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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1046.57052" ID-GBIF-Dataset="348e46a4-8819-4e3c-b4b9-b5e88bee10d7" ID-PMC="PMC8238928" ID-Pensoft-Pub="1313-2970-1046-1" ID-Pensoft-UUID="DBC42C98E4DA5074B86525CF2FB2FA64" ID-PubMed="34239337" ID-ZooBank="66E61C4EFAFE45DE9145DB38199BDEC3" ModsDocID="1313-2970-1046-1" checkinTime="1624307367557" checkinUser="pensoft" docAuthor="Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott &amp; Meyer, Marc De" docDate="2021" docId="D68C114EABBE50678F1529F49480749B" docLanguage="en" docName="ZooKeys 1046: 1-141" docOrigin="ZooKeys 1046" docPubDate="2021-06-21" docSource="http://dx.doi.org/10.3897/zookeys.1046.57052" docTitle="Mesembrius madagascariensis Keiser 1971" docType="treatment" docVersion="4" id="DBC42C98E4DA5074B86525CF2FB2FA64" lastPageNumber="1" masterDocId="DBC42C98E4DA5074B86525CF2FB2FA64" masterDocTitle="Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data" masterLastPageNumber="141" masterPageNumber="1" pageNumber="1" updateTime="1668150539886" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jordaens, Kurt</mods:namePart>
<mods:affiliation>Royal Museum for Central Africa, Invertebrates Section and JEMU, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium</mods:affiliation>
<mods:nameIdentifier type="email">kurt.jordaens@africamuseum.be</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Goergen, Georg</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4496-0495</mods:nameIdentifier>
<mods:affiliation>International Institute for Tropical Agriculture (IITA), Biodiversity Centre, 08 BP 0932 Tri Postal, Cotonou, Benin</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Skevington, Jeffrey H.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1445-9870</mods:nameIdentifier>
<mods:affiliation>Canadian National Collection of Insects, Arachnids and Nematodes, Agriculture and Agri-Food Canada, K. W. Neatby Building, 960 Carling Avenue, Ottawa, ON K 1 A 0 C 6, Canada</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kelso, Scott</mods:namePart>
<mods:affiliation>Canadian National Collection of Insects, Arachnids and Nematodes, Agriculture and Agri-Food Canada, K. W. Neatby Building, 960 Carling Avenue, Ottawa, ON K 1 A 0 C 6, Canada</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Meyer, Marc De</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0755-2898</mods:nameIdentifier>
<mods:affiliation>Royal Museum for Central Africa, Invertebrates Section and JEMU, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-06-21</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1046</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>141</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1046.57052</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1046.57052</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1046-1</mods:identifier>
<mods:identifier type="ZooBank">66E61C4EFAFE45DE9145DB38199BDEC3</mods:identifier>
<mods:identifier type="Pensoft-UUID">DBC42C98E4DA5074B86525CF2FB2FA64</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="183114799" LSID="urn:lsid:plazi:treatment:D68C114EABBE50678F1529F49480749B" httpUri="http://treatment.plazi.org/id/D68C114EABBE50678F1529F49480749B" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="D68C114E-ABBE-5067-8F15-29F49480749B" authority="Keiser, 1971" authorityName="Keiser" authorityYear="1971" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius madagascariensis" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="madagascariensis">Mesembrius madagascariensis Keiser, 1971</taxonomicName>
<figureCitation captionStart="Figures 13, 14" captionStartId="F7" captionText="Figures 13, 14. Mesembrius spp., habitus, lateral view 13 M. madagascariensis Keiser (♂) 14 M. minor (Bezzi) (♂)." figureDoi="10.3897/zookeys.1046.57052.figures13-14" httpUri="https://binary.pensoft.net/fig/557468" pageId="0" pageNumber="1">Figs 13</figureCitation>
<figureCitation captionStart="Figures 33, 34" captionStartId="F17" captionText="Figures 33, 34. Mesembrius spp., habitus, lateral view 33 M. maculifer Hull (♀) 34 M. madagascariensis Keiser (♀)." figureDoi="10.3897/zookeys.1046.57052.figures33-34" httpUri="https://binary.pensoft.net/fig/557478" pageId="0" pageNumber="1">, 34</figureCitation>
<figureCitation captionStart="Figures 5257" captionStartId="F25" captionText="Figures 52 - 57. Mesembrius spp., head, frontal view 52 M. copelandi sp. nov. (♂) 53 M. cyanipennis (Bezzi) (♂) 54 M. ingratus (Loew) (♂) 55 M. longipilosus sp. nov. (♂) 56 M. madagascariensis Keiser (♂) 57 M. minor (Bezzi) (♂)." figureDoi="10.3897/zookeys.1046.57052.figures52-57" httpUri="https://binary.pensoft.net/fig/557486" pageId="0" pageNumber="1">, 56</figureCitation>
<figureCitation captionStart="Figures 8994" captionStartId="F32" captionText="Figures 89 - 94. Mesembrius spp., abdomen, dorsal view 89 M. copelandi sp. nov. (♂) 90 M. cyanipennis (Bezzi) (♂) 91 M. ingratus (Loew) (♂) 92 M. longipilosus sp. nov. (♂) 93 M. madagascariensis Keiser (♂) 94 M. minor (Bezzi) (♂)." figureDoi="10.3897/zookeys.1046.57052.figures89-94" httpUri="https://binary.pensoft.net/fig/557493" pageId="0" pageNumber="1">, 93</figureCitation>
<figureCitation captionStart="Figures 113118" captionStartId="F36" captionText="Figures 113 - 118. Mesembrius spp., abdomen, dorsal view 113 M. cyanipennis (Bezzi) (♀) 114 M. maculifer Hull (♀) 115 M. madagascariensis Keiser (♀) 116 M. minor (Bezzi) (♀) 117 M. morio (Bezzi) (♀) 118 M. platytarsis Curran syn. nov. (♀)." figureDoi="10.3897/zookeys.1046.57052.figures113-118" httpUri="https://binary.pensoft.net/fig/557497" pageId="0" pageNumber="1">, 115</figureCitation>
<figureCitation captionStart="Figures 135138" captionStartId="F41" captionText="Figures 135 - 138. Mesembrius spp., right wing 135 M. maculifer Hull (♀) 136 M. madagascariensis Keiser (♂) 137 M. minor (Bezzi) (♂) 138 M. morio (Bezzi) (♀)." figureDoi="10.3897/zookeys.1046.57052.figures135-138" httpUri="https://binary.pensoft.net/fig/557502" pageId="0" pageNumber="1">, 136</figureCitation>
