120 lines
13 KiB
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120 lines
13 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/dez.70.107322" ID-Pensoft-Pub="1860-1324-2-291" ID-Pensoft-UUID="DD34414053665BD69D8E11CB57724004" ID-ZooBank="52692B916EE24AA0B9D639BDF3107D92" ModsDocID="1860-1324-70-2-291" checkinTime="1692351740246" checkinUser="pensoft" docAuthor="Sasakawa, Koji & Mitsuduka, Yoshiji" docDate="2023" docId="083E24067E0F50DB82FF6898327C2997" docLanguage="en" docName="DeutEntomolZeit 70(2): 291-310" docOrigin="Deutsche Entomologische Zeitschrift 70 (2)" docPubDate="2023-08-17" docSource="http://dx.doi.org/10.3897/dez.70.107322" docTitle="Pterostichus (Nialoe) asahinus Habu & Baba 1960" docType="treatment" docVersion="1" id="DD34414053665BD69D8E11CB57724004" lastPageNumber="291" masterDocId="DD34414053665BD69D8E11CB57724004" masterDocTitle="A morphology-based revision and phylogenetic analysis of the Pterostichus macrogenys species group (Coleoptera, Carabidae) and implications for differentiation of the species group" masterLastPageNumber="310" masterPageNumber="291" pageNumber="291" updateTime="1692351740246" updateUser="pensoft">
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<mods:title id="5A2DC9EDC649EE1DE6255343A73ECA1B">A morphology-based revision and phylogenetic analysis of the Pterostichus macrogenys species group (Coleoptera, Carabidae) and implications for differentiation of the species group</mods:title>
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<mods:roleTerm id="4537F8B80A098F3285E34800DFAA8C52">Author</mods:roleTerm>
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<mods:namePart id="25E6BA58F1BCE68B97235790BAACC8A9">Sasakawa, Koji</mods:namePart>
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<mods:affiliation id="1F077E5E593BBC502CE73BF85DB14B29">Laboratory of Zoology, Department of Science Education, Faculty of Education, Chiba University, 1 - 33 Yayoi-cho, Inage-ku, Chiba-shi, Chiba 263 - 8522, Japan</mods:affiliation>
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<mods:nameIdentifier id="5442C3A336182BD91F6B0D8F8E0F816A" type="email">ksasa@chiba-u.jp</mods:nameIdentifier>
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<mods:namePart id="166347DAD7BE22203D019FCC63574E71">Mitsuduka, Yoshiji</mods:namePart>
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<mods:affiliation id="B9ABF33B00AD045B3913680E63C2D778">2 - 18 - 24 Hanatate Yamagata-shi, Yamagata 990 - 0067, Japan</mods:affiliation>
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<mods:typeOfResource id="8475531F7D72686148F122F32F0D98AA">text</mods:typeOfResource>
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<mods:title id="BE45362D9F8CB94C52C15E0441D0E685">Deutsche Entomologische Zeitschrift</mods:title>
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<mods:date id="415E7FF8A2F83D62F09593356E0A2845">2023</mods:date>
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<mods:number id="CD50FA8D38333DE34FF725C77988027A">2023-08-17</mods:number>
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<mods:number id="73C2241990729B5CE5680DA25C788506">70</mods:number>
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<mods:number id="A4B40E973B0CB756051B917288FCE0E1">2</mods:number>
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<mods:classification id="DD34887B7875C657D718C1625CEE7439">journal article</mods:classification>
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<mods:identifier id="A36D1C2E7F78631434C1D52228C98C7A" type="DOI">http://dx.doi.org/10.3897/dez.70.107322</mods:identifier>
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<mods:identifier id="CD17D361A2BE9975C8BD0375179E83A3" type="Pensoft-Pub">1860-1324-2-291</mods:identifier>
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<treatment id="083E24067E0F50DB82FF6898327C2997" LSID="urn:lsid:plazi:treatment:083E24067E0F50DB82FF6898327C2997" httpUri="http://treatment.plazi.org/id/083E24067E0F50DB82FF6898327C2997" lastPageNumber="291" pageId="0" pageNumber="291">
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<subSubSection id="FD6013AD1728EB3724B0D6975DC04211" pageId="0" pageNumber="291" type="nomenclature">
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<paragraph id="E10DFAC975779850689FB0FEFD58C5A2" pageId="0" pageNumber="291">
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<taxonomicName id="D029CB9FCBAEE6B9B5A25EA83B78B380" ID-CoL="8MHQC" LSID="083E2406-7E0F-50DB-82FF-6898327C2997" authority="Habu & Baba, 1960" authorityName="Habu & Baba" authorityYear="1960" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Nialoe) asahinus" order="Coleoptera" pageId="0" pageNumber="291" phylum="Arthropoda" rank="species" species="asahinus" subGenus="Nialoe">Pterostichus (Nialoe) asahinus Habu & Baba, 1960</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="16AA70ADDCD8036D83E05CA31DA5B51B" pageId="0" pageNumber="291" type="description">
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<paragraph id="150E2CFD708A7E91D4A1F31F25646A24" pageId="0" pageNumber="291">
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<figureCitation id="28A4386D5873C601A86B824FA0F01C50" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus dorsal view of the Pterostichus macrogenys species group. A. Pterostichus namahage sp. nov. holotype male from Haraikawa; B. P. kamurosanus sp. nov. holotype male from Ariya; C. P. atsumidakensis sp. nov. holotype male from Take-no-koshi; D. P. asahinus male from Momoyake." figureDoi="10.3897/dez.70.107322.figure2" httpUri="https://binary.pensoft.net/fig/893610" pageId="0" pageNumber="291">Figs 2D</figureCitation>
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<figureCitation id="817F1AC49548FD0C0C9FF03F01EBFD5E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Endophallus of Pterostichus namahage sp. nov. holotype male from Haraikawa (A-D), P. kamurosanus sp. nov. holotype male from Ariya (E-H), P. atsumidakensis sp. nov. holotype male from Take-no-koshi (I-L) and P. asahinus male from Momoyake (M-P). Fully inflated endophallus in left lateral (A, E, I, M), apical (B, F, J, N), right lateral (C, G, K, O) and basal part in dorsal and apical part in ventral (D, H, L, P) views. Abbreviations: go - gonopore; lal - left apical lobe; lpb - left pigmented band; lpl - left pre-apical lobe; rpl - right pre-apical lobe." figureDoi="10.3897/dez.70.107322.figure4" httpUri="https://binary.pensoft.net/fig/893612" pageId="0" pageNumber="291">, 4M-P</figureCitation>
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<figureCitation id="FD115FD0FE21650E97D0E55FC7C8EE2A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Right paramere of Pterostichus namahage sp. nov. holotype male from Haraikawa (A), P. kamurosanus sp. nov. holotype male from Ariya (B), P. atsumidakensis sp. nov. holotype male from Take-no-koshi (C-E), P. asahinus male from Momoyake (F-H), P. chokaisanus male from Momoyake (I), P. chokaisanus male from Zao-chuo-kogen (J), P. kitakamisanus male from Odateminami (K, L) and P. kitakamisanus holotype male from Genbeidaira (M). Left lateral (A-C, F, I-K), right lateral (L, M), apical part in apical and basal part in ventral (D, G) and apical part in dorsal (E, H) views." figureDoi="10.3897/dez.70.107322.figure6" httpUri="https://binary.pensoft.net/fig/893614" pageId="0" pageNumber="291">, 6F-H</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="AB28AB3A63F8A936CB616F7D690292C9" pageId="0" pageNumber="291" type="materials_examined">
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<paragraph id="0FD23FFC3A27330E05B9D1895EF611D5" pageId="0" pageNumber="291">Materials examined.</paragraph>
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<paragraph id="8621EFC5CD8F46E8C1A44D4AD8985E24" pageId="0" pageNumber="291">
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<materialsCitation id="43E2C39007924DC46D03E00D389A10E4" accessionNumber="SVJ26680" collectingDate="2021-06-24" collectingDateMax="2021-07-15" collectingDateMin="2021-06-24" collectingMethod="subterranean baited trap" collectorName="Y. Mitsuduka" country="Japan" elevation="1030" latitude="39.10911" location="Momoyake" longLatPrecision="1" longitude="140.15198" municipality="Yurihonjo-shi" specimenCount="1" specimenCount-male="1" stateProvince="Akita Prefecture">
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<specimenCount id="78A358F9CCE7B8D1D68961774FE26AA3" type="male">1♂</specimenCount>
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,
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<location id="4499BE3473455B138004EEBC90533515" LSID="urn:lsid:plazi:treatment:083E24067E0F50DB82FF6898327C2997:4499BE3473455B138004EEBC90533515" country="Japan" latitude="39.10911" longLatPrecision="1" longitude="140.15198" municipality="Yurihonjo-shi" name="Momoyake" stateProvince="Akita Prefecture">Momoyake</location>
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, alt.
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<quantity id="3A28DA75F13992C6A4981512A9BCE875" metricMagnitude="3" metricUnit="m" metricValue="1.03" unit="m" value="1030.0">
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<elevation id="6CF9ECC62E1171D2E3EA34FB4E2E4D67" metricMagnitude="3" metricUnit="m" metricValue="1.03" unit="m" value="1030.0">1030 m</elevation>
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</quantity>
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,
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<normalizedToken id="8104147CC176B491CA58837DC24B29A6" originalValue="Yurihonjô-shi">
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<collectingMunicipality id="1B7065A22EA031E451F7C5031A7D220A">Yurihonjo-shi</collectingMunicipality>
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</normalizedToken>
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,
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<collectingRegion id="C0D99123AF860F35C660FE9047389FF1" country="Japan" name="Akita">Akita Prefecture</collectingRegion>
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,
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<collectingCountry id="EE70AC1A111F813B832D09ED744C3095" name="Japan">Japan</collectingCountry>
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(MGRS: 54
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<accessionNumber id="48CE64989FE866BB6D7D16D2324D44A3" httpUri="http://www.ncbi.nlm.nih.gov/protein/SVJ26680">SVJ26680</accessionNumber>
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29227;
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<geoCoordinate id="F62EAD8F73DA20F9ABB8A935BB2C495B" degrees="39.1091115" direction="north" orientation="latitude" precision="1" value="39.10911">39.1091115°N</geoCoordinate>
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,
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<geoCoordinate id="3FE4100EE9E7BE2755398664A6C81029" degrees="140.15197046" direction="east" orientation="longitude" precision="1" value="140.15198">140.15197046°E</geoCoordinate>
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),
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<collectingDate id="8D80D9882052B13BD07ABE5849B0C2B9" value="2021-06-24" valueMax="2021-07-15" valueMin="2021-06-24">24. vi-15. vii. 2021</collectingDate>
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,
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<collectingMethod id="7CCC6891EEDBD4A99213C067E415FDA8">subterranean baited trap</collectingMethod>
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,
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<collectorName id="DEAA8FCA92CD7015E5712B68544E508D">Y. Mitsuduka</collectorName>
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leg.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="01BEB34ED92BA5E50C10C0955CBE419E" pageId="0" pageNumber="291" type="notes">
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<paragraph id="F421BBF40EB9E05646CBC887F4D51E24" pageId="0" pageNumber="291">Notes.</paragraph>
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<paragraph id="F1616EF1BBE9F46E8B4B269B1218242C" pageId="0" pageNumber="291">
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The body lengths (mm) of the specimen are BLm 16.38, BLl 14.60 and BLc 14.06. This specimen was collected together with
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<taxonomicName id="5E198668B600A0F4AA82C2CEE6EACD2B" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="P. chokaisanus" order="Coleoptera" pageId="0" pageNumber="291" rank="species" species="chokaisanus">
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<emphasis id="EAC5BA8EB291A867298E7B0AD79EC137" italics="true" pageId="0" pageNumber="291">P. chokaisanus</emphasis>
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</taxonomicName>
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, which is to be recorded below. This is the first report of the sympatric occurrence of this species pair and provides evidence that they are reproductively isolated distinct species. Although the comparison was based on a single male for each species, a body-size difference was observed between the two species, with
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<taxonomicName id="1A6A6F075A436D25895F0190EE1CD197" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="P. asahinus" order="Coleoptera" pageId="0" pageNumber="291" rank="species" species="asahinus">
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<emphasis id="0CB83000A91968C6C455AC835EEAE395" italics="true" pageId="0" pageNumber="291">P. asahinus</emphasis>
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</taxonomicName>
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larger than
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<taxonomicName id="619304900B40D037FC2BFCC3CA45CC67" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="P. chokaisanus" order="Coleoptera" pageId="0" pageNumber="291" rank="species" species="chokaisanus">
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<emphasis id="7C0FE1A1374E50D6CAC239A741D31822" italics="true" pageId="0" pageNumber="291">P. chokaisanus</emphasis>
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</taxonomicName>
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. This collection record is the northernmost distribution record of the species and the first record for Akita Prefecture.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |