386 lines
40 KiB
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386 lines
40 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/alpento.6.97595" ID-GBIF-Dataset="4c4bf0af-a375-477b-bd50-e6ffdb0d475a" ID-Pensoft-Pub="2535-0889-6-147" ID-Pensoft-UUID="E924B3F1494452F192E8D36AA3256A1B" ID-ZooBank="74F9780D4AEC4CDFAC5168F827CB87BB" ModsDocID="2535-0889-6-147" checkinTime="1670417673427" checkinUser="pensoft" docAuthor="Burckhardt, Daniel" docDate="2022" docId="B380492DADBD5C59A4E88983207BAFF7" docLanguage="en" docName="AlpEntomol 6: 147-151" docOrigin="Alpine Entomology 6" docPubDate="2022-12-06" docSource="http://dx.doi.org/10.3897/alpento.6.97595" docTitle="Eryngiofaga perrara Burckhardt 2022, sp. nov." docType="treatment" docUuid="153CB420-A251-4F01-A70D-6B9787EF2520" docUuidSource="ZooBank" docVersion="2" id="E924B3F1494452F192E8D36AA3256A1B" lastPageNumber="147" masterDocId="E924B3F1494452F192E8D36AA3256A1B" masterDocTitle="Eryngiofaga perrara sp. nov. (Hemiptera, Psylloidea) from Mount Pilatus (Obwalden), a new species of a genus previously unknown from Switzerland or the Alps" masterLastPageNumber="151" masterPageNumber="147" pageNumber="147" updateTime="1670418276596" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Eryngiofaga perrara sp. nov. (Hemiptera, Psylloidea) from Mount Pilatus (Obwalden), a new species of a genus previously unknown from Switzerland or the Alps</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Burckhardt, Daniel</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8368-5268</mods:nameIdentifier>
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<mods:affiliation>Naturhistorisches Museum, Augustinergasse 2, 4001 Basel, Switzerland</mods:affiliation>
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<mods:nameIdentifier type="email">daniel.burckhardt@bs.ch</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Alpine Entomology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-12-06</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>6</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>147</mods:start>
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<mods:end>151</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/alpento.6.97595</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/alpento.6.97595</mods:identifier>
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<mods:identifier type="Pensoft-Pub">2535-0889-6-147</mods:identifier>
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<mods:identifier type="ZooBank">74F9780D4AEC4CDFAC5168F827CB87BB</mods:identifier>
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<mods:identifier type="Pensoft-UUID">E924B3F1494452F192E8D36AA3256A1B</mods:identifier>
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</mods:mods>
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<subSection pageId="0" pageNumber="147" type="results">
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<treatment LSID="urn:lsid:zoobank.org:act:153CB420-A251-4F01-A70D-6B9787EF2520" httpUri="http://treatment.plazi.org/id/B380492DADBD5C59A4E88983207BAFF7" lastPageNumber="147" pageId="0" pageNumber="147">
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<subSubSection pageId="0" pageNumber="147" type="nomenclature">
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<paragraph pageId="0" pageNumber="147">
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<taxonomicName LSID="https://zoobank.org/153CB420-A251-4F01-A70D-6B9787EF2520" authority="Burckhardt, 2022" authorityName="Burckhardt" authorityYear="2022" class="Insecta" family="Triozidae" genus="Eryngiofaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eryngiofaga perrara" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="perrara" status="sp. nov.">Eryngiofaga perrara</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="147">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="147" type="description">
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<paragraph pageId="0" pageNumber="147">
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<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Eryngiofaga perrara sp. nov. 1 - 3. Habitus, adults; 1. Male, in lateral view; 2. Male, in dorsal view; 3. Female in lateral view; 4. Head, in dorsal view. Scale bars: 0.5 mm (1 - 3); 0.2 mm (4)." figureDoi="10.3897/alpento.6.97595.figures1-4" httpUri="https://binary.pensoft.net/fig/778232" pageId="0" pageNumber="147">Figs 1-4</figureCitation>
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<figureCitation captionStart="Figures 5–8" captionStartId="F2" captionText="Figures 5 - 8. Eryngiofaga perrara sp. nov., terminalia. 5. Male terminalia, in lateral view; 6. Inner face of paramere; 7. Distal portion of aedeagus; 8. Female terminalia, in profile. Scale bars: 0.1 mm (5, 8); 0.05 mm (6, 7)." figureDoi="10.3897/alpento.6.97595.figures5-8" httpUri="https://binary.pensoft.net/fig/778233" pageId="0" pageNumber="147">, 5-8</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="147" type="type locality">
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<paragraph pageId="0" pageNumber="147">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="147">
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Switzerland, Obwalden, Pilatus, Chilchsteine,
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<geoCoordinate degrees="46.9758" direction="north" orientation="latitude" precision="5" value="46.9758">46.9758°N</geoCoordinate>
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,
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<geoCoordinate degrees="8.2542" direction="east" orientation="longitude" precision="5" value="8.2542">8.2542°E</geoCoordinate>
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, 1850 m.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="147" type="holotype">
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<paragraph pageId="0" pageNumber="147">Holotype.</paragraph>
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<paragraph pageId="0" pageNumber="147">
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Male. Switzerland: Obwalden, Pilatus, Chilchsteine,
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<geoCoordinate degrees="46.9758" direction="north" orientation="latitude" precision="5" value="46.9758">46.9758°N</geoCoordinate>
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,
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<geoCoordinate degrees="8.2542" direction="east" orientation="longitude" precision="5" value="8.2542">8.2542°E</geoCoordinate>
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, 1850 m, 12.vii.2010, D. Burckhardt & I.
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<normalizedToken originalValue="Zürcher">Zuercher</normalizedToken>
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// herbaceous vegetation with
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<taxonomicName class="Magnoliopsida" family="Rosaceae" genus="Alchemilla" kingdom="Plantae" lsidName="Alchemilla" order="Rosales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="147">Alchemilla</emphasis>
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</taxonomicName>
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,
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<taxonomicName class="Magnoliopsida" family="Apiaceae" genus="Astrantia" kingdom="Plantae" lsidName="Astrantia" order="Apiales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="147">Astrantia</emphasis>
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</taxonomicName>
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,
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<taxonomicName class="Magnoliopsida" family="Apiaceae" genus="Bupleurum" kingdom="Plantae" lsidName="Bupleurum" order="Apiales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="147">Bupleurum</emphasis>
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</taxonomicName>
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,
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<taxonomicName class="Magnoliopsida" family="Caryophyllaceae" genus="Cerastium" kingdom="Plantae" lsidName="Cerastium" order="Caryophyllales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="147">Cerastium</emphasis>
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</taxonomicName>
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//
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<taxonomicName authorityName="Burckhardt" authorityYear="2022" class="Insecta" family="Triozidae" genus="Eryngiofaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eryngiofaga perrara" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="perrara">
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<emphasis italics="true" pageId="0" pageNumber="147">Eryngiofaga perrara</emphasis>
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</taxonomicName>
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sp. nov., holotype, det. D. Burckhardt, 2022 // NMB-PSYLL0007231 // NHMB, dry.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="147" type="paratype">
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<paragraph pageId="0" pageNumber="147">Paratype.</paragraph>
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<paragraph pageId="0" pageNumber="147">Switzerland, 1 female, Obwalden, same data as holotype but 18.vii.2000, D. Burckhardt // herbaceous vegetation // PSYLL NHMB 00002446 // NHMB, dry.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="147" type="diagnosis">
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<paragraph pageId="0" pageNumber="147">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="147">
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Adult yellow to orange (Figs
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<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Eryngiofaga perrara sp. nov. 1 - 3. Habitus, adults; 1. Male, in lateral view; 2. Male, in dorsal view; 3. Female in lateral view; 4. Head, in dorsal view. Scale bars: 0.5 mm (1 - 3); 0.2 mm (4)." figureDoi="10.3897/alpento.6.97595.figures1-4" httpUri="https://binary.pensoft.net/fig/778232" pageId="0" pageNumber="147">1-3</figureCitation>
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); genal processes light apically (Fig.
