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<document id="B39832F3DD1A0049B29B022F5163706C" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D48920EFE6DF9A2C9C72E60" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius opatroides Arrow 1941" docType="treatment" docVersion="10" lastPageNumber="86" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="85" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="3B0AEE53207445780A27098CAFCC5B1B">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
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<mods:namePart id="A4ACFDA57168F4DCF83E2FFFEF0BC92F">Telnov, Dmitry</mods:namePart>
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<mods:title id="D0C1011DEDE8A601E4F07F82741438EC">European Journal of Taxonomy</mods:title>
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<mods:date id="BEE94B84111C225B616C3DC5A37DE137">2020</mods:date>
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<mods:classification id="BEFC7980BBE6D82BFF12313DCF54ED91">journal article</mods:classification>
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<mods:identifier id="C045DA33F610A1FB24290D4284ADF904" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
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<treatment id="1A3787F69D48920EFE6DF9A2C9C72E60" ID-DOI="http://doi.org/10.5281/zenodo.4344497" ID-GBIF-Taxon="168648159" ID-Zenodo-Dep="4344497" LSID="urn:lsid:plazi:treatment:1A3787F69D48920EFE6DF9A2C9C72E60" httpUri="http://treatment.plazi.org/id/1A3787F69D48920EFE6DF9A2C9C72E60" lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85">
<subSubSection id="DA84656B9D48920FFE6DF9A2CC602E55" box="[510,1078,1564,1591]" pageId="84" pageNumber="85" type="nomenclature">
<paragraph id="922136E09D48920FFE6DF9A2CC602E55" blockId="84.[510,1078,1564,1628]" box="[510,1078,1564,1591]" pageId="84" pageNumber="85">
<heading id="C969818C9D48920FFE6DF9A2CC602E55" box="[510,1078,1564,1591]" centered="true" fontSize="11" level="2" pageId="84" pageNumber="85" reason="2">
<taxonomicName id="559E4D639D48920FFE6DF9A2CBFB2E55" ID-CoL="7X3XC" authority="Arrow, 1941" authorityName="Arrow" authorityYear="1941" box="[510,941,1564,1591]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="opatroides">
<emphasis id="A0EAEAF29D48920FFE6DF9A2CB462E55" bold="true" box="[510,784,1564,1591]" italics="true" pageId="84" pageNumber="85">Maechidius opatroides</emphasis>
<bibRefCitation id="F60F4B119D48920FFC8BF9A3CBFB2E55" author="Arrow G. E." box="[792,941,1565,1591]" pageId="84" pageNumber="85" pagination="448 - 464" refId="ref97558" refString="Arrow G. E. 1941. XXIX. - Some undescribed species of melolonthid Coleoptera from Western New Guinea and the adjacent islands of Waigeu and Japen. Annals and Magazine of Natural History, Series 11 7 (41): 448 - 464. https: // doi. org / 10.1080 / 03745481.1941.9727945" type="journal article" year="1941">Arrow, 1941</bibRefCitation>
</taxonomicName>
comb. rest.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D48920FFDC8F9FFCB8E2E39" box="[603,984,1601,1628]" pageId="84" pageNumber="85" type="description">
<paragraph id="922136E09D48920FFDC8F9FFCB8E2E39" blockId="84.[510,1078,1564,1628]" box="[603,984,1601,1628]" pageId="84" pageNumber="85">
<figureCitation id="0AA52A659D48920FFDC8F9FFCAE32E39" box="[603,693,1601,1627]" captionStart="Figs 70–73" captionStartId="93.[189,241,1896,1922]" captionTargetBox="[327,1247,269,1869]" captionTargetId="figure-67@93.[327,1254,265,1878]" captionTargetPageId="93" captionText="Figs 70–73. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 70. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 71. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 72. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 73. M. peregrinus Lansberge, 1886, ♂ (DTC). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122158" httpUri="https://zenodo.org/record/4122158/files/figure.png" pageId="84" pageNumber="85">Figs 70</figureCitation>
,
<figureCitation id="0AA52A659D48920FFD52F9FFCAA62E3E" box="[705,752,1601,1628]" captionStart="Figs 145–156" captionStartId="141.[189,241,1787,1813]" captionTargetBox="[328,1254,265,1740]" captionTargetId="figure-124@141.[328,1260,265,1749]" captionTargetPageId="141" captionText="Figs 145–156. Papuan Maechidius Macleay, 1819, head, dorsal view. 145. M. merdeka sp. nov., holotype, ♂ (BMNH). 146. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 147. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 148. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 149. Same, ♂ (SNSD). 150. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 151. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 152. M. nepenthephilus sp. nov., holotype, ♂ (NME). 153. Same, paratype, ♀ (DTC). 154. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 155. Same, paratype, ♀ (BMNH). 156. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122181" httpUri="https://zenodo.org/record/4122181/files/figure.png" pageId="84" pageNumber="85">150</figureCitation>
,
<figureCitation id="0AA52A659D48920FFD6FF9FFCB7D2E39" box="[764,811,1601,1627]" captionStart="Figs 226–241" captionStartId="147.[189,241,1681,1707]" captionTargetBox="[391,1192,268,1647]" captionTargetId="figure-170@147.[391,1196,265,1656]" captionTargetPageId="147" captionText="Figs 226–241. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 226. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 227. M. lobaticeps Frey, 1969, ♂ (DTC). 228. M. longipes sp. nov., holotype, ♂ (BMNH). 229. M. luniceps Fairmaire, 1883, ♂ (BMNH). 230. M. mailu sp. nov., holotype, ♂ (NHMB). 231. M. maleo sp. nov., holotype, ♂ (IRSN). 232. M. merdeka sp. nov., holotype, ♂ (BMNH). 233. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 234. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 235. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 236. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 237. M. nepenthephilus sp. nov., holotype, ♂ (NME). 238. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 239. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 240. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 241. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122193" httpUri="https://zenodo.org/record/4122193/files/figure.png" pageId="84" pageNumber="85">238</figureCitation>
,
<figureCitation id="0AA52A659D48920FFCA4F9FFCB302E39" box="[823,870,1601,1627]" captionStart="Figs 317–332" captionStartId="154.[189,241,1681,1707]" captionTargetBox="[401,1187,265,1655]" captionTargetId="figure-172@154.[401,1187,265,1669]" captionTargetPageId="154" captionText="Figs 317–332. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 317. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 318. M. lobaticeps Frey, 1969, ♂ (DTC). 319. M. longipes sp. nov., holotype, ♂ (BMNH). 320. M. luniceps Fairmaire, 1883, ♂ (BMNH). 321. M. mailu sp. nov., holotype, ♂ (NHMB). 322. M. maleo sp. nov., holotype, ♂ (IRSN). 323. M. merdeka sp. nov., holotype, ♂ (BMNH). 324. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 325. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 326. M. muticusArrow, 1941, syntype, ♂ (BMNH). 327. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 328. M. nepenthephilus sp. nov., holotype, ♂ (NME). 329. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 330. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 331. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 332. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122210" httpUri="https://zenodo.org/record/4122210/files/figure.png" pageId="84" pageNumber="85">329</figureCitation>
, 402, 498
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D48920EFF2EF93DCD722914" lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85" type="materials_examined">
<paragraph id="922136E09D48920FFF2EF93DC93A2EFF" blockId="84.[189,364,1667,1693]" box="[189,364,1667,1693]" pageId="84" pageNumber="85">
<heading id="C969818C9D48920FFF2EF93DC93A2EFF" bold="true" box="[189,364,1667,1693]" fontSize="11" level="3" pageId="84" pageNumber="85" reason="3">
<emphasis id="A0EAEAF29D48920FFF2EF93DC93A2EFF" bold="true" box="[189,364,1667,1693]" pageId="84" pageNumber="85">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D48920FFF79F90DCA3A2EAF" blockId="84.[189,1399,1714,1886]" box="[234,620,1714,1741]" pageId="84" pageNumber="85">
<emphasis id="A0EAEAF29D48920FFF79F90DC9302EAF" bold="true" box="[234,358,1715,1741]" pageId="84" pageNumber="85">
<typeStatus id="4D2588429D48920FFF79F90DC9302EAF" box="[234,358,1715,1741]" pageId="84" pageNumber="85" type="lectotype">Lectotype</typeStatus>
</emphasis>
[herewith designated]
</paragraph>
<paragraph id="922136E09D48920FFF2EF969CADD2F3C" blockId="84.[189,1399,1714,1886]" pageId="84" pageNumber="85">
<materialsCitation id="22F63CBD9D48920FFF2EF969CADD2F3C" ID-GBIF-Occurrence="2898878496" collectingDate="1938-04" collectionCode="BMNH" collectorName="L. E. Cheesman." country="Indonesia" elevation="762" location="INDONESIA" pageId="84" pageNumber="85" specimenCount="1" specimenCount-female="1" typeStatus="syntype">
<collectingCountry id="EA8976709D48920FFF2EF969C90D2E93" box="[189,347,1751,1777]" name="Indonesia" pageId="84" pageNumber="85">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D48920FFEEBF969C9D92E93" box="[376,399,1751,1777]" pageId="84" pageNumber="85" type="female">♀</specimenCount>
; “Type [p, label circular, red frame] //
<typeStatus id="4D2588429D48920FFCC4F969CB8D2E93" box="[855,987,1751,1777]" pageId="84" pageNumber="85" type="syntype">SYNTYPE</typeStatus>
[p, label circular, blue frame] // N. DUTCH NEW GUINEA: Waigeu. Camp Nok.
<quantity id="55669B059D48920FFCB1F945CB282F74" box="[802,894,1787,1814]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="84" pageNumber="85" unit="ft" value="2500.0">
<elevation id="19B3D1D39D48920FFCB1F945CB282F74" box="[802,894,1787,1814]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="84" pageNumber="85" unit="ft" value="2500.0">2,500ft.</elevation>
</quantity>
<date id="E62010209D48920FFC1DF945CBBF2F77" box="[910,1001,1787,1813]" pageId="84" pageNumber="85" value="1938-04">
<collectingDate id="F664E9C89D48920FFC1DF945CBBF2F77" box="[910,1001,1787,1813]" pageId="84" pageNumber="85" value="1938-04">iv.1938</collectingDate>
</date>
.
<collectorName id="3F6B53369D48920FFC6EF945CCE42F77" box="[1021,1202,1787,1813]" pageId="84" pageNumber="85">L.E.Cheesman.</collectorName>
B.M.1938-593. [p] //
<emphasis id="A0EAEAF29D48920FFF64F8A1CA1B2F5B" box="[247,589,1823,1849]" italics="true" pageId="84" pageNumber="85">
Coniomaechidius
<taxonomicName id="559E4D639D48920FFE42F8A1CA1B2F5B" authorityName="Arrow" authorityYear="1941" box="[465,589,1823,1849]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="opatroides">opatroides</taxonomicName>
</emphasis>
,
<typeStatus id="4D2588429D48920FFDF1F89ECACA2F58" box="[610,668,1824,1850]" pageId="84" pageNumber="85">Type</typeStatus>
Arrow [h]”;
<collectionCode id="F48FAE259D48920FFCD5F89ECBC82F58" box="[838,926,1824,1850]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="84" pageNumber="85">BMNH</collectionCode>
[the name “
<emphasis id="A0EAEAF29D48920FFBD3F8A1CD5B2F5B" box="[1088,1293,1823,1849]" italics="true" pageId="84" pageNumber="85">Coniomaechidius</emphasis>
”, which appears on label, was never published].
</materialsCitation>
</paragraph>
<paragraph id="922136E09D48920FFF79F838C9F42FC2" blockId="84.[189,1399,1926,2025]" box="[234,418,1926,1952]" pageId="84" pageNumber="85">
<emphasis id="A0EAEAF29D48920FFF79F838C9F42FC2" bold="true" box="[234,418,1926,1952]" pageId="84" pageNumber="85">
<typeStatus id="4D2588429D48920FFF79F838C9F42FC2" box="[234,418,1926,1952]" pageId="84" pageNumber="85" type="paralectotype">Paralectotypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D48920FFF2EF814CC7F2F8B" blockId="84.[189,1399,1926,2025]" pageId="84" pageNumber="85">
<materialsCitation id="22F63CBD9D48920FFF2EF814CCC72FA7" ID-GBIF-Occurrence="2898878312" box="[189,1169,1961,1989]" collectionCode="DUTCH, NEW" country="Indonesia" location="INDONESIA" pageId="84" pageNumber="85" specimenCount="3" specimenCount-female="3" typeStatus="syntype">
<collectingCountry id="EA8976709D48920FFF2EF814C90D2FA6" box="[189,347,1962,1988]" name="Indonesia" pageId="84" pageNumber="85">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D48920FFEE0F814C9E32FA7" box="[371,437,1962,1989]" pageId="84" pageNumber="85" type="female">3 ♀♀</specimenCount>
; “
<typeStatus id="4D2588429D48920FFE5EF814CA022FA6" box="[461,596,1962,1988]" pageId="84" pageNumber="85" type="syntype">SYNTYPE</typeStatus>
[p, label circular, blue frame] // N.
<collectionCode id="F48FAE259D48920FFC4CF814CC1E2FA6" box="[991,1096,1962,1988]" pageId="84" pageNumber="85">DUTCH</collectionCode>
<collectionCode id="F48FAE259D48920FFBDCF815CCC72FA7" box="[1103,1169,1963,1989]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="84" pageNumber="85">NEW</collectionCode>
</materialsCitation>
<materialsCitation id="22F63CBD9D48920FFB04F814CC732F8B" ID-GBIF-Occurrence="2898878600" collectingDate="1938-04" collectionCode="BMNH" collectorName="L. E. Cheesman." country="Guinea-Bissau" elevation="762" location="GUINEA" pageId="84" pageNumber="85" specimenCount="1" typeStatus="paralectotype">
<collectingCountry id="EA8976709D48920FFB04F814CD5A2FA6" box="[1175,1292,1962,1988]" name="Guinea-Bissau" pageId="84" pageNumber="85">GUINEA</collectingCountry>
: Waigeu. Camp Nok.
<quantity id="55669B059D48920FFEDFF870C9F12F8B" box="[332,423,1998,2025]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="84" pageNumber="85" unit="ft" value="2500.0">
<elevation id="19B3D1D39D48920FFEDFF870C9F12F8B" box="[332,423,1998,2025]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="84" pageNumber="85" unit="ft" value="2500.0">2,500ft.</elevation>
</quantity>
<date id="E62010209D48920FFE3DF870CA5F2F8A" box="[430,521,1998,2024]" pageId="84" pageNumber="85" value="1938-04">
<collectingDate id="F664E9C89D48920FFE3DF870CA5F2F8A" box="[430,521,1998,2024]" pageId="84" pageNumber="85" value="1938-04">iv.1938</collectingDate>
</date>
.
<collectorName id="3F6B53369D48920FFD87F870CA9E2F8A" box="[532,712,1998,2024]" pageId="84" pageNumber="85">L.E.Cheesman.</collectorName>
B.M.1938-593. [p]”;
<collectionCode id="F48FAE259D48920FFC5AF871CC732F8B" box="[969,1061,1999,2025]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="84" pageNumber="85">BMNH</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="922136E09D49920EFF2EFEB7C9302940" blockId="85.[189,358,264,291]" box="[189,358,264,291]" pageId="85" pageNumber="86">
<heading id="C969818C9D49920EFF2EFEB7C9302940" bold="true" box="[189,358,264,291]" fontSize="11" level="3" pageId="85" pageNumber="86" reason="3">
<emphasis id="A0EAEAF29D49920EFF2EFEB7C9302940" bold="true" box="[189,358,264,291]" pageId="85" pageNumber="86">New material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D49920EFF2EFE86CD722914" blockId="85.[189,1399,312,375]" pageId="85" pageNumber="86">
<materialsCitation id="22F63CBD9D49920EFF2EFE86CA9C2930" ID-GBIF-Occurrence="2898878625" box="[189,714,312,338]" collectionCode="DUTCH, NEW" country="Indonesia" location="INDONESIA" pageId="85" pageNumber="86" specimenCount="1" specimenCount-female="1">
<collectingCountry id="EA8976709D49920EFF2EFE86C90D2930" box="[189,347,312,338]" name="Indonesia" pageId="85" pageNumber="86">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D49920EFE1CFE86C99F2930" box="[399,457,312,338]" pageId="85" pageNumber="86" type="female">1 ♀</specimenCount>
; “N.DUTCH NEW
</materialsCitation>
<materialsCitation id="22F63CBD9D49920EFD4CFE86CD762914" ID-GBIF-Occurrence="2898878642" collectingDate="1938-05" collectionCode="BMNH" collectorName="Waigeu. Camp I. Mt. Nok. &amp; L. E. Cheesman." country="Guinea-Bissau" determinerName="M. E. Bacchus" elevation="762" location="Mt. Nok." pageId="85" pageNumber="86" specimenCount="1" typeStatus="syntype">
GUINEA:
<collectorName id="3F6B53369D49920EFCFFFE86CCCA2930" box="[876,1180,312,338]" pageId="85" pageNumber="86">Waigeu.Camp I.Mt.Nok.</collectorName>
<quantity id="55669B059D49920EFB23FE86CD5A2931" box="[1200,1292,312,339]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="85" pageNumber="86" unit="ft" value="2500.0">
<elevation id="19B3D1D39D49920EFB23FE86CD5A2931" box="[1200,1292,312,339]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="85" pageNumber="86" unit="ft" value="2500.0">2,500ft.</elevation>
</quantity>
<date id="E62010209D49920EFAB3FE86CD252930" box="[1312,1395,312,338]" pageId="85" pageNumber="86" value="1938-05">
<collectingDate id="F664E9C89D49920EFAB3FE86CD252930" box="[1312,1395,312,338]" pageId="85" pageNumber="86" value="1938-05">v.1938</collectingDate>
</date>
.
<collectorName id="3F6B53369D49920EFF2EFEE2C9272914" box="[189,369,348,374]" pageId="85" pageNumber="86">L.E.Cheesman.</collectorName>
B.M.1938-593. [p] // not a
<typeStatus id="4D2588429D49920EFD3CFEE2CB5C2914" box="[687,778,348,374]" pageId="85" pageNumber="86" type="syntype">syntype</typeStatus>
[h]
<determinerName id="67FDDA249D49920EFCA8FEE2CBB12914" box="[827,999,348,374]" pageId="85" pageNumber="86">M. E. Bacchus</determinerName>
det. 196[p] 9[h]”;
<collectionCode id="F48FAE259D49920EFB57FEE2CD762914" box="[1220,1312,348,374]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="85" pageNumber="86">BMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D49920EFF2EFE20CCBF2A10" pageId="85" pageNumber="86" type="discussion">
<paragraph id="922136E09D49920EFF2EFE20C97929DA" blockId="85.[189,303,414,440]" box="[189,303,414,440]" pageId="85" pageNumber="86">
<heading id="C969818C9D49920EFF2EFE20C97929DA" bold="true" box="[189,303,414,440]" fontSize="11" level="3" pageId="85" pageNumber="86" reason="3">
<emphasis id="A0EAEAF29D49920EFF2EFE20C97929DA" bold="true" box="[189,303,414,440]" pageId="85" pageNumber="86">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D49920EFF2EFE73C9AF2A6E" blockId="85.[189,1399,461,524]" pageId="85" pageNumber="86">
The
<typeStatus id="4D2588429D49920EFF66FE73C9372985" box="[245,353,461,487]" pageId="85" pageNumber="86" type="lectotype">lectotype</typeStatus>
is designated in order to enhance the stability of nomenclature and fix the specimen I based my redescription on.
</paragraph>
<paragraph id="922136E09D49920EFF2EFD8ACCBF2A10" blockId="85.[189,1399,564,626]" pageId="85" pageNumber="86">
<bibRefCitation id="F60F4B119D49920EFF2EFD8AC9CE2A2D" author="Arrow G. E." box="[189,408,564,591]" pageId="85" pageNumber="86" pagination="448 - 464" refId="ref97558" refString="Arrow G. E. 1941. XXIX. - Some undescribed species of melolonthid Coleoptera from Western New Guinea and the adjacent islands of Waigeu and Japen. Annals and Magazine of Natural History, Series 11 7 (41): 448 - 464. https: // doi. org / 10.1080 / 03745481.1941.9727945" type="journal article" year="1941">Arrow (1941: 454)</bibRefCitation>
described this species from four female specimens, not designating the
<typeStatus id="4D2588429D49920EFB46FD8ACD162A2C" box="[1237,1344,564,590]" pageId="85" pageNumber="86" type="holotype">holotype</typeStatus>
. The BMNH specimen with Arrow’s handwriting “type” is herewith designated as the
<typeStatus id="4D2588429D49920EFBE6FDE6CCB22A10" box="[1141,1252,600,626]" pageId="85" pageNumber="86" type="lectotype">lectotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D49920EFF2EFD24C9C72E60" pageId="85" pageNumber="86" type="description">
<paragraph id="922136E09D49920EFF2EFD24C9182AD6" blockId="85.[189,334,666,692]" box="[189,334,666,692]" pageId="85" pageNumber="86">
<heading id="C969818C9D49920EFF2EFD24C9182AD6" bold="true" box="[189,334,666,692]" fontSize="11" level="3" pageId="85" pageNumber="86" reason="3">
<emphasis id="A0EAEAF29D49920EFF2EFD24C9182AD6" bold="true" box="[189,334,666,692]" pageId="85" pageNumber="86">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D49920EFF2EFD74C9292DF2" blockId="85.[189,1399,713,1425]" pageId="85" pageNumber="86">
Dorsum densely covered with dirty yellowish microscopical velvety pubescence. Female labroclypeus (
<figureCitation id="0AA52A659D49920EFF55FD50C97D2B6A" box="[198,299,749,776]" captionStart="Figs 145–156" captionStartId="141.[189,241,1787,1813]" captionTargetBox="[328,1254,265,1740]" captionTargetId="figure-124@141.[328,1260,265,1749]" captionTargetPageId="141" captionText="Figs 145–156. Papuan Maechidius Macleay, 1819, head, dorsal view. 145. M. merdeka sp. nov., holotype, ♂ (BMNH). 146. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 147. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 148. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 149. Same, ♂ (SNSD). 150. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 151. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 152. M. nepenthephilus sp. nov., holotype, ♂ (NME). 153. Same, paratype, ♀ (DTC). 154. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 155. Same, paratype, ♀ (BMNH). 156. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122181" httpUri="https://zenodo.org/record/4122181/files/figure.png" pageId="85" pageNumber="86">Fig. 150</figureCitation>
) broadly emarginate on anterior margin, its lateral margins nearly straight in dorsal, slightly sinuous in lateral view. Anterolateral angles of labroclypeus acute, rather strongly protruding. Head dorsal punctures dense, circular to ovoid, deep. Intervening spaces generally smaller than punctures. Setae scale-like, suberect, clavate, small to rather long. Pronotum anterior and basal margins sinuous in dorsal view. Lateral margin of pronotum somewhat flattened and laterally expanding in anterior half (in dorsal view), strongly sinuous in dorsal view, broadly rounded in anterior ¾, emarginate prebasally and again widened at base. Lateral margin of pronotum nearly straight in lateral view, delicately crenulate all along (
<figureCitation id="0AA52A659D49920EFEA6FC54C9C92C66" box="[309,415,1002,1028]" captionStart="Figs 226–241" captionStartId="147.[189,241,1681,1707]" captionTargetBox="[391,1192,268,1647]" captionTargetId="figure-170@147.[391,1196,265,1656]" captionTargetPageId="147" captionText="Figs 226–241. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 226. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 227. M. lobaticeps Frey, 1969, ♂ (DTC). 228. M. longipes sp. nov., holotype, ♂ (BMNH). 229. M. luniceps Fairmaire, 1883, ♂ (BMNH). 230. M. mailu sp. nov., holotype, ♂ (NHMB). 231. M. maleo sp. nov., holotype, ♂ (IRSN). 232. M. merdeka sp. nov., holotype, ♂ (BMNH). 233. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 234. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 235. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 236. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 237. M. nepenthephilus sp. nov., holotype, ♂ (NME). 238. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 239. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 240. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 241. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122193" httpUri="https://zenodo.org/record/4122193/files/figure.png" pageId="85" pageNumber="86">Fig. 238</figureCitation>
). A scale-like curved clavate seta present between every two crenulae. Hypomeron flange-like protruding, emarginate opposite compound eye. Antennal pocket deep. Pronotal punctures elongate ovoid, deep. Intervening spaces generally larger than punctures. Lateral sides of pronotum covered with dense brush-like clusters of setae reminiscent of mammal fur (
<figureCitation id="0AA52A659D49920EFBC2FBE8CCEC2C12" box="[1105,1210,1110,1136]" captionStart="Figs 226–241" captionStartId="147.[189,241,1681,1707]" captionTargetBox="[391,1192,268,1647]" captionTargetId="figure-170@147.[391,1196,265,1656]" captionTargetPageId="147" captionText="Figs 226–241. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 226. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 227. M. lobaticeps Frey, 1969, ♂ (DTC). 228. M. longipes sp. nov., holotype, ♂ (BMNH). 229. M. luniceps Fairmaire, 1883, ♂ (BMNH). 230. M. mailu sp. nov., holotype, ♂ (NHMB). 231. M. maleo sp. nov., holotype, ♂ (IRSN). 232. M. merdeka sp. nov., holotype, ♂ (BMNH). 233. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 234. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 235. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 236. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 237. M. nepenthephilus sp. nov., holotype, ♂ (NME). 238. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 239. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 240. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 241. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122193" httpUri="https://zenodo.org/record/4122193/files/figure.png" pageId="85" pageNumber="86">Fig. 238</figureCitation>
), dorsal surface fully hidden. Setae scale-like, clavate, suberect; each seta rises from anterior margin of corresponding puncture, not surpassing its length. Elytral punctures elongate, irregularly sized and shaped, deep (
<figureCitation id="0AA52A659D49920EFF55FB7CC9792CBE" box="[198,303,1218,1244]" captionStart="Figs 317–332" captionStartId="154.[189,241,1681,1707]" captionTargetBox="[401,1187,265,1655]" captionTargetId="figure-172@154.[401,1187,265,1669]" captionTargetPageId="154" captionText="Figs 317–332. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 317. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 318. M. lobaticeps Frey, 1969, ♂ (DTC). 319. M. longipes sp. nov., holotype, ♂ (BMNH). 320. M. luniceps Fairmaire, 1883, ♂ (BMNH). 321. M. mailu sp. nov., holotype, ♂ (NHMB). 322. M. maleo sp. nov., holotype, ♂ (IRSN). 323. M. merdeka sp. nov., holotype, ♂ (BMNH). 324. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 325. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 326. M. muticusArrow, 1941, syntype, ♂ (BMNH). 327. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 328. M. nepenthephilus sp. nov., holotype, ♂ (NME). 329. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 330. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 331. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 332. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122210" httpUri="https://zenodo.org/record/4122210/files/figure.png" pageId="85" pageNumber="86">Fig. 329</figureCitation>
). Intervening spaces much larger than punctures. Elytral setae scale-like, clavate, appressed, not surpassing length of corresponding punctures. Longer suberect scale-like clavate setae arranged in irregular longitudinal rows – single sutural and two doubled rows. Female pygidium covered with furlike pubescence, deeply densely punctate with ovoid punctures (
<figureCitation id="0AA52A659D49920EFC3AFA90CC462D2A" box="[937,1040,1326,1352]" captionStart="Figs487–503" captionStartId="165.[189,241,1681,1707]" captionTargetBox="[422,1160,265,1655]" captionTargetId="figure-171@165.[421,1167,265,1658]" captionTargetPageId="165" captionText="Figs487–503.Papuan andWallaceanMaechidius Macleay, 1819,pygidium, dorsalview.487.M.lobaticeps Frey, 1969, ♂ (DTC). 488. M. longipes sp. nov., holotype, ♂ (BMNH). 489. M. luniceps Fairmaire, 1883, ♂ (BMNH). 490. M. mailu sp. nov., holotype, ♂ (NHMB). 491. M. maleo sp. nov., holotype, ♂ (IRSN). 492. M. merdeka sp. nov., holotype, ♂ (BMNH). 493. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 494. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 495. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 496. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 497. M. nepenthephilus sp. nov., holotype, ♂ (NME). 498. M. opatroidesArrow, 1941 comb. rest., lectotype, ♀ (BMNH). 499. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 500. Same, paratype, ♀ (BMNH). 501. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 502. Same, different aspect. 503. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122226" httpUri="https://zenodo.org/record/4122226/files/figure.png" pageId="85" pageNumber="86">Fig. 498</figureCitation>
). Setae of pygidium scale-like, elongate, suberect, rather long. Female protibia with two distal teeth on external margin, both nearly acute (Fig. 402).
</paragraph>
<paragraph id="922136E09D49920EFF2EFA06C9E62DB0" blockId="85.[189,432,1464,1490]" box="[189,432,1464,1490]" pageId="85" pageNumber="86">
<heading id="C969818C9D49920EFF2EFA06C9E62DB0" bold="true" box="[189,432,1464,1490]" fontSize="11" level="3" pageId="85" pageNumber="86" reason="3">
<emphasis id="A0EAEAF29D49920EFF2EFA06C9E62DB0" bold="true" box="[189,432,1464,1490]" pageId="85" pageNumber="86">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D49920EFF2EFA56C9C72E60" blockId="85.[189,401,1512,1538]" box="[189,401,1512,1538]" pageId="85" pageNumber="86">Male is unknown.</paragraph>
</subSubSection>
</treatment>
</document>
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