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<bibRefCitation author="Latreille, P. A." box="[285,488,220,244]" pageId="3" pageNumber="404" refId="ref15370" refString="Latreille, P. A. (1829) Les crustaces, les arachnides et les insectes: distribues en familles naturelles. Vol. 2. Chez Deterville, Paris, 556 pp. https: // doi. org / 10.5962 / bhl. title. 11575" type="book" year="1829">Latreille, 1829: 194</bibRefCitation>
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(not
<treatmentCitation author="Latreille, P. A." box="[545,750,219,244]" page="574" pageId="3" pageNumber="404" year="1828">
<bibRefCitation author="Latreille, P. A." box="[545,750,219,244]" pageId="3" pageNumber="404" refId="ref15309" refString="Latreille, P. A. (1828) Pimeliaires. Pimeliariae. In: Bory de Saint-Vincent, M. (Ed.), Dictionnaire classique d'histoire naturelle. Vol. 13. Rey et Gravier, Paris, pp. 573 - 583. https: // doi. org / 10.5962 / bhl. title. 33901" type="book" year="1828">Latreille, 1828: 574</bibRefCitation>
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, nor
<treatmentCitation author="Latreille, P. A." box="[804,997,220,244]" page="18" pageId="3" pageNumber="404" year="1829">
<bibRefCitation author="Latreille, P. A." box="[804,997,220,244]" pageId="3" pageNumber="404" refId="ref15370" refString="Latreille, P. A. (1829) Les crustaces, les arachnides et les insectes: distribues en familles naturelles. Vol. 2. Chez Deterville, Paris, 556 pp. https: // doi. org / 10.5962 / bhl. title. 11575" type="book" year="1829">Latreille, 1829: 18</bibRefCitation>
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);
<treatmentCitation author="Laporte, F. L." box="[1018,1198,219,244]" page="86" pageId="3" pageNumber="404" year="1833">
<bibRefCitation author="Laporte, F. L." box="[1018,1198,219,244]" pageId="3" pageNumber="404" pagination="17 - 88" refId="ref15273" refString="Laporte, F. L. (1833) Essai d'une classification systematique de l'orde des Hemipteres (Hemipteres-Heteropteres Latr.). Magasin de Zoologie, 2 (1), 17 - 88." type="journal article" year="1833">Laporte 1833: 86</bibRefCitation>
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;
<treatmentCitation author="Stal, C." box="[1210,1351,219,244]" page="31" pageId="3" pageNumber="404" year="1870">
<bibRefCitation author="Stal, C." box="[1210,1351,219,244]" pageId="3" pageNumber="404" pagination="1 - 232" refId="ref16840" refString="Stal, C. (1870) Enumeratio Hemipterorum. Bidrag till en f ˆ rteckning ˆ fver alla hittills kanda Hemiptera, jemte systematiska meddelanden. 1. Kongliga Svenska Vetenskaps-Akademiens Handlingar, 9 (1), 1 - 232." type="journal article" year="1870">Stål 1870: 31</bibRefCitation>
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;
<bibRefCitation author="Distant, W. L." box="[1363,1436,220,244]" pageId="3" pageNumber="404" refId="ref14323" refString="Distant, W. L. (1880) Insecta. Rhynchota, Hemiptera-Heteroptera. In: Godman, F. D. &amp; Salvin, O. (Eds.), Biologia Centrali Americana. Vol. 1. R. H. Porter, London, pp. 1 - 88." type="book" year="1880">Distant</bibRefCitation>
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1880: 33;
<treatmentCitation author="Distant, W. L." box="[303,476,250,274]" page="55" pageId="3" pageNumber="404" year="1900">
<bibRefCitation author="Distant, W. L." box="[303,476,250,274]" pageId="3" pageNumber="404" pagination="55 - 64" refId="ref14378" refString="Distant, W. L. (1900) Rhynchotal notes. - V. Heteroptera: Asopinae and Tessaratominae. Journal of Natural History, 6 (31), 55 - 64. https: // doi. org / 10.1080 / 00222930008678340" type="journal article" year="1900">Distant 1900: 55</bibRefCitation>
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;
<treatmentCitation author="Schouteden, H." box="[487,704,250,274]" page="18" pageId="3" pageNumber="404" year="1907">
<bibRefCitation author="Schouteden, H." box="[487,704,250,274]" pageId="3" pageNumber="404" refId="ref16706" refString="Schouteden, H. (1907) Heteroptera, Fam. Pentatomidae, Subfam. Asopinae (Amyoteinae). Genera Insectorum 52. P. Wytsman, Bruxelles, 82 pp." type="book" year="1907">Schouteden 1907: 18</bibRefCitation>
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;
<treatmentCitation author="Kormilev, N. A." box="[716,912,250,274]" page="40" pageId="3" pageNumber="404" year="1951">
<bibRefCitation author="Kormilev, N. A." box="[716,912,250,274]" pageId="3" pageNumber="404" pagination="35 - 45" refId="ref15240" refString="Kormilev, N. A. (1951) Notas sobre hemipteros bolivianos con la descripcion de un genero y tres especies nuevos. Folia Universitaria, 5, 35 - 45." type="journal article" year="1951">Kormilev 1951: 40</bibRefCitation>
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;
<treatmentCitation author="Piran, A. A." box="[923,1076,250,274]" page="19" pageId="3" pageNumber="404" year="1954">
<bibRefCitation author="Piran, A. A." box="[923,1076,250,274]" pageId="3" pageNumber="404" pagination="19 - 21" refId="ref15561" refString="Piran, A. A. (1954) Dos especies nuevas de Heteroscelis de las faunas de la Argentina y Paraguay (Hemiptera, Pentatomidae, Asopinae). Revista de la Sociedad Entomologica Argentina, 17 (1 - 2), 19 - 21." type="journal article" year="1954">Pirán 1954: 19</bibRefCitation>
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;
<treatmentCitation author="Piran, A. A." box="[1087,1240,250,274]" page="84" pageId="3" pageNumber="404" year="1961">
<bibRefCitation author="Piran, A. A." box="[1087,1240,250,274]" pageId="3" pageNumber="404" pagination="83 - 99" refId="ref15609" refString="Piran, A. A. (1961) Sinopsis del genero Heteroscelis Latreille 1829 (Hem. Pentatomidae) con la descripcion de cinco especies nuevas. Revista de Investigaciones Agricolas, 15, 83 - 99." type="journal article" year="1961">Pirán 1961: 84</bibRefCitation>
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;
<treatmentCitation author="Thomas, D. B." box="[1251,1432,250,274]" page="45" pageId="3" pageNumber="404" year="1992">
<bibRefCitation author="Thomas, D. B." box="[1251,1432,250,274]" pageId="3" pageNumber="404" refId="ref16883" refString="Thomas, D. B. (1992) Taxonomic synopsis of the asopine Pentatomidae (Heteroptera) of the Western Hemisphere. Vol. 16. Thomas Say Foundation Monographs. Entomological Society of America, New York, New York, 156 pp." type="book" year="1992">Thomas 1992: 45</bibRefCitation>
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;
<treatmentCitation author="Grazia, J. &amp; Schwertner, C. F." box="[199,527,281,305]" page="713" pageId="3" pageNumber="404" year="2011">
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<bibRefCitation author="Grazia, J. &amp; Panizzi, A. R. &amp; Greve, C. &amp; Schwertner, C. F. &amp; Campos, L. A. &amp; Garbelotto, T. A. &amp; Fernandes, J. A. M." box="[538,777,281,305]" pageId="3" pageNumber="404" refId="ref14821" refString="Grazia, J., Panizzi, A. R., Greve, C., Schwertner, C. F., Campos, L. A., Garbelotto, T. A. &amp; Fernandes, J. A. M. (2015) Stink bugs (Pentatomidae). In: Panizzi, A. R. &amp; Grazia, J. (Eds.), True bugs (Heteroptera) of the neotropics. Vol. 2. Springer, Dordrecht, pp. 681 - 756. https: // doi. org / 10.1007 / 978 - 94 - 017 - 9861 - 7 _ 22" type="book" year="2015">
Grazia
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. 2015: 686
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<bibRefCitation author="Cazorla, D." box="[788,966,281,305]" pageId="3" pageNumber="404" pagination="3 - 133" refId="ref14158" refString="Cazorla, D. (2021) Pentatomidae (Hemiptera: Heteroptera) de Venezuela. Revista Nicarag ¸ ense de Entomologia, 234, 3 - 133." type="journal article" year="2021">Cazorla 2021: 10</bibRefCitation>
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Castro-Huertas
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2022: 9
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<treatmentCitation author="Spinola, M." box="[151,493,342,366]" page="306" pageId="3" pageNumber="404" year="1837">
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<emphasis box="[151,292,342,366]" italics="true" pageId="3" pageNumber="404">Phyllocheirus</emphasis>
<bibRefCitation author="Spinola, M." box="[298,493,342,366]" pageId="3" pageNumber="404" refId="ref16739" refString="Spinola, M. (1837) Essai sur les genres d'insectes appartenants a l'ordre des hemipteres, Lin., ou rhyngotes, Fab., et a la section des heteropteres, Dufour. Chez Yves Gravier, Geneva, 382 pp." type="book" year="1837">Spinola, 1837: 306</bibRefCitation>
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<treatmentCitation author="Kirkaldy, G. W." box="[504,692,342,366]" page="30" pageId="3" pageNumber="404" year="1909">
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(syn.).
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<emphasis box="[151,280,373,397]" italics="true" pageId="3" pageNumber="404">Phyllochirus</emphasis>
<bibRefCitation author="Amyot, C. J. B. &amp; Serville, A." box="[287,570,373,397]" pageId="3" pageNumber="404" refId="ref13461" refString="Amyot, C. J. B. &amp; Serville, A. (1843) Histoire naturelle des insectes: Hemipteres. Vol. 1. Librairie encyclopedique de Roret, Paris, 675 pp." type="book" year="1843">Amyot &amp; Serville, 1843: 80</bibRefCitation>
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(emendation);
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<bibRefCitation author="Stal, C." box="[725,863,373,397]" pageId="3" pageNumber="404" pagination="81 - 118" refId="ref16811" refString="Stal, C. (1862) Hemiptera Mexicana enumeravit speciesque novas descripsit. Stettiner Entomologische Zeitung, 23 (1 - 2), 81 - 118." type="journal article" year="1862">Stål 1862: 93</bibRefCitation>
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;
<treatmentCitation author="Stal, C." box="[874,1009,373,397]" page="31" pageId="3" pageNumber="404" year="1870">
<bibRefCitation author="Stal, C." box="[874,1009,373,397]" pageId="3" pageNumber="404" pagination="1 - 232" refId="ref16840" refString="Stal, C. (1870) Enumeratio Hemipterorum. Bidrag till en f ˆ rteckning ˆ fver alla hittills kanda Hemiptera, jemte systematiska meddelanden. 1. Kongliga Svenska Vetenskaps-Akademiens Handlingar, 9 (1), 1 - 232." type="journal article" year="1870">Stål 1870: 31</bibRefCitation>
</treatmentCitation>
(syn.);
<treatmentCitation author="Kirkaldy, G. W." box="[1087,1276,373,397]" page="30" pageId="3" pageNumber="404" year="1909">
<bibRefCitation author="Kirkaldy, G. W." box="[1087,1276,373,397]" pageId="3" pageNumber="404" refId="ref15076" refString="Kirkaldy, G. W. (1909) Catalogue of the Hemiptera (Heteroptera) with biological and anatomical references, lists of foodplants and parasites, etc. Vol. I Cimicidae. F. L. Dames, Berlin, xl + 392 pp. https: // doi. org / 10.5962 / bhl. title. 15205" type="book" year="1909">Kirkaldy 1909: 30</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="3.[151,1437,219,520]" box="[151,1040,403,428]" pageId="3" pageNumber="404">
<treatmentCitationGroup box="[151,1040,403,428]" pageId="3" pageNumber="404">
<treatmentCitation author="Stal, C." box="[151,396,403,428]" page="434" pageId="3" pageNumber="404" year="1859">
<taxonomicName authority="Stal, 1859: 434" authorityName="Stal" authorityPageNumber="434" authorityYear="1859" box="[151,396,403,428]" class="Insecta" genus="Agerrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis box="[151,233,404,427]" italics="true" pageId="3" pageNumber="404">Agerrus</emphasis>
<bibRefCitation author="Stal, C." box="[239,396,403,428]" pageId="3" pageNumber="404" pagination="219 - 298" refId="ref16784" refString="Stal, C. (1859) Hemiptera. Species novas descripsit. Konglinga Svenska Fregatten Eugenies Resa Omkring Jorden, 3, 219 - 298." type="journal article" year="1859">Stål, 1859: 434</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation author="Stal, C." box="[407,541,403,428]" page="93" pageId="3" pageNumber="404" year="1862">
<bibRefCitation author="Stal, C." box="[407,541,403,428]" pageId="3" pageNumber="404" pagination="81 - 118" refId="ref16811" refString="Stal, C. (1862) Hemiptera Mexicana enumeravit speciesque novas descripsit. Stettiner Entomologische Zeitung, 23 (1 - 2), 81 - 118." type="journal article" year="1862">Stål 1862: 93</bibRefCitation>
</treatmentCitation>
(syn.);
<treatmentCitation author="Schouteden, H." box="[620,836,404,428]" page="19" pageId="3" pageNumber="404" year="1907">
<bibRefCitation author="Schouteden, H." box="[620,836,404,428]" pageId="3" pageNumber="404" refId="ref16706" refString="Schouteden, H. (1907) Heteroptera, Fam. Pentatomidae, Subfam. Asopinae (Amyoteinae). Genera Insectorum 52. P. Wytsman, Bruxelles, 82 pp." type="book" year="1907">Schouteden 1907: 19</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Kirkaldy, G. W." box="[847,1036,404,428]" page="30" pageId="3" pageNumber="404" year="1909">
<bibRefCitation author="Kirkaldy, G. W." box="[847,1036,404,428]" pageId="3" pageNumber="404" refId="ref15076" refString="Kirkaldy, G. W. (1909) Catalogue of the Hemiptera (Heteroptera) with biological and anatomical references, lists of foodplants and parasites, etc. Vol. I Cimicidae. F. L. Dames, Berlin, xl + 392 pp. https: // doi. org / 10.5962 / bhl. title. 15205" type="book" year="1909">Kirkaldy 1909: 30</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="3.[151,1437,219,520]" box="[151,1437,434,458]" pageId="3" pageNumber="404">
<treatmentCitationGroup box="[151,1437,434,458]" pageId="3" pageNumber="404">
<treatmentCitation author="Walker, F." box="[151,432,434,458]" page="119" pageId="3" pageNumber="404" year="1867">
<taxonomicName authority="Walker, 1867: 119" authorityName="Walker" authorityPageNumber="119" authorityYear="1867" box="[151,432,434,458]" class="Insecta" genus="Bodetria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis box="[151,240,434,458]" italics="true" pageId="3" pageNumber="404">Bodetria</emphasis>
<bibRefCitation author="Walker, F." box="[245,432,434,458]" pageId="3" pageNumber="404" refId="ref16929" refString="Walker, F. (1867) Catalogue of the specimens of Hemiptera Heteroptera in the collection of the British Museum. Part I. E. Newman, London, 240 pp." type="book" year="1867">Walker, 1867: 119</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation author="Walker, F." box="[441,623,434,458]" page="528" pageId="3" pageNumber="404" year="1868">
<bibRefCitation author="Walker, F." box="[441,623,434,458]" pageId="3" pageNumber="404" pagination="418 - 599" refId="ref16963" refString="Walker, F. (1868) s. n. In: Catalogue of the specimens of Hemiptera Heteroptera in the collection of the British Museum. Part III. E. Newman, London, pp. 418 - 599." type="book chapter" year="1868">Walker 1868: 528</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Stal, C." box="[632,765,434,458]" page="31" pageId="3" pageNumber="404" year="1870">
<bibRefCitation author="Stal, C." box="[632,765,434,458]" pageId="3" pageNumber="404" pagination="1 - 232" refId="ref16840" refString="Stal, C. (1870) Enumeratio Hemipterorum. Bidrag till en f ˆ rteckning ˆ fver alla hittills kanda Hemiptera, jemte systematiska meddelanden. 1. Kongliga Svenska Vetenskaps-Akademiens Handlingar, 9 (1), 1 - 232." type="journal article" year="1870">Stål 1870: 31</bibRefCitation>
</treatmentCitation>
(syn.);
<treatmentCitation author="Distant, W. L." box="[842,1012,434,458]" page="63" pageId="3" pageNumber="404" year="1900">
<bibRefCitation author="Distant, W. L." box="[842,1012,434,458]" pageId="3" pageNumber="404" pagination="55 - 64" refId="ref14378" refString="Distant, W. L. (1900) Rhynchotal notes. - V. Heteroptera: Asopinae and Tessaratominae. Journal of Natural History, 6 (31), 55 - 64. https: // doi. org / 10.1080 / 00222930008678340" type="journal article" year="1900">Distant 1900: 63</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Schouteden, H." box="[1021,1236,434,458]" page="19" pageId="3" pageNumber="404" year="1907">
<bibRefCitation author="Schouteden, H." box="[1021,1236,434,458]" pageId="3" pageNumber="404" refId="ref16706" refString="Schouteden, H. (1907) Heteroptera, Fam. Pentatomidae, Subfam. Asopinae (Amyoteinae). Genera Insectorum 52. P. Wytsman, Bruxelles, 82 pp." type="book" year="1907">Schouteden 1907: 19</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Kirkaldy, G. W." box="[1246,1433,434,458]" page="30" pageId="3" pageNumber="404" year="1909">
<bibRefCitation author="Kirkaldy, G. W." box="[1246,1433,434,458]" pageId="3" pageNumber="404" refId="ref15076" refString="Kirkaldy, G. W. (1909) Catalogue of the Hemiptera (Heteroptera) with biological and anatomical references, lists of foodplants and parasites, etc. Vol. I Cimicidae. F. L. Dames, Berlin, xl + 392 pp. https: // doi. org / 10.5962 / bhl. title. 15205" type="book" year="1909">Kirkaldy 1909: 30</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="3.[151,1437,219,520]" box="[151,990,465,489]" pageId="3" pageNumber="404">
<treatmentCitationGroup box="[151,990,465,489]" pageId="3" pageNumber="404">
<treatmentCitation author="Schouteden, H." box="[151,543,465,489]" page="19" pageId="3" pageNumber="404" year="1907">
<taxonomicName authority="Schouteden, 1907: 19" authorityName="Schouteden" authorityPageNumber="19" authorityYear="1907" box="[151,543,465,489]" class="Insecta" family="Pentatomidae" genus="Heterosceloides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis box="[151,313,465,489]" italics="true" pageId="3" pageNumber="404">Heterosceloides</emphasis>
<bibRefCitation author="Schouteden, H." box="[320,543,465,489]" pageId="3" pageNumber="404" refId="ref16706" refString="Schouteden, H. (1907) Heteroptera, Fam. Pentatomidae, Subfam. Asopinae (Amyoteinae). Genera Insectorum 52. P. Wytsman, Bruxelles, 82 pp." type="book" year="1907">Schouteden, 1907: 19</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation author="Kirkaldy, G. W." box="[553,739,465,489]" page="30" pageId="3" pageNumber="404" year="1909">
<bibRefCitation author="Kirkaldy, G. W." box="[553,739,465,489]" pageId="3" pageNumber="404" refId="ref15076" refString="Kirkaldy, G. W. (1909) Catalogue of the Hemiptera (Heteroptera) with biological and anatomical references, lists of foodplants and parasites, etc. Vol. I Cimicidae. F. L. Dames, Berlin, xl + 392 pp. https: // doi. org / 10.5962 / bhl. title. 15205" type="book" year="1909">Kirkaldy 1909: 30</bibRefCitation>
</treatmentCitation>
(syn.);
<bibRefCitation author="McDonald, F. J. D." box="[817,986,465,489]" pageId="3" pageNumber="404" pagination="7 - 150" refId="ref15463" refString="McDonald, F. J. D. (1966) The genitalia of North American Pentatomoidea (Hemiptera: Heteroptera). Quaestiones Entomologicae, 2, 7 - 150." type="journal article" year="1966">
<collectingCountry box="[817,925,465,489]" name="Heard Island and McDonald Islands" pageId="3" pageNumber="404">McDonald</collectingCountry>
1966
</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection box="[151,1190,495,520]" pageId="3" pageNumber="404" type="type_taxon">
<paragraph blockId="3.[151,1437,219,520]" box="[151,1190,495,520]" pageId="3" pageNumber="404">
<typeStatus box="[151,201,496,520]" pageId="3" pageNumber="404">Type</typeStatus>
species:
<taxonomicName authority="Laporte, 1833" authorityName="Laporte" authorityYear="1833" box="[294,655,496,520]" class="Insecta" family="Pentatomidae" genus="Heteroscelis" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="servillii">
<emphasis box="[294,503,496,520]" italics="true" pageId="3" pageNumber="404">Heteroscelis servillii</emphasis>
<bibRefCitation author="Laporte, F. L." box="[510,655,496,520]" pageId="3" pageNumber="404" pagination="17 - 88" refId="ref15273" refString="Laporte, F. L. (1833) Essai d'une classification systematique de l'orde des Hemipteres (Hemipteres-Heteropteres Latr.). Magasin de Zoologie, 2 (1), 17 - 88." type="journal article" year="1833">Laporte, 1833</bibRefCitation>
</taxonomicName>
, by subsequent monotypy (
<bibRefCitation author="ICZN &amp; London, XXIX" box="[931,1052,496,520]" pageId="3" pageNumber="404" refId="ref15048" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. The International Trust for Zoological Nomenclature, London, XXIX + 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
: Article 69.3)
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="404" type="diagnosis">
<paragraph blockId="3.[151,1437,565,2032]" pageId="3" pageNumber="404">
<emphasis bold="true" box="[151,274,565,591]" pageId="3" pageNumber="404">Diagnosis.</emphasis>
Head elongated, with lateral post-ocular projections; proportion of labiomeres: I&lt;II&gt;III&lt;IV; pronotal depression anterior to the cicatrices forming a collar; humeral angles laterally projected, bifid, with the anterior portion longer than the posterior; pro-, meso- and metasternum covered by small setae; prosternum flat, mesosternum with a low longitudinal carina dilated anteriorly and posteriorly, metasternum slightly elevated with a shallow longitudinal groove.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="404" type="description">
<paragraph blockId="3.[151,1437,565,2032]" box="[199,365,745,771]" pageId="3" pageNumber="404">
<emphasis bold="true" box="[199,365,745,771]" pageId="3" pageNumber="404">Redescription</emphasis>
</paragraph>
<paragraph blockId="3.[151,1437,565,2032]" box="[199,788,782,808]" pageId="3" pageNumber="404">BODY: Elongated oval, with variable color patterns.</paragraph>
<paragraph blockId="3.[151,1437,565,2032]" pageId="3" pageNumber="404">HEAD: Rectangular, longer than wide, longitudinally convex, slightly punctured; mandibular plates longer than clypeus, with the anterior margin rounded; bucculae low, rounded at apex and evanescent posteriorly; eyes separated from the anterolateral angles of pronotum by at least half the eye diameter; post-ocular lateral triangular projection contiguous to the eyes; antennomere I shorter and wider than II, which is thinner than the others; antennomeres III, IV and V similar in length and more setose than the others.</paragraph>
<paragraph blockId="3.[151,1437,565,2032]" pageId="3" pageNumber="404">THORAX: Pronotum trapezoidal, uniformly and deeply punctured, twice as wide as long; scutellum uniform and shallowly punctured; frenal constriction width more than half the basal width, postfrenal lobe with parallel margins, broadly rounded at apex; coria with posterior margin sinuous; mesopleural evaporatoria occupying the entire posterior margin and the anterolateral mesopleural angle; metapleural evaporatoria surrounding the disc-shaped peritreme, occupying more than half of the metapleural width; ostiole guttiform in ventral view, laterally oriented; peritremal furrow extending more than half the length of the peritreme; dorsal and ventral foretibial expansions present, the dorsal expansion wider than the ventral.</paragraph>
<paragraph blockId="3.[151,1437,565,2032]" pageId="3" pageNumber="404">ABDOMEN: Uniformly punctured; mid tubercle on urosternite III; posterior angles of urosternites III to VII ending in minute spine; male bearing ventrolateral glandular setose patches, extending from near the anterior margin of urosternite V to the anterior margin of urosternite VII.</paragraph>
<paragraph blockId="3.[151,1437,565,2032]" box="[199,260,1358,1384]" pageId="3" pageNumber="404">
<emphasis bold="true" box="[199,260,1358,1384]" pageId="3" pageNumber="404">Male</emphasis>
</paragraph>
<paragraph blockId="3.[151,1437,565,2032]" pageId="3" pageNumber="404">GENITALIA: Pygophore bowl-shaped, with setae on the entire surface, ventral rim slightly sinuous; inferior layer excavated; posterolateral angles rounded; segment X trapezoidal, with rounded apex, ventrally directed, dorsally sclerotized, setose on apex; pseudoclaspers concave, sculptured, each placed between the paramere and the lateral portion of the dorsal rim; parameres sculptured, head triangular, spatulate, occupying in dorsal view most of the space between the segment X and the posterolateral angles. Phallus: phallotheca divided in a basal theca and thecal shield; ejaculatory reservoir contained inside basal theca; basal foramen elliptical, reinforced by basal plates; thecal shield longer and wider than basal theca; vesica oval in dorsal and lateral view; ductus seminis with uniform diameter, running between filaments of vesica, ending on a secondary gonopore, ventrally directed; conjuntiva with: two ventral lobes with sclerotized apex, and one dorsal lobe; all lobes posteriorly directed.</paragraph>
<paragraph blockId="3.[151,1437,565,2032]" box="[199,286,1718,1744]" pageId="3" pageNumber="404">
<emphasis bold="true" box="[199,286,1718,1744]" pageId="3" pageNumber="404">Female</emphasis>
</paragraph>
<paragraph blockId="3.[151,1437,565,2032]" pageId="3" pageNumber="404">GENITALIA: Valvifers VIII longer than wide, subtriangular, posterior margins sinuous, sutural margins straight, juxtaposed; laterotergites VIII subtriangular; exposed portion of valvifer IX trapezoidal, wider than long, juxtaposed to the laterotergites IX; exposed portion of laterotergites IX digitiform, attaining the mediotergite VIII; segment X trapezoidal with posterior margin straight; thickening of vaginal intima elongated; pars intermedialis small, narrower than the median duct of vesicular area; capsula seminalis longer and wider than pars intermedialis.</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="404" type="distribution">
<paragraph blockId="3.[151,1437,565,2032]" pageId="3" pageNumber="404">
<emphasis bold="true" box="[199,355,1933,1959]" pageId="3" pageNumber="404">Distribution:</emphasis>
<collectingCountry box="[366,662,1934,1960]" name="United States of America" pageId="3" pageNumber="404">United States of America</collectingCountry>
(
<collectingRegion box="[682,752,1934,1960]" country="United States of America" name="Texas" pageId="3" pageNumber="404">Texas</collectingRegion>
),
<collectingCountry box="[777,865,1934,1960]" name="Mexico" pageId="3" pageNumber="404">Mexico</collectingCountry>
,
<collectingCountry box="[881,1005,1934,1960]" name="Guatemala" pageId="3" pageNumber="404">Guatemala</collectingCountry>
,
<collectingCountry box="[1021,1133,1934,1960]" name="Honduras" pageId="3" pageNumber="404">Honduras</collectingCountry>
,
<collectingCountry box="[1149,1286,1934,1960]" name="El Salvador" pageId="3" pageNumber="404">El Salvador</collectingCountry>
,
<collectingCountry box="[1302,1432,1934,1960]" name="Costa Rica" pageId="3" pageNumber="404">Costa Rica</collectingCountry>
,
<collectingCountry box="[151,243,1970,1996]" name="Panama" pageId="3" pageNumber="404">Panama</collectingCountry>
,
<collectingCountry box="[257,370,1970,1996]" name="Colombia" pageId="3" pageNumber="404">Colombia</collectingCountry>
,
<collectingCountry box="[384,477,1970,1996]" name="Ecuador" pageId="3" pageNumber="404">Ecuador</collectingCountry>
(new record),
<collectingCountry box="[651,892,1970,1996]" name="Trinidad and Tobago" pageId="3" pageNumber="404">Trinidad and Tobago</collectingCountry>
,
<collectingCountry box="[906,1024,1970,1996]" name="Venezuela" pageId="3" pageNumber="404">Venezuela</collectingCountry>
,
<collectingCountry box="[1038,1128,1970,1996]" name="Guyana" pageId="3" pageNumber="404">Guyana</collectingCountry>
,
<collectingCountry box="[1142,1240,1970,1996]" name="Suriname" pageId="3" pageNumber="404">Surinam</collectingCountry>
, French
<collectingCountry box="[1342,1432,1970,1996]" name="Guyana" pageId="3" pageNumber="404">Guyana</collectingCountry>
,
<collectingCountry box="[151,222,2006,2032]" name="Brazil" pageId="3" pageNumber="404">Brazil</collectingCountry>
,
<collectingCountry box="[234,288,2006,2032]" name="Peru" pageId="3" pageNumber="404">Peru</collectingCountry>
,
<collectingCountry box="[299,384,2006,2032]" name="Bolivia" pageId="3" pageNumber="404">Bolivia</collectingCountry>
,
<collectingCountry box="[395,499,2006,2032]" name="Paraguay" pageId="3" pageNumber="404">Paraguay</collectingCountry>
and
<collectingCountry box="[552,663,2006,2032]" name="Argentina" pageId="3" pageNumber="404">Argentina</collectingCountry>
(
<figureCitation box="[679,745,2006,2032]" captionStart="FIGURE 1" captionStartId="4.[152,255,1061,1085]" captionTargetBox="[186,1402,187,1031]" captionTargetId="figure-26@4.[177,1410,181,1037]" captionTargetPageId="4" captionText="FIGURE 1. Geographic distribution of Heteroscelis Latreille, 1829. The countries highlighted for each species on the small maps are based on literature data. The points marked for each species are from the specimens examined in this paper, and from records retrieved from iNaturalist." figureDoi="http://doi.org/10.5281/zenodo.7906377" httpUri="https://zenodo.org/record/7906377/files/figure.png" pageId="3" pageNumber="404">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>