571 lines
73 KiB
XML
571 lines
73 KiB
XML
<document id="B7D1C57EE6B814BB88F6CDA75BD3BE5A" ID-CLB-Dataset="45873" ID-DOI="10.5281/zenodo.206642" ID-GBIF-Dataset="db6cee80-6ba7-427f-9f9a-ce86dfb84f6b" ID-ISSN="1175-5326" ID-Zenodo-Dep="206642" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460415443680" checkinUser="plazi" docAuthor="Hartke, Tamara R., Koenemann, Stefan & Yager, Jill" docDate="2011" docId="039234658D1DFE4EDDE50A10FBDBBCE0" docLanguage="en" docName="zt03115p028.pdf" docOrigin="Zootaxa 3115" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Speleonectes williamsi Hartke, Koenemann & Yager, 2011, new species" docType="treatment" docVersion="11" lastPageNumber="26" masterDocId="FFAB4C1D8D1DFE4BDD720D0EFFDDBB14" masterDocTitle="Speleonectes williamsi, a new species of Remipedia (Crustacea) from the Bahamas" masterLastPageNumber="28" masterPageNumber="21" pageNumber="21" updateTime="1698328676377" updateUser="ExternalLinkService">
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<mods:mods id="8C8105857175F3598035E288753612A7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="3E562A2684320C089E3EA86E2E280039">
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<mods:title id="3CED9010532138A2062848479F6F6E5F">Speleonectes williamsi, a new species of Remipedia (Crustacea) from the Bahamas</mods:title>
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</mods:titleInfo>
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<mods:name id="C51F9880C8E0328176379CEC7DAA82F6" type="personal">
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<mods:role id="FBE07277C4B542817B6A100B47335D66">
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<mods:roleTerm id="5DFDCF43B7EFF9646A4EFB60DC77A554">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E6CCA34AE2FA34D746C66E5BE427D594">Hartke, Tamara R.</mods:namePart>
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</mods:name>
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<mods:name id="B92F64881E1E1FDC81C84CAC7BB2FA0E" type="personal">
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<mods:role id="677A503DED134F59994B16687BE7EDB0">
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<mods:roleTerm id="302621A197C8960633DC4EBE08B60390">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A6564BBC261DAD1B4694625A3A6F6CDF">Koenemann, Stefan</mods:namePart>
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</mods:name>
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<mods:name id="469D38A6675F594C316D23066DF39B59" type="personal">
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<mods:role id="698E48C5A1C7460B832A6A1E30B4563C">
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<mods:roleTerm id="DF004CDC215A6B2B435037E3CC823607">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3E0BFE02A0D6733E35EEB1C22E72C18B">Yager, Jill</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="9F3186C97299435E8A24B620436F74FB">text</mods:typeOfResource>
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<mods:relatedItem id="0735892D62A77734C9A95847EC319219" type="host">
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<mods:titleInfo id="42922D1D863E5E6CACC97B6C59B6B6A9">
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<mods:title id="5293ABB0909FCAF1AB9AF005FA77DCBA">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="D6767B55CFF6C995C25BEADA1FEF9689">
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<mods:date id="0C39B993A88F3104E43169892B340DEF">2011</mods:date>
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<mods:detail id="1DD4AF293FBACD2A04D21C665EB68DB1" type="volume">
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<mods:number id="711253A30B0FCF6976C3B9FB6A63212A">3115</mods:number>
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</mods:detail>
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<mods:extent id="995885B523FF16459929347D66C4250A" unit="page">
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<mods:start id="5BF245E892CBD15D756D260A3AD97A3F">21</mods:start>
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<mods:end id="F0C7A1743F6300E7FA21EC8557B96A32">28</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="4A882355D9C35201F5F25A554C4C31F1">journal article</mods:classification>
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<mods:identifier id="BAB691EDD6C3E379D28F9570F29C97FA" type="CLB-Dataset">45873</mods:identifier>
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<mods:identifier id="B71D77C3918290F954C3BA62DF8FE6A8" type="DOI">10.5281/zenodo.206642</mods:identifier>
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<mods:identifier id="DF947635BCCF7CB0544B1D4E4D112391" type="GBIF-Dataset">db6cee80-6ba7-427f-9f9a-ce86dfb84f6b</mods:identifier>
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<mods:identifier id="A428A92D9347B1DB730A8F0EEDFD6247" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="1506732AA007E4D558A9E0E7FA429F4B" type="Zenodo-Dep">206642</mods:identifier>
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</mods:mods>
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<treatment id="039234658D1DFE4EDDE50A10FBDBBCE0" ID-DOI="http://doi.org/10.5281/zenodo.5685649" ID-GBIF-Taxon="119424783" ID-Zenodo-Dep="5685649" LSID="urn:lsid:plazi:treatment:039234658D1DFE4EDDE50A10FBDBBCE0" httpUri="http://treatment.plazi.org/id/039234658D1DFE4EDDE50A10FBDBBCE0" lastPageId="5" lastPageNumber="26" pageId="0" pageNumber="21">
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<subSubSection id="C321D6F88D1DFE4BDDE50A10FD99BC2C" box="[151,580,1822,1848]" pageId="0" pageNumber="21" type="nomenclature">
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||
<paragraph id="8B8485738D1DFE4BDDE50A10FD99BC2C" blockId="0.[151,580,1822,1848]" box="[151,580,1822,1848]" pageId="0" pageNumber="21">
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<heading id="D0CC321F8D1DFE4BDDE50A10FD99BC2C" bold="true" box="[151,580,1822,1848]" fontSize="11" level="1" pageId="0" pageNumber="21" reason="1">
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||
<emphasis id="B94F59618D1DFE4BDDE50A10FD99BC2C" bold="true" box="[151,580,1822,1848]" pageId="0" pageNumber="21">
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||
<taxonomicName id="4C3BFEF08D1DFE4BDDE50A10FE7BBC2C" box="[151,422,1822,1848]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="williamsi" status="sp. nov.">
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<emphasis id="B94F59618D1DFE4BDDE50A10FE7BBC2C" bold="true" box="[151,422,1822,1848]" italics="true" pageId="0" pageNumber="21">Speleonectes williamsi</emphasis>
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||
</taxonomicName>
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||
,
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<taxonomicNameLabel id="A27CE41A8D1DFE4BDCC70A10FD99BC2C" box="[437,580,1822,1848]" pageId="0" pageNumber="21" rank="species">new species</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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</subSubSection>
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||
<subSubSection id="C321D6F88D1DFE4ADDE50A6BFDD5BA08" lastPageId="1" lastPageNumber="22" pageId="0" pageNumber="21" type="materials_examined">
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||
<paragraph id="8B8485738D1DFE4BDDE50A6BFB25BC68" blockId="0.[151,1437,1892,2025]" box="[151,1272,1892,1917]" pageId="0" pageNumber="21">
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||
<emphasis id="B94F59618D1DFE4BDDE50A6BFEE8BC69" bold="true" box="[151,309,1892,1917]" pageId="0" pageNumber="21">
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<typeStatus id="54803BD18D1DFE4BDDE50A6BFF0FBC69" box="[151,210,1893,1917]" pageId="0" pageNumber="21">Type</typeStatus>
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locality.
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</emphasis>
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Sagittarius Cave, Sweetings Cay (
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<geoCoordinate id="EE0FE3B48D1DFE4BDFCB0A6AFCD4BC68" box="[697,777,1892,1916]" direction="north" orientation="latitude" pageId="0" pageNumber="21" precision="5555" value="26.9">N 26.9</geoCoordinate>
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, W -77.8), Grand
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<collectingCountry id="F32CC5E38D1DFE4BDEA60A6AFBEDBC68" box="[980,1072,1892,1916]" name="Bahamas" pageId="0" pageNumber="21">Bahama</collectingCountry>
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Island,
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<collectingCountry id="F32CC5E38D1DFE4BD9FB0A6AFB2EBC68" box="[1161,1267,1892,1916]" name="Bahamas" pageId="0" pageNumber="21">Bahamas</collectingCountry>
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.
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</paragraph>
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<paragraph id="8B8485738D1DFE4ADDB50A87FDD5BA08" blockId="0.[151,1437,1892,2025]" lastBlockId="1.[151,1437,151,860]" lastPageId="1" lastPageNumber="22" pageId="0" pageNumber="21">
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<emphasis id="B94F59618D1DFE4BDDB50A87FEA8BCB5" bold="true" box="[199,373,1928,1953]" pageId="0" pageNumber="21">
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<typeStatus id="54803BD18D1DFE4BDDB50A87FEDFBCB5" box="[199,258,1929,1953]" pageId="0" pageNumber="21">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54803BD18D1DFE4BDC0E0A86FE39BCB4" box="[380,484,1928,1952]" pageId="0" pageNumber="21" type="holotype">Holotype</typeStatus>
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(
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<collectingCountry id="F32CC5E38D1DFE4BDC810A86FDC5BCB4" box="[499,536,1928,1952]" name="United States of America" pageId="0" pageNumber="21">US</collectingCountry>
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Natural History Museum 1155294),
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<quantity id="4CC328968D1DFE4BDECB0A86FBCEBCB4" box="[953,1043,1928,1952]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="0" pageNumber="21" unit="mm" value="8.9">8.9 mm</quantity>
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.
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<typeStatus id="54803BD18D1DFE4BD96C0A87FB5DBCB5" box="[1054,1152,1929,1953]" pageId="0" pageNumber="21" type="paratype">Paratype</typeStatus>
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1 (private collection SK, ID: BH 51),
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<quantity id="4CC328968D1DFE4BDC540AA2FE50BCD0" box="[294,397,1964,1988]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="0" pageNumber="21" unit="mm" value="10.2">10.2 mm</quantity>
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||
, dissected for description.
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<typeStatus id="54803BD18D1DFE4BDFC90AA3FCC0BCD1" box="[699,797,1965,1989]" pageId="0" pageNumber="21" type="paratype">Paratype</typeStatus>
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2 (private collection SK, ID: BH 52),
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<quantity id="4CC328968D1DFE4BD9A30AA2FAF4BCD0" box="[1233,1321,1964,1988]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="0" pageNumber="21" unit="mm" value="8.5">8.5 mm</quantity>
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||
, dissected for description.
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<typeStatus id="54803BD18D1DFE4BDC380ADFFE71BCFD" box="[330,428,2001,2025]" pageId="0" pageNumber="21" type="paratype">Paratype</typeStatus>
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3 (private collection SK, ID: BH 53), 8.0 mm.
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<typeStatus id="54803BD18D1DFE4BDEB60ADFFBFBBCFD" box="[964,1062,2001,2025]" pageId="0" pageNumber="21" type="paratype">Paratype</typeStatus>
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4 (ID: BH 54),
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<quantity id="4CC328968D1DFE4BD9A90ADEFAE9BCFC" box="[1243,1332,2000,2024]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="0" pageNumber="21" unit="mm" value="8.1">8.1 mm</quantity>
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||
, used for
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<collectionCode id="ED2A1DB68D1CFE4ADDE50D96FF09BBA4" LSID="urn:lsid:biocol.org:col:15536" box="[151,212,152,176]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="1" pageNumber="22">DNA</collectionCode>
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extraction.
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<typeStatus id="54803BD18D1CFE4ADC2A0D96FE67BBA4" box="[344,442,152,176]" pageId="1" pageNumber="22" type="paratype">Paratype</typeStatus>
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5 (private collection SK, ID: BH 55), 9.0 mm.
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<typeStatus id="54803BD18D1CFE4ADEBA0D96FBF7BBA4" box="[968,1066,152,176]" pageId="1" pageNumber="22" type="paratype">Paratype</typeStatus>
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6 (
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<collectionCode id="ED2A1DB68D1CFE4AD93F0D99FB69BBBB" box="[1101,1204,151,175]" pageId="1" pageNumber="22">USNHM</collectionCode>
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||
1155295),
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||
<quantity id="4CC328968D1CFE4AD8400D99FA45BBBB" box="[1330,1432,151,175]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="1" pageNumber="22" unit="mm" value="11.2">11.2 mm</quantity>
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.
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<typeStatus id="54803BD18D1CFE4ADDE50DB3FF24BBC1" box="[151,249,189,213]" pageId="1" pageNumber="22" type="paratype">Paratype</typeStatus>
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7 (
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<collectionCode id="ED2A1DB68D1CFE4ADC500DB2FE54BBC0" box="[290,393,188,212]" pageId="1" pageNumber="22">USNHM</collectionCode>
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||
1155296),
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||
<quantity id="4CC328968D1CFE4ADF7C0DB2FDAABBC0" box="[526,631,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="1" pageNumber="22" unit="mm" value="10.6">10.6 mm</quantity>
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, 20 trunk segments.
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<typeStatus id="54803BD18D1CFE4ADE1A0DB3FC08BBC1" box="[872,981,189,213]" pageId="1" pageNumber="22" type="paratype">Paratypes</typeStatus>
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||
1–5 with 22 trunk segments;
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<typeStatus id="54803BD18D1CFE4AD8410DB2FA45BBC0" box="[1331,1432,188,212]" pageId="1" pageNumber="22" type="holotype">holotype</typeStatus>
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||
,
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<typeStatus id="54803BD18D1CFE4ADDE50DEEFEDFBBEC" box="[151,258,224,248]" pageId="1" pageNumber="22" type="paratype">paratypes</typeStatus>
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6 and 7 with 20 trunk segments.
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<typeStatus id="54803BD18D1CFE4ADF040DD1FD03BBE3" box="[630,734,223,247]" pageId="1" pageNumber="22" type="holotype">Holotype</typeStatus>
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and
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<typeStatus id="54803BD18D1CFE4ADE670DEEFC5DBBEC" box="[789,896,224,248]" pageId="1" pageNumber="22" type="paratype">paratypes</typeStatus>
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were collected from the
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<typeStatus id="54803BD18D1CFE4AD9E50DEEFB1ABBEC" box="[1175,1223,224,248]" pageId="1" pageNumber="22">type</typeStatus>
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||
locality by Dennis Williams on
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<date id="FF85A3B38D1CFE4ADC540C0AFDDEBA08" box="[294,515,260,284]" pageId="1" pageNumber="22" value="1987-09-12">12 September 1987</date>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C321D6F88D1CFE4ADDB50C26FEC2BA93" pageId="1" pageNumber="22" type="etymology">
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<paragraph id="8B8485738D1CFE4ADDB50C26FEC2BA93" blockId="1.[151,1437,151,860]" pageId="1" pageNumber="22">
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<emphasis id="B94F59618D1CFE4ADDB50C26FE93BA54" bold="true" box="[199,334,296,320]" pageId="1" pageNumber="22">Etymology.</emphasis>
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The specific epithet
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<taxonomicName id="4C3BFEF08D1CFE4ADF3B0C26FD72BA54" box="[585,687,296,320]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="1" pageNumber="22" phylum="Arthropoda" rank="species" species="williamsi">
|
||
<emphasis id="B94F59618D1CFE4ADF3B0C26FD72BA54" box="[585,687,296,320]" italics="true" pageId="1" pageNumber="22">williamsi</emphasis>
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||
</taxonomicName>
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is chosen to honor the memory of cave diver Dennis Williams (
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<date id="FF85A3B38D1CFE4ADDED0C42FEFFBA70" box="[159,290,332,356]" pageId="1" pageNumber="22" value="1943" valueMax="2005">1943–2005</date>
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), who explored numerous anchialine cave systems and assisted with discoveries of remipede crustaceans.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C321D6F88D1CFE4ADDB50C9DFD98BAE1" pageId="1" pageNumber="22" type="diagnosis">
|
||
<paragraph id="8B8485738D1CFE4ADDB50C9DFD98BAE1" blockId="1.[151,1437,151,860]" pageId="1" pageNumber="22">
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||
<emphasis id="B94F59618D1CFE4ADDB50C9DFE9CBAB8" bold="true" box="[199,321,403,428]" pageId="1" pageNumber="22">Diagnosis.</emphasis>
|
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A comparatively small and slender species. Prehensile cephalic limbs sparsely setose, maxillules well developed. Sternal bars isomorphic. Antennules short, about 1/5 of body length; dorsal ramus with 10–11 segments, ventral ramus with 5 segments.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C321D6F88D1CFE4EDDB50CF1FBDBBCE0" lastPageId="5" lastPageNumber="26" pageId="1" pageNumber="22" type="description">
|
||
<paragraph id="8B8485738D1CFE4ADDB50CF1FBE7B9B3" blockId="1.[151,1437,151,860]" pageId="1" pageNumber="22">
|
||
<emphasis id="B94F59618D1CFE4ADDB50CF1FE84B90C" bold="true" box="[199,345,511,536]" pageId="1" pageNumber="22">Description.</emphasis>
|
||
Body slender, up to
|
||
<quantity id="4CC328968D1CFE4ADF310CF1FD7BB903" box="[579,678,511,535]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="1" pageNumber="22" unit="mm" value="11.5">11.5 mm</quantity>
|
||
and 22 trunk segments (
|
||
<figureCitation id="130099F68D1CFE4ADEC90CF1FC20B903" box="[955,1021,511,535]" captionStart="FIGURE 1" captionStartId="1.[151,249,1293,1316]" captionTargetBox="[268,1302,896,1287]" captionTargetId="figure@1.[268,1302,894,1289]" captionTargetPageId="1" captionText="FIGURE 1. Speleonectes williamsi n. sp. Photograph of paratype 1 (10.2 mm), dorsal view." httpUri="https://zenodo.org/record/206643/files/figure.png" pageId="1" pageNumber="22">Fig. 1</figureCitation>
|
||
). Pleurae of tergites well developed, with rounded distolateral corners, becoming slightly pointed around mid-trunk. Head shield subrectangular, tapering anteriorly, with concave lateral margins (
|
||
<figureCitation id="130099F68D1CFE4ADFEA0F49FD00B94B" box="[664,733,583,607]" captionStart="FIGURE 2" captionStartId="2.[151,249,1915,1938]" captionTargetBox="[188,1407,193,1892]" captionTargetId="figure@2.[181,1415,193,1898]" captionTargetPageId="2" captionText="FIGURE 2. Speleonectes williamsi n. sp., paratype 1 (10.2 mm). A: Head shield and anterior trunk segments (dorsal view). B: Frontal filaments. C: Antennule. D: Antenna. E: Labrum. F: Enlarged posterior margin of labrum. G: Left mandible. H: Lacinia mobilis (left) and incisor process (right) of left mandible. I: Lacinia mobilis (left) and incisor process (right) of right mandible. Scale bars C – G = 0.1 mm." httpUri="https://zenodo.org/record/206644/files/figure.png" pageId="1" pageNumber="22">Fig. 2</figureCitation>
|
||
A). First trunk segment partially covered by head shield (
|
||
<figureCitation id="130099F68D1CFE4AD81D0F49FF7BB990" captionStart="FIGURE 2" captionStartId="2.[151,249,1915,1938]" captionTargetBox="[188,1407,193,1892]" captionTargetId="figure@2.[181,1415,193,1898]" captionTargetPageId="2" captionText="FIGURE 2. Speleonectes williamsi n. sp., paratype 1 (10.2 mm). A: Head shield and anterior trunk segments (dorsal view). B: Frontal filaments. C: Antennule. D: Antenna. E: Labrum. F: Enlarged posterior margin of labrum. G: Left mandible. H: Lacinia mobilis (left) and incisor process (right) of left mandible. I: Lacinia mobilis (left) and incisor process (right) of right mandible. Scale bars C – G = 0.1 mm." httpUri="https://zenodo.org/record/206644/files/figure.png" pageId="1" pageNumber="22">Fig. 2</figureCitation>
|
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A). Sternites unmodified. Sternal bars isomorphic. Female gonopores on protopods of seventh trunk limbs; male gonopores behind cone-like cuticular projections on protopods of 14th trunk limbs.
|
||
</paragraph>
|
||
<paragraph id="8B8485738D1CFE4ADDB50FBAFB82B9D8" blockId="1.[151,1437,151,860]" box="[199,1119,692,716]" pageId="1" pageNumber="22">
|
||
Frontal filaments with medial process inserted near tip of main filament (
|
||
<figureCitation id="130099F68D1CFE4ADE880FBAFBE3B9D8" box="[1018,1086,692,716]" captionStart="FIGURE 2" captionStartId="2.[151,249,1915,1938]" captionTargetBox="[188,1407,193,1892]" captionTargetId="figure@2.[181,1415,193,1898]" captionTargetPageId="2" captionText="FIGURE 2. Speleonectes williamsi n. sp., paratype 1 (10.2 mm). A: Head shield and anterior trunk segments (dorsal view). B: Frontal filaments. C: Antennule. D: Antenna. E: Labrum. F: Enlarged posterior margin of labrum. G: Left mandible. H: Lacinia mobilis (left) and incisor process (right) of left mandible. I: Lacinia mobilis (left) and incisor process (right) of right mandible. Scale bars C – G = 0.1 mm." httpUri="https://zenodo.org/record/206644/files/figure.png" pageId="1" pageNumber="22">Fig. 2</figureCitation>
|
||
B).
|
||
</paragraph>
|
||
<paragraph id="8B8485738D1CFE4ADDB50FD9FD12B848" blockId="1.[151,1437,151,860]" pageId="1" pageNumber="22">
|
||
Antennules (
|
||
<figureCitation id="130099F68D1CFE4ADC250FD9FE41B9FB" box="[343,412,727,751]" captionStart="FIGURE 2" captionStartId="2.[151,249,1915,1938]" captionTargetBox="[188,1407,193,1892]" captionTargetId="figure@2.[181,1415,193,1898]" captionTargetPageId="2" captionText="FIGURE 2. Speleonectes williamsi n. sp., paratype 1 (10.2 mm). A: Head shield and anterior trunk segments (dorsal view). B: Frontal filaments. C: Antennule. D: Antenna. E: Labrum. F: Enlarged posterior margin of labrum. G: Left mandible. H: Lacinia mobilis (left) and incisor process (right) of left mandible. I: Lacinia mobilis (left) and incisor process (right) of right mandible. Scale bars C – G = 0.1 mm." httpUri="https://zenodo.org/record/206644/files/figure.png" pageId="1" pageNumber="22">Fig. 2</figureCitation>
|
||
C): Peduncle with small field of long ribbon-like aesthetascs. Dorsal ramus about 20–25% of body length, composed of 10–11 segments. Ventral ramus approximately half the length of dorsal ramus, composed of 5 segments; proximal-most segment elongate. Segments of both rami with 2–3 distal compound aesthetascs and several fine, short setae along margin.
|
||
</paragraph>
|
||
<caption id="DF44D5FB8D1CFE4ADDE50803FBE0BE37" box="[151,1085,1293,1316]" httpUri="https://zenodo.org/record/206643/files/figure.png" pageId="1" pageNumber="22" targetBox="[268,1302,896,1287]" targetPageId="1">
|
||
<paragraph id="8B8485738D1CFE4ADDE50803FBE0BE37" blockId="1.[151,1085,1293,1316]" box="[151,1085,1293,1316]" pageId="1" pageNumber="22">
|
||
<emphasis id="B94F59618D1CFE4ADDE50803FECFBE37" bold="true" box="[151,274,1293,1316]" pageId="1" pageNumber="22">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C3BFEF08D1CFE4ADC6A0800FE22BE30" box="[280,511,1294,1316]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="1" pageNumber="22" phylum="Arthropoda" rank="species" species="williamsi" status="sp. nov.">
|
||
<emphasis id="B94F59618D1CFE4ADC6A0800FE22BE30" box="[280,511,1294,1316]" italics="true" pageId="1" pageNumber="22">Speleonectes williamsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94F59618D1CFE4ADF760800FDE3BE37" bold="true" box="[516,574,1294,1315]" pageId="1" pageNumber="22">
|
||
<taxonomicNameLabel id="A27CE41A8D1CFE4ADF760800FDE3BE37" box="[516,574,1294,1315]" pageId="1" pageNumber="22" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Photograph of paratype 1 (10.2 mm), dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8485738D1CFE4ADDB50845FA41BEB9" blockId="1.[151,1437,1355,2013]" pageId="1" pageNumber="22">
|
||
Antennae (
|
||
<figureCitation id="130099F68D1CFE4ADC330845FE58BE77" box="[321,389,1355,1379]" captionStart="FIGURE 2" captionStartId="2.[151,249,1915,1938]" captionTargetBox="[188,1407,193,1892]" captionTargetId="figure@2.[181,1415,193,1898]" captionTargetPageId="2" captionText="FIGURE 2. Speleonectes williamsi n. sp., paratype 1 (10.2 mm). A: Head shield and anterior trunk segments (dorsal view). B: Frontal filaments. C: Antennule. D: Antenna. E: Labrum. F: Enlarged posterior margin of labrum. G: Left mandible. H: Lacinia mobilis (left) and incisor process (right) of left mandible. I: Lacinia mobilis (left) and incisor process (right) of right mandible. Scale bars C – G = 0.1 mm." httpUri="https://zenodo.org/record/206644/files/figure.png" pageId="1" pageNumber="22">Fig. 2</figureCitation>
|
||
D): Proximal segment of protopod with 3 setae, distal segment with 6 setae. Exopod 1.5 times longer than distal segment of protopod, with about 26 setae along margin. Endopod slightly curved; proximal and medial segments each with 5 setae; distal segment with 19 setae, of which 12 originate as pairs. All setae plumose.
|
||
</paragraph>
|
||
<paragraph id="8B8485738D1CFE4ADDB508B5FCBABEEC" blockId="1.[151,1437,1355,2013]" pageId="1" pageNumber="22">
|
||
Labrum (
|
||
<figureCitation id="130099F68D1CFE4ADC4208B5FEA9BEC7" box="[304,372,1467,1491]" captionStart="FIGURE 2" captionStartId="2.[151,249,1915,1938]" captionTargetBox="[188,1407,193,1892]" captionTargetId="figure@2.[181,1415,193,1898]" captionTargetPageId="2" captionText="FIGURE 2. Speleonectes williamsi n. sp., paratype 1 (10.2 mm). A: Head shield and anterior trunk segments (dorsal view). B: Frontal filaments. C: Antennule. D: Antenna. E: Labrum. F: Enlarged posterior margin of labrum. G: Left mandible. H: Lacinia mobilis (left) and incisor process (right) of left mandible. I: Lacinia mobilis (left) and incisor process (right) of right mandible. Scale bars C – G = 0.1 mm." httpUri="https://zenodo.org/record/206644/files/figure.png" pageId="1" pageNumber="22">Fig. 2</figureCitation>
|
||
E, F) prominent, bearing fields of fine setules on posterolateral corners and medial lobes; funnel-shaped medial cavity with unilateral field of sharp denticles.
|
||
</paragraph>
|
||
<paragraph id="8B8485738D1CFE4ADDB50B0BFB4ABD7C" blockId="1.[151,1437,1355,2013]" pageId="1" pageNumber="22">
|
||
Mandibles asymmetrical (
|
||
<figureCitation id="130099F68D1CFE4ADC9E0B0BFDF2BD09" box="[492,559,1541,1565]" captionStart="FIGURE 2" captionStartId="2.[151,249,1915,1938]" captionTargetBox="[188,1407,193,1892]" captionTargetId="figure@2.[181,1415,193,1898]" captionTargetPageId="2" captionText="FIGURE 2. Speleonectes williamsi n. sp., paratype 1 (10.2 mm). A: Head shield and anterior trunk segments (dorsal view). B: Frontal filaments. C: Antennule. D: Antenna. E: Labrum. F: Enlarged posterior margin of labrum. G: Left mandible. H: Lacinia mobilis (left) and incisor process (right) of left mandible. I: Lacinia mobilis (left) and incisor process (right) of right mandible. Scale bars C – G = 0.1 mm." httpUri="https://zenodo.org/record/206644/files/figure.png" pageId="1" pageNumber="22">Fig. 2</figureCitation>
|
||
G–I). Left incisor process with 4 large denticles; left lacinia mobilis v-shaped or 2-cusped, with indistinct denticles. Right incisor process and lacinia mobilis each with 3 denticles. Molar processes of both mandibles well-defined, apical surface equipped with field of plumose setae.
|
||
</paragraph>
|
||
<paragraph id="8B8485738D1CFE4ADDB70B7BFD95BCC9" blockId="1.[151,1437,1355,2013]" pageId="1" pageNumber="22">
|
||
Maxillules (
|
||
<figureCitation id="130099F68D1CFE4ADC3C0B7BFE4CBD99" box="[334,401,1653,1677]" captionStart="FIGURE 3" captionStartId="3.[151,249,1975,1998]" captionTargetBox="[355,1255,201,1943]" captionTargetId="figure@3.[328,1259,193,1954]" captionTargetPageId="3" captionText="FIGURE 3. Speleonectes williamsi n. sp., paratype 1 (10.2 mm). A: Maxillule, arrow pointing at enlarged endite of segment 1. B: Maxilla. C: Maxilliped, with enlarged terminal claw. Scale bar A – C = 0.1 mm." httpUri="https://zenodo.org/record/206645/files/figure.png" pageId="1" pageNumber="22">Fig. 3</figureCitation>
|
||
A) much more robust than maxillae and maxillipeds. Segment 1 with long, slender medial endite bearing 7 apical spine-like setae, posterior-most seta long and robust, anterior-most serrate. Segment 2 with broad plate-like endite, bearing a row of 7 spine-like setae along posterior apical margin and many small simple subterminal setae, anterior-most seta serrate; 1 plumose seta and several short simple setae along posterior margin; 4 long and several short setae on anterior apical margin. Segment 3 short, with reduced conical endite bearing 2 robust sclerotized setae with serrations on distal half and 2 slender simple setae. Segment 4 (lacertus) robust, with conical medial margin, bearing 2 robust sclerotized setae and 3–4 long simple setae. Segment 5 well developed, expanded, with a cluster of few setae on distomedial margin. Segment 6 greatly reduced, about 1/5 as long as segment 5, with clusters of setae on distomedial and distolateral margins. Terminal claw slightly longer than segment 6, with 2 clusters of long setae at base.
|
||
</paragraph>
|
||
<caption id="DF44D5FB8D1FFE49DDE50A75FE79BCFA" httpUri="https://zenodo.org/record/206644/files/figure.png" pageId="2" pageNumber="23" targetBox="[188,1407,193,1892]" targetPageId="2">
|
||
<paragraph id="8B8485738D1FFE49DDE50A75FE79BCFA" blockId="2.[151,1437,1915,2030]" pageId="2" pageNumber="23">
|
||
<emphasis id="B94F59618D1FFE49DDE50A75FECFBC85" bold="true" box="[151,274,1915,1938]" pageId="2" pageNumber="23">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C3BFEF08D1FFE49DC6A0A72FE22BC86" box="[280,511,1916,1938]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="williamsi" status="sp. nov.">
|
||
<emphasis id="B94F59618D1FFE49DC6A0A72FE22BC86" box="[280,511,1916,1938]" italics="true" pageId="2" pageNumber="23">Speleonectes williamsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94F59618D1FFE49DF770A72FDE3BC85" bold="true" box="[517,574,1916,1937]" pageId="2" pageNumber="23">
|
||
<taxonomicNameLabel id="A27CE41A8D1FFE49DF770A72FDE3BC85" box="[517,574,1916,1937]" pageId="2" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype 1 (10.2 mm). A: Head shield and anterior trunk segments (dorsal view). B: Frontal filaments. C: Antennule. D: Antenna. E: Labrum. F: Enlarged posterior margin of labrum. G: Left mandible. H: Lacinia mobilis (left) and incisor process (right) of left mandible. I: Lacinia mobilis (left) and incisor process (right) of right mandible. Scale bars C–G = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF44D5FB8D1EFE48DDE50AB9FC09BCF8" httpUri="https://zenodo.org/record/206645/files/figure.png" pageId="3" pageNumber="24" targetBox="[355,1255,201,1943]" targetPageId="3">
|
||
<paragraph id="8B8485738D1EFE48DDE50AB9FC09BCF8" blockId="3.[151,1436,1975,2028]" pageId="3" pageNumber="24">
|
||
<emphasis id="B94F59618D1EFE48DDE50AB9FECFBCD9" bold="true" box="[151,274,1975,1998]" pageId="3" pageNumber="24">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C3BFEF08D1EFE48DC6A0AB9FE23BCD9" box="[280,510,1975,1997]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="3" pageNumber="24" phylum="Arthropoda" rank="species" species="williamsi" status="sp. nov.">
|
||
<emphasis id="B94F59618D1EFE48DC6A0AB9FE23BCD9" box="[280,510,1975,1997]" italics="true" pageId="3" pageNumber="24">Speleonectes williamsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94F59618D1EFE48DF760AB6FDE0BCD9" bold="true" box="[516,573,1976,1997]" pageId="3" pageNumber="24">
|
||
<taxonomicNameLabel id="A27CE41A8D1EFE48DF760AB6FDE0BCD9" box="[516,573,1976,1997]" pageId="3" pageNumber="24" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype 1 (10.2 mm). A: Maxillule, arrow pointing at enlarged endite of segment 1. B: Maxilla. C: Maxilliped, with enlarged terminal claw. Scale bar A–C = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF44D5FB8D19FE4FDDE50FCDFE52B9EC" httpUri="https://zenodo.org/record/206646/files/figure.png" pageId="4" pageNumber="25" targetBox="[240,1346,181,700]" targetPageId="4">
|
||
<paragraph id="8B8485738D19FE4FDDE50FCDFE52B9EC" blockId="4.[151,1436,707,760]" pageId="4" pageNumber="25">
|
||
<emphasis id="B94F59618D19FE4FDDE50FCDFEC9B9CC" bold="true" box="[151,276,707,730]" pageId="4" pageNumber="25">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C3BFEF08D19FE4FDC6F0FCDFDDBB9CD" box="[285,518,707,729]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="williamsi" status="sp. nov.">
|
||
<emphasis id="B94F59618D19FE4FDC6F0FCDFDDBB9CD" box="[285,518,707,729]" italics="true" pageId="4" pageNumber="25">Speleonectes williamsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94F59618D19FE4FDF7C0FCAFD97B9CD" bold="true" box="[526,586,708,729]" pageId="4" pageNumber="25">
|
||
<taxonomicNameLabel id="A27CE41A8D19FE4FDF7C0FCAFD97B9CD" box="[526,586,708,729]" pageId="4" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype 7. Claw of maxilla. A: Anterior view, showing putative sensory pit. B: Enlarged posterior view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8485738D19FE4FDDB50E11FE19B8D6" blockId="4.[151,1437,799,2038]" pageId="4" pageNumber="25">
|
||
Maxillae (
|
||
<figureCitation id="130099F68D19FE4FDC480E11FEA3B823" box="[314,382,799,823]" captionStart="FIGURE 3" captionStartId="3.[151,249,1975,1998]" captionTargetBox="[355,1255,201,1943]" captionTargetId="figure@3.[328,1259,193,1954]" captionTargetPageId="3" captionText="FIGURE 3. Speleonectes williamsi n. sp., paratype 1 (10.2 mm). A: Maxillule, arrow pointing at enlarged endite of segment 1. B: Maxilla. C: Maxilliped, with enlarged terminal claw. Scale bar A – C = 0.1 mm." httpUri="https://zenodo.org/record/206645/files/figure.png" pageId="4" pageNumber="25">Fig. 3</figureCitation>
|
||
B): Segment 1 with three long digitiform endites each with 1 robust sclerotized curved spinelike seta, with 1–2 short setae and 1–2 long subterminal setae. Segment 2 short with conical medial endite bearing 1 or 2 stout spine-like setae and several subapical long and short setae. Segment 3 long, slightly expanded medially, sparsely setose with 3–4 long setae. Segment 4 trapezoidal with clusters of several distal setae. Point of flexure between segments 3 and 4.
|
||
</paragraph>
|
||
<paragraph id="8B8485738D19FE4FDDB70EC3FE0EBF7B" blockId="4.[151,1437,799,2038]" pageId="4" pageNumber="25">
|
||
Segments 5 and 6 both nearly square, bearing clusters of short and long distal setae and several setae on distomedial half of each segment. Segment 5 about 1/2 as long as segment 4. Segment 6 slightly smaller than segment 5 and more setose. Distal claw complex consisting of an arc of approximately 8–10 incised denticles with robust anterior and posterior, subterminal spines, a large anterior subterminal pit (putative sensory pore; see
|
||
<figureCitation id="130099F68D19FE4FD875093BFA97BF59" box="[1287,1354,1077,1101]" captionStart="FIGURE 4" captionStartId="4.[151,249,707,730]" captionTargetBox="[240,1346,181,700]" captionTargetId="figure@4.[240,1346,181,700]" captionTargetPageId="4" captionText="FIGURE 4. Speleonectes williamsi n. sp., paratype 7. Claw of maxilla. A: Anterior view, showing putative sensory pit. B: Enlarged posterior view." httpUri="https://zenodo.org/record/206646/files/figure.png" pageId="4" pageNumber="25">Fig. 4</figureCitation>
|
||
), and a short setose thumb-like pad.
|
||
</paragraph>
|
||
<paragraph id="8B8485738D19FE4FDDB50974FD77BE91" blockId="4.[151,1437,799,2038]" pageId="4" pageNumber="25">
|
||
Maxillipeds (
|
||
<figureCitation id="130099F68D19FE4FDC2E0974FE40BF86" box="[348,413,1146,1170]" captionStart="FIGURE 3" captionStartId="3.[151,249,1975,1998]" captionTargetBox="[355,1255,201,1943]" captionTargetId="figure@3.[328,1259,193,1954]" captionTargetPageId="3" captionText="FIGURE 3. Speleonectes williamsi n. sp., paratype 1 (10.2 mm). A: Maxillule, arrow pointing at enlarged endite of segment 1. B: Maxilla. C: Maxilliped, with enlarged terminal claw. Scale bar A – C = 0.1 mm." httpUri="https://zenodo.org/record/206645/files/figure.png" pageId="4" pageNumber="25">Fig. 3</figureCitation>
|
||
C) approximately 1.5 times as long as maxillae; 9-segmented with point of flexure between segments 5 and 6. Segments 1–3 forming convoluted joint complex. Segment 1 short with a few medial setae. Segment 2 oblique, with several medial setae. Segment 3 slightly expanded medially with a few medial setae. Segment 4 (lacertus) elongate, rounded medially with about 4 long and several short medial setae. Segment 5 slightly shorter than segment 4, slightly expanded, with several long distomedial setae. Segment 6 slightly shorter than segment 5 with distomedial and distolateral clusters of setae. Segment 7 slightly shorter than segment 6, with distal clusters of setae. Segment 8 shorter than segment 7, bearing a medial row of setae and cluster of many distolateral setae. Terminal claw complex similar to that of maxillae.
|
||
</paragraph>
|
||
<paragraph id="8B8485738D19FE4FDDB7089EFCC2BD26" blockId="4.[151,1437,799,2038]" pageId="4" pageNumber="25">
|
||
Limbs of first post-cephalic trunk somite biramous, with relatively slender rami (
|
||
<figureCitation id="130099F68D19FE4FD901089EFB65BEBC" box="[1139,1208,1424,1448]" captionStart="FIGURE 5" captionStartId="6.[151,249,1975,1998]" captionTargetBox="[218,1367,199,1943]" captionTargetId="figure@6.[218,1370,193,1954]" captionTargetPageId="6" captionText="FIGURE 5. Speleonectes williamsi n. sp., paratype 1 (10.2 mm). A: Limb of first trunk somite. B: Limb of fourteenth trunk somite, with enlarged short, stout plumose (C) and short, stout serrate seta (D). E: Anal somite. All scale bars = 0.1 mm." httpUri="https://zenodo.org/record/206647/files/figure.png" pageId="4" pageNumber="25">Fig. 5</figureCitation>
|
||
A). Endopod 4-segmented; endopods of all trunk limbs slightly shorter and narrower than exopods; exopod 3-segmented; both rami bearing long, plumose setae on margins. Limbs of mid-trunk region with broad, expanded segments (
|
||
<figureCitation id="130099F68D19FE4FD98C08DBFA9CBEF9" box="[1278,1345,1493,1517]" captionStart="FIGURE 5" captionStartId="6.[151,249,1975,1998]" captionTargetBox="[218,1367,199,1943]" captionTargetId="figure@6.[218,1370,193,1954]" captionTargetPageId="6" captionText="FIGURE 5. Speleonectes williamsi n. sp., paratype 1 (10.2 mm). A: Limb of first trunk somite. B: Limb of fourteenth trunk somite, with enlarged short, stout plumose (C) and short, stout serrate seta (D). E: Anal somite. All scale bars = 0.1 mm." httpUri="https://zenodo.org/record/206647/files/figure.png" pageId="4" pageNumber="25">Fig. 5</figureCitation>
|
||
B); exopod with up to 5 short serrate setae on distolateral corners; endopod bearing both short plumose and short serrate setae on distolateral and distomedial corners (
|
||
<figureCitation id="130099F68D19FE4FDFE50B14FD04BD26" box="[663,729,1562,1586]" captionStart="FIGURE 5" captionStartId="6.[151,249,1975,1998]" captionTargetBox="[218,1367,199,1943]" captionTargetId="figure@6.[218,1370,193,1954]" captionTargetPageId="6" captionText="FIGURE 5. Speleonectes williamsi n. sp., paratype 1 (10.2 mm). A: Limb of first trunk somite. B: Limb of fourteenth trunk somite, with enlarged short, stout plumose (C) and short, stout serrate seta (D). E: Anal somite. All scale bars = 0.1 mm." httpUri="https://zenodo.org/record/206647/files/figure.png" pageId="4" pageNumber="25">Fig. 5</figureCitation>
|
||
C, D).
|
||
</paragraph>
|
||
<paragraph id="8B8485738D19FE4FDDB70B33FACBBD42" blockId="4.[151,1437,799,2038]" box="[197,1302,1597,1622]" pageId="4" pageNumber="25">
|
||
Anal segment nearly as wide as it is long (
|
||
<figureCitation id="130099F68D19FE4FDFED0B33FD3CBD41" box="[671,737,1597,1621]" captionStart="FIGURE 5" captionStartId="6.[151,249,1975,1998]" captionTargetBox="[218,1367,199,1943]" captionTargetId="figure@6.[218,1370,193,1954]" captionTargetPageId="6" captionText="FIGURE 5. Speleonectes williamsi n. sp., paratype 1 (10.2 mm). A: Limb of first trunk somite. B: Limb of fourteenth trunk somite, with enlarged short, stout plumose (C) and short, stout serrate seta (D). E: Anal somite. All scale bars = 0.1 mm." httpUri="https://zenodo.org/record/206647/files/figure.png" pageId="4" pageNumber="25">Fig. 5</figureCitation>
|
||
E). Caudal rami slightly longer than anal segment.
|
||
</paragraph>
|
||
<paragraph id="8B8485738D19FE4FDDB70B51FAB7BC16" blockId="4.[151,1437,799,2038]" pageId="4" pageNumber="25">
|
||
<emphasis id="B94F59618D19FE4FDDB70B51FE3BBD6C" bold="true" box="[197,486,1631,1656]" pageId="4" pageNumber="25">Additional observations</emphasis>
|
||
. Most specimens of
|
||
<taxonomicName id="4C3BFEF08D19FE4FDFAF0B6EFCBEBD6C" box="[733,867,1632,1656]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="williamsi" status="sp. nov.">
|
||
<emphasis id="B94F59618D19FE4FDFAF0B6EFCBEBD6C" box="[733,867,1632,1656]" italics="true" pageId="4" pageNumber="25">S. williamsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94F59618D19FE4FDE1C0B6EFC6DBD6C" bold="true" box="[878,944,1632,1656]" pageId="4" pageNumber="25">
|
||
<taxonomicNameLabel id="A27CE41A8D19FE4FDE1C0B6EFC6DBD6C" box="[878,944,1632,1656]" pageId="4" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
have two very visible rust-colored glands extending from the head to the second trunk segment (
|
||
<figureCitation id="130099F68D19FE4FDE740B8CFC91BD8E" box="[774,844,1666,1690]" captionStart="FIGURE 6" captionStartId="7.[151,249,804,827]" captionTargetBox="[414,1172,219,787]" captionTargetId="figure@7.[414,1172,219,788]" captionTargetPageId="7" captionText="FIGURE 6. Speleonectes williamsi n. sp., paratype 6. Detail of head and first trunk segments showing conspicuous glandular structures (see also Yager, 1991)." httpUri="https://zenodo.org/record/206648/files/figure.png" pageId="4" pageNumber="25">Fig. 6</figureCitation>
|
||
).
|
||
<bibRefCitation id="EFAAF8828D19FE4FDE110B8DFC2ABD8E" author="Yager" box="[867,1015,1666,1691]" pageId="4" pageNumber="25" refString="Yager, J. (1991) The Remipedia (Crustacea): Recent investigations of their biology and phylogeny. Verhandlungen der Deutschen Zoologischen Gesellschaft, 84, 261 - 269." type="journal article" year="1991">Yager (1991)</bibRefCitation>
|
||
previously described these glands in the specimen known here as
|
||
<typeStatus id="54803BD18D19FE4FDCAB0BA8FDE6BDAA" box="[473,571,1702,1726]" pageId="4" pageNumber="25" type="paratype">Paratype</typeStatus>
|
||
6. The presence of these glands was documented in other species of Remipedia by
|
||
<bibRefCitation id="EFAAF8828D19FE4FDD900BC6FDDEBDF4" author="Schram" box="[226,515,1736,1760]" pageId="4" pageNumber="25" refString="Schram F. R. & Lewis C. A. (1989) Functional morphology of feeding in Nectiopoda. In Felgenhauer B. E., Watling L. & Thistle A. B. (Eds), Functional Morphology of Feeding and Grooming in Crustacea. A. A. Balkema, Rotterdam, pp. 115 - 122." type="book chapter" year="1989">Schram and Lewis (1989)</bibRefCitation>
|
||
and van der
|
||
<bibRefCitation id="EFAAF8828D19FE4FDFE00BC7FC01BDF4" author="Ham" box="[658,988,1736,1760]" pageId="4" pageNumber="25" refString="van der Ham, J. L. & Felgenhauer, B. E. (2007) The functional morphology of the putative injecting apparatus of Speleonectes tanumekes (Remipedia). Journal of Crustacean Biology, 27, 1 - 9." type="journal article" year="2007">Ham and Felgenhauer (2007)</bibRefCitation>
|
||
. However, no other species described to date has such distinct glands and ducts visible in the head and anterior trunk segments as those of
|
||
<emphasis id="B94F59618D19FE4FD9AD0BE5FAB7BC16" box="[1247,1386,1770,1794]" italics="true" pageId="4" pageNumber="25">
|
||
<taxonomicName id="4C3BFEF08D19FE4FD9AD0BE5FAB8BC16" box="[1247,1381,1770,1794]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="williamsi">S. williamsi</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B8485738D19FE4FDDB50A02FD25BCE1" blockId="4.[151,1437,799,2038]" pageId="4" pageNumber="25">
|
||
<emphasis id="B94F59618D19FE4FDDB50A02FD2DBC31" bold="true" box="[199,752,1804,1829]" pageId="4" pageNumber="25">Morphological comparison with other species.</emphasis>
|
||
<taxonomicName id="4C3BFEF08D19FE4FDF8A0A03FC2ABC31" box="[760,1015,1805,1829]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="williamsi" status="sp. nov.">
|
||
<emphasis id="B94F59618D19FE4FDF8A0A03FC2ABC31" box="[760,1015,1805,1829]" italics="true" pageId="4" pageNumber="25">Speleonectes williamsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94F59618D19FE4FDE8D0A03FBE2BC31" bold="true" box="[1023,1087,1805,1829]" pageId="4" pageNumber="25">
|
||
<taxonomicNameLabel id="A27CE41A8D19FE4FDE8D0A03FBE2BC31" box="[1023,1087,1805,1829]" pageId="4" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
does not share any of the apomorphies that define the families
|
||
<taxonomicName id="4C3BFEF08D19FE4FDF6F0A3EFC49BC5C" authority="Schram et al., 1986" authorityName="Schram et al." authorityYear="1986" box="[541,916,1840,1864]" class="Remipedia" family="Godzilliidae" kingdom="Animalia" order="Nectiopoda" pageId="4" pageNumber="25" phylum="Arthropoda" rank="family">Godzilliidae Schram et al., 1986</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C3BFEF08D19FE4FDEB20A3EFA45BC5C" authority="Koenemann et al., 2007" authorityName="Koenemann et al." authorityYear="2007" box="[960,1432,1840,1864]" class="Remipedia" family="Micropacteridae" kingdom="Animalia" order="Nectiopoda" pageId="4" pageNumber="25" phylum="Arthropoda" rank="family">
|
||
Micropacteridae
|
||
<bibRefCitation id="EFAAF8828D19FE4FD9F30A3FFA45BC5C" author="Koenemann" box="[1153,1432,1840,1864]" pageId="4" pageNumber="25" refString="Lorentzen, D., Koenemann, S. & Iliffe, T. M. (2007) Speleonectes emersoni, a new species of Remipedia (Crustacea) from the Dominican Republic. Zootaxa, 1543, 61 - 68." type="journal article" year="2007">Koenemann et al., 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
, including in particular the reduction and modification of the post-mandibular cephalic limbs. The prehensile cephalic limbs of
|
||
<taxonomicName id="4C3BFEF08D19FE4FDC2D0A78FE3EBC99" box="[351,483,1909,1933]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="williamsi">
|
||
<emphasis id="B94F59618D19FE4FDC2D0A78FEB0BC99" box="[351,365,1910,1933]" italics="true" pageId="4" pageNumber="25">S</emphasis>
|
||
.
|
||
<emphasis id="B94F59618D19FE4FDC0E0A7BFE3EBC99" box="[380,483,1909,1933]" italics="true" pageId="4" pageNumber="25">williamsi</emphasis>
|
||
</taxonomicName>
|
||
also lack conspicuous characters such as the presence of discoid organs (in
|
||
<taxonomicName id="4C3BFEF08D19FE4FD84A0A78FE5ABCA4" authority="Yager, 1987" authorityName="Yager" authorityYear="1987" class="Remipedia" family="Cryptocorynetidae" genus="Cryptocorynetes" kingdom="Animalia" order="Nectiopoda" pageId="4" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94F59618D19FE4FD84A0A78FF2BBCA4" italics="true" pageId="4" pageNumber="25">Cryptocorynetes</emphasis>
|
||
<bibRefCitation id="EFAAF8828D19FE4FDD8F0A97FE5ABCA4" author="Yager" box="[253,391,1944,1969]" pageId="4" pageNumber="25" refString="Yager, J. (1987) Cryptocorynetes haptodiscus, new genus, new species, and Speleonectes benjamini, new species, of remipede crustaceans from anchialine caves in the Bahamas, with remarks on distribution and ecology. Proceedings of the Biological Society of Washington, 100, 302 - 320." type="journal article" year="1987">Yager, 1987</bibRefCitation>
|
||
</taxonomicName>
|
||
), dense fields of plumose setae (in
|
||
<taxonomicName id="4C3BFEF08D19FE4FDE670A96FB43BCA4" authority="Koenemann et al., 2004" authorityName="Koenemann et al." authorityYear="2004" box="[789,1182,1944,1968]" class="Remipedia" family="Cryptocorynetidae" genus="Kaloketos" kingdom="Animalia" order="Nectiopoda" pageId="4" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94F59618D19FE4FDE670A96FC58BCA4" box="[789,901,1944,1968]" italics="true" pageId="4" pageNumber="25">Kaloketos</emphasis>
|
||
<bibRefCitation id="EFAAF8828D19FE4FDEFF0A97FB43BCA4" author="Koenemann" box="[909,1182,1944,1968]" pageId="4" pageNumber="25" refString="Koenemann, S., Iliffe, T. M. & Yager, J. (2004) Kaloketos pilosus, a new genus and species of Remipedia (Crustacea) from the Turks and Caicos Islands. Zootaxa, 618, 1 - 12." type="journal article" year="2004">Koenemann et al., 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
), or irregularly shaped terminal claws (in
|
||
<taxonomicName id="4C3BFEF08D19FE4FDC180AB5FCCDBCC6" authority="Yager and Schram, 1986" authorityName="Yager and Schram" authorityYear="1986" box="[362,784,1978,2003]" class="Remipedia" family="Speleonectidae" genus="Lasionectes" kingdom="Animalia" order="Nectiopoda" pageId="4" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94F59618D19FE4FDC180AB5FE33BCC6" box="[362,494,1979,2002]" italics="true" pageId="4" pageNumber="25">Lasionectes</emphasis>
|
||
Yager and Schram, 1986
|
||
</taxonomicName>
|
||
). Therefore, the new species is placed in the genus
|
||
<taxonomicName id="4C3BFEF08D19FE4FD8240AB4FEA5BCE1" authority="Yager, 1981" authorityName="Yager" authorityYear="1981" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="4" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94F59618D19FE4FD8240AB4FF36BCE1" italics="true" pageId="4" pageNumber="25">Speleonectes</emphasis>
|
||
<bibRefCitation id="EFAAF8828D19FE4FDD830AD0FEA5BCE1" author="Yager" box="[241,376,2013,2038]" pageId="4" pageNumber="25" refString="Yager, J. (1981) A new class of Crustacea from a marine cave in the Bahamas. Journal of Crustacean Biology, 1, 328 - 333." type="journal article" year="1981">Yager, 1981</bibRefCitation>
|
||
</taxonomicName>
|
||
within the family
|
||
<taxonomicName id="4C3BFEF08D19FE4FDF350AD3FD29BCE1" box="[583,756,2013,2037]" class="Remipedia" family="Speleonectidae" kingdom="Animalia" order="Nectiopoda" pageId="4" pageNumber="25" phylum="Arthropoda" rank="family">Speleonectidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8485738D18FE4EDDB50D96FCC7BA03" blockId="5.[151,1437,151,1113]" pageId="5" pageNumber="26">
|
||
From among the 12 species currently assigned to the genus
|
||
<taxonomicName id="4C3BFEF08D18FE4EDEFD0D96FBFDBBA4" box="[911,1056,152,176]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94F59618D18FE4EDEFD0D96FBFDBBA4" box="[911,1056,152,176]" italics="true" pageId="5" pageNumber="26">Speleonectes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3BFEF08D18FE4ED9400D96FB67BBA4" box="[1074,1210,152,176]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="williamsi">
|
||
<emphasis id="B94F59618D18FE4ED9400D96FB9DBBBB" box="[1074,1088,152,175]" italics="true" pageId="5" pageNumber="26">S</emphasis>
|
||
.
|
||
<emphasis id="B94F59618D18FE4ED9210D96FB67BBA4" box="[1107,1210,152,176]" italics="true" pageId="5" pageNumber="26">williamsi</emphasis>
|
||
</taxonomicName>
|
||
is morphologically closely related to
|
||
<taxonomicName id="4C3BFEF08D18FE4EDC2F0DB5FD03BBC6" authority="Lorentzen et al., 2007" authorityName="Lorentzen et al." authorityYear="2007" box="[349,734,186,211]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="emersoni">
|
||
<emphasis id="B94F59618D18FE4EDC2F0DB5FEB6BBC6" box="[349,363,187,210]" italics="true" pageId="5" pageNumber="26">S</emphasis>
|
||
.
|
||
<emphasis id="B94F59618D18FE4EDC0B0DB2FE3DBBC7" box="[377,480,188,211]" italics="true" pageId="5" pageNumber="26">emersoni</emphasis>
|
||
<bibRefCitation id="EFAAF8828D18FE4EDC940DB5FD03BBC6" author="Lorentzen" box="[486,734,186,210]" pageId="5" pageNumber="26" refString="Lorentzen, D., Koenemann, S. & Iliffe, T. M. (2007) Speleonectes emersoni, a new species of Remipedia (Crustacea) from the Dominican Republic. Zootaxa, 1543, 61 - 68." type="journal article" year="2007">Lorentzen et al., 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
. Both species are relatively small, with a nearly identical habitus. They both have isomorphic sternal bars, and at a first glance, all appendages exhibit a striking resemblance in shape, relative sizes, and setal and segmentation patterns.
|
||
</paragraph>
|
||
<paragraph id="8B8485738D18FE4EDDB70C2DFB8CB95B" blockId="5.[151,1437,151,1113]" pageId="5" pageNumber="26">
|
||
Upon closer examination, however, we found a number of morphological differences in the antennules, antennae, prehensile cephalic limbs and the anal somite (
|
||
<tableCitation id="C6B9B0C88D18FE4EDFA90C4BFCFCBA49" box="[731,801,325,349]" captionStart="TABLE 1" captionStartId="5.[151,235,1157,1178]" captionTargetBox="[159,1378,1371,1951]" captionTargetPageId="5" captionText="TABLE 1. Comparison of morphological similarities and differences between S. williamsi n. sp. and S. emersoni." httpUri="http://table.plazi.org/id/DF44D5FB8D18FE4EDDE5098BFACABF8F" pageId="5" pageNumber="26" tableUuid="DF44D5FB8D18FE4EDDE5098BFACABF8F">Tab. 1</tableCitation>
|
||
). The ventral flagellum of the antennule has 5 segments in
|
||
<taxonomicName id="4C3BFEF08D18FE4EDDC60C66FEEABA94" box="[180,311,360,384]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="williamsi">
|
||
<emphasis id="B94F59618D18FE4EDDC60C66FEEABA94" box="[180,311,360,384]" italics="true" pageId="5" pageNumber="26">S. williamsi</emphasis>
|
||
</taxonomicName>
|
||
but
|
||
<quantity id="4CC328968D18FE4EDC180C69FE48BA6B" box="[362,405,359,383]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="5" pageNumber="26" unit="in" value="6.0">6 in</quantity>
|
||
<taxonomicName id="4C3BFEF08D18FE4EDCEE0C66FDC2BA94" box="[412,543,360,384]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="emersoni">
|
||
<emphasis id="B94F59618D18FE4EDCEE0C66FDC2BA94" box="[412,543,360,384]" italics="true" pageId="5" pageNumber="26">S. emersoni</emphasis>
|
||
</taxonomicName>
|
||
. The setae on the expods of the antennae in
|
||
<taxonomicName id="4C3BFEF08D18FE4ED9660C66FB4ABA94" box="[1044,1175,360,384]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="williamsi">
|
||
<emphasis id="B94F59618D18FE4ED9660C66FB4ABA94" box="[1044,1175,360,384]" italics="true" pageId="5" pageNumber="26">S. williamsi</emphasis>
|
||
</taxonomicName>
|
||
are numerous (26) and all long, while
|
||
<taxonomicName id="4C3BFEF08D18FE4EDC4C0C85FE1CBAB7" box="[318,449,395,419]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="emersoni">
|
||
<emphasis id="B94F59618D18FE4EDC4C0C85FE1CBAB7" box="[318,449,395,419]" italics="true" pageId="5" pageNumber="26">S. emersoni</emphasis>
|
||
</taxonomicName>
|
||
possesses a total of 18 long and short setae. The outer denticle of the right lacinia mobilis of the mandible has a flattened apical surface in
|
||
<taxonomicName id="4C3BFEF08D18FE4EDFA90CA0FC80BAD1" box="[731,861,429,453]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="williamsi">
|
||
<emphasis id="B94F59618D18FE4EDFA90CA0FC80BAD1" box="[731,861,429,453]" italics="true" pageId="5" pageNumber="26">S. williamsi</emphasis>
|
||
</taxonomicName>
|
||
but a pointed tip in
|
||
<taxonomicName id="4C3BFEF08D18FE4ED9300CA0FB1BBAD1" box="[1090,1222,430,453]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="emersoni">
|
||
<emphasis id="B94F59618D18FE4ED9300CA0FB1BBAD1" box="[1090,1222,430,453]" italics="true" pageId="5" pageNumber="26">S. emersoni</emphasis>
|
||
</taxonomicName>
|
||
. The fourth maxillary segment in
|
||
<taxonomicName id="4C3BFEF08D18FE4EDC3D0CDEFE09BAFC" box="[335,468,464,488]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="williamsi">
|
||
<emphasis id="B94F59618D18FE4EDC3D0CDEFE09BAFC" box="[335,468,464,488]" italics="true" pageId="5" pageNumber="26">S. williamsi</emphasis>
|
||
</taxonomicName>
|
||
is as long as the lacertus and only slightly expanded, however that of
|
||
<taxonomicName id="4C3BFEF08D18FE4ED98E0CDEFA5CBAFC" box="[1276,1409,464,488]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="emersoni">
|
||
<emphasis id="B94F59618D18FE4ED98E0CDEFA5CBAFC" box="[1276,1409,464,488]" italics="true" pageId="5" pageNumber="26">S. emersoni</emphasis>
|
||
</taxonomicName>
|
||
is shorter than the lacertus and is distinctly expanded distally. The terminal claws of maxillae and maxilipeds of
|
||
<taxonomicName id="4C3BFEF08D18FE4ED8F50CFDFF23B939" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="williamsi">
|
||
<emphasis id="B94F59618D18FE4ED8F50CFDFF23B939" italics="true" pageId="5" pageNumber="26">S. williamsi</emphasis>
|
||
</taxonomicName>
|
||
all have deeply incised denticles whereas those of
|
||
<taxonomicName id="4C3BFEF08D18FE4EDE4F0F18FC1CB939" box="[829,961,534,557]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="emersoni">
|
||
<emphasis id="B94F59618D18FE4EDE4F0F18FC1CB939" box="[829,961,534,557]" italics="true" pageId="5" pageNumber="26">S. emersoni</emphasis>
|
||
</taxonomicName>
|
||
are small (fused). In
|
||
<taxonomicName id="4C3BFEF08D18FE4ED9C00F18FAE8B939" box="[1202,1333,533,557]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="williamsi">
|
||
<emphasis id="B94F59618D18FE4ED9C00F18FAE8B939" box="[1202,1333,533,557]" italics="true" pageId="5" pageNumber="26">S. williamsi</emphasis>
|
||
</taxonomicName>
|
||
, the anal somite is nearly square, while the anal somite of
|
||
<taxonomicName id="4C3BFEF08D18FE4EDFC80F36FCE0B944" box="[698,829,568,592]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="emersoni">
|
||
<emphasis id="B94F59618D18FE4EDFC80F36FCE0B944" box="[698,829,568,592]" italics="true" pageId="5" pageNumber="26">S. emersoni</emphasis>
|
||
</taxonomicName>
|
||
is longer than it is wide.
|
||
</paragraph>
|
||
<paragraph id="8B8485738D18FE4EDDB50F55FA40B856" blockId="5.[151,1437,151,1113]" pageId="5" pageNumber="26">
|
||
At the
|
||
<typeStatus id="54803BD18D18FE4EDC600F55FE9FB967" box="[274,322,603,627]" pageId="5" pageNumber="26">type</typeStatus>
|
||
locality,
|
||
<taxonomicName id="4C3BFEF08D18FE4EDCDB0F55FDF1B967" box="[425,556,603,627]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="williamsi">
|
||
<emphasis id="B94F59618D18FE4EDCDB0F55FDF1B967" box="[425,556,603,627]" italics="true" pageId="5" pageNumber="26">S. williamsi</emphasis>
|
||
</taxonomicName>
|
||
occurs sympatrically with
|
||
<taxonomicName id="4C3BFEF08D18FE4EDE280F55FB50B967" box="[858,1165,603,627]" class="Remipedia" family="Godzilliidae" genus="Godzilliognomus" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="frondosus">
|
||
<emphasis id="B94F59618D18FE4EDE280F55FB50B967" box="[858,1165,603,627]" italics="true" pageId="5" pageNumber="26">Godzilliognomus frondosus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3BFEF08D18FE4ED9E90F55FF6BB981" class="Remipedia" family="Pleomothridae" genus="Pleomothra" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="apletocheles">
|
||
<emphasis id="B94F59618D18FE4ED9E90F55FF6BB981" italics="true" pageId="5" pageNumber="26">Pleomothra apletocheles</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3BFEF08D18FE4EDDB70F70FDDFB981" box="[197,514,637,661]" class="Remipedia" family="Speleonectidae" genus="Cryptocoynetes" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="haptodiscus">
|
||
<emphasis id="B94F59618D18FE4EDDB70F70FDDFB981" box="[197,514,637,661]" italics="true" pageId="5" pageNumber="26">Cryptocoynetes haptodiscus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C3BFEF08D18FE4EDF310F70FD0CB981" box="[579,721,637,661]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="benjamini">
|
||
<emphasis id="B94F59618D18FE4EDF310F70FD0CB981" box="[579,721,637,661]" italics="true" pageId="5" pageNumber="26">S. benjamini</emphasis>
|
||
</taxonomicName>
|
||
. Grand
|
||
<collectingCountry id="F32CC5E38D18FE4EDE5C0F73FC57B981" box="[814,906,637,661]" name="Bahamas" pageId="5" pageNumber="26">Bahama</collectingCountry>
|
||
Island is more than
|
||
<quantity id="4CC328968D18FE4ED9040F73FB07B981" box="[1142,1242,637,661]" metricMagnitude="6" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="26" unit="km" value="1100.0">1100 km</quantity>
|
||
northwest of the only known populations of
|
||
<taxonomicName id="4C3BFEF08D18FE4EDCA40FAEFD86B9AC" box="[470,603,672,696]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="emersoni">
|
||
<emphasis id="B94F59618D18FE4EDCA40FAEFD86B9AC" box="[470,603,672,696]" italics="true" pageId="5" pageNumber="26">S. emersoni</emphasis>
|
||
</taxonomicName>
|
||
, in anchialine caves near the southern coast of
|
||
<collectingCountry id="F32CC5E38D18FE4ED9F70F91FAF0B9A3" box="[1157,1325,671,695]" name="Dominican Republic" pageId="5" pageNumber="26">the Dominican</collectingCountry>
|
||
Republic (island of Hispaniola). Recognition of
|
||
<taxonomicName id="4C3BFEF08D18FE4EDF3B0FCDFD16B9CF" box="[585,715,707,731]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="williamsi">
|
||
<emphasis id="B94F59618D18FE4EDF3B0FCDFD16B9CF" box="[585,715,707,731]" italics="true" pageId="5" pageNumber="26">S. williamsi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3BFEF08D18FE4EDE710FCDFC5AB9CF" box="[771,903,707,731]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="emersoni">
|
||
<emphasis id="B94F59618D18FE4EDE710FCDFC5AB9CF" box="[771,903,707,731]" italics="true" pageId="5" pageNumber="26">S. emersoni</emphasis>
|
||
</taxonomicName>
|
||
as distinct species, in spite of their high degree of morphological similarity, suggests that the islands on which they are found may be inhabited by a series of cryptic or pseudo-cryptic species. Unfortunately, we were unable to extract usable DNA from
|
||
<typeStatus id="54803BD18D18FE4ED9D00E07FADFB835" box="[1186,1282,777,801]" pageId="5" pageNumber="26" type="paratype">paratype</typeStatus>
|
||
4. Remipede dispersal and colonization patterns, based on DNA sequence analyses, remain the subject of ongoing investigation.
|
||
</paragraph>
|
||
<paragraph id="8B8485738D18FE4EDDB50E43FAFCBF4C" blockId="5.[151,1437,151,1113]" pageId="5" pageNumber="26">
|
||
The discovery of two apparently closely related but geographically disjunct species highlights our ignorance of the evolutionary trajectories within Remipedia. It also emphasizes our uncertainty about which traits are truly diagnostic and which are plastic within a species. In the 30 years since their discovery, the boundaries between intraspecific and interspecific variation in remipedes remain unclear due to the very small number of individuals that have been collected for most species. The difficulty of accessing remipede habitat and the desire to avoid depleting what may be small populations within a given cave ensure that this will remain a problem for some time to come. The case of
|
||
<taxonomicName id="4C3BFEF08D18FE4EDD9D0910FEA9BF21" box="[239,372,1053,1077]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="williamsi">
|
||
<emphasis id="B94F59618D18FE4EDD9D0910FEA9BF21" box="[239,372,1053,1077]" italics="true" pageId="5" pageNumber="26">S. williamsi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3BFEF08D18FE4EDCDF0910FDECBF21" box="[429,561,1054,1077]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="emersoni">
|
||
<emphasis id="B94F59618D18FE4EDCDF0910FDECBF21" box="[429,561,1054,1077]" italics="true" pageId="5" pageNumber="26">S. emersoni</emphasis>
|
||
</taxonomicName>
|
||
suggests that expanding the character set to include molecular in addition to morphological variation may result in a more complete picture of remipede diversity and biogeography.
|
||
</paragraph>
|
||
<caption id="DF44D5FB8D18FE4EDDE5098BFACABF8F" ID-Table-UUID="DF44D5FB8D18FE4EDDE5098BFACABF8F" box="[151,1303,1157,1179]" httpUri="http://table.plazi.org/id/DF44D5FB8D18FE4EDDE5098BFACABF8F" pageId="5" pageNumber="26" targetBox="[159,1378,1371,1951]" targetIsTable="true" targetPageId="5">
|
||
<paragraph id="8B8485738D18FE4EDDE5098BFACABF8F" blockId="5.[151,1303,1157,1311]" box="[151,1303,1157,1179]" pageId="5" pageNumber="26">
|
||
<emphasis id="B94F59618D18FE4EDDE5098BFED9BF8E" bold="true" box="[151,260,1157,1178]" pageId="5" pageNumber="26">TABLE 1.</emphasis>
|
||
Comparison of morphological similarities and differences between
|
||
<taxonomicName id="4C3BFEF08D18FE4EDEC60988FBF6BF8F" box="[948,1067,1157,1179]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="williamsi" status="sp. nov.">
|
||
<emphasis id="B94F59618D18FE4EDEC60988FBF6BF8F" box="[948,1067,1157,1179]" italics="true" pageId="5" pageNumber="26">S. williamsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94F59618D18FE4ED9430988FBB7BF8F" bold="true" box="[1073,1130,1158,1179]" pageId="5" pageNumber="26">
|
||
<taxonomicNameLabel id="A27CE41A8D18FE4ED9430988FBB7BF8F" box="[1073,1130,1158,1179]" pageId="5" pageNumber="26" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C3BFEF08D18FE4ED9E90988FACCBF8F" box="[1179,1297,1158,1179]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="emersoni">
|
||
<emphasis id="B94F59618D18FE4ED9E90988FB7ABF8F" box="[1179,1191,1158,1179]" italics="true" pageId="5" pageNumber="26">S</emphasis>
|
||
.
|
||
<emphasis id="B94F59618D18FE4ED9C60988FACCBF8F" box="[1204,1297,1158,1179]" italics="true" pageId="5" pageNumber="26">emersoni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8485738D18FE4EDF7E09B9FB8EBFE2" blockId="5.[151,1303,1157,1311]" pageId="5" pageNumber="26">
|
||
<taxonomicName id="4C3BFEF08D18FE4EDF7E09B9FD2FBFD9" box="[524,754,1207,1229]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="williamsi" status="sp. nov.">
|
||
<emphasis id="B94F59618D18FE4EDF7E09B9FD2FBFD9" box="[524,754,1207,1229]" italics="true" pageId="5" pageNumber="26">Speleonectes williamsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94F59618D18FE4EDF8A09B9FCECBFD8" bold="true" box="[760,817,1207,1228]" pageId="5" pageNumber="26">
|
||
<taxonomicNameLabel id="A27CE41A8D18FE4EDF8A09B9FCECBFD8" box="[760,817,1207,1228]" pageId="5" pageNumber="26" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
<taxonomicName id="4C3BFEF08D18FE4EDEBF09B9FB6EBFD9" box="[973,1203,1207,1229]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" kingdom="Animalia" order="Nectiopoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="emersoni">
|
||
<emphasis id="B94F59618D18FE4EDEBF09B9FB6EBFD9" box="[973,1203,1207,1229]" italics="true" pageId="5" pageNumber="26">Speleonectes emersoni</emphasis>
|
||
</taxonomicName>
|
||
Adult body length
|
||
<quantity id="4CC328968D18FE4EDF7E09EEFD4FBFE2" box="[524,658,1248,1270]" metricMagnitude="-3" metricUnit="m" metricValue="9.649999999999999" metricValueMax="11.2" metricValueMin="8.1" pageId="5" pageNumber="26" unit="mm" value="9.649999999999999" valueMax="11.2" valueMin="8.1">8.1–11.2 mm</quantity>
|
||
<quantity id="4CC328968D18FE4EDEBF09EEFB8EBFE2" box="[973,1107,1248,1270]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.25" metricValueMin="0.95" pageId="5" pageNumber="26" unit="mm" value="11.0" valueMax="12.5" valueMin="9.5">9.5–12.5 mm</quantity>
|
||
</paragraph>
|
||
<paragraph id="8B8485738D18FE4EDDED0807FBD1BE0B" blockId="5.[151,1303,1157,1311]" box="[159,1036,1289,1311]" pageId="5" pageNumber="26">Number of trunk somites, adults 20–22 19–21</paragraph>
|
||
<paragraph id="8B8485738D18FE4EDDED0852FB70BC8B" pageId="5" pageNumber="26">
|
||
<table id="F93B77D38D1801B4DDED0855FABFBC8B" box="[159,1378,1371,1951]" gridcols="3" gridrows="12" pageId="5" pageNumber="26">
|
||
<tr id="350B87318D1801B4DDED0855FABFBE66" box="[159,1378,1371,1394]" gridrow="0" pageId="5" pageNumber="26">
|
||
<th id="76DAEE4D8D1801B4DDED0855FE3FBE66" box="[159,482,1371,1394]" gridcol="0" gridrow="0" pageId="5" pageNumber="26">A1, dorsal ramus</th>
|
||
<th id="76DAEE4D8D1801B4DF7E0855FC66BE66" box="[524,955,1371,1394]" gridcol="1" gridrow="0" pageId="5" pageNumber="26">10–11 segments</th>
|
||
<th id="76DAEE4D8D1801B4DEBF0855FABFBE66" box="[973,1378,1371,1394]" gridcol="2" gridrow="0" pageId="5" pageNumber="26">11 segments</th>
|
||
</tr>
|
||
<tr id="350B87318D1801B4DDED088BFABFBE8F" box="[159,1378,1413,1435]" gridrow="1" pageId="5" pageNumber="26">
|
||
<th id="76DAEE4D8D1801B4DDED088BFE3FBE8F" box="[159,482,1413,1435]" gridcol="0" gridrow="1" pageId="5" pageNumber="26">A1, ventral flagellum</th>
|
||
<td id="76DAEE4D8D1801B4DF7E088BFC66BE8F" box="[524,955,1413,1435]" gridcol="1" gridrow="1" pageId="5" pageNumber="26">5 segments</td>
|
||
<td id="76DAEE4D8D1801B4DEBF088BFABFBE8F" box="[973,1378,1413,1435]" gridcol="2" gridrow="1" pageId="5" pageNumber="26">6 segments</td>
|
||
</tr>
|
||
<tr id="350B87318D1801B4DDED08A0FABFBED0" box="[159,1378,1454,1476]" gridrow="2" pageId="5" pageNumber="26">
|
||
<th id="76DAEE4D8D1801B4DDED08A0FE3FBED0" box="[159,482,1454,1476]" gridcol="0" gridrow="2" pageId="5" pageNumber="26">A1, peduncular aesthetascs</th>
|
||
<td id="76DAEE4D8D1801B4DF7E08A0FC66BED0" box="[524,955,1454,1476]" gridcol="1" gridrow="2" pageId="5" pageNumber="26">ca. 20</td>
|
||
<td id="76DAEE4D8D1801B4DEBF08A0FABFBED0" box="[973,1378,1454,1476]" gridcol="2" gridrow="2" pageId="5" pageNumber="26">ca. 30</td>
|
||
</tr>
|
||
<tr id="350B87318D1801B4DDED08D6FABFBEFA" box="[159,1378,1496,1518]" gridrow="3" pageId="5" pageNumber="26">
|
||
<th id="76DAEE4D8D1801B4DDED08D6FE3FBEFA" box="[159,482,1496,1518]" gridcol="0" gridrow="3" pageId="5" pageNumber="26">Setae of A2, exopod</th>
|
||
<td id="76DAEE4D8D1801B4DF7E08D6FC66BEFA" box="[524,955,1496,1518]" gridcol="1" gridrow="3" pageId="5" pageNumber="26">26 long</td>
|
||
<td id="76DAEE4D8D1801B4DEBF08D6FABFBEFA" box="[973,1378,1496,1518]" gridcol="2" gridrow="3" pageId="5" pageNumber="26">15 long, 3 short</td>
|
||
</tr>
|
||
<tr id="350B87318D1801B4DDED0B0FFABFBD03" box="[159,1378,1537,1559]" gridrow="4" pageId="5" pageNumber="26">
|
||
<th id="76DAEE4D8D1801B4DDED0B0FFE3FBD03" box="[159,482,1537,1559]" gridcol="0" gridrow="4" pageId="5" pageNumber="26">FF, main filament</th>
|
||
<td id="76DAEE4D8D1801B4DF7E0B0FFC66BD03" box="[524,955,1537,1559]" gridcol="1" gridrow="4" pageId="5" pageNumber="26">3.1 times longer than wide</td>
|
||
<td id="76DAEE4D8D1801B4DEBF0B0FFABFBD03" box="[973,1378,1537,1559]" gridcol="2" gridrow="4" pageId="5" pageNumber="26">5 times longer than wide</td>
|
||
</tr>
|
||
<tr id="350B87318D1801B4DDED0B24FABFBD54" box="[159,1378,1578,1600]" gridrow="5" pageId="5" pageNumber="26">
|
||
<th id="76DAEE4D8D1801B4DDED0B24FE3FBD54" box="[159,482,1578,1600]" gridcol="0" gridrow="5" pageId="5" pageNumber="26">FF, medial process</th>
|
||
<td id="76DAEE4D8D1801B4DF7E0B24FC66BD54" box="[524,955,1578,1600]" gridcol="1" gridrow="5" pageId="5" pageNumber="26">40% length of main filament</td>
|
||
<td id="76DAEE4D8D1801B4DEBF0B24FABFBD54" box="[973,1378,1578,1600]" gridcol="2" gridrow="5" pageId="5" pageNumber="26">32% length of main filament</td>
|
||
</tr>
|
||
<tr id="350B87318D1801B4DDED0B5DFABFBD7E" box="[159,1378,1619,1642]" gridrow="6" pageId="5" pageNumber="26">
|
||
<th id="76DAEE4D8D1801B4DDED0B5DFE3FBD7E" box="[159,482,1619,1642]" gridcol="0" gridrow="6" pageId="5" pageNumber="26">Mandibles, right lacinia mobilis</th>
|
||
<td id="76DAEE4D8D1801B4DF7E0B5DFC66BD7E" box="[524,955,1619,1642]" gridcol="1" gridrow="6" pageId="5" pageNumber="26">outer denticle with flattened apical surface</td>
|
||
<td id="76DAEE4D8D1801B4DEBF0B5DFABFBD7E" box="[973,1378,1619,1642]" gridcol="2" gridrow="6" pageId="5" pageNumber="26">outer denticle with pointed tip</td>
|
||
</tr>
|
||
<tr id="350B87318D1801B4DDED0BA8FABFBDCF" box="[159,1378,1702,1755]" gridrow="7" pageId="5" pageNumber="26">
|
||
<th id="76DAEE4D8D1801B4DDED0BA8FE3FBDCF" box="[159,482,1702,1755]" gridcol="0" gridrow="7" pageId="5" pageNumber="26">Max, segment 4</th>
|
||
<td id="76DAEE4D8D1801B4DF7E0BA8FC66BDCF" box="[524,955,1702,1755]" gridcol="1" gridrow="7" pageId="5" pageNumber="26">as long as lacertus, slightly expanded</td>
|
||
<td id="76DAEE4D8D1801B4DEBF0BA8FABFBDCF" box="[973,1378,1702,1755]" gridcol="2" gridrow="7" pageId="5" pageNumber="26">shorter than lacertus, with distinct distal expansion</td>
|
||
</tr>
|
||
<tr id="350B87318D1801B4DDED0BE0FABFBC10" box="[159,1378,1774,1796]" gridrow="8" pageId="5" pageNumber="26">
|
||
<th id="76DAEE4D8D1801B4DDED0BE0FE3FBC10" box="[159,482,1774,1796]" gridcol="0" gridrow="8" pageId="5" pageNumber="26">Max, segment 6</th>
|
||
<td id="76DAEE4D8D1801B4DF7E0BE0FC66BC10" box="[524,955,1774,1796]" gridcol="1" gridrow="8" pageId="5" pageNumber="26">shorter than segment 5</td>
|
||
<td id="76DAEE4D8D1801B4DEBF0BE0FABFBC10" box="[973,1378,1774,1796]" gridcol="2" gridrow="8" pageId="5" pageNumber="26">as long as segment 5</td>
|
||
</tr>
|
||
<tr id="350B87318D1801B4DDED0A16FABFBC3A" box="[159,1378,1816,1838]" gridrow="9" pageId="5" pageNumber="26">
|
||
<th id="76DAEE4D8D1801B4DDED0A16FE3FBC3A" box="[159,482,1816,1838]" gridcol="0" gridrow="9" pageId="5" pageNumber="26">Max, Mxp, terminal claws</th>
|
||
<td id="76DAEE4D8D1801B4DF7E0A16FC66BC3A" box="[524,955,1816,1838]" gridcol="1" gridrow="9" pageId="5" pageNumber="26">with ca. 8 deeply incised denticles</td>
|
||
<td id="76DAEE4D8D1801B4DEBF0A16FABFBC3A" box="[973,1378,1816,1838]" gridcol="2" gridrow="9" pageId="5" pageNumber="26">with 7–10 small (fused) denticles</td>
|
||
</tr>
|
||
<tr id="350B87318D1801B4DDED0A4FFABFBC62" box="[159,1378,1857,1910]" gridrow="10" pageId="5" pageNumber="26">
|
||
<th id="76DAEE4D8D1801B4DDED0A4FFE3FBC62" box="[159,482,1857,1910]" gridcol="0" gridrow="10" pageId="5" pageNumber="26">Trunk limbs, short stout corner setae</th>
|
||
<td id="76DAEE4D8D1801B4DF7E0A4FFC66BC62" box="[524,955,1857,1910]" gridcol="1" gridrow="10" pageId="5" pageNumber="26">plumose and serrate types</td>
|
||
<td id="76DAEE4D8D1801B4DEBF0A4FFABFBC62" box="[973,1378,1857,1910]" gridcol="2" gridrow="10" pageId="5" pageNumber="26">serrate type</td>
|
||
</tr>
|
||
<tr id="350B87318D1801B4DDED0A87FABFBC8B" box="[159,1378,1929,1951]" gridrow="11" pageId="5" pageNumber="26">
|
||
<th id="76DAEE4D8D1801B4DDED0A87FE3FBC8B" box="[159,482,1929,1951]" gridcol="0" gridrow="11" pageId="5" pageNumber="26">Trunk limbs, shape</th>
|
||
<td id="76DAEE4D8D1801B4DF7E0A87FC66BC8B" box="[524,955,1929,1951]" gridcol="1" gridrow="11" pageId="5" pageNumber="26">sturdy, robust</td>
|
||
<td id="76DAEE4D8D1801B4DEBF0A87FABFBC8B" box="[973,1378,1929,1951]" gridcol="2" gridrow="11" pageId="5" pageNumber="26">comparatively slender</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8B8485738D18FE4EDDE50ACEFBDBBCE0" blockId="5.[151,1436,1984,2036]" pageId="5" pageNumber="26">Abbreviations: A1 = antennule (first antenna); A2 = antenna (second antenna); FF = frontal filament; Max = maxilla (second maxilla); Mxp = maxilliped. See Morphological Comparisons for further explanations.</paragraph>
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</subSubSection>
|
||
</treatment>
|
||
</document> |