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<mods:title id="424A4DD947FDCD976B2B730577A91C34">Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species</mods:title>
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<heading id="F82081B9FFF0E6084F9CFE717318B8F6" bold="true" box="[151,817,463,490]" fontSize="11" level="2" pageId="26" pageNumber="27" reason="4">
<taxonomicName id="64D74D56FFF0E6084F9CFE717318B8F6" ID-CoL="79a78ad9-a452-45dc-911a-ca2c718f36bb" authority="Tirmizi &amp; Javed, 1980" authorityName="Tirmizi &amp; Javed" authorityYear="1980" box="[151,817,463,490]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="bengalensis">
<emphasis id="91A3EAC7FFF0E6084F9CFE717318B8F6" bold="true" box="[151,817,463,490]" pageId="26" pageNumber="27">
<emphasis id="91A3EAC7FFF0E6084F9CFE71723BB8F5" bold="true" box="[151,530,463,490]" italics="true" pageId="26" pageNumber="27">Phylladiorhynchus bengalensis</emphasis>
<bibRefCitation id="C7464B24FFF0E6084D12FE717318B8F6" author="Tirmizi, N. M. &amp; Javed, W." box="[537,817,463,490]" pageId="26" pageNumber="27" pagination="255 - 262" refId="ref119830" refString="Tirmizi, N. M. &amp; Javed, W. (1980) Two new species and one new record of Phylladiorhynchus Baba from the Indian Ocean (Decapoda, Galatheidae) Crustaceana, 39, 255 - 262. https: // doi. org / 10.1163 / 156854080 X 00715" type="journal article" year="1980">Tirmizi &amp; Javed, 1980</bibRefCitation>
</emphasis>
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<emphasis id="91A3EAC7FFF0E6084F9CFDAA71FDBB33" box="[151,468,532,556]" italics="true" pageId="26" pageNumber="27">Phylladiorhynchus bengalensis</emphasis>
<bibRefCitation id="C7464B24FFF0E6084ED1FDAA72E8BB33" author="Tirmizi, N. M. &amp; Javed, W." box="[474,705,532,556]" pageId="26" pageNumber="27" pagination="255 - 262" refId="ref119830" refString="Tirmizi, N. M. &amp; Javed, W. (1980) Two new species and one new record of Phylladiorhynchus Baba from the Indian Ocean (Decapoda, Galatheidae) Crustaceana, 39, 255 - 262. https: // doi. org / 10.1163 / 156854080 X 00715" type="journal article" year="1980">Tirmizi &amp; Javed, 1980</bibRefCitation>
a: 258
</taxonomicName>
, fig. 2 (Andaman Sea,
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).—
<treatmentCitation id="227610C4FFF0E6084B34FDAA7566BB33" author="Tirmizi, N. M. &amp; Javed, W." box="[1087,1359,532,556]" page="31" pageId="26" pageNumber="27" year="1993">
<bibRefCitation id="C7464B24FFF0E6084B34FDAA7566BB33" author="Tirmizi, N. M. &amp; Javed, W." box="[1087,1359,532,556]" pageId="26" pageNumber="27" refId="ref119883" refString="Tirmizi, N. M. &amp; Javed, W. (1993) Indian Ocean galatheids (Crustacea: Anomura) Marine Reference Collection and Resource Centre, University of Karachi, Karachi, 147 pp" type="book" year="1993">Tirmizi &amp; Javed, 1993: 31</bibRefCitation>
</treatmentCitation>
, fig. 14 (redescription).—
<treatmentCitation id="227610C4FFF0E6084E75FD8D7208BB54" author="Baba, K." box="[382,545,562,587]" page="304" pageId="26" pageNumber="27" year="2005">
<bibRefCitation id="C7464B24FFF0E6084E75FD8D7208BB54" author="Baba, K." box="[382,545,562,587]" pageId="26" pageNumber="27" pagination="1 - 317" refId="ref115445" refString="Baba, K. (2005) Deep-sea chirostylid and galatheid crustaceans (Decapoda: Anomura) from the Indo-West Pacific, with a list of species. Galathea Reports, 20, 1 - 317." type="journal article" year="2005">Baba, 2005: 304</bibRefCitation>
</treatmentCitation>
(key, synonymies).—
<treatmentCitation id="227610C4FFF0E6084DF5FD8D73F5BB55" author="Baba, K. &amp; Macpherson, E. &amp; Poore, G. C. B. &amp; Ahyong, S. T. &amp; Bermudez, A. &amp; Cabezas, P. &amp; Lin, C. W. &amp; Nizinski, M. &amp; Rodrigues, C. &amp; Schnabel, K. E." box="[766,988,562,587]" page="175" pageId="26" pageNumber="27" year="2008">
<bibRefCitation id="C7464B24FFF0E6084DF5FD8D73F5BB55" author="Baba, K. &amp; Macpherson, E. &amp; Poore, G. C. B. &amp; Ahyong, S. T. &amp; Bermudez, A. &amp; Cabezas, P. &amp; Lin, C. W. &amp; Nizinski, M. &amp; Rodrigues, C. &amp; Schnabel, K. E." box="[766,988,562,587]" pageId="26" pageNumber="27" pagination="1 - 220" refId="ref115482" refString="Baba, K., Macpherson, E., Poore, G. C. B., Ahyong, S. T., Bermudez, A., Cabezas, P., Lin, C. W., Nizinski, M., Rodrigues, C. &amp; Schnabel, K. E. (2008) Catalogue of squat lobsters of the world (Crustacea: Decapoda: Anomura-families Chirostylidae, Galatheidae and Kiwaidae). Zootaxa, 1905 (1), 1 - 220. https: // doi. org / 10.11646 / zootaxa. 1905.1.1" type="journal article" year="2008">
Baba
<emphasis id="91A3EAC7FFF0E6084C32FD8D734DBB54" box="[825,868,563,587]" italics="true" pageId="26" pageNumber="27">et al</emphasis>
., 2008: 175
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(compilation).—
<treatmentCitation id="227610C4FFF0E6084B82FD8D70C4BB76" author="Schnabel, K. E. &amp; Ahyong, S. T." page="304" pageId="26" pageNumber="27" year="2019">
<bibRefCitation id="C7464B24FFF0E6084B82FD8D70C4BB76" author="Schnabel, K. E. &amp; Ahyong, S. T." pageId="26" pageNumber="27" pagination="301 - 347" refId="ref119703" refString="Schnabel, K. E. &amp; Ahyong, S. T. (2019) The squat lobster genus Phylladiorhynchus Baba, 1969 in New Zealand and eastern Australia, with description of six new species. Zootaxa, 4688 (3), 301 - 347. https: // doi. org / 10.11646 / zootaxa. 4688.3.1" type="journal article" year="2019">Schnabel &amp; Ahyong, 2019: 304</bibRefCitation>
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(key).
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<emphasis id="91A3EAC7FFF0E6084F9CFD29713BBBAE" bold="true" box="[151,274,663,689]" pageId="26" pageNumber="27">Diagnosis.</emphasis>
Rostrum triangular, subapical spines absent or, if present, obsolescent, lateral margins straight. Carapace with epigastric ridge armed with 5 spines; anterior branchial margin with 3 spines; anterior protogastric ridge not interrupted medially, without scale behind median epigastric spine; anterior mesogastric and metagastric ridges not medially interrupted. Anterior margin of thoracic sternite 3 nearly transverse with median and lateral projections. Antennule article 1 with 5 distal spines; distomesial spine minute. Antennal article 2 with distolateral spine longer than distomesial; article 3 with minute distomesial spine. Flexor margin of Mxp3 merus with 2 prominent spines. P24 dactylus extensor margin without upright spines at bases of movable spines.
</paragraph>
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<subSubSection id="EBCD655EFFF0E6084FCCFC2D73D5BAB1" box="[199,1020,915,942]" pageId="26" pageNumber="27" type="distribution">
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<emphasis id="91A3EAC7FFF0E6084FCCFC2D7148BAB2" bold="true" box="[199,353,915,941]" pageId="26" pageNumber="27">Distribution.</emphasis>
Only known from the
<typeStatus id="7C6C8877FFF0E6084D6BFC2A72B9BAB1" box="[608,656,916,942]" pageId="26" pageNumber="27">type</typeStatus>
locality, Andaman Sea, at
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<quantity id="642F9B30FFF0E6084CB7FC2A73D1BAB1" box="[956,1016,916,942]" metricMagnitude="1" metricUnit="m" metricValue="7.7" pageId="26" pageNumber="27" unit="m" value="77.0">77 m</quantity>
.
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<subSubSection id="EBCD655EFFF0E6084F9CFC06744DBD7D" pageId="26" pageNumber="27" type="discussion">
<paragraph id="A36836D5FFF0E6084F9CFC06744DBD7D" blockId="26.[151,1437,663,1122]" pageId="26" pageNumber="27">
<emphasis id="91A3EAC7FFF0E6084F9CFC067122BACD" bold="true" box="[151,267,952,978]" pageId="26" pageNumber="27">Remarks.</emphasis>
The species is only known by the
<typeStatus id="7C6C8877FFF0E6084D9CFC0972D0BACE" box="[663,761,951,977]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
male and it is characterized by the presence of 5 epigastric spines, 1 small hepatic spine, and 3 spines on the anterior branchial margin. The closest relative is
<taxonomicName id="64D74D56FFF0E6084BD3FC62716DBD05" authority="(Miyake &amp; Baba, 1965)" baseAuthorityName="Miyake &amp; Baba" baseAuthorityYear="1965" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ikedai">
<emphasis id="91A3EAC7FFF0E6084BD3FC62751EBAEA" box="[1240,1335,987,1013]" italics="true" pageId="26" pageNumber="27">P. ikedai</emphasis>
(
<bibRefCitation id="C7464B24FFF0E6084A4EFC657112BD05" author="Miyake, S. &amp; Baba, K." pageId="26" pageNumber="27" pagination="585 - 593" refId="ref118214" refString="Miyake, S. &amp; Baba, K. (1965) Some galatheids obtained from the Bonin Islands (Crustacea, Anomura). Journal of the Faculty of Agriculture, Kyushu University, 13, 585 - 593. https: // doi. org / 10.5109 / 22735" type="journal article" year="1965">Miyake &amp; Baba, 1965</bibRefCitation>
)
</taxonomicName>
from Bonin Islands,
<collectingCountry id="DBC07645FFF0E6084D35FBBE72A8BD05" box="[574,641,1024,1050]" name="Japan" pageId="26" pageNumber="27">Japan</collectingCountry>
. However, they can be easily differentiated by the shape of thoracic sternite 3, with the anterior margin nearly transverse, and median and lateral projections, in
<emphasis id="91A3EAC7FFF0E6084B9FFB9A7513BD22" box="[1172,1338,1059,1085]" italics="true" pageId="26" pageNumber="27">
<taxonomicName id="64D