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<document id="608D1781E778303DE69AAACF5251A072" ID-DOI="10.11646/zootaxa.3847.3.1" ID-GBIF-Dataset="624e0a0e-4766-436b-a552-8aad3b411999" ID-ISSN="1175-5326" ID-Zenodo-Dep="253194" ID-ZooBank="19E58554-5B31-496B-B851-A26DEE4B7929" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460471539568" checkinUser="plazi" docAuthor="Anisyutkin, Leonid N." docDate="2014" docId="03CA2572FFD6FF8785801409C0F2B67A" docLanguage="en" docName="zt03847p332.pdf" docOrigin="Zootaxa 3847 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Morphna lucida Anisyutkin, sp. nov." docType="treatment" docVersion="11" lastPageNumber="311" masterDocId="FFF35D0AFFDEFF8D85171669C30FB71E" masterDocTitle="On cockroaches of the subfamily Epilamprinae (Dictyoptera: Blaberidae) from South India and Sri Lanka, with descriptions of new taxa" masterLastPageNumber="332" masterPageNumber="301" pageNumber="309" updateTime="1698305926497" updateUser="plazi">
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<mods:title id="34A12A1793988D4065525D66938FD7EC">On cockroaches of the subfamily Epilamprinae (Dictyoptera: Blaberidae) from South India and Sri Lanka, with descriptions of new taxa</mods:title>
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<mods:namePart id="90E6009BBB62D536131DD1E4C7BD73E5">Anisyutkin, Leonid N.</mods:namePart>
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<mods:title id="8F0F817C37D1294B1D1FB3DEE1CDED2D">Zootaxa</mods:title>
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<mods:date id="3D1F82019C40BE82355CCE2E44DE1573">2014</mods:date>
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<treatment id="03CA2572FFD6FF8785801409C0F2B67A" ID-DOI="http://doi.org/10.5281/zenodo.3501517" ID-GBIF-Taxon="119515389" ID-Zenodo-Dep="3501517" LSID="urn:lsid:plazi:treatment:03CA2572FFD6FF8785801409C0F2B67A" httpUri="http://treatment.plazi.org/id/03CA2572FFD6FF8785801409C0F2B67A" lastPageId="10" lastPageNumber="311" pageId="8" pageNumber="309">
<subSubSection id="C379C7EFFFD6FF8585801409C156B564" box="[151,601,608,634]" pageId="8" pageNumber="309" type="nomenclature">
<paragraph id="8BDC9464FFD6FF8585801409C156B564" blockId="8.[151,601,608,667]" box="[151,601,608,634]" pageId="8" pageNumber="309">
<heading id="D0942308FFD6FF8585801409C156B564" bold="true" box="[151,601,608,634]" fontSize="11" level="1" pageId="8" pageNumber="309" reason="1">
<emphasis id="B9174876FFD6FF8585801409C156B564" bold="true" box="[151,601,608,634]" pageId="8" pageNumber="309">
<taxonomicName id="4C63EFE7FFD6FF8585801409C2E0B564" ID-CoL="6S2SL" authority="Anisyutkin" authorityName="Anisyutkin" box="[151,495,608,634]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="lucida" status="sp. nov.">
<emphasis id="B9174876FFD6FF8585801409C254B564" bold="true" box="[151,347,608,634]" italics="true" pageId="8" pageNumber="309">Morphna lucida</emphasis>
Anisyutkin
</taxonomicName>
,
<taxonomicNameLabel id="A224F50DFFD6FF8584EC1409C156B564" box="[507,601,608,634]" pageId="8" pageNumber="309" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C379C7EFFFD6FF85858014EBC2BFB585" box="[151,432,642,667]" pageId="8" pageNumber="309" type="description">
<paragraph id="8BDC9464FFD6FF85858014EBC2BFB585" blockId="8.[151,601,608,667]" box="[151,432,642,667]" pageId="8" pageNumber="309">
(
<figureCitation id="135888E1FFD6FF85858814EBC20DB585" box="[159,258,642,667]" captionStart="FIGURE 1. A, D, F" captionStartId="5.[151,250,1921,1943]" captionTargetBox="[170,1400,200,1869]" captionTargetId="figure@5.[167,1406,193,1876]" captionTargetPageId="5" captionText="FIGURE 1. A, D, F, H—facial part of head; B, E, G, I, J—general view from above; C—anterior part of body from below. A C— Indoapterolampra rugosiuscula sp. nov. (male, holotype); D, E— Morphna lucida sp. nov. (male, holotype); F, G— M. plana (Brunner von Wattenwyl, 1865) (male); H J— M. decolyi (Bolivar, 1897) (H, I—male, J—female). In Figs. B, I, J abdominal apex removed. All specimens are pinned." httpUri="https://zenodo.org/record/253195/files/figure.png" pageId="8" pageNumber="309">Figs. 1D</figureCitation>
, E,
<figureCitation id="135888E1FFD6FF85843914EAC233B585" box="[302,316,643,667]" captionStart="FIGURE 4" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[188,1401,208,1824]" captionTargetId="figure@9.[175,1411,193,1830]" captionTargetPageId="9" captionText="FIGURE 4. Morphna lucida sp. nov., male, holotype. A—facial part of head; B—pronotum and head from above; C—left hind metatarsus from outside; D—abdominal apex from above; E—paraprocts from below; F—hypandrium from below; G, H—right phallomere from below, schematically (G) and above (H); I—sclerite L2D from above; J—apical part of sclerite L2D from above; KM—sclerite L3. Dotted area shows dark color (A, B) or membranous parts (C, E, F, HM). Abbreviations: c.p.R1T; d.o.; f.s.; l.s.a.; R1T; R2; R3; R4; R5; s.t.; spi.—see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = KM." httpUri="https://zenodo.org/record/253198/files/figure.png" pageId="8" pageNumber="309">4</figureCitation>
AM,
<figureCitation id="135888E1FFD6FF85849214EAC29CB585" box="[389,403,643,667]" captionStart="FIGURE 9" captionStartId="17.[151,250,1670,1692]" captionTargetBox="[151,1427,200,1638]" captionTargetId="figure@17.[151,1436,193,1648]" captionTargetPageId="17" captionText="FIGURE 9. Tegmina (A, B) and wing (C) from above. A—Morphna lucida sp. nov. (male, holotype); B, C Rhabdoblatta praecipua (Walker, 1868) (male). Abbreviations: 1pl.; A1; CuA; CuP; M; R; R+M; RA; RS; Sc—see text. Membrane reticulations omitted. Scale bar 1 mm: a = A; b = B; c = C." httpUri="https://zenodo.org/record/253203/files/figure.png" pageId="8" pageNumber="309">9</figureCitation>
A)
</paragraph>
</subSubSection>
<subSubSection id="C379C7EFFFD6FF85858014A3C267B419" pageId="8" pageNumber="309" type="materials_examined">
<paragraph id="8BDC9464FFD6FF85858014A3C267B419" blockId="8.[151,1437,714,2035]" pageId="8" pageNumber="309">
<emphasis id="B9174876FFD6FF85858014A3C3F0B5FD" bold="true" box="[151,255,714,739]" pageId="8" pageNumber="309">Material</emphasis>
.
<materialsCitation id="3B0B9E39FFD6FF85841A14A3C267B419" collectingDate="1961-05" collectionCode="MMUM" collectorName="P. S. Nathan" country="India" elevation="975" location="Devala" municipality="Nilgiri Hills" pageId="8" pageNumber="309" specimenCount="1" specimenCount-male="1" stateProvince="Tamil Nadu" typeStatus="holotype">
<typeStatus id="54D82AC6FFD6FF85841A14A3C276B5FD" box="[269,377,714,739]" pageId="8" pageNumber="309" type="holotype">Holotype</typeStatus>
<specimenCount id="9D655FEDFFD6FF85848414A3C2C4B5FD" box="[403,459,714,739]" pageId="8" pageNumber="309" type="male">male</specimenCount>
, SOUTH
<collectingCountry id="F374D4F4FFD6FF85872E14A2C184B5FD" box="[569,651,715,739]" name="India" pageId="8" pageNumber="309">INDIA</collectingCountry>
, state
<collectingRegion id="49A75A86FFD6FF8587C614A3C058B5FD" box="[721,855,714,739]" country="India" name="Tamil Nadu" pageId="8" pageNumber="309">Tamil Nadu</collectingRegion>
,
<collectingMunicipality id="6BB80E1EFFD6FF85867414A3C0E2B5FD" box="[867,1005,714,739]" pageId="8" pageNumber="309">Nilgiri Hills</collectingMunicipality>
,
<location id="8EBCC2BFFFD6FF8586EE14A3C744B5FD" LSID="urn:lsid:plazi:treatment:03CA2572FFD6FF8785801409C0F2B67A:8EBCC2BFFFD6FF8586EE14A3C744B5FD" box="[1017,1099,714,739]" country="India" municipality="Nilgiri Hills" name="Devala" pageId="8" pageNumber="309" stateProvince="Tamil Nadu">Devala</location>
,
<quantity id="4C9B3981FFD6FF85814114A2C7A0B5FD" box="[1110,1199,714,739]" metricMagnitude="2" metricUnit="m" metricValue="9.7536" pageId="8" pageNumber="309" unit="ft" value="3200.0">
<elevation id="004E7357FFD6FF85814114A2C7A0B5FD" box="[1110,1199,714,739]" metricMagnitude="2" metricUnit="m" metricValue="9.7536" pageId="8" pageNumber="309" unit="ft" value="3200.0">3200 ft.</elevation>
</quantity>
,
<date id="FFDDB2A4FFD6FF8581AD14A2C621B5FD" box="[1210,1326,714,739]" pageId="8" pageNumber="309" value="1961-05">
<collectingDate id="EF994B4CFFD6FF8581AD14A2C621B5FD" box="[1210,1326,714,739]" pageId="8" pageNumber="309" value="1961-05">May 1961</collectingDate>
</date>
, coll.
<collectorName id="2696F1B2FFD6FF85806514A2C3E7B419" pageId="8" pageNumber="309">P.S. Nathan</collectorName>
(
<collectionCode id="ED720CA1FFD6FF8585E01486C252B419" box="[247,349,751,775]" collectionName="MMUM" country="United Kingdom" httpUri="http://grbio.org/cool/c9j8-pjt7" name="University of Manchester Museum, Zoology and Entomology" pageId="8" pageNumber="309" type="Museum">MMUM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C379C7EFFFD6FF8585D0157BC7B5B099" pageId="8" pageNumber="309" type="description">
<paragraph id="8BDC9464FFD6FF8585D0157BC627B101" blockId="8.[151,1437,714,2035]" pageId="8" pageNumber="309">
<emphasis id="B9174876FFD6FF8585D0157BC25EB435" bold="true" box="[199,337,786,811]" pageId="8" pageNumber="309">Description</emphasis>
. Male (
<typeStatus id="54D82AC6FFD6FF8584BB157BC11BB435" box="[428,532,786,811]" pageId="8" pageNumber="309" type="holotype">holotype</typeStatus>
). General color light yellowish, with scattered small brown spots (
<figureCitation id="135888E1FFD6FF858007157BC67AB435" box="[1296,1397,786,811]" captionStart="FIGURE 1. A, D, F" captionStartId="5.[151,250,1921,1943]" captionTargetBox="[170,1400,200,1869]" captionTargetId="figure@5.[167,1406,193,1876]" captionTargetPageId="5" captionText="FIGURE 1. A, D, F, H—facial part of head; B, E, G, I, J—general view from above; C—anterior part of body from below. A C— Indoapterolampra rugosiuscula sp. nov. (male, holotype); D, E— Morphna lucida sp. nov. (male, holotype); F, G— M. plana (Brunner von Wattenwyl, 1865) (male); H J— M. decolyi (Bolivar, 1897) (H, I—male, J—female). In Figs. B, I, J abdominal apex removed. All specimens are pinned." httpUri="https://zenodo.org/record/253195/files/figure.png" pageId="8" pageNumber="309">Figs. 1D</figureCitation>
, E, 4A, B); facial part of head yellow, with small brown dots located mostly between eyes and on the vertex (
<figureCitation id="135888E1FFD6FF858023155FC698B451" box="[1332,1431,822,847]" captionStart="FIGURE 1. A, D, F" captionStartId="5.[151,250,1921,1943]" captionTargetBox="[170,1400,200,1869]" captionTargetId="figure@5.[167,1406,193,1876]" captionTargetPageId="5" captionText="FIGURE 1. A, D, F, H—facial part of head; B, E, G, I, J—general view from above; C—anterior part of body from below. A C— Indoapterolampra rugosiuscula sp. nov. (male, holotype); D, E— Morphna lucida sp. nov. (male, holotype); F, G— M. plana (Brunner von Wattenwyl, 1865) (male); H J— M. decolyi (Bolivar, 1897) (H, I—male, J—female). In Figs. B, I, J abdominal apex removed. All specimens are pinned." httpUri="https://zenodo.org/record/253195/files/figure.png" pageId="8" pageNumber="309">Figs. 1D</figureCitation>
,
<figureCitation id="135888E1FFD6FF8585801532C3A9B46D" box="[151,166,859,883]" captionStart="FIGURE 4" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[188,1401,208,1824]" captionTargetId="figure@9.[175,1411,193,1830]" captionTargetPageId="9" captionText="FIGURE 4. Morphna lucida sp. nov., male, holotype. A—facial part of head; B—pronotum and head from above; C—left hind metatarsus from outside; D—abdominal apex from above; E—paraprocts from below; F—hypandrium from below; G, H—right phallomere from below, schematically (G) and above (H); I—sclerite L 2 D from above; J—apical part of sclerite L 2 D from above; K M—sclerite L 3. Dotted area shows dark color (A, B) or membranous parts (C, E, F, H M). Abbreviations: c. p. R 1 T; d. o.; f. s.; l. s. a.; R 1 T; R 2; R 3; R 4; R 5; s. t.; spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K M." httpUri="https://zenodo.org/record/253198/files/figure.png" pageId="8" pageNumber="309">4</figureCitation>
A); eyes blackish brown; antennae and mouthparts yellowish; pronotum yellowish, lateral parts nearly translucent, with small brown spots at margins (
<figureCitation id="135888E1FFD6FF8587D21517C01CB489" box="[709,787,894,919]" captionStart="FIGURE 1. A, D, F" captionStartId="5.[151,250,1921,1943]" captionTargetBox="[170,1400,200,1869]" captionTargetId="figure@5.[167,1406,193,1876]" captionTargetPageId="5" captionText="FIGURE 1. A, D, F, H—facial part of head; B, E, G, I, J—general view from above; C—anterior part of body from below. A C— Indoapterolampra rugosiuscula sp. nov. (male, holotype); D, E— Morphna lucida sp. nov. (male, holotype); F, G— M. plana (Brunner von Wattenwyl, 1865) (male); H J— M. decolyi (Bolivar, 1897) (H, I—male, J—female). In Figs. B, I, J abdominal apex removed. All specimens are pinned." httpUri="https://zenodo.org/record/253195/files/figure.png" pageId="8" pageNumber="309">Figs. 1</figureCitation>
E, 4B); tegmina yellowish, with not numerous scattered small brown spots, a pair of larger ones (one on each tegmen) located at the middle of tegmina (
<figureCitation id="135888E1FFD6FF8581E015CBC636B4A5" box="[1271,1337,930,955]" captionStart="FIGURE 1. A, D, F" captionStartId="5.[151,250,1921,1943]" captionTargetBox="[170,1400,200,1869]" captionTargetId="figure@5.[167,1406,193,1876]" captionTargetPageId="5" captionText="FIGURE 1. A, D, F, H—facial part of head; B, E, G, I, J—general view from above; C—anterior part of body from below. A C— Indoapterolampra rugosiuscula sp. nov. (male, holotype); D, E— Morphna lucida sp. nov. (male, holotype); F, G— M. plana (Brunner von Wattenwyl, 1865) (male); H J— M. decolyi (Bolivar, 1897) (H, I—male, J—female). In Figs. B, I, J abdominal apex removed. All specimens are pinned." httpUri="https://zenodo.org/record/253195/files/figure.png" pageId="8" pageNumber="309">Fig. 1</figureCitation>
E); legs,
<taxonomicName id="4C63EFE7FFD6FF85858015AFC3D0B4C1" box="[151,223,966,991]" class="Insecta" family="Blaberidae" genus="Thorax" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="332" phylum="Arthropoda" rank="genus">thorax</taxonomicName>
and abdomen from below yellowish. Surfaces smooth and lustrous, distal parts of antennae (approximately from 1819th segments) and 5th segment of maxillary palps dull; fine punctation present only in basal parts of tegmina; facial part of head with transverse wrinkles between eyes, antennal sockets and above clypeus (
<figureCitation id="135888E1FFD6FF8580241267C698B339" box="[1331,1431,1038,1063]" captionStart="FIGURE 1. A, D, F" captionStartId="5.[151,250,1921,1943]" captionTargetBox="[170,1400,200,1869]" captionTargetId="figure@5.[167,1406,193,1876]" captionTargetPageId="5" captionText="FIGURE 1. A, D, F, H—facial part of head; B, E, G, I, J—general view from above; C—anterior part of body from below. A C— Indoapterolampra rugosiuscula sp. nov. (male, holotype); D, E— Morphna lucida sp. nov. (male, holotype); F, G— M. plana (Brunner von Wattenwyl, 1865) (male); H J— M. decolyi (Bolivar, 1897) (H, I—male, J—female). In Figs. B, I, J abdominal apex removed. All specimens are pinned." httpUri="https://zenodo.org/record/253195/files/figure.png" pageId="8" pageNumber="309">Figs. 1D</figureCitation>
,
<figureCitation id="135888E1FFD6FF858580125AC3A9B355" box="[151,166,1075,1099]" captionStart="FIGURE 4" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[188,1401,208,1824]" captionTargetId="figure@9.[175,1411,193,1830]" captionTargetPageId="9" captionText="FIGURE 4. Morphna lucida sp. nov., male, holotype. A—facial part of head; B—pronotum and head from above; C—left hind metatarsus from outside; D—abdominal apex from above; E—paraprocts from below; F—hypandrium from below; G, H—right phallomere from below, schematically (G) and above (H); I—sclerite L 2 D from above; J—apical part of sclerite L 2 D from above; K M—sclerite L 3. Dotted area shows dark color (A, B) or membranous parts (C, E, F, H M). Abbreviations: c. p. R 1 T; d. o.; f. s.; l. s. a.; R 1 T; R 2; R 3; R 4; R 5; s. t.; spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K M." httpUri="https://zenodo.org/record/253198/files/figure.png" pageId="8" pageNumber="309">4</figureCitation>
A). Head longer than wide (
<figureCitation id="135888E1FFD6FF8584F1125BC144B355" box="[486,587,1074,1099]" captionStart="FIGURE 1. A, D, F" captionStartId="5.[151,250,1921,1943]" captionTargetBox="[170,1400,200,1869]" captionTargetId="figure@5.[167,1406,193,1876]" captionTargetPageId="5" captionText="FIGURE 1. A, D, F, H—facial part of head; B, E, G, I, J—general view from above; C—anterior part of body from below. A C— Indoapterolampra rugosiuscula sp. nov. (male, holotype); D, E— Morphna lucida sp. nov. (male, holotype); F, G— M. plana (Brunner von Wattenwyl, 1865) (male); H J— M. decolyi (Bolivar, 1897) (H, I—male, J—female). In Figs. B, I, J abdominal apex removed. All specimens are pinned." httpUri="https://zenodo.org/record/253195/files/figure.png" pageId="8" pageNumber="309">Figs. 1D</figureCitation>
,
<figureCitation id="135888E1FFD6FF85874E125AC167B355" box="[601,616,1075,1099]" captionStart="FIGURE 4" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[188,1401,208,1824]" captionTargetId="figure@9.[175,1411,193,1830]" captionTargetPageId="9" captionText="FIGURE 4. Morphna lucida sp. nov., male, holotype. A—facial part of head; B—pronotum and head from above; C—left hind metatarsus from outside; D—abdominal apex from above; E—paraprocts from below; F—hypandrium from below; G, H—right phallomere from below, schematically (G) and above (H); I—sclerite L 2 D from above; J—apical part of sclerite L 2 D from above; K M—sclerite L 3. Dotted area shows dark color (A, B) or membranous parts (C, E, F, H M). Abbreviations: c. p. R 1 T; d. o.; f. s.; l. s. a.; R 1 T; R 2; R 3; R 4; R 5; s. t.; spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K M." httpUri="https://zenodo.org/record/253198/files/figure.png" pageId="8" pageNumber="309">4</figureCitation>
A); ocellar spots large; facial part with weak impressions between eyes; distance between eyes 0.7 times of the eye length; distance between antennal sockets 1.5 times of the scape length (
<quantity id="4C9B3981FFD6FF8585881213C3F5B38D" box="[159,250,1146,1171]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="309" unit="mm" value="1.5">1.5 mm</quantity>
); approximate length ratio of 3rd5th segments of maxillary palps 1: 1: 1.2. Pronotum as in
<figureCitation id="135888E1FFD6FF85801D1213C65CB38D" box="[1290,1363,1146,1171]" captionStart="FIGURE 1. A, D, F" captionStartId="5.[151,250,1921,1943]" captionTargetBox="[170,1400,200,1869]" captionTargetId="figure@5.[167,1406,193,1876]" captionTargetPageId="5" captionText="FIGURE 1. A, D, F, H—facial part of head; B, E, G, I, J—general view from above; C—anterior part of body from below. A C— Indoapterolampra rugosiuscula sp. nov. (male, holotype); D, E— Morphna lucida sp. nov. (male, holotype); F, G— M. plana (Brunner von Wattenwyl, 1865) (male); H J— M. decolyi (Bolivar, 1897) (H, I—male, J—female). In Figs. B, I, J abdominal apex removed. All specimens are pinned." httpUri="https://zenodo.org/record/253195/files/figure.png" pageId="8" pageNumber="309">Figs. 1</figureCitation>
E, 4B. Tegmina and wings completely developed, surpassed abdominal apex, truncated apically (
<figureCitation id="135888E1FFD6FF85818512F7C7D2B3A9" box="[1170,1245,1182,1207]" captionStart="FIGURE 1. A, D, F" captionStartId="5.[151,250,1921,1943]" captionTargetBox="[170,1400,200,1869]" captionTargetId="figure@5.[167,1406,193,1876]" captionTargetPageId="5" captionText="FIGURE 1. A, D, F, H—facial part of head; B, E, G, I, J—general view from above; C—anterior part of body from below. A C— Indoapterolampra rugosiuscula sp. nov. (male, holotype); D, E— Morphna lucida sp. nov. (male, holotype); F, G— M. plana (Brunner von Wattenwyl, 1865) (male); H J— M. decolyi (Bolivar, 1897) (H, I—male, J—female). In Figs. B, I, J abdominal apex removed. All specimens are pinned." httpUri="https://zenodo.org/record/253195/files/figure.png" pageId="8" pageNumber="309">Figs. 1</figureCitation>
E, 9A). Tegmina with emarginated apex (
<figureCitation id="135888E1FFD6FF8584A512ABC10EB3C5" box="[434,513,1218,1243]" captionStart="FIGURE 1. A, D, F" captionStartId="5.[151,250,1921,1943]" captionTargetBox="[170,1400,200,1869]" captionTargetId="figure@5.[167,1406,193,1876]" captionTargetPageId="5" captionText="FIGURE 1. A, D, F, H—facial part of head; B, E, G, I, J—general view from above; C—anterior part of body from below. A C— Indoapterolampra rugosiuscula sp. nov. (male, holotype); D, E— Morphna lucida sp. nov. (male, holotype); F, G— M. plana (Brunner von Wattenwyl, 1865) (male); H J— M. decolyi (Bolivar, 1897) (H, I—male, J—female). In Figs. B, I, J abdominal apex removed. All specimens are pinned." httpUri="https://zenodo.org/record/253195/files/figure.png" pageId="8" pageNumber="309">Figs. 1</figureCitation>
E, 9A); weakly sclerotized; venation distinct; costal field wider and shorter as compared with those structures of
<taxonomicName id="4C63EFE7FFD6FF858727128FC049B3E0" box="[560,838,1254,1279]" class="Insecta" family="Blaberidae" genus="Rhabdoblatta" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="praecipua">
<emphasis id="B9174876FFD6FF858727128FC049B3E0" box="[560,838,1254,1279]" italics="true" pageId="8" pageNumber="309">Rhabdoblatta praecipua</emphasis>
</taxonomicName>
(
<figureCitation id="135888E1FFD6FF85864E128FC0AFB3E1" box="[857,928,1254,1279]" captionStart="FIGURE 9" captionStartId="17.[151,250,1670,1692]" captionTargetBox="[151,1427,200,1638]" captionTargetId="figure@17.[151,1436,193,1648]" captionTargetPageId="17" captionText="FIGURE 9. Tegmina (A, B) and wing (C) from above. A— Morphna lucida sp. nov. (male, holotype); B, C Rhabdoblatta praecipua (Walker, 1868) (male). Abbreviations: 1 pl.; A 1; CuA; CuP; M; R; R + M; RA; RS; Sc — see text. Membrane reticulations omitted. Scale bar 1 mm: a = A; b = B; c = C." httpUri="https://zenodo.org/record/253203/files/figure.png" pageId="8" pageNumber="309">Fig. 9</figureCitation>
B) and well sclerotized;
<emphasis id="B9174876FFD6FF8581D3128EC7D0B3E0" box="[1220,1247,1255,1278]" italics="true" pageId="8" pageNumber="309">Sc</emphasis>
thickened (well visible on ventral side of tegmen);
<emphasis id="B9174876FFD6FF85873E1365C135B23D" box="[553,570,1292,1315]" italics="true" pageId="8" pageNumber="309">R</emphasis>
,
<emphasis id="B9174876FFD6FF85875D1365C16EB23D" box="[586,609,1292,1315]" italics="true" pageId="8" pageNumber="309">M</emphasis>
and
<emphasis id="B9174876FFD6FF85878B1362C1C1B23C" box="[668,718,1291,1314]" italics="true" pageId="8" pageNumber="309">CuA</emphasis>
stems separated;
<emphasis id="B9174876FFD6FF85868D1362C0C3B23C" box="[922,972,1291,1314]" italics="true" pageId="8" pageNumber="309">CuP</emphasis>
distinct. Anterior margin of fore femora armed according to the type B, with 7 spines, including 2 apical ones. Structure of hind tarsi: metatarsus slightly shorter than other segments combined, with euplantula more than 3/4 of segment length (
<figureCitation id="135888E1FFD6FF85819B133BC7DDB275" box="[1164,1234,1362,1387]" captionStart="FIGURE 4" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[188,1401,208,1824]" captionTargetId="figure@9.[175,1411,193,1830]" captionTargetPageId="9" captionText="FIGURE 4. Morphna lucida sp. nov., male, holotype. A—facial part of head; B—pronotum and head from above; C—left hind metatarsus from outside; D—abdominal apex from above; E—paraprocts from below; F—hypandrium from below; G, H—right phallomere from below, schematically (G) and above (H); I—sclerite L 2 D from above; J—apical part of sclerite L 2 D from above; K M—sclerite L 3. Dotted area shows dark color (A, B) or membranous parts (C, E, F, H M). Abbreviations: c. p. R 1 T; d. o.; f. s.; l. s. a.; R 1 T; R 2; R 3; R 4; R 5; s. t.; spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K M." httpUri="https://zenodo.org/record/253198/files/figure.png" pageId="8" pageNumber="309">Fig. 4</figureCitation>
C); euplantulae of 2nd-4th segments large; spines absent, with exception of several delicate spinules located on metatarsal euplantula (
<figureCitation id="135888E1FFD6FF85858813F3C3EDB2AD" box="[159,226,1434,1459]" captionStart="FIGURE 4" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[188,1401,208,1824]" captionTargetId="figure@9.[175,1411,193,1830]" captionTargetPageId="9" captionText="FIGURE 4. Morphna lucida sp. nov., male, holotype. A—facial part of head; B—pronotum and head from above; C—left hind metatarsus from outside; D—abdominal apex from above; E—paraprocts from below; F—hypandrium from below; G, H—right phallomere from below, schematically (G) and above (H); I—sclerite L 2 D from above; J—apical part of sclerite L 2 D from above; K M—sclerite L 3. Dotted area shows dark color (A, B) or membranous parts (C, E, F, H M). Abbreviations: c. p. R 1 T; d. o.; f. s.; l. s. a.; R 1 T; R 2; R 3; R 4; R 5; s. t.; spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K M." httpUri="https://zenodo.org/record/253198/files/figure.png" pageId="8" pageNumber="309">Fig. 4</figureCitation>
C,
<emphasis id="B9174876FFD6FF85841613F5C226B2AD" box="[257,297,1436,1459]" italics="true" pageId="8" pageNumber="309">spi.</emphasis>
); claws symmetrical, simple; arolium about half of claw length. Anal plate (tergite X) widely rounded caudally, with small triangular median incision (
<figureCitation id="135888E1FFD6FF8587D513D7C008B2C9" box="[706,775,1470,1495]" captionStart="FIGURE 4" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[188,1401,208,1824]" captionTargetId="figure@9.[175,1411,193,1830]" captionTargetPageId="9" captionText="FIGURE 4. Morphna lucida sp. nov., male, holotype. A—facial part of head; B—pronotum and head from above; C—left hind metatarsus from outside; D—abdominal apex from above; E—paraprocts from below; F—hypandrium from below; G, H—right phallomere from below, schematically (G) and above (H); I—sclerite L 2 D from above; J—apical part of sclerite L 2 D from above; K M—sclerite L 3. Dotted area shows dark color (A, B) or membranous parts (C, E, F, H M). Abbreviations: c. p. R 1 T; d. o.; f. s.; l. s. a.; R 1 T; R 2; R 3; R 4; R 5; s. t.; spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K M." httpUri="https://zenodo.org/record/253198/files/figure.png" pageId="8" pageNumber="309">Fig. 4</figureCitation>
D). Cerci long, flatten, with segments distinctly separated (
<figureCitation id="135888E1FFD6FF858588138BC3E8B2E5" box="[159,231,1506,1531]" captionStart="FIGURE 4" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[188,1401,208,1824]" captionTargetId="figure@9.[175,1411,193,1830]" captionTargetPageId="9" captionText="FIGURE 4. Morphna lucida sp. nov., male, holotype. A—facial part of head; B—pronotum and head from above; C—left hind metatarsus from outside; D—abdominal apex from above; E—paraprocts from below; F—hypandrium from below; G, H—right phallomere from below, schematically (G) and above (H); I—sclerite L 2 D from above; J—apical part of sclerite L 2 D from above; K M—sclerite L 3. Dotted area shows dark color (A, B) or membranous parts (C, E, F, H M). Abbreviations: c. p. R 1 T; d. o.; f. s.; l. s. a.; R 1 T; R 2; R 3; R 4; R 5; s. t.; spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K M." httpUri="https://zenodo.org/record/253198/files/figure.png" pageId="8" pageNumber="309">Fig. 4</figureCitation>
D). Paraprocts of blaberid-type (
<figureCitation id="135888E1FFD6FF858749138BC1AAB2E5" box="[606,677,1506,1531]" captionStart="FIGURE 4" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[188,1401,208,1824]" captionTargetId="figure@9.[175,1411,193,1830]" captionTargetPageId="9" captionText="FIGURE 4. Morphna lucida sp. nov., male, holotype. A—facial part of head; B—pronotum and head from above; C—left hind metatarsus from outside; D—abdominal apex from above; E—paraprocts from below; F—hypandrium from below; G, H—right phallomere from below, schematically (G) and above (H); I—sclerite L 2 D from above; J—apical part of sclerite L 2 D from above; K M—sclerite L 3. Dotted area shows dark color (A, B) or membranous parts (C, E, F, H M). Abbreviations: c. p. R 1 T; d. o.; f. s.; l. s. a.; R 1 T; R 2; R 3; R 4; R 5; s. t.; spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K M." httpUri="https://zenodo.org/record/253198/files/figure.png" pageId="8" pageNumber="309">Fig. 4</figureCitation>
E). Hypandrium nearly symmetrical (
<figureCitation id="135888E1FFD6FF858142138BC793B2E5" box="[1109,1180,1506,1531]" captionStart="FIGURE 4" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[188,1401,208,1824]" captionTargetId="figure@9.[175,1411,193,1830]" captionTargetPageId="9" captionText="FIGURE 4. Morphna lucida sp. nov., male, holotype. A—facial part of head; B—pronotum and head from above; C—left hind metatarsus from outside; D—abdominal apex from above; E—paraprocts from below; F—hypandrium from below; G, H—right phallomere from below, schematically (G) and above (H); I—sclerite L 2 D from above; J—apical part of sclerite L 2 D from above; K M—sclerite L 3. Dotted area shows dark color (A, B) or membranous parts (C, E, F, H M). Abbreviations: c. p. R 1 T; d. o.; f. s.; l. s. a.; R 1 T; R 2; R 3; R 4; R 5; s. t.; spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K M." httpUri="https://zenodo.org/record/253198/files/figure.png" pageId="8" pageNumber="309">Fig. 4</figureCitation>
F); antero-lateral parts short (
<figureCitation id="135888E1FFD6FF8585C9106FC22EB101" box="[222,289,1542,1567]" captionStart="FIGURE 4" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[188,1401,208,1824]" captionTargetId="figure@9.[175,1411,193,1830]" captionTargetPageId="9" captionText="FIGURE 4. Morphna lucida sp. nov., male, holotype. A—facial part of head; B—pronotum and head from above; C—left hind metatarsus from outside; D—abdominal apex from above; E—paraprocts from below; F—hypandrium from below; G, H—right phallomere from below, schematically (G) and above (H); I—sclerite L 2 D from above; J—apical part of sclerite L 2 D from above; K M—sclerite L 3. Dotted area shows dark color (A, B) or membranous parts (C, E, F, H M). Abbreviations: c. p. R 1 T; d. o.; f. s.; l. s. a.; R 1 T; R 2; R 3; R 4; R 5; s. t.; spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K M." httpUri="https://zenodo.org/record/253198/files/figure.png" pageId="8" pageNumber="309">Fig. 4</figureCitation>
F,
<emphasis id="B9174876FFD6FF85842B106FC27EB101" box="[316,369,1542,1567]" italics="true" pageId="8" pageNumber="309">l.s.a.</emphasis>
); caudal margin between styli rounded, with distinct median incision; styli fusiform.
</paragraph>
<paragraph id="8BDC9464FFD6FF8585D01043C0D6B005" blockId="8.[151,1437,714,2035]" pageId="8" pageNumber="309">
Male genitalia (
<figureCitation id="135888E1FFD6FF8584601043C2B5B15D" box="[375,442,1578,1603]" captionStart="FIGURE 4" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[188,1401,208,1824]" captionTargetId="figure@9.[175,1411,193,1830]" captionTargetPageId="9" captionText="FIGURE 4. Morphna lucida sp. nov., male, holotype. A—facial part of head; B—pronotum and head from above; C—left hind metatarsus from outside; D—abdominal apex from above; E—paraprocts from below; F—hypandrium from below; G, H—right phallomere from below, schematically (G) and above (H); I—sclerite L 2 D from above; J—apical part of sclerite L 2 D from above; K M—sclerite L 3. Dotted area shows dark color (A, B) or membranous parts (C, E, F, H M). Abbreviations: c. p. R 1 T; d. o.; f. s.; l. s. a.; R 1 T; R 2; R 3; R 4; R 5; s. t.; spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K M." httpUri="https://zenodo.org/record/253198/files/figure.png" pageId="8" pageNumber="309">Fig. 4</figureCitation>
GM). Right phallomere (R+N) with caudal part of sclerite R1T subrectangular in shape (
<figureCitation id="135888E1FFD6FF8585881027C3E8B179" box="[159,231,1614,1639]" captionStart="FIGURE 4" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[188,1401,208,1824]" captionTargetId="figure@9.[175,1411,193,1830]" captionTargetPageId="9" captionText="FIGURE 4. Morphna lucida sp. nov., male, holotype. A—facial part of head; B—pronotum and head from above; C—left hind metatarsus from outside; D—abdominal apex from above; E—paraprocts from below; F—hypandrium from below; G, H—right phallomere from below, schematically (G) and above (H); I—sclerite L 2 D from above; J—apical part of sclerite L 2 D from above; K M—sclerite L 3. Dotted area shows dark color (A, B) or membranous parts (C, E, F, H M). Abbreviations: c. p. R 1 T; d. o.; f. s.; l. s. a.; R 1 T; R 2; R 3; R 4; R 5; s. t.; spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K M." httpUri="https://zenodo.org/record/253198/files/figure.png" pageId="8" pageNumber="309">Fig. 4</figureCitation>
G, H,
<emphasis id="B9174876FFD6FF8584241026C285B179" box="[307,394,1615,1639]" italics="true" pageId="8" pageNumber="309">c.p.R1T</emphasis>
), densely covered with bristles; R2 weakly curved (
<figureCitation id="135888E1FFD6FF8586EA1027C74AB179" box="[1021,1093,1614,1639]" captionStart="FIGURE 4" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[188,1401,208,1824]" captionTargetId="figure@9.[175,1411,193,1830]" captionTargetPageId="9" captionText="FIGURE 4. Morphna lucida sp. nov., male, holotype. A—facial part of head; B—pronotum and head from above; C—left hind metatarsus from outside; D—abdominal apex from above; E—paraprocts from below; F—hypandrium from below; G, H—right phallomere from below, schematically (G) and above (H); I—sclerite L 2 D from above; J—apical part of sclerite L 2 D from above; K M—sclerite L 3. Dotted area shows dark color (A, B) or membranous parts (C, E, F, H M). Abbreviations: c. p. R 1 T; d. o.; f. s.; l. s. a.; R 1 T; R 2; R 3; R 4; R 5; s. t.; spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K M." httpUri="https://zenodo.org/record/253198/files/figure.png" pageId="8" pageNumber="309">Fig. 4</figureCitation>
G, H); R3 distinctly widened caudally (
<figureCitation id="135888E1FFD6FF858411101BC247B195" box="[262,328,1650,1675]" captionStart="FIGURE 4" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[188,1401,208,1824]" captionTargetId="figure@9.[175,1411,193,1830]" captionTargetPageId="9" captionText="FIGURE 4. Morphna lucida sp. nov., male, holotype. A—facial part of head; B—pronotum and head from above; C—left hind metatarsus from outside; D—abdominal apex from above; E—paraprocts from below; F—hypandrium from below; G, H—right phallomere from below, schematically (G) and above (H); I—sclerite L 2 D from above; J—apical part of sclerite L 2 D from above; K M—sclerite L 3. Dotted area shows dark color (A, B) or membranous parts (C, E, F, H M). Abbreviations: c. p. R 1 T; d. o.; f. s.; l. s. a.; R 1 T; R 2; R 3; R 4; R 5; s. t.; spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K M." httpUri="https://zenodo.org/record/253198/files/figure.png" pageId="8" pageNumber="309">Fig. 4</figureCitation>
G, H); R4 large (
<figureCitation id="135888E1FFD6FF858713101BC147B195" box="[516,584,1650,1675]" captionStart="FIGURE 4" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[188,1401,208,1824]" captionTargetId="figure@9.[175,1411,193,1830]" captionTargetPageId="9" captionText="FIGURE 4. Morphna lucida sp. nov., male, holotype. A—facial part of head; B—pronotum and head from above; C—left hind metatarsus from outside; D—abdominal apex from above; E—paraprocts from below; F—hypandrium from below; G, H—right phallomere from below, schematically (G) and above (H); I—sclerite L 2 D from above; J—apical part of sclerite L 2 D from above; K M—sclerite L 3. Dotted area shows dark color (A, B) or membranous parts (C, E, F, H M). Abbreviations: c. p. R 1 T; d. o.; f. s.; l. s. a.; R 1 T; R 2; R 3; R 4; R 5; s. t.; spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K M." httpUri="https://zenodo.org/record/253198/files/figure.png" pageId="8" pageNumber="309">Fig. 4</figureCitation>
H); R5 large, fused with sclerite R3 (
<figureCitation id="135888E1FFD6FF8586FF101BC724B195" box="[1000,1067,1650,1675]" captionStart="FIGURE 4" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[188,1401,208,1824]" captionTargetId="figure@9.[175,1411,193,1830]" captionTargetPageId="9" captionText="FIGURE 4. Morphna lucida sp. nov., male, holotype. A—facial part of head; B—pronotum and head from above; C—left hind metatarsus from outside; D—abdominal apex from above; E—paraprocts from below; F—hypandrium from below; G, H—right phallomere from below, schematically (G) and above (H); I—sclerite L 2 D from above; J—apical part of sclerite L 2 D from above; K M—sclerite L 3. Dotted area shows dark color (A, B) or membranous parts (C, E, F, H M). Abbreviations: c. p. R 1 T; d. o.; f. s.; l. s. a.; R 1 T; R 2; R 3; R 4; R 5; s. t.; spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K M." httpUri="https://zenodo.org/record/253198/files/figure.png" pageId="8" pageNumber="309">Fig. 4</figureCitation>
G, H). Sclerite L2D (L1) divided into basal and apical parts (
<figureCitation id="135888E1FFD6FF8584D110FFC105B1B1" box="[454,522,1686,1711]" captionStart="FIGURE 4" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[188,1401,208,1824]" captionTargetId="figure@9.[175,1411,193,1830]" captionTargetPageId="9" captionText="FIGURE 4. Morphna lucida sp. nov., male, holotype. A—facial part of head; B—pronotum and head from above; C—left hind metatarsus from outside; D—abdominal apex from above; E—paraprocts from below; F—hypandrium from below; G, H—right phallomere from below, schematically (G) and above (H); I—sclerite L 2 D from above; J—apical part of sclerite L 2 D from above; K M—sclerite L 3. Dotted area shows dark color (A, B) or membranous parts (C, E, F, H M). Abbreviations: c. p. R 1 T; d. o.; f. s.; l. s. a.; R 1 T; R 2; R 3; R 4; R 5; s. t.; spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K M." httpUri="https://zenodo.org/record/253198/files/figure.png" pageId="8" pageNumber="309">
Fig.
<date id="FFDDB2A4FFD6FF8584ED10FEC105B1B1" box="[506,522,1687,1711]" pageId="8" pageNumber="309">4</date>
</figureCitation>
I); basal part rod-like, moderately widened cranially (
<figureCitation id="135888E1FFD6FF85814C10FFC790B1B1" box="[1115,1183,1686,1711]" captionStart="FIGURE 4" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[188,1401,208,1824]" captionTargetId="figure@9.[175,1411,193,1830]" captionTargetPageId="9" captionText="FIGURE 4. Morphna lucida sp. nov., male, holotype. A—facial part of head; B—pronotum and head from above; C—left hind metatarsus from outside; D—abdominal apex from above; E—paraprocts from below; F—hypandrium from below; G, H—right phallomere from below, schematically (G) and above (H); I—sclerite L 2 D from above; J—apical part of sclerite L 2 D from above; K M—sclerite L 3. Dotted area shows dark color (A, B) or membranous parts (C, E, F, H M). Abbreviations: c. p. R 1 T; d. o.; f. s.; l. s. a.; R 1 T; R 2; R 3; R 4; R 5; s. t.; spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K M." httpUri="https://zenodo.org/record/253198/files/figure.png" pageId="8" pageNumber="309">
Fig.
<date id="FFDDB2A4FFD6FF85819810FEC790B1B1" box="[1167,1183,1687,1711]" pageId="8" pageNumber="309">4</date>
</figureCitation>
I); apical part rounded, densely covered with recumbent bristles (
<figureCitation id="135888E1FFD6FF85877A10D3C1BDB1CD" box="[621,690,1722,1747]" captionStart="FIGURE 4" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[188,1401,208,1824]" captionTargetId="figure@9.[175,1411,193,1830]" captionTargetPageId="9" captionText="FIGURE 4. Morphna lucida sp. nov., male, holotype. A—facial part of head; B—pronotum and head from above; C—left hind metatarsus from outside; D—abdominal apex from above; E—paraprocts from below; F—hypandrium from below; G, H—right phallomere from below, schematically (G) and above (H); I—sclerite L 2 D from above; J—apical part of sclerite L 2 D from above; K M—sclerite L 3. Dotted area shows dark color (A, B) or membranous parts (C, E, F, H M). Abbreviations: c. p. R 1 T; d. o.; f. s.; l. s. a.; R 1 T; R 2; R 3; R 4; R 5; s. t.; spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K M." httpUri="https://zenodo.org/record/253198/files/figure.png" pageId="8" pageNumber="309">Fig. 4</figureCitation>
J); &quot;dorsal outgrows&quot; flat (
<figureCitation id="135888E1FFD6FF8586CB10D3C72FB1CD" box="[988,1056,1722,1747]" captionStart="FIGURE 4" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[188,1401,208,1824]" captionTargetId="figure@9.[175,1411,193,1830]" captionTargetPageId="9" captionText="FIGURE 4. Morphna lucida sp. nov., male, holotype. A—facial part of head; B—pronotum and head from above; C—left hind metatarsus from outside; D—abdominal apex from above; E—paraprocts from below; F—hypandrium from below; G, H—right phallomere from below, schematically (G) and above (H); I—sclerite L 2 D from above; J—apical part of sclerite L 2 D from above; K M—sclerite L 3. Dotted area shows dark color (A, B) or membranous parts (C, E, F, H M). Abbreviations: c. p. R 1 T; d. o.; f. s.; l. s. a.; R 1 T; R 2; R 3; R 4; R 5; s. t.; spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K M." httpUri="https://zenodo.org/record/253198/files/figure.png" pageId="8" pageNumber="309">Fig. 4</figureCitation>
J,
<emphasis id="B9174876FFD6FF85812E10D3C76CB1CD" box="[1081,1123,1722,1747]" italics="true" pageId="8" pageNumber="309">d.o.</emphasis>
). Sclerite L3 (L2d) without basal subsclerite; &quot;folded structure&quot; distinct, without bristles (
<figureCitation id="135888E1FFD6FF85865910B7C09CB1E9" box="[846,915,1758,1783]" captionStart="FIGURE 4" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[188,1401,208,1824]" captionTargetId="figure@9.[175,1411,193,1830]" captionTargetPageId="9" captionText="FIGURE 4. Morphna lucida sp. nov., male, holotype. A—facial part of head; B—pronotum and head from above; C—left hind metatarsus from outside; D—abdominal apex from above; E—paraprocts from below; F—hypandrium from below; G, H—right phallomere from below, schematically (G) and above (H); I—sclerite L 2 D from above; J—apical part of sclerite L 2 D from above; K M—sclerite L 3. Dotted area shows dark color (A, B) or membranous parts (C, E, F, H M). Abbreviations: c. p. R 1 T; d. o.; f. s.; l. s. a.; R 1 T; R 2; R 3; R 4; R 5; s. t.; spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K M." httpUri="https://zenodo.org/record/253198/files/figure.png" pageId="8" pageNumber="309">Fig. 4</figureCitation>
K, L,
<emphasis id="B9174876FFD6FF8586C310B6C0F9B1E9" box="[980,1014,1759,1783]" italics="true" pageId="8" pageNumber="309">f.s.</emphasis>
); apex of L3 with &quot;small tooth&quot; (
<figureCitation id="135888E1FFD6FF85807910B7C3AAB005" captionStart="FIGURE 4" captionStartId="9.[151,250,1851,1873]" captionTargetBox="[188,1401,208,1824]" captionTargetId="figure@9.[175,1411,193,1830]" captionTargetPageId="9" captionText="FIGURE 4. Morphna lucida sp. nov., male, holotype. A—facial part of head; B—pronotum and head from above; C—left hind metatarsus from outside; D—abdominal apex from above; E—paraprocts from below; F—hypandrium from below; G, H—right phallomere from below, schematically (G) and above (H); I—sclerite L 2 D from above; J—apical part of sclerite L 2 D from above; K M—sclerite L 3. Dotted area shows dark color (A, B) or membranous parts (C, E, F, H M). Abbreviations: c. p. R 1 T; d. o.; f. s.; l. s. a.; R 1 T; R 2; R 3; R 4; R 5; s. t.; spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K M." httpUri="https://zenodo.org/record/253198/files/figure.png" pageId="8" pageNumber="309">Fig. 4</figureCitation>
KM,
<emphasis id="B9174876FFD6FF8585F9116DC201B005" box="[238,270,1796,1819]" italics="true" pageId="8" pageNumber="309">s.t.</emphasis>
), groove hge absent. Sclerite L4U (L3d) distinct and triangular.
</paragraph>
<paragraph id="8BDC9464FFD6FF8585D0114FC29FB021" blockId="8.[151,1437,714,2035]" box="[199,400,1830,1855]" pageId="8" pageNumber="309">Female unknown.</paragraph>
<paragraph id="8BDC9464FFD6FF8585D01122C7B5B099" blockId="8.[151,1437,714,2035]" pageId="8" pageNumber="309">
Measurements (mm). Head length 5.7, head width 5.4, pronotum length 9.2, pronotum width 12.5, tegmen length 40.0, tegmen width (in place where
<emphasis id="B9174876FFD6FF8587611106C1A7B098" box="[630,680,1903,1926]" italics="true" pageId="8" pageNumber="309">CuP</emphasis>
running into posterior margin of tegmen) 13.0.
</paragraph>
</subSubSection>
<subSubSection id="C379C7EFFFD6FF8785D011FBC0F2B67A" lastPageId="10" lastPageNumber="311" pageId="8" pageNumber="309" type="diagnosis">
<paragraph id="8BDC9464FFD6FF8785D011FBC074B7CB" blockId="8.[151,1437,714,2035]" lastBlockId="10.[151,1437,151,357]" lastPageId="10" lastPageNumber="311" pageId="8" pageNumber="309">
<emphasis id="B9174876FFD6FF8585D011FBC26EB0B5" bold="true" box="[199,353,1938,1963]" pageId="8" pageNumber="309">Comparison.</emphasis>
The new species can readily be distinguished from
<taxonomicName id="4C63EFE7FFD6FF8586A811FDC753B0B5" box="[959,1116,1938,1963]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="auriculata">
<emphasis id="B9174876FFD6FF8586A811FDC753B0B5" box="[959,1116,1938,1963]" italics="true" pageId="8" pageNumber="309">M. auriculata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C63EFE7FFD6FF85817A11FDC7DAB0B5" box="[1133,1237,1938,1963]" class="Insecta" family="Blaberidae" genus="Epilampra" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="badia">
<emphasis id="B9174876FFD6FF85817A11FDC7DAB0B5" box="[1133,1237,1938,1963]" italics="true" pageId="8" pageNumber="309">M. badia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C63EFE7FFD6FF8581F111FDC662B0B5" box="[1254,1389,1938,1963]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="clypeata">
<emphasis id="B9174876FFD6FF8581F111FDC662B0B5" box="[1254,1389,1938,1963]" italics="true" pageId="8" pageNumber="309">M. clypeata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C63EFE7FFD6FF85806A11FDC3D0B0D1" class="Insecta" family="Blaberidae" genus="Epilampra" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="dotata">
<emphasis id="B9174876FFD6FF85806A11FDC3D0B0D1" italics="true" pageId="8" pageNumber="309">M. dotata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C63EFE7FFD6FF8585FA11DEC28BB0D1" box="[237,388,1974,1999]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="B9174876FFD6FF8585FA11DEC28BB0D1" box="[237,388,1974,1999]" italics="true" pageId="8" pageNumber="309">M. humeralis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C63EFE7FFD6FF85848511DEC130B0D0" box="[402,575,1975,1998]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="imperatoria">
<emphasis id="B9174876FFD6FF85848511DEC130B0D0" box="[402,575,1975,1998]" italics="true" pageId="8" pageNumber="309">M. imperatoria</emphasis>
</taxonomicName>
,
<taxonomicName id="4C63EFE7FFD6FF85875A11DEC1D3B0D1" box="[589,732,1974,1999]" class="Insecta" family="Blaberidae" genus="Epilampra" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B9174876FFD6FF85875A11DEC1D3B0D1" box="[589,732,1974,1999]" italics="true" pageId="8" pageNumber="309">M. maculata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C63EFE7FFD6FF8587FC11DEC06BB0D1" box="[747,868,1974,1999]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="moloch">
<emphasis id="B9174876FFD6FF8587FC11DEC06BB0D1" box="[747,868,1974,1999]" italics="true" pageId="8" pageNumber="309">M. moloch</emphasis>
</taxonomicName>
,
<taxonomicName id="4C63EFE7FFD6FF85866511DEC0D7B0D1" box="[882,984,1974,1999]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="plana">
<emphasis id="B9174876FFD6FF85866511DEC0D7B0D1" box="[882,984,1974,1999]" italics="true" pageId="8" pageNumber="309">M. plana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C63EFE7FFD6FF85810711DEC791B0D1" box="[1040,1182,1974,1999]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="pustulata">
<emphasis id="B9174876FFD6FF85810711DEC791B0D1" box="[1040,1182,1974,1999]" italics="true" pageId="8" pageNumber="309">M. pustulata</emphasis>
</taxonomicName>
by its nearly uniform, light yellowish coloration, with brown spots being small and sparse.
<taxonomicName id="4C63EFE7FFD6FF85868E11B3C741B0ED" box="[921,1102,2010,2035]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="lucida" status="sp. nov.">
<emphasis id="B9174876FFD6FF85868E11B3C741B0ED" box="[921,1102,2010,2035]" italics="true" pageId="8" pageNumber="309">Morphna lucida</emphasis>
</taxonomicName>
<emphasis id="B9174876FFD6FF85814211B3C7A1B0ED" bold="true" box="[1109,1198,2010,2035]" pageId="8" pageNumber="309">
<taxonomicNameLabel id="A224F50DFFD6FF85814211B3C7A1B0ED" box="[1109,1198,2010,2035]" pageId="8" pageNumber="309" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="4C63EFE7FFD4FF8785C016F0C25FB7AE" box="[215,336,151,176]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="decolyi">
<emphasis id="B9174876FFD4FF8785C016F0C25FB7AE" box="[215,336,151,176]" italics="true" pageId="10" pageNumber="311">M. decolyi</emphasis>
</taxonomicName>
by the fully developed tegmina and wings surpassing the abdominal apex; these structures are strongly abbreviated in
<taxonomicName id="4C63EFE7FFD4FF8784B716D4C119B7CB" box="[416,534,188,213]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="decolyi">
<emphasis id="B9174876FFD4FF8784B716D4C119B7CB" box="[416,534,188,213]" italics="true" pageId="10" pageNumber="311">M. decolyi</emphasis>
</taxonomicName>
(see below for the description).
</paragraph>
<caption id="DF1CC4ECFFD7FF8485801152C27EB0F4" httpUri="https://zenodo.org/record/253198/files/figure.png" pageId="9" pageNumber="310" targetBox="[188,1401,208,1824]" targetPageId="9">
<paragraph id="8BDC9464FFD7FF8485801152C27EB0F4" blockId="9.[151,1437,1851,2026]" pageId="9" pageNumber="310">
<emphasis id="B9174876FFD7FF8485801152C21AB04E" bold="true" box="[151,277,1851,1873]" pageId="9" pageNumber="310">FIGURE 4.</emphasis>
<taxonomicName id="4C63EFE7FFD7FF84840A1152C2CCB04F" box="[285,451,1851,1873]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="lucida" status="sp. nov.">
<emphasis id="B9174876FFD7FF84840A1152C2CCB04F" box="[285,451,1851,1873]" italics="true" pageId="9" pageNumber="310">Morphna lucida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A224F50DFFD7FF8484DB1152C110B04E" box="[460,543,1851,1872]" pageId="9" pageNumber="310" rank="species">
<emphasis id="B9174876FFD7FF8484DB1152C110B04E" bold="true" box="[460,543,1851,1872]" pageId="9" pageNumber="310">sp. nov.</emphasis>
</taxonomicNameLabel>
, male, holotype. A—facial part of head; B—pronotum and head from above; C—left hind metatarsus from outside; D—abdominal apex from above; E—paraprocts from below; F—hypandrium from below; G, H—right phallomere from below, schematically (G) and above (H); I—sclerite L2D from above; J—apical part of sclerite L2D from above; KM—sclerite L3. Dotted area shows dark color (A, B) or membranous parts (C, E, F, HM). Abbreviations:
<emphasis id="B9174876FFD7FF84858011DFC3EAB0D2" box="[151,229,1974,1996]" italics="true" pageId="9" pageNumber="310">c.p.R1T</emphasis>
;
<emphasis id="B9174876FFD7FF8485E611DFC217B0D2" box="[241,280,1974,1996]" italics="true" pageId="9" pageNumber="310">d.o.</emphasis>
;
<emphasis id="B9174876FFD7FF84843311DFC24CB0D2" box="[292,323,1974,1996]" italics="true" pageId="9" pageNumber="310">f.s.</emphasis>
;
<emphasis id="B9174876FFD7FF84845811DFC28FB0D2" box="[335,384,1974,1996]" italics="true" pageId="9" pageNumber="310">l.s.a.</emphasis>
;
<emphasis id="B9174876FFD7FF84849B11DFC2BAB0D2" box="[396,437,1974,1996]" italics="true" pageId="9" pageNumber="310">R1T</emphasis>
;
<emphasis id="B9174876FFD7FF8484D411DFC2D0B0D2" box="[451,479,1974,1996]" italics="true" pageId="9" pageNumber="310">R2</emphasis>
;
<emphasis id="B9174876FFD7FF8484FB11DFC107B0D2" box="[492,520,1974,1996]" italics="true" pageId="9" pageNumber="310">R3</emphasis>
;
<emphasis id="B9174876FFD7FF84870211DFC13EB0D2" box="[533,561,1974,1996]" italics="true" pageId="9" pageNumber="310">R4</emphasis>
;
<emphasis id="B9174876FFD7FF84872911DEC155B0D2" box="[574,602,1975,1996]" italics="true" pageId="9" pageNumber="310">R5</emphasis>
;
<emphasis id="B9174876FFD7FF84877F11DEC18AB0D2" box="[616,645,1975,1996]" italics="true" pageId="9" pageNumber="310">s.t.</emphasis>
;
<emphasis id="B9174876FFD7FF84878511DEC1B8B0D2" box="[658,695,1975,1996]" italics="true" pageId="9" pageNumber="310">spi.</emphasis>
—see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = KM.
</paragraph>
</caption>
<paragraph id="8BDC9464FFD4FF8785D01689C0F2B67A" blockId="10.[151,1437,151,357]" pageId="10" pageNumber="311">
By its light coloration, the new species is somewhat similar to
<taxonomicName id="4C63EFE7FFD4FF8786961688C720B7E6" box="[897,1071,223,248]" class="Insecta" family="Blaberidae" genus="Epilampra" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="amplipennis">
<emphasis id="B9174876FFD4FF8786961688C720B7E6" box="[897,1071,223,248]" italics="true" pageId="10" pageNumber="311">M. amplipennis</emphasis>
</taxonomicName>
, but differs from it in having the comparatively smaller pronotum [the ratio length of tegmen/length of pronotum in
<taxonomicName id="4C63EFE7FFD4FF878129176CC7A5B603" box="[1086,1194,260,285]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="lucida" status="sp. nov.">
<emphasis id="B9174876FFD4FF878129176CC7A5B603" box="[1086,1194,260,285]" italics="true" pageId="10" pageNumber="311">M. lucida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A224F50DFFD4FF8781A6176DC604B603" box="[1201,1291,260,285]" pageId="10" pageNumber="311" rank="species">
<emphasis id="B9174876FFD4FF8781A6176DC604B603" bold="true" box="[1201,1291,260,285]" pageId="10" pageNumber="311">sp. nov.</emphasis>
</taxonomicNameLabel>
is about 4.3, while in
<taxonomicName id="4C63EFE7FFD4FF8785EB1740C2A3B65E" box="[252,428,295,320]" class="Insecta" family="Blaberidae" genus="Epilampra" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="amplipennis">
<emphasis id="B9174876FFD4FF8785EB1740C2A3B65E" box="[252,428,295,320]" italics="true" pageId="10" pageNumber="311">M. amplipennis</emphasis>
</taxonomicName>
—3.6], the distinctly elongated abdomen and the color pattern of pronotum and tegmina [cf.
<figureCitation id="135888E1FFD4FF8785D31725C209B67A" box="[196,262,332,357]" captionStart="FIGURE 1. A, D, F" captionStartId="5.[151,250,1921,1943]" captionTargetBox="[170,1400,200,1869]" captionTargetId="figure@5.[167,1406,193,1876]" captionTargetPageId="5" captionText="FIGURE 1. A, D, F, H—facial part of head; B, E, G, I, J—general view from above; C—anterior part of body from below. A C— Indoapterolampra rugosiuscula sp. nov. (male, holotype); D, E— Morphna lucida sp. nov. (male, holotype); F, G— M. plana (Brunner von Wattenwyl, 1865) (male); H J— M. decolyi (Bolivar, 1897) (H, I—male, J—female). In Figs. B, I, J abdominal apex removed. All specimens are pinned." httpUri="https://zenodo.org/record/253195/files/figure.png" pageId="10" pageNumber="311">Figs 1</figureCitation>
E, 4B and photos of
<taxonomicName id="4C63EFE7FFD4FF8784F81724C192B67B" box="[495,669,332,357]" class="Insecta" family="Blaberidae" genus="Epilampra" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="amplipennis">
<emphasis id="B9174876FFD4FF8784F81724C192B67B" box="[495,669,332,357]" italics="true" pageId="10" pageNumber="311">M. amplipennis</emphasis>
</taxonomicName>
<typeStatus id="54D82AC6FFD4FF8787B31725C009B67B" box="[676,774,332,357]" pageId="10" pageNumber="311" type="holotype">holotype</typeStatus>
in
<bibRefCitation id="EFF2E995FFD4FF87863D1725C0FEB67A" accessDate="2013-05-24" author="Beccaloni, G. W." box="[810,1009,332,357]" firstAuthor="Beccaloni" pageId="10" pageNumber="311" publicationUrl="http://Blattodea.SpeciesFile.org" refId="ref19486" refString="Beccaloni, G. W. (2007) Blattodea Species File Online. Version 1.2 / 4.0. Available from: http: // Blattodea. SpeciesFile. org (accessed 24 May 2013)" title="Blattodea Species File Online. Version 1.2 / 4.0." type="journal article" year="2007">Beccaloni (2007)</bibRefCitation>
].
</paragraph>
</subSubSection>
</treatment>
</document>