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<document id="B3054C286FC9DFA15955C4A8D866EE66" ID-CLB-Dataset="26262" ID-DOI="10.11646/zootaxa.4629.4.4" ID-GBIF-Dataset="5cad254d-d641-4448-a8b7-34d1bdfa17f2" ID-ISSN="1175-5326" ID-Zenodo-Dep="3274140" ID-ZooBank="986375B5-931C-4284-8AAA-CABB7D36C038" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1562654103261" checkinUser="plazi" docAuthor="Azarkina, Galina N. &amp; Trilikauskas, Laimonas A." docDate="2019" docId="039887D3B703FFCDFF25FB7366FBE6DD" docLanguage="en" docName="zootaxa.4629.4.4.pdf" docOrigin="Zootaxa 4629 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Halocosa Azarkina &amp; Trilikauskas 2019, gen. n." docType="treatment" docVersion="10" lastPageNumber="557" masterDocId="FFA1FFABB702FFCFFFB2FFB56738E279" masterDocTitle="Halocosa gen. n., a new genus of Lycosidae (Araneae) from the Palaearctic, with a redescription of H. cereipes (L. Koch, 1878)" masterLastPageNumber="570" masterPageNumber="555" pageNumber="556" updateTime="1698739575921" updateUser="ExternalLinkService">
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<mods:title id="1B5186646F6915E1C24F4D5A79DD62E2">Halocosa gen. n., a new genus of Lycosidae (Araneae) from the Palaearctic, with a redescription of H. cereipes (L. Koch, 1878)</mods:title>
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<mods:namePart id="CCC6832D4E48EFE8AAB25E68F1F6D82F">Azarkina, Galina N.</mods:namePart>
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<paragraph id="8B8E36C5B703FFCEFF25FB73665BE698" blockId="1.[151,355,1222,1249]" box="[151,355,1222,1249]" pageId="1" pageNumber="556">
<heading id="D0C681A9B703FFCEFF25FB73665BE698" bold="true" box="[151,355,1222,1249]" fontSize="11" level="1" pageId="1" pageNumber="556" reason="1">
<emphasis id="B945EAD7B703FFCEFF25FB73665BE698" bold="true" box="[151,355,1222,1249]" pageId="1" pageNumber="556">
<taxonomicName id="4C314D46B703FFCEFF25FB736631E698" ID-CoL="4SVT" ID-ENA="2713039" authority="Azarkina &amp; Trilikauskas, 2019" authorityName="Azarkina &amp; Trilikauskas" authorityYear="2019" box="[151,265,1222,1249]" class="Arachnida" family="Lycosidae" genus="Halocosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="556" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B945EAD7B703FFCEFF25FB736631E698" bold="true" box="[151,265,1222,1249]" italics="true" pageId="1" pageNumber="556">Halocosa</emphasis>
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<taxonomicNameLabel id="A27657ACB703FFCEFEA3FB72665BE698" box="[273,355,1223,1249]" pageId="1" pageNumber="556" rank="genus">gen. n.</taxonomicNameLabel>
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<paragraph id="8B8E36C5B703FFCEFF25FABE655CE75A" blockId="1.[151,612,1290,1315]" box="[151,612,1290,1315]" pageId="1" pageNumber="556">
<emphasis id="B945EAD7B703FFCEFF25FABE6626E75A" bold="true" box="[151,286,1290,1315]" pageId="1" pageNumber="556">
<typeStatus id="548A8867B703FFCEFF25FABE67F4E75A" box="[151,204,1291,1315]" pageId="1" pageNumber="556">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="4C314D46B703FFCEFE98FABE655CE75A" authority="L. Koch, 1878" authorityName="L. Koch" authorityYear="1878" box="[298,612,1290,1315]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="556" phylum="Arthropoda" rank="species" species="cereipes">
<emphasis id="B945EAD7B703FFCEFE98FABE66F2E75B" box="[298,458,1291,1314]" italics="true" pageId="1" pageNumber="556">Lycosa cereipes</emphasis>
L.
<bibRefCitation id="EFA04B34B703FFCEFE5FFABF655CE75A" author="Koch, L." box="[493,612,1290,1315]" pageId="1" pageNumber="556" pagination="36 - 71" refId="ref6715" refString="Koch, L. (1878) Kaukasische Arachnoiden. In: Schneider, O. (Ed.), Naturwissenschaftliche Beitrage zur Kenntniss der Kaukasuslander. Dresden, 3, 36 - 71." type="journal article" year="1878">Koch, 1878</bibRefCitation>
</taxonomicName>
</paragraph>
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<subSubSection id="C32B654EB703FFCEFF25FAE463D1E7F6" pageId="1" pageNumber="556" type="etymology">
<paragraph id="8B8E36C5B703FFCEFF25FAE463D1E7F6" blockId="1.[151,1437,1361,2035]" pageId="1" pageNumber="556">
<emphasis id="B945EAD7B703FFCEFF25FAE46620E712" bold="true" box="[151,280,1361,1387]" pageId="1" pageNumber="556">Etymology</emphasis>
. The generic name refers to saline habitats of members of the new genus, a derivate of the Latinized Greek word “
<emphasis id="B945EAD7B703FFCEFE82FAC0665AE7F6" box="[304,354,1397,1423]" italics="true" pageId="1" pageNumber="556">halo</emphasis>
—“salt”, and “-
<emphasis id="B945EAD7B703FFCEFDA2FAC3657BE7F6" box="[528,579,1398,1423]" italics="true" pageId="1" pageNumber="556">cosa</emphasis>
” a common ending of
<taxonomicName id="4C314D46B703FFCEFCF7FAC0648FE7F6" authorityName="Sundevall" authorityYear="1833" box="[837,951,1397,1423]" class="Arachnida" family="Lycosidae" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="556" phylum="Arthropoda" rank="family">Lycosidae</taxonomicName>
genera. Gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="C32B654EB703FFCEFF75FA2D652BE4D6" pageId="1" pageNumber="556" type="diagnosis">
<paragraph id="8B8E36C5B703FFCEFF75FA2D652BE4D6" blockId="1.[151,1437,1361,2035]" pageId="1" pageNumber="556">
<emphasis id="B945EAD7B703FFCEFF75FA2D6602E7CA" bold="true" box="[199,314,1432,1459]" pageId="1" pageNumber="556">Diagnosis</emphasis>
.
<taxonomicName id="4C314D46B703FFCEFEFBFA2C668CE7CA" authority="Azarkina &amp; Trilikauskas, 2019" authorityName="Azarkina &amp; Trilikauskas" authorityYear="2019" box="[329,436,1433,1459]" class="Arachnida" family="Lycosidae" genus="Halocosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="556" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B945EAD7B703FFCEFEFBFA2C668CE7CA" box="[329,436,1433,1459]" italics="true" pageId="1" pageNumber="556">Halocosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657ACB703FFCEFE0EFA2C6533E7CA" box="[444,523,1433,1459]" pageId="1" pageNumber="556" rank="genus">
<emphasis id="B945EAD7B703FFCEFE0EFA2C6533E7CA" bold="true" box="[444,523,1433,1459]" pageId="1" pageNumber="556">gen. n.</emphasis>
</taxonomicNameLabel>
belongs to the subfamily Lycosinae
<bibRefCitation id="EFA04B34B703FFCEFC1AFA2C6366E7CA" author="Sundevall, C. J." box="[936,1118,1433,1459]" pageId="1" pageNumber="556" pagination="172 - 272" refId="ref8156" refString="Sundevall, C. J. (1833 a) Svenska spindlarnes beskrifning. Fortsattning och slut. Bihang till Kongliga Svenska Vetenskaps-Akademiens Handlingar 1832, 172 - 272. https: // doi. org / 10.5962 / bhl. title. 66046" type="journal article" year="1833">Sundevall, 1833</bibRefCitation>
(
<emphasis id="B945EAD7B703FFCEFBDDFA2F6395E7CA" box="[1135,1197,1434,1459]" italics="true" pageId="1" pageNumber="556">sensu</emphasis>
<bibRefCitation id="EFA04B34B703FFCEFB07FA2C627FE7CA" author="Zyuzin, A. A." box="[1205,1351,1432,1459]" pageId="1" pageNumber="556" pagination="40 - 51" refId="ref8373" refString="Zyuzin, A. A. (1985) Generic and subfamilial criteria in the systematics of the spider family Lycosidae (Aranei), with the description of a new genus and two new subfamilies. Trudy Zoologieskogo Instituta Akademija Nauk SSSR, 139, 40 - 51. [in Russian]" type="journal article" year="1985">Zyuzin 1985</bibRefCitation>
). It differs from other genera of this subfamily in the following characters: (1) tegular apophysis triangular, narrowing apically, its apical part is bifurcated and has a groove (it seems that TgA is rolled up—Figs 3, 5); (2) TgA is situated retrolatero-ventrally, Ɔ-shaped (as seen from retrolateral view of left palp—Figs 3, 27, 29, 31); (3) synembolus is connected with terminal apophysis by a membrane (
<figureCitation id="130A2A40B703FFCEFD69F99C641AE43A" box="[731,802,1577,1603]" captionStart="FIGURES 18" captionStartId="4.[151,264,1610,1635]" captionTargetBox="[158,1423,189,1580]" captionTargetId="figure@4.[151,1435,181,1590]" captionTargetPageId="4" captionText="FIGURES 18. Halocosa cereipes comb. n. 1. male left palp, ventral view (ISEA, 001.7980); 2. bulbus, ventral view; 3. ditto, retrolateral view; 4. epigyne, ventral view (ISEA, 001.7980); 5. tegular apophysis, retrolateral view; 6.Apex of embolus, ventral view; 7. embolic division, ventral view; 8. ditto, ventro-apical view. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/3274142/files/figure.png" pageId="1" pageNumber="556">Figs 8</figureCitation>
,
<figureCitation id="130A2A40B703FFCEFC9CF99C6440E43A" box="[814,888,1577,1603]" captionStart="FIGURES 3240" captionStartId="8.[151,264,1924,1949]" captionTargetBox="[154,1431,246,1827]" captionTargetId="figure@8.[151,1436,236,1839]" captionTargetPageId="8" captionText="FIGURES 3240. Halocosa cereipes comb. n. 3236. male left palp, ventral view (3233 ISEA 001.8186 and ISEA 001.8185, 34 ISEA 001.8303, 3435 ISEA 001.8188 and ISEA 001.8189); 3739. terminal part with embolus, ventral view (37 neotype ZMB/Arach49123, 38 ISEA, 001.8187, 39 ISEA, 001.7984); 40. bulbus, apical view (ISEA, 001.7979). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/3274150/files/figure.png" pageId="1" pageNumber="556">3739</figureCitation>
); (4) spermathecae with long, tube-shaped accessory glands widening apically (
<figureCitation id="130A2A40B703FFCEFE45F9F8657DE41E" box="[503,581,1613,1639]" captionStart="FIGURES 4149" captionStartId="9.[151,264,1289,1314]" captionTargetBox="[153,1433,250,1229]" captionTargetId="figure@9.[151,1436,229,1244]" captionTargetPageId="9" captionText="FIGURES 4149. Halocosa cereipes comb. n. 41, 43. epigyne, ventral view (ISEA, 001.7983 and PSU, 7668); 42, 4446. vulva, dorsal view (42, 4546 ISEA, 001.7983, 44 PSU, 7668); 4748. scheme of insemination duct; 49. vulva, lateral view (ISEA, 001.7980). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/3274152/files/figure.png" pageId="1" pageNumber="556">Fig 49</figureCitation>
); (5) septum and septal pedicel wide, epigyne apically with a wide epigynal atrium, which is bordered laterally by epigynal “wings” (
<figureCitation id="130A2A40B703FFCEFC94F9C46443E4F2" box="[806,891,1649,1675]" captionStart="FIGURES 4149" captionStartId="9.[151,264,1289,1314]" captionTargetBox="[153,1433,250,1229]" captionTargetId="figure@9.[151,1436,229,1244]" captionTargetPageId="9" captionText="FIGURES 4149. Halocosa cereipes comb. n. 41, 43. epigyne, ventral view (ISEA, 001.7983 and PSU, 7668); 42, 4446. vulva, dorsal view (42, 4546 ISEA, 001.7983, 44 PSU, 7668); 4748. scheme of insemination duct; 49. vulva, lateral view (ISEA, 001.7980). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/3274152/files/figure.png" pageId="1" pageNumber="556">Fig. 41</figureCitation>
); and (6) the upper part of the copulatory ducts swollen, forming a loop (
<figureCitation id="130A2A40B703FFCEFE00F920653EE4D6" box="[434,518,1685,1711]" captionStart="FIGURES 4149" captionStartId="9.[151,264,1289,1314]" captionTargetBox="[153,1433,250,1229]" captionTargetId="figure@9.[151,1436,229,1244]" captionTargetPageId="9" captionText="FIGURES 4149. Halocosa cereipes comb. n. 41, 43. epigyne, ventral view (ISEA, 001.7983 and PSU, 7668); 42, 4446. vulva, dorsal view (42, 4546 ISEA, 001.7983, 44 PSU, 7668); 4748. scheme of insemination duct; 49. vulva, lateral view (ISEA, 001.7980). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/3274152/files/figure.png" pageId="1" pageNumber="556">Fig. 49</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32B654EB703FFCEFF75F90D645DE58A" pageId="1" pageNumber="556" type="description">
<paragraph id="8B8E36C5B703FFCEFF75F90D6472E5FE" blockId="1.[151,1437,1361,2035]" pageId="1" pageNumber="556">
<emphasis id="B945EAD7B703FFCEFF75F90D6669E4AA" bold="true" box="[199,337,1720,1747]" pageId="1" pageNumber="556">Description</emphasis>
. Medium to large spiders, with body length ranging from
<quantity id="4CC99B20B703FFCEFC5FF90C6393E4AA" box="[1005,1195,1721,1747]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" metricValueMax="9.8" metricValueMin="6.4" pageId="1" pageNumber="556" unit="mm" value="8.1" valueMax="9.8" valueMin="6.4">6.40 to 9.80 mm</quantity>
in males and
<quantity id="4CC99B20B703FFCEFAFFF90C6632E48E" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.26" metricValueMin="0.84" pageId="1" pageNumber="556" unit="mm" value="10.5" valueMax="12.6" valueMin="8.4">8.40 to 12.60 mm</quantity>
in females. Sexes are similar in general body shape and coloration. Carapace low, without a transverse depression that is typical of
<taxonomicName id="4C314D46B703FFCEFE6FF8B76514E562" authorityName="Simon" authorityYear="1882" box="[477,556,1794,1819]" class="Arachnida" family="Lycosidae" genus="Evippa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="556" phylum="Arthropoda" rank="genus">
<emphasis id="B945EAD7B703FFCEFE6FF8B76514E562" box="[477,556,1794,1819]" italics="true" pageId="1" pageNumber="556">Evippa</emphasis>
</taxonomicName>
(cf.
<figureCitation id="130A2A40B703FFCEFDD6F8B46584E562" box="[612,700,1793,1819]" captionStart="FIGURES 918" captionStartId="5.[151,264,1935,1960]" captionTargetBox="[199,1388,166,1906]" captionTargetId="figure@5.[199,1388,166,1915]" captionTargetPageId="5" captionText="FIGURES 918. Arctosa leopardus (910), Evippa jocquei (11-14) and Halocosa cereipes comb. n. (1518) 9, 11, 15 male carapace, dorsal view; 10, 12, 16 male “face”; 13, 17 male carapace, lateral view; 14, 18 male first leg, retrolateral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3274144/files/figure.png" pageId="1" pageNumber="556">Figs 13</figureCitation>
, arrowed and 17 for
<taxonomicName id="4C314D46B703FFCEFC1DF8B7627EE562" authority="Alderweireldt, 1991" authorityName="Alderweireldt" authorityYear="1991" box="[943,1350,1793,1819]" class="Arachnida" family="Lycosidae" genus="Evippa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="556" phylum="Arthropoda" rank="species" species="jocquei">
<emphasis id="B945EAD7B703FFCEFC1DF8B76361E562" box="[943,1113,1794,1819]" italics="true" pageId="1" pageNumber="556">Evippa jocquei</emphasis>
<bibRefCitation id="EFA04B34B703FFCEFBD1F8B4627EE562" author="Alderweireldt, M." box="[1123,1350,1793,1819]" pageId="1" pageNumber="556" pagination="359 - 381" refId="ref6168" refString="Alderweireldt, M. (1991) A revision of the African representatives of the wolf spider genus Evippa Simon, 1882 (Araneae, Lycosidae) with notes on allied species and genera. Journal of Natural History, 25, 359 - 381. https: // doi. org / 10.1080 / 00222939100770261" type="journal article" year="1991">Alderweireldt, 1991</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C314D46B703FFCEFA33F8B767CBE546" authorityName="L. Koch" authorityYear="1878" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="556" phylum="Arthropoda" rank="species" species="cereipes">
<emphasis id="B945EAD7B703FFCEFA33F8B767CBE546" italics="true" pageId="1" pageNumber="556">H. cereipes</emphasis>
</taxonomicName>
respectivelly). Carapace coloration of
<taxonomicName id="4C314D46B703FFCEFD06F8906427E546" authority="Azarkina &amp; Trilikauskas, 2019" authorityName="Azarkina &amp; Trilikauskas" authorityYear="2019" box="[692,799,1829,1855]" class="Arachnida" family="Lycosidae" genus="Halocosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="556" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B945EAD7B703FFCEFD06F8906427E546" box="[692,799,1829,1855]" italics="true" pageId="1" pageNumber="556">Halocosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657ACB703FFCEFC9BF8906443E546" box="[809,891,1829,1855]" pageId="1" pageNumber="556" rank="genus">
<emphasis id="B945EAD7B703FFCEFC9BF8906443E546" bold="true" box="[809,891,1829,1855]" pageId="1" pageNumber="556">gen. n.</emphasis>
</taxonomicNameLabel>
is similar to that of some
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<emphasis id="B945EAD7B703FFCEFB02F890623EE546" box="[1200,1286,1829,1855]" italics="true" pageId="1" pageNumber="556">Arctosa</emphasis>
</taxonomicName>
species (e.g.
<bibRefCitation id="EFA04B34B703FFCEFF25F8FC66FCE51A" author="Lugetti, G. &amp; Tongiorgi, P." box="[151,452,1864,1891]" pageId="1" pageNumber="556" pagination="165 - 229" refId="ref6845" refString="Lugetti, G. &amp; Tongiorgi, P. (1965) Revisione delle specie italiane dei generi Arctosa C. L. Koch e Tricca Simon con note su una Acantholycosa delle Alpi Giulie (Araneae-Lycosidae). Redia, 49, 165 - 229." type="journal article" year="1965">Lugetti &amp; Tongiorgi 1965</bibRefCitation>
, fig.
<date id="FF8F1005B703FFCEFDB3F8FC6527E51A" box="[513,543,1865,1891]" pageId="1" pageNumber="556">I.1</date>
and
<figureCitation id="130A2A40B703FFCEFDEEF8FC659BE51A" box="[604,675,1865,1891]" captionStart="FIGURES 918" captionStartId="5.[151,264,1935,1960]" captionTargetBox="[199,1388,166,1906]" captionTargetId="figure@5.[199,1388,166,1915]" captionTargetPageId="5" captionText="FIGURES 918. Arctosa leopardus (910), Evippa jocquei (11-14) and Halocosa cereipes comb. n. (1518) 9, 11, 15 male carapace, dorsal view; 10, 12, 16 male “face”; 13, 17 male carapace, lateral view; 14, 18 male first leg, retrolateral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3274144/files/figure.png" pageId="1" pageNumber="556">Fig. 9</figureCitation>
):
<emphasis id="B945EAD7B703FFCEFD09F8FF645AE51A" box="[699,866,1865,1891]" italics="true" pageId="1" pageNumber="556">pars thoracica</emphasis>
medially with a butterfly-shaped patch and a Wshaped yellow patch in distal part of
<emphasis id="B945EAD7B703FFCEFD86F8DB65E2E5FE" box="[564,730,1901,1927]" italics="true" pageId="1" pageNumber="556">pars cephalica</emphasis>
(
<figureCitation id="130A2A40B703FFCEFD5BF8D86404E5FE" box="[745,828,1900,1927]" captionStart="FIGURES 918" captionStartId="5.[151,264,1935,1960]" captionTargetBox="[199,1388,166,1906]" captionTargetId="figure@5.[199,1388,166,1915]" captionTargetPageId="5" captionText="FIGURES 918. Arctosa leopardus (910), Evippa jocquei (11-14) and Halocosa cereipes comb. n. (1518) 9, 11, 15 male carapace, dorsal view; 10, 12, 16 male “face”; 13, 17 male carapace, lateral view; 14, 18 male first leg, retrolateral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3274144/files/figure.png" pageId="1" pageNumber="556">Fig. 15</figureCitation>
).
</paragraph>
<paragraph id="8B8E36C5B703FFCEFF75F824645DE58A" blockId="1.[151,1437,1361,2035]" pageId="1" pageNumber="556">
Eyes in three rows. The second row as wide as the first one, AME 1.5 times as big as ALE (
<figureCitation id="130A2A40B703FFCEFB7AF8246223E5D5" box="[1224,1307,1937,1964]" captionStart="FIGURES 918" captionStartId="5.[151,264,1935,1960]" captionTargetBox="[199,1388,166,1906]" captionTargetId="figure@5.[199,1388,166,1915]" captionTargetPageId="5" captionText="FIGURES 918. Arctosa leopardus (910), Evippa jocquei (11-14) and Halocosa cereipes comb. n. (1518) 9, 11, 15 male carapace, dorsal view; 10, 12, 16 male “face”; 13, 17 male carapace, lateral view; 14, 18 male first leg, retrolateral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3274144/files/figure.png" pageId="1" pageNumber="556">Fig. 16</figureCitation>
); in
<taxonomicName id="4C314D46B703FFCEFAFFF82762A4E5D2" authorityName="Simon" authorityYear="1882" box="[1357,1436,1938,1963]" class="Arachnida" family="Lycosidae" genus="Evippa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="556" phylum="Arthropoda" rank="genus">
<emphasis id="B945EAD7B703FFCEFAFFF82762A4E5D2" box="[1357,1436,1938,1963]" italics="true" pageId="1" pageNumber="556">Evippa</emphasis>
</taxonomicName>
the second eye row is wider than the first one, and AME twice as big as ALE (
<figureCitation id="130A2A40B703FFCEFBBAF8006363E5B6" box="[1032,1115,1973,1999]" captionStart="FIGURES 918" captionStartId="5.[151,264,1935,1960]" captionTargetBox="[199,1388,166,1906]" captionTargetId="figure@5.[199,1388,166,1915]" captionTargetPageId="5" captionText="FIGURES 918. Arctosa leopardus (910), Evippa jocquei (11-14) and Halocosa cereipes comb. n. (1518) 9, 11, 15 male carapace, dorsal view; 10, 12, 16 male “face”; 13, 17 male carapace, lateral view; 14, 18 male first leg, retrolateral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3274144/files/figure.png" pageId="1" pageNumber="556">Fig. 12</figureCitation>
); in
<taxonomicName id="4C314D46B703FFCEFB3CF80063DCE5B6" authorityName="C.L.Koch" authorityYear="1847" box="[1166,1252,1973,1999]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="556" phylum="Arthropoda" rank="genus">
<emphasis id="B945EAD7B703FFCEFB3CF80063DCE5B6" box="[1166,1252,1973,1999]" italics="true" pageId="1" pageNumber="556">Arctosa</emphasis>
</taxonomicName>
the width of the first and second eye rows is identical, and AME=ALE (
<figureCitation id="130A2A40B703FFCEFCB6F86C6460E58A" box="[772,856,2009,2035]" captionStart="FIGURES 918" captionStartId="5.[151,264,1935,1960]" captionTargetBox="[199,1388,166,1906]" captionTargetId="figure@5.[199,1388,166,1915]" captionTargetPageId="5" captionText="FIGURES 918. Arctosa leopardus (910), Evippa jocquei (11-14) and Halocosa cereipes comb. n. (1518) 9, 11, 15 male carapace, dorsal view; 10, 12, 16 male “face”; 13, 17 male carapace, lateral view; 14, 18 male first leg, retrolateral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3274144/files/figure.png" pageId="1" pageNumber="556">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32B654EB700FFCDFF75FF2C6657E33A" pageId="2" pageNumber="557" type="discussion">
<paragraph id="8B8E36C5B700FFCDFF75FF2C6657E33A" blockId="2.[151,1437,152,1188]" pageId="2" pageNumber="557">
Tibia I with 6 (3 pairs) ventral spines, metatarsus I with 5 ventral spines, tarsal claws of legs III of usual length, with eight teeth, those of legs IIIIV long, with 10 teeth and long pretarsal hairs, tarsus without pseudo-articulation (
<figureCitation id="130A2A40B700FFCDFF12FF5467CAE282" box="[160,242,225,251]" captionStart="FIGURES 918" captionStartId="5.[151,264,1935,1960]" captionTargetBox="[199,1388,166,1906]" captionTargetId="figure@5.[199,1388,166,1915]" captionTargetPageId="5" captionText="FIGURES 918. Arctosa leopardus (910), Evippa jocquei (11-14) and Halocosa cereipes comb. n. (1518) 9, 11, 15 male carapace, dorsal view; 10, 12, 16 male “face”; 13, 17 male carapace, lateral view; 14, 18 male first leg, retrolateral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3274144/files/figure.png" pageId="2" pageNumber="557">Fig. 18</figureCitation>
). In
<taxonomicName id="4C314D46B700FFCDFE96FF57664BE282" authorityName="Simon" authorityYear="1882" box="[292,371,226,251]" class="Arachnida" family="Lycosidae" genus="Evippa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="557" phylum="Arthropoda" rank="genus">
<emphasis id="B945EAD7B700FFCDFE96FF57664BE282" box="[292,371,226,251]" italics="true" pageId="2" pageNumber="557">Evippa</emphasis>
</taxonomicName>
, the number of ventral spines on tibia
<date id="FF8F1005B700FFCDFCAFFF546467E282" box="[797,863,225,251]" pageId="2" pageNumber="557" value="2010-01">I—10</date>
(5 pairs) or more, tarsal claws long, with numerous teeth (from
<quantity id="4CC99B20B700FFCDFEAEFEB06670E366" box="[284,328,261,287]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="2" pageNumber="557" unit="in" value="7.0">7 in</quantity>
<taxonomicName id="4C314D46B700FFCDFEFDFEB36534E366" authorityName="Alredweireldt et Jocque" authorityYear="2017" box="[335,524,261,287]" class="Arachnida" family="Lycosidae" genus="Evippa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="557" phylum="Arthropoda" rank="species" species="brevicymbium">
<emphasis id="B945EAD7B700FFCDFEFDFEB36534E366" box="[335,524,261,287]" italics="true" pageId="2" pageNumber="557">E. brevicymbium</emphasis>
</taxonomicName>
to
<quantity id="4CC99B20B700FFCDFD82FEB06552E366" box="[560,618,261,287]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="2" pageNumber="557" unit="in" value="14.0">14 in</quantity>
<taxonomicName id="4C314D46B700FFCDFDC3FEB365DBE366" authorityName="Alderweireldt" authorityYear="1991" box="[625,739,262,287]" class="Arachnida" family="Lycosidae" genus="Evippa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="557" phylum="Arthropoda" rank="species" species="jocquei">
<emphasis id="B945EAD7B700FFCDFDC3FEB365DBE366" box="[625,739,262,287]" italics="true" pageId="2" pageNumber="557">E. jocquei</emphasis>
</taxonomicName>
), tarsus with pseudo-articulation (
<bibRefCitation id="EFA04B34B700FFCDFBD6FEB0627AE366" author="Alderweireldt, M." box="[1124,1346,261,287]" pageId="2" pageNumber="557" pagination="359 - 381" refId="ref6168" refString="Alderweireldt, M. (1991) A revision of the African representatives of the wolf spider genus Evippa Simon, 1882 (Araneae, Lycosidae) with notes on allied species and genera. Journal of Natural History, 25, 359 - 381. https: // doi. org / 10.1080 / 00222939100770261" type="journal article" year="1991">Alderweireldt 1991</bibRefCitation>
, p. 360, fig. 3, and
<figureCitation id="130A2A40B700FFCDFEA2FE9C665AE33A" box="[272,354,297,323]" captionStart="FIGURES 918" captionStartId="5.[151,264,1935,1960]" captionTargetBox="[199,1388,166,1906]" captionTargetId="figure@5.[199,1388,166,1915]" captionTargetPageId="5" captionText="FIGURES 918. Arctosa leopardus (910), Evippa jocquei (11-14) and Halocosa cereipes comb. n. (1518) 9, 11, 15 male carapace, dorsal view; 10, 12, 16 male “face”; 13, 17 male carapace, lateral view; 14, 18 male first leg, retrolateral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3274144/files/figure.png" pageId="2" pageNumber="557">Fig. 14</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32B654EB700FFCDFF75FEF8649CE38E" pageId="2" pageNumber="557" type="nomenclature">
<paragraph id="8B8E36C5B700FFCDFF75FEF8649CE38E" blockId="2.[151,1437,152,1188]" pageId="2" pageNumber="557">
Embolic division with terminal apophysis (Ta) and synembolus (Se) connected by a membrane, embolus long and thin, embolic base is situated latero-medially (
<figureCitation id="130A2A40B700FFCDFD7DFEC4640CE3F2" box="[719,820,369,395]" captionStart="FIGURES 18" captionStartId="4.[151,264,1610,1635]" captionTargetBox="[158,1423,189,1580]" captionTargetId="figure@4.[151,1435,181,1590]" captionTargetPageId="4" captionText="FIGURES 18. Halocosa cereipes comb. n. 1. male left palp, ventral view (ISEA, 001.7980); 2. bulbus, ventral view; 3. ditto, retrolateral view; 4. epigyne, ventral view (ISEA, 001.7980); 5. tegular apophysis, retrolateral view; 6.Apex of embolus, ventral view; 7. embolic division, ventral view; 8. ditto, ventro-apical view. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/3274142/files/figure.png" pageId="2" pageNumber="557">Figs 68</figureCitation>
,
<figureCitation id="130A2A40B700FFCDFCF2FEC464B3E3F2" box="[832,907,369,395]" captionStart="FIGURES 3240" captionStartId="8.[151,264,1924,1949]" captionTargetBox="[154,1431,246,1827]" captionTargetId="figure@8.[151,1436,236,1839]" captionTargetPageId="8" captionText="FIGURES 3240. Halocosa cereipes comb. n. 3236. male left palp, ventral view (3233 ISEA 001.8186 and ISEA 001.8185, 34 ISEA 001.8303, 3435 ISEA 001.8188 and ISEA 001.8189); 3739. terminal part with embolus, ventral view (37 neotype ZMB/Arach49123, 38 ISEA, 001.8187, 39 ISEA, 001.7984); 40. bulbus, apical view (ISEA, 001.7979). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/3274150/files/figure.png" pageId="2" pageNumber="557">3739</figureCitation>
); while in
<taxonomicName id="4C314D46B700FFCDFBB0FEC763F6E3F2" ID-CoL="4HND" ID-ENA="2584324" authority=", Ta and Se" authorityName="Ta and Se" box="[1026,1230,369,395]" class="Arachnida" family="Lycosidae" genus="Evippa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="557" phylum="Arthropoda" rank="genus">
<emphasis id="B945EAD7B700FFCDFBB0FEC76369E3F2" box="[1026,1105,370,395]" italics="true" pageId="2" pageNumber="557">Evippa</emphasis>
, Ta and Se
</taxonomicName>
are not connected by a membrane and embolic base is situated meso-apically (
<figureCitation id="130A2A40B700FFCDFC84FE206481E3D6" box="[822,953,405,431]" captionStart="FIGURES 1925" captionStartId="6.[151,264,1592,1617]" captionTargetBox="[159,1406,186,1543]" captionTargetId="figure@6.[151,1436,181,1572]" captionTargetPageId="6" captionText="FIGURES 1925. Evippa jocquei 19. male left palp, ventral view; 20. ditto, retrolateral view; 21. tegular apophysis, retrolateral view; 22. terminal part with embolus, ventral view; 23. epigyne, ventral view; 24. vulva, dorsal view; 25. scheme of insemination duct. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/3274146/files/figure.png" pageId="2" pageNumber="557">Figs 19, 22</figureCitation>
). Epigynal septum and septal pedicel wide, epigyne apically with a wide epigynal atrium (
<figureCitation id="130A2A40B700FFCDFD2FFE0C65C8E3AA" box="[669,752,441,467]" captionStart="FIGURES 4149" captionStartId="9.[151,264,1289,1314]" captionTargetBox="[153,1433,250,1229]" captionTargetId="figure@9.[151,1436,229,1244]" captionTargetPageId="9" captionText="FIGURES 4149. Halocosa cereipes comb. n. 41, 43. epigyne, ventral view (ISEA, 001.7983 and PSU, 7668); 42, 4446. vulva, dorsal view (42, 4546 ISEA, 001.7983, 44 PSU, 7668); 4748. scheme of insemination duct; 49. vulva, lateral view (ISEA, 001.7980). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/3274152/files/figure.png" pageId="2" pageNumber="557">Fig. 41</figureCitation>
). The upper part of copulatory ducts swollen, forming a loop, spermathecae with long accessory glands widening apically (
<figureCitation id="130A2A40B700FFCDFCF1FE6864A0E38E" box="[835,920,477,503]" captionStart="FIGURES 4149" captionStartId="9.[151,264,1289,1314]" captionTargetBox="[153,1433,250,1229]" captionTargetId="figure@9.[151,1436,229,1244]" captionTargetPageId="9" captionText="FIGURES 4149. Halocosa cereipes comb. n. 41, 43. epigyne, ventral view (ISEA, 001.7983 and PSU, 7668); 42, 4446. vulva, dorsal view (42, 4546 ISEA, 001.7983, 44 PSU, 7668); 4748. scheme of insemination duct; 49. vulva, lateral view (ISEA, 001.7980). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/3274152/files/figure.png" pageId="2" pageNumber="557">Fig. 49</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32B654EB700FFCDFF75FDB565F6E126" pageId="2" pageNumber="557" type="materials_examined">
<paragraph id="8B8E36C5B700FFCDFF75FDB565F6E126" blockId="2.[151,1437,152,1188]" pageId="2" pageNumber="557">
<emphasis id="B945EAD7B700FFCDFF75FDB56667E062" bold="true" box="[199,351,512,539]" pageId="2" pageNumber="557">Composition</emphasis>
.
<taxonomicName id="4C314D46B700FFCDFEDDFDB466E4E062" authority="Azarkina &amp; Trilikauskas, 2019" authorityName="Azarkina &amp; Trilikauskas" authorityYear="2019" box="[367,476,513,539]" class="Arachnida" family="Lycosidae" genus="Halogosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="557" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B945EAD7B700FFCDFEDDFDB466E4E062" box="[367,476,513,539]" italics="true" pageId="2" pageNumber="557">Halogosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657ACB700FFCDFE56FDB4650CE062" box="[484,564,513,539]" pageId="2" pageNumber="557" rank="genus">
<emphasis id="B945EAD7B700FFCDFE56FDB4650CE062" bold="true" box="[484,564,513,539]" pageId="2" pageNumber="557">gen. n.</emphasis>
</taxonomicNameLabel>
consists of three species, of which only
<taxonomicName id="4C314D46B700FFCDFBB3FDB763B8E062" authorityName="Azarkina &amp; Trilikauskas" authorityYear="2019" box="[1025,1152,514,539]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="557" phylum="Arthropoda" rank="species" species="cereipes" status="comb. nov.">
<emphasis id="B945EAD7B700FFCDFBB3FDB763B8E062" box="[1025,1152,514,539]" italics="true" pageId="2" pageNumber="557">H. cereipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657ACB700FFCDFB3AFDB463D7E062" box="[1160,1263,513,539]" pageId="2" pageNumber="557" rank="species">
<emphasis id="B945EAD7B700FFCDFB3AFDB463D7E062" bold="true" box="[1160,1263,513,539]" pageId="2" pageNumber="557">comb. n.</emphasis>
</taxonomicNameLabel>
is known from both sexes. Two
<taxonomicName id="4C314D46B700FFCDFEE8FD906680E046" authorityName="C. L. Koch" authorityYear="1847" box="[346,440,549,575]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="557" phylum="Arthropoda" rank="genus">
<emphasis id="B945EAD7B700FFCDFEE8FD906680E046" box="[346,440,549,575]" italics="true" pageId="2" pageNumber="557">Pardosa</emphasis>
</taxonomicName>
species (
<taxonomicName id="4C314D46B700FFCDFD97FD936583E046" authorityName="Azarkina &amp; Trilikauskas" authorityYear="2019" baseAuthorityName="Azarkina &amp; Trilikauskas" baseAuthorityYear="2019" box="[549,699,549,575]" class="Arachnida" family="Lycosidae" genus="Halocosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="557" phylum="Arthropoda" rank="species" species="hatanensis">
<emphasis id="B945EAD7B700FFCDFD97FD936583E046" box="[549,699,549,575]" italics="true" pageId="2" pageNumber="557">P. hatanensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C314D46B700FFCDFD44FD936467E046" authorityName="Azarkina &amp; Trilikauskas" authorityYear="2019" baseAuthorityName="Azarkina &amp; Trilikauskas" baseAuthorityYear="2019" box="[758,863,550,575]" class="Arachnida" family="Lycosidae" genus="Halocosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="557" phylum="Arthropoda" rank="species" species="jartica">
<emphasis id="B945EAD7B700FFCDFD44FD936467E046" box="[758,863,550,575]" italics="true" pageId="2" pageNumber="557">P. jartica</emphasis>
</taxonomicName>
) described from
<collectingRegion id="49F5F827B700FFCDFB94FD9063E0E046" box="[1062,1240,549,575]" country="China" name="Nei Mongol" pageId="2" pageNumber="557">Inner Mongolia</collectingRegion>
(
<collectingCountry id="F3267655B700FFCDFB5BFD906217E039" box="[1257,1327,549,576]" name="China" pageId="2" pageNumber="557">China</collectingCountry>
) are here transferred to
<taxonomicName id="4C314D46B700FFCDFE8AFDFC669BE01A" authority="Azarkina &amp; Trilikauskas, 2019" authorityName="Azarkina &amp; Trilikauskas" authorityYear="2019" box="[312,419,585,611]" class="Arachnida" family="Lycosidae" genus="Halocosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="558" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B945EAD7B700FFCDFE8AFDFC669BE01A" box="[312,419,585,611]" italics="true" pageId="2" pageNumber="557">Halocosa</emphasis>
</taxonomicName>
. Although we failed to obtain and re-examine their
<typeStatus id="548A8867B700FFCDFC4AFDFC6310E01A" box="[1016,1064,585,611]" pageId="2" pageNumber="557">type</typeStatus>
series, based on the original descriptions and illustrations (see
<bibRefCitation id="EFA04B34B700FFCDFE4BFDD8642CE0FE" author="Urita &amp; Tang, G. M. &amp; Song, D. X." box="[505,788,621,647]" pageId="2" pageNumber="557" pagination="46 - 49" refId="ref8274" refString="Urita, Tang, G. M. &amp; Song, D. X. (1993) Two new species of the genus Pardosa from Inner Mongolia, China (Araneae, Lycosidae). Journal of Inner Mongolia Normal University (Nat. Sci. Ed.), 3, 46 - 49. [in Chinese and English]" type="journal article" year="1993">Urita, Tang et Song 1993</bibRefCitation>
, figs 12), both species belong to the newly erected genus
<taxonomicName id="4C314D46B700FFCDFF25FD24663AE0D2" authority="Azarkina &amp; Trilikauskas, 2019" authorityName="Azarkina &amp; Trilikauskas" authorityYear="2019" box="[151,258,657,683]" class="Arachnida" family="Lycosidae" genus="Halocosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="557" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B945EAD7B700FFCDFF25FD24663AE0D2" box="[151,258,657,683]" italics="true" pageId="2" pageNumber="557">Halocosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657ACB700FFCDFEBAFD24666DE0D2" box="[264,341,657,683]" pageId="2" pageNumber="557" rank="genus">
<emphasis id="B945EAD7B700FFCDFEBAFD246677E0D2" bold="true" box="[264,335,657,683]" pageId="2" pageNumber="557">gen. n</emphasis>
.
