C162FDDBFF49FD24FEB1" box="[573,750,254,277]" italics="true" pageId="5" pageNumber="302">H. sarawakensis</emphasis>
<bibRefCitation id="EFC60C92FF81C162FF89FEA9FEC3FE91" author="Franz, H" box="[111,265,286,309]" pageId="5" pageNumber="302" pagination="69 - 76" refId="ref4571" refString="Franz, H., 1992, Scydmaeniden aus Malaysia und Sulawesi (Coleoptera: Scydmaenidae). Koleopterologische Rundschau, 62: 69 - 76." type="journal article" year="1992">Franz, 1992</bibRefCitation>
</taxonomicName>
(from W
<collectingCountry id="F34031F3FF81C162FE71FEA9FDC0FE91" box="[407,522,286,309]" name="Malaysia" pageId="5" pageNumber="302">Malaysia</collectingCountry>
and Borneo),
<taxonomicName id="4C570AE0FF81C162FD37FEA9FE5BFEF1" authority="Franz, 1984" authorityName="Franz" authorityYear="1984" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="302" phylum="Arthropoda" rank="species" species="valdepunctatus">
<emphasis id="B923AD71FF81C162FD37FEA9FEDAFEF1" italics="true" pageId="5" pageNumber="302">H. valdepunctatus</emphasis>
<bibRefCitation id="EFC60C92FF81C162FEF2FE89FE5BFEF1" author="Franz, H" box="[276,401,318,341]" pageId="5" pageNumber="302" pagination="89 - 142" refId="ref4499" refString="Franz, H., 1984. Beitrag zur Kenntnis der Scydmaeniden von Sumatra und benachbarter Gebiete. Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, Mathematisch- Naturwissenschaftliche Klasse, Abt. I, 193: 89 - 142." type="journal article" year="1984">Franz, 1984</bibRefCitation>
</taxonomicName>
(W
<collectingCountry id="F34031F3FF81C162FE5DFE89FDEBFEF1" box="[443,545,318,341]" name="Malaysia" pageId="5" pageNumber="302">Malaysia</collectingCountry>
),
<taxonomicName id="4C570AE0FF81C162FDD7FE89FF32FED1" authority="Franz, 1992" authorityName="Franz" authorityYear="1992" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="302" phylum="Arthropoda" rank="species" species="punctatissimus">
<emphasis id="B923AD71FF81C162FDD7FE89FD24FEF1" box="[561,750,318,341]" italics="true" pageId="5" pageNumber="302">H. punctatissimus</emphasis>
<bibRefCitation id="EFC60C92FF81C162FF89FEE9FF32FED1" author="Franz, H" box="[111,248,350,373]" pageId="5" pageNumber="302" pagination="69 - 76" refId="ref4571" refString="Franz, H., 1992, Scydmaeniden aus Malaysia und Sulawesi (Coleoptera: Scydmaenidae). Koleopterologische Rundschau, 62: 69 - 76." type="journal article" year="1992">Franz, 1992</bibRefCitation>
</taxonomicName>
(Borneo, Sumatra and Siberut Is.), and
<taxonomicName id="4C570AE0FF81C162FD35FEE9FF32FE31" authorityName="ski" authorityYear="2009" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="302" phylum="Arthropoda" rank="species" species="samosirensis">
<emphasis id="B923AD71FF81C162FD35FEE9FF32FE31" italics="true" pageId="5" pageNumber="302">H. samosirensis</emphasis>
</taxonomicName>
described below. Together with
<taxonomicName id="4C570AE0FF81C162FDB7FEC9FD2DFE31" authorityName="ski" authorityYear="2009" box="[593,743,382,405]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="302" phylum="Arthropoda" rank="species" species="fakfakensis">
<emphasis id="B923AD71FF81C162FDB7FEC9FD2DFE31" box="[593,743,382,405]" italics="true" pageId="5" pageNumber="302">H. fakfakensis</emphasis>
</taxonomicName>
, these taxa share a similar body form, with very broad, nearly circular pronotum, and stout elytra.
<taxonomicName id="4C570AE0FF81C162FDD9FE09FF2CFE51" authorityName="ski" authorityYear="2009" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="302" phylum="Arthropoda" rank="species" species="fakfakensis">
<emphasis id="B923AD71FF81C162FDD9FE09FF2CFE51" italics="true" pageId="5" pageNumber="302">Horaeomorphus fakfakensis</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C570AE0FF81C162FDF4FE69FD0EFE51" authorityName="Franz" authorityYear="1992" box="[530,708,478,501]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="302" phylum="Arthropoda" rank="species" species="sarawakensis">
<emphasis id="B923AD71FF81C162FDF4FE69FD0EFE51" box="[530,708,478,501]" italics="true" pageId="5" pageNumber="302">H. sarawakensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C570AE0FF81C162FD32FE69FEC4FDB1" authorityName="Franz" authorityYear="1992" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="302" phylum="Arthropoda" rank="species" species="punctatissimus">
<emphasis id="B923AD71FF81C162FD32FE69FEC4FDB1" italics="true" pageId="5" pageNumber="302">H. punctatissimus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C570AE0FF81C162FEA1FE49FE3EFDB1" authorityName="ski" authorityYear="2009" box="[327,500,510,533]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="302" phylum="Arthropoda" rank="species" species="samosirensis">
<emphasis id="B923AD71FF81C162FEA1FE49FE3EFDB1" box="[327,500,510,533]" italics="true" pageId="5" pageNumber="302">H. samosirensis</emphasis>
</taxonomicName>
on the basis of nearly straight metatibiae, while those in the three species are strongly bent.
<taxonomicName id="4C570AE0FF81C162FEFFFD89FD4FFDF1" authorityName="Franz" authorityYear="1984" box="[281,645,574,597]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="302" phylum="Arthropoda" rank="species" species="valdepunctatus">
<emphasis id="B923AD71FF81C162FEFFFD89FD4FFDF1" box="[281,645,574,597]" italics="true" pageId="5" pageNumber="302">Horaeomorphus valdepunctatus</emphasis>
</taxonomicName>
, in turn, has only slightly curved metatibiae, but metatrochanters in males are strongly modified, forming a long projection; metatrochanters in
<taxonomicName id="4C570AE0FF81C162FEA0FD29FE2EFD11" authorityName="ski" authorityYear="2009" box="[326,484,670,693]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="302" phylum="Arthropoda" rank="species" species="fakfakensis">
<emphasis id="B923AD71FF81C162FEA0FD29FE2EFD11" box="[326,484,670,693]" italics="true" pageId="5" pageNumber="302">H. fakfakensis</emphasis>
</taxonomicName>
are non-modified. This species is the first member of the genus known to occur in New
<collectingCountry id="F34031F3FF81C162FF4CFD69FF31FD51" box="[170,251,734,757]" name="Guinea-Bissau" pageId="5" pageNumber="302">Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>