<figureCitation captionStart="Figures 177180" captionStartId="F51" captionText="Figures 177 - 180. Mesembrius spp., mesoleg, dorsal view 177 M. tibialis sp. nov. (♂) 178 M. caffer (Loew) (♂) 179 M. capensis (Macquart) (♂) 180 M. madagascariensis Keiser (♂). Abbreviations: bt-basitarsus, fem-femur, tib-tibia." figureDoi="10.3897/zookeys.1046.57052.figures177-180" httpUri="https://binary.pensoft.net/fig/557511" pageId="0" pageNumber="1">, 180</figureCitation>
<figureCitation captionStart="Figures 205216" captionStartId="F58" captionText="Figures 205 - 216. Mesembrius spp., male genitalia, lateral view 205 M. arcuatus sp. nov. 206 M. caffer (Loew) (nominal morph) 207 M. caffer (Loew) (spined morph) 208 M. ctenifer Hull syn. nov. 209 M. capensis (Macquart) 210 M. chapini Curran 211 M. copelandi sp. nov. 212 M. cyanipennis (Bezzi) 213 M. ingratus (Loew) 214 M. longipilosus sp. nov. 215 M. madagascariensis Keiser 216 M. minor (Bezzi)." figureDoi="10.3897/zookeys.1046.57052.figures205-216" httpUri="https://binary.pensoft.net/fig/557518" pageId="0" pageNumber="1">, 215</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Keiser" authorityYear="1971" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius madagascariensis" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="madagascariensis">Mesembrius madagascariensis</taxonomicName>
Keiser, 1971: 261.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Keiser" authorityYear="1971" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius madagascariensis" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="madagascariensis">Mesembrius madagascariensis</taxonomicName>
-
<bibRefCitation author="Smith, KGV" editor="Crosskey, RW" journalOrPublisher="British Museum (Natural History), London" pageId="0" pageNumber="1" pagination="488 - 510" refId="B49" refString="Smith, KGV, Vockeroth, JR, 1980. 38. Family Syrphidae. In: Crosskey, RW, Ed., Catalogue of the Diptera of the Afrotropical Region. British Museum (Natural History), London: 488 - 510" title="38. Family Syrphidae." volumeTitle="Catalogue of the Diptera of the Afrotropical Region." year="1980">Smith and Vockeroth (1980)</bibRefCitation>
: 504.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Keiser" authorityYear="1971" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius madagascariensis" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">Mesembrius madagascariensis</emphasis>
</taxonomicName>
males lack an apical pile brush on the profemur and have an unmodified metatibia. The profemur is dorsally flattened and the metabasitarsus does not have a lobe as in some males of
<taxonomicName lsidName="M. simplicipes" pageId="0" pageNumber="1" rank="species" species="simplicipes">
<emphasis italics="true" pageId="0" pageNumber="1">M. simplicipes</emphasis>
</taxonomicName>
. The male of
<taxonomicName lsidName="M. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">M. madagascariensis</emphasis>
</taxonomicName>
cannot be confused with any other species by the strong difference in the pile colour and length between the proximal and distal part of the mesotibia which is long and yellow pile in the proximal 2/3 and short and black in the distal 1/3. The female of
<taxonomicName lsidName="M. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">M. madagascariensis</emphasis>
</taxonomicName>
can be distinguished from any other species (except from
<taxonomicName lsidName="M. simplicipes" pageId="0" pageNumber="1" rank="species" species="simplicipes">
<emphasis italics="true" pageId="0" pageNumber="1">M. simplicipes</emphasis>
</taxonomicName>
) by the nearly black abdomen (clearly yellow to orange and black in other species). It can be distinguished from
<taxonomicName lsidName="M. simplicipes" pageId="0" pageNumber="1" rank="species" species="simplicipes">
<emphasis italics="true" pageId="0" pageNumber="1">M. simplicipes</emphasis>
</taxonomicName>
by the pro- and mesolegs which are extensively brown and black (reddish-brown in
<taxonomicName lsidName="M. simplicipes" pageId="0" pageNumber="1" rank="species" species="simplicipes">
<emphasis italics="true" pageId="0" pageNumber="1">M. simplicipes</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Examined material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Keiser" authorityYear="1971" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius madagascariensis" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">Mesembrius madagascariensis</emphasis>
</taxonomicName>
<materialsCitation collectingDate="1957-12-13" collectorName="F. Keiser" country="Madagascar" location="Antananarivo" specimenCount="1" stateProvince="Antananarivo" typeStatus="Holotype">
Keiser:
<typeStatus>Holotype</typeStatus>
, male,
<collectingCountry name="Madagascar">Madagascar</collectingCountry>
,
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
, M,
<collectingDate value="1957-12-13">13 Dec 1957</collectingDate>
,
<collectorName>F. Keiser</collectorName>
(MNHN: type not studied; see comments).
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Keiser" authorityYear="1971" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius madagascariensis" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">Mesembrius madagascariensis</emphasis>
</taxonomicName>
<materialsCitation collectingDate="1957-12-13" country="Madagascar" location="Tananarive" specimenCount="1" typeStatus="Allotype">
Keiser:
<typeStatus>Allotype</typeStatus>
, female, &quot;ALLO // TYPUS&quot; &quot;
<collectingCountry name="Madagascar">Madagascar</collectingCountry>
. TAN. //
<location LSID="urn:lsid:plazi:treatment:D68C114EABBE50678F1529F49480749B:5706D6010FB5E7673FEE1E5C560A5D33" country="Madagascar" name="Tananarive">Tananarive</location>
//
<collectingDate value="1957-12-13">13.XII.1957</collectingDate>
// F. KEISER&quot; &quot;
<taxonomicName authorityName="Rondani" authorityYear="1857" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Mesembrius</taxonomicName>
// madagascar- // iensis&quot; [NMB].
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="1975-12-13" collectorName="F. Keiser" country="Madagascar" location="Tananarive" specimenCount="28" specimenCount-female="21" specimenCount-male="7" typeStatus="Paratypes">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">
<typeStatus>Paratypes</typeStatus>
</emphasis>
:
</emphasis>
<collectingCountry name="Madagascar">Madagascar</collectingCountry>
<specimenCount type="male">7♂♂</specimenCount>
<specimenCount type="female">21♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:D68C114EABBE50678F1529F49480749B:E88DF365A89EA3842FFA12A615E6199A" country="Madagascar" name="Tananarive">Tananarive</location>
;
<collectingDate value="1975-12-13">13 Dec 1975</collectingDate>
;
<collectorName>F. Keiser</collectorName>
leg.; NMB
</materialsCitation>
<materialsCitation collectingDate="1958-03-20" collectorName="F. Keiser" country="Madagascar" location="Ampefy" municipality="Lac Kavitaha" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount type="female">1♀</specimenCount>
;
<collectingMunicipality>Lac Kavitaha</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:D68C114EABBE50678F1529F49480749B:68CDA4BFA62AA2026A5DED5CD3BC6F23" country="Madagascar" municipality="Lac Kavitaha" name="Ampefy">Ampefy</location>
;
<collectingDate value="1958-03-20">20 Mar 1958</collectingDate>
;
<collectorName>F. Keiser</collectorName>
; NMB
</materialsCitation>
<materialsCitation collectingDate="1958-01-28" collectingDateMax="1958-01-29" collectingDateMin="1958-01-28" collectorName="F. Keiser" country="Madagascar" location="Ambalavao" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Fianarantsoa" typeStatus="paratype">
<specimenCount type="male">4♂♂</specimenCount>
<specimenCount type="female">2♀♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:D68C114EABBE50678F1529F49480749B:35B702E4BFA06D98160A7E9DCCE6E281" country="Madagascar" name="Ambalavao" stateProvince="Fianarantsoa">Ambalavao</location>
,
<collectingRegion country="Madagascar" name="Fianarantsoa">Fianarantsoa</collectingRegion>
;
<collectingDate value="1958-01-28" valueMax="1958-01-29" valueMin="1958-01-28">28-29 Jan 1958</collectingDate>
;
<collectorName>F. Keiser</collectorName>
; NMB
</materialsCitation>
<materialsCitation collectingDate="1968-02-07" collectingDateMax="1968-02-08" collectingDateMin="1968-02-07" collectorName="F. Keiser" country="Madagascar" location="Tzimbazaza Park" specimenCount="11" specimenCount-female="2" specimenCount-male="9" stateProvince="Antananarivo" typeStatus="paratype">
<specimenCount type="male">9♂♂</specimenCount>
<specimenCount type="female">2♀♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:D68C114EABBE50678F1529F49480749B:A87CEACF78989F612848642F73DCD0A7" country="Madagascar" name="Tzimbazaza Park" stateProvince="Antananarivo">Tzimbazaza Park</location>
,
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
;
<collectingDate value="1968-02-07" valueMax="1968-02-08" valueMin="1968-02-07">7-8 Feb 1968</collectingDate>
;
<collectorName>F. Keiser</collectorName>
leg.; NMB
</materialsCitation>
<materialsCitation collectingDate="1957-12-24" collectorName="B. R. Stuckenberg" country="Madagascar" county="Station Agric." location="District Anbatondrakzaka" municipality="Alaotra" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount type="female">1♀</specimenCount>
;
<collectingCounty>Station Agric.</collectingCounty>
,
<collectingMunicipality>Alaotra</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:D68C114EABBE50678F1529F49480749B:21108A7A74DC0CCE9D84EE034839A136" country="Madagascar" county="Station Agric." municipality="Alaotra" name="District Anbatondrakzaka">District Anbatondrakzaka</location>
;
<collectingDate value="1957-12-24">24 Dec 1957</collectingDate>
;
<collectorName>B.R. Stuckenberg</collectorName>
leg.; NMSA
</materialsCitation>
<materialsCitation collectingDate="1958-01-01" collectorName="B. R. Stuckenberg" country="Madagascar" location="Antananarivo" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Antananarivo" typeStatus="paratype">
<specimenCount type="male">4♂♂</specimenCount>
<specimenCount type="female">2♀♀</specimenCount>
;
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
;
<collectingDate value="1958-01-01">1 Jan 1958</collectingDate>
;
<collectorName>B.R. Stuckenberg</collectorName>
leg.; NMSA
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="1993-10-16" collectorName="M. Hauser" country="Madagascar" location="Tsimbazaza" specimenCount="1" specimenCount-female="1" stateProvince="Antananarivo">
<collectingCountry name="Madagascar">Madagascar</collectingCountry>
<specimenCount type="female">1♀</specimenCount>
;
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:D68C114EABBE50678F1529F49480749B:57F8456FFAD77C42D2F15549D3CA6922" country="Madagascar" name="Tsimbazaza" stateProvince="Antananarivo">Tsimbazaza</location>
;
<collectingDate value="1993-10-16">16 Oct 1993</collectingDate>
;
<collectorName>M. Hauser</collectorName>
leg.; CAS
</materialsCitation>
<materialsCitation collectingDate="1993-10-16" collectingDateMax="1993-10-22" collectingDateMin="1993-10-16" collectorName="C. Kassebeer" country="Madagascar" location="Tsimbazaza" specimenCount="1" specimenCount-male="1" stateProvince="Antananarivo">
<specimenCount type="male">1♂</specimenCount>
;
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:D68C114EABBE50678F1529F49480749B:77DD2BBF64FB4CB05DDF732867471BF4" country="Madagascar" name="Tsimbazaza" stateProvince="Antananarivo">Tsimbazaza</location>
;
<collectingDate value="1993-10-16" valueMax="1993-10-22" valueMin="1993-10-16">16-22 Oct 1993</collectingDate>
;
<collectorName>C. Kassebeer</collectorName>
leg.; CAS
</materialsCitation>
<materialsCitation collectingDate="1993-10-16" collectingDateMax="1993-10-22" collectingDateMin="1993-10-16" collectorName="C. Kassebeer" country="Madagascar" location="Tsimbazaza" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Antananarivo">
<specimenCount type="male">1♂</specimenCount>
<specimenCount type="female">1♀</specimenCount>
;
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:D68C114EABBE50678F1529F49480749B:51D644B9271296E4EEBBDB91EC08C4F5" country="Madagascar" name="Tsimbazaza" stateProvince="Antananarivo">Tsimbazaza</location>
;
<collectingDate value="1993-10-16" valueMax="1993-10-22" valueMin="1993-10-16">16-22 Oct 1993</collectingDate>
;
<collectorName>C. Kassebeer</collectorName>
leg.; CNC
</materialsCitation>
<materialsCitation collectingDate="1993-10-14" collectorName="C. Kassebeer" country="Madagascar" location="Tsimbazaza" specimenCount="1" specimenCount-male="1" stateProvince="Antananarivo">
<specimenCount type="male">1♂</specimenCount>
;
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:D68C114EABBE50678F1529F49480749B:A984A1FBC1512EEBB131F413D9C03299" country="Madagascar" name="Tsimbazaza" stateProvince="Antananarivo">Tsimbazaza</location>
;
<collectingDate value="1993-10-14">14 Oct 1993</collectingDate>
;
<collectorName>C. Kassebeer</collectorName>
leg.; NHMUK
</materialsCitation>
<materialsCitation collectingDate="1993-10-26" collectorName="C. Kassebeer" country="Madagascar" location="Tsimbazaza" specimenCount="1" specimenCount-female="1" stateProvince="Antananarivo">
<specimenCount type="female">1♀</specimenCount>
;
<collectingRegion country="Madagascar" name="Antananarivo">Antananarivo</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:D68C114EABBE50678F1529F49480749B:53151EF5DB0839BE4AF77F0F0925BAC7" country="Madagascar" name="Tsimbazaza" stateProvince="Antananarivo">Tsimbazaza</location>
;
<collectingDate value="1993-10-26">26 Oct 1993</collectingDate>
;
<collectorName>C. Kassebeer</collectorName>
leg.; NHMUK
</materialsCitation>
<materialsCitation collectingDate="2012-07-06" collectingDateMax="2012-07-16" collectingDateMin="2012-07-06" collectorName="M. Irwin, R. Harin'Hala" country="Madagascar" location="Andrefana" municipality="Atsimo" specimenCount="1" specimenCount-female="1">
<specimenCount type="female">1♀</specimenCount>
;
<collectingMunicipality>Atsimo</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:D68C114EABBE50678F1529F49480749B:E48C4C004073DCC69A659261D55C764C" country="Madagascar" municipality="Atsimo" name="Andrefana">Andrefana</location>
;
<collectingDate value="2012-07-06" valueMax="2012-07-16" valueMin="2012-07-06">6-16 Jul 2012</collectingDate>
;
<collectorName>M. Irwin</collectorName>
and
<collectorName>
R.
<normalizedToken originalValue="HarinHala">Harin'Hala</normalizedToken>
</collectorName>
leg.; CAS
</materialsCitation>
<materialsCitation collectingDate="2012-05-31" collectorName="M. Irwin, R. Harin'Hala" country="Madagascar" location="Andrefana" municipality="Atsimo" specimenCount="1" specimenCount-female="1">
<specimenCount type="female">1♀</specimenCount>
;
<collectingMunicipality>Atsimo</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:D68C114EABBE50678F1529F49480749B:B3D1D3774B2D484EFFF755415A70BF83" country="Madagascar" municipality="Atsimo" name="Andrefana">Andrefana</location>
;
<collectingDate value="2012-05-31">31 May 2012</collectingDate>
;
<collectorName>M. Irwin</collectorName>
and
<collectorName>
R.
<normalizedToken originalValue="HarinHala">Harin'Hala</normalizedToken>
</collectorName>
leg.; CAS
</materialsCitation>
<materialsCitation collectingDate="1957-12-13" collectorName="F. Keiser" country="Madagascar" location="Tananarive" specimenCount="1" specimenCount-female="1">
<specimenCount type="female">1♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:D68C114EABBE50678F1529F49480749B:E0197B360E3B8C13C4E230E7ABD56635" country="Madagascar" name="Tananarive">Tananarive</location>
;
<collectingDate value="1957-12-13">13 Dec 1957</collectingDate>
;
<collectorName>F. Keiser</collectorName>
leg.; NMB
</materialsCitation>
<materialsCitation collectingDate="1957-12-15" collectorName="F. Keiser" country="Madagascar" location="Tananarive" specimenCount="1" specimenCount-female="1">
<specimenCount type="female">1♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:D68C114EABBE50678F1529F49480749B:0BA8D23D8ED96A1DDBFE47ADC509FB34" country="Madagascar" name="Tananarive">Tananarive</location>
;
<collectingDate value="1957-12-15">15 Dec 1957</collectingDate>
;
<collectorName>F. Keiser</collectorName>
leg.; NMB
</materialsCitation>
<materialsCitation collectingDate="1957-12-29" collectorName="F. Keiser" country="Madagascar" location="Tananarive" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount type="male">1♂</specimenCount>
<specimenCount type="female">1♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:D68C114EABBE50678F1529F49480749B:4D483B5D7BC06973C566D967ECD915B2" country="Madagascar" name="Tananarive">Tananarive</location>
;
<collectingDate value="1957-12-29">29 Dec 1957</collectingDate>
;
<collectorName>F. Keiser</collectorName>
leg.; NMB
</materialsCitation>
<materialsCitation collectingDate="1958-04-17" collectorName="F. Keiser" country="Madagascar" location="Tananarive" specimenCount="1" specimenCount-female="1">
<specimenCount type="female">1♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:D68C114EABBE50678F1529F49480749B:729751462442ED3DCFFFE16DEEBE3110" country="Madagascar" name="Tananarive">Tananarive</location>
;
<collectingDate value="1958-04-17">17 Apr 1958</collectingDate>
;
<collectorName>F. Keiser</collectorName>
leg.; NMB
</materialsCitation>
<materialsCitation collectingDate="1968-04-26" collectingDateMax="1968-04-30" collectingDateMin="1968-04-26" collectorName="K. M. Guichard" country="Madagascar" location="Tananarive" specimenCount="2" specimenCount-female="2">
<specimenCount type="female">2♀♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:D68C114EABBE50678F1529F49480749B:DA445541C9F534DEDCB8369C4CA479A6" country="Madagascar" name="Tananarive">Tananarive</location>
;
<collectingDate value="1968-04-26" valueMax="1968-04-30" valueMin="1968-04-26">26-30 Apr 1968</collectingDate>
;
<collectorName>K.M. Guichard</collectorName>
leg.; NHMUK
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Re-description male</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
(Fig.
<figureCitation captionStart="Figures 13, 14" captionStartId="F7" captionText="Figures 13, 14. Mesembrius spp., habitus, lateral view 13 M. madagascariensis Keiser (♂) 14 M. minor (Bezzi) (♂)." figureDoi="10.3897/zookeys.1046.57052.figures13-14" httpUri="https://binary.pensoft.net/fig/557468" pageId="0" pageNumber="1">13</figureCitation>
).
</emphasis>
Body length: 13.7-14.6 mm. Wing length: 9.6-10.5 mm.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
(Fig.
<figureCitation captionStart="Figures 5257" captionStartId="F25" captionText="Figures 52 - 57. Mesembrius spp., head, frontal view 52 M. copelandi sp. nov. (♂) 53 M. cyanipennis (Bezzi) (♂) 54 M. ingratus (Loew) (♂) 55 M. longipilosus sp. nov. (♂) 56 M. madagascariensis Keiser (♂) 57 M. minor (Bezzi) (♂)." figureDoi="10.3897/zookeys.1046.57052.figures52-57" httpUri="https://binary.pensoft.net/fig/557486" pageId="0" pageNumber="1">56</figureCitation>
). Eyes bare; holoptic, length of eye contiguity approx. length of ocellar triangle. Face white with dark medial vitta; white pilose; white pollinose. Vertical triangle black pilose on ventral half and at ocellar triangle, yellow pilose on dorsal half; yellow pollinose until just before anterior ocellus. Lateral ocellus touching eye margin. Occiput yellow; yellow pilose; yellow and white pollinose. Frontal prominence shiny black; black pilose. Antenna, scape and pedicel reddish-brown; postpedicel black; antennal arista reddish-brown.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Thorax.</emphasis>
Scutum black; dorsally with a pair of well-demarcated yellow vittae and a faint yellow medial line; vittae and medial line become faint posteriorly; yellow vague lateral vitta; yellow pilose. Scutellum uniformly yellow-brown; yellow pilose.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Legs.</emphasis>
All legs dark chocolate-brown to black, except for pro- and mesotibiae and basitarsi which are reddish-brown. Proleg: Femur dark chocolate-brown to black; without apical pile brush; pile inconspicuous, except for long yellow pile on the anterodorsal side which, dorsally, is bordered with a row of largely spaced black pile. Tibia reddish-brown; yellow pilose with some short black pile on distal end dorsally. Basitarsus orange-brown; black pilose with some longer yellow pile posteriorly. Other tarsi black; black pilose, with some longer yellow pile posteriorly, except on the two most distal tarsi. Mesoleg (Fig.
<figureCitation captionStart="Figures 177180" captionStartId="F51" captionText="Figures 177 - 180. Mesembrius spp., mesoleg, dorsal view 177 M. tibialis sp. nov. (♂) 178 M. caffer (Loew) (♂) 179 M. capensis (Macquart) (♂) 180 M. madagascariensis Keiser (♂). Abbreviations: bt-basitarsus, fem-femur, tib-tibia." figureDoi="10.3897/zookeys.1046.57052.figures177-180" httpUri="https://binary.pensoft.net/fig/557511" pageId="0" pageNumber="1">180</figureCitation>
): Femur with predominantly long yellow pile on ventral and posteroventral side, with shorter black pile on anterodorsal side. Tibia reddish-brown; proximal half with a comb of long, curved yellow pile on proximal half; with much shorter, yellow and black pile on distal half. Basitarsus orange-brown; black pilose with some longer yellow pile posteriorly. Other tarsi black; black pilose, with some longer yellow pile posteriorly, except on the two most distal tarsi. Metaleg: Femur dark chocolate-brown to black; with long, thin yellow pile, especially anteriorly; ventrally with less black pile. Tibia dark chocolate-brown to black; unmodified; with short, black pile throughout and short, yellow pile on anteroproximal half. Basitarsus and second tarsomere orange-brown, other tarsi black; black pilose.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Wing</emphasis>
(Fig.
<figureCitation captionStart="Figures 135138" captionStartId="F41" captionText="Figures 135 - 138. Mesembrius spp., right wing 135 M. maculifer Hull (♀) 136 M. madagascariensis Keiser (♂) 137 M. minor (Bezzi) (♂) 138 M. morio (Bezzi) (♀)." figureDoi="10.3897/zookeys.1046.57052.figures135-138" httpUri="https://binary.pensoft.net/fig/557502" pageId="0" pageNumber="1">136</figureCitation>
). Entire wing uniformly microtrichose.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
(Fig.
<figureCitation captionStart="Figures 8994" captionStartId="F32" captionText="Figures 89 - 94. Mesembrius spp., abdomen, dorsal view 89 M. copelandi sp. nov. (♂) 90 M. cyanipennis (Bezzi) (♂) 91 M. ingratus (Loew) (♂) 92 M. longipilosus sp. nov. (♂) 93 M. madagascariensis Keiser (♂) 94 M. minor (Bezzi) (♂)." figureDoi="10.3897/zookeys.1046.57052.figures89-94" httpUri="https://binary.pensoft.net/fig/557493" pageId="0" pageNumber="1">93</figureCitation>
). Tergite II with a pair of yellow-orange, rounded maculae; black marking broadly hourglass-shaped; yellow pilose, except for posterior half of posterior black marking where pile is black; with a stretch of yellow pollinosity on anterior part of black marking. Tergite III with a pair of small, anterolateral maculae; yellow-white pilose; white pollinose on posterior half. Tergite IV dark chocolate-brown; yellow-white pilose; white pollinose.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Genitalia</emphasis>
(Fig.
<figureCitation captionStart="Figures 205216" captionStartId="F58" captionText="Figures 205 - 216. Mesembrius spp., male genitalia, lateral view 205 M. arcuatus sp. nov. 206 M. caffer (Loew) (nominal morph) 207 M. caffer (Loew) (spined morph) 208 M. ctenifer Hull syn. nov. 209 M. capensis (Macquart) 210 M. chapini Curran 211 M. copelandi sp. nov. 212 M. cyanipennis (Bezzi) 213 M. ingratus (Loew) 214 M. longipilosus sp. nov. 215 M. madagascariensis Keiser 216 M. minor (Bezzi)." figureDoi="10.3897/zookeys.1046.57052.figures205-216" httpUri="https://binary.pensoft.net/fig/557518" pageId="0" pageNumber="1">215</figureCitation>
). Epandrium: Dorsal lobe of surstylus club-shaped; distal end densely covered with black spines; proximal half (
<normalizedToken originalValue="stalk">'stalk'</normalizedToken>
) long yellow pilose dorsally and with some shorter, thicker black pile laterally. Ventral lobe of surstylus with one very long and thick black setula distally.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Re-description female</paragraph>
<paragraph pageId="0" pageNumber="1">
(Fig.
<figureCitation captionStart="Figures 33, 34" captionStartId="F17" captionText="Figures 33, 34. Mesembrius spp., habitus, lateral view 33 M. maculifer Hull (♀) 34 M. madagascariensis Keiser (♀)." figureDoi="10.3897/zookeys.1046.57052.figures33-34" httpUri="https://binary.pensoft.net/fig/557478" pageId="0" pageNumber="1">34</figureCitation>
). Body length: 12.5-13.5 mm. Wing length: 8.7-9.3 mm.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
Eyes bare; dichoptic. Face white with dark medial vitta; white pilose, white pollinose. Frons black, but strongly white pollinose on ventral 4/5; black pilose on ocellar triangle and just ventral of ocellar triangle; white pilose. Distance between lateral ocellus and eye margin approx.
<normalizedToken originalValue="1½">11/2</normalizedToken>
x width of ocellus. Occiput yellow-white; yellow-white pilose; yellow-white pollinose. Frontal prominence shiny black, orange-brown at distal end. Antenna, scape and pedicel orange-brown; postpedicel black, white pollinose; antennal arista yellow-orange.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Thorax.</emphasis>
Scutum dark brown to black with a pair of dorsolateral white-grey pollinose vittae; with lateral grey pollinose vague vitta; yellow and black pilose. Scutellum yellow-brown; yellow and black pilose.
</paragraph>
<caption doi="10.3897/zookeys.1046.57052.figures46-51" httpUri="https://binary.pensoft.net/fig/557485" pageId="0" pageNumber="1" start="Figures 4651" startId="F24">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 46-51.</emphasis>
<taxonomicName authorityName="Rondani" authorityYear="1857" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Mesembrius</emphasis>
</taxonomicName>
spp., head, frontal view
<emphasis bold="true" pageId="0" pageNumber="1">46</emphasis>
<taxonomicName lsidName="M. arcuatus" pageId="0" pageNumber="1" rank="species" species="arcuatus">
<emphasis italics="true" pageId="0" pageNumber="1">M. arcuatus</emphasis>
</taxonomicName>
sp. nov. (♂)
<emphasis bold="true" pageId="0" pageNumber="1">47</emphasis>
<taxonomicName lsidName="M. caffer" pageId="0" pageNumber="1" rank="species" species="caffer">
<emphasis italics="true" pageId="0" pageNumber="1">M. caffer</emphasis>
</taxonomicName>
(Loew) (nominal morph) (♂)
<emphasis bold="true" pageId="0" pageNumber="1">48</emphasis>
<taxonomicName lsidName="M. caffer" pageId="0" pageNumber="1" rank="species" species="caffer">
<emphasis italics="true" pageId="0" pageNumber="1">M. caffer</emphasis>
</taxonomicName>
(Loew) (spined morph) (♂)
<emphasis bold="true" pageId="0" pageNumber="1">49</emphasis>
<taxonomicName lsidName="M. ctenifer" pageId="0" pageNumber="1" rank="species" species="ctenifer">
<emphasis italics="true" pageId="0" pageNumber="1">M. ctenifer</emphasis>
</taxonomicName>
Hull syn. nov. (♂)
<emphasis bold="true" pageId="0" pageNumber="1">50</emphasis>
<taxonomicName lsidName="M. capensis" pageId="0" pageNumber="1" rank="species" species="capensis">
<emphasis italics="true" pageId="0" pageNumber="1">M. capensis</emphasis>
</taxonomicName>
(Macquart) (♂)
<emphasis bold="true" pageId="0" pageNumber="1">51</emphasis>
<taxonomicName lsidName="M. chapini" pageId="0" pageNumber="1" rank="species" species="chapini">
<emphasis italics="true" pageId="0" pageNumber="1">M. chapini</emphasis>
</taxonomicName>
Curran (♂).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Legs.</emphasis>
Pro- and mesoleg: Femur black, distal end reddish-brown; short black and longer yellow pilose. Tibia reddish-brown; yellow and black pilose dorsally; yellow pilose ventrally. Basitarsus reddish-brown; black pilose dorsally; orange-yellow pilose ventrally. Other tarsi black; black pilose dorsally, orange-yellow pilose ventrally. Metaleg: Femur black, distal 1/5 reddish-brown; yellow-white pilose, with scarce very short black pile ventrally. Tibia reddish-brown; white pilose, short black pilose posteriorly. Basitarsus dark reddish-brown; black pilose dorsally, yellow and black pilose ventrally. Other tarsi black; predominantly black pilose dorsally, densely yellow-orange pilose ventrally.
</paragraph>
<caption doi="10.3897/zookeys.1046.57052.figures52-57" httpUri="https://binary.pensoft.net/fig/557486" pageId="0" pageNumber="1" start="Figures 5257" startId="F25">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 52-57.</emphasis>
<taxonomicName authorityName="Rondani" authorityYear="1857" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Mesembrius</emphasis>
</taxonomicName>
spp., head, frontal view
<emphasis bold="true" pageId="0" pageNumber="1">52</emphasis>
<taxonomicName lsidName="M. copelandi" pageId="0" pageNumber="1" rank="species" species="copelandi">
<emphasis italics="true" pageId="0" pageNumber="1">M. copelandi</emphasis>
</taxonomicName>
sp. nov. (♂)
<emphasis bold="true" pageId="0" pageNumber="1">53</emphasis>
<taxonomicName lsidName="M. cyanipennis" pageId="0" pageNumber="1" rank="species" species="cyanipennis">
<emphasis italics="true" pageId="0" pageNumber="1">M. cyanipennis</emphasis>
</taxonomicName>
(Bezzi) (♂)
<emphasis bold="true" pageId="0" pageNumber="1">54</emphasis>
<taxonomicName lsidName="M. ingratus" pageId="0" pageNumber="1" rank="species" species="ingratus">
<emphasis italics="true" pageId="0" pageNumber="1">M. ingratus</emphasis>
</taxonomicName>
(Loew) (♂)
<emphasis bold="true" pageId="0" pageNumber="1">55</emphasis>
<taxonomicName lsidName="M. longipilosus" pageId="0" pageNumber="1" rank="species" species="longipilosus">
<emphasis italics="true" pageId="0" pageNumber="1">M. longipilosus</emphasis>
</taxonomicName>
sp. nov. (♂)
<emphasis bold="true" pageId="0" pageNumber="1">56</emphasis>
<taxonomicName lsidName="M. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">M. madagascariensis</emphasis>
</taxonomicName>
Keiser (♂)
<emphasis bold="true" pageId="0" pageNumber="1">57</emphasis>
<taxonomicName lsidName="M. minor" pageId="0" pageNumber="1" rank="species" species="minor">
<emphasis italics="true" pageId="0" pageNumber="1">M. minor</emphasis>
</taxonomicName>
(Bezzi) (♂).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Wing.</emphasis>
Entire wing uniformly microtrichose.
</paragraph>
<caption doi="10.3897/zookeys.1046.57052.figures58-63" httpUri="https://binary.pensoft.net/fig/557487" pageId="0" pageNumber="1" start="Figures 5863" startId="F26">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 58-63.</emphasis>
<taxonomicName authorityName="Rondani" authorityYear="1857" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Mesembrius</emphasis>
</taxonomicName>
spp., head, frontal view
<emphasis bold="true" pageId="0" pageNumber="1">58</emphasis>
<taxonomicName lsidName="M. nigriceps" pageId="0" pageNumber="1" rank="species" species="nigriceps">
<emphasis italics="true" pageId="0" pageNumber="1">M. nigriceps</emphasis>
</taxonomicName>
Curran (♂)
<emphasis bold="true" pageId="0" pageNumber="1">59</emphasis>
<taxonomicName lsidName="M. perforatus" pageId="0" pageNumber="1" rank="species" species="perforatus">
<emphasis italics="true" pageId="0" pageNumber="1">M. perforatus</emphasis>
</taxonomicName>
(Speiser) (♂)
<emphasis bold="true" pageId="0" pageNumber="1">60</emphasis>
<taxonomicName lsidName="M. platytarsis" pageId="0" pageNumber="1" rank="species" species="platytarsis">
<emphasis italics="true" pageId="0" pageNumber="1">M. platytarsis</emphasis>
</taxonomicName>
Curran syn. nov. (♂)
<emphasis bold="true" pageId="0" pageNumber="1">61</emphasis>
<taxonomicName lsidName="M. regulus" pageId="0" pageNumber="1" rank="species" species="regulus">
<emphasis italics="true" pageId="0" pageNumber="1">M. regulus</emphasis>
</taxonomicName>
(Hull) (♂)
<emphasis bold="true" pageId="0" pageNumber="1">62</emphasis>
<taxonomicName lsidName="M. rex" pageId="0" pageNumber="1" rank="species" species="rex">
<emphasis italics="true" pageId="0" pageNumber="1">M. rex</emphasis>
</taxonomicName>
Curran (♂)
<emphasis bold="true" pageId="0" pageNumber="1">63</emphasis>
<taxonomicName lsidName="M. senegalensis" pageId="0" pageNumber="1" rank="species" species="senegalensis">
<emphasis italics="true" pageId="0" pageNumber="1">M. senegalensis</emphasis>
</taxonomicName>
(Macquart) (♂).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
(Fig.
<figureCitation captionStart="Figures 113118" captionStartId="F36" captionText="Figures 113 - 118. Mesembrius spp., abdomen, dorsal view 113 M. cyanipennis (Bezzi) (♀) 114 M. maculifer Hull (♀) 115 M. madagascariensis Keiser (♀) 116 M. minor (Bezzi) (♀) 117 M. morio (Bezzi) (♀) 118 M. platytarsis Curran syn. nov. (♀)." figureDoi="10.3897/zookeys.1046.57052.figures113-118" httpUri="https://binary.pensoft.net/fig/557497" pageId="0" pageNumber="1">115</figureCitation>
). Dark brown to black; largely white pollinose, except for anterior border and medial area of tergite II and posterior half of tergite V; short black and white pilose, except for non-pollinose areas with white pilose only.
</paragraph>
<caption doi="10.3897/zookeys.1046.57052.figures64-69" httpUri="https://binary.pensoft.net/fig/557488" pageId="0" pageNumber="1" start="Figures 6469" startId="F27">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 64-69.</emphasis>
<taxonomicName authorityName="Rondani" authorityYear="1857" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Mesembrius</emphasis>
</taxonomicName>
spp., head, frontal view
<emphasis bold="true" pageId="0" pageNumber="1">64</emphasis>
<taxonomicName lsidName="M. simplicipes" pageId="0" pageNumber="1" rank="species" species="simplicipes">
<emphasis italics="true" pageId="0" pageNumber="1">M. simplicipes</emphasis>
</taxonomicName>
Curran (♂)
<emphasis bold="true" pageId="0" pageNumber="1">65</emphasis>
<taxonomicName lsidName="M. strigilatus" pageId="0" pageNumber="1" rank="species" species="strigilatus">
<emphasis italics="true" pageId="0" pageNumber="1">M. strigilatus</emphasis>
</taxonomicName>
(Bezzi) (♂)
<emphasis bold="true" pageId="0" pageNumber="1">66</emphasis>
<taxonomicName lsidName="M. sulcus" pageId="0" pageNumber="1" rank="species" species="sulcus">
<emphasis italics="true" pageId="0" pageNumber="1">M. sulcus</emphasis>
</taxonomicName>
sp. nov. (♂)
<emphasis bold="true" pageId="0" pageNumber="1">67</emphasis>
<taxonomicName lsidName="M. tarsatus" pageId="0" pageNumber="1" rank="species" species="tarsatus">
<emphasis italics="true" pageId="0" pageNumber="1">M. tarsatus</emphasis>
</taxonomicName>
(Bigot) (♂)
<emphasis bold="true" pageId="0" pageNumber="1">68</emphasis>
<taxonomicName lsidName="M. tibialis" pageId="0" pageNumber="1" rank="species" species="tibialis">
<emphasis italics="true" pageId="0" pageNumber="1">M. tibialis</emphasis>
</taxonomicName>
sp. nov. (♂)
<emphasis bold="true" pageId="0" pageNumber="1">69</emphasis>
<taxonomicName lsidName="M. vockerothi" pageId="0" pageNumber="1" rank="species" species="vockerothi">
<emphasis italics="true" pageId="0" pageNumber="1">M. vockerothi</emphasis>
</taxonomicName>
sp. nov. (♂).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">Madagascar.</paragraph>
<caption doi="10.3897/zookeys.1046.57052.figures70-74" httpUri="https://binary.pensoft.net/fig/557489" pageId="0" pageNumber="1" start="Figures 7074" startId="F28">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 70-74.</emphasis>
<taxonomicName authorityName="Rondani" authorityYear="1857" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Mesembrius</emphasis>
</taxonomicName>
spp., head, frontal view
<emphasis bold="true" pageId="0" pageNumber="1">70</emphasis>
<taxonomicName lsidName="M. capensis" pageId="0" pageNumber="1" rank="species" species="capensis">
<emphasis italics="true" pageId="0" pageNumber="1">M. capensis</emphasis>
</taxonomicName>
(Macquart) (♀)
<emphasis bold="true" pageId="0" pageNumber="1">71</emphasis>
<taxonomicName lsidName="M. cyanipennis" pageId="0" pageNumber="1" rank="species" species="cyanipennis">
<emphasis italics="true" pageId="0" pageNumber="1">M. cyanipennis</emphasis>
</taxonomicName>
(Bezzi) (♀)
<emphasis bold="true" pageId="0" pageNumber="1">72</emphasis>
<taxonomicName lsidName="M. chapini" pageId="0" pageNumber="1" rank="species" species="chapini">
<emphasis italics="true" pageId="0" pageNumber="1">M. chapini</emphasis>
</taxonomicName>
Curran (♀)
<emphasis bold="true" pageId="0" pageNumber="1">73</emphasis>
<taxonomicName lsidName="M. maculifer" pageId="0" pageNumber="1" rank="species" species="maculifer">
<emphasis italics="true" pageId="0" pageNumber="1">M. maculifer</emphasis>
</taxonomicName>
Hull (♀)
<emphasis bold="true" pageId="0" pageNumber="1">74</emphasis>
<taxonomicName lsidName="M. morio" pageId="0" pageNumber="1" rank="species" species="morio">
<emphasis italics="true" pageId="0" pageNumber="1">M. morio</emphasis>
</taxonomicName>
(Bezzi) (♀).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="comments">
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph pageId="0" pageNumber="1">
The type series comprises more than 50 specimens of both sexes collected from a dozen of sites from the central and eastern domains of Madagascar (
<bibRefCitation author="Keiser, F" journalOrPublisher="Verhandlungen der Naturforschenden Gesellschaft in Basel" pageId="0" pageNumber="1" pagination="223 - 318" refId="B34" refString="Keiser, F, 1971. Syrphidae von Madagascar (Dipt.). Verhandlungen der Naturforschenden Gesellschaft in Basel 81: 223 - 318" title="Syrphidae von Madagascar (Dipt.)." volume="81" year="1971">Keiser 1971</bibRefCitation>
). The holotype of the species should be in the collection at MNHN (
<bibRefCitation author="Keiser, F" journalOrPublisher="Verhandlungen der Naturforschenden Gesellschaft in Basel" pageId="0" pageNumber="1" pagination="223 - 318" refId="B34" refString="Keiser, F, 1971. Syrphidae von Madagascar (Dipt.). Verhandlungen der Naturforschenden Gesellschaft in Basel 81: 223 - 318" title="Syrphidae von Madagascar (Dipt.)." volume="81" year="1971">Keiser 1971</bibRefCitation>
), but neither could we trace the holotype in the MNHN collection, nor is it listed on the MNHN entomology collection webpages.
</paragraph>
<caption doi="10.3897/zookeys.1046.57052.figures75-78" httpUri="https://binary.pensoft.net/fig/557490" pageId="0" pageNumber="1" start="Figures 7578" startId="F29">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 75-78.</emphasis>
<taxonomicName authorityName="Rondani" authorityYear="1857" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Mesembrius</emphasis>
</taxonomicName>
spp., head, frontal view
<emphasis bold="true" pageId="0" pageNumber="1">75</emphasis>
<taxonomicName lsidName="M. platytarsis" pageId="0" pageNumber="1" rank="species" species="platytarsis">
<emphasis italics="true" pageId="0" pageNumber="1">M. platytarsis</emphasis>
</taxonomicName>
Curran syn. nov. (♀)
<emphasis bold="true" pageId="0" pageNumber="1">76</emphasis>
<taxonomicName lsidName="M. regulus" pageId="0" pageNumber="1" rank="species" species="regulus">
<emphasis italics="true" pageId="0" pageNumber="1">M. regulus</emphasis>
</taxonomicName>
(Hull) (♂)
<emphasis bold="true" pageId="0" pageNumber="1">77</emphasis>
<taxonomicName lsidName="M. rex" pageId="0" pageNumber="1" rank="species" species="rex">
<emphasis italics="true" pageId="0" pageNumber="1">M. rex</emphasis>
</taxonomicName>
Curran (♂)
<emphasis bold="true" pageId="0" pageNumber="1">78</emphasis>
<taxonomicName lsidName="M. senegalensis" pageId="0" pageNumber="1" rank="species" species="senegalensis">
<emphasis italics="true" pageId="0" pageNumber="1">M. senegalensis</emphasis>
</taxonomicName>
(Macquart) (♂).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1046.57052.figures79-82" httpUri="https://binary.pensoft.net/fig/557491" pageId="0" pageNumber="1" start="Figures 7982" startId="F30">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 79-82.</emphasis>
<taxonomicName authorityName="Rondani" authorityYear="1857" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Mesembrius</emphasis>
</taxonomicName>
spp., head, frontal view
<emphasis bold="true" pageId="0" pageNumber="1">79</emphasis>
<taxonomicName lsidName="M. simplicipes" pageId="0" pageNumber="1" rank="species" species="simplicipes">
<emphasis italics="true" pageId="0" pageNumber="1">M. simplicipes</emphasis>
</taxonomicName>
Curran (♂)
<emphasis bold="true" pageId="0" pageNumber="1">80</emphasis>
<taxonomicName lsidName="M. sulcus" pageId="0" pageNumber="1" rank="species" species="sulcus">
<emphasis italics="true" pageId="0" pageNumber="1">M. sulcus</emphasis>
</taxonomicName>
sp. nov. (♀)
<emphasis bold="true" pageId="0" pageNumber="1">81</emphasis>
<taxonomicName lsidName="M. tarsatus" pageId="0" pageNumber="1" rank="species" species="tarsatus">
<emphasis italics="true" pageId="0" pageNumber="1">M. tarsatus</emphasis>
</taxonomicName>
(Bigot) (♀)
<emphasis bold="true" pageId="0" pageNumber="1">82</emphasis>
<taxonomicName lsidName="M. vockerothi" pageId="0" pageNumber="1" rank="species" species="vockerothi">
<emphasis italics="true" pageId="0" pageNumber="1">M. vockerothi</emphasis>
</taxonomicName>
sp. nov. (♀).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>