</taxonomicNameLabel>
, based on general view of palp and shape of TgA in males, and shape of epigyne and spermathecae with long, tube-shaped accessory glands in females. The two species,
<taxonomicName id="4C314D46B700FFCDFC1AFD03637BE0B6" authorityName="Azarkina &amp; Trilikauskas" authorityYear="2019" baseAuthorityName="Azarkina &amp; Trilikauskas" baseAuthorityYear="2019" box="[936,1091,693,719]" class="Arachnida" family="Lycosidae" genus="Halocosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="557" phylum="Arthropoda" rank="species" species="hatanensis" status="comb. nov.">
<emphasis id="B945EAD7B700FFCDFC1AFD03637BE0B6" box="[936,1091,693,719]" italics="true" pageId="2" pageNumber="557">H. hatanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657ACB700FFCDFBF8FD006391E0B6" box="[1098,1193,693,719]" pageId="2" pageNumber="557" rank="species">
<emphasis id="B945EAD7B700FFCDFBF8FD006399E0B6" bold="true" box="[1098,1185,693,719]" pageId="2" pageNumber="557">comb.n</emphasis>
.
</taxonomicNameLabel>
and
<taxonomicName id="4C314D46B700FFCDFB52FD036275E0B6" authorityName="Azarkina &amp; Trilikauskas" authorityYear="2019" baseAuthorityName="Azarkina &amp; Trilikauskas" baseAuthorityYear="2019" box="[1248,1357,694,719]" class="Arachnida" family="Lycosidae" genus="Halocosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="557" phylum="Arthropoda" rank="species" species="jartica" status="comb. nov.">
<emphasis id="B945EAD7B700FFCDFB52FD036275E0B6" box="[1248,1357,694,719]" italics="true" pageId="2" pageNumber="557">H. jartica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657ACB700FFCDFAE6FD006795E08A" pageId="2" pageNumber="557" rank="species">
<emphasis id="B945EAD7B700FFCDFAE6FD00679FE08A" bold="true" pageId="2" pageNumber="557">comb. n</emphasis>
.
</taxonomicNameLabel>
, remain known only from male and female, respectively.
<taxonomicName id="4C314D46B700FFCDFC89FD6F6490E08A" authorityName="Azarkina &amp; Trilikauskas" authorityYear="2019" baseAuthorityName="Azarkina &amp; Trilikauskas" baseAuthorityYear="2019" box="[827,936,730,755]" class="Arachnida" family="Lycosidae" genus="Halocosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="557" phylum="Arthropoda" rank="species" species="jartica" status="comb. nov.">
<emphasis id="B945EAD7B700FFCDFC89FD6F6490E08A" box="[827,936,730,755]" italics="true" pageId="2" pageNumber="557">H. jartica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657ACB700FFCDFC02FD6C6337E08A" box="[944,1039,729,755]" pageId="2" pageNumber="557" rank="species">
<emphasis id="B945EAD7B700FFCDFC02FD6C633FE08A" bold="true" box="[944,1031,729,755]" pageId="2" pageNumber="557">comb.n</emphasis>
.
</taxonomicNameLabel>
is highly likely to represent the female of
<taxonomicName id="4C314D46B700FFCDFF41FD4B66B5E16E" authorityName="Azarkina &amp; Trilikauskas" authorityYear="2019" baseAuthorityName="Azarkina &amp; Trilikauskas" baseAuthorityYear="2019" box="[243,397,765,791]" class="Arachnida" family="Lycosidae" genus="Halocosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="557" phylum="Arthropoda" rank="species" species="hatanensis" status="comb. nov.">
<emphasis id="B945EAD7B700FFCDFF41FD4B66B5E16E" box="[243,397,765,791]" italics="true" pageId="2" pageNumber="557">H. hatanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657ACB700FFCDFE26FD4866CAE16E" box="[404,498,765,791]" pageId="2" pageNumber="557" rank="species">
<emphasis id="B945EAD7B700FFCDFE26FD4866D4E16E" bold="true" box="[404,492,765,791]" pageId="2" pageNumber="557">comb.n</emphasis>
.
</taxonomicNameLabel>
, and the name
<taxonomicName id="4C314D46B700FFCDFD2CFD4B6433E16E" authorityName="Azarkina &amp; Trilikauskas" authorityYear="2019" baseAuthorityName="Azarkina &amp; Trilikauskas" baseAuthorityYear="2019" box="[670,779,766,791]" class="Arachnida" family="Lycosidae" genus="Halocosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="557" phylum="Arthropoda" rank="species" species="jartica" status="comb. nov.">
<emphasis id="B945EAD7B700FFCDFD2CFD4B6433E16E" box="[670,779,766,791]" italics="true" pageId="2" pageNumber="557">H. jartica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657ACB700FFCDFCA0FD486449E16E" box="[786,881,765,791]" pageId="2" pageNumber="557" rank="species">
<emphasis id="B945EAD7B700FFCDFCA0FD486452E16E" bold="true" box="[786,874,765,791]" pageId="2" pageNumber="557">comb.n</emphasis>
.
</taxonomicNameLabel>
could be synonymised with
<taxonomicName id="4C314D46B700FFCDFB01FD4B6275E16E" authorityName="Azarkina &amp; Trilikauskas" authorityYear="2019" baseAuthorityName="Azarkina &amp; Trilikauskas" baseAuthorityYear="2019" box="[1203,1357,765,791]" class="Arachnida" family="Lycosidae" genus="Halocosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="557" phylum="Arthropoda" rank="species" species="hatanensis" status="comb. nov.">
<emphasis id="B945EAD7B700FFCDFB01FD4B6275E16E" box="[1203,1357,765,791]" italics="true" pageId="2" pageNumber="557">H. hatanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657ACB700FFCDFAE6FD486796E142" pageId="2" pageNumber="557" rank="species">
<emphasis id="B945EAD7B700FFCDFAE6FD48679FE142" bold="true" pageId="2" pageNumber="557">comb. n</emphasis>
.
</taxonomicNameLabel>
A formal synonymy is postponed until the
<typeStatus id="548A8867B700FFCDFD29FC9465F3E142" box="[667,715,801,827]" pageId="2" pageNumber="557">type</typeStatus>
and additional material on both names have been (re)examined (to date we have been unable to borrow the
<typeStatus id="548A8867B700FFCDFD30FCF065FAE126" box="[642,706,837,863]" pageId="2" pageNumber="557">types</typeStatus>
).
</paragraph>
</subSubSection>
<subSubSection id="C32B654EB700FFCDFF75FCDD66FBE6DD" pageId="2" pageNumber="557" type="distribution">
<paragraph id="8B8E36C5B700FFCDFF75FCDD66FBE622" blockId="2.[151,1437,152,1188]" pageId="2" pageNumber="557">
<emphasis id="B945EAD7B700FFCDFF75FCDD6661E1FA" bold="true" box="[199,345,872,899]" pageId="2" pageNumber="557">Distribution</emphasis>
.
<collectingCountry id="F3267655B700FFCDFEDBFCDC66DCE1FA" box="[361,484,873,899]" name="Azerbaijan" pageId="2" pageNumber="557">Azerbaijan</collectingCountry>
(
<bibRefCitation id="EFA04B34B700FFCDFE47FCDC65D8E1FA" author="Marusik, Y. M. &amp; Guseinov, E. F. &amp; Koponen, S." box="[501,736,873,899]" pageId="2" pageNumber="557" pagination="47 - 65" refId="ref6916" refString="Marusik, Y. M., Guseinov, E. F. &amp; Koponen, S. (2003) Spiders (Arachnida: Aranei) of Azerbaijan. 2. Critical survey of wolf spiders (Lycosidae) found in the country with description of three new species and brief review of Palaearctic Evippa Simon, 1885. Arthropoda Selecta, 12 (1), 47 - 65." type="journal article" year="2003">
Marusik
<emphasis id="B945EAD7B700FFCDFDECFCDF65A1E1FA" box="[606,665,873,899]" italics="true" pageId="2" pageNumber="557">et al.</emphasis>
2003
</bibRefCitation>
); South
<collectingCountry id="F3267655B700FFCDFCF0FCDC64A4E1FA" box="[834,924,873,899]" name="Ukraine" pageId="2" pageNumber="557">Ukraine</collectingCountry>
(
<bibRefCitation id="EFA04B34B700FFCDFC1FFCDC63B4E1FA" author="Polchaninova, N. Y." box="[941,1164,873,899]" pageId="2" pageNumber="557" pagination="148 - 149" refId="ref7211" refString="Polchaninova, N. Y. (2001) Role of protected areas for the conservation of steppe araneocomplexes in southern Ukraine. Faltseinnovski chytannia: Zbirnik naukovykh prats. Proceedings of international scientific conference (Kherson, 25 - 27 kvitnia 2001). Kherson: Terra, 148 - 149. [in Russian]" type="book chapter" year="2001">Polchaninova 2001</bibRefCitation>
,
<bibRefCitation id="EFA04B34B700FFCDFB28FCDC63EEE1FA" author="Polchaninova, N. Y." box="[1178,1238,873,899]" pageId="2" pageNumber="557" pagination="85 - 108" refId="ref7268" refString="Polchaninova, N. Y. (2012) A checklist of spiders (Araneae) of the Chernomorskyi Biosphere Reserve (Ukraine). Pryrodnychyi Almanakh. Biolohichni nauky. PAT &quot; Khersonska miska drukarnia &quot;, Kherson, 85 - 108. [in Russian, with English and Ukrai- nian summary]" type="book chapter" year="2012">2012</bibRefCitation>
;
<bibRefCitation id="EFA04B34B700FFCDFB56FCDC665AE1DE" author="Polchaninova, N. Y. &amp; Prokopenko, E. V." pageId="2" pageNumber="557" pagination="269 - 280" refId="ref7323" refString="Polchaninova, N. Y. &amp; Prokopenko, E. V. (2005) History of study and a brief survey of the araneofauna of the Left-Bank Ukraine (Araneae). In: Deltshev, C. &amp; Stoev, P. (Eds.), European Arachnology 2005. Acta Zoologica Bulgarica, Supplement 1, 269 - 280." type="journal article" year="2005">Polchaninova &amp; Prokopenko 2005</bibRefCitation>
,
<bibRefCitation id="EFA04B34B700FFCDFEDDFC386691E1DE" author="Polchaninova, N. Y. &amp; Prokopenko, E. V." box="[367,425,909,935]" pageId="2" pageNumber="557" refId="ref7388" refString="Polchaninova, N. Y. &amp; Prokopenko, E. V. (2011) Spiders (Araneae) of the seaside saline marshes of the Left-Bank Ukraine. Abstracts of 26 th European Congress of Arachnology. Israel, 4 - 8 September 2011, 119." type="journal volume" year="2011">2011</bibRefCitation>
,
<bibRefCitation id="EFA04B34B700FFCDFE04FC3866C9E1DE" author="Polchaninova, N. Y. &amp; Prokopenko, E. V." box="[438,497,909,935]" pageId="2" pageNumber="557" refId="ref7437" refString="Polchaninova, N. Y. &amp; Prokopenko, E. V. (2013) Catalogue of the spiders (Arachnida, Aranei) of Left-Bank Ukraine. Arthropoda Selecta. Supplement 2. KMK Scientific Press, Moscow, 268 pp." type="book" year="2013">2013</bibRefCitation>
,
<bibRefCitation id="EFA04B34B700FFCDFE4FFC386502E1DE" author="Polchaninova, N. Y. &amp; Prokopenko, E. V." box="[509,570,909,935]" pageId="2" pageNumber="557" pagination="60 - 64" refId="ref7481" refString="Polchaninova, N. Y. &amp; Prokopenko, E. V. (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen. 57, 60 - 64. https: // doi. org / 10.30963 / aramit 5711" type="journal article" year="2019">2019</bibRefCitation>
); southern regions of European
<collectingCountry id="F3267655B700FFCDFC15FC3864CAE1DE" box="[935,1010,909,935]" name="Russia" pageId="2" pageNumber="557">Russia</collectingCountry>
(
<bibRefCitation id="EFA04B34B700FFCDFBB0FC3863FFE1DE" author="Ponomarev, A. V." box="[1026,1223,909,935]" pageId="2" pageNumber="557" pagination="96 - 104" refId="ref7531" refString="Ponomarev, A. V. (2002) Spiders (Aranei) of &quot; Rostovskii &quot; reserve (first data of study on fauna). Trudy gosudarstvennogo zapovednika &quot; Rostovskii &quot;, Rostov-on-Don, 1, 96 - 104. [in Russian]" type="journal article" year="2002">Ponomarev 2002</bibRefCitation>
,
<bibRefCitation id="EFA04B34B700FFCDFB66FC386228E1DE" author="Ponomarev, A. V." box="[1236,1296,909,935]" pageId="2" pageNumber="557" pagination="105 - 125" refId="ref7577" refString="Ponomarev, A. V. (2010) Spiders (Arachnida: Aranei) of &quot; Rostovskii &quot; reserve: species inventory and specification of fauna. Problemy izucheniya nazemnykh bespozvonochnykh. Trudy FGU &quot; Gosudarstvennyi prirodnyi zapovednik ' Rostovskii' &quot;, 4, 105 - 125. [in Russian]" type="journal article" year="2010">2010</bibRefCitation>
;
<bibRefCitation id="EFA04B34B700FFCDFAAFFC386666E1B2" author="Ponomarev, A. V. &amp; Tsvetkov, A. S." pageId="2" pageNumber="557" pagination="84 - 104" refId="ref8010" refString="Ponomarev, A. V. &amp; Tsvetkov, A. S. (2004) The generalized data on spiders (Aranei) of the Nature Research &quot; Rostovski &quot;. Trudy Gosudarstvennogo Zapovednika &quot; Rostovskii &quot;, 3, 84 - 104. [in Russian]" type="journal article" year="2004">Ponomarev &amp; Tsvetkov 2004</bibRefCitation>
;
<bibRefCitation id="EFA04B34B700FFCDFEDBFC046553E1B2" author="Ponomarev, A. V. &amp; Belosludtsev, E. A. &amp; Dvadnenko, K. V." box="[361,619,945,971]" pageId="2" pageNumber="557" pagination="163 - 185" refId="ref7697" refString="Ponomarev, A. V., Belosludtsev, E. A. &amp; Dvadnenko, K. V. (2008) Spiders (Aranei) of the Lower Volga Region (Astrakhan and Volgograd areas of Russia) with the description of new taxa. Caucasian Entomological Bulletin, 4 (2), 163 - 185. [in Rus- sian] https: // doi. org / 10.23885 / 1814 - 3326 - 2008 - 4 - 2 - 163 - 185" type="journal article" year="2008">
Ponomarev
<emphasis id="B945EAD7B700FFCDFE42FC076511E1B2" box="[496,553,945,971]" italics="true" pageId="2" pageNumber="557">et al.</emphasis>
2008
</bibRefCitation>
;
<bibRefCitation id="EFA04B34B700FFCDFDC5FC04642DE1B2" author="Kuzmin, E. A." box="[631,789,945,971]" pageId="2" pageNumber="557" pagination="53 - 62" refId="ref6751" refString="Kuzmin, E. A. (2014) Epigeic spiders (Aranei) of salineferous biotopes of Bogdinsko-Baskunchakskii Reserve. Biraznoobrazie aridnykh system. FGBU &quot; Gosudarstvennyi zapovednik &quot; Bogdinsko-Baskunchakskii &quot;. Planeta, Moscow, pp. 53 - 62. [in Russian]" type="book chapter" year="2014">Kuzmin 2014</bibRefCitation>
;
<bibRefCitation id="EFA04B34B700FFCDFC92FC0463FBE1B2" author="Ponomarev, A. V. &amp; Abdurakhmanov, G. M." box="[800,1219,945,971]" pageId="2" pageNumber="557" pagination="76 - 121" refId="ref7630" refString="Ponomarev, A. V. &amp; Abdurakhmanov, G. M. (2014) Spiders (Aranei) of the North Caspian coast and islands. The South of Russia: ecology, development, 1, 76 - 121. [in Russian] https: // doi. org / 10.18470 / 1992 - 1098 - 2014 - 1 - 76 - 121" type="journal article" year="2014">Ponomarev &amp; Abdurakhmanov 2014</bibRefCitation>
;
<bibRefCitation id="EFA04B34B700FFCDFB7CFC04660BE196" author="Ponomarev, A. V. &amp; Prishutova, Z. G." pageId="2" pageNumber="557" pagination="60 - 65" refId="ref7939" refString="Ponomarev, A. V. &amp; Prishutova, Z. G. (2017) Terrestrial spiders (Aranei) of Vodnyi Island (Manych-Gudilo Lake). Science in the South of Russia, 13 (2), 60 - 65. [in Russian] https: // doi. org / 10.23885 / 2500 - 0640 - 2017 - 13 - 2 - 60 - 65" type="journal article" year="2017">Ponomarev &amp; Prishutova 2017</bibRefCitation>
); steppe zone of south of Western Siberia (
<bibRefCitation id="EFA04B34B700FFCDFCA8FC606343E196" author="Fyodorov, I. V. &amp; Trilikauskas, L. A." box="[794,1147,981,1007]" pageId="2" pageNumber="557" pagination="233 - 242" refId="ref6658" refString="Fyodorov, I. V. &amp; Trilikauskas, L. A. (2013) Spider population of a neogenic ecosystem of the salt lake arid zone in Kulunda, West Siberia, Russia. Euroasian Entomological Journal, 12 (3), 233 - 242. [in Russian, with English summary]" type="journal article" year="2013">Fyodorov &amp; Trilikauskas 2013</bibRefCitation>
;
<bibRefCitation id="EFA04B34B700FFCDFB3AFC6062AFE196" author="Mordkovich, V. G. &amp; Dudko, R. Y. &amp; Trilikauskas, L. A. &amp; Lyubechanskii, I. I." box="[1160,1431,980,1007]" pageId="2" pageNumber="557" pagination="447 - 454" refId="ref7072" refString="Mordkovich, V. G., Dudko, R. Y., Trilikauskas, L. A. &amp; Lyubechanskii, I. I. (2015) Carabid beetles (Coleoptera, Carabidae) and spiders (Aranei) are part of the soil fauna of the shore of a salt lake in South Siberia, Russia. Euroasian Entomological Journal, 14 (5), 447 - 454. [in Russian, with English summary]" type="journal article" year="2015">
Mordkovich
<emphasis id="B945EAD7B700FFCDFAA9FC636275E196" box="[1307,1357,981,1007]" italics="true" pageId="2" pageNumber="557">et al</emphasis>
. 2015
</bibRefCitation>
;
<bibRefCitation id="EFA04B34B700FFCDFF25FC4C6693E66A" author="Azarkina, G. N. &amp; Lyubechanskii, I. I. &amp; Trilikauskas, L. A. &amp; Dudko, R. Yu. &amp; Bespalov, A. N. &amp; Mordkovich, V. G." box="[151,427,1017,1043]" pageId="2" pageNumber="557" pagination="73 - 93" refId="ref6254" refString="Azarkina, G. N., Lyubechanskii, I. I., Trilikauskas, L. A., Dudko, R. Yu., Bespalov, A. N. &amp; Mordkovich, V. G. (2018) A check-list and zoogeographic analysis of the spider fauna (Arachnida: Aranei) of Novosibirsk Area (West Siberia, Russia). Arthropoda Selecta, 27 (1), 73 - 93. https: // doi. org / 10.15298 / arthsel. 27.1.11" type="journal article" year="2018">
Azarkina
<emphasis id="B945EAD7B700FFCDFEB7FC4F660FE66A" box="[261,311,1017,1043]" italics="true" pageId="2" pageNumber="557">et al</emphasis>
. 2018: 84
</bibRefCitation>
),
<collectingCountry id="F3267655B700FFCDFE72FC4C657BE66A" box="[448,579,1017,1043]" name="Kazakhstan" pageId="2" pageNumber="557">Kazakhstan</collectingCountry>
(
<bibRefCitation id="EFA04B34B700FFCDFDE0FC4C65C8E66A" author="Bragina, T. M." box="[594,752,1017,1043]" pageId="2" pageNumber="557" refId="ref6346" refString="Bragina, T. M. (2012) Some data about the fauna of spiders (arachnida, Aranei) of Naurzum Reserve. The materials of the sixth international symposium and of the eighth international school-seminar of young scientists &quot; Geoenviromental problems of the steppe regions &quot;. Orenburg, IPK &quot; Gazprompechat &quot; OOO &quot; Orenburggazpromservis &quot;, 940 pp." type="book" year="2012">Bragina 2012</bibRefCitation>
;
<bibRefCitation id="EFA04B34B700FFCDFD49FC4C6302E66A" author="Ponomarev, A. V. &amp; Bragina, T. M." box="[763,1082,1016,1043]" pageId="2" pageNumber="557" pagination="46 - 57" refId="ref7784" refString="Ponomarev, A. V. &amp; Bragina, T. M. (2015) Preliminary data on spider fauna (Aranei) of protected areas in Kostanay Region (Kazakhstan). The South of Russia: ecology, development, 4, 46 - 57. [in Russian] https: // doi. org / 10.18470 / 1992 - 1098 - 2014 - 4 - 46 - 57" type="journal article" year="2015">Ponomarev &amp; Bragina 2015</bibRefCitation>
; Ponomarev
<emphasis id="B945EAD7B700FFCDFB7FFC4F623EE66A" box="[1229,1286,1017,1043]" italics="true" pageId="2" pageNumber="557">et al.</emphasis>
2017);
<collectingCountry id="F3267655B700FFCDFAEFFC4C67C4E64E" name="Turkmenistan" pageId="2" pageNumber="557">Turkmenistan</collectingCountry>
(L.
<bibRefCitation id="EFA04B34B700FFCDFE9FFBA86696E64E" author="Koch, L." box="[301,430,1053,1079]" pageId="2" pageNumber="557" pagination="36 - 71" refId="ref6715" refString="Koch, L. (1878) Kaukasische Arachnoiden. In: Schneider, O. (Ed.), Naturwissenschaftliche Beitrage zur Kenntniss der Kaukasuslander. Dresden, 3, 36 - 71." type="journal article" year="1878">Koch 1878</bibRefCitation>
);
<collectingCountry id="F3267655B700FFCDFE70FBA8650CE64E" box="[450,564,1053,1079]" name="Tajikistan" pageId="2" pageNumber="557">Tajikistan</collectingCountry>
;
<collectingCountry id="F3267655B700FFCDFDF2FBA865BBE64E" box="[576,643,1053,1079]" name="China" pageId="2" pageNumber="557">China</collectingCountry>
(
<collectingRegion id="49F5F827B700FFCDFD20FBA8647EE641" box="[658,838,1053,1080]" country="China" name="Nei Mongol" pageId="2" pageNumber="557">Inner Mongolia</collectingRegion>
) (
<bibRefCitation id="EFA04B34B700FFCDFCEEFBA86345E64E" author="Urita &amp; Tang, G. M. &amp; Song, D. X." box="[860,1149,1053,1079]" pageId="2" pageNumber="557" pagination="46 - 49" refId="ref8274" refString="Urita, Tang, G. M. &amp; Song, D. X. (1993) Two new species of the genus Pardosa from Inner Mongolia, China (Araneae, Lycosidae). Journal of Inner Mongolia Normal University (Nat. Sci. Ed.), 3, 46 - 49. [in Chinese and English]" type="journal article" year="1993">Urita, Tang et Song 1993</bibRefCitation>
, sub.
<taxonomicName id="4C314D46B700FFCDFB0DFBA862A4E64E" authorityName="Urita, Tang &amp; Song" authorityYear="1993" box="[1215,1436,1053,1079]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="557" phylum="Arthropoda" rank="species" species="hatanensis">
<emphasis id="B945EAD7B700FFCDFB0DFBA862A4E64E" box="[1215,1436,1053,1079]" italics="true" pageId="2" pageNumber="557">Pardosa hatanensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C314D46B700FFCDFF75FBF76615E622" authorityName="Azarkina &amp; Trilikauskas" authorityYear="2019" baseAuthorityName="Azarkina &amp; Trilikauskas" baseAuthorityYear="2019" box="[199,301,1090,1115]" class="Arachnida" family="Lycosidae" genus="Halocosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="557" phylum="Arthropoda" rank="species" species="jartica">
<emphasis id="B945EAD7B700FFCDFF75FBF76615E622" box="[199,301,1090,1115]" italics="true" pageId="2" pageNumber="557">P. jartica</emphasis>
</taxonomicName>
) (Figure 72).
</paragraph>
<paragraph id="8B8E36C5B700FFCDFF75FBD166FBE6DD" blockId="2.[151,1437,152,1188]" pageId="2" pageNumber="557">
<emphasis id="B945EAD7B700FFCDFF75FBD16616E606" bold="true" box="[199,302,1124,1151]" pageId="2" pageNumber="557">Habitats</emphasis>
. The species occurs on (tidal) salt marshes along sea/lake shores, where spiders live in burrows made in sandy soil (Figs 6871).
</paragraph>
</subSubSection>
</treatment>
</document>