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<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Eryngiofaga perrara sp. nov. 1 - 3. Habitus, adults; 1. Male, in lateral view; 2. Male, in dorsal view; 3. Female in lateral view; 4. Head, in dorsal view. Scale bars: 0.5 mm (1 - 3); 0.2 mm (4)." figureDoi="10.3897/alpento.6.97595.figures1-4" httpUri="https://binary.pensoft.net/fig/778232" pageId="0" pageNumber="147">4</figureCitation>
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); with more or less distinct greyish brown longitudinal medial stripe on head and thorax. Antennal segments 1-3 yellow, segments 4-10 almost black. Forewing transparent, colourless. - Genal processes conical, subacute apically, 0.3-0.4 times as long as vertex along midline. Antenna 1.9-2.2 times as long as head width. Forewing (Figs
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<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Eryngiofaga perrara sp. nov. 1 - 3. Habitus, adults; 1. Male, in lateral view; 2. Male, in dorsal view; 3. Female in lateral view; 4. Head, in dorsal view. Scale bars: 0.5 mm (1 - 3); 0.2 mm (4)." figureDoi="10.3897/alpento.6.97595.figures1-4" httpUri="https://binary.pensoft.net/fig/778232" pageId="0" pageNumber="147">1</figureCitation>
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,
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<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Eryngiofaga perrara sp. nov. 1 - 3. Habitus, adults; 1. Male, in lateral view; 2. Male, in dorsal view; 3. Female in lateral view; 4. Head, in dorsal view. Scale bars: 0.5 mm (1 - 3); 0.2 mm (4)." figureDoi="10.3897/alpento.6.97595.figures1-4" httpUri="https://binary.pensoft.net/fig/778232" pageId="0" pageNumber="147">3</figureCitation>
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) 4.3-4.4 times as long as head width, 2.8 times as long as wide; costal margin strongly, evenly curved, anal margin almost straight; vein Rs short, distinctly concave; surface spinules lacking apart from base of wing. Paramere bifid (Figs
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<figureCitation captionStart="Figures 5–8" captionStartId="F2" captionText="Figures 5 - 8. Eryngiofaga perrara sp. nov., terminalia. 5. Male terminalia, in lateral view; 6. Inner face of paramere; 7. Distal portion of aedeagus; 8. Female terminalia, in profile. Scale bars: 0.1 mm (5, 8); 0.05 mm (6, 7)." figureDoi="10.3897/alpento.6.97595.figures5-8" httpUri="https://binary.pensoft.net/fig/778233" pageId="0" pageNumber="147">5</figureCitation>
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,
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<figureCitation captionStart="Figures 5–8" captionStartId="F2" captionText="Figures 5 - 8. Eryngiofaga perrara sp. nov., terminalia. 5. Male terminalia, in lateral view; 6. Inner face of paramere; 7. Distal portion of aedeagus; 8. Female terminalia, in profile. Scale bars: 0.1 mm (5, 8); 0.05 mm (6, 7)." figureDoi="10.3897/alpento.6.97595.figures5-8" httpUri="https://binary.pensoft.net/fig/778233" pageId="0" pageNumber="147">6</figureCitation>
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); with narrow inner anterior process which is sclerotised apically bearing each a small anterior and posterior toothlet, and with broader outer posterior lobe which is irregularly narrowing to strongly sclerotised apex forming apical tooth. Distal segment of aedeagus (Fig.
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<figureCitation captionStart="Figures 5–8" captionStartId="F2" captionText="Figures 5 - 8. Eryngiofaga perrara sp. nov., terminalia. 5. Male terminalia, in lateral view; 6. Inner face of paramere; 7. Distal portion of aedeagus; 8. Female terminalia, in profile. Scale bars: 0.1 mm (5, 8); 0.05 mm (6, 7)." figureDoi="10.3897/alpento.6.97595.figures5-8" httpUri="https://binary.pensoft.net/fig/778233" pageId="0" pageNumber="147">7</figureCitation>
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), hardly expanded basally, weakly expanded apically with small subapical hook ventrally. Female terminalia (Fig.
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<figureCitation captionStart="Figures 5–8" captionStartId="F2" captionText="Figures 5 - 8. Eryngiofaga perrara sp. nov., terminalia. 5. Male terminalia, in lateral view; 6. Inner face of paramere; 7. Distal portion of aedeagus; 8. Female terminalia, in profile. Scale bars: 0.1 mm (5, 8); 0.05 mm (6, 7)." figureDoi="10.3897/alpento.6.97595.figures5-8" httpUri="https://binary.pensoft.net/fig/778233" pageId="0" pageNumber="147">8</figureCitation>
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) short; proctiger ending in small apical tubercle.
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</paragraph>
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<caption doi="10.3897/alpento.6.97595.figures1-4" httpUri="https://binary.pensoft.net/fig/778232" pageId="0" pageNumber="147" start="Figures 1–4" startId="F1">
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<paragraph pageId="0" pageNumber="147">
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<emphasis bold="true" pageId="0" pageNumber="147">Figures 1-4.</emphasis>
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<taxonomicName authorityName="Burckhardt" authorityYear="2022" class="Insecta" family="Triozidae" genus="Eryngiofaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eryngiofaga perrara" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="perrara">
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<emphasis italics="true" pageId="0" pageNumber="147">Eryngiofaga perrara</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="147">1-3.</emphasis>
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Habitus, adults;
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<emphasis bold="true" pageId="0" pageNumber="147">1.</emphasis>
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Male, in lateral view;
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<emphasis bold="true" pageId="0" pageNumber="147">2.</emphasis>
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Male, in dorsal view;
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<emphasis bold="true" pageId="0" pageNumber="147">3.</emphasis>
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Female in lateral view;
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<emphasis bold="true" pageId="0" pageNumber="147">4.</emphasis>
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Head, in dorsal view. Scale bars: 0.5 mm (
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<emphasis bold="true" pageId="0" pageNumber="147">1-3</emphasis>
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); 0.2 mm (
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<emphasis bold="true" pageId="0" pageNumber="147">4</emphasis>
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).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="147" type="description">
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<paragraph pageId="0" pageNumber="147">Description.</paragraph>
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<paragraph pageId="0" pageNumber="147">
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Adult (Figs
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<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Eryngiofaga perrara sp. nov. 1 - 3. Habitus, adults; 1. Male, in lateral view; 2. Male, in dorsal view; 3. Female in lateral view; 4. Head, in dorsal view. Scale bars: 0.5 mm (1 - 3); 0.2 mm (4)." figureDoi="10.3897/alpento.6.97595.figures1-4" httpUri="https://binary.pensoft.net/fig/778232" pageId="0" pageNumber="147">1</figureCitation>
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-
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<figureCitation captionStart="Figures 5–8" captionStartId="F2" captionText="Figures 5 - 8. Eryngiofaga perrara sp. nov., terminalia. 5. Male terminalia, in lateral view; 6. Inner face of paramere; 7. Distal portion of aedeagus; 8. Female terminalia, in profile. Scale bars: 0.1 mm (5, 8); 0.05 mm (6, 7)." figureDoi="10.3897/alpento.6.97595.figures5-8" httpUri="https://binary.pensoft.net/fig/778233" pageId="0" pageNumber="147">8</figureCitation>
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). Colouration. Yellow to orange (Figs
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<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Eryngiofaga perrara sp. nov. 1 - 3. Habitus, adults; 1. Male, in lateral view; 2. Male, in dorsal view; 3. Female in lateral view; 4. Head, in dorsal view. Scale bars: 0.5 mm (1 - 3); 0.2 mm (4)." figureDoi="10.3897/alpento.6.97595.figures1-4" httpUri="https://binary.pensoft.net/fig/778232" pageId="0" pageNumber="147">1</figureCitation>
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,
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<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Eryngiofaga perrara sp. nov. 1 - 3. Habitus, adults; 1. Male, in lateral view; 2. Male, in dorsal view; 3. Female in lateral view; 4. Head, in dorsal view. Scale bars: 0.5 mm (1 - 3); 0.2 mm (4)." figureDoi="10.3897/alpento.6.97595.figures1-4" httpUri="https://binary.pensoft.net/fig/778232" pageId="0" pageNumber="147">3</figureCitation>
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). Head and thorax with a greyish brown longitudinal medial stripe, very light in male (Fig.
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<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Eryngiofaga perrara sp. nov. 1 - 3. Habitus, adults; 1. Male, in lateral view; 2. Male, in dorsal view; 3. Female in lateral view; 4. Head, in dorsal view. Scale bars: 0.5 mm (1 - 3); 0.2 mm (4)." figureDoi="10.3897/alpento.6.97595.figures1-4" httpUri="https://binary.pensoft.net/fig/778232" pageId="0" pageNumber="147">2</figureCitation>
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), darker in female. Tips of genal processes light (Fig.
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<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Eryngiofaga perrara sp. nov. 1 - 3. Habitus, adults; 1. Male, in lateral view; 2. Male, in dorsal view; 3. Female in lateral view; 4. Head, in dorsal view. Scale bars: 0.5 mm (1 - 3); 0.2 mm (4)." figureDoi="10.3897/alpento.6.97595.figures1-4" httpUri="https://binary.pensoft.net/fig/778232" pageId="0" pageNumber="147">4</figureCitation>
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). Eyes red to greyish. Antennal segments 1-3 yellow, segments 4-10 almost black (Figs
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<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Eryngiofaga perrara sp. nov. 1 - 3. Habitus, adults; 1. Male, in lateral view; 2. Male, in dorsal view; 3. Female in lateral view; 4. Head, in dorsal view. Scale bars: 0.5 mm (1 - 3); 0.2 mm (4)." figureDoi="10.3897/alpento.6.97595.figures1-4" httpUri="https://binary.pensoft.net/fig/778232" pageId="0" pageNumber="147">1-3</figureCitation>
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). Legs yellow with apical tarsal segments dark brown; pro- and mesofemora dark brown laterally; pro- and mesotibiae yellow in basal quarter, greyish brown otherwise; basal segment of pro- and mesotarsi greyish brown. Forewing (Figs
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<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Eryngiofaga perrara sp. nov. 1 - 3. Habitus, adults; 1. Male, in lateral view; 2. Male, in dorsal view; 3. Female in lateral view; 4. Head, in dorsal view. Scale bars: 0.5 mm (1 - 3); 0.2 mm (4)." figureDoi="10.3897/alpento.6.97595.figures1-4" httpUri="https://binary.pensoft.net/fig/778232" pageId="0" pageNumber="147">1</figureCitation>
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,
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<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Eryngiofaga perrara sp. nov. 1 - 3. Habitus, adults; 1. Male, in lateral view; 2. Male, in dorsal view; 3. Female in lateral view; 4. Head, in dorsal view. Scale bars: 0.5 mm (1 - 3); 0.2 mm (4)." figureDoi="10.3897/alpento.6.97595.figures1-4" httpUri="https://binary.pensoft.net/fig/778232" pageId="0" pageNumber="147">3</figureCitation>
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) transparent, colourless with yellow to light brown veins. Hindwing whitish, transparent. Abdominal tergites in male ochreous, in female brown. Male with slightly less expanded and distinct dark colour.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="147" type="structure">
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<paragraph pageId="0" pageNumber="147">Structure.</paragraph>
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<paragraph pageId="0" pageNumber="147">
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Conforming to the generic description of
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<bibRefCitation author="Loginova, MM" journalOrPublisher="Zoologicheskii Zhurnal" pageId="0" pageNumber="147" pagination="510 - 521" refId="B12" refString="Loginova, MM, 1977. A review of the genus Eryngiofaga Klimasz. (Triozidae, Homoptera). Zoologicheskii Zhurnal 56 (4): 510 - 521" title="A review of the genus Eryngiofaga Klimasz. (Triozidae, Homoptera)." volume="56" year="1977">Loginova (1977)</bibRefCitation>
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. Body length ♂ 2.5 mm, ♀ 2.7 mm (1 ♂, 1 ♀). Head hardly inclined from longitudinal body axis (Figs
|
||
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Eryngiofaga perrara sp. nov. 1 - 3. Habitus, adults; 1. Male, in lateral view; 2. Male, in dorsal view; 3. Female in lateral view; 4. Head, in dorsal view. Scale bars: 0.5 mm (1 - 3); 0.2 mm (4)." figureDoi="10.3897/alpento.6.97595.figures1-4" httpUri="https://binary.pensoft.net/fig/778232" pageId="0" pageNumber="147">1</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Eryngiofaga perrara sp. nov. 1 - 3. Habitus, adults; 1. Male, in lateral view; 2. Male, in dorsal view; 3. Female in lateral view; 4. Head, in dorsal view. Scale bars: 0.5 mm (1 - 3); 0.2 mm (4)." figureDoi="10.3897/alpento.6.97595.figures1-4" httpUri="https://binary.pensoft.net/fig/778232" pageId="0" pageNumber="147">3</figureCitation>
|
||
). Vertex subtrapezoidal (Fig.
|
||
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Eryngiofaga perrara sp. nov. 1 - 3. Habitus, adults; 1. Male, in lateral view; 2. Male, in dorsal view; 3. Female in lateral view; 4. Head, in dorsal view. Scale bars: 0.5 mm (1 - 3); 0.2 mm (4)." figureDoi="10.3897/alpento.6.97595.figures1-4" httpUri="https://binary.pensoft.net/fig/778232" pageId="0" pageNumber="147">4</figureCitation>
|
||
). Genal processes conical, subacute apically, 0.3-0.4 times as long as vertex along midline (Fig.
|
||
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Eryngiofaga perrara sp. nov. 1 - 3. Habitus, adults; 1. Male, in lateral view; 2. Male, in dorsal view; 3. Female in lateral view; 4. Head, in dorsal view. Scale bars: 0.5 mm (1 - 3); 0.2 mm (4)." figureDoi="10.3897/alpento.6.97595.figures1-4" httpUri="https://binary.pensoft.net/fig/778232" pageId="0" pageNumber="147">4</figureCitation>
|
||
). Antenna 1.9-2.2 times as long as head width; relative length of flagellar segments as 1.0: 0.3: 0.3: 0.3: 0.3: 0.2: 0.2: 0.2; segment 3 longer than segments 4-6 together; relative length of segment 10 and terminal setae as 1.0: 0.7: 0.4. Rostrum short, only tip of apical segment visible in lateral view. Metatibia 0.9-1.2 times as long as head width. Forewing (Figs
|
||
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Eryngiofaga perrara sp. nov. 1 - 3. Habitus, adults; 1. Male, in lateral view; 2. Male, in dorsal view; 3. Female in lateral view; 4. Head, in dorsal view. Scale bars: 0.5 mm (1 - 3); 0.2 mm (4)." figureDoi="10.3897/alpento.6.97595.figures1-4" httpUri="https://binary.pensoft.net/fig/778232" pageId="0" pageNumber="147">1</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Eryngiofaga perrara sp. nov. 1 - 3. Habitus, adults; 1. Male, in lateral view; 2. Male, in dorsal view; 3. Female in lateral view; 4. Head, in dorsal view. Scale bars: 0.5 mm (1 - 3); 0.2 mm (4)." figureDoi="10.3897/alpento.6.97595.figures1-4" httpUri="https://binary.pensoft.net/fig/778232" pageId="0" pageNumber="147">3</figureCitation>
|
||
) 4.3-4.4 times as long as head width, 2.8 times as long as wide; costal margin strongly, evenly curved, anal margin almost straight; wing widest near the middle; subacute apically, wing apex lying in cell r2; vein Rs short, distinctly concave; m1 cell value 1.2-1.5, cell cu1 value 1.7-1.9; surface spinules lacking apart from base of wing; radular spinules present in cells m1, m2 and cu1. Hindwing two thirds length of forewing, membranous; costal setae not grouped. Terminalia as in Figs
|
||
<figureCitation captionStart="Figures 5–8" captionStartId="F2" captionText="Figures 5 - 8. Eryngiofaga perrara sp. nov., terminalia. 5. Male terminalia, in lateral view; 6. Inner face of paramere; 7. Distal portion of aedeagus; 8. Female terminalia, in profile. Scale bars: 0.1 mm (5, 8); 0.05 mm (6, 7)." figureDoi="10.3897/alpento.6.97595.figures5-8" httpUri="https://binary.pensoft.net/fig/778233" pageId="0" pageNumber="147">5-8</figureCitation>
|
||
. Male proctiger (Fig.
|
||
<figureCitation captionStart="Figures 5–8" captionStartId="F2" captionText="Figures 5 - 8. Eryngiofaga perrara sp. nov., terminalia. 5. Male terminalia, in lateral view; 6. Inner face of paramere; 7. Distal portion of aedeagus; 8. Female terminalia, in profile. Scale bars: 0.1 mm (5, 8); 0.05 mm (6, 7)." figureDoi="10.3897/alpento.6.97595.figures5-8" httpUri="https://binary.pensoft.net/fig/778233" pageId="0" pageNumber="147">5</figureCitation>
|
||
) 0.5 times as long as head width, sparsely beset with long setae, weakly produced posteriorly; posterior margin slightly angular in basal third. Paramere (Figs
|
||
<figureCitation captionStart="Figures 5–8" captionStartId="F2" captionText="Figures 5 - 8. Eryngiofaga perrara sp. nov., terminalia. 5. Male terminalia, in lateral view; 6. Inner face of paramere; 7. Distal portion of aedeagus; 8. Female terminalia, in profile. Scale bars: 0.1 mm (5, 8); 0.05 mm (6, 7)." figureDoi="10.3897/alpento.6.97595.figures5-8" httpUri="https://binary.pensoft.net/fig/778233" pageId="0" pageNumber="147">5</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 5–8" captionStartId="F2" captionText="Figures 5 - 8. Eryngiofaga perrara sp. nov., terminalia. 5. Male terminalia, in lateral view; 6. Inner face of paramere; 7. Distal portion of aedeagus; 8. Female terminalia, in profile. Scale bars: 0.1 mm (5, 8); 0.05 mm (6, 7)." figureDoi="10.3897/alpento.6.97595.figures5-8" httpUri="https://binary.pensoft.net/fig/778233" pageId="0" pageNumber="147">6</figureCitation>
|
||
) bifid; in lateral view, with narrow inner anterior process which is sclerotised apically, bearing each a small anterior and posterior toothlet, and with broader outer posterior lobe which is irregularly narrowing to strongly sclerotised apex forming an apical tooth; the outer face bears long setae mostly on the outer posterior lobe; the inner face with long setae along the anterior margin in basal half and on the outer posterior lobe in apical two thirds. Distal segment of aedeagus (Fig.
|
||
<figureCitation captionStart="Figures 5–8" captionStartId="F2" captionText="Figures 5 - 8. Eryngiofaga perrara sp. nov., terminalia. 5. Male terminalia, in lateral view; 6. Inner face of paramere; 7. Distal portion of aedeagus; 8. Female terminalia, in profile. Scale bars: 0.1 mm (5, 8); 0.05 mm (6, 7)." figureDoi="10.3897/alpento.6.97595.figures5-8" httpUri="https://binary.pensoft.net/fig/778233" pageId="0" pageNumber="147">7</figureCitation>
|
||
) hardly expanded basally, weakly expanded apically, bearing small subapical hook ventrally; sclerotised end tube of ductus ejaculatorius moderately long, weakly sinuous. Female terminalia (Fig.
|
||
<figureCitation captionStart="Figures 5–8" captionStartId="F2" captionText="Figures 5 - 8. Eryngiofaga perrara sp. nov., terminalia. 5. Male terminalia, in lateral view; 6. Inner face of paramere; 7. Distal portion of aedeagus; 8. Female terminalia, in profile. Scale bars: 0.1 mm (5, 8); 0.05 mm (6, 7)." figureDoi="10.3897/alpento.6.97595.figures5-8" httpUri="https://binary.pensoft.net/fig/778233" pageId="0" pageNumber="147">8</figureCitation>
|
||
) short; proctiger 0.8 times as long as head width, sparsely beset with long setae in the middle and short setae apically; dorsal outline, in lateral view, strongly narrowing towards apex which forms a small tubercle; circumanal ring oval, 0.6 times as long as proctiger, consisting of two unequal rows of pores. Female subgenital plate 0.6 times as long as proctiger, acute apically; sparsely beset with moderately long setae laterally. Dorsal valvulae, in lateral view, triangular; ventral valvulae straight, lacking teeth.
|
||
</paragraph>
|
||
<caption doi="10.3897/alpento.6.97595.figures5-8" httpUri="https://binary.pensoft.net/fig/778233" pageId="0" pageNumber="147" start="Figures 5–8" startId="F2">
|
||
<paragraph pageId="0" pageNumber="147">
|
||
<emphasis bold="true" pageId="0" pageNumber="147">Figures 5-8.</emphasis>
|
||
<taxonomicName authorityName="Burckhardt" authorityYear="2022" class="Insecta" family="Triozidae" genus="Eryngiofaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eryngiofaga perrara" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="perrara">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">Eryngiofaga perrara</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., terminalia.
|
||
<emphasis bold="true" pageId="0" pageNumber="147">5.</emphasis>
|
||
Male terminalia, in lateral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="147">6.</emphasis>
|
||
Inner face of paramere;
|
||
<emphasis bold="true" pageId="0" pageNumber="147">7.</emphasis>
|
||
Distal portion of aedeagus;
|
||
<emphasis bold="true" pageId="0" pageNumber="147">8.</emphasis>
|
||
Female terminalia, in profile. Scale bars: 0.1 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="147">5, 8</emphasis>
|
||
); 0.05 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="147">6, 7</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="147" type="description">
|
||
<paragraph pageId="0" pageNumber="147">Measurements</paragraph>
|
||
<paragraph pageId="0" pageNumber="147">
|
||
<emphasis bold="true" pageId="0" pageNumber="147">(in mm; 1 ♂, 1 ♀).</emphasis>
|
||
Head width 0.46-0.50; antenna length 0.96-1.00; forewing length 1.98-2.20; male proctiger length 0.24; paramere length 0.16; length of distal segment of aedeagus 0.22; female proctiger length 0.40.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="147" type="immature">
|
||
<paragraph pageId="0" pageNumber="147">Immature</paragraph>
|
||
<paragraph pageId="0" pageNumber="147">unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="147" type="etymology">
|
||
<paragraph pageId="0" pageNumber="147">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="147">From Latin perrarus = very rare, in reference to the paucity of available material.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="147" type="distribution">
|
||
<paragraph pageId="0" pageNumber="147">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="147">Known only from Mount Pilatus (Switzerland, Obwalden).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="147" type="host">
|
||
<paragraph pageId="0" pageNumber="147">Host plant unknown.</paragraph>
|
||
<paragraph pageId="0" pageNumber="147">
|
||
As host plants of
|
||
<taxonomicName authorityName="Klimaszewski" authorityYear="1968" class="Insecta" family="Triozidae" genus="Eryngiofaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eryngiofaga" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">Eryngiofaga</emphasis>
|
||
</taxonomicName>
|
||
species are restricted to the genera
|
||
<taxonomicName class="Magnoliopsida" family="Apiaceae" genus="Bupleurum" kingdom="Plantae" lsidName="Bupleurum" order="Apiales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">Bupleurum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Magnoliopsida" family="Apiaceae" genus="Eryngium" kingdom="Plantae" lsidName="Eryngium" order="Apiales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">Eryngium</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName class="Magnoliopsida" family="Apiaceae" kingdom="Plantae" lsidName="" order="Apiales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="family">Apiaceae</taxonomicName>
|
||
) a likely host of
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. perrara" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="perrara">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">E. perrara</emphasis>
|
||
</taxonomicName>
|
||
is
|
||
<taxonomicName class="Magnoliopsida" family="Apiaceae" genus="Bupleurum" kingdom="Plantae" lsidName="Bupleurum ranunculoides" order="Apiales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="species" species="ranunculoides">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">Bupleurum ranunculoides</emphasis>
|
||
</taxonomicName>
|
||
L. which grows at the site
|
||
<normalizedToken originalValue="“Chilchsteine”">"Chilchsteine"</normalizedToken>
|
||
on Mount Pilatus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="147" type="comments">
|
||
<paragraph pageId="0" pageNumber="147">Comments.</paragraph>
|
||
<paragraph pageId="0" pageNumber="147">
|
||
The morphology of the parameres places
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. perrara" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="perrara">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">E. perrara</emphasis>
|
||
</taxonomicName>
|
||
in the
|
||
<taxonomicName baseAuthorityName="Loginova" baseAuthorityYear="1966" class="Insecta" family="Triozidae" genus="Eryngiofaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eryngiofaga congenita" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="congenita">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">Eryngiofaga congenita</emphasis>
|
||
</taxonomicName>
|
||
group. It differs from
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. armeniaca" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="armeniaca">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">E. armeniaca</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. maga" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="maga">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">E. maga</emphasis>
|
||
</taxonomicName>
|
||
in the basally more slender distal segment of the aedeagus, and from
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. congenita" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="congenita">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">E. congenita</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. refuga" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="refuga">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">E. refuga</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. hungarica" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="hungarica">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">E. hungarica</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. matura" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="matura">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">E. matura</emphasis>
|
||
</taxonomicName>
|
||
in the much shorter subapical ventral hook on the distal segment of the aedeagus. It differs also from the other species of the
|
||
<taxonomicName class="Magnoliopsida" family="Apiaceae" genus="Eryngium" kingdom="Plantae" lsidName="Eryngium congenita" order="Apiales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="species" species="congenita">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">Eryngium congenita</emphasis>
|
||
</taxonomicName>
|
||
group in details of the paramere. In
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. hungarica" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="hungarica">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">E. hungarica</emphasis>
|
||
</taxonomicName>
|
||
the anterior process of the paramere is broad (slender in all the other species); in
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. congenita" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="congenita">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">E. congenita</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. maga" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="maga">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">E. maga</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. matura" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="matura">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">E. matura</emphasis>
|
||
</taxonomicName>
|
||
the posterior lobe of the paramere, in lateral view, is narrowly triangular and distinctly longer than the anterior one (broadly triangular and only slightly longer in the other species); in
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. armeniaca" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="armeniaca">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">E. armeniaca</emphasis>
|
||
</taxonomicName>
|
||
the posterior lobe is big and the incision between anterior and posterior lobes is shallow not reaching the apical quarter of paramere (posterior lobe narrower and incision deep reaching basal third in
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. perrara" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="perrara">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">E. perrara</emphasis>
|
||
</taxonomicName>
|
||
); in
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. refuga" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="refuga">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">E. refuga</emphasis>
|
||
</taxonomicName>
|
||
the anterior process of the paramere is strongly curved along fore margin terminating in a posteriorly directed tooth (anterior process weakly curved anteriorly and terminating in each a forward and backward directed toothlet in
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. perrara" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="perrara">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">E. perrara</emphasis>
|
||
</taxonomicName>
|
||
). The head, thorax and abdomen of
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. congenita" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="congenita">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">E. congenita</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. refuga" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="refuga">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">E. refuga</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. hungarica" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="hungarica">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">E. hungarica</emphasis>
|
||
</taxonomicName>
|
||
are dark brown or almost black, but yellow, ochreous or, at most, light brown in
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. perrara" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="perrara">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">E. perrara</emphasis>
|
||
</taxonomicName>
|
||
. The new species also differs markedly in the distribution from its putatively closest relatives (based on paramere shape): Switzerland (
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. perrara" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="perrara">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">E. perrara</emphasis>
|
||
</taxonomicName>
|
||
) versus Caucasus, Kazakhstan, Siberia and Mongolia (
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. armeniaca" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="armeniaca">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">E. armeniaca</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. congenita" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="congenita">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">E. congenita</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. matura" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="matura">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">E. matura</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. refuga" order="Hemiptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="refuga">
|
||
<emphasis italics="true" pageId="0" pageNumber="147">E. refuga</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |