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<document id="CC723F52845C6E8A1BA0B8C8E5AD22C6" ID-CLB-Dataset="29885" ID-DOI="10.11646/zootaxa.4434.1.2" ID-GBIF-Dataset="876d1781-7086-46ee-b7d6-94753804c4b2" ID-ISSN="1175-5326" ID-Zenodo-Dep="1290936" ID-ZooBank="BBD0A5B9-B363-4802-8F2B-78C0209C08D5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1529082724990" checkinUser="plazi" docAuthor="Grados, Juan" docDate="2018" docId="03CA35558876D17051A423C4FE68F873" docLanguage="en" docName="zootaxa.4434.1.2.pdf" docOrigin="Zootaxa 4434 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paranerita maculata subsp. sandeepani Grados 2018, ssp. nov." docType="treatment" docVersion="7" lastPageNumber="42" masterDocId="FFF34D2D887DD17D51332630FFB1FFB3" masterDocTitle="Four new species and one new subspecies of Arctiinae (Lepidoptera: Erebidae) from the Tambopata river, Madre de Dios, Peru" masterLastPageNumber="48" masterPageNumber="29" pageNumber="40" updateTime="1698549631496" updateUser="ExternalLinkService">
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<mods:title id="90A450EBC55BBAD8B0EB555144E1594A">Four new species and one new subspecies of Arctiinae (Lepidoptera: Erebidae) from the Tambopata river, Madre de Dios, Peru</mods:title>
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<mods:namePart id="6ECEF11D267DA45E62A037F7C1C4FE99">Grados, Juan</mods:namePart>
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<mods:date id="3CCCA324C0AEF45835B60A4865CE014F">2018</mods:date>
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<mods:number id="6018182A4DE575CAD97D5EF95DACD64F">2018-06-14</mods:number>
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<treatment id="03CA35558876D17051A423C4FE68F873" ID-DOI="http://doi.org/10.5281/zenodo.6485738" ID-GBIF-Taxon="145966337" ID-Zenodo-Dep="6485738" LSID="urn:lsid:plazi:treatment:03CA35558876D17051A423C4FE68F873" httpUri="http://treatment.plazi.org/id/03CA35558876D17051A423C4FE68F873" lastPageId="13" lastPageNumber="42" pageId="11" pageNumber="40">
<subSubSection id="C379D7C88876D17651A423C4FE9FF99C" pageId="11" pageNumber="40" type="nomenclature">
<paragraph id="8BDC84438876D17651A423C4FD21F9BD" blockId="11.[151,656,1524,1584]" box="[151,656,1524,1550]" pageId="11" pageNumber="40">
<heading id="D094332F8876D17651A423C4FD21F9BD" bold="true" box="[151,656,1524,1550]" fontSize="11" level="1" pageId="11" pageNumber="40" reason="1">
<emphasis id="B91758518876D17651A423C4FD21F9BD" bold="true" box="[151,656,1524,1550]" pageId="11" pageNumber="40">
<taxonomicName id="4C63FFC08876D17651A423C4FD91F9BD" authority="Grados, 2018" authorityName="Grados" authorityYear="2018" box="[151,544,1524,1550]" class="Insecta" family="Arctiidae" genus="Paranerita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="40" phylum="Arthropoda" rank="subSpecies" species="maculata" status="subsp. nov." subSpecies="sandeepani">
<emphasis id="B91758518876D17651A423C4FD91F9BD" bold="true" box="[151,544,1524,1550]" italics="true" pageId="11" pageNumber="40">Paranerita maculata sandeepani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A224E52A8876D176531423C4FD21F9BD" box="[551,656,1524,1550]" pageId="11" pageNumber="40" rank="subSpecies">
<taxonomicName id="4C63FFC08876D176531423C4FD3AF9BD" box="[551,651,1524,1550]" pageId="11" pageNumber="40" rank="subSpecies" subSpecies="nov">ssp. nov</taxonomicName>
.
</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDC84438876D17651A42027FE9FF99C" blockId="11.[151,656,1524,1584]" box="[151,302,1559,1584]" pageId="11" pageNumber="40">
(
<figureCitation id="135898C68876D17651AC2027FE97F99C" box="[159,294,1559,1584]" captionStart-0="FIGURES 3134" captionStart-1="FIGURES 3538" captionStartId-0="13.[151,264,742,765]" captionStartId-1="13.[151,264,1352,1375]" captionTargetBox-0="[340,1247,195,721]" captionTargetBox-1="[359,1247,851,1315]" captionTargetId-0="figure@13.[340,1247,195,721]" captionTargetId-1="figure@13.[340,1247,845,1331]" captionTargetPageId-0="13" captionTargetPageId-1="13" captionText-0="FIGURES 3134. Paranerita maculata sandeepani Grados, new subespecies. 31. Holotype male, dorsal. 32. Holotype male, ventral. 33. Paratype female, dorsal (Albergue Refugio Amazonas). 34. Paratype female, ventral. Scales=5 mm." captionText-1="FIGURES 3538. Genitalia of Paranerita maculata sandeepani Grados, new species. (Genitalia # JGA762). 35. Dorsal view. 36. Ventral view. 37. Lateral view. 38. Aedeagus. Scales= 1 mm." httpUri-0="https://zenodo.org/record/1290952/files/figure.png" httpUri-1="https://zenodo.org/record/1290954/files/figure.png" pageId="11" pageNumber="40">Figs. 3138</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C379D7C88876D17651A4206FFD75F957" pageId="11" pageNumber="40" type="materials_examined">
<paragraph id="8BDC84438876D17651A4206FFD75F957" blockId="11.[151,1437,1631,2016]" pageId="11" pageNumber="40">
<materialsCitation id="3B0B8E1E8876D17651A4206FFD13F92F" ID-GBIF-Occurrence="1849198248" collectingDate="2016-10-22" collectionCode="PERU" collectorName="J. Grados &amp; D. Couceiro" country="Peru" elevation="231" latitude="-12.875" location="Madre de Dios" longLatPrecision="21" longitude="-69.40972" pageId="11" pageNumber="40" specimenCount="1" stateProvince="Madre de Dios" typeStatus="holotype">
<emphasis id="B91758518876D17651A4206FFEF9F9CB" bold="true" box="[151,328,1631,1656]" pageId="11" pageNumber="40">
<typeStatus id="54D83AE18876D17651A4206FFEB5F9CB" box="[151,260,1631,1656]" pageId="11" pageNumber="40" type="holotype">Holotype</typeStatus>
male
</emphasis>
(
<figureCitation id="135898C68876D1765069206FFE59F9C4" box="[346,488,1631,1656]" captionStart="FIGURES 3134" captionStartId="13.[151,264,742,765]" captionTargetBox="[340,1247,195,721]" captionTargetId="figure@13.[340,1247,195,721]" captionTargetPageId="13" captionText="FIGURES 3134. Paranerita maculata sandeepani Grados, new subespecies. 31. Holotype male, dorsal. 32. Holotype male, ventral. 33. Paratype female, dorsal (Albergue Refugio Amazonas). 34. Paratype female, ventral. Scales=5 mm." httpUri="https://zenodo.org/record/1290952/files/figure.png" pageId="11" pageNumber="40">Figs. 3132</figureCitation>
):
<emphasis id="B91758518876D17650CD206FFCA6F9CB" bold="true" box="[510,791,1631,1656]" pageId="11" pageNumber="40">
<collectionCode id="ED721C868876D17650CD206FFDFDF9CB" box="[510,588,1631,1656]" country="Italy" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14398" name="Universit<69> di Perugia" pageId="11" pageNumber="40">
<collectingCountry id="F374C4D38876D17650CD206FFDFDF9CB" box="[510,588,1631,1656]" name="Peru" pageId="11" pageNumber="40">PERU</collectingCountry>
</collectionCode>
,
<collectingRegion id="49A74AA18876D1765368206FFCA2F9CB" box="[603,787,1631,1656]" country="Peru" name="Madre de Dios" pageId="11" pageNumber="40">Madre de Dios</collectingRegion>
,
</emphasis>
Albergue Refugio
<collectingRegion id="49A74AA18876D17652CB2050FBC0F9CB" box="[1016,1137,1632,1656]" country="Peru" name="Amazonas" pageId="11" pageNumber="40">Amazonas</collectingRegion>
,
<geoCoordinate id="EE57E2848876D17655B3206FFAB1F9CB" box="[1152,1280,1631,1656]" direction="south" orientation="latitude" pageId="11" pageNumber="40" precision="15" value="-12.875">12°52'30''S</geoCoordinate>
,
<geoCoordinate id="EE57E2848876D176543C206FFA26F9CB" box="[1295,1431,1631,1656]" direction="west" orientation="longitude" pageId="11" pageNumber="40" precision="15" value="-69.40972">69°24'35''W</geoCoordinate>
,
<elevation id="004E63708876D17651A420B4FF68F92F" box="[151,217,1668,1692]" metricMagnitude="2" metricUnit="m" metricValue="2.31" pageId="11" pageNumber="40" unit="m" value="231.0">
<quantity id="4C9B29A68876D17651A420B4FF68F92F" box="[151,217,1668,1692]" metricMagnitude="2" metricUnit="m" metricValue="2.31" pageId="11" pageNumber="40" unit="m" value="231.0">231m</quantity>
</elevation>
,
<collectingDate id="EF995B6B8876D17651DA20B3FEE8F92F" box="[233,345,1667,1692]" pageId="11" pageNumber="40" value="2016-10-22">
<date id="FFDDA2838876D17651DA20B3FEE8F92F" box="[233,345,1667,1692]" pageId="11" pageNumber="40" value="2016-10-22">22.x.2016</date>
</collectingDate>
(
<collectorName id="2696E1958876D176505E20B4FE6AF92F" box="[365,475,1667,1692]" pageId="11" pageNumber="40">J. Grados</collectorName>
&amp;
<collectorName id="2696E1958876D176533420B4FD2BF92F" box="[519,666,1667,1692]" pageId="11" pageNumber="40">D. Couceiro</collectorName>
)
</materialsCitation>
.
<materialsCitation id="3B0B8E1E8876D176538320B4FACDF92F" ID-GBIF-Occurrence="1849198250" box="[688,1404,1667,1692]" collectionCode="PERU" country="Peru" location="Madre de Dios" pageId="11" pageNumber="40" specimenCount="6" specimenCount-female="1" specimenCount-male="2" stateProvince="Madre de Dios" typeStatus="paratype">
<specimenCount id="9D654FCA8876D176538320B4FC85F92F" box="[688,820,1668,1692]" pageId="11" pageNumber="40" type="generic" typeStatus="paratypes">3 paratypes</specimenCount>
(
<specimenCount id="9D654FCA8876D176527B20B4FC12F92F" box="[840,931,1667,1692]" pageId="11" pageNumber="40" type="male">2 males</specimenCount>
and
<specimenCount id="9D654FCA8876D17652D120B4FBFBF92F" box="[994,1098,1667,1692]" pageId="11" pageNumber="40" type="female">1 female</specimenCount>
):
<emphasis id="B91758518876D176555520B3FACDF92F" bold="true" box="[1126,1404,1667,1692]" pageId="11" pageNumber="40">
<collectionCode id="ED721C868876D176555520B3FB05F92F" box="[1126,1204,1667,1692]" country="Italy" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14398" name="Universit<69> di Perugia" pageId="11" pageNumber="40">
<collectingCountry id="F374C4D38876D176555520B3FB05F92F" box="[1126,1204,1667,1692]" name="Peru" pageId="11" pageNumber="40">PERU</collectingCountry>
</collectionCode>
,
<collectingRegion id="49A74AA18876D17655F720B3FACDF92F" box="[1220,1404,1667,1692]" country="Peru" name="Madre de Dios" pageId="11" pageNumber="40">Madre de Dios</collectingRegion>
</emphasis>
</materialsCitation>
,
<materialsCitation id="3B0B8E1E8876D17654BD20B4FB44F973" ID-GBIF-Occurrence="1849198324" collectingDate="2016-03-15" collectorName="D. Couceiro" country="Peru" elevation="231" latitude="-12.875" location="Amazonas" longLatPrecision="21" longitude="-69.40972" pageId="11" pageNumber="40" specimenCount="1" specimenCount-female="1" stateProvince="Amazonas" typeStatus="paratype">
<specimenCount id="9D654FCA8876D17654BD20B4FF57F973" pageId="11" pageNumber="40" type="female">1 female</specimenCount>
, Albergue Refugio
<collectingRegion id="49A74AA18876D17650FF2098FDF4F973" box="[460,581,1704,1728]" country="Peru" name="Amazonas" pageId="11" pageNumber="40">Amazonas</collectingRegion>
,
<geoCoordinate id="EE57E2848876D17653672097FD64F973" box="[596,725,1703,1728]" direction="south" orientation="latitude" pageId="11" pageNumber="40" precision="15" value="-12.875">12°52'30''S</geoCoordinate>
,
<geoCoordinate id="EE57E2848876D17653D72097FCDDF973" box="[740,876,1703,1728]" direction="west" orientation="longitude" pageId="11" pageNumber="40" precision="15" value="-69.40972">69°24'35''W</geoCoordinate>
,
<elevation id="004E63708876D176524F2097FC0FF90C" box="[892,958,1703,1727]" metricMagnitude="2" metricUnit="m" metricValue="2.31" pageId="11" pageNumber="40" unit="m" value="231.0">
<quantity id="4C9B29A68876D176524F2097FC0FF90C" box="[892,958,1703,1727]" metricMagnitude="2" metricUnit="m" metricValue="2.31" pageId="11" pageNumber="40" unit="m" value="231.0">231m</quantity>
</elevation>
,
<collectingDate id="EF995B6B8876D17652FE2097FBF7F973" box="[973,1094,1703,1728]" pageId="11" pageNumber="40" value="2016-03-15">
<date id="FFDDA2838876D17652FE2097FBF7F973" box="[973,1094,1703,1728]" pageId="11" pageNumber="40" value="2016-03-15">15.iii.2016</date>
</collectingDate>
(
<collectorName id="2696E1958876D17655682097FB5CF973" box="[1115,1261,1703,1728]" pageId="11" pageNumber="40">D. Couceiro</collectorName>
)
</materialsCitation>
; 1 male, idem except,
<date id="FFDDA2838876D17651DD20FBFED1F957" box="[238,352,1739,1764]" pageId="11" pageNumber="40" value="2016-05-20">20.v.2016</date>
; 1male, idem except
<date id="FFDDA2838876D176537F20FBFD71F957" box="[588,704,1739,1764]" pageId="11" pageNumber="40" value="2016-10-02">02.x.2016</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C379D7C88876D17651F420DFFBACF8C7" pageId="11" pageNumber="40" type="diagnosis">
<paragraph id="8BDC84438876D17651F420DFFBACF8C7" blockId="11.[151,1437,1631,2016]" pageId="11" pageNumber="40">
<emphasis id="B91758518876D17651F420DFFE8BF8BB" bold="true" box="[199,314,1775,1800]" pageId="11" pageNumber="40">Diagnosis</emphasis>
(male): Small species (
<quantity id="4C9B29A68876D176537620DFFD29F8B4" box="[581,664,1775,1799]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="40" unit="mm" value="11.0">11 mm</quantity>
), with yellowish red head. Thorax brown with yellow and red spots. Forewings brown with red edged yellow spots. Hindwings red with brown external margin which does not reach the anal angle. An expansion of the tornus heads towards the back and presents a fold in the inner margin. The internal margin also bears an elongated patch of brown scales on its ventral side.
</paragraph>
</subSubSection>
<subSubSection id="C379D7C88876D17151F4214FFDBDFA1E" lastPageId="12" lastPageNumber="41" pageId="11" pageNumber="40" type="description">
<paragraph id="8BDC84438876D17151F4214FFDBDFA1E" blockId="11.[151,1437,1631,2016]" lastBlockId="12.[151,1437,151,1633]" lastPageId="12" lastPageNumber="41" pageId="11" pageNumber="40">
<emphasis id="B91758518876D17651F4214FFE25F82B" bold="true" box="[199,404,1919,1944]" pageId="11" pageNumber="40">Description male</emphasis>
(
<figureCitation id="135898C68876D1765097214FFD9DF824" box="[420,556,1919,1944]" captionStart="FIGURES 3134" captionStartId="13.[151,264,742,765]" captionTargetBox="[340,1247,195,721]" captionTargetId="figure@13.[340,1247,195,721]" captionTargetPageId="13" captionText="FIGURES 3134. Paranerita maculata sandeepani Grados, new subespecies. 31. Holotype male, dorsal. 32. Holotype male, ventral. 33. Paratype female, dorsal (Albergue Refugio Amazonas). 34. Paratype female, ventral. Scales=5 mm." httpUri="https://zenodo.org/record/1290952/files/figure.png" pageId="11" pageNumber="40">Figs. 3132</figureCitation>
) and female (
<figureCitation id="135898C68876D17653F9214FFCE4F824" box="[714,853,1919,1944]" captionStart="FIGURES 3134" captionStartId="13.[151,264,742,765]" captionTargetBox="[340,1247,195,721]" captionTargetId="figure@13.[340,1247,195,721]" captionTargetPageId="13" captionText="FIGURES 3134. Paranerita maculata sandeepani Grados, new subespecies. 31. Holotype male, dorsal. 32. Holotype male, ventral. 33. Paratype female, dorsal (Albergue Refugio Amazonas). 34. Paratype female, ventral. Scales=5 mm." httpUri="https://zenodo.org/record/1290952/files/figure.png" pageId="11" pageNumber="40">Figs. 3334</figureCitation>
). Forewing span:
<quantity id="4C9B29A68876D1765512214FFBC1F824" box="[1057,1136,1919,1943]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="40" unit="mm" value="11.0">11 mm</quantity>
in males (n=3),
<quantity id="4C9B29A68876D176541D214FFACFF824" box="[1326,1406,1919,1943]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="40" unit="mm" value="12.0">12 mm</quantity>
in female (n=1.
<emphasis id="B91758518876D17650072193FECCF80F" bold="true" box="[308,381,1955,1980]" pageId="11" pageNumber="40">Head:</emphasis>
Frontoclypeus yellow, with a brown cross line. At the base of the antennae, with scattered red scales. Vertex yellow, with posterolateral parts with red spots, which project forward until reaching the base of the antennae. Palpi yellow, with red scales on the external side of the first segment and on the dorsolateral side of the three proximal quarters of the length of the second segment. Antennae with rami reduced with scape and pedicel both yellow. Red scales in the proximal fifth and, in the distal third with white scales.
<emphasis id="B91758518871D171557C26EFFB00FF4B" bold="true" box="[1103,1201,223,248]" pageId="12" pageNumber="41">Thorax:</emphasis>
Brown. Patagia with red scales on the anterior part, from the middle of the area and towards the mesal margin. Tegula with red scales towards the posterior part. Two yellow spots edged in red, located in the central and mesal part of the mesoscutum. Presence of yellow and red hairs between the metascutum and metascutellum. First pair of legs yellow, with the proximal half of the dorsal side of the femur with a red line and the distal end with a red dot. Dorsal side and mesal middle margin red. Tibia dorsal side red; and on the ectal margin, almost from the middle to the distal end brown. Second and third pair of legs, yellow.
<emphasis id="B91758518871D171536B2787FD63FE63" bold="true" box="[600,722,439,464]" pageId="12" pageNumber="41">Forewing:</emphasis>
Dorsal surface brown with cream spots and red contours: an oblong spot near the base of the wing and below the basal part of the Cu, projecting obliquely, being narrower, towards the outside of the wing, reaching the basal part of the discal cell, crosses the R obliquely towards the base of the wing, reaching the costal margin; a larger spot below the intersection of Cu and Cu1 extends toward the base of cell M3-Cu1, from where it divides, extending the first branch toward the termen and covering the proximal two thirds of cell M2-M3 and which connects with a small spot that is at the distal end of the cell M3-Cu1 and reaches the termen; the other branch ascends with the discal cells internal limit, reaches the intersection of R and M1, where it becomes narrower, and then reaches the costal margin, from where a thin red line arises and goes along the margin and towards the apex without reaching it; a rounded spot in the center of the anal cell which reaches the posterior margin; a small spot on the distal end of Cu1-Cu2; a tiny spot at the distal end of cell M1-M2, reaching the termen; a somewhat elongated red spot on the distal end of cell R4-R5 and another near the base of cell R5-M1. An elongated red spot at the base of anal cell. The apex presents yellow and reddish scales. Ventral surface with the same pattern as the dorsal side with the brown background and the spots of red contours, only these are yellow instead of creamy; mesal area and posterior margin yellow. Unlike the dorsal side, the following is not observed: the rounded spot in the center of anal cell, which reaches the posterior margin; the elongated red spot at the base of anal cell; the elongated red spot on the distal end of cell R4-R5 and the another near the base of cell R5-M1.
<emphasis id="B91758518871D17155FA25D2FAF9FC48" bold="true" box="[1225,1352,994,1019]" pageId="12" pageNumber="41">Hindwing:</emphasis>
Dorsal surface red, with costal margin yellow and brown termen In females, the brown band in the termen is wider. Tornus expanded posteriorly with a lobulated shape, and a fold in the inner margin. In females, does not show the expansion. Ventral surface yellow, with an elongated oval patch of brown scales in the fold of the inner margin. In females, does not have the elongated oval patch.
<emphasis id="B91758518871D17153F62244FCF2FB3E" bold="true" box="[709,835,1140,1165]" pageId="12" pageNumber="41">Abdomen:</emphasis>
Red, with yellow scales. In the distal part of the last
<emphasis id="B91758518871D17151A422AAFF5AFB02" box="[151,235,1178,1201]" italics="true" pageId="12" pageNumber="41">sternite</emphasis>
with yellow scales. Creamy yellow ventral side.
<emphasis id="B91758518871D171520322A8FC6EFB02" bold="true" box="[816,991,1176,1201]" pageId="12" pageNumber="41">Male genitalia</emphasis>
(
<figureCitation id="135898C68871D17152C122A8FBCDFB02" box="[1010,1148,1176,1201]" captionStart="FIGURES 3538" captionStartId="13.[151,264,1352,1375]" captionTargetBox="[359,1247,851,1315]" captionTargetId="figure@13.[340,1247,845,1331]" captionTargetPageId="13" captionText="FIGURES 3538. Genitalia of Paranerita maculata sandeepani Grados, new species. (Genitalia # JGA762). 35. Dorsal view. 36. Ventral view. 37. Lateral view. 38. Aedeagus. Scales= 1 mm." httpUri="https://zenodo.org/record/1290954/files/figure.png" pageId="12" pageNumber="41">Figs. 3538</figureCitation>
) (Genitalia # JGA 762): Saccus wide and slightly curved towards the right side in dorsal view. Tegumen wide towards posterior end, with each side of triangular shape. On the external posterior side, it forms an angle of almost ninety degrees with the distal margin, which is wide and where the uncus emerges. On the lateral sides of this distal margin, it presents two small sclerosed pointed projections. Uncus slender, curved towards the ventral side. The end has a small curvature. Long valvae with the edges of the sacculus and the costa parallel, thinning towards the distal end. Juxta wide in the anterior part and thinning towards the end. Aedeagus elongated and curved upward dorsally. Caecum penis small. Vesical short with small spicules.
</paragraph>
</subSubSection>
<subSubSection id="C379D7C88871D17151F42388FD1FFA46" pageId="12" pageNumber="41" type="etymology">
<paragraph id="8BDC84438871D17151F42388FD1FFA46" blockId="12.[151,1437,151,1633]" pageId="12" pageNumber="41">
<emphasis id="B91758518871D17151F42388FEF9FA62" bold="true" box="[199,328,1464,1489]" pageId="12" pageNumber="41">Etymology</emphasis>
:
<taxonomicName id="4C63FFC08871D171506A2388FE69FA62" authorityName="Grados" authorityYear="2018" box="[345,472,1464,1489]" class="Insecta" family="Arctiidae" genus="Paranerita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="subSpecies" species="maculata" subSpecies="sandeepani">
<emphasis id="B91758518871D171506A2388FE69FA62" box="[345,472,1464,1489]" italics="true" pageId="12" pageNumber="41">sandeepani</emphasis>
</taxonomicName>
is a singular genitive noun, dedicated to Dileep Sandeepan for having collected the first specimen at the Refugio
<collectingRegion id="49A74AA18871D17150D223EDFDE6FA46" box="[481,599,1501,1525]" country="Peru" name="Amazonas" pageId="12" pageNumber="41">Amazonas</collectingRegion>
Lodge.
</paragraph>
</subSubSection>
<subSubSection id="C379D7C88871D17151F42030FCC4F9D2" pageId="12" pageNumber="41" type="distribution">
<paragraph id="8BDC84438871D17151F42030FAF3F9AA" blockId="12.[151,1437,151,1633]" box="[199,1346,1536,1561]" pageId="12" pageNumber="41">
<emphasis id="B91758518871D17151F42030FED3F9AA" bold="true" box="[199,354,1536,1561]" pageId="12" pageNumber="41">Distribution:</emphasis>
Known only from the department of
<collectingRegion id="49A74AA18871D17152362030FC1EF9AA" box="[773,943,1536,1561]" country="Peru" name="Madre de Dios" pageId="12" pageNumber="41">Madre de Dios</collectingRegion>
, southeast of the Peruvian Amazon.
</paragraph>
<paragraph id="8BDC84438871D17151F42014FCC4F9D2" blockId="12.[151,1437,151,1633]" pageId="12" pageNumber="41">
<emphasis id="B91758518871D17151F42014FEFDF98E" bold="true" box="[199,332,1572,1597]" pageId="12" pageNumber="41">Barcoding:</emphasis>
The access number to Genbank is
<accessionNumber id="943019A08871D17153E62014FCE8F98E" box="[725,857,1572,1597]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MG837876" pageId="12" pageNumber="41">MG837876</accessionNumber>
. The mitochondrial DNA sequence (COI) of one of the
<typeStatus id="54D83AE18871D17151F32079FE9AF9D2" box="[192,299,1609,1633]" pageId="12" pageNumber="41" type="paratype">paratypes</typeStatus>
is as follows (Voucher MUSM
<taxonomicName id="4C63FFC08871D17153A62078FD4FF9D2" box="[661,766,1608,1633]" class="Insecta" family="Erebidae" genus="Arctiinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="47" phylum="Arthropoda" rank="genus">Arctiinae</taxonomicName>
VB # 67):
</paragraph>
</subSubSection>
<subSubSection id="C379D7C88871D17151A420A1FDD4F85E" box="[151,1433,1681,2029]" pageId="12" pageNumber="41" type="description">
<paragraph id="8BDC84438871D17151A420A1FDD4F85E" blockId="12.[151,1433,1681,2029]" box="[151,1433,1681,2029]" pageId="12" pageNumber="41">AACTTTATATTTTATTTTTGGTATTTGAGCTGGAATAGTAGGAACTTCATTAAGTTTATTAATTCGAGCTG AATTAGGAAATCCTGGATCCTTAATTGGAGATGATCAAATTTATAATACTATTGTAACAGCACATGCTTT CATTATAATTTTCTTTATAGTTATACCAATTATAATTGGAGGATTTGGTAATTGATTAGTTCCATTAATATT AGGAGCCCCAGATATAGCCTTCCCTCGAATAAATAATATAAGTTTTTGATTATTACCCCCTTCCTTAACT CTTTTAATTTCAAGAAGAATTGTAGAAAACGGTGCTGGAACTGGATGAACTGTTTACCCCCCTCTTTC TTCTAATATTGCCCATGGAGGAAGATCAGTAGATCTAGCTATTTTTTCCCTTCATTTAGCTGGAATTTCT TCAATTTTAGGAGCAATTAATTTCATTACTACAATTATTAATATACGATTAAATAACTTATCATTTGATCA AATACCTCTATTTGTGTGAGCTGTAGGAATTACAGCATTTTTATTATTACTTTCATTACCTGTTTTAGCTG GAGCTATTACTATGTTATTAACTGATCGAAATTTAAATACTTCTTTTTTCGACCCTGCTGGGGGAGGAG ATCCAATTCTTTATCAACACTTATTT</paragraph>
</subSubSection>
<caption id="DF1CD4CB8870D17051A424D6FAB3FCA8" httpUri="https://zenodo.org/record/1290952/files/figure.png" pageId="13" pageNumber="42" startId="13.[151,264,742,765]" targetBox="[340,1247,195,721]" targetPageId="13">
<paragraph id="8BDC84438870D17051A424D6FAB3FCA8" blockId="13.[151,1436,742,796]" pageId="13" pageNumber="42">
<emphasis id="B91758518870D17051A424D6FEFFFD4F" bold="true" box="[151,334,742,765]" pageId="13" pageNumber="42">FIGURES 3134</emphasis>
.
<taxonomicName id="4C63FFC08870D170506F24D8FD4EFD4F" authority="Grados" authorityName="Grados" authorityYear="2018" box="[348,767,742,765]" class="Insecta" family="Arctiidae" genus="Paranerita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="subSpecies" species="maculata" subSpecies="sandeepani">
<emphasis id="B91758518870D170506F24D8FD1AFD4F" box="[348,683,742,765]" italics="true" pageId="13" pageNumber="42">Paranerita maculata sandeepani</emphasis>
Grados
</taxonomicName>
, new subespecies.
<emphasis id="B91758518870D17052F324D7FC68FD4F" bold="true" box="[960,985,743,764]" pageId="13" pageNumber="42">31</emphasis>
. Holotype male, dorsal.
<emphasis id="B91758518870D17055EB24D7FB40FD4F" bold="true" box="[1240,1265,743,764]" pageId="13" pageNumber="42">32</emphasis>
. Holotype male, ventral.
<emphasis id="B91758518870D17051D92535FEB2FCA9" bold="true" box="[234,259,773,794]" pageId="13" pageNumber="42">33</emphasis>
. Paratype female, dorsal (Albergue Refugio Amazonas).
<emphasis id="B91758518870D170527A2535FCD3FCA9" bold="true" box="[841,866,773,794]" pageId="13" pageNumber="42">34</emphasis>
. Paratype female, ventral. Scales=5 mm.
</paragraph>
</caption>
<caption id="DF1CD4CB8870D17051A42378FC97FACE" httpUri="https://zenodo.org/record/1290954/files/figure.png" pageId="13" pageNumber="42" startId="13.[151,264,1352,1375]" targetBox="[359,1247,851,1315]" targetPageId="13">
<paragraph id="8BDC84438870D17051A42378FC97FACE" blockId="13.[151,1437,1352,1405]" pageId="13" pageNumber="42">
<emphasis id="B91758518870D17051A42378FEFDFAEE" bold="true" box="[151,332,1352,1375]" pageId="13" pageNumber="42">FIGURES 3538</emphasis>
. Genitalia of
<taxonomicName id="4C63FFC08870D17050E52379FCC4FAED" authority="Grados" authorityName="Grados" authorityYear="2018" box="[470,885,1352,1374]" class="Insecta" family="Arctiidae" genus="Paranerita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="subSpecies" species="maculata" status="sp. nov." subSpecies="sandeepani">
<emphasis id="B91758518870D17050E52379FC92FAED" box="[470,803,1352,1374]" italics="true" pageId="13" pageNumber="42">Paranerita maculata sandeepani</emphasis>
Grados
</taxonomicName>
,
<taxonomicNameLabel id="A224E52A8870D170524D2379FC4BFAED" box="[894,1018,1352,1374]" pageId="13" pageNumber="42" rank="species">new species</taxonomicNameLabel>
. (Genitalia # JGA762).
<emphasis id="B91758518870D17055C42378FAA1FAEE" bold="true" box="[1271,1296,1352,1373]" pageId="13" pageNumber="42">35</emphasis>
. Dorsal view.
<emphasis id="B91758518870D17051A42357FF01FACF" bold="true" box="[151,176,1383,1404]" pageId="13" pageNumber="42">36</emphasis>
. Ventral view.
<emphasis id="B91758518870D170507B2357FED0FACF" bold="true" box="[328,353,1383,1404]" pageId="13" pageNumber="42">37</emphasis>
. Lateral view.
<emphasis id="B91758518870D17050CA2357FDA3FACF" bold="true" box="[505,530,1383,1404]" pageId="13" pageNumber="42">38</emphasis>
. Aedeagus. Scales= 1 mm.
</paragraph>
</caption>
<subSubSection id="C379D7C88870D17051F42394FDDDF9FE" pageId="13" pageNumber="42" type="discussion">
<paragraph id="8BDC84438870D17051F42394FDDDF9FE" blockId="13.[151,1436,1444,1984]" pageId="13" pageNumber="42">
<emphasis id="B91758518870D17051F42394FE8CFA0E" bold="true" box="[199,317,1444,1469]" pageId="13" pageNumber="42">Remarks:</emphasis>
As I pointed out for the previous species, reviewing the genitalia morphology of
<taxonomicName id="4C63FFC08870D17054122395FEE6FA53" authority="Druce" authorityName="Druce" authorityYear="1883" class="Insecta" family="Arctiidae" genus="Paranerita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="polyxenus">
<emphasis id="B91758518870D17054122395FEB6FA52" italics="true" pageId="13" pageNumber="42">Paranerita polyxenus</emphasis>
Druce
</taxonomicName>
and
<taxonomicName id="4C63FFC08870D17050A423F8FD78FA52" authority="Cramer" authorityName="Cramer" authorityYear="1777" box="[407,713,1480,1505]" class="Insecta" family="Erebidae" genus="Trichromia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="onytes">
<emphasis id="B91758518870D17050A423F8FDD7FA53" box="[407,614,1480,1505]" italics="true" pageId="13" pageNumber="42">Trichromia onytes</emphasis>
Cramer
</taxonomicName>
, and being these the
<typeStatus id="54D83AE18870D17052FA23F9FC48FA52" box="[969,1017,1481,1505]" pageId="13" pageNumber="42">type</typeStatus>
species for each respective genera, significant differences can be observed, making it unlikely that they belong to the same genus, hence I do not consider both genera to be synonyms. For these reasons, I maintain the species
<taxonomicName id="4C63FFC08870D17055522020FB78F99A" authorityName="Rothschild" authorityYear="1909" box="[1121,1225,1552,1577]" class="Insecta" family="Arctiidae" genus="Paranerita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B91758518870D17055522020FB78F99A" box="[1121,1225,1552,1577]" italics="true" pageId="13" pageNumber="42">maculata</emphasis>
</taxonomicName>
within the genus considered by
<bibRefCitation id="EFF2F9B28870D17050092005FDD9F9FE" box="[314,616,1588,1613]" pageId="13" pageNumber="42" refId="ref15136" refString="Watson, A. &amp; Goodger, D. (1986) Catalogue of the Neotropical Tiger - moths. Ocassional Papers on Systematic Entomology, 1, 1 - 71." type="journal article">Watson &amp; Goodger (1986)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C379D7C88870D17051F42068FE68F873" pageId="13" pageNumber="42" type="reference_group">
<paragraph id="8BDC84438870D17051F42068FE68F873" blockId="13.[151,1436,1444,1984]" pageId="13" pageNumber="42">
Regarding
<taxonomicName id="4C63FFC08870D17050742069FCE0F9C2" authority="maculata (Rothschild, 1909)" authorityName="Rothschild" authorityYear="1909" box="[327,849,1624,1649]" class="Insecta" family="Erebidae" genus="Evius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="subSpecies" species="maculata" subSpecies="maculata">
<emphasis id="B91758518870D17050742069FDDEF9C2" box="[327,623,1624,1649]" italics="true" pageId="13" pageNumber="42">Evius maculata maculata</emphasis>
(
<bibRefCitation id="EFF2F9B28870D17053B12068FCF9F9C2" box="[642,840,1624,1649]" pageId="13" pageNumber="42" refId="ref14263" refString="Rothschild, W. (1909) Descriptions of new South-American Arctianae. Annals and Magazine of Natural History, Series 8, 4 (21), 205 - 229. https: // doi. org / 10.1080 / 00222930908692664" type="journal article">Rothschild, 1909</bibRefCitation>
)
</taxonomicName>
it was described from a female from Fonte Boa,
<collectingRegion id="49A74AA18870D17051A4204DFEBCF926" box="[151,269,1661,1685]" country="Peru" name="Amazonas" pageId="13" pageNumber="42">Amazonas</collectingRegion>
(
<collectingCountry id="F374C4D38870D170502D204CFED4F926" box="[286,357,1660,1685]" name="Brazil" pageId="13" pageNumber="42">Brazil</collectingCountry>
) and transferred to
<taxonomicName id="4C63FFC08870D170537A204DFD75F927" authorityName="Hampson" authorityYear="1901" box="[585,708,1661,1684]" class="Insecta" family="Arctiidae" genus="Paranerita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="B91758518870D170537A204DFD75F927" box="[585,708,1661,1684]" italics="true" pageId="13" pageNumber="42">Paranerita</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFF2F9B28870D17053C1204DFB97F926" box="[754,1062,1660,1685]" pageId="13" pageNumber="42" refId="ref15136" refString="Watson, A. &amp; Goodger, D. (1986) Catalogue of the Neotropical Tiger - moths. Ocassional Papers on Systematic Entomology, 1, 1 - 71." type="journal article">Watson &amp; Goodger (1986)</bibRefCitation>
. Based on females, we observed that the
<taxonomicName id="4C63FFC08870D17051C32091FDD0F90A" authorityName="Grados" authorityYear="2018" box="[240,609,1696,1721]" class="Insecta" family="Arctiidae" genus="Paranerita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="subSpecies" species="maculata" subSpecies="sandeepani">
<emphasis id="B91758518870D17051C32091FDD0F90A" box="[240,609,1696,1721]" italics="true" pageId="13" pageNumber="42">Paranerita maculata sandeepani</emphasis>
</taxonomicName>
has the large, creamy spot that starts below the intersection of Cu and Cu1 extending towards the base of cell Cu1, where it divides. The first branch extends towards the termen, covering the proximal two thirds of the cell M2 and connects with a small spot situated on distal end of cell M3 and which reaches the term, without interruptions until reaching the termen. In
<taxonomicName id="4C63FFC08870D170528B2124FB22F89E" authority="maculata" box="[952,1171,1812,1837]" class="Insecta" family="Arctiidae" genus="Paranerita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="subSpecies" species="maculata" subSpecies="maculata">
<emphasis id="B91758518870D170528B2124FB22F89E" box="[952,1171,1812,1837]" italics="true" pageId="13" pageNumber="42">maculata maculata</emphasis>
</taxonomicName>
the large spot appears discontinuously, formed by three spots separated by brown areas. The main spot emerging at the intersection of Cu and Cu1, as it ascends has a constriction at the lower angle of the discal cell and does not extend into cell M3. The pattern of the
<taxonomicName id="4C63FFC08870D170500821B5FD01F82F" authorityName="Grados" authorityYear="2018" box="[315,688,1923,1948]" class="Insecta" family="Arctiidae" genus="Paranerita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="subSpecies" species="maculata" subSpecies="sandeepani">
<emphasis id="B91758518870D170500821B5FD01F82F" box="[315,688,1923,1948]" italics="true" pageId="13" pageNumber="42">Paranerita maculata sandeepani</emphasis>
</taxonomicName>
is also true for males, this being the first time that males of this species have been described.
</paragraph>
</subSubSection>
</treatment>
</document>