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<document id="485D704317529A4260211609C351A0E9" ID-CLB-Dataset="25520" ID-DOI="10.11646/zootaxa.4671.2.10" ID-GBIF-Dataset="0010c4e6-f37b-40c6-a2fc-f03234cd0a1e" ID-ISSN="1175-5326" ID-Zenodo-Dep="3442338" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1568800482310" checkinUser="plazi" docAuthor="Frolov, Andrey, Akhmetova, Lilia &amp; Montreuil, Olivier" docDate="2019" docId="0392500BFFABB55BFF3F85D124FFAC32" docLanguage="en" docName="zootaxa.4671.2.10.pdf" docOrigin="Zootaxa 4671 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hybalus servulus Normand 1949" docType="treatment" docVersion="8" lastPageNumber="293" masterDocId="FFAB2873FFA8B558FFA885482738AE06" masterDocTitle="A review of Hybalus Dejean from Tunisia (Coleoptera: Scarabaeidae: Orphninae) with description of a new species" masterLastPageNumber="294" masterPageNumber="290" pageNumber="293" updateTime="1698755172623" updateUser="ExternalLinkService">
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<mods:title id="82BD1797959194026AB3084E61DF0753">A review of Hybalus Dejean from Tunisia (Coleoptera: Scarabaeidae: Orphninae) with description of a new species</mods:title>
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<mods:namePart id="FDBC9864DC91D2FACF88F21289B122E6">Frolov, Andrey</mods:namePart>
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<mods:namePart id="90816FD5182049BB34D20B98DC7A40C2">Akhmetova, Lilia</mods:namePart>
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<paragraph id="8B84E11DFFABB55BFF3F85D1251AAEB5" blockId="3.[151,1437,153,564]" box="[151,546,153,180]" pageId="3" pageNumber="293">
<heading id="D0CC5671FFABB55BFF3F85D1251AAEB5" bold="true" box="[151,546,153,180]" fontSize="11" level="1" pageId="3" pageNumber="293" reason="1">
<taxonomicName id="4C3B9A9EFFABB55BFF3F85D1251AAEB5" authority="Normand, 1949" authorityName="Normand" authorityYear="1949" box="[151,546,153,180]" class="Insecta" family="Orphnidae" genus="Hybalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="species" species="servulus">
<emphasis id="B94F3D0FFFABB55BFF3F85D1251AAEB5" bold="true" box="[151,546,153,180]" pageId="3" pageNumber="293">
<emphasis id="B94F3D0FFFABB55BFF3F85D1265CAEB2" bold="true" box="[151,356,153,180]" italics="true" pageId="3" pageNumber="293">Hybalus servulus</emphasis>
Normand, 1949
</emphasis>
</taxonomicName>
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<subSubSection id="C321B296FFABB55BFF3F85F22463AEF6" pageId="3" pageNumber="293" type="reference_group">
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<taxonomicName id="4C3B9A9EFFABB55BFF3F85F2253BAED4" authority="Normand, 1949: 86" authorityName="Normand" authorityPageNumber="86" authorityYear="1949" box="[151,515,185,210]" class="Insecta" family="Orphnidae" genus="Hybalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="species" species="servulus">
<emphasis id="B94F3D0FFFABB55BFF3F85F2267AAED4" box="[151,322,186,210]" italics="true" pageId="3" pageNumber="293">Hybalus servulus</emphasis>
Normand, 1949: 86
</taxonomicName>
</paragraph>
<paragraph id="8B84E11DFFABB55BFF3F85902463AEF6" blockId="3.[151,1437,153,564]" box="[151,859,216,241]" pageId="3" pageNumber="293">
<treatmentCitationGroup id="AB2BC633FFABB55BFF3F85902463AEF6" box="[151,859,216,241]" pageId="3" pageNumber="293">
<taxonomicName id="4C3B9A9EFFABB55BFF3F85902532AEF7" authority="Baraud, 1980: 61" authorityName="Baraud" authorityPageNumber="61" authorityYear="1980" box="[151,522,216,241]" class="Insecta" family="Orphnidae" genus="Hybalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="species" species="demoflysi">
<emphasis id="B94F3D0FFFABB55BFF3F8590266BAEF6" box="[151,339,216,240]" italics="true" pageId="3" pageNumber="293">Hybalus demoflysi</emphasis>
<treatmentCitation id="0A9AC70CFFABB55BFEF185902532AEF7" author="Baraud, J." box="[345,522,216,241]" page="61" pageId="3" pageNumber="293" year="1980">
<bibRefCitation id="EFAA9CECFFABB55BFEF185902532AEF7" author="Baraud, J." box="[345,522,216,241]" pageId="3" pageNumber="293" pagination="59 - 62" refId="ref2142" refString="Baraud, J. (1980) Les Hybalus Brulle de Tunisie. Nouvelle Revue d'Entomologie, 10, 59 - 62." type="journal article" year="1980">Baraud, 1980: 61</bibRefCitation>
</treatmentCitation>
</taxonomicName>
, synonymy by
<treatmentCitation id="0A9AC70CFFABB55BFD0D8590246FAEF6" author="Baraud, J." box="[677,855,216,241]" page="23" pageId="3" pageNumber="293" year="1991">
<bibRefCitation id="EFAA9CECFFABB55BFD0D8590246FAEF6" author="Baraud, J." box="[677,855,216,241]" pageId="3" pageNumber="293" pagination="3 - 28" refId="ref2165" refString="Baraud, J. (1991) Revision du genre Hybalus Brulle, 1834 (Coleoptera, Scarabaeoidea, Orphninae). Annales de la Societe Entomologique de France, New Series, 27, 3 - 28." type="journal article" year="1991">Baraud, 1991: 23</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C321B296FFABB55BFF6F85B524FFAC32" pageId="3" pageNumber="293" type="materials_examined">
<paragraph id="8B84E11DFFABB55BFF6F85B524A7AF86" blockId="3.[151,1437,153,564]" pageId="3" pageNumber="293">
<emphasis id="B94F3D0FFFABB55BFF6F85B526EFAF12" bold="true" box="[199,471,252,277]" pageId="3" pageNumber="293">Type material examined.</emphasis>
<taxonomicName id="4C3B9A9EFFABB55BFE4985B425A6AF12" authorityName="Baraud" authorityYear="1980" box="[481,670,252,276]" class="Insecta" family="Orphnidae" genus="Hybalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="species" species="demoflysi">
<emphasis id="B94F3D0FFFABB55BFE4985B425A6AF12" box="[481,670,252,276]" italics="true" pageId="3" pageNumber="293">Hybalus demoflysi</emphasis>
</taxonomicName>
:
<materialsCitation id="3B53EB40FFABB55BFD0685B4222AAF12" ID-GBIF-Occurrence="2413160972" box="[686,1298,252,276]" collectingDate="1939-02" collectionCode="MNHN" collectorName="R. Demoflys" pageId="3" pageNumber="293" specimenCount="1" typeStatus="holotype">
<typeStatus id="54805FBFFFABB55BFD0685B42430AF12" box="[686,776,252,276]" pageId="3" pageNumber="293" type="holotype">holotype</typeStatus>
, male: “Sidi Tabet,
<date id="FF85C7DDFFABB55BFC7185B42333AF12" box="[985,1035,252,276]" pageId="3" pageNumber="293" value="1939-02">
<collectingDate id="EFC13E35FFABB55BFC7185B42333AF12" box="[985,1035,252,276]" pageId="3" pageNumber="293" value="1939-02">II.39</collectingDate>
</date>
,
<collectorName id="26CE84CBFFABB55BFBB085B523A7AF12" box="[1048,1183,252,276]" pageId="3" pageNumber="293">R. Démoflys</collectorName>
” (
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)
</materialsCitation>
,
<typeStatus id="54805FBFFFABB55BFAB785B522BBAF13" box="[1311,1411,253,277]" pageId="3" pageNumber="293" type="paratype">paratypes</typeStatus>
: 1 male and
<specimenCount id="9D3D2A94FFABB55BFF528468266AAF3E" box="[250,338,288,312]" pageId="3" pageNumber="293" type="female">1 female</specimenCount>
with the same data (
<collectionCode id="ED2A79D8FFABB55BFD8A8468255DAF3E" box="[546,613,288,312]" pageId="3" pageNumber="293">CJBH</collectionCode>
). The
<typeStatus id="54805FBFFFABB55BFD0B846925E0AF3F" box="[675,728,289,313]" pageId="3" pageNumber="293">types</typeStatus>
of
<taxonomicName id="4C3B9A9EFFABB55BFD5284692455AF3E" authorityName="Normand" authorityYear="1949" box="[762,877,288,312]" class="Insecta" family="Orphnidae" genus="Hybalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="species" species="servulus">
<emphasis id="B94F3D0FFFABB55BFD5284692455AF3E" box="[762,877,288,312]" italics="true" pageId="3" pageNumber="293">H. servulus</emphasis>
</taxonomicName>
are not available for examination (Synda Kheder, per- sonal communication) therefore we follow Barauds interpretation of this species. The photographs of the specimens from the Normands collection, provided by Synda Kheder, support this interpretation.
</paragraph>
<paragraph id="8B84E11DFFABB55BFF6F84C426F9AFEA" blockId="3.[151,1437,153,564]" pageId="3" pageNumber="293">
<emphasis id="B94F3D0FFFABB55BFF6F84C4252AAFA2" bold="true" box="[199,530,396,420]" pageId="3" pageNumber="293">Additional material examined.</emphasis>
<materialsCitation id="3B53EB40FFABB55BFDB184C423B9AFA2" ID-GBIF-Occurrence="2413160965" box="[537,1153,396,420]" collectingDate="2012-03-04" collectionCode="ZIN" collectorName="Djebba, W" country="Tunisia" location="Beja" pageId="3" pageNumber="293" specimenCount="2" specimenCount-male="2" stateProvince="Beja">
<emphasis id="B94F3D0FFFABB55BFDB184C42548AFA2" bold="true" box="[537,624,396,420]" pageId="3" pageNumber="293">
<collectingCountry id="F32CA18DFFABB55BFDB184C42554AFA2" box="[537,620,396,420]" name="Tunisia" pageId="3" pageNumber="293">Tunisia</collectingCountry>
:
</emphasis>
<collectingRegion id="49FF2FFFFFABB55BFDDF84C42590AFA2" box="[631,680,396,420]" country="Tunisia" name="Beja" pageId="3" pageNumber="293">Beja</collectingRegion>
,
<collectorName id="26CE84CBFFABB55BFD1B84C4241AAFA2" box="[691,802,396,420]" pageId="3" pageNumber="293">Djebba, W</collectorName>
. Ziegler
<date id="FF85C7DDFFABB55BFCD484C424D9AFA2" box="[892,993,396,420]" pageId="3" pageNumber="293" value="2012-03-04">
<collectingDate id="EFC13E35FFABB55BFCD484C424D9AFA2" box="[892,993,396,420]" pageId="3" pageNumber="293" value="2012-03-04">4.III.2012</collectingDate>
</date>
,
<specimenCount id="9D3D2A94FFABB55BFC4584C42304AFA2" box="[1005,1084,396,420]" pageId="3" pageNumber="293" type="male">2 males</specimenCount>
(
<collectionCode id="ED2A79D8FFABB55BFBE384C42342AFA2" box="[1099,1146,396,420]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="3" pageNumber="293" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B53EB40FFABB55BFB2584C422A1AFA2" ID-GBIF-Occurrence="2413160971" box="[1165,1433,396,420]" collectorName="Teboursouk Pass, W" country="Tunisia" location="Beja" pageId="3" pageNumber="293" specimenCount="1" stateProvince="Beja">
<collectingRegion id="49FF2FFFFFABB55BFB2584C42386AFA2" box="[1165,1214,396,420]" country="Tunisia" name="Beja" pageId="3" pageNumber="293">Beja</collectingRegion>
,
<collectorName id="26CE84CBFFABB55BFB6084C422A1AFA2" box="[1224,1433,396,420]" pageId="3" pageNumber="293">Teboursouk Pass, W</collectorName>
</materialsCitation>
. Ziegler
<date id="FF85C7DDFFABB55BFF4E84F82606AFCE" box="[230,318,432,456]" pageId="3" pageNumber="293" value="2012-03-01">1.3.2012</date>
/
<geoCoordinate id="EE0F87DAFFABB55BFEFA84E726F0AFCF" box="[338,456,431,457]" degrees="36" direction="north" minutes="29" orientation="latitude" pageId="3" pageNumber="293" precision="15" seconds="48" value="36.496666">N36°29ʹ48ʺ</geoCoordinate>
<geoCoordinate id="EE0F87DAFFABB55BFE6784F92571AFCF" box="[463,585,431,457]" degrees="09" direction="east" minutes="09" orientation="longitude" pageId="3" pageNumber="293" precision="15" seconds="38" value="9.160555">E 09°09ʹ38ʺ</geoCoordinate>
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Pass,
<specimenCount id="9D3D2A94FFABB55BFD6784F82425AFCE" box="[719,797,432,456]" pageId="3" pageNumber="293" type="male">2 males</specimenCount>
and
<specimenCount id="9D3D2A94FFABB55BFCE684F82497AFCE" box="[846,943,432,456]" pageId="3" pageNumber="293" type="female">2 females</specimenCount>
(
<collectionCode id="ED2A79D8FFABB55BFC1584F824C3AFCE" box="[957,1019,432,456]" pageId="3" pageNumber="293">CWZ</collectionCode>
);
<materialsCitation id="3B53EB40FFABB55BFBA584F822A1AFCE" ID-GBIF-Occurrence="2413160969" box="[1037,1433,432,456]" elevation="850" location="Ort" pageId="3" pageNumber="293" specimenCount="1" stateProvince="Kef">
<collectingRegion id="49FF2FFFFFABB55BFBA584F82369AFCE" box="[1037,1105,432,456]" country="Tunisia" name="Kef" pageId="3" pageNumber="293">El Kef</collectingRegion>
,
<quantity id="4CC34CF8FFABB55BFBF484F823A5AFCE" box="[1116,1181,432,456]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="293" unit="km" value="10.0">10 km</quantity>
NO vom
<location id="8EE4B7C6FFABB55BFAAA84F8221EAFCE" LSID="urn:lsid:plazi:treatment:0392500BFFABB55BFF3F85D124FFAC32:8EE4B7C6FFABB55BFAAA84F8221EAFCE" box="[1282,1318,432,456]" name="Ort" pageId="3" pageNumber="293" stateProvince="Kef">Ort</location>
,
<quantity id="4CC34CF8FFABB55BFA9884F82249AFCE" box="[1328,1393,432,456]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="3" pageNumber="293" unit="m" value="850.0">
<elevation id="0016062EFFABB55BFA9884F82249AFCE" box="[1328,1393,432,456]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="3" pageNumber="293" unit="m" value="850.0">850 m</elevation>
</quantity>
leg
</materialsCitation>
. Esser
<date id="FF85C7DDFFABB55BFF7C849C2619AFEA" box="[212,289,468,492]" pageId="3" pageNumber="293" value="13-03-26">26.3.13</date>
,
<specimenCount id="9D3D2A94FFABB55BFE83849C2641AFEA" box="[299,377,468,492]" pageId="3" pageNumber="293" type="male">2 males</specimenCount>
(
<collectionCode id="ED2A79D8FFABB55BFE2E849C268DAFEA" box="[390,437,468,492]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="3" pageNumber="293" type="Museum">ZIN</collectionCode>
).
</paragraph>
<paragraph id="8B84E11DFFABB55BFF6F84B024FFAC32" blockId="3.[151,1437,153,564]" pageId="3" pageNumber="293">
This species was described from Fondouk-Djedid and known also from Sidi Tabet, Djebba, Teboursouk Pass and
<collectingRegion id="49FF2FFFFFABB55BFAFF84B022A5AC16" box="[1367,1437,504,528]" country="Tunisia" name="Kef" pageId="3" pageNumber="293">El Kef</collectingRegion>
(
<figureCitation id="1300FD98FFABB55BFF37875427E4AC32" box="[159,220,540,564]" captionStart="FIGURE 2" captionStartId="2.[151,250,1939,1964]" captionTargetBox="[156,1427,997,1907]" captionTargetId="figure@2.[151,1435,992,1914]" captionTargetPageId="2" captionText="FIGURE 2. Hybalus Dejean, 1833 spp. Map of locality records in Tunisia." figureDoi="http://doi.org/10.5281/zenodo.3442342" httpUri="https://zenodo.org/record/3442342/files/figure.png" pageId="3" pageNumber="293">Fig. 2</figureCitation>
). Record from Radès (
<bibRefCitation id="EFAA9CECFFABB55BFE178754257AAC32" author="Baraud, J." box="[447,578,540,564]" pageId="3" pageNumber="293" pagination="3 - 28" refId="ref2165" refString="Baraud, J. (1991) Revision du genre Hybalus Brulle, 1834 (Coleoptera, Scarabaeoidea, Orphninae). Annales de la Societe Entomologique de France, New Series, 27, 3 - 28." type="journal article" year="1991">Baraud 1991</bibRefCitation>
) belongs to
<taxonomicName id="4C3B9A9EFFABB55BFD1687552479AC32" authority="Frolov &amp; Akhmetova &amp; Montreuil, 2019" authorityName="Frolov &amp; Akhmetova &amp; Montreuil" authorityYear="2019" box="[702,833,540,564]" class="Insecta" family="Orphnidae" genus="Hybalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="293" phylum="Arthropoda" rank="species" species="bellmanni" status="sp. nov.">
<emphasis id="B94F3D0FFFABB55BFD1687552479AC32" box="[702,833,540,564]" italics="true" pageId="3" pageNumber="293">H. bellmanni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C8074FFABB55BFCEF875524FBAC32" box="[839,963,540,564]" pageId="3" pageNumber="293" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
</treatment>
</document>