412 lines
55 KiB
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412 lines
55 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1167.102840" ID-Pensoft-Pub="1313-2970-1167-89" ID-Pensoft-UUID="650453C68D1958238E34CEEB5261F02B" ID-ZooBank="6E397DD67B6B4837BF0C5C84720DCF55" ModsDocID="1313-2970-1167-89" checkinTime="1686777970495" checkinUser="pensoft" docAuthor="Jochum, Adrienne, Bochud, Estee, Haberthuer, David, Lee, Harry G., Hlushchuk, Ruslan & Portell, Roger W." docDate="2023" docId="A5A9E77D85925E979C4DA11BDD6722A1" docLanguage="en" docName="ZooKeys 1167: 89-107" docOrigin="ZooKeys 1167" docPubDate="2023-06-14" docSource="http://dx.doi.org/10.3897/zookeys.1167.102840" docTitle="Carychium floridanum G. H. Clapp 1918" docType="treatment" docVersion="1" id="650453C68D1958238E34CEEB5261F02B" lastPageNumber="89" masterDocId="650453C68D1958238E34CEEB5261F02B" masterDocTitle="Fossil Carychiidae (Eupulmonata, Ellobioidea) from the Lower Pleistocene Nashua Formation of Florida, with the description of a new species" masterLastPageNumber="107" masterPageNumber="89" pageNumber="89" updateTime="1686777970495" updateUser="pensoft">
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<mods:title id="7029ABEBC03A53CA92E886BD6A895A80">Fossil Carychiidae (Eupulmonata, Ellobioidea) from the Lower Pleistocene Nashua Formation of Florida, with the description of a new species</mods:title>
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<mods:namePart id="65731E7413D2216F818AE3A10B3FF118">Jochum, Adrienne</mods:namePart>
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<mods:nameIdentifier id="5FF2EE896CC09B2943515F1EA48A363C" type="ORCID">https://orcid.org/0000-0002-6624-6412</mods:nameIdentifier>
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<mods:affiliation id="996F28B1535BF1436DBAA496895E5235">Natural History Museum Bern, Bernastrasse 15, CH- 3005 Bern, Switzerland & Institute of Ecology and Evolution, University of Bern, Baltzerstrasse 6, 3012 Bern, Switzerland & Senckenberg Research Institute and Natural History Museum, Senckenberganlage 25, 60325 Frankfurt am Main, Germany</mods:affiliation>
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<mods:nameIdentifier id="C96E4E8A876A84DB32EFB903D60A68DD" type="email">adrienne.jochum@gmail.com</mods:nameIdentifier>
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<mods:namePart id="C618F14E180A3F1DD3C14647E6F871C6">Bochud, Estee</mods:namePart>
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<mods:affiliation id="F747710DEE9A88F8734BD3BFBCB8A76E">Natural History Museum Bern, Bernastrasse 15, CH- 3005 Bern, Switzerland & Institute of Ecology and Evolution, University of Bern, Baltzerstrasse 6, 3012 Bern, Switzerland</mods:affiliation>
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<mods:namePart id="1B7F7531CAE8C9725429E3F76F88E7FC">Haberthuer, David</mods:namePart>
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<mods:nameIdentifier id="376F9999E5E63B3ED869180649D93636" type="ORCID">https://orcid.org/0000-0003-3388-9187</mods:nameIdentifier>
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<mods:affiliation id="F14D5CE0802EF74660C52A6E34BEE2F7">Institute of Ecology and Evolution, University of Bern, Baltzerstrasse 6, 3012 Bern, Switzerland</mods:affiliation>
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<mods:namePart id="78AFBF47C8B7D608EF415A958A29004B">Lee, Harry G.</mods:namePart>
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<mods:affiliation id="60C9C063E333F933D89560BA0C54655E">Institute of Anatomy, University of Bern, Baltzerstrasse 2, CH- 3012 Bern, Switzerland</mods:affiliation>
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<mods:namePart id="ECB778C2E93F26D4B4658396F3C5D465">Hlushchuk, Ruslan</mods:namePart>
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<mods:affiliation id="8F8963AF5C1BFD8A9A2254DF2B5AA4AC">Institute of Ecology and Evolution, University of Bern, Baltzerstrasse 6, 3012 Bern, Switzerland</mods:affiliation>
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<mods:namePart id="1F63BB5C561B82F4F6D4BDFDBE793F33">Portell, Roger W.</mods:namePart>
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<mods:affiliation id="0C2399E2E8901311BE15D5B762B976C3">Institute of Anatomy, University of Bern, Baltzerstrasse 2, CH- 3012 Bern, Switzerland</mods:affiliation>
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<treatment id="A5A9E77D85925E979C4DA11BDD6722A1" LSID="urn:lsid:plazi:treatment:A5A9E77D85925E979C4DA11BDD6722A1" httpUri="http://treatment.plazi.org/id/A5A9E77D85925E979C4DA11BDD6722A1" lastPageNumber="89" pageId="0" pageNumber="89">
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<subSubSection id="840D3562885256144DAE8901AE6CED9C" pageId="0" pageNumber="89" type="nomenclature">
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<paragraph id="B20C3CC96CE38B8E340AEEEF5F5C3049" pageId="0" pageNumber="89">
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<taxonomicName id="D01BE00818FB2CEEF1F74A6C208D66A6" ID-CoL="RHYJ" ID-ENA="904307" LSID="A5A9E77D-8592-5E97-9C4D-A11BDD6722A1" authority="G. H. Clapp, 1918" authorityName="G. H. Clapp" authorityYear="1918" class="Gastropoda" family="Ellobiidae" genus="Carychium" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Carychium floridanum" order="Ellobiida" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="floridanum">Carychium floridanum G. H. Clapp, 1918</taxonomicName>
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</paragraph>
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<subSubSection id="8925D8224B4A4378F546E21620297BA6" pageId="0" pageNumber="89" type="description">
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<paragraph id="1AF5C49B113652F7925AFEC2786BAAF1" pageId="0" pageNumber="89">
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<figureCitation id="72AC9B9152F23D178BE8E4A80E1CB779" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Fossil Carychium floridanum G. H. Clapp, 1918 (NMBE 577015 - 577016). Light microscopic images showing different perspectives A-G (NMBE 577015) H-N (NMBE 577016). Peristome thickly callused with apertural barriers. Grey and orange colours indicate taphonomic and sedimentary processes, repectively. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1167.102840.figure4" httpUri="https://binary.pensoft.net/fig/863995" pageId="0" pageNumber="89">Figs 4</figureCitation>
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<figureCitation id="9D4F0C0341DDCCB7184580E5ED6AAA7F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. 3 D visualizations of X-ray micro-CT data of fossil Carychium floridanum G. H. Clapp, 1918 (NMBE 577015) A apertural view showing extent of lamellar fold B dorsal view C umbilical view showing thick peristome and parietalis D aperture facing right view showing tongue-like flexion of the lamella E aperture facing left view F apical view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1167.102840.figure5" httpUri="https://binary.pensoft.net/fig/863996" pageId="0" pageNumber="89">, 5</figureCitation>
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<figureCitation id="BAE52D4B4D51BA82E630DC5C96C11985" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. 3 D visualizations of X-ray micro-CT data of fossil Carychium floridanum G. H. Clapp, 1918 (NMBE 577016) A apertural view B dorsal view C umbilical view showing thick peristome and parietalis D aperture facing right view E aperture facing left view F apical view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1167.102840.figure6" httpUri="https://binary.pensoft.net/fig/863997" pageId="0" pageNumber="89">, 6</figureCitation>
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<figureCitation id="4B27A7507E4F4990B07D28253108BC2F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Recent Carychium floridanum G. H. Clapp, 1918 (NMBE 572256) from Wakulla Springs, Florida. Light microscopic images showing A apertural view with thick peristome, parietalis and columellaris B dorsal view showing axial ribbing C aperture facing right view D aperture facing left view E umbilical view showing thick peristome and parietalis F apical view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1167.102840.figure7" httpUri="https://binary.pensoft.net/fig/863998" pageId="0" pageNumber="89">, 7</figureCitation>
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<figureCitation id="4BEF64DA0E10901BCBA0FB596825B696" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. 3 D visualizations of X-ray micro-CT data of extant Carychium floridanum G. H. Clapp, 1918 (NMBE 572256) from Wakulla Springs, Florida A apertural view B dorsal view C aperture facing right view D aperture facing left view E cross section of D showing lamellar flexion F cross section of B showing full columellar extension G apical view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1167.102840.figure8" httpUri="https://binary.pensoft.net/fig/863999" pageId="0" pageNumber="89">, 8</figureCitation>
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</paragraph>
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<subSubSection id="B143F45FD77E02008B0AE60F47930DA3" pageId="0" pageNumber="89" type="reference_group">
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<paragraph id="292444D16FE5BFC172512DA8ED2F6FE3" pageId="0" pageNumber="89">
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<taxonomicName id="7B53CFB0BF5AA54D138B4E34B2D569D4" authorityName="G. H. Clapp" authorityYear="1918" class="Gastropoda" family="Ellobiidae" genus="Carychium" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Carychium exiguum subsp. floridanum" order="Ellobiida" pageId="0" pageNumber="89" phylum="Mollusca" rank="subSpecies" species="exiguum" subSpecies="floridanum">Carychium exiguum floridanum</taxonomicName>
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G. H. Clapp, 1918: 73-75, pl. 8.
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</paragraph>
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<paragraph id="A6E51B5D7C17CDC8EB1C4BD8114061E3" pageId="0" pageNumber="89">
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<taxonomicName id="B328E688F61EE60CBB91D3AE5ACAFA7D" authorityName="G. H. Clapp" authorityYear="1918" class="Gastropoda" family="Ellobiidae" genus="Carychium" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Carychium floridanum" order="Ellobiida" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="floridanum">Carychium floridanum</taxonomicName>
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G. H. Clapp, 1918:
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<bibRefCitation id="027E9260202876A7489DD21404EE3440" DOI="https://doi.org/10.1186/1471-2148-13-18" author="Weigand, AM" journalOrPublisher="BMC Evolutionary Biology" pageId="0" pageNumber="89" pagination=": 18" refId="B29" refString="Weigand, AM, Jochum, A, Slapnik, R, Schnitzler, J, Zarza, E, Klussmann-Kolb, A, 2013. Evolution of microgastropods (Ellobioidea, Carychiidae): Integrating taxonomic, phylogenetic and evolutionary hypotheses. BMC Evolutionary Biology 13 (1): 18, DOI: https://doi.org/10.1186/1471-2148-13-18" title="Evolution of microgastropods (Ellobioidea, Carychiidae): Integrating taxonomic, phylogenetic and evolutionary hypotheses." url="https://doi.org/10.1186/1471-2148-13-18" volume="13" year="2013">Weigand et al. 2013</bibRefCitation>
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: 3, fig. 1 24|C5; Seq. ID: BARCA032-10, BARCA033-10, BARCA034-10, BARCA035-10, BARCA037-10.
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</paragraph>
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<paragraph id="80B6F587A929B05AD7E0974604E5A85D" pageId="0" pageNumber="89">
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<taxonomicName id="F3B1A4BD97723C5220428A9A82701873" authorityName="G. H. Clapp" authorityYear="1918" class="Gastropoda" family="Ellobiidae" genus="Carychium" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Carychium floridanum" order="Ellobiida" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="floridanum">Carychium floridanum</taxonomicName>
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G. H. Clapp, 1918:
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<bibRefCitation id="087F451294CC6205A8B86DD817D86985" DOI="https://doi.org/10.3897/zookeys.675.12453" author="Jochum, A" journalOrPublisher="ZooKeys" pageId="0" pageNumber="89" pagination="97 - 127" refId="B12" refString="Jochum, A, Weigand, AM, Bochud, E, Inaebnit, T, Doerge, D, Ruthensteiner, B, Favre, A, Martels, G, Kampschulte, M, 2017. Three new species of Carychium O. F. Mueller, 1773 from the southeastern USA, Belize, and Panama are described using computer tomography (CT) (Eupulmonata, Ellobioidea, Carychiidae). ZooKeys 675: 97 - 127, DOI: https://doi.org/10.3897/zookeys.675.12453" title="Three new species of Carychium O. F. Mueller, 1773 from the southeastern USA, Belize, and Panama are described using computer tomography (CT) (Eupulmonata, Ellobioidea, Carychiidae)." url="https://doi.org/10.3897/zookeys.675.12453" volume="675" year="2017">Jochum et al. 2017</bibRefCitation>
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: 110-111, figs 6, 7.
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</paragraph>
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</subSubSection>
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<subSubSection id="DF89287C37291399AC5316E904A584F5" pageId="0" pageNumber="89" type="materials_examined">
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<paragraph id="F02AC3F4D17BD458851CE0236535D076" pageId="0" pageNumber="89">Material examined.</paragraph>
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<paragraph id="70D98C7DE82D0B6C115E9760819416CC" pageId="0" pageNumber="89">
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<materialsCitation id="8E0467B9CD140E900A4EFBE438F1BBF0" collectingDate="2021-10" collectorName="R. Portell, H. Means" country="USA" county="Orange County" elevation="500" latitude="28.4489" location="Orlando" longLatPrecision="6" longitude="-81.0375" specimenCount="1" stateProvince="Florida">
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<collectingCountry id="F0F53FB44C3FFB157D8458261106C4EB" name="United States of America">USA</collectingCountry>
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,
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<collectingRegion id="0B05087F92AD520646C6C277C952606B" country="United States of America" name="Florida">Florida</collectingRegion>
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•
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<collectingCounty id="362334218EE818211FBD010D05863319">Orange County</collectingCounty>
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,
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<location id="A4D0F36DA1B40004EC485652B7C43915" LSID="urn:lsid:plazi:treatment:A5A9E77D85925E979C4DA11BDD6722A1:A4D0F36DA1B40004EC485652B7C43915" country="USA" county="Orange County" latitude="28.4489" longLatPrecision="6" longitude="-81.0375" name="Orlando" stateProvince="Florida">Orlando</location>
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;
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<geoCoordinate id="BF5FD11146B6768A2E8A49505F071AFC" degrees="28.4489" direction="north" orientation="latitude" precision="5" value="28.4489">28.4489</geoCoordinate>
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,
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<geoCoordinate id="E061C8D1F3FCCCE34BDF5F48AACCBB39" degrees="81.0375" direction="west" orientation="longitude" precision="5" value="-81.0375">-81.0375</geoCoordinate>
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(WGS84) encompassing
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<quantity id="9EBB8D452EE17792EFC733F48BFD3DF3" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">
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<elevation id="9E1F24BC4C49C9852EC62550C470207D" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">500 m</elevation>
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</quantity>
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radius;
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<collectingDate id="501B822E95A2DDF4452A40B9CD6450E3" value="2021-10">Oct. 2021</collectingDate>
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;
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<collectorName id="78BC43CBE87C16ECD35D0670BC1986FC">R. Portell</collectorName>
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and
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<collectorName id="E19E246913BD73E0271FE63DFEE90B9E">H. Means</collectorName>
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leg.; NMBE 577015-577016 (2 fossil specimens) (Figs
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<figureCitation id="2D06B309603E63FA8E7D24C4E2FE1703" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Fossil Carychium floridanum G. H. Clapp, 1918 (NMBE 577015 - 577016). Light microscopic images showing different perspectives A-G (NMBE 577015) H-N (NMBE 577016). Peristome thickly callused with apertural barriers. Grey and orange colours indicate taphonomic and sedimentary processes, repectively. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1167.102840.figure4" httpUri="https://binary.pensoft.net/fig/863995" pageId="0" pageNumber="89">4</figureCitation>
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-
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<figureCitation id="D55D80E3814F2FFE27CCC2A32CA97154" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. 3 D visualizations of X-ray micro-CT data of fossil Carychium floridanum G. H. Clapp, 1918 (NMBE 577016) A apertural view B dorsal view C umbilical view showing thick peristome and parietalis D aperture facing right view E aperture facing left view F apical view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1167.102840.figure6" httpUri="https://binary.pensoft.net/fig/863997" pageId="0" pageNumber="89">6</figureCitation>
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)
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</materialsCitation>
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.
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</paragraph>
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<caption id="5848A3AB1C8C6D69582693E06DBBCC79" doi="10.3897/zookeys.1167.102840.figure4" httpUri="https://binary.pensoft.net/fig/863995" pageId="0" pageNumber="89" start="Figure 4" startId="F4">
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<paragraph id="3CC7D1307104DDB87068F700FF7570C4" pageId="0" pageNumber="89">
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<emphasis id="0F729502D2B8A44BA1049418D7AA2EED" bold="true" pageId="0" pageNumber="89">Figure 4.</emphasis>
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Fossil
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<taxonomicName id="F4F71281E17DCB65C11ABBA8972C3610" authorityName="G. H. Clapp" authorityYear="1918" class="Gastropoda" family="Ellobiidae" genus="Carychium" kingdom="Animalia" lsidName="Carychium floridanum" order="Ellobiida" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="floridanum">
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|
<emphasis id="D3FAD16E195A5F05E738EE908F91922F" italics="true" pageId="0" pageNumber="89">Carychium floridanum</emphasis>
|
|
</taxonomicName>
|
|
G. H. Clapp, 1918 (NMBE 577015-577016). Light microscopic images showing different perspectives
|
|
<emphasis id="2A9ABAF4FC5AF5145FEE9EA4B4370FAE" bold="true" pageId="0" pageNumber="89">A-G</emphasis>
|
|
(NMBE 577015)
|
|
<emphasis id="FBA4F43CD66671D1723C26E1554A18F1" bold="true" pageId="0" pageNumber="89">H-N</emphasis>
|
|
(NMBE 577016). Peristome thickly callused with apertural barriers. Grey and orange colours indicate taphonomic and sedimentary processes, repectively. Scale bar:
|
|
<quantity id="10F64A5AD7441BD220A5E4984FE6C0A3" metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
|
|
.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="F67BA3634C721D3FC7CE57B94C05923B" doi="10.3897/zookeys.1167.102840.figure5" httpUri="https://binary.pensoft.net/fig/863996" pageId="0" pageNumber="89" start="Figure 5" startId="F5">
|
|
<paragraph id="A6A428FD5D178605CF3C8B7FAE4955E0" pageId="0" pageNumber="89">
|
|
<emphasis id="A4F02CE6EE46AB17CCEDA3BAB1CD4024" bold="true" pageId="0" pageNumber="89">Figure 5.</emphasis>
|
|
3D visualizations of X-ray micro-CT data of fossil
|
|
<taxonomicName id="324F6B0B4F84D2C036C05003383DF2D4" authorityName="G. H. Clapp" authorityYear="1918" class="Gastropoda" family="Ellobiidae" genus="Carychium" kingdom="Animalia" lsidName="Carychium floridanum" order="Ellobiida" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="floridanum">
|
|
<emphasis id="381442578E0D157DC3C4059B639C769A" italics="true" pageId="0" pageNumber="89">Carychium floridanum</emphasis>
|
|
</taxonomicName>
|
|
G. H. Clapp, 1918 (NMBE 577015)
|
|
<emphasis id="AF0E457CE307B2AC34698358AE0E976B" bold="true" pageId="0" pageNumber="89">A</emphasis>
|
|
apertural view showing extent of lamellar fold
|
|
<emphasis id="35C1686C13ADE94CBDEBD355C301C141" bold="true" pageId="0" pageNumber="89">B</emphasis>
|
|
dorsal view
|
|
<emphasis id="7035A8EF1F1A8C5CC11EBACCB6C89126" bold="true" pageId="0" pageNumber="89">C</emphasis>
|
|
umbilical view showing thick peristome and parietalis
|
|
<emphasis id="0F37A6DB68E8462D98E81B8E71FD1A7D" bold="true" pageId="0" pageNumber="89">D</emphasis>
|
|
aperture facing right view showing tongue-like flexion of the lamella
|
|
<emphasis id="3C4D2497CFEE62FD2036DA50A08B942C" bold="true" pageId="0" pageNumber="89">E</emphasis>
|
|
aperture facing left view
|
|
<emphasis id="ECF6204D2C0317E13150E1BAA1480F9E" bold="true" pageId="0" pageNumber="89">F</emphasis>
|
|
apical view. Scale bar:
|
|
<quantity id="86F97833B7667079056F5E47964CB912" metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
|
|
.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="02D020ACFD3F9AABDAFF4EF1BC49DD2F" doi="10.3897/zookeys.1167.102840.figure6" httpUri="https://binary.pensoft.net/fig/863997" pageId="0" pageNumber="89" start="Figure 6" startId="F6">
|
|
<paragraph id="4FDA334E7C916716BB77EDC19AB64CF3" pageId="0" pageNumber="89">
|
|
<emphasis id="ECEDD359FC2397FF101960A37DE26046" bold="true" pageId="0" pageNumber="89">Figure 6.</emphasis>
|
|
3D visualizations of X-ray micro-CT data of fossil
|
|
<taxonomicName id="41C0A0935DC42655B3F62EA093F6489B" authorityName="G. H. Clapp" authorityYear="1918" class="Gastropoda" family="Ellobiidae" genus="Carychium" kingdom="Animalia" lsidName="Carychium floridanum" order="Ellobiida" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="floridanum">
|
|
<emphasis id="0C160B4F7B547F9D28F32030814DB25C" italics="true" pageId="0" pageNumber="89">Carychium floridanum</emphasis>
|
|
</taxonomicName>
|
|
G. H. Clapp, 1918 (NMBE 577016)
|
|
<emphasis id="77223E5090D33D58491C51D236E9A05E" bold="true" pageId="0" pageNumber="89">A</emphasis>
|
|
apertural view
|
|
<emphasis id="E9E1F25023DCEE677242D87E14415295" bold="true" pageId="0" pageNumber="89">B</emphasis>
|
|
dorsal view
|
|
<emphasis id="F3662035442A24B0349C4D6107FBAC8D" bold="true" pageId="0" pageNumber="89">C</emphasis>
|
|
umbilical view showing thick peristome and parietalis
|
|
<emphasis id="696DE0CA9B7140BCB4BAB9446F0AB9AE" bold="true" pageId="0" pageNumber="89">D</emphasis>
|
|
aperture facing right view
|
|
<emphasis id="BAAABA361D16078054269A483EC15472" bold="true" pageId="0" pageNumber="89">E</emphasis>
|
|
aperture facing left view
|
|
<emphasis id="6817FC1D4C1099F34032E21B62C7576C" bold="true" pageId="0" pageNumber="89">F</emphasis>
|
|
apical view. Scale bar:
|
|
<quantity id="33EAA1EE6ED10F19FD2E040C660DC297" metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1mm</quantity>
|
|
.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="65EC7CEB7D6D31CD378F45982FAE7E74" pageId="0" pageNumber="89">
|
|
The two shells were compared to the
|
|
<typeStatus id="C01BF03D77EFA0F6F664F2A1E96066EA">syntype</typeStatus>
|
|
shell from Snapper Creek Hammock, Miami, FL (CM 46540) presented in
|
|
<bibRefCitation id="427F3FA52E0604C8D5EE4FBFA6D4D4FE" DOI="https://doi.org/10.3897/zookeys.675.12453" author="Jochum, A" journalOrPublisher="ZooKeys" pageId="0" pageNumber="89" pagination="97 - 127" refId="B12" refString="Jochum, A, Weigand, AM, Bochud, E, Inaebnit, T, Doerge, D, Ruthensteiner, B, Favre, A, Martels, G, Kampschulte, M, 2017. Three new species of Carychium O. F. Mueller, 1773 from the southeastern USA, Belize, and Panama are described using computer tomography (CT) (Eupulmonata, Ellobioidea, Carychiidae). ZooKeys 675: 97 - 127, DOI: https://doi.org/10.3897/zookeys.675.12453" title="Three new species of Carychium O. F. Mueller, 1773 from the southeastern USA, Belize, and Panama are described using computer tomography (CT) (Eupulmonata, Ellobioidea, Carychiidae)." url="https://doi.org/10.3897/zookeys.675.12453" volume="675" year="2017">Jochum et al. (2017</bibRefCitation>
|
|
, figs 6, 7) and additionally, to a Recent shell from the Wakulla Springs, FL population [NMBE 572256 (ex. AJC 1446)] [
|
|
<bibRefCitation id="C57968B9101BC0F68D553D2F7A6073CC" DOI="https://doi.org/10.1186/1471-2148-13-18" author="Weigand, AM" journalOrPublisher="BMC Evolutionary Biology" pageId="0" pageNumber="89" pagination=": 18" refId="B29" refString="Weigand, AM, Jochum, A, Slapnik, R, Schnitzler, J, Zarza, E, Klussmann-Kolb, A, 2013. Evolution of microgastropods (Ellobioidea, Carychiidae): Integrating taxonomic, phylogenetic and evolutionary hypotheses. BMC Evolutionary Biology 13 (1): 18, DOI: https://doi.org/10.1186/1471-2148-13-18" title="Evolution of microgastropods (Ellobioidea, Carychiidae): Integrating taxonomic, phylogenetic and evolutionary hypotheses." url="https://doi.org/10.1186/1471-2148-13-18" volume="13" year="2013">Weigand et al. 2013</bibRefCitation>
|
|
, Evolutionary Lineage (EL) C5] (Figs
|
|
<figureCitation id="D6106F988BE50E0B14E4608FFA909D75" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Recent Carychium floridanum G. H. Clapp, 1918 (NMBE 572256) from Wakulla Springs, Florida. Light microscopic images showing A apertural view with thick peristome, parietalis and columellaris B dorsal view showing axial ribbing C aperture facing right view D aperture facing left view E umbilical view showing thick peristome and parietalis F apical view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1167.102840.figure7" httpUri="https://binary.pensoft.net/fig/863998" pageId="0" pageNumber="89">7</figureCitation>
|
|
,
|
|
<figureCitation id="DB45AA340C4D60B47D26B38857A6FE84" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. 3 D visualizations of X-ray micro-CT data of extant Carychium floridanum G. H. Clapp, 1918 (NMBE 572256) from Wakulla Springs, Florida A apertural view B dorsal view C aperture facing right view D aperture facing left view E cross section of D showing lamellar flexion F cross section of B showing full columellar extension G apical view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1167.102840.figure8" httpUri="https://binary.pensoft.net/fig/863999" pageId="0" pageNumber="89">8</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption id="634D462BE04780E5ECA55A81A55D3980" doi="10.3897/zookeys.1167.102840.figure7" httpUri="https://binary.pensoft.net/fig/863998" pageId="0" pageNumber="89" start="Figure 7" startId="F7">
|
|
<paragraph id="753E26969766365B4A454340EE2CF321" pageId="0" pageNumber="89">
|
|
<emphasis id="A642B57A366FA34DB0C7A6F4C1E1658F" bold="true" pageId="0" pageNumber="89">Figure 7.</emphasis>
|
|
Recent
|
|
<taxonomicName id="556ACA5359388CD057B2F1D2BC89F50E" authorityName="G. H. Clapp" authorityYear="1918" class="Gastropoda" family="Ellobiidae" genus="Carychium" kingdom="Animalia" lsidName="Carychium floridanum" order="Ellobiida" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="floridanum">
|
|
<emphasis id="72FB2A6D74D5E137C7791A14F13BBFA1" italics="true" pageId="0" pageNumber="89">Carychium floridanum</emphasis>
|
|
</taxonomicName>
|
|
G. H. Clapp, 1918 (NMBE 572256) from Wakulla Springs,
|
|
<collectingRegion id="BE7290C1E72898B1F9C3B09849244504" country="United States of America" name="Florida">Florida</collectingRegion>
|
|
. Light microscopic images showing
|
|
<emphasis id="23711D9C70D9B6985AF4A57CAA64F522" bold="true" pageId="0" pageNumber="89">A</emphasis>
|
|
apertural view with thick peristome, parietalis and columellaris
|
|
<emphasis id="BBD55495AB76F013D379C55FF690DCB6" bold="true" pageId="0" pageNumber="89">B</emphasis>
|
|
dorsal view showing axial ribbing
|
|
<emphasis id="FEDF11EA5ABED867732227ABC7C97980" bold="true" pageId="0" pageNumber="89">C</emphasis>
|
|
aperture facing right view
|
|
<emphasis id="53A49FF78A5DDF0E6E0016F5CBA0EB61" bold="true" pageId="0" pageNumber="89">D</emphasis>
|
|
aperture facing left view
|
|
<emphasis id="C71CCBBC4D7D53E76CA9D046E1553AD0" bold="true" pageId="0" pageNumber="89">E</emphasis>
|
|
umbilical view showing thick peristome and parietalis
|
|
<emphasis id="C7D3ADA74832863948C0CBDA73F5DA44" bold="true" pageId="0" pageNumber="89">F</emphasis>
|
|
apical view. Scale bar:
|
|
<quantity id="98A33AB83186F011A18E26FFAC018B52" metricMagnitude="-4" metricUnit="m" metricValue="5.0" unit="mm" value="0.5">0.5 mm</quantity>
|
|
.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="761880C87B13505E2986CFC5C3313BFD" doi="10.3897/zookeys.1167.102840.figure8" httpUri="https://binary.pensoft.net/fig/863999" pageId="0" pageNumber="89" start="Figure 8" startId="F8">
|
|
<paragraph id="C30E44014EF0506D5247DB0BB4CCE49D" pageId="0" pageNumber="89">
|
|
<emphasis id="AAAD1C0731B46D5821A0C52D9126E1DF" bold="true" pageId="0" pageNumber="89">Figure 8.</emphasis>
|
|
3D visualizations of X-ray micro-CT data of extant
|
|
<taxonomicName id="E389BB2B611AD287E62073BE759811A9" authorityName="G. H. Clapp" authorityYear="1918" class="Gastropoda" family="Ellobiidae" genus="Carychium" kingdom="Animalia" lsidName="Carychium floridanum" order="Ellobiida" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="floridanum">
|
|
<emphasis id="41ACD2EC4054961DD020AC6B1E607E55" italics="true" pageId="0" pageNumber="89">Carychium floridanum</emphasis>
|
|
</taxonomicName>
|
|
G. H. Clapp, 1918 (NMBE 572256) from Wakulla Springs,
|
|
<collectingRegion id="B1F3F376A430B5F4365FFE265928C6AE" country="United States of America" name="Florida">Florida</collectingRegion>
|
|
<emphasis id="1E0E3A68595EBEF5183B799A294B912F" bold="true" pageId="0" pageNumber="89">A</emphasis>
|
|
apertural view
|
|
<emphasis id="10316A9C6BC9F01B4A9400CE2FF6643E" bold="true" pageId="0" pageNumber="89">B</emphasis>
|
|
dorsal view
|
|
<emphasis id="B168C4D7F5E028D347E06E75812C4177" bold="true" pageId="0" pageNumber="89">C</emphasis>
|
|
aperture facing right view
|
|
<emphasis id="7BB5A2F7F3BB8647AC409FA932FA94E4" bold="true" pageId="0" pageNumber="89">D</emphasis>
|
|
aperture facing left view
|
|
<emphasis id="A280C34F743004F11F4AE075E2E90EFF" bold="true" pageId="0" pageNumber="89">E</emphasis>
|
|
cross section of
|
|
<emphasis id="8B0103DBD70036F4AC309921CF8783B0" bold="true" pageId="0" pageNumber="89">D</emphasis>
|
|
showing lamellar flexion
|
|
<emphasis id="D4AD3BDD1BA576BF6FCBE77E9C73BCFD" bold="true" pageId="0" pageNumber="89">F</emphasis>
|
|
cross section of
|
|
<emphasis id="4B91EBE1990F761463B612E31B46C922" bold="true" pageId="0" pageNumber="89">B</emphasis>
|
|
showing full columellar extension
|
|
<emphasis id="AAD2B3897CC532D41DB134BC9E01C911" bold="true" pageId="0" pageNumber="89">G</emphasis>
|
|
apical view. Scale bar:
|
|
<quantity id="20494F772F55126D00F2DBBCAC1AF04F" metricMagnitude="-4" metricUnit="m" metricValue="5.0" unit="mm" value="0.5">0.5 mm</quantity>
|
|
.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="E9C65B95FE356C1F32F602DF89E3298B" pageId="0" pageNumber="89" type="stratigraphic occurrence">
|
|
<paragraph id="EA52C9F9493AABA68F4464121BBF7E56" pageId="0" pageNumber="89">Stratigraphic occurrence.</paragraph>
|
|
<paragraph id="C3826514C102D85377AD5A579147EC46" pageId="0" pageNumber="89">Freshwater Marl Bed (middle nonmarine layer) Florida Museum of Natural History Invertebrate Paleontology (FLMNH IP) locality OR022, Lower Pleistocene Nashua Formation.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="8F92424262F9108C2190DA0C34D92C09" pageId="0" pageNumber="89" type="description">
|
|
<paragraph id="98CBFB5EBF5B3BC2BE3457E31BBEB508" pageId="0" pageNumber="89">Description.</paragraph>
|
|
<paragraph id="FCFA2DFA33825C189BEF333BC0F68CBA" pageId="0" pageNumber="89">
|
|
Shell medium-sized for genus (SH: 1.6-1.8 mm), pupiform in shape, glossy; teleoconch with occasional, fine threadlike ribs, with 4.5-5 tumid whorls. Suture deep. Protoconch smooth, bulbous, shiny, and distinct from teleoconch. Whorl profile strongly convex, aperture higher than wide, elliptical, with 2 barriers, parietal robust with the columellaris constituting a low swollen hump deep in the shell. Peristome doubled and heavily callused, expanded in palatal, basal and columellar regions; largely reflected at the base. Palatal lip weakly sigmoid and buttressed with heavy callus inside and marked with a concavity or line at mid-section behind the apertural rim, extending 2/3 the body whorl in aperture facing left profile view (clearly visible on the grey shell and not detectable on the yellowish one) (Fig.
|
|
<figureCitation id="63687658D0302A3A85D5A56D08FB8C3F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Fossil Carychium floridanum G. H. Clapp, 1918 (NMBE 577015 - 577016). Light microscopic images showing different perspectives A-G (NMBE 577015) H-N (NMBE 577016). Peristome thickly callused with apertural barriers. Grey and orange colours indicate taphonomic and sedimentary processes, repectively. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1167.102840.figure4" httpUri="https://binary.pensoft.net/fig/863995" pageId="0" pageNumber="89">4</figureCitation>
|
|
). PH 38% of SH. The upper palatal part of the peristome is narrower causing the callused mid-section to project beyond the apertural rim (in aperture facing left view). Umbilicus rimate (Figs
|
|
<figureCitation id="2D7B2F6F9C4BEECCDB568A09A50FAB20" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. 3 D visualizations of X-ray micro-CT data of fossil Carychium floridanum G. H. Clapp, 1918 (NMBE 577015) A apertural view showing extent of lamellar fold B dorsal view C umbilical view showing thick peristome and parietalis D aperture facing right view showing tongue-like flexion of the lamella E aperture facing left view F apical view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1167.102840.figure5" httpUri="https://binary.pensoft.net/fig/863996" pageId="0" pageNumber="89">5D</figureCitation>
|
|
,
|
|
<figureCitation id="9E8E6084B2DADDFBF151271E3F07038B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. 3 D visualizations of X-ray micro-CT data of fossil Carychium floridanum G. H. Clapp, 1918 (NMBE 577016) A apertural view B dorsal view C umbilical view showing thick peristome and parietalis D aperture facing right view E aperture facing left view F apical view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1167.102840.figure6" httpUri="https://binary.pensoft.net/fig/863997" pageId="0" pageNumber="89">6D</figureCitation>
|
|
). Internally, a single lamella spirals down the columella for at least the last whorl and a half, terminating in the parietalis of the aperture, which is partially visible and positioned near the columellar side of the inner margin of the peristome in umbilical view (Figs
|
|
<figureCitation id="D395081F114773152E6A180050ABD171" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. 3 D visualizations of X-ray micro-CT data of fossil Carychium floridanum G. H. Clapp, 1918 (NMBE 577015) A apertural view showing extent of lamellar fold B dorsal view C umbilical view showing thick peristome and parietalis D aperture facing right view showing tongue-like flexion of the lamella E aperture facing left view F apical view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1167.102840.figure5" httpUri="https://binary.pensoft.net/fig/863996" pageId="0" pageNumber="89">5C</figureCitation>
|
|
,
|
|
<figureCitation id="078C636FC9136B42FCC0A03B0C0DA345" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. 3 D visualizations of X-ray micro-CT data of fossil Carychium floridanum G. H. Clapp, 1918 (NMBE 577016) A apertural view B dorsal view C umbilical view showing thick peristome and parietalis D aperture facing right view E aperture facing left view F apical view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1167.102840.figure6" httpUri="https://binary.pensoft.net/fig/863997" pageId="0" pageNumber="89">6C</figureCitation>
|
|
). The lamella is concave, broadly expanded and wing-like, and strongly flexes down immediately before the aperture, forming the robust parietalis (Figs
|
|
<figureCitation id="8DE8E85AAE9C07A91B53F7874482505C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. 3 D visualizations of X-ray micro-CT data of fossil Carychium floridanum G. H. Clapp, 1918 (NMBE 577015) A apertural view showing extent of lamellar fold B dorsal view C umbilical view showing thick peristome and parietalis D aperture facing right view showing tongue-like flexion of the lamella E aperture facing left view F apical view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1167.102840.figure5" httpUri="https://binary.pensoft.net/fig/863996" pageId="0" pageNumber="89">5A, E</figureCitation>
|
|
,
|
|
<figureCitation id="207467D18C9B30B111A19302DA0E0E1E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. 3 D visualizations of X-ray micro-CT data of fossil Carychium floridanum G. H. Clapp, 1918 (NMBE 577016) A apertural view B dorsal view C umbilical view showing thick peristome and parietalis D aperture facing right view E aperture facing left view F apical view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1167.102840.figure6" httpUri="https://binary.pensoft.net/fig/863997" pageId="0" pageNumber="89">6A, E</figureCitation>
|
|
). The surface of the lamella is broad (in dorsal view) with the edge sloping slightly oblique to the columella, flipping over forming a thickened, tongue-like structure in aperture facing right view (Figs
|
|
<figureCitation id="9A4735205A73ED2C885B0B0DC37F2A8D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. 3 D visualizations of X-ray micro-CT data of fossil Carychium floridanum G. H. Clapp, 1918 (NMBE 577015) A apertural view showing extent of lamellar fold B dorsal view C umbilical view showing thick peristome and parietalis D aperture facing right view showing tongue-like flexion of the lamella E aperture facing left view F apical view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1167.102840.figure5" httpUri="https://binary.pensoft.net/fig/863996" pageId="0" pageNumber="89">5B, D</figureCitation>
|
|
,
|
|
<figureCitation id="FDD73E5FF180617E123A006D757987D0" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. 3 D visualizations of X-ray micro-CT data of fossil Carychium floridanum G. H. Clapp, 1918 (NMBE 577016) A apertural view B dorsal view C umbilical view showing thick peristome and parietalis D aperture facing right view E aperture facing left view F apical view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1167.102840.figure6" httpUri="https://binary.pensoft.net/fig/863997" pageId="0" pageNumber="89">6B, D</figureCitation>
|
|
). The early lamella starts tightly spiraling down the columella at about 2/3 the height of the penultimate whorl (Figs
|
|
<figureCitation id="B048B92D43F4343180484EB20F2FF1CA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. 3 D visualizations of X-ray micro-CT data of fossil Carychium floridanum G. H. Clapp, 1918 (NMBE 577015) A apertural view showing extent of lamellar fold B dorsal view C umbilical view showing thick peristome and parietalis D aperture facing right view showing tongue-like flexion of the lamella E aperture facing left view F apical view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1167.102840.figure5" httpUri="https://binary.pensoft.net/fig/863996" pageId="0" pageNumber="89">5B</figureCitation>
|
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,
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|
<figureCitation id="287666592604C67A3537ECF267A2CBE6" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. 3 D visualizations of X-ray micro-CT data of fossil Carychium floridanum G. H. Clapp, 1918 (NMBE 577016) A apertural view B dorsal view C umbilical view showing thick peristome and parietalis D aperture facing right view E aperture facing left view F apical view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1167.102840.figure6" httpUri="https://binary.pensoft.net/fig/863997" pageId="0" pageNumber="89">6B</figureCitation>
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|
), expanding rapidly to its fullest winged extension in the middle of the penultimate whorl (Figs
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|
<figureCitation id="EC64763B505C28900B84195BD08562A3" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. 3 D visualizations of X-ray micro-CT data of fossil Carychium floridanum G. H. Clapp, 1918 (NMBE 577015) A apertural view showing extent of lamellar fold B dorsal view C umbilical view showing thick peristome and parietalis D aperture facing right view showing tongue-like flexion of the lamella E aperture facing left view F apical view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1167.102840.figure5" httpUri="https://binary.pensoft.net/fig/863996" pageId="0" pageNumber="89">5A</figureCitation>
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,
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|
<figureCitation id="9C02C6BF612D12F54A92CF3326296192" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. 3 D visualizations of X-ray micro-CT data of fossil Carychium floridanum G. H. Clapp, 1918 (NMBE 577016) A apertural view B dorsal view C umbilical view showing thick peristome and parietalis D aperture facing right view E aperture facing left view F apical view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1167.102840.figure6" httpUri="https://binary.pensoft.net/fig/863997" pageId="0" pageNumber="89">6A</figureCitation>
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|
). The callused swollen peristome is flush with the convexity of the penultimate whorl in aperture facing left view (Figs
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|
<figureCitation id="F48D8B7947447706D6FD123D8E168BB8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. 3 D visualizations of X-ray micro-CT data of fossil Carychium floridanum G. H. Clapp, 1918 (NMBE 577015) A apertural view showing extent of lamellar fold B dorsal view C umbilical view showing thick peristome and parietalis D aperture facing right view showing tongue-like flexion of the lamella E aperture facing left view F apical view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1167.102840.figure5" httpUri="https://binary.pensoft.net/fig/863996" pageId="0" pageNumber="89">5E</figureCitation>
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,
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|
<figureCitation id="2BD96F45B0FA6A6942B84DA41835D9A0" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. 3 D visualizations of X-ray micro-CT data of fossil Carychium floridanum G. H. Clapp, 1918 (NMBE 577016) A apertural view B dorsal view C umbilical view showing thick peristome and parietalis D aperture facing right view E aperture facing left view F apical view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1167.102840.figure6" httpUri="https://binary.pensoft.net/fig/863997" pageId="0" pageNumber="89">6E</figureCitation>
|
|
). The regularly coiled spire is broad with the penultimate whorl expanding almost completely over the body whorl in apical view (Figs
|
|
<figureCitation id="BF59E638CEFE4A38A1D5DFDE50A48D8A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. 3 D visualizations of X-ray micro-CT data of fossil Carychium floridanum G. H. Clapp, 1918 (NMBE 577015) A apertural view showing extent of lamellar fold B dorsal view C umbilical view showing thick peristome and parietalis D aperture facing right view showing tongue-like flexion of the lamella E aperture facing left view F apical view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1167.102840.figure5" httpUri="https://binary.pensoft.net/fig/863996" pageId="0" pageNumber="89">5F</figureCitation>
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,
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|
<figureCitation id="D6C8B5FAF932F40110F6F026B29FA373" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. 3 D visualizations of X-ray micro-CT data of fossil Carychium floridanum G. H. Clapp, 1918 (NMBE 577016) A apertural view B dorsal view C umbilical view showing thick peristome and parietalis D aperture facing right view E aperture facing left view F apical view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1167.102840.figure6" httpUri="https://binary.pensoft.net/fig/863997" pageId="0" pageNumber="89">6F</figureCitation>
|
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).
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</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="A69FB1C58A24DBF9AACDD587D71C3FF0" pageId="0" pageNumber="89" type="description">
|
|
<paragraph id="D2D8469B93E598F2422BAA912BFD661C" pageId="0" pageNumber="89">Measurements.</paragraph>
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<paragraph id="AEA6BB7CE73F37F5967C242DFAF54A0E" pageId="0" pageNumber="89">
|
|
SH = 1.6-1.8 mm; SW = 0.81-0.92 mm; PH = 0.60-0.7 mm; PW = 0.61-0.67 mm (
|
|
<emphasis id="5A15F8A244884AA4E0BD653889900A88" italics="true" pageId="0" pageNumber="89">N</emphasis>
|
|
= 3).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="2ACF4011739A2CA3873F53CA25EC79F0" pageId="0" pageNumber="89" type="occurrence">
|
|
<paragraph id="816081076EB46DBB7ECDE85C0617E3B7" pageId="0" pageNumber="89">Occurrence.</paragraph>
|
|
<paragraph id="28C9D709C235A78F5FFA0A2DFA2C68AF" pageId="0" pageNumber="89">A frequent find in the bulk samples of the Freshwater Marl Bed (middle nonmarine layer).</paragraph>
|
|
<paragraph id="A23E54B2C18F46E47FBCCF981389A212" pageId="0" pageNumber="89">
|
|
Living
|
|
<taxonomicName id="20B950A7C8D88680F9E16A3C78ABFED0" kingdom="Animalia" lsidName="C. floridanum" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="floridanum">
|
|
<emphasis id="4B47716837DBF9FD6A93643F8DD7B829" italics="true" pageId="0" pageNumber="89">C. floridanum</emphasis>
|
|
</taxonomicName>
|
|
are known today from central Florida up to Wakulla Springs (
|
|
<bibRefCitation id="26809A6319087AFEE9544B4D44BB7A30" DOI="https://doi.org/10.1186/1471-2148-13-18" author="Weigand, AM" journalOrPublisher="BMC Evolutionary Biology" pageId="0" pageNumber="89" pagination=": 18" refId="B29" refString="Weigand, AM, Jochum, A, Slapnik, R, Schnitzler, J, Zarza, E, Klussmann-Kolb, A, 2013. Evolution of microgastropods (Ellobioidea, Carychiidae): Integrating taxonomic, phylogenetic and evolutionary hypotheses. BMC Evolutionary Biology 13 (1): 18, DOI: https://doi.org/10.1186/1471-2148-13-18" title="Evolution of microgastropods (Ellobioidea, Carychiidae): Integrating taxonomic, phylogenetic and evolutionary hypotheses." url="https://doi.org/10.1186/1471-2148-13-18" volume="13" year="2013">Weigand et al. 2013</bibRefCitation>
|
|
). Earlier collections at the beginning of the 20th century extended as far south as the type locality of Miami in Dade County and in southwestern Florida at Cape Sable, Monroe County (
|
|
<bibRefCitation id="AA3D7ED6E84D5AE2AE849E67CA07594A" author="Clapp, GH" journalOrPublisher="The Nautilus" pageId="0" pageNumber="89" pagination="73 - 74" refId="B3" refString="Clapp, GH, 1918. New southern forms of Carychium and Thysanophora. The Nautilus 31 (3): 73 - 74" title="New southern forms of Carychium and Thysanophora." volume="31" year="1918">Clapp 1918</bibRefCitation>
|
|
). Though human encroachment, canalization practices and habitat modification have severely impacted its occurrence in southern Florida, the paucity of recent records in south Florida is likely the result of limited sampling and not extirpation of the species in this part of the state.
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|
</paragraph>
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|
</subSubSection>
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|
<subSubSection id="CB46AF7B2FF9E0D27782AF4B5352B21C" pageId="0" pageNumber="89" type="materials_examined">
|
|
<paragraph id="229C90D9A337077C8C1CD5BC50CC5126" pageId="0" pageNumber="89">Recent material for comparison.</paragraph>
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|
<paragraph id="3EAD28B830F7C03382EFE8B106DFD402" pageId="0" pageNumber="89">
|
|
<taxonomicName id="D226C0EB8923D3BCD3C606AE630BD8EC" authorityName="G. H. Clapp" authorityYear="1918" class="Gastropoda" family="Ellobiidae" genus="Carychium" kingdom="Animalia" lsidName="Carychium floridanum" order="Ellobiida" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="floridanum">
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|
<emphasis id="F30912F3519A894C267D077CF2C98AFF" italics="true" pageId="0" pageNumber="89">Carychium floridanum</emphasis>
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|
</taxonomicName>
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|
<materialsCitation id="0A5B36E9D166A8E3AA66105DE9BCC0F2" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="G. H. Clapp, Wakulla Springs, Wakulla County, FL" county="Wakulla County" latitude="30.2355" location="Wakulla County" longLatPrecision="6" longitude="-84.3031" specimenCount="1">
|
|
<collectorName id="0C9168563207A6F7A4EF95A49202852E">G. H. Clapp</collectorName>
|
|
, 1918,
|
|
<collectorName id="FC1FC8985EB492AA9A46FCB84882C9E5">Wakulla Springs</collectorName>
|
|
,
|
|
<collectorName id="6B069E91BD7C5DCEB1D8284620BE8AB3">
|
|
<collectingCounty id="DF40C6D1F821AAB16B7E6A7D3ACFAE38">Wakulla County</collectingCounty>
|
|
, FL
|
|
</collectorName>
|
|
;
|
|
<geoCoordinate id="A5E55504ED13400AB93C7ABDD50950BC" degrees="30.2355" direction="north" orientation="latitude" precision="5" value="30.2355">30.2355</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="04070DD2B1F3911790E30A7985EA8133" degrees="84.3031" direction="west" orientation="longitude" precision="5" value="-84.3031">-84.3031</geoCoordinate>
|
|
; BARCA032-10; [NMBE 572256 (ex. AJC 1446)] (
|
|
<emphasis id="A1D07AD2F73B4C364C79E69E4D026A57" italics="true" pageId="0" pageNumber="89">N</emphasis>
|
|
= 1) (
|
|
<bibRefCitation id="7EDC18C75561269A0DC8A2C92CB7F7F5" DOI="https://doi.org/10.1186/1471-2148-13-18" author="Weigand, AM" journalOrPublisher="BMC Evolutionary Biology" pageId="0" pageNumber="89" pagination=": 18" refId="B29" refString="Weigand, AM, Jochum, A, Slapnik, R, Schnitzler, J, Zarza, E, Klussmann-Kolb, A, 2013. Evolution of microgastropods (Ellobioidea, Carychiidae): Integrating taxonomic, phylogenetic and evolutionary hypotheses. BMC Evolutionary Biology 13 (1): 18, DOI: https://doi.org/10.1186/1471-2148-13-18" title="Evolution of microgastropods (Ellobioidea, Carychiidae): Integrating taxonomic, phylogenetic and evolutionary hypotheses." url="https://doi.org/10.1186/1471-2148-13-18" volume="13" year="2013">Weigand et al. 2013</bibRefCitation>
|
|
) (Figs
|
|
<figureCitation id="652302031C76A1F7ED20EE1844797B3D" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Recent Carychium floridanum G. H. Clapp, 1918 (NMBE 572256) from Wakulla Springs, Florida. Light microscopic images showing A apertural view with thick peristome, parietalis and columellaris B dorsal view showing axial ribbing C aperture facing right view D aperture facing left view E umbilical view showing thick peristome and parietalis F apical view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1167.102840.figure7" httpUri="https://binary.pensoft.net/fig/863998" pageId="0" pageNumber="89">7</figureCitation>
|
|
,
|
|
<figureCitation id="EEE9644BEB91AFC604D18C6A2C408743" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. 3 D visualizations of X-ray micro-CT data of extant Carychium floridanum G. H. Clapp, 1918 (NMBE 572256) from Wakulla Springs, Florida A apertural view B dorsal view C aperture facing right view D aperture facing left view E cross section of D showing lamellar flexion F cross section of B showing full columellar extension G apical view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1167.102840.figure8" httpUri="https://binary.pensoft.net/fig/863999" pageId="0" pageNumber="89">8</figureCitation>
|
|
)
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="2CF87BF66813D154FAC4E677E99BC33C" pageId="0" pageNumber="89" type="description">
|
|
<paragraph id="C7CD65BD40E3EC54D00D5531B2EA6666" pageId="0" pageNumber="89">Description and comparison.</paragraph>
|
|
<paragraph id="C38F57C6EFFC15F8FA07D3BFB98E0996" pageId="0" pageNumber="89">Shell pupiform, translucent with fine threadlike ribbing on the teleoconch. 4.5 tumid whorls. Aperture elliptical with 2 barriers; parietalis points downwards and columellaris is stronger than in the fossil shells. The fossil shells show greater whorl convexity and a bulbous protoconch. Peristome unevenly shaped and thickly callused, parietal shield more extensive than in fossil shells. Measurements (NMBE 572256): SH = 1.67 mm; SW= 0.84 mm; PH = 0.68 mm; PW= 0.65 mm.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="4E45B97A1C0EC029F1753E45F6E119B3" pageId="0" pageNumber="89" type="remarks">
|
|
<paragraph id="BBFF38F63238CDBEA4881EDDD21CA38C" pageId="0" pageNumber="89">Remarks.</paragraph>
|
|
<paragraph id="D8D94C1C0E1713C740D20D70579506D0" pageId="0" pageNumber="89">
|
|
Considering that
|
|
<taxonomicName id="23E1AD13350AA7E90EA4CEE61802F714" kingdom="Animalia" lsidName="C. floridanum" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="floridanum">
|
|
<emphasis id="051D828B4CB3B7449A8CD255A3E95B10" italics="true" pageId="0" pageNumber="89">C. floridanum</emphasis>
|
|
</taxonomicName>
|
|
is clearly genetically and morphologically distinct from its SE North American congeners [EL C5 in
|
|
<bibRefCitation id="09682DA5104A7B5D8C02BDBCA52D98FA" DOI="https://doi.org/10.1186/1471-2148-13-18" author="Weigand, AM" journalOrPublisher="BMC Evolutionary Biology" pageId="0" pageNumber="89" pagination=": 18" refId="B29" refString="Weigand, AM, Jochum, A, Slapnik, R, Schnitzler, J, Zarza, E, Klussmann-Kolb, A, 2013. Evolution of microgastropods (Ellobioidea, Carychiidae): Integrating taxonomic, phylogenetic and evolutionary hypotheses. BMC Evolutionary Biology 13 (1): 18, DOI: https://doi.org/10.1186/1471-2148-13-18" title="Evolution of microgastropods (Ellobioidea, Carychiidae): Integrating taxonomic, phylogenetic and evolutionary hypotheses." url="https://doi.org/10.1186/1471-2148-13-18" volume="13" year="2013">Weigand et al (2013)</bibRefCitation>
|
|
], the internal morphology of the northern Wakulla Springs material is congruent with that of the fossil shells here but not fully congruent with that of the southern syntype material (CM 46540) in
|
|
<bibRefCitation id="6215E9668EDDC70C7AC8CB2C72C0FB48" DOI="https://doi.org/10.3897/zookeys.675.12453" author="Jochum, A" journalOrPublisher="ZooKeys" pageId="0" pageNumber="89" pagination="97 - 127" refId="B12" refString="Jochum, A, Weigand, AM, Bochud, E, Inaebnit, T, Doerge, D, Ruthensteiner, B, Favre, A, Martels, G, Kampschulte, M, 2017. Three new species of Carychium O. F. Mueller, 1773 from the southeastern USA, Belize, and Panama are described using computer tomography (CT) (Eupulmonata, Ellobioidea, Carychiidae). ZooKeys 675: 97 - 127, DOI: https://doi.org/10.3897/zookeys.675.12453" title="Three new species of Carychium O. F. Mueller, 1773 from the southeastern USA, Belize, and Panama are described using computer tomography (CT) (Eupulmonata, Ellobioidea, Carychiidae)." url="https://doi.org/10.3897/zookeys.675.12453" volume="675" year="2017">Jochum et al. (2017)</bibRefCitation>
|
|
. The possibility that the Wakulla Springs population and the fossil shells in this study could instead constitute a northern-central Floridian lineage designated
|
|
<taxonomicName id="2388F500E27C3887BFED301C93E80659" kingdom="Animalia" lsidName="C. floridanum" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="floridanum">
|
|
<emphasis id="8C3CAFAACEBB2A82992BC354A3B0C8CD" italics="true" pageId="0" pageNumber="89">C. floridanum</emphasis>
|
|
</taxonomicName>
|
|
by
|
|
<bibRefCitation id="E31A7B523B206DE96E28FE17E833AD6E" DOI="https://doi.org/10.1186/1471-2148-13-18" author="Weigand, AM" journalOrPublisher="BMC Evolutionary Biology" pageId="0" pageNumber="89" pagination=": 18" refId="B29" refString="Weigand, AM, Jochum, A, Slapnik, R, Schnitzler, J, Zarza, E, Klussmann-Kolb, A, 2013. Evolution of microgastropods (Ellobioidea, Carychiidae): Integrating taxonomic, phylogenetic and evolutionary hypotheses. BMC Evolutionary Biology 13 (1): 18, DOI: https://doi.org/10.1186/1471-2148-13-18" title="Evolution of microgastropods (Ellobioidea, Carychiidae): Integrating taxonomic, phylogenetic and evolutionary hypotheses." url="https://doi.org/10.1186/1471-2148-13-18" volume="13" year="2013">Weigand et al. (2013)</bibRefCitation>
|
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has yet to be investigated. Ultimately, until genetic data from
|
|
<taxonomicName id="1A25D15CD7BBF18D9B0AFA4B1B7B6961" kingdom="Animalia" lsidName="C. floridanum" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="floridanum">
|
|
<emphasis id="C2672A1BB74BD5952D3FA32E4B3EFFC7" italics="true" pageId="0" pageNumber="89">C. floridanum</emphasis>
|
|
</taxonomicName>
|
|
from southern Florida is available to prove otherwise, we maintain that the fossil and the Wakulla Springs members are
|
|
<taxonomicName id="2BA85C774F2E270AAAFCCB79CDEC2024" kingdom="Animalia" lsidName="C. floridanum" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="floridanum">
|
|
<emphasis id="393B41606709B395BEACB6361AAEC6BD" italics="true" pageId="0" pageNumber="89">C. floridanum</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |