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<document id="02D72B8E9C33E95D99C4867918B09771" ID="10.11646/zootaxa.4144.2.8" ID-DOI="10.11646/zootaxa.4144.2.8" ID-GBIF-Dataset="39648a3c-8d3a-4d55-b613-1b56452667eb" ID-ISSN="1175-5326" ID-Zenodo-Dep="261230" ID-ZooBank="5FEA38D2-96BA-496A-9EC7-C5C020F39D5B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1469603138887" checkinUser="plazi" docAuthor="Ning, Xin, Kompier, Tom, Yu, Xin &amp; Bu, Wenjun" docDate="2016" docId="03BA6003A64C0E4BFF12F4E94AF945E3" docLanguage="en" docName="zootaxa.4144.2.8.pdf" docOrigin="Zootaxa 4144 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paracercion ambiguum Kompier &amp; Yu, sp. nov." docType="treatment" docVersion="8" lastPageNumber="273" masterDocId="FF83187BA6490E40FF85F4354B3C451B" masterDocTitle="Paracercion ambiguum sp. nov. from Lang Son, Vietnam (Zygoptera: Coenagrionidae)" masterLastPageNumber="275" masterPageNumber="263" pageNumber="268" updateTime="1698366315559" updateUser="plazi">
<mods:mods id="B30FFE202DFD29BC7A715D8D6BF1B582" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0D036FC33DB4A13263EB46FEFAF8150B">
<mods:title id="F7A1870ADFF97C6C89C67096F957F345">Paracercion ambiguum sp. nov. from Lang Son, Vietnam (Zygoptera: Coenagrionidae)</mods:title>
</mods:titleInfo>
<mods:name id="A80CE357934FF7FE52CD7235ADE77EAC" type="personal">
<mods:role id="E2EF54DE20E6E4D90DA75C58757FBF9E">
<mods:roleTerm id="471EB9937891EECFE7EC050A49A34982">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="52127A18484500AAEA012F9506130624">Ning, Xin</mods:namePart>
</mods:name>
<mods:name id="6C68762D0EA24F98EB4EA37E5B663F12" type="personal">
<mods:role id="5171EC520F5A4DEF8BBC87A23815E331">
<mods:roleTerm id="9275EC88DF690386407D5E8E74A1E8E1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CB423268368E102C6B8AD150BACE9754">Kompier, Tom</mods:namePart>
</mods:name>
<mods:name id="5B17D7005262EAFA46650E68F30073FD" type="personal">
<mods:role id="1A1E27247F00C7842206E2A1FE3B8B33">
<mods:roleTerm id="90E70F32482CE8291EFDD9DE266FCD70">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="ACAEF9676D4D0B090A2FA64079E444CA">Yu, Xin</mods:namePart>
</mods:name>
<mods:name id="C858B04CF1C90D5A7BC8C41E3A11F88E" type="personal">
<mods:role id="2768D2DE8A86D72EAF30D1BAA12A6F12">
<mods:roleTerm id="AD63AC6538941FD0CC2D59AF8526E030">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B15FCE371862A1628280DF7CCA115506">Bu, Wenjun</mods:namePart>
</mods:name>
<mods:typeOfResource id="4ABB9DCD4BE6E053EA9DD5B128EC23CF">text</mods:typeOfResource>
<mods:relatedItem id="592B25DC6DA1C1CD9755C5988B707114" type="host">
<mods:titleInfo id="D7A6DD6F2974CA5AA58F5864206BA17E">
<mods:title id="BEE1C42B6F9E042C60682247579FF1EE">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D958272E35603AE6C15407EC2742BDD3">
<mods:date id="70209E3FDE1E70D8AD16D739589A55BD">2016</mods:date>
<mods:detail id="A4F3554736FD2FF04FB934621C57AAAD" type="volume">
<mods:number id="A3398231664C9EEC62C94CA1D28E7CB9">4144</mods:number>
</mods:detail>
<mods:detail id="8F0BFBBD4F4611847A82D823C58D2DDD" type="issue">
<mods:number id="3E5C41E2B1EC2093AFBBD50591E5D42B">2</mods:number>
</mods:detail>
<mods:extent id="55CDECC0BE0C35831DB40D63195FA8B4" unit="page">
<mods:start id="57E7634AE7BEDA79B413EADB29BE7AC9">263</mods:start>
<mods:end id="92ADEF1F8AD50F04CEBA52097379617B">275</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="9CC4BF475220575BDE4740DC07DF7F7D">journal article</mods:classification>
<mods:identifier id="40B12FBB66723A572D56B2C759404FC5" type="DOI">10.11646/zootaxa.4144.2.8</mods:identifier>
<mods:identifier id="49DAB7F620D368C377E3DE29A3189B90" type="GBIF-Dataset">39648a3c-8d3a-4d55-b613-1b56452667eb</mods:identifier>
<mods:identifier id="A44E36FD5A45D6B19B6A367FB350ECAE" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A590EF8A7B05A5E21F94BF2949180933" type="Zenodo-Dep">261230</mods:identifier>
<mods:identifier id="21D26B561A3FDE7BB89722E4A0EBC7F6" type="ZooBank">5FEA38D2-96BA-496A-9EC7-C5C020F39D5B</mods:identifier>
</mods:mods>
<treatment id="03BA6003A64C0E4BFF12F4E94AF945E3" ID-DOI="http://doi.org/10.5281/zenodo.5678525" ID-GBIF-Taxon="123207770" ID-Zenodo-Dep="5678525" LSID="urn:lsid:plazi:treatment:03BA6003A64C0E4BFF12F4E94AF945E3" httpUri="http://treatment.plazi.org/id/03BA6003A64C0E4BFF12F4E94AF945E3" lastPageId="11" lastPageNumber="273" pageId="5" pageNumber="268">
<subSubSection id="C309829EA64C0E45FF12F4E949E745EC" box="[151,731,220,247]" pageId="5" pageNumber="268" type="nomenclature">
<paragraph id="8BACD115A64C0E45FF12F4E949E745EC" blockId="5.[151,731,220,247]" box="[151,731,220,247]" pageId="5" pageNumber="268">
<heading id="D0E46679A64C0E45FF12F4E949E745EC" bold="true" box="[151,731,220,247]" fontSize="11" level="1" pageId="5" pageNumber="268" reason="1">
<emphasis id="B9670D07A64C0E45FF12F4E949E745EC" bold="true" box="[151,731,220,247]" pageId="5" pageNumber="268">
<taxonomicName id="4C13AA96A64C0E45FF12F4E9494A45EC" ID-CoL="9WVP4" authority="Kompier &amp; Yu" authorityName="Kompier &amp; Yu" box="[151,630,220,247]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="268" phylum="Arthropoda" rank="species" species="ambiguum" status="sp. nov.">
<emphasis id="B9670D07A64C0E45FF12F4E94A8845ED" bold="true" box="[151,436,220,246]" italics="true" pageId="5" pageNumber="268">Paracercion ambiguum</emphasis>
Kompier &amp; Yu
</taxonomicName>
<taxonomicNameLabel id="A254B07CA64C0E45FDF8F4E849E745EC" box="[637,731,221,247]" pageId="5" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C309829EA64C0E45FF12F5114AFE44D6" pageId="5" pageNumber="268" type="materials_examined">
<paragraph id="8BACD115A64C0E45FF12F5114AFE44D6" blockId="5.[151,1437,292,784]" pageId="5" pageNumber="268">
<materialsCitation id="3B7BDB48A64C0E45FF12F5114951447B" ID-GBIF-Occurrence="1288102700" collectingDate="2013-12-01" country="Vietnam" location="Tom Kompier" municipality="Huu Lien Nature Reserve" pageId="5" pageNumber="268" specimenCount="1" stateProvince="Lang Son" typeStatus="holotype">
<emphasis id="B9670D07A64C0E45FF12F5114AB54426" bold="true" box="[151,393,292,317]" pageId="5" pageNumber="268">
Material.
<typeStatus id="54A86FB7A64C0E45FE96F5114AB84426" box="[275,388,292,317]" pageId="5" pageNumber="268" type="holotype">Holotype</typeStatus>
:
</emphasis>
male (GenBank accession number:
<accessionNumber id="94404CF6A64C0E45FCBBF51148834426" box="[830,959,292,317]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KX263718" pageId="5" pageNumber="268">KX263718</accessionNumber>
),
<collectingCountry id="F3049185A64C0E45FC5DF5114F064426" box="[984,1082,292,317]" name="Vietnam" pageId="5" pageNumber="268">Vietnam</collectingCountry>
,
<collectingRegion id="49D71FF7A64C0E45FBCEF5104FFC4427" box="[1099,1216,292,317]" country="Vietnam" name="Lang Son" pageId="5" pageNumber="268">Lang Son</collectingRegion>
,
<collectingMunicipality id="6BC84B6FA64C0E45FB57F5104BC84444" pageId="5" pageNumber="268">Huu Lien Nature Reserve</collectingMunicipality>
,
<collectingDate id="EFE90E3DA64C0E45FE80F57D4ABB447B" box="[261,391,328,352]" pageId="5" pageNumber="268" value="2013-12-01">
<date id="FFADF7D5A64C0E45FE80F57D4ABB447B" box="[261,391,328,352]" pageId="5" pageNumber="268" value="2013-12-01">1-XII-2013</date>
</collectingDate>
,
<location id="8ECC87CEA64C0E45FE12F57D4907447B" LSID="urn:lsid:plazi:treatment:03BA6003A64C0E4BFF12F4E94AF945E3:8ECC87CEA64C0E45FE12F57D4907447B" box="[407,571,327,352]" country="Vietnam" municipality="Huu Lien Nature Reserve" name="Tom Kompier" pageId="5" pageNumber="268" stateProvince="Lang Son">Tom Kompier</location>
leg
</materialsCitation>
.
<emphasis id="B9670D07A64C0E45FDF8F57249C2447B" bold="true" box="[637,766,327,352]" pageId="5" pageNumber="268">Paratypes:</emphasis>
1 male (GenBank accession number:
<accessionNumber id="94404CF6A64C0E45FB4CF5724E76447B" box="[1225,1354,327,352]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KX263719" pageId="5" pageNumber="268">KX263719</accessionNumber>
), same information but
<date id="FFADF7D5A64C0E45FECAF5594AEA449F" box="[335,470,364,388]" pageId="5" pageNumber="268" value="2013-11-23">23-XI-2013</date>
;
<materialsCitation id="3B7BDB48A64C0E45FE66F5594B8144B3" ID-GBIF-Occurrence="1288102702" collectingDate="2013-12-07" country="Vietnam" location="Huu Lien Nature Reserve" pageId="5" pageNumber="268" specimenCount="1" specimenCount-female="1" stateProvince="Lang Son" typeStatus="holotype">
<specimenCount id="9D151A9CA64C0E45FE66F5594974449E" box="[483,584,364,389]" pageId="5" pageNumber="268" type="female">1 female</specimenCount>
,
<collectingCountry id="F3049185A64C0E45FDD1F5594989449E" box="[596,693,364,389]" name="Vietnam" pageId="5" pageNumber="268">Vietnam</collectingCountry>
,
<collectingRegion id="49D71FF7A64C0E45FD47F558480D449F" box="[706,817,364,389]" country="Vietnam" name="Lang Son" pageId="5" pageNumber="268">Lang Son</collectingRegion>
,
<location id="8ECC87CEA64C0E45FCBBF5584F5F449F" LSID="urn:lsid:plazi:treatment:03BA6003A64C0E4BFF12F4E94AF945E3:8ECC87CEA64C0E45FCBBF5584F5F449F" box="[830,1123,364,389]" country="Vietnam" name="Huu Lien Nature Reserve" pageId="5" pageNumber="268" stateProvince="Lang Son">Huu Lien Nature Reserve</location>
,
<collectingDate id="EFE90E3DA64C0E45FBEAF5594FCE449F" box="[1135,1266,364,388]" pageId="5" pageNumber="268" value="2013-12-07">
<date id="FFADF7D5A64C0E45FBEAF5594FCE449F" box="[1135,1266,364,388]" pageId="5" pageNumber="268" value="2013-12-07">7-XII-2013</date>
</collectingDate>
, Tom Kompier leg
</materialsCitation>
.
<materialsCitation id="3B7BDB48A64C0E45FF48F5BA4AFE44D6" ID-GBIF-Occurrence="1288102701" country="China" county="Institute" location="Life Sciences College" municipality="Entomology" pageId="5" pageNumber="268" specimenCount="2" typeStatus="holotype">
<typeStatus id="54A86FB7A64C0E45FF48F5BA4A0944B3" box="[205,309,399,424]" pageId="5" pageNumber="268" type="holotype">Holotype</typeStatus>
and
<typeStatus id="54A86FB7A64C0E45FEFDF5A54ADF44B3" box="[376,483,400,424]" pageId="5" pageNumber="268" type="paratype">paratypes</typeStatus>
will be deposited in
<collectingCounty id="62CDA999A64C0E45FD68F5BA487544B3" box="[749,841,399,424]" pageId="5" pageNumber="268">Institute</collectingCounty>
of
<collectingMunicipality id="6BC84B6FA64C0E45FCFFF5BA4F3A44B3" box="[890,1030,399,424]" pageId="5" pageNumber="268">Entomology</collectingMunicipality>
,
<location id="8ECC87CEA64C0E45FB92F5BA4E2544B3" LSID="urn:lsid:plazi:treatment:03BA6003A64C0E4BFF12F4E94AF945E3:8ECC87CEA64C0E45FB92F5BA4E2544B3" box="[1047,1305,399,424]" country="China" county="Institute" municipality="Entomology" name="Life Sciences College" pageId="5" pageNumber="268">Life Sciences College</location>
of
<location id="8ECC87CEA64C0E45FACEF5BA4A3344D6" LSID="urn:lsid:plazi:treatment:03BA6003A64C0E4BFF12F4E94AF945E3:8ECC87CEA64C0E45FACEF5BA4A3344D6" country="China" county="Institute" municipality="Entomology" name="Nankai University" pageId="5" pageNumber="268">Nankai University</location>
,
<collectingCountry id="F3049185A64C0E45FE9CF5814A5044D6" box="[281,364,436,461]" name="China" pageId="5" pageNumber="268">Tianjin</collectingCountry>
,
<collectingCountry id="F3049185A64C0E45FEF2F5814A8244D6" box="[375,446,436,461]" name="China" pageId="5" pageNumber="268">China</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C309829EA64C0E45FF42F5E249E0470E" pageId="5" pageNumber="268" type="etymology">
<paragraph id="8BACD115A64C0E45FF42F5E249E0470E" blockId="5.[151,1437,292,784]" pageId="5" pageNumber="268">
<emphasis id="B9670D07A64C0E45FF42F5E24A7244EB" bold="true" box="[199,334,471,496]" pageId="5" pageNumber="268">Etymology.</emphasis>
The Latin word
<taxonomicName id="4C13AA96A64C0E45FDA4F5ED49AB44EB" box="[545,663,472,496]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="268" phylum="Arthropoda" rank="species" species="ambiguum">
<emphasis id="B9670D07A64C0E45FDA4F5ED49AB44EB" box="[545,663,472,496]" italics="true" pageId="5" pageNumber="268">ambiguum</emphasis>
</taxonomicName>
means puzzling or doubtful, which emphasizes the difficulty on deciding the taxonomic position of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C309829EA64C0E49FF42F62A4BF44245" lastPageId="9" lastPageNumber="272" pageId="5" pageNumber="268" type="description">
<paragraph id="8BACD115A64C0E45FF42F62A4FE5460B" blockId="5.[151,1437,292,784]" pageId="5" pageNumber="268">
<emphasis id="B9670D07A64C0E45FF42F62A4AEA4723" bold="true" box="[199,470,543,568]" pageId="5" pageNumber="268">
Description.
<typeStatus id="54A86FB7A64C0E45FEE7F62A4AED4723" box="[354,465,543,568]" pageId="5" pageNumber="268" type="holotype">Holotype</typeStatus>
.
</emphasis>
Labium cream-white. Bases of mandibles and labrum light blue, labrum with black dorsolateral corners and central black spot on upper margin. Anteclypeus light blue, postclypeus black with light blue anterior edge and two elongated light-blue spots on posterior edge. Antefrons light blue with black anterior edge, postfrons light blue with large central black spot, narrowly connected with black to cranium. Antennal base and scape light blue, remainder of antenna black. Top of head black with two light-blue lines extending from postclypeus onto it near base of antennae. Postocular spots large, triangular, blue color. No occipital bar. Compound eyes blackish dorsally, greenish blue anteriorly and pale blue ventrally in life (
<figureCitation id="1328CD90A64C0E45FB02F6C24FF7460B" box="[1159,1227,759,784]" captionStart="FIGURE 8" captionStartId="9.[151,250,1764,1786]" captionTargetBox="[195,1418,1046,1721]" captionTargetId="figure@9.[167,1419,1046,1742]" captionTargetPageId="9" captionText="FIGURE 8. Face of male Paracercion ambiguum." httpUri="https://zenodo.org/record/261237/files/figure.png" pageId="5" pageNumber="268">Fig. 8</figureCitation>
).
</paragraph>
<caption id="DF6C819DA64C0E45FF12F3AE489642AA" box="[151,938,1947,1969]" httpUri="https://zenodo.org/record/261232/files/figure.png" pageId="5" pageNumber="268" targetBox="[151,1325,915,1866]" targetPageId="5">
<paragraph id="8BACD115A64C0E45FF12F3AE489642AA" blockId="5.[151,938,1947,1969]" box="[151,938,1947,1969]" pageId="5" pageNumber="268">
<emphasis id="B9670D07A64C0E45FF12F3AE4A2E42AB" bold="true" box="[151,274,1947,1969]" pageId="5" pageNumber="268">FIGURE 2.</emphasis>
Male caudal appendages of
<taxonomicName id="4C13AA96A64C0E45FDB6F3AE481E42AB" box="[563,802,1947,1968]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="268" phylum="Arthropoda" rank="species" species="ambiguum">
<emphasis id="B9670D07A64C0E45FDB6F3AE481E42AB" box="[563,802,1947,1968]" italics="true" pageId="5" pageNumber="268">Paracercion ambiguum</emphasis>
</taxonomicName>
, dorsal view.
</paragraph>
</caption>
<caption id="DF6C819DA64F0E46FF12F17D48974045" box="[151,939,1352,1374]" httpUri="https://zenodo.org/record/261233/files/figure.png" pageId="6" pageNumber="269" targetBox="[151,1363,193,1271]" targetPageId="6">
<paragraph id="8BACD115A64F0E46FF12F17D48974045" blockId="6.[151,939,1352,1374]" box="[151,939,1352,1374]" pageId="6" pageNumber="269">
<emphasis id="B9670D07A64F0E46FF12F17D4A2E4046" bold="true" box="[151,274,1352,1374]" pageId="6" pageNumber="269">FIGURE 3.</emphasis>
Male caudal appendages of
<taxonomicName id="4C13AA96A64F0E46FDB6F17C481E4046" box="[563,802,1352,1374]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="269" phylum="Arthropoda" rank="species" species="ambiguum">
<emphasis id="B9670D07A64F0E46FDB6F17C481E4046" box="[563,802,1352,1374]" italics="true" pageId="6" pageNumber="269">Paracercion ambiguum</emphasis>
</taxonomicName>
, lateral view.
</paragraph>
</caption>
<paragraph id="8BACD115A64F0E46FF42F1B0491F4381" blockId="6.[151,1437,1413,1978]" pageId="6" pageNumber="269">
Prothorax black with light-blue lateral edges. Anterior lobe largely light blue, lateral lobes light-blue with interior edge black, posterior lobe black with light-blue posterior edge. Mesostigmal lamina black with light-blue lateral edges. Synthorax black dorsally with broad light-blue antehumeral stripe. Laterally blue with black markings as follows: broad black stripe over humeral suture; distinct black stripe over first lateral suture from posterior end extending almost to midpoint, arrow-shaped and broadest at anterior end; thin black line over second lateral suture that expands into half-oval at posterior end forming black spot. Legs cream-white, with blackish extensor surfaces of femora and blackish spines. Wings hyaline, with 10–11 postnodals and brown pterostigma covering one cell in length (
<figureCitation id="1328CD90A64F0E46FE54F2B449294381" box="[465,533,1665,1690]" captionStart="FIGURE 9" captionStartId="10.[151,250,1643,1665]" captionTargetBox="[151,1435,733,1621]" captionTargetId="figure@10.[151,1435,733,1621]" captionTargetPageId="10" captionText="FIGURE 9. Field photo of male Paracercion ambiguum." httpUri="https://zenodo.org/record/261238/files/figure.png" pageId="6" pageNumber="269">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="8BACD115A64F0E46FF42F29049364269" blockId="6.[151,1437,1413,1978]" pageId="6" pageNumber="269">
Abdomen S1 light-blue with black dorsum, black narrowest just before posterior end; S2–4 black dorsally and blue laterally, end of S3–4 almost wholly black; S5–7 black and S8–9 blue; S10 blue with basal and distal margins black dorsally; intersegment rings covered with small black spines. Caudal appendages black. Cerci a little longer than S10 and three times length of paraprocts, apically dilated and with large basal tooth. Paraproct slanting upwards at 45 degree angle, simple in lateral view (
<figureCitation id="1328CD90A64F0E46FD5CF300487B4255" box="[729,839,1845,1870]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="5.[151,250,1947,1969]" captionStartId-1="6.[151,250,1352,1374]" captionStartId-2="7.[151,250,1377,1399]" captionTargetBox-0="[151,1325,915,1866]" captionTargetBox-1="[151,1363,193,1271]" captionTargetBox-2="[257,1236,193,1346]" captionTargetId-0="figure@5.[151,1436,837,1925]" captionTargetId-1="figure@6.[151,1435,193,1331]" captionTargetId-2="figure@7.[257,1330,193,1355]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURE 2. Male caudal appendages of Paracercion ambiguum, dorsal view." captionText-1="FIGURE 3. Male caudal appendages of Paracercion ambiguum, lateral view." captionText-2="FIGURE 4. Inner view of the right cercus and paraproct of Paracercion ambiguum, with left cercus and paraproct removed." httpUri-0="https://zenodo.org/record/261232/files/figure.png" httpUri-1="https://zenodo.org/record/261233/files/figure.png" httpUri-2="https://zenodo.org/record/261234/files/figure.png" pageId="6" pageNumber="269">Figs. 2–4</figureCitation>
). Head of genital ligula shield-like, one pair of apical lobes broad and long (
<figureCitation id="1328CD90A64F0E46FE15F36C4AC14269" box="[400,509,1881,1906]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="8.[151,250,1993,2015]" captionStartId-1="9.[151,250,974,996]" captionTargetBox-0="[176,1435,196,1917]" captionTargetBox-1="[178,1436,214,953]" captionTargetId-0="figure@8.[151,1435,1023,1971]" captionTargetId-1="figure@9.[150,1437,193,953]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 6. Male genital ligula of Paracercion ambiguum, ventral view." captionText-1="FIGURE 7. Male genital ligula of Paracercion ambiguum, lateral view." httpUri-0="https://zenodo.org/record/261235/files/figure.png" httpUri-1="https://zenodo.org/record/261236/files/figure.png" pageId="6" pageNumber="269">Figs. 6–7</figureCitation>
).
</paragraph>
<paragraph id="8BACD115A64F0E46FF42F34848CB42A1" blockId="6.[151,1437,1413,1978]" pageId="6" pageNumber="269">
<emphasis id="B9670D07A64F0E46FF42F3484AF3428D" bold="true" box="[199,463,1917,1942]" pageId="6" pageNumber="269">
Variation in
<typeStatus id="54A86FB7A64F0E46FED9F3484AF6428D" box="[348,458,1917,1942]" pageId="6" pageNumber="269" type="paratype">paratype</typeStatus>
.
</emphasis>
The
<typeStatus id="54A86FB7A64F0E46FD82F34A495B428C" box="[519,615,1919,1943]" pageId="6" pageNumber="269" type="paratype">paratype</typeStatus>
male is extremely similar to the
<typeStatus id="54A86FB7A64F0E46FC50F3484F0B428D" box="[981,1079,1917,1942]" pageId="6" pageNumber="269" type="holotype">holotype</typeStatus>
male. However, the wings have 11–12 postnodals and S10 is decorated with an I-shaped black mark dorsally.
</paragraph>
<caption id="DF6C819DA64E0E47FF12F1544EB8406C" box="[151,1412,1377,1399]" httpUri="https://zenodo.org/record/261234/files/figure.png" pageId="7" pageNumber="270" targetBox="[257,1236,193,1346]" targetPageId="7">
<paragraph id="8BACD115A64E0E47FF12F1544EB8406C" blockId="7.[151,1412,1377,1399]" box="[151,1412,1377,1399]" pageId="7" pageNumber="270">
<emphasis id="B9670D07A64E0E47FF12F1544A2E406D" bold="true" box="[151,274,1377,1399]" pageId="7" pageNumber="270">FIGURE 4.</emphasis>
Inner view of the right cercus and paraproct of
<taxonomicName id="4C13AA96A64E0E47FD73F15748D9406D" box="[758,997,1377,1399]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="270" phylum="Arthropoda" rank="species" species="ambiguum">
<emphasis id="B9670D07A64E0E47FD73F15748D9406D" box="[758,997,1377,1399]" italics="true" pageId="7" pageNumber="270">Paracercion ambiguum</emphasis>
</taxonomicName>
, with left cercus and paraproct removed.
</paragraph>
</caption>
<paragraph id="8BACD115A64E0E47FF42F1AB4AD542F5" blockId="7.[151,1437,1438,2030]" pageId="7" pageNumber="270">
Female. Facial pattern identical to male, but postocular spots somewhat smaller and light-blue paler and somewhat greenish. Eyes brown dorsally and green ventrally. Prothorax similar in pattern to male, but central lobe with two pale longitudinal lines along center and color of light parts of central lobe, lateral lobes and posterior lobe mixture of pale reddish brown and greenish blue, posterior lobe without horns, its hind ridge bow-shaped, straight centrally, then slightly curving forward and towards the lateral sides slightly curving caudad again. Anterior lobe largely light blue as in male. Synthorax with basic pattern as male, but light-blue more greenish, especially towards dorsum. Dorsum more brownish and with thin paler lines along dorsal carina. Black antehumeral stripe with two pale brown lines nested in, one on either side of humeral suture, as in several other
<taxonomicName id="4C13AA96A64E0E47FBD0F2A24FDC43B5" box="[1109,1248,1687,1710]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="270" phylum="Arthropoda" rank="genus">
<emphasis id="B9670D07A64E0E47FBD0F2A24FDC43B5" box="[1109,1248,1687,1710]" italics="true" pageId="7" pageNumber="270">Paracercion</emphasis>
</taxonomicName>
species. Arrowshaped line along first lateral suture distinct as in male, but thin line over second lateral suture absent. Only halfoval spot towards posterior end of second lateral suture present. Legs and wings as male (
<figureCitation id="1328CD90A64E0E47FB07F2E84FE943ED" box="[1154,1237,1757,1782]" captionStart="FIGURE 10" captionStartId="11.[151,250,1178,1200]" captionTargetBox="[150,1435,303,1155]" captionTargetId="figure@11.[150,1436,303,1156]" captionTargetPageId="11" captionText="FIGURE 10. Head and thorax of female Paracercion ambiguum in life." httpUri="https://zenodo.org/record/261239/files/figure.png" pageId="7" pageNumber="270">Fig. 10</figureCitation>
). The abdomen is thicker than that of the male, giving a more robust appearance. Abdominal segments S1–7 pale blue laterally, progressively less distinct towards S7, and blackish along dorsum. Black widest at posterior end of segment along intersegmental ring, forming incomplete rings, and blue extending towards dorsum at anterior end of segment along intersegmental ring. S8 almost entirely black, but with clear blue lateral markings, especially towards posterior edge. S9 blue with small black markings on dorsum at anterior end, S10 all blue. Cerci whitish, two-thirds the length of S10 and slanting downwards, simple conical. Paraproct whitish. Ovipositor reaching just past middle of S10, cream-white (
<figureCitation id="1328CD90A64E0E47FE0FF3E04AE742F5" box="[394,475,2005,2030]" captionStart="FIGURE 11" captionStartId="11.[151,250,1887,1909]" captionTargetBox="[151,1435,1250,1865]" captionTargetId="figure@11.[151,1436,1250,1866]" captionTargetPageId="11" captionText="FIGURE 11. Field photo of female Paracercion ambiguum." httpUri="https://zenodo.org/record/261240/files/figure.png" pageId="7" pageNumber="270">Fig. 11</figureCitation>
).
</paragraph>
<caption id="DF6C819DA6410E48FF12F3FC484542C4" box="[151,889,1993,2015]" httpUri="https://zenodo.org/record/261235/files/figure.png" pageId="8" pageNumber="271" targetBox="[176,1435,196,1917]" targetPageId="8">
<paragraph id="8BACD115A6410E48FF12F3FC484542C4" blockId="8.[151,889,1993,2015]" box="[151,889,1993,2015]" pageId="8" pageNumber="271">
<emphasis id="B9670D07A6410E48FF12F3FC4A2E42C5" bold="true" box="[151,274,1993,2015]" pageId="8" pageNumber="271">FIGURE 6.</emphasis>
Male genital ligula of
<taxonomicName id="4C13AA96A6410E48FE7FF3FF49D542C5" box="[506,745,1993,2015]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="271" phylum="Arthropoda" rank="species" species="ambiguum">
<emphasis id="B9670D07A6410E48FE7FF3FF49D542C5" box="[506,745,1993,2015]" italics="true" pageId="8" pageNumber="271">Paracercion ambiguum</emphasis>
</taxonomicName>
, ventral view.
</paragraph>
</caption>
<caption id="DF6C819DA6400E49FF12F7FB484E46FF" box="[151,882,974,996]" httpUri="https://zenodo.org/record/261236/files/figure.png" pageId="9" pageNumber="272" targetBox="[178,1436,214,953]" targetPageId="9">
<paragraph id="8BACD115A6400E49FF12F7FB484E46FF" blockId="9.[151,882,974,996]" box="[151,882,974,996]" pageId="9" pageNumber="272">
<emphasis id="B9670D07A6400E49FF12F7FB4A2E46FF" bold="true" box="[151,274,974,996]" pageId="9" pageNumber="272">FIGURE 7.</emphasis>
Male genital ligula of
<taxonomicName id="4C13AA96A6400E49FE7FF7FA49D546FF" box="[506,745,975,996]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="272" phylum="Arthropoda" rank="species" species="ambiguum">
<emphasis id="B9670D07A6400E49FE7FF7FA49D546FF" box="[506,745,975,996]" italics="true" pageId="9" pageNumber="272">Paracercion ambiguum</emphasis>
</taxonomicName>
, lateral view.
</paragraph>
</caption>
<caption id="DF6C819DA6400E49FF12F2D149AB43E1" box="[151,663,1764,1786]" httpUri="https://zenodo.org/record/261237/files/figure.png" pageId="9" pageNumber="272" targetBox="[195,1418,1046,1721]" targetPageId="9">
<paragraph id="8BACD115A6400E49FF12F2D149AB43E1" blockId="9.[151,663,1764,1786]" box="[151,663,1764,1786]" pageId="9" pageNumber="272">
<emphasis id="B9670D07A6400E49FF12F2D14A2E43E2" bold="true" box="[151,274,1764,1786]" pageId="9" pageNumber="272">FIGURE 8.</emphasis>
Face of male
<taxonomicName id="4C13AA96A6400E49FE27F2D049AD43E2" box="[418,657,1764,1786]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="272" phylum="Arthropoda" rank="species" species="ambiguum">
<emphasis id="B9670D07A6400E49FE27F2D049AD43E2" box="[418,657,1764,1786]" italics="true" pageId="9" pageNumber="272">Paracercion ambiguum</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BACD115A6400E49FF42F3174BF44245" blockId="9.[151,1436,1825,2030]" pageId="9" pageNumber="272">Measurements (mm): male abdomen + appendages 30.5; hind wing 19.0. Female abdomen 33.0; hind wing 21.5</paragraph>
</subSubSection>
<subSubSection id="C309829EA6400E4AFF42F35C499947BC" lastPageId="10" lastPageNumber="273" pageId="9" pageNumber="272" type="diagnosis">
<paragraph id="8BACD115A6400E49FF42F35C4EA642F5" blockId="9.[151,1436,1825,2030]" pageId="9" pageNumber="272">
<emphasis id="B9670D07A6400E49FF42F35C4A7D4299" bold="true" box="[199,321,1897,1922]" pageId="9" pageNumber="272">Diagnosis.</emphasis>
<taxonomicName id="4C13AA96A6400E49FECAF35F4AD44299" box="[335,488,1898,1922]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="272" phylum="Arthropoda" rank="species" species="ambiguum">
<emphasis id="B9670D07A6400E49FECAF35F4AD44299" box="[335,488,1898,1922]" italics="true" pageId="9" pageNumber="272">P. ambiguum</emphasis>
</taxonomicName>
can be separated from
<taxonomicName id="4C13AA96A6400E49FC91F35F489F4299" box="[788,931,1898,1922]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="B9670D07A6400E49FC91F35F489F4299" box="[788,931,1898,1922]" italics="true" pageId="9" pageNumber="272">Pseudagrion</emphasis>
</taxonomicName>
species within whose range it occurs by characteristics of patterning (for instance, in the male a black line on the first lateral suture and in females the double lines bordering the humeral suture). The female also lacks the horns on the pronotum that are characteristic of
<taxonomicName id="4C13AA96A6400E49FF33F3E34A7942F5" box="[182,325,2006,2030]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="B9670D07A6400E49FF33F3E34A7942F5" box="[182,325,2006,2030]" italics="true" pageId="9" pageNumber="272">Pseudagrion</emphasis>
</taxonomicName>
females. The long cerci of the male separate it from all other
<taxonomicName id="4C13AA96A6400E49FB84F3E24FB042F5" box="[1025,1164,2007,2030]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="B9670D07A6400E49FB84F3E24FB042F5" box="[1025,1164,2007,2030]" italics="true" pageId="9" pageNumber="272">Paracercion</emphasis>
</taxonomicName>
species, but not from
<emphasis id="B9670D07A6400E49FA02F3E24EA642F5" box="[1415,1434,2007,2030]" italics="true" pageId="9" pageNumber="272">P.</emphasis>
</paragraph>
<paragraph id="8BACD115A6430E4AFF12F4AD499947BC" blockId="10.[151,1437,151,680]" pageId="10" pageNumber="273">
<taxonomicName id="4C13AA96A6430E4AFF12F4AD4BC745AB" box="[151,251,152,176]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="dorothea">
<emphasis id="B9670D07A6430E4AFF12F4AD4BC745AB" box="[151,251,152,176]" italics="true" pageId="10" pageNumber="273">dorothea</emphasis>
</taxonomicName>
and
<taxonomicName id="4C13AA96A6430E4AFEB7F4AC4AFD45AB" box="[306,449,152,176]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="plagiosum">
<emphasis id="B9670D07A6430E4AFEB7F4AC4AFD45AB" box="[306,449,152,176]" italics="true" pageId="10" pageNumber="273">P. plagiosum</emphasis>
</taxonomicName>
, which also have cerci at least as long as S10. However,
<taxonomicName id="4C13AA96A6430E4AFBC1F4AC4FE945AB" box="[1092,1237,152,176]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="ambiguum">
<emphasis id="B9670D07A6430E4AFBC1F4AC4FE945AB" box="[1092,1237,152,176]" italics="true" pageId="10" pageNumber="273">P. ambiguum</emphasis>
</taxonomicName>
differs from both in lateral view by having the ventral tooth at the base of the cerci where the base is so large that it extends twothirds of the length of the appendage. The tooth is only half as long as it is wide. In
<taxonomicName id="4C13AA96A6430E4AFBD6F4D44FE845E3" box="[1107,1236,224,248]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="dorothea">
<emphasis id="B9670D07A6430E4AFBD6F4D44FE845E3" box="[1107,1236,224,248]" italics="true" pageId="10" pageNumber="273">P. dorothea</emphasis>
</taxonomicName>
the basal tooth is long and slender and in
<taxonomicName id="4C13AA96A6430E4AFE30F53049754407" box="[437,585,260,284]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="plagiosum">
<emphasis id="B9670D07A6430E4AFE30F53049754407" box="[437,585,260,284]" italics="true" pageId="10" pageNumber="273">P. plagiosum</emphasis>
</taxonomicName>
it is similarly longer than wide. In
<taxonomicName id="4C13AA96A6430E4AFC6AF5304FBE4407" box="[1007,1154,260,284]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="plagiosum">
<emphasis id="B9670D07A6430E4AFC6AF5304FBE4407" box="[1007,1154,260,284]" italics="true" pageId="10" pageNumber="273">P. plagiosum</emphasis>
</taxonomicName>
the apex of the cerci is tapering and in
<taxonomicName id="4C13AA96A6430E4AFEC9F51C4AF6445B" box="[332,458,296,320]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="dorothea">
<emphasis id="B9670D07A6430E4AFEC9F51C4AF6445B" box="[332,458,296,320]" italics="true" pageId="10" pageNumber="273">P. dorothea</emphasis>
</taxonomicName>
it appears bifid, butin
<taxonomicName id="4C13AA96A6430E4AFD51F51C4854445B" box="[724,872,296,320]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="ambiguum">
<emphasis id="B9670D07A6430E4AFD51F51C4854445B" box="[724,872,296,320]" italics="true" pageId="10" pageNumber="273">P. ambiguum</emphasis>
</taxonomicName>
the apex is robust and rounded. The paraproct is robust and upward slanting in
<taxonomicName id="4C13AA96A6430E4AFE70F57849B4447F" box="[501,648,332,356]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="ambiguum">
<emphasis id="B9670D07A6430E4AFE70F57849B4447F" box="[501,648,332,356]" italics="true" pageId="10" pageNumber="273">P. ambiguum</emphasis>
</taxonomicName>
, close and parallel in lateral view to the basal spine of the cerci and extending to about one third of the length of the cerci. In
<emphasis id="B9670D07A6430E4AFCA0F544489F4493" box="[805,931,368,392]" italics="true" pageId="10" pageNumber="273">P. do ro t h e a</emphasis>
the paraproct is short, triangular and carries two spines. In
<taxonomicName id="4C13AA96A6430E4AFEBFF5A04AF644B7" box="[314,458,404,428]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="plagiosum">
<emphasis id="B9670D07A6430E4AFEBFF5A04AF644B7" box="[314,458,404,428]" italics="true" pageId="10" pageNumber="273">P. plagiosum</emphasis>
</taxonomicName>
the paraproct has a short, curved spine, not nearly as robust as in
<taxonomicName id="4C13AA96A6430E4AFB2CF5A04E0744B7" box="[1193,1339,404,428]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="ambiguum">
<emphasis id="B9670D07A6430E4AFB2CF5A04E0744B7" box="[1193,1339,404,428]" italics="true" pageId="10" pageNumber="273">P. ambiguum</emphasis>
</taxonomicName>
, and not following the line of the basal tooth of the cerci closely. In addition, the body of
<taxonomicName id="4C13AA96A6430E4AFB9CF58C4F9644CB" box="[1049,1194,440,464]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="ambiguum">
<emphasis id="B9670D07A6430E4AFB9CF58C4F9644CB" box="[1049,1194,440,464]" italics="true" pageId="10" pageNumber="273">P. ambiguum</emphasis>
</taxonomicName>
is obviously slimmer than
<taxonomicName id="4C13AA96A6430E4AFF4BF5E84A7244EF" box="[206,334,476,500]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="dorothea">
<emphasis id="B9670D07A6430E4AFF4BF5E84A7244EF" box="[206,334,476,500]" italics="true" pageId="10" pageNumber="273">P. dorothea</emphasis>
</taxonomicName>
as well as all other
<taxonomicName id="4C13AA96A6430E4AFDABF5E8498544EF" box="[558,697,477,500]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="B9670D07A6430E4AFDABF5E8498544EF" box="[558,697,477,500]" italics="true" pageId="10" pageNumber="273">Paracercion</emphasis>
</taxonomicName>
species examined. The female can be identified by the hind ridge of the pronotum, which differs from those of both
<taxonomicName id="4C13AA96A6430E4AFD49F63448774703" box="[716,843,512,536]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="dorothea">
<emphasis id="B9670D07A6430E4AFD49F63448774703" box="[716,843,512,536]" italics="true" pageId="10" pageNumber="273">P. dorothea</emphasis>
</taxonomicName>
and
<taxonomicName id="4C13AA96A6430E4AFC07F6344F2D4703" box="[898,1041,512,536]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="plagiosum">
<emphasis id="B9670D07A6430E4AFC07F6344F2D4703" box="[898,1041,512,536]" italics="true" pageId="10" pageNumber="273">P. plagiosum</emphasis>
</taxonomicName>
by its bow-shaped outline, straight in the middle, then curving first forward then backward again towards its lateral ends. In
<emphasis id="B9670D07A6430E4AFB2EF6104E174727" box="[1195,1323,548,572]" italics="true" pageId="10" pageNumber="273">P. do ro t h e a</emphasis>
there are foliate outgrowths on either side of the hind ridge and in
<taxonomicName id="4C13AA96A6430E4AFCA2F67C4885477B" box="[807,953,584,608]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="plagiosum">
<emphasis id="B9670D07A6430E4AFCA2F67C4885477B" box="[807,953,584,608]" italics="true" pageId="10" pageNumber="273">P. plagiosum</emphasis>
</taxonomicName>
the hind ridge is undulating, but lacks the straight middle. This straight middle section is at least lacking too in
<emphasis id="B9670D07A6430E4AFC2AF6584AA147B3" italics="true" pageId="10" pageNumber="273">
<taxonomicName id="4C13AA96A6430E4AFC2AF6584F72479F" box="[943,1102,620,644]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="calamorum">P. calamorum</taxonomicName>
,
<taxonomicName id="4C13AA96A6430E4AFBDEF6584FD4479F" box="[1115,1256,620,644]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="barbatum">P. barbatum</taxonomicName>
,
<taxonomicName id="4C13AA96A6430E4AFB71F6584E46479F" box="[1268,1402,621,644]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="v-nigrum">P. v-nigrum</taxonomicName>
,
<taxonomicName id="4C13AA96A6430E4AFA02F6584A2A47B3" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="melanotum">P. melanotum</taxonomicName>
,
<taxonomicName id="4C13AA96A6430E4AFEA4F6A44AA147B3" box="[289,413,656,680]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="sieboldii">P. sieboldii</taxonomicName>
</emphasis>
and
<taxonomicName id="4C13AA96A6430E4AFE51F6A449A247B3" box="[468,670,656,680]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="hieroglyphicum">
<emphasis id="B9670D07A6430E4AFE51F6A449A247B3" box="[468,670,656,680]" italics="true" pageId="10" pageNumber="273">P. hieroglyphicum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="DF6C819DA6430E4AFF12F25E49E7439A" box="[151,731,1643,1665]" httpUri="https://zenodo.org/record/261238/files/figure.png" pageId="10" pageNumber="273" targetBox="[151,1435,733,1621]" targetPageId="10">
<paragraph id="8BACD115A6430E4AFF12F25E49E7439A" blockId="10.[151,731,1643,1665]" box="[151,731,1643,1665]" pageId="10" pageNumber="273">
<emphasis id="B9670D07A6430E4AFF12F25E4A2E439B" bold="true" box="[151,274,1643,1665]" pageId="10" pageNumber="273">FIGURE 9.</emphasis>
Field photo of male
<taxonomicName id="4C13AA96A6430E4AFE63F25E49E9439B" box="[486,725,1643,1664]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="ambiguum">
<emphasis id="B9670D07A6430E4AFE63F25E49E9439B" box="[486,725,1643,1664]" italics="true" pageId="10" pageNumber="273">Paracercion ambiguum</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C309829EA6430E4BFF42F2984AF945E3" lastPageId="11" lastPageNumber="274" pageId="10" pageNumber="273" type="description">
<paragraph id="8BACD115A6430E4BFF42F2984AF945E3" blockId="10.[151,1436,1709,2022]" lastBlockId="11.[151,1436,151,248]" lastPageId="11" lastPageNumber="274" pageId="10" pageNumber="273">
<emphasis id="B9670D07A6430E4AFF42F298497643DD" bold="true" box="[199,586,1709,1734]" pageId="10" pageNumber="273">Notes on habitat and behavior.</emphasis>
Unlike most
<taxonomicName id="4C13AA96A6430E4AFD77F29B484143DE" box="[754,893,1710,1733]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="B9670D07A6430E4AFD77F29B484143DE" box="[754,893,1710,1733]" italics="true" pageId="10" pageNumber="273">Paracercion</emphasis>
</taxonomicName>
species,
<taxonomicName id="4C13AA96A6430E4AFC6BF29B4FB943DE" box="[1006,1157,1709,1733]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="ambiguum">
<emphasis id="B9670D07A6430E4AFC6BF29B4FB943DE" box="[1006,1157,1709,1733]" italics="true" pageId="10" pageNumber="273">P. ambiguum</emphasis>
</taxonomicName>
behaves much like the
<taxonomicName id="4C13AA96A6430E4AFF12F2E74A1A43F1" box="[151,294,1746,1770]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="B9670D07A6430E4AFF12F2E74A1A43F1" box="[151,294,1746,1770]" italics="true" pageId="10" pageNumber="273">Pseudagrion</emphasis>
</taxonomicName>
species it occurs side-by-side with at the
<typeStatus id="54A86FB7A6430E4AFC87F2E7480E43F1" box="[770,818,1746,1770]" pageId="10" pageNumber="273">type</typeStatus>
locality. Males typically hang from tips of vegetation, between just a few cm and
<quantity id="4CEB7CF0A6430E4AFE43F2C349314216" box="[454,525,1782,1806]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="273" unit="cm" value="30.0">30 cm</quantity>
, over the water surface along the edge of a slow streaming river, about
<quantity id="4CEB7CF0A6430E4AFAA3F2C34E644215" box="[1318,1368,1782,1806]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="273" unit="m" value="10.0">10m</quantity>
wide, or along the edges of the reservoir from which the stream originates. Males are easily found, even under less promising cloudy conditions, but females are rarely observed and only visit to oviposit, presumably foraging at some distance from the water. Males were observed to interact with
<taxonomicName id="4C13AA96A6430E4AFC08F3574F4B4262" box="[909,1143,1890,1914]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="spencei">
<emphasis id="B9670D07A6430E4AFC08F3574F4B4262" box="[909,1143,1890,1914]" italics="true" pageId="10" pageNumber="273">Pseudagrion spencei</emphasis>
</taxonomicName>
and
<taxonomicName id="4C13AA96A6430E4AFB29F3574E494261" box="[1196,1397,1890,1914]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="microcephalum">
<emphasis id="B9670D07A6430E4AFB29F3574E494261" box="[1196,1397,1890,1914]" italics="true" pageId="10" pageNumber="273">P. microcephalum</emphasis>
</taxonomicName>
for positions on leaves. Other coenagrionid species common along the same stream include
<taxonomicName id="4C13AA96A6430E4AFB0CF3B34EA04286" box="[1161,1436,1925,1949]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="calamorum">
<emphasis id="B9670D07A6430E4AFB0CF3B34EA04286" box="[1161,1436,1925,1949]" italics="true" pageId="10" pageNumber="273">Paracercion calamorum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C13AA96A6430E4AFF42F39F4AF542D9" box="[199,457,1962,1986]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="rubriceps">
<emphasis id="B9670D07A6430E4AFF42F39F4AF542D9" box="[199,457,1962,1986]" italics="true" pageId="10" pageNumber="273">Pseudagrion rubriceps</emphasis>
</taxonomicName>
, with
<taxonomicName id="4C13AA96A6430E4AFD97F39F481E42D9" box="[530,802,1962,1986]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="melanotum">
<emphasis id="B9670D07A6430E4AFD97F39F481E42D9" box="[530,802,1962,1986]" italics="true" pageId="10" pageNumber="273">Paracercion melanotum</emphasis>
</taxonomicName>
scarcer.
<taxonomicName id="4C13AA96A6430E4AFC03F39F4F2542D9" box="[902,1049,1962,1986]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="species" species="ambiguum">
<emphasis id="B9670D07A6430E4AFC03F39F4F2542D9" box="[902,1049,1962,1986]" italics="true" pageId="10" pageNumber="273">P. ambiguum</emphasis>
</taxonomicName>
has not been seen to interact with these other
<taxonomicName id="4C13AA96A6430E4AFE9EF3FB4A9A42FE" box="[283,422,1998,2021]" class="Insecta" family="Coenagrionidae" genus="Paracercion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="B9670D07A6430E4AFE9EF3FB4A9A42FE" box="[283,422,1998,2021]" italics="true" pageId="10" pageNumber="273">Paracercion</emphasis>
</taxonomicName>
species, which display rather different behaviour, perching on floating weeds rather than hanging from vegetation along the water’s edge. Observations have been made with earliest date
<date id="FFADF7D5A6420E4BFB7EF4A24E5645AB" box="[1275,1386,151,176]" pageId="11" pageNumber="274" value="1922-03">March 22</date>
and latest
<date id="FFADF7D5A6420E4BFF59F4894A5645CF" box="[220,362,188,213]" pageId="11" pageNumber="274">December 7</date>
, but the species may well occur year round. It is relatively common, with at least
<specimenCount id="9D151A9CA6420E4BFAB1F4894EA045CE" box="[1332,1436,188,213]" pageId="11" pageNumber="274" type="male">10 males</specimenCount>
observed during each visit.
</paragraph>
</subSubSection>
</treatment>
</document>
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