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<document id="BE0187F22B216342D7F805D27D4A44CA" ID-DOI="10.11646/zootaxa.4006.2.1" ID-GBIF-Dataset="78b1e800-beb2-470b-91a3-1f670551351a" ID-ISSN="1175-5326" ID-Zenodo-Dep="243797" ID-ZooBank="B154E526-0C08-4A92-868E-A36E45F2F1A1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461204948217" checkinUser="plazi" docAuthor="Li, Xuankun, Marshall, Stephen A. &amp; Yang, Ding" docDate="2015" docId="03ED4575FFE6FFB5759DF99CD80C77C0" docLanguage="en" docName="zt04006p246.pdf" docOrigin="Zootaxa 4006 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cothornobata viriata Enderlein 1922" docType="treatment" docVersion="8" lastPageNumber="244" masterDocId="FFD43D0DFFCFFF9E750AFFA8DB7A7238" masterDocTitle="A review of the Oriental species of Cothornobata Czerny (Diptera, Micropezidae, Eurybatinae)" masterLastPageNumber="246" masterPageNumber="201" pageNumber="242" updateTime="1698601325199" updateUser="plazi">
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<mods:title id="433EADD4119E74C6EA369C4315B2A126">A review of the Oriental species of Cothornobata Czerny (Diptera, Micropezidae, Eurybatinae)</mods:title>
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<mods:namePart id="9B2E432F00E35B50F1CF01F0DE864F96">Li, Xuankun</mods:namePart>
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<mods:namePart id="65075921959E4395DF7C79783BB6E3AE">Marshall, Stephen A.</mods:namePart>
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<mods:namePart id="70012328B9F4BAD0053501871ED43514">Yang, Ding</mods:namePart>
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<mods:date id="FC3F8D6C6728EC45DBA10E7F98652082">2015</mods:date>
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<treatment id="03ED4575FFE6FFB5759DF99CD80C77C0" ID-DOI="http://doi.org/10.5281/zenodo.6094547" ID-GBIF-Taxon="119630154" ID-Zenodo-Dep="6094547" LSID="urn:lsid:plazi:treatment:03ED4575FFE6FFB5759DF99CD80C77C0" httpUri="http://treatment.plazi.org/id/03ED4575FFE6FFB5759DF99CD80C77C0" lastPageId="43" lastPageNumber="244" pageId="41" pageNumber="242">
<subSubSection id="C35EA7E8FFE6FFB7759DF99CD9E57476" box="[151,671,1587,1614]" pageId="41" pageNumber="242" type="nomenclature">
<paragraph id="8BFBF463FFE6FFB7759DF99CD9E57476" blockId="41.[151,671,1587,1648]" box="[151,671,1587,1614]" pageId="41" pageNumber="242">
<heading id="D0B3430FFFE6FFB7759DF99CD9E57476" bold="true" box="[151,671,1587,1614]" fontSize="11" level="1" pageId="41" pageNumber="242" reason="1">
<emphasis id="B9302871FFE6FFB7759DF99CD9E57476" bold="true" box="[151,671,1587,1614]" pageId="41" pageNumber="242">
19.
<taxonomicName id="4C448FE0FFE6FFB775C9F99BD9E57476" ID-CoL="YXG8" authority="Enderlein, 1922" authorityName="Enderlein" authorityYear="1922" box="[195,671,1587,1614]" class="Insecta" family="Micropezidae" genus="Cothornobata" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="242" phylum="Arthropoda" rank="species" species="viriata">
<emphasis id="B9302871FFE6FFB775C9F99BDAC57475" bold="true" box="[195,447,1587,1613]" italics="true" pageId="41" pageNumber="242">Cothornobata viriata</emphasis>
(
<bibRefCitation id="EFD58992FFE6FFB774C4F99CD9ED7476" author="Enderlein" box="[462,663,1588,1614]" pageId="41" pageNumber="242" refString="Enderlein, G. (1922) Klassifikation der Micropeziden. Archiv fuer Naturgeschichte Berlin Abt A, 88 (5), 140 - 299." type="journal article" year="1922">Enderlein, 1922</bibRefCitation>
)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35EA7E8FFE6FFB7759DF9FFDB837457" box="[151,249,1623,1648]" pageId="41" pageNumber="242" type="description">
<paragraph id="8BFBF463FFE6FFB7759DF9FFDB837457" blockId="41.[151,671,1587,1648]" box="[151,249,1623,1648]" pageId="41" pageNumber="242">
(
<figureCitation id="137FE8E6FFE6FFB77595F9FFDB8B7457" box="[159,241,1623,1648]" captionStart="FIGURE 24" captionStartId="43.[151,250,1077,1099]" captionTargetBox="[151,1434,193,1055]" captionTargetId="figure@43.[151,1435,193,1055]" captionTargetPageId="43" captionText="FIGURE 24. Cothornobata viriata Enderlein, type male, left lateral." httpUri="https://zenodo.org/record/243821/files/figure.png" pageId="41" pageNumber="242">Fig. 24</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35EA7E8FFE6FFB7759DF908D9CD74CA" pageId="41" pageNumber="242" type="reference_group">
<paragraph id="8BFBF463FFE6FFB7759DF908D9BA74EB" blockId="41.[151,908,1695,1778]" pageId="41" pageNumber="242">
<treatmentCitationGroup id="AB54D34DFFE6FFB7759DF908D9BA74EB" pageId="41" pageNumber="242">
<taxonomicName id="4C448FE0FFE6FFB7759DF908D927748D" ID-CoL="8C8N3" authority="Enderlein, 1922: 174" authorityName="Enderlein" authorityPageNumber="174" authorityYear="1922" box="[151,605,1695,1717]" class="Insecta" family="Micropezidae" genus="Grammicomyia" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="242" phylum="Arthropoda" rank="species" species="viriata">
<emphasis id="B9302871FFE6FFB7759DF908DA05748D" box="[151,383,1696,1717]" italics="true" pageId="41" pageNumber="242">Grammicomyia viriata</emphasis>
<treatmentCitation id="0AE5D272FFE6FFB7748EF937D927748D" author="Enderlein" box="[388,605,1695,1717]" page="174" pageId="41" pageNumber="242" year="1922">
<bibRefCitation id="EFD58992FFE6FFB7748EF937D950748D" author="Enderlein" box="[388,554,1695,1717]" pageId="41" pageNumber="242" refString="Enderlein, G. (1922) Klassifikation der Micropeziden. Archiv fuer Naturgeschichte Berlin Abt A, 88 (5), 140 - 299." type="journal article" year="1922">Enderlein, 1922</bibRefCitation>
: 174
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="54FF4AC1FFE6FFB7776CF908D9E2748D" box="[614,664,1696,1717]" pageId="41" pageNumber="242">Type</typeStatus>
locality: Deli, Sumatra.
<taxonomicName id="4C448FE0FFE6FFB7759DF915DA8874EB" authority="Enderlein" authorityName="Enderlein" box="[151,498,1725,1748]" class="Insecta" family="Micropezidae" genus="Trepidarioides" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="242" phylum="Arthropoda" rank="species" species="viriata">
<emphasis id="B9302871FFE6FFB7759DF915DA0F74EC" box="[151,373,1725,1748]" italics="true" pageId="41" pageNumber="242">Trepidarioides viriata</emphasis>
(Enderlein)
</taxonomicName>
:
<treatmentCitation id="0AE5D272FFE6FFB774F1F915D9C174EB" author="Hennig" box="[507,699,1725,1747]" page="307" pageId="41" pageNumber="242" year="1935">
<bibRefCitation id="EFD58992FFE6FFB774F1F915D9F274EB" author="Hennig" box="[507,648,1725,1747]" pageId="41" pageNumber="242" refString="Hennig, W. (1935) Revision der Tyliden (Dipt., Acalypt.). II. Teil: Die ausseramerikanischen Taeniapterinae, die Trepidariinae und Tylinae. Allgemeines uber die Tyliden. Zugleich ein Beitrag zu den Ergebnissen der Sunda-expedition Rensch, 1927 [part]. Konowia, 14, 192 - 216, 289 - 310." type="journal article" year="1935">Hennig, 1935</bibRefCitation>
: 307
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFBF463FFE6FFB7759DF974D9CD74CA" blockId="41.[151,908,1695,1778]" box="[151,695,1756,1778]" pageId="41" pageNumber="242">
<treatmentCitationGroup id="AB54D34DFFE6FFB7759DF974D9CD74CA" box="[151,695,1756,1778]" pageId="41" pageNumber="242">
<taxonomicName id="4C448FE0FFE6FFB7759DF974DA9674CA" ID-CoL="YXG8" authority="Enderlein" authorityName="Enderlein" box="[151,492,1756,1778]" class="Insecta" family="Micropezidae" genus="Cothornobata" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="242" phylum="Arthropoda" rank="species" species="viriata">
<emphasis id="B9302871FFE6FFB7759DF974DA1574CA" box="[151,367,1756,1778]" italics="true" pageId="41" pageNumber="242">Cothornobata viriata</emphasis>
(Enderlein)
</taxonomicName>
:
<treatmentCitation id="0AE5D272FFE6FFB774FFF974D9C974CA" author="Steyskal" box="[501,691,1756,1778]" page="13" pageId="41" pageNumber="242" year="1977">
<bibRefCitation id="EFD58992FFE6FFB774FFF974D9F774CA" author="Steyskal" box="[501,653,1756,1778]" pageId="41" pageNumber="242" refString="Steyskal, G. C. (1977) Family Micropezidae. In: Delfinado, M. D. &amp; Hardy, D. E. (Eds.), A catalog of the Diptera of the Oriental region, 3, pp. 13 - 20. [The University Press of Hawaii]" type="journal article" year="1977">Steyskal, 1977</bibRefCitation>
: 13
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35EA7E8FFE6FFB7759DF8B7D9BA7500" box="[151,704,1823,1848]" pageId="41" pageNumber="242" type="diagnosis">
<paragraph id="8BFBF463FFE6FFB7759DF8B7D9BA7500" blockId="41.[151,1437,1823,2028]" box="[151,704,1823,1848]" pageId="41" pageNumber="242">
<emphasis id="B9302871FFE6FFB7759DF8B7DA707500" bold="true" box="[151,266,1823,1848]" pageId="41" pageNumber="242">Diagnosis</emphasis>
(From photo and original description).
</paragraph>
</subSubSection>
<subSubSection id="C35EA7E8FFE6FFB575CDF8EBDBAC76E0" lastPageId="43" lastPageNumber="244" pageId="41" pageNumber="242" type="description">
<paragraph id="8BFBF463FFE6FFB575CDF8EBDBAC76E0" blockId="41.[151,1437,1823,2028]" lastBlockId="43.[151,1437,1138,1528]" lastPageId="43" lastPageNumber="244" pageId="41" pageNumber="242">
<emphasis id="B9302871FFE6FFB775CDF8EBDAE57564" bold="true" box="[199,415,1859,1884]" pageId="41" pageNumber="242">Male and Female.</emphasis>
Head brownish. Head with 1 frontal seta and 1 orbital seta. Antenna dark yellow; pedicel with 1 ventral apical seta nearly half length of first flagellomere; first flagellomere 1.7X as long as broad; arista brown and bare, nearly 3.6X length of first flagellomere. Proboscis brownish-yellow. Thorax dull orange. Scutellum slightly convex, angled dorsally. Legs mostly yellow, with coxae brownish-yellow; fore femur brownish-yellow on apical half; fore tibia pale yellow. Wing clear, with a light brown apical band and a light brown middle band. Bm-cu distal to CuA2, A1+CuA2
<date id="FFFAD2A3FFE4FFB577AFFBDAD9B276B2" box="[677,712,1138,1162]" pageId="43" pageNumber="244">3X</date>
as long as CuA2; distance between apex of R2+3 and R4+5 half as long as M1 beyond dm-cu. Halter pale yellow. Abdomen dark brown. Male genital fork yellowish brown, relatively short.
</paragraph>
</subSubSection>
<caption id="DF3BA4EBFFE5FFB4759DF830DA2175D3" httpUri="https://zenodo.org/record/243820/files/figure.png" pageId="42" pageNumber="243" targetBox="[181,1401,220,1895]" targetPageId="42">
<paragraph id="8BFBF463FFE5FFB4759DF830DA2175D3" blockId="42.[151,1437,1944,2027]" pageId="42" pageNumber="243">
<emphasis id="B9302871FFE5FFB4759DF830DA587595" bold="true" box="[151,290,1944,1966]" pageId="42" pageNumber="243">FIGURE 23.</emphasis>
<taxonomicName id="4C448FE0FFE5FFB47420F830D9397596" box="[298,579,1944,1966]" class="Insecta" family="Micropezidae" genus="Cothornobata" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="243" phylum="Arthropoda" rank="species" species="vietnamensis" status="sp. nov.">
<emphasis id="B9302871FFE5FFB47420F830D9397596" box="[298,579,1944,1966]" italics="true" pageId="42" pageNumber="243">Cothornobata vietnamensis</emphasis>
</taxonomicName>
<emphasis id="B9302871FFE5FFB47746F830D9E57595" bold="true" box="[588,671,1944,1965]" pageId="42" pageNumber="243">
<taxonomicNameLabel id="A203950AFFE5FFB47746F830D9E57595" box="[588,671,1944,1965]" pageId="42" pageNumber="243" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male.
<emphasis id="B9302871FFE5FFB477E4F830D87D7595" bold="true" box="[750,775,1944,1965]" pageId="42" pageNumber="243">A,</emphasis>
left lateral;
<emphasis id="B9302871FFE5FFB47683F830D8DA7595" bold="true" box="[905,928,1944,1965]" pageId="42" pageNumber="243">B,</emphasis>
head;
<emphasis id="B9302871FFE5FFB476E2F830DF787596" bold="true" box="[1000,1026,1944,1966]" pageId="42" pageNumber="243">C,</emphasis>
wing;
<emphasis id="B9302871FFE5FFB47146F830DF1E7595" bold="true" box="[1100,1124,1944,1965]" pageId="42" pageNumber="243">D,</emphasis>
genital fork;
<emphasis id="B9302871FFE5FFB471FEF830DE717595" bold="true" box="[1268,1291,1944,1965]" pageId="42" pageNumber="243">E,</emphasis>
genitalia, left lateral;
<emphasis id="B9302871FFE5FFB475E9F81FDB8E75F4" bold="true" box="[227,244,1975,1996]" pageId="42" pageNumber="243">F,</emphasis>
genitalia, posterior;
<emphasis id="B9302871FFE5FFB474C2F81EDAA675F4" bold="true" box="[456,476,1974,1996]" pageId="42" pageNumber="243">G,</emphasis>
aedeagus and associated structures, left lateral;
<emphasis id="B9302871FFE5FFB476CEF81FD8A475F4" bold="true" box="[964,990,1975,1996]" pageId="42" pageNumber="243">H,</emphasis>
aedeagus and associated structures, ventral. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="DF3BA4EBFFE4FFB5759DFB9DD82B7672" box="[151,849,1076,1099]" httpUri="https://zenodo.org/record/243821/files/figure.png" pageId="43" pageNumber="244" targetBox="[151,1434,193,1055]" targetPageId="43">
<paragraph id="8BFBF463FFE4FFB5759DFB9DD82B7672" blockId="43.[151,849,1076,1099]" box="[151,849,1076,1099]" pageId="43" pageNumber="244">
<emphasis id="B9302871FFE4FFB5759DFB9DDA5A7672" bold="true" box="[151,288,1077,1099]" pageId="43" pageNumber="244">FIGURE 24.</emphasis>
<taxonomicName id="4C448FE0FFE4FFB5742FFB9CD9137672" authority="Enderlein" authorityName="Enderlein" box="[293,617,1076,1099]" class="Insecta" family="Micropezidae" genus="Cothornobata" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="244" phylum="Arthropoda" rank="species" species="viriata">
<emphasis id="B9302871FFE4FFB5742FFB9CDA847673" box="[293,510,1076,1099]" italics="true" pageId="43" pageNumber="244">Cothornobata viriata</emphasis>
Enderlein
</taxonomicName>
, type male, left lateral.
</paragraph>
</caption>
<subSubSection id="C35EA7E8FFE4FFB575CDFB4BDAD8777C" pageId="43" pageNumber="244" type="materials_examined">
<paragraph id="8BFBF463FFE4FFB575CDFB4BDAD8777C" blockId="43.[151,1437,1138,1528]" pageId="43" pageNumber="244">
<emphasis id="B9302871FFE4FFB575CDFB4BDACA76C4" bold="true" box="[199,432,1251,1276]" pageId="43" pageNumber="244">Material examined.</emphasis>
<typeStatus id="54FF4AC1FFE4FFB574BCFB4CD92176C4" box="[438,603,1252,1276]" pageId="43" pageNumber="244" type="lectotype">LECTOTYPE</typeStatus>
: SUMATRA, OberLangkat, Deli, 1894. M.Ude S. (♂,
<collectionCode id="ED556CA6FFE4FFB571CDFB4CDE6776C4" box="[1223,1309,1252,1276]" pageId="43" pageNumber="244">MNBG</collectionCode>
) (Male and fenale cotypes were examined and the male
<typeStatus id="54FF4AC1FFE4FFB5779CFAA0D9BC7718" box="[662,710,1288,1312]" pageId="43" pageNumber="244">type</typeStatus>
photographed by the second author in 2002). The male is here designated as
<typeStatus id="54FF4AC1FFE4FFB5743EFA83DAE5777C" box="[308,415,1323,1348]" pageId="43" pageNumber="244" type="lectotype">lectotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C35EA7E8FFE4FFB575CDFAE7DA1277B4" pageId="43" pageNumber="244" type="distribution">
<paragraph id="8BFBF463FFE4FFB575CDFAE7DA1277B4" blockId="43.[151,1437,1138,1528]" pageId="43" pageNumber="244">
<emphasis id="B9302871FFE4FFB575CDFAE7DAA57750" bold="true" box="[199,479,1359,1384]" pageId="43" pageNumber="244">Distribution. Oriental:</emphasis>
Sumatra. Previous records from New
<collectingCountry id="F353B4F3FFE4FFB576CEFAE7DF6F7750" box="[964,1045,1359,1384]" name="Guinea-Bissau" pageId="43" pageNumber="244">Guinea</collectingCountry>
(
<bibRefCitation id="EFD58992FFE4FFB57127FAE7DF9B7750" author="Evenhuis" box="[1069,1249,1359,1384]" pageId="43" pageNumber="244" refString="Evenhuis, N. L. (1989) 54. Family Micropezidae. In: Evenhuis, N. L. (Ed.), Catalog of the Diptera of the Australasian and Oceanian Regions. Bishop Museum Special Publication, 86, pp. 464 - 467." type="journal article" year="1989">Evenhuis 1989</bibRefCitation>
) were in error (
<bibRefCitation id="EFD58992FFE4FFB57594FADBDA2677B4" author="McAlpine" box="[158,348,1395,1420]" pageId="43" pageNumber="244" refString="McAlpine, D. K. (1998) Review of the Australian stilt flies (Diptera: Micropezidae) with a phylogenetic analysis of the family. Invertebrate Taxonomy, 12 (1), 55 - 134. http: // dx. doi. org / 10.1071 / IT 96018" type="journal article" year="1998">McAlpine, 1998</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35EA7E8FFE4FFB575CDFA3FD80C77C0" pageId="43" pageNumber="244" type="discussion">
<paragraph id="8BFBF463FFE4FFB575CDFA3FD80C77C0" blockId="43.[151,1437,1138,1528]" pageId="43" pageNumber="244">
<emphasis id="B9302871FFE4FFB575CDFA3FDA417788" bold="true" box="[199,315,1431,1456]" pageId="43" pageNumber="244">Remarks.</emphasis>
This species differs from all other Oriental
<taxonomicName id="4C448FE0FFE4FFB57622FA3FD8BF7788" box="[808,965,1431,1456]" class="Insecta" family="Micropezidae" genus="Cothornobata" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="244" phylum="Arthropoda" rank="genus">
<emphasis id="B9302871FFE4FFB57622FA3FD8BF7788" box="[808,965,1431,1456]" italics="true" pageId="43" pageNumber="244">Cothornobata</emphasis>
</taxonomicName>
in its yellow legs and pale reddish-brown thorax. The
<typeStatus id="54FF4AC1FFE4FFB57429FA13DAF377EC" box="[291,393,1467,1492]" pageId="43" pageNumber="244" type="lectotype">lectotype</typeStatus>
was not dissected as it is the only known male. Relationships of this species should be reevaluated when additional males become available for dissection.
</paragraph>
</subSubSection>
</treatment>
</document>