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<document id="4EC4625A1E82594F6D33C98F9F909738" ID-CLB-Dataset="300349" ID-DOI="10.3897/vz.74.e124752" ID-GBIF-Dataset="b637a981-80a3-4475-99c3-f213940e174d" ID-ZooBank="154CE236-EFA8-4411-834B-234A9B45A63F" ID-publisher-id="124752" URI-arpha="EA0088AE-2D74-569C-900B-1A82CA6D5784" URI-zoobank="http://zoobank.org/154CE236-EFA8-4411-834B-234A9B45A63F" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" article-type="research-article" checkinTime="1722264855187" checkinUser="pensoft" docAuthor="Boruah, Bitupan, Narayanan, Surya, Aravind, Neelavar Ananthram, Lalronunga, Samuel, Deepak, V. &amp; Das, Abhijit" docDate="2024" docId="739EA53FD4D75FBFB2DB62ECAC41A8D3" docLanguage="en" docName="Vertebrate Zoology 74: 453-486" docOrigin="Vertebrate Zoology 74" docSource="https://vertebrate-zoology.arphahub.com/article/124752/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Cyrtodactylus manipurensis Boruah, Narayanan, Deepak &amp; Das 2024, sp. nov." docType="treatment" docUuid="9AEE0E40-7983-444E-AC19-53AA8CE709D7" docUuidSource="ZooBank" docVersion="4" dtd-version="3.0" lastPageNumber="486" masterDocId="EA0088AE2D74569C900B1A82CA6D5784" masterDocTitle="Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India" masterLastPageNumber="486" masterPageNumber="453" pageNumber="453" updateTime="1722279389842" updateUser="ExternalLinkService">
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<mods:titleInfo id="90CE2FB877001F5AB9CF5AFC5D888C1E">
<mods:title id="838814733FF4F9D5996F00F60C9FA48A">Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India</mods:title>
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<mods:name id="E294B06F3D02973AE49D1DF998871D65" type="personal">
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<mods:namePart id="7BA8A66493041B5D29B67C5FBE795418">Boruah, Bitupan</mods:namePart>
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<mods:affiliation id="68BB575D30479CC9464A2AF1B4E7015F">Wildlife Institute of India, Chandrabani, Dehradun, Uttarakhand 248001, India</mods:affiliation>
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<mods:namePart id="D191892F9FD72D220B9299E6CD22B108">Narayanan, Surya</mods:namePart>
<mods:nameIdentifier id="4CCFAB72931EE5849727C4AC293297F3" type="ORCID">0000-0001-9359-2815</mods:nameIdentifier>
<mods:affiliation id="BCF2196F635F458686B2CFE5DAA1C7CD">SM Sehgal Foundation Center for Biodiversity and Conservation, Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Bangalore, Karnataka 560064, India</mods:affiliation>
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<mods:namePart id="110EFF26EEBEAFA2D672FC91EE530371">Aravind, Neelavar Ananthram</mods:namePart>
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<mods:affiliation id="538DCA5E4542D35DADDEFAB715A4F90D">SM Sehgal Foundation Center for Biodiversity and Conservation, Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Bangalore, Karnataka 560064, India &amp; Yenepoya Research Centre, Yenepoya University, Derlakatte, Mangalore, India</mods:affiliation>
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<mods:namePart id="FA6C8B7268E2D9F04B889F53BFC51386">Lalronunga, Samuel</mods:namePart>
<mods:nameIdentifier id="9EC5B5ADDF24C688780BD80BA088126B" type="ORCID">0000-0002-7542-4815</mods:nameIdentifier>
<mods:affiliation id="E3186FF29F9E3D397558EE815B72E4C6">Holy Child Society, Nalkata, Tripura 799263, India</mods:affiliation>
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<mods:namePart id="00968A16EA910BA191E09BC33DB7CD8C">Deepak, V.</mods:namePart>
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<mods:affiliation id="80F91B3E50F4B5B7C32CD42610D4221C">Department of Life Sciences, The Natural History Museum, London SW 7 5 BD, UK &amp; School of Natural and Environmental Sciences, Newcastle University, Newcastle upon Tyne NE 1 7 RU, UK</mods:affiliation>
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<mods:namePart id="83DF925A4B3C6CD8952560645AE27E5A">Das, Abhijit</mods:namePart>
<mods:nameIdentifier id="6EFC1446DF0E93FE4F712A46D5C856FB" type="ORCID">0000-0002-5851-8457</mods:nameIdentifier>
<mods:affiliation id="E448802819882BB0057BDB78497FB874">Wildlife Institute of India, Chandrabani, Dehradun, Uttarakhand 248001, India</mods:affiliation>
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<mods:title id="35EBB58EAD4134CA7BCBB48A18722998">Vertebrate Zoology</mods:title>
</mods:titleInfo>
<mods:part id="CBCFE1D7DD7C34CDCEDCE5E8A04B4461">
<mods:date id="62CA26472914F6394670B6F490B536E4">2024</mods:date>
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<mods:number id="141F783044EE9258952A795569ECB69C">2024-07-29</mods:number>
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<mods:number id="19CB911F406FE5B6D79DF0CAF90538A8">74</mods:number>
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<mods:identifier id="D32C6AFFE2590D4C73AE1304E9AA014A" type="DOI">10.3897/vz.74.e124752</mods:identifier>
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<treatment id="739EA53FD4D75FBFB2DB62ECAC41A8D3" ID-DOI="http://doi.org/10.5281/zenodo.13124029" ID-Zenodo-Dep="13124029" ID-arpha="739EA53F-D4D7-5FBF-B2DB-62ECAC41A8D3" ID-zoobank="https://zoobank.org/9AEE0E40-7983-444E-AC19-53AA8CE709D7" LSID="urn:lsid:zoobank.org:act:9AEE0E40-7983-444E-AC19-53AA8CE709D7" LSID-ZBK="urn:lsid:zoobank.org:act:9AEE0E40-7983-444E-AC19-53AA8CE709D7" httpUri="http://treatment.plazi.org/id/739EA53FD4D75FBFB2DB62ECAC41A8D3">
<subSubSection id="0620ACE781A8C5C6502441820332457C" type="nomenclature">
<paragraph id="9AD0A380D7E710BABE78DAA9F73C7314">
<taxonomicName id="A262AFE602D052D548130A495EF28B85" authority="Boruah, Narayanan, Deepak &amp; Das" authorityName="Boruah, Narayanan, Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="manipurensis" status="sp. nov.">
<emphasis id="5F6825074104D3D05E622FB1A649480D" italics="true">Cyrtodactylus manipurensis</emphasis>
Boruah, Narayanan, Deepak &amp; Das
</taxonomicName>
<taxonomicNameLabel id="254E1AFE97EDCE19ECA6EE7F51E2706E" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="D2161ED2478E1BB0BF02AFD40429C881" type="description">
<paragraph id="26DA8AA246533D811DAB7CC584707F37">
<figureCitation id="BD8A1F61B3ED25E46954B11637FDF38A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Holotype (WII-ADR 1596) of Cyrtodactylus manipurensis sp. nov. in preserved condition. A dorsal and B ventral view of the whole body; C ventral, D dorsal, and E lateral view of head; F dorsal enlarged tubercles on trunk; G ventral view of right manus and H of right pes. Scale bar: 10 mm." figureDoi="10.3897/vz.74.e124752.figure6" httpUri="https://binary.pensoft.net/fig/1102272">Figure 6</figureCitation>
;
<tableCitation id="BB9F8FA5D98B21FC30C1FD9304EE523D" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparative table of summary morphological characteristics among the members of the khasiensis group including the seven new species. NA- data not available; “ * ” data from the literature is not comparable with our data as the count method is different." httpUri="http://table.plazi.org/id/4AAE94DC4D30D8EB1A84D336CC270F8B" tableUuid="4AAE94DC4D30D8EB1A84D336CC270F8B">Tables 2, S 2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4JBK" type="Holotype">
<paragraph id="3B20C361D030E972EFF1AD88FD7767D1">
<heading id="FD983ACF905F03882FCCFEE8703D4B86" reason="title">
<typeStatus id="68FD44D14155E64934AA7EEFD57140A0" type="holotype">Holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="7C7FC2740E2792CE77E38D3F0305C7F6">
<materialsCitation id="71FDF20A7AA196EF57454C30D8C25E2E" collectingDate="2022-07-25" collectorName="Bitupan Boruah" country="India" elevation="1240" latitude="24.5954" location="Lamdan Kabui village" longLatPrecision="7" longitude="93.7085" municipality="Churachandpur District" specimenCode="WII-ADR 1596" specimenCount="1" specimenCount-male="1" stateProvince="Manipur" typeStatus="holotype">
<specimenCount id="5BE6018FE05E1CC889099339E615AA02" count="1" type="male">Adult male</specimenCount>
(
<specimenCode id="81A0426388BC24F2A8A071B0EC6CA921">WII-ADR 1596</specimenCode>
), collected near
<location id="8F00AAB846E65BB43BBD8234773AACF6" LSID="urn:lsid:plazi:treatment:739EA53FD4D75FBFB2DB62ECAC41A8D3:8F00AAB846E65BB43BBD8234773AACF6" country="India" latitude="24.5954" longLatPrecision="7" longitude="93.7085" municipality="Churachandpur District" name="Lamdan Kabui village" stateProvince="Manipur">Lamdan Kabui village</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EGKBK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[93.708500,24.595400]}">
<geoCoordinate id="FC129D6F8811F998299FAFA55751AAAF" degrees="24.5954" direction="north" orientation="latitude" precision="5" value="24.5954">24.5954 ° N</geoCoordinate>
;
<geoCoordinate id="135C4070C755FF39B31244D5A0A83E7F" degrees="93.7085" direction="east" orientation="longitude" precision="5" value="93.7085">93.7085 ° E</geoCoordinate>
</named-content>
;
<elevation id="F457AF4BC5E3225C609CAE2D90D14A26" metricMagnitude="3" metricUnit="m" metricValue="1.24" unit="m" value="1240.0">
elevation
<quantity id="5EEDD9284A49F4519CDE640A64782588" metricMagnitude="3" metricUnit="m" metricValue="1.24" unit="m" value="1240.0">1240 m</quantity>
a. s. l.
</elevation>
) (Fig.
<figureCitation id="E2D6F7413813B8D1C4E1C4454B925817" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Map showing geographic distribution of the clades within the khasiensis group. A type localities of the members of the four clades in the Indo-Burma region: 1 Cyrtodactylus septentrionalis, 2 C. guwahatiensis, 3 C. urbanus, 4 C. kazirangaensis, 5 C. siangensis sp. nov., 6 C. cayuensis, 7 C. namdaphaensis sp. nov., 8 C. bapme, 9 C. karsticola, 10 C. agarwali, 11 C. exercitus, 12 C. jaintiaensis, 13 C. khasiensis, 14 C. barailensis sp. nov., 15 C. nagalandensis, 16 C. kiphire sp. nov., 17 C. namtiram, 18 C. vairengtensis, 19 C. manipurensis sp. nov., 20 C. mombergi, 21 C. dianxiensis, 22 C. tripuraensis, 23 C. montanus, 24 C. aaronbaueri, 25 C. bengkhuaiai, 26 C. ngopensis, 27 C. lungleiensis, 28 C. ngengpuiensis sp. nov., 29 C. siahaensis, 30 C. gansi, 31 C. myaleiktaung, 32 C. brevidactylus, 33 C. aunglini, 34 C. chrysopylos, 35 C. ayeyarwadyensis; B cayuensis clade, green star (type locality of C. siangensis sp. nov.), pink star (type locality of C. cayuensis), pink triangle (C. “ arunachalensis ”), pink circle (C. cayuensis recorded in this study) (numbers correspond to the locality given in Table 1)." figureDoi="10.3897/vz.74.e124752.figure3" httpUri="https://binary.pensoft.net/fig/1102269">3 A</figureCitation>
),
<collectingMunicipality id="1CF99C873034C1D5EA7ED5DA665E0CD6">Churachandpur District</collectingMunicipality>
,
<collectingRegion id="DFFDA889672F15D6C6D1CBB4725D107E" country="India" name="Manipur">Manipur</collectingRegion>
,
<collectingCountry id="CAEC4F48F5B0B8EA15B9E2FE2E56797E" name="India">India</collectingCountry>
collected by
<collectorName id="FD5A199EA7E7D5A89B5F6F8BD100A6D7">Bitupan Boruah</collectorName>
on
<date id="20C7547AA0FDA64DADCCCD6E2DF26872" value="2022-07-25">
<collectingDate id="120B6E55E61EC70876242F49B817DD55" value="2022-07-25">25 July 2022</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<caption id="F16245DA920B26476EC6EEA9E97726F5" ID-DOI="10.3897/vz.74.e124752.figure6" ID-arpha="6008AF31-8C27-58D6-A030-7C0FDA0091C3" httpUri="https://binary.pensoft.net/fig/1102272" startId="F6">
<paragraph id="D808E1DEB1C90A739CDDB307A90F9EF2">
<label id="7525362A0A7DCABCD6DA88E762F1751B">Figure 6.</label>
</paragraph>
<paragraph id="207D35A611B5CB03FBCCAF9B9E250833">
Holotype (
<specimenCode id="9A4F742410F32F495C26CBD0B4C2F5EA">WII-ADR 1596</specimenCode>
) of
<taxonomicName id="8FEB927D81B4E1AE8772B450D6D5AA61" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="manipurensis" status="sp. nov.">
<emphasis id="CD2B94B8D0A25A362A3004FFAE821B27" italics="true">Cyrtodactylus manipurensis</emphasis>
</taxonomicName>
<emphasis id="291F3E7F758A04A143EF013E70034DBE" bold="true">
<taxonomicNameLabel id="07FC4F52168AA403C160DC265AF56220" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in preserved condition.
<emphasis id="B26A7BE1011DAD81B8242043CE20E981" bold="true">A</emphasis>
dorsal and
<emphasis id="20A07F716706B71D89C3E9098E0C1DA5" bold="true">B</emphasis>
ventral view of the whole body;
<emphasis id="94DB13A12AD8E161CDE10A682AF37D9F" bold="true">C</emphasis>
ventral,
<emphasis id="AB2A952086B3F04C4423B0C7C4DC2A91" bold="true">D</emphasis>
dorsal, and
<emphasis id="B0F6A2623A3C113E40409F7E07D8A9CE" bold="true">E</emphasis>
lateral view of head;
<emphasis id="6A8D129ABF12DDFEA063673D88C726BB" bold="true">F</emphasis>
dorsal enlarged tubercles on trunk;
<emphasis id="3A9CE39224AFF71BF911C8667B92054B" bold="true">G</emphasis>
ventral view of right manus and
<emphasis id="08043CFDAFEA7D2A0554FE392FBFF5B2" bold="true">H</emphasis>
of right pes. Scale bar: 10 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EZLBK" type="diagnosis">
<paragraph id="F9598474E104AA50B5DF13F9541C4A6A">
<heading id="ABA52E5444DE6B3B9D5B520164CC8052" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="9C85782A7E52A0A02DBE879347D4A0E6">
Medium-sized gecko,
<abbrev id="ABBRID0E6LBK" xlink_title="snout to vent length">SVL</abbrev>
at least
<quantity id="13A2A17A165DFF12D28FD943ADB36F4C" metricMagnitude="-2" metricUnit="m" metricValue="5.95" unit="mm" value="59.5">59.5 mm</quantity>
in adult male; 10 supralabials; eight or nine infralabials; 21 bluntly conical and feebly keeled tubercles across midbody; 37 paravertebral tubercles; 36 mid-ventral scales between ventrolateral folds; 11 or 12 subdigital lamellae beneath digit IV of manus; 1316 subdigital lamellae beneath digit IV of pes; seven precloacal pores arranged in a continuous series; six irregular shaped dark-brown cross bands on back between axilla and groin; tail dorsally with eight dark-brown and seven pale-brown bands arranged alternatively.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDMBK" type="description">
<paragraph id="BBF5D4BCEEEC3221FF515998BD98A122">
<heading id="E748ECAA9B232EF4C991092CF2C5E0EC" reason="title">
Description of the
<typeStatus id="6E492EB195EC776A6973633F66FC3CC6" type="holotype">holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="7805DF5F0F726AF71D35CD90E823F0BC">
Specimen well preserved except an incision below left axilla ventrolaterally. Snout-vent length
<quantity id="346813F6713831BC1E8F6B20C29202DD" metricMagnitude="-2" metricUnit="m" metricValue="5.95" unit="mm" value="59.5">59.5 mm</quantity>
. Head moderately large (
<abbrev id="ABBRID0EJMBK" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
/
<abbrev id="ABBRID0ENMBK" xlink_title="snout to vent length">SVL</abbrev>
= 0.26), oval, dorsoventrally depressed, longer than width (
<abbrev id="ABBRID0ERMBK" xlink_title="maximum head width">HW</abbrev>
/
<abbrev id="ABBRID0EVMBK" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
= 0.71), distinct from neck, broader at occipital region; snout tip rounded in both dorsal and lateral view; loreal region convex; canthus rostralis rounded, indistinct; interorbital space flat; a longitudinal furrow on dorsal surface of the snout; snout short (
<abbrev id="ABBRID0EZMBK" xlink_title="distance between the snout tip and the anterior bony orbital border">SO</abbrev>
/
<abbrev id="ABBRID0E4MBK" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
= 0.39), longer than orbit (
<abbrev id="ABBRID0EBNBK" xlink_title="orbit diameter, taken between the anterior and posterior bony orbital borders">OD</abbrev>
/
<abbrev id="ABBRID0EFNBK" xlink_title="distance between the snout tip and the anterior bony orbital border">SO</abbrev>
= 0.68); nostril nearly rounded, opening directed posterolaterally; ear opening rounded; scales on head heterogeneous, largest on snout and loreal region, posteriorly smaller in upper eyelid, interorbital space and occipital region, granular juxtaposed; scales on upper eyelids heterogeneous; supraciliaries outwardly sharp giving serrated appearance in dorsal view, size anteriorly and posteriorly decreases, largest at the anterodorsal region; rostral wide, a short groove at the middle on top, rostral connected with nasals, supranasals, an internasal and first supralabials, two scales between the supranasals paced longitudinally, granular scales at parietal, occipital and temporal region intermixed with slightly large rounded granular tubercles starting from the level of posterior margin of the upper eyelids, those on temporal region are slightly larger than that of occipital region, size of the tubercles increases towards nape; 10 supralabial scales on both sides, size decreases towards angle of jaw; 8 supralabials up to midorbit on both sides; mental nearly triangular, connected with first infralabials, inner postmentals; nine infralabials on right side and eight on left side, size decreases towards angle of jaw; first infralabials connected with mental, second infralabial, inner and outer postmentals; inner pair of postmentals are larger than the outer postmentals; two rows of slightly enlarged scales along the infralabials starting below the outer postmentals, posteriorly size of those decreases; rest of the gular scales are small, granular juxtaposed, nearly homogeneous, size increases towards the throat where they become imbricate.
</paragraph>
<paragraph id="30FFD2E2B8F4D3D2DF344688D63DB837">
Habitus slender (
<abbrev id="ABBRID0ELNBK" xlink_title="maximum body width">BW</abbrev>
/
<abbrev id="ABBRID0EPNBK" xlink_title="snout to vent length">SVL</abbrev>
= 0.16,
<abbrev id="ABBRID0ETNBK" xlink_title="trunk length, between the axilla and groin">TRL</abbrev>
/
<abbrev id="ABBRID0EXNBK" xlink_title="snout to vent length">SVL</abbrev>
= 0.49), dorsoventrally depressed, dorsal scales granular, rounded, heterogeneous, intermixed with rounded, weakly keeled and bluntly conical tubercles irregularly arranged, up to third segment of the tail, size increases towards posterior body and pronounced at the base of tail; 21 dorsal tubercles across mid dorsum; 37 paravertebral tubercles; ventrolateral fold weak; ventral scales larger than those of dorsal, flat, smooth, cycloid subimbricate to imbricate; 36 mid-ventral scales between ventrolateral fold; seven precloacal pores arranged in an inverted “
<collectionCode id="8C1D06DE5B66CCDEEDFA7078CCC7410A" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
” shaped continuous series, followed by a series of five unpored enlarged scales below it, largest at the apex, an unpored scale equal size to
<abbrev id="ABBRID0E2NBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
present in continuous with the
<abbrev id="ABBRID0E6NBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
in both ends; scales above the
<abbrev id="ABBRID0EDOBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
larger than those of belly scales.
</paragraph>
<paragraph id="05AC5AEBC4111AD726E81BD0C90F7DDB">
Forelimbs and hindlimbs slender (
<abbrev id="ABBRID0EJOBK" xlink_title="forearm length, taken from the elbow to the wrist">FL</abbrev>
/
<abbrev id="ABBRID0ENOBK" xlink_title="snout to vent length">SVL</abbrev>
= 0.14,
<abbrev id="ABBRID0EROBK" xlink_title="crus length, taken from the knee to the base of the foot">CL</abbrev>
/
<abbrev id="ABBRID0EVOBK" xlink_title="snout to vent length">SVL</abbrev>
= 0.17), digits strongly inflected at the joints, all bearing large recurved claw, enlarged subdigital lamellae; lamellae beneath digit IV of right and left manus (given as basal + distal) is 6 + 5 and 6 + 6 respectively; lamellae beneath digit IV of both right and left pes (given as basal + distal) is 6 + 7 and 7 + 9 respectively; dorsal scales on forelimbs heterogeneous is size, mostly granular; proximal scales on upper arm are smaller than that of lower arm; scales near elbow on lower arm smooth and subimbricate; upper arm scales granular at proximal part and at the distal end it is smooth, cycloid and imbricate; dorsal scales of hindlimbs heterogeneous, intermixed with slightly enlarged, feebly keeled and bluntly conical tubercles; scales on inner lateral side of the thighs are smooth, large and subimbricate, those on dorsal side are small granular; scales on tibia are small, granular juxtaposed; ventral scales of forelimbs granular, juxtaposed, heterogeneous; scales on palm heterogeneous in shape and size, granular juxtaposed; ventral scales on hindlimbs heterogeneous; most of the thigh scales are smooth, cycloid and subimbricate, but on the knee and below the level of precloacal pores, scales are smaller and granular; scales above the vent granular; tibia scales smooth, nearly homogeneous, cycloid and subimbricate; scales on soles heterogeneous, granular, juxtaposed to subimbricate.
</paragraph>
<paragraph id="ABAF3048690B3A2467BE741056B7DC7D">
Tail complete (
<abbrev id="ABBRID0E2OBK" xlink_title="tail length, from the cloaca to the tail tip">TL</abbrev>
=
<quantity id="8C62E474DCF3E2E573890F7D39D5431B" metricMagnitude="-2" metricUnit="m" metricValue="6.7" unit="mm" value="67.0">67 mm</quantity>
), slender, gradually tapering towards tip, segments indistinct, dorsal scales small, granular, juxtaposed at the base, posteriorly size increases, flat, smooth, subimbricate, heterogeneous in shape and size, large feebly keeled scales up to third segment of the tail, those on basal segment are pronounced; subcaudal scales smooth, subimbricate, wider than that of dorsal, heterogeneous in shape and size; no enlarged plate like series of subcaudal scales; three bluntly conical spurs on both sides of the tail base.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6OBK" type="Colouration in preservative">
<paragraph id="85DE975F549A46DA3873F69FF1F400A7">
<heading id="DF8D82EED3EFA575FB1656D564E8E475" reason="title">Colouration in preservative.</heading>
</paragraph>
<paragraph id="034F666B53845C87330C3623536B8E92">Head on top and laterally brown; upper eyelids grey; dorsal ground colour of neck and back slightly paler than that of head; dark-brown irregular patches on neck up to fore limb insertion level; six irregular shaped dark-brown cross bands on back between axilla and groin, mid-dorsally interrupted; one dark-brown band on sacrum; tail dorsally with eight dark-brown and seven pale-brown bands arranged alternatively; dark bands are comparatively broader than the pale bands; the first dark-brown band on tail base broken mid-dorsally into two enlarged elongated spots. Ventrally head, neck, trunk and limbs light-brown; tail with irregular dark-brown specks.</paragraph>
</subSubSection>
<subSubSection id="SECID0EEPBK" type="comparisons">
<paragraph id="BFF96A897D806F58673E87C3E2CADD37">
<heading id="23B5D12D1F725938AD8863BFE7EEE5AF" reason="title">Comparison.</heading>
</paragraph>
<paragraph id="08A32520CFD2707E32F1986E5A7C653B">
<taxonomicName id="F2C8697266240254000C0F76CF95A23D" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="manipurensis" status="sp. nov.">
<emphasis id="FAAE0E34454F8289D46AE0E4D9AEADA9" italics="true">Cyrtodactylus manipurensis</emphasis>
</taxonomicName>
<emphasis id="9F8D96EAC9F5D6E6C56E47FC59059042" bold="true">
<taxonomicNameLabel id="6A431120FBE5C4BEA78660482502D655" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="D21A32565BBA539E7D61EC8ABDE728D5" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="aaronbaueri">
<emphasis id="A4956D154ADE6751699FD8607E485BE6" italics="true">C. aaronbaueri</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0ECQBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
21 (vs.
<abbrev id="ABBRID0EGQBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="C8BD8C3A8533C2393BD50C7FD47F9A5F">DTR</collectionCode>
</abbrev>
2228); differs from
<taxonomicName id="BC404615932E38FE5E76DDD3BD6F1436" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="aunglini">
<emphasis id="649A3634CE861C7DB67F44D3BF41B02B" italics="true">C. aunglini</emphasis>
</taxonomicName>
by fewer mid-ventral scale rows,
<abbrev id="ABBRID0EVQBK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="73CD4B4C06EFA043009EA6CFE3991F0B">MVSR</collectionCode>
</abbrev>
36 (vs.
<abbrev id="ABBRID0EZQBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="149FB05B122FF4FAEA0D7650AC160502">DTR</collectionCode>
</abbrev>
4749), by fewer precloacal pores,
<abbrev id="ABBRID0E4QBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
7 (vs.
<abbrev id="ABBRID0EBRBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
12 or 13); differs from
<taxonomicName id="27B5F42F5D270AAB251A8ADFB4C5047A" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="barailensis" status="sp. nov.">
<emphasis id="A47058AF00962AEF68E4F8CBC82D0854" italics="true">C. barailensis</emphasis>
</taxonomicName>
<emphasis id="DE449E1CD91A90F3DB56E83D3BEAC7C8" bold="true">
<taxonomicNameLabel id="B2348BDB2D072179BABCE6E55EB008FF" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by number of dorsal tubercle rows,
<abbrev id="ABBRID0ESRBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
21 (vs.
<abbrev id="ABBRID0EWRBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="26CFFF408CA30237CDB7B2F5FC116A68">DTR</collectionCode>
</abbrev>
17), number of paravertebral tubercle rows,
<abbrev id="ABBRID0E1RBK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">PVT</abbrev>
37 (vs.
<abbrev id="ABBRID0E5RBK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">
<collectionCode id="5EC03ABB24555729C29EFE0CF40CF9D2">PVT</collectionCode>
</abbrev>
32); differs from
<taxonomicName id="24F89A370325CC81C6BB9F1E5AB8CD98" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="bengkhuaiai">
<emphasis id="2C11DC776872C5982B45EF267D9251F6" italics="true">C. bengkhuaiai</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0ENSBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
21 (vs.
<abbrev id="ABBRID0ERSBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="B49923EE72542A772EFC25BF4A2493F5">DTR</collectionCode>
</abbrev>
2226); differs from
<taxonomicName id="3154D65BD6A875B1FDD3E08580D69309" authorityName="Bauer" authorityYear="2002" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="brevidactylus">
<emphasis id="24B3BF80F3DE1D0FEC8846873975F9BC" italics="true">C. brevidactylus</emphasis>
</taxonomicName>
by having fewer dorsal tubercle rows,
<abbrev id="ABBRID0EATBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
21 (vs.
<abbrev id="ABBRID0EETBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="9C4A4F6AEB43725F476340579123C3A7">DTR</collectionCode>
</abbrev>
2730), enlarged chocolate-brown patches on head and back absent (vs. present); differs from
<taxonomicName id="2AC5C3524ACBE6CCB5E4AF5A87DAE35A" authorityName="Bauer" authorityYear="2003" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="chrysopylos">
<emphasis id="C52360A5CA627FCA244064F44B58BCAD" italics="true">C. chrysopylos</emphasis>
</taxonomicName>
by much smaller body size,
<abbrev id="ABBRID0ETTBK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="E743E589A53ABB4EB0752BACCAE47C3C" metricMagnitude="-2" metricUnit="m" metricValue="5.95" unit="mm" value="59.5">59.5 mm</quantity>
(vs.
<abbrev id="ABBRID0EXTBK" xlink_title="snout to vent length">
<collectionCode id="D6622687119C1A338A6F27F9B2C6AC43">SVL</collectionCode>
</abbrev>
<quantity id="342C0ECD33F49391AA69E28BA8FB4BE9" metricMagnitude="-2" metricUnit="m" metricValue="7.2" metricValueMax="7.909999999999999" metricValueMin="6.49" unit="mm" value="72.0" valueMax="79.1" valueMin="64.9">64.979.1 mm</quantity>
in male), fewer precloacal pores,
<abbrev id="ABBRID0E2TBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
7 (vs
<abbrev id="ABBRID0E6TBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
813); differs from
<taxonomicName id="18C5317C08FB0DE488DAF2AD7722C212" authorityName="Liu &amp; Rao" authorityYear="2021" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="dianxiensis">
<emphasis id="9793AE215740729FC5DCAB227BF3FCF9" italics="true">C. dianxiensis</emphasis>
</taxonomicName>
by smaller body size,
<abbrev id="ABBRID0EOUBK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="79EC13178443A19605C780E536412309" metricMagnitude="-2" metricUnit="m" metricValue="5.95" unit="mm" value="59.5">59.5 mm</quantity>
(vs.
<abbrev id="ABBRID0ESUBK" xlink_title="snout to vent length">
<collectionCode id="780AB5955949C8E03DF8033C5A2CD97A">SVL</collectionCode>
</abbrev>
<quantity id="B94A7FE1669E5BDECF03502894DA6366" metricMagnitude="-2" metricUnit="m" metricValue="7.685" metricValueMax="7.99" metricValueMin="7.38" unit="mm" value="76.85" valueMax="79.9" valueMin="73.8">73.879.9 mm</quantity>
), 21 rows of dorsal tubercle rows (vs.
<abbrev id="ABBRID0EWUBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="6B8802CEE14E236A88A9589EFAA65313">DTR</collectionCode>
</abbrev>
18 or 19); differs from
<taxonomicName id="F92AE2BDA49D0E05D1350AB272BED886" authorityName="Bauer" authorityYear="2003" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="gansi">
<emphasis id="445860ABF17DF766B5A41DA44D456CCF" italics="true">C. gansi</emphasis>
</taxonomicName>
by fewer precloacal pores,
<abbrev id="ABBRID0EFVBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
7 (vs.
<abbrev id="ABBRID0EJVBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
1629), ventrolateral fold present on trunk (vs. absent); differs from
<taxonomicName id="954E22EF8E3D538C4EE78440EC7B76A4" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="jaintiaensis">
<emphasis id="8FEB6E60965B5C171910AA8B3C0F1985" italics="true">C. jaintiaensis</emphasis>
</taxonomicName>
by much smaller body size,
<abbrev id="ABBRID0EYVBK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="FC5101E69C6AD649DCBD3D27FAA4A436" metricMagnitude="-2" metricUnit="m" metricValue="5.95" unit="mm" value="59.5">59.5 mm</quantity>
(vs.
<abbrev id="ABBRID0E3VBK" xlink_title="snout to vent length">
<collectionCode id="BA5FF1B68466A96765F81818789D0BBE">SVL</collectionCode>
</abbrev>
87.0
<quantity id="288EF0CD8B9433AF692717A943233F3D" metricMagnitude="-2" metricUnit="m" metricValue="8.83" unit="mm" value="88.3">88.3 mm</quantity>
in male), fewer precloacal pores in male,
<abbrev id="ABBRID0EAWBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
7 (vs.
<abbrev id="ABBRID0EEWBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
11 or 12); differs from
<taxonomicName id="326C2C53733A585F8E1BE009097240DB" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="kiphire" status="sp. nov.">
<emphasis id="20679E38A689B23EB4BBFC20830FE339" italics="true">C. kiphire</emphasis>
</taxonomicName>
<emphasis id="4A496A8F8AA822EA6444F8C6ADC8CC35" bold="true">
<taxonomicNameLabel id="5C46630D0A80FC02E9B4D7D90DBCA073" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by smaller body size,
<abbrev id="ABBRID0EVWBK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="179348035F6C676C48C904A2A067700C" metricMagnitude="-2" metricUnit="m" metricValue="5.95" unit="mm" value="59.5">59.5 mm</quantity>
(
<abbrev id="ABBRID0EZWBK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="B67EFF2C9AA3FF7850289E4DAA9EA09F" metricMagnitude="-2" metricUnit="m" metricValue="6.43" metricValueMax="6.47" metricValueMin="6.39" unit="mm" value="64.3" valueMax="64.7" valueMin="63.9">63.964.7 mm</quantity>
), number of dorsal tubercle rows,
<abbrev id="ABBRID0E4WBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
21 (vs.
<abbrev id="ABBRID0EBXBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="BEC4CD02DEF3BE237EE966F9FD08C0A4">DTR</collectionCode>
</abbrev>
16), number of paravertebral tubercle rows,
<abbrev id="ABBRID0EFXBK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">PVT</abbrev>
37 (vs.
<abbrev id="ABBRID0EJXBK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">
<collectionCode id="6A688FBFD62BB82C8F4F5E85110395A4">PVT</collectionCode>
</abbrev>
26 or 29); differs from
<taxonomicName id="239C2FCBD42D744120D8CA20DDDEC6B4" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="lungleiensis">
<emphasis id="A1CE98F6EE16ECCEAFC75C32A0969F7D" italics="true">C. lungleiensis</emphasis>
</taxonomicName>
by higher number of precloacal pores,
<abbrev id="ABBRID0EYXBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
7 (vs.
<abbrev id="ABBRID0E3XBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
<quantity id="B0F27D983C18812485F5B65A61B76CE6" metricMagnitude="-1" metricUnit="m" metricValue="1.016" metricValueMax="1.27" metricValueMin="0.762" unit="in" value="4.0" valueMax="5.0" valueMin="3.0">35 in</quantity>
male), fewer dorsal tubercle rows,
<abbrev id="ABBRID0EAYBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
21 (vs. 2428); differs from
<taxonomicName id="942C21A573571D22879667502953DC50" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="montanus">
<emphasis id="0417EBB504BD1F9860D5DB9B1D0C6A6A" italics="true">C. montanus</emphasis>
</taxonomicName>
by larger body size,
<abbrev id="ABBRID0EPYBK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="1A803DA34AE1607445548588E5A288DC" metricMagnitude="-2" metricUnit="m" metricValue="5.95" unit="mm" value="59.5">59.5 mm</quantity>
(vs. 53.655.0 mm in male); differs from
<taxonomicName id="8C4476CEF25977CF4AF9734C3C56C052" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="myaleiktaung">
<emphasis id="0626093DBCA48BBF875CE2023F0B1BC5" italics="true">C. myaleiktaung</emphasis>
</taxonomicName>
by fewer mid-ventral scale rows,
<abbrev id="ABBRID0E5YBK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="64317A82276EA08210050DADB603C5F8">MVSR</collectionCode>
</abbrev>
36 (vs.
<abbrev id="ABBRID0ECZBK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="07A2C3AB1BCE576B1335ED5D98C2CFD5">MVSR</collectionCode>
</abbrev>
57), precloacal pores present (vs.
<abbrev id="ABBRID0EGZBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
absent), broad regular dark bands absent on dorsum (vs. present); differs from
<taxonomicName id="1587C6320E6F9D7205868D8A9B176B0F" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="nagalandensis">
<emphasis id="FE7B3FF8D2ABA32823D88A776217E646" italics="true">C. nagalandensis</emphasis>
</taxonomicName>
by higher number of dorsal tubercle rows,
<abbrev id="ABBRID0EVZBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
21 (vs.
<abbrev id="ABBRID0EZZBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="DBF5EE27CCAD33CEE116B2D83B3F03D1">DTR</collectionCode>
</abbrev>
1618); differs from
<taxonomicName id="DD92636A43960E6F56645EB61E3570D1" authorityName="Mahony &amp; Kamei" authorityYear="2021" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="namtiram">
<emphasis id="8105F1FAAE33942DFB5848D2EE578168" italics="true">C. namtiram</emphasis>
</taxonomicName>
by having fewer precloacal pores,
<abbrev id="ABBRID0EI1BK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
7 (vs.
<abbrev id="ABBRID0EM1BK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
12); differs from
<taxonomicName id="784386F859640E9085523CBA733884E2" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="ngopensis">
<emphasis id="8BF7E6901C7535C4ED37B63A305103E1" italics="true">C. ngopensis</emphasis>
</taxonomicName>
by the number of precloacal pores,
<abbrev id="ABBRID0E21BK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
7 (vs.
<abbrev id="ABBRID0E61BK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
6); differs from
<taxonomicName id="75D0F1876C856C6B0078E7123B23230B" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="siahaensis">
<emphasis id="B623C3E85FADB9FD306808E36F6E93A5" italics="true">C. siahaensis</emphasis>
</taxonomicName>
by be fewer subdigital lamellae, 11 or 12 lamellae under fourth finger (vs. 13 or 14 lamellae under fourth finger), dark-brown broad bands consists of two series of enlarged spots on dorsum (vs. dark-brown blotches in the form of reticulation on dorsum); differs from
<taxonomicName id="5823E40797750B7F8CBEEB74A25DC822" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="vairengtensis">
<emphasis id="C092F38654F54858C9E4CB8AB77E80F8" italics="true">C. vairengtensis</emphasis>
</taxonomicName>
by fewer precloacal pores,
<abbrev id="ABBRID0EZ2BK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
7 (vs.
<abbrev id="ABBRID0E42BK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
911). Morphological differences with other members of
<taxonomicName id="12DB307EA7EA1DC6E7E220E0031DC732" baseAuthorityName="Jerdon" baseAuthorityYear="1870" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="khasiensis">
<emphasis id="C259E8D5EF271F9332EA7E67EBA34B30" italics="true">khasiensis</emphasis>
</taxonomicName>
group is presented in Table
<tableCitation id="1FE329B200BE8F8E2208046872CC71D7" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparative table of summary morphological characteristics among the members of the khasiensis group including the seven new species. NA- data not available; “ * ” data from the literature is not comparable with our data as the count method is different." httpUri="http://table.plazi.org/id/4AAE94DC4D30D8EB1A84D336CC270F8B" tableUuid="4AAE94DC4D30D8EB1A84D336CC270F8B">2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EP3BK" type="Sequence divergence">
<paragraph id="E11F7423E607E8B3E25EF97290D90A14">
<heading id="CE6B432ACA643337FB4AB0E7AF68D504" reason="title">Sequence divergence.</heading>
</paragraph>
<paragraph id="8E198900797B5288C374D675D9339C43">
<taxonomicName id="C92DFBFCB13F8CDDD1D41EDCA4242DDE" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="manipurensis" status="sp. nov.">
<emphasis id="A96FD26C492C02A422321EA0A4E0CB18" italics="true">Cyrtodactylus manipurensis</emphasis>
</taxonomicName>
<emphasis id="A410B67C0B1420DDC5DCDE889ED63C5C" bold="true">
<taxonomicNameLabel id="1F31FC271704EA45EEF504D74B5905D1" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a high genetic divergence of 11.211.3 % from its closely related
<taxonomicName id="014FC2C4D1AE0D7D13EBA48090D00858" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="ngopensis">
<emphasis id="817D0220941C0471869BAE3FA598E251" italics="true">C. ngopensis</emphasis>
</taxonomicName>
, 1010.3 % from
<taxonomicName id="C93CD769FAE042ADD49C62180ED9A53D" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="aaronbaueri">
<emphasis id="B84B98217FF80B4715C85AD0F1DC625C" italics="true">C. aaronbaueri</emphasis>
</taxonomicName>
, 10.210.5 % from
<taxonomicName id="B16371B8E905CFDAF6891C10C8B9DDFD" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="vairengtensis">
<emphasis id="189B270AC549CC7D703AD3BE17385383" italics="true">C. vairengtensis</emphasis>
</taxonomicName>
and 9.711.1 % from the
<taxonomicName id="0115F927457EE26D5B6BE03FA16AE5A6" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="montanus">
<emphasis id="EE94C4B9A5C45C87EBD78DB462D44195" italics="true">C. montanus</emphasis>
</taxonomicName>
. With other members of the clade,
<taxonomicName id="394A09A3E2E92E46EE95EB84996FA459" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="kiphire" status="sp. nov.">
<emphasis id="26C85D73B2D18FFCDD3830A05C170BDF" italics="true">C. kiphire</emphasis>
</taxonomicName>
<emphasis id="4E4CD84F441E1E9804404055F139113B" bold="true">
<taxonomicNameLabel id="A401388F9AFAF8D6D424E1EEA0130D14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a genetic divergence of 8.6 % and 23 % in the ND 2 gene.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E25BK" type="etymology">
<paragraph id="8EB34D7DE05395680F819129CCFF3559">
<heading id="47DC0361EC668A9B95E3D341D24FB14C" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="9F9D2A627E60E5F35A20C0D598482CE4">
This species is named after
<collectingRegion id="7196F82B0F88285AA3D4B49E13632965" country="India" name="Manipur">Manipur state</collectingRegion>
in
<collectingCountry id="B7D9FD37A9D115EECEAD5244B71CFAA7" name="India">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EA6BK" type="Suggested common name">
<paragraph id="D2D035E27C25B69A3B28C7EF91F8973A">
<heading id="46D344BE69F1B1C03A096AC3D848F597" reason="title">Suggested common name.</heading>
</paragraph>
<paragraph id="9F61F7917C160037EACD18ABE1A2FC52">
<collectingRegion id="DBEB75D324CB5A0DF5E6319CB3C67A07" country="India" name="Manipur">Manipur</collectingRegion>
bent-toed gecko.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EF6BK" type="distribution">
<paragraph id="95A0D7DC0F06A37507DFA1EB28737E9B">
<heading id="C0321E2E9C5BC2E7C462089782050778" reason="title">Distribution and natural history.</heading>
</paragraph>
<paragraph id="C7E4C8B5541D29B294BF28958ABBB55A">
<taxonomicName id="F6931D89F9FAF8C5FBB519E2F19D66DD" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="manipurensis" status="sp. nov.">
<emphasis id="789F958F4A278FDB9C2FD4A1B5F1A719" italics="true">Cyrtodactylus manipurensis</emphasis>
</taxonomicName>
<emphasis id="BC311887EA894D357CB38CD93FB57B61" bold="true">
<taxonomicNameLabel id="4659FDB93C760BFACEBA6D278EC578EE" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is currently only recorded from the type locality. We collected a single individual at 18: 00 hrs on
<date id="2A81CCF026A391872CDC8CC0E4C271F2" value="2022-07-25">25 July 2022</date>
near Lamdan Kabui village. It was perched on a shrub at a height of approximately
<quantity id="A35F8395BB9B318B8A37F1FB7E2D4EE7" metricMagnitude="0" metricUnit="m" metricValue="1.5" unit="m" value="1.5">1.5 m</quantity>
, on the road connecting Leimatak and Charoikhullen. The habitat is secondary forest, with
<taxonomicName id="CCAEC143F246A4BBEEF1FD3A43B59A26" class="Liliopsida" family="Zingiberaceae" genus="Zingiber" kingdom="Plantae" order="Zingiberales" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="A8D395E30503CF911D2663EED735AED6" italics="true">Zingiber</emphasis>
sp.
</taxonomicName>
cultivation and settlements.
</paragraph>
</subSubSection>
</treatment>
</document>