164 lines
19 KiB
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164 lines
19 KiB
XML
<document id="0F108A28E9600D25F1E3DC269286E498" ID-DOI="10.11646/zootaxa.5025.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5258835" ID-ZooBank="35E6C8FA-4078-4C53-9B74-F9618D5E6E60" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629835009263" checkinUser="plazi" docAuthor="Harzhauser, Mathias & Landau, Bernard M." docDate="2021" docId="256F7D50E263FFE8ACD63197FE89B013" docLanguage="en" docName="zootaxa.5025.1.1.pdf" docOrigin="Zootaxa 5025 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5025.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Graphicomassa Iredale 1929" docType="treatment" docVersion="5" lastPageNumber="9" masterDocId="D9560528E26BFFE0AC413244FFB9B617" masterDocTitle="The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea — striking diversity of a negelected group" masterLastPageNumber="75" masterPageNumber="1" pageNumber="9" updateTime="1699118969203" updateUser="plazi">
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<mods:titleInfo id="BC2C148ECD44A0B24E7C65B70A54438D">
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<mods:title id="AF683BB9AFB7E09B69B662B4BFBE1C49">The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea — striking diversity of a negelected group</mods:title>
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<mods:name id="9A26E1A2DD36E3BEDC5D5BA5674CFC18" type="personal">
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<mods:namePart id="C32E0C0106BBBFC262060881CE202FDA">Harzhauser, Mathias</mods:namePart>
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<mods:nameIdentifier id="2FD73263A96297040A501F3D5F9BFA84" type="email">mathias.harzhauser@nhm-wien.ac.at</mods:nameIdentifier>
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<mods:namePart id="076D5EA3729CCCA02A37E00542E583FC">Landau, Bernard M.</mods:namePart>
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<mods:nameIdentifier id="9D03DADD9992D05BAAF13C24B5CC2E81" type="ORCID">0000-0002-4471-6655</mods:nameIdentifier>
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<mods:nameIdentifier id="F51077958F66F54E304D32C9A8A3FFCC" type="email">mathias.harzhauser@nhm-wien.ac.at</mods:nameIdentifier>
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<mods:typeOfResource id="42317F4314EFDD71CD2D97A950BC6517">text</mods:typeOfResource>
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<mods:title id="C50B78973641083E152CC37F45104665">Zootaxa</mods:title>
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<mods:part id="493E3E74C8C0DEC4F2AACBF8C63B6474">
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<mods:date id="1C2F8DCD1B2B35AFD1DB651CAFAD014D">2021</mods:date>
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<mods:number id="88113D8F806C1BE5C21BDFA6A5B3323B">2021-08-23</mods:number>
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<mods:number id="372EC80F6880FC1CE4330F474A3990CA">5025</mods:number>
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<mods:detail id="76C68AEE3A96064092E7FAE17BFF8C7D" type="issue">
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<mods:number id="5222C29C584688623D614C2EAD81C19A">1</mods:number>
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<mods:start id="09E7B0E5B288544B69D7DBE26DB61003">1</mods:start>
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<mods:end id="DB7D27C9061B3544CBD73A902FA81D6D">75</mods:end>
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<mods:url id="9D8226A9CCC5286986955D8AEBD98912">http://dx.doi.org/10.11646/zootaxa.5025.1.1</mods:url>
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<mods:classification id="5D1DB859B6097935FC8771BE62152456">journal article</mods:classification>
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<mods:identifier id="9433A639F22E6E94C34E47C3620E0AD8" type="DOI">10.11646/zootaxa.5025.1.1</mods:identifier>
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<treatment id="256F7D50E263FFE8ACD63197FE89B013" LSID="urn:lsid:plazi:treatment:256F7D50E263FFE8ACD63197FE89B013" httpUri="http://treatment.plazi.org/id/256F7D50E263FFE8ACD63197FE89B013" lastPageNumber="9" pageId="8" pageNumber="9">
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<subSubSection id="E5DC9FCDE263FFE8ACD63197FDEDB5FB" box="[151,596,978,1005]" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph id="AD79CC46E263FFE8ACD63197FDEDB5FB" blockId="8.[151,596,978,1005]" box="[151,596,978,1005]" pageId="8" pageNumber="9">
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<heading id="F6317B2AE263FFE8ACD63197FDEDB5FB" bold="true" box="[151,596,978,1005]" fontSize="11" level="2" pageId="8" pageNumber="9" reason="4">
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<emphasis id="9FB21054E263FFE8ACD63197FDEDB5FB" bold="true" box="[151,596,978,1005]" pageId="8" pageNumber="9">
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<emphasis id="9FB21054E263FFE8ACD63197FF5CB5FA" bold="true" box="[151,229,979,1005]" italics="true" pageId="8" pageNumber="9">Genus</emphasis>
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<taxonomicName id="6AC6B7C5E263FFE8ACAC3196FDEDB5FB" ID-CoL="7P4KD" authority="Iredale, 1929" authorityName="Iredale" authorityYear="1929" box="[237,596,978,1005]" class="Gastropoda" family="Columbellidae" genus="Graphicomassa" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
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<emphasis id="9FB21054E263FFE8ACAC3196FE13B5FA" bold="true" box="[237,426,978,1005]" italics="true" pageId="8" pageNumber="9">Graphicomassa</emphasis>
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<bibRefCitation id="C957B1B7E263FFE8ADF03197FDEDB5FB" author="Iredale, T." box="[433,596,978,1005]" pageId="8" pageNumber="9" pagination="261 - 297" refId="ref59638" refString="Iredale, T. (1929) Queensland molluscan notes. No. 1. Memoirs of the Queensland Museum, 9, 261 - 297. [https: // www. biodiversitylibrary. org / page / 52430224 # page / 61 / mode / 1 up]" type="journal article" year="1929">Iredale, 1929</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="E5DC9FCDE263FFE8ACD63653FA9BB238" box="[151,1314,1047,1071]" pageId="8" pageNumber="9" type="type_taxon">
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<paragraph id="AD79CC46E263FFE8ACD63653FA9BB238" blockId="8.[151,1314,1047,1071]" box="[151,1314,1047,1071]" pageId="8" pageNumber="9">
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<emphasis id="9FB21054E263FFE8ACD63653FE9AB238" bold="true" box="[151,291,1047,1071]" pageId="8" pageNumber="9">
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<typeStatus id="727D72E4E263FFE8ACD63653FF75B238" box="[151,204,1047,1071]" pageId="8" pageNumber="9">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="6AC6B7C5E263FFE8AD6B3653FDCBB238" ID-CoL="XBV6" authority="Duclos, 1840" authorityName="Duclos" authorityYear="1840" box="[298,626,1047,1071]" class="Gastropoda" family="Columbellidae" genus="Columbella" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="ligula">
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<emphasis id="9FB21054E263FFE8AD6B3653FE58B238" box="[298,481,1047,1071]" italics="true" pageId="8" pageNumber="9">Columbella ligula</emphasis>
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<bibRefCitation id="C957B1B7E263FFE8ADA93653FDCBB238" author="Duclos P. L." box="[488,626,1047,1071]" pageId="8" pageNumber="9" refId="ref56480" refString="Duclos P. L. (1840) Histoire naturelle generale et particuliere de tous les genres de coquilles univalves marines a l'etat vivant et fossile, publiee par monographie. Genre Colombelle. Didot, Paris, 13 pls. [http: // biodiversitylibrary. org / page / 45200570]" type="url" year="1840">Duclos, 1840</bibRefCitation>
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</taxonomicName>
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; original designation by
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<bibRefCitation id="C957B1B7E263FFE8AF2D3653FB8AB238" author="Iredale, T." box="[876,1075,1047,1071]" pageId="8" pageNumber="9" pagination="261 - 297" refId="ref59638" refString="Iredale, T. (1929) Queensland molluscan notes. No. 1. Memoirs of the Queensland Museum, 9, 261 - 297. [https: // www. biodiversitylibrary. org / page / 52430224 # page / 61 / mode / 1 up]" type="journal article" year="1929">Iredale (1929: 289)</bibRefCitation>
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. Present-day,
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<collectingCountry id="D5D18CD6E263FFE8A8FF3653FAA7B238" box="[1214,1310,1047,1071]" name="Australia" pageId="8" pageNumber="9">Australia</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="E5DC9FCDE263FFE8ACD63619FE89B013" pageId="8" pageNumber="9" type="discussion">
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<paragraph id="AD79CC46E263FFE8ACD63619FE89B013" blockId="8.[151,1437,1117,1540]" pageId="8" pageNumber="9">
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<emphasis id="9FB21054E263FFE8ACD63619FEA5B260" bold="true" box="[151,284,1117,1143]" pageId="8" pageNumber="9">Discussion.</emphasis>
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We place
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<taxonomicName id="6AC6B7C5E263FFE8ADCA361AFCCBB260" authority="Pusch, 1837" authorityName="Pusch" authorityYear="1837" box="[395,882,1117,1143]" class="Gastropoda" family="Muricidae" genus="Nassa" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="subSpecies" species="columbelloides" subSpecies="polonica">
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<emphasis id="9FB21054E263FFE8ADCA361AFD5CB260" box="[395,741,1117,1143]" italics="true" pageId="8" pageNumber="9">Nassa columbelloides polonica</emphasis>
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Pusch, 1837
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</taxonomicName>
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and
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<taxonomicName id="6AC6B7C5E263FFE8AFE4361AFAA1B26F" authority="Baluk, 1995" authorityName="Baluk" authorityYear="1995" box="[933,1304,1117,1144]" class="Gastropoda" family="Columbellidae" genus="Pyrene" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="korytnicensis">
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<emphasis id="9FB21054E263FFE8AFE4361AFB35B260" box="[933,1164,1117,1143]" italics="true" pageId="8" pageNumber="9">Pyrene korytnicensis</emphasis>
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<bibRefCitation id="C957B1B7E263FFE8A8D03619FAA1B26F" author="Baluk, W." box="[1169,1304,1117,1144]" pageId="8" pageNumber="9" pagination="153 - 255" refId="ref54455" refString="Baluk, W. (1995) Middle Miocene (Badenian) gastropods from Korytnica, Poland; Part II. Acta Geologica Polonica, 45, 153 - 255. [https: // geojournals. pgi. gov. pl / agp / article / view / 13663]" type="journal article" year="1995">
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Bałuk,
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<quantity id="6A3E61A3E263FFE8A8A13619FAA1B26F" box="[1248,1304,1117,1144]" metricMagnitude="1" metricUnit="m" metricValue="5.0673" pageId="8" pageNumber="9" unit="in" value="1995.0">1995</quantity>
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</bibRefCitation>
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</taxonomicName>
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in the Indo- West Pacific genus
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<taxonomicName id="6AC6B7C5E263FFE8AD3736C5FD9EB28C" authorityName="Iredale" authorityYear="1929" box="[374,551,1153,1179]" class="Gastropoda" family="Columbellidae" genus="Graphicomassa" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
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<emphasis id="9FB21054E263FFE8AD3736C5FD9EB28C" box="[374,551,1153,1179]" italics="true" pageId="8" pageNumber="9">Graphicomassa</emphasis>
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</taxonomicName>
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, based on their subcylindrical last whorls, presence of a subsutural concavity, giving the adapical portion of the last whorl a slightly pinched aspect, slightly gradate spires and absence of prominent axial sculpture. The absence of an internal row of columellar denticles allows a separation from
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<taxonomicName id="6AC6B7C5E263FFE8A8B9368DFAC2B2F4" box="[1272,1403,1225,1251]" class="Gastropoda" family="Columbellidae" genus="Columbella" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
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<emphasis id="9FB21054E263FFE8A8B9368DFAC2B2F4" box="[1272,1403,1225,1251]" italics="true" pageId="8" pageNumber="9">Columbella</emphasis>
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</taxonomicName>
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. A common feature of both Miocene species is a decollate early spire and the development of a low, spirally formed plug. The loss of the apex is known from several extant
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<taxonomicName id="6AC6B7C5E263FFE8AF583755FC78B33C" authorityName="Swainson" authorityYear="1840" box="[793,961,1297,1323]" class="Gastropoda" family="Columbellidae" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="family">Columbellidae</taxonomicName>
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, such as
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<taxonomicName id="6AC6B7C5E263FFE8A8683755FA25B33C" authority="(Gmelin, 1791)" baseAuthorityName="Gmelin" baseAuthorityYear="1791" box="[1065,1436,1297,1323]" class="Magnoliopsida" family="Annonaceae" genus="Mitrella" kingdom="Plantae" order="Magnoliales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="ocellata">
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<emphasis id="9FB21054E263FFE8A8683755FB5FB33C" box="[1065,1254,1297,1323]" italics="true" pageId="8" pageNumber="9">Mitrella ocellata</emphasis>
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(
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<bibRefCitation id="C957B1B7E263FFE8A8B73755FA2DB33C" author="Gmelin, J. F." box="[1270,1428,1297,1323]" pageId="8" pageNumber="9" pagination="3021 - 3910" refId="ref57083" refString="Gmelin, J. F. (1791) Vermes. In: Gmelin, J. F. (Ed.), Caroli a Linnaei Systema Naturae per Regna Tria Naturae. Tome 1 (6). 13 th Edition. G. E. Beer, Lipsiae, pp. 3021 - 3910. [http: // www. biodiversitylibrary. org / item / 83098 # 5]" type="book chapter" year="1791">Gmelin, 1791</bibRefCitation>
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)
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</taxonomicName>
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and even in some species of the deep water genus
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<taxonomicName id="6AC6B7C5E263FFE8AE9B3772FC91B347" authorityName="H. Adams & A. Adams" authorityYear="1853" box="[730,808,1334,1360]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
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<emphasis id="9FB21054E263FFE8AE9B3772FC91B347" box="[730,808,1334,1360]" italics="true" pageId="8" pageNumber="9">Astyris</emphasis>
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</taxonomicName>
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, such as
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<taxonomicName id="6AC6B7C5E263FFE8AFD53772FF76B364" authority="Araya, Catalan & Aliaga, 2016" authorityName="Araya" authorityYear="2016" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="atacamensis">
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<emphasis id="9FB21054E263FFE8AFD53772FBCDB358" box="[916,1140,1334,1360]" italics="true" pageId="8" pageNumber="9">Astyris atacamensis</emphasis>
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<bibRefCitation id="C957B1B7E263FFE8A83D3772FF76B364" author="Araya, J. F. & Catalan, R. & Aliaga, J. A." pageId="8" pageNumber="9" pagination="140 - 144" refId="ref54219" refString="Araya, J. F., Catalan, R., Aliaga, J. A. (2016) A new deep-water Astyris species (Buccinoidea: Columbellidae) from the southeastern Pacific. Zootaxa, 4139 (1), 140 - 144. https: // doi. org / 10.11646 / zootaxa. 4139.1.11" type="journal article" year="2016">Araya, Catalán & Aliaga, 2016</bibRefCitation>
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</taxonomicName>
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(pers. com. Marta deMaintenon,
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<date id="D978EA86E263FFE8AE0E371EFD6DB364" box="[591,724,1369,1396]" pageId="8" pageNumber="9" value="2020-11-07">7. 11. 2020</date>
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). An abraded apex is frequently seen in
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<taxonomicName id="6AC6B7C5E263FFE8A8E0371DFA25B364" baseAuthorityName="Duclos" baseAuthorityYear="1840" box="[1185,1436,1369,1395]" class="Gastropoda" family="Columbellidae" genus="Graphicomassa" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="ligula">
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<emphasis id="9FB21054E263FFE8A8E0371DFA25B364" box="[1185,1436,1369,1395]" italics="true" pageId="8" pageNumber="9">Graphicomassa ligula</emphasis>
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</taxonomicName>
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(e.g.
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<bibRefCitation id="C957B1B7E263FFE8AC933739FE10B380" author="deMaintenon, M." box="[210,425,1405,1431]" pageId="8" pageNumber="9" pagination="43 - 48" refId="ref56227" refString="deMaintenon, M. (2016) On the identity of Graphicomassa albina (Kiener, 1841) (Gastropoda: Columbellidae). Zoosystema, 38, 43 - 48. https: // doi. org / 10.5252 / z 2016 n 1 a 2" type="journal article" year="2016">deMaintenon 2016</bibRefCitation>
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, figs 1B, 1C, 1E, 1F). The genus was represented in Europe during the early and middle Miocene by several species:
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<taxonomicName id="6AC6B7C5E263FFE8AD9B37E5FC35B3AC" authority="(Sacco, 1890)" baseAuthorityName="Sacco" baseAuthorityYear="1890" box="[474,908,1441,1467]" class="Gastropoda" family="Columbellidae" genus="Graphicomassa" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="ringens">
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<emphasis id="9FB21054E263FFE8AD9B37E5FD5EB3AC" box="[474,743,1441,1467]" italics="true" pageId="8" pageNumber="9">Graphicomassa ringens</emphasis>
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(Sacco, 1890)
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</taxonomicName>
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,
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<taxonomicName id="6AC6B7C5E263FFE8AFD937E6FB73B3AC" authority="(Sacco, 1891)" baseAuthorityName="Sacco" baseAuthorityYear="1891" box="[920,1226,1441,1468]" class="Gastropoda" family="Columbellidae" genus="Graphicomassa" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="scalarata">
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<emphasis id="9FB21054E263FFE8AFD937E6FB9CB3AC" box="[920,1061,1441,1468]" italics="true" pageId="8" pageNumber="9">G. scalarata</emphasis>
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(Sacco, 1891)
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</taxonomicName>
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,
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<taxonomicName id="6AC6B7C5E263FFE8A89737E6FDB6B3C8" authority="(Sacco, 1890) (Proto-Mediterranean Sea)" baseAuthorityName="Sacco" baseAuthorityYear="1890" class="Gastropoda" family="Columbellidae" genus="Graphicomassa" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="inflata">
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<emphasis id="9FB21054E263FFE8A89737E6FAFBB3AC" box="[1238,1346,1441,1468]" italics="true" pageId="8" pageNumber="9">G. inflata</emphasis>
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(Sacco, 1890) (Proto-Mediterranean Sea)
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</taxonomicName>
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and
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|
<taxonomicName id="6AC6B7C5E263FFE8AE093782FCCCB3F7" authority="(Mayer, 1869)" baseAuthorityName="Mayer" baseAuthorityYear="1869" box="[584,885,1477,1504]" class="Gastropoda" family="Columbellidae" genus="Graphicomassa" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="turonica">
|
|
<emphasis id="9FB21054E263FFE8AE093782FD73B3C8" box="[584,714,1478,1504]" italics="true" pageId="8" pageNumber="9">G. turonica</emphasis>
|
|
(
|
|
<bibRefCitation id="C957B1B7E263FFE8AE9A3781FCD4B3F7" author="Mayer, C." box="[731,877,1477,1504]" pageId="8" pageNumber="9" pagination="282 - 287" refId="ref61172" refString="Mayer, C. (1869) Description de Coquilles nouvelles des etages superieurs des terrains tertiaires. Journal de Conchyliologie, 17, 82 - 88, 282 - 287. [https: // www. biodiversitylibrary. org / item / 53431 # page / 286 / mode / 1 up]" type="journal article" year="1869">Mayer, 1869</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
(northeastern Atlantic) (see
|
|
<bibRefCitation id="C957B1B7E263FFE8A8FA3781FAF6B3C8" author="Sacco, F." box="[1211,1359,1477,1503]" pageId="8" pageNumber="9" refId="ref63411" refString="Sacco, F. (1890 b) I molluschi dei terreni terziari del Piemonte e della Liguria. Parte 6. (Volutidae, Marginellidae, Columbellidae). C. Clausen, Torino, 76 pp., 2 pls. [https: // www. biodiversitylibrary. org / item / 45259 # page / 9 / mode / 1 up]" type="book" year="1890">Sacco 1890b</bibRefCitation>
|
|
, 1891;
|
|
<bibRefCitation id="C957B1B7E263FFE8ACD637AEFE9AB013" author="Mayer, C." box="[151,291,1513,1540]" pageId="8" pageNumber="9" pagination="282 - 287" refId="ref61172" refString="Mayer, C. (1869) Description de Coquilles nouvelles des etages superieurs des terrains tertiaires. Journal de Conchyliologie, 17, 82 - 88, 282 - 287. [https: // www. biodiversitylibrary. org / item / 53431 # page / 286 / mode / 1 up]" type="journal article" year="1869">Mayer 1869</bibRefCitation>
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|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
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</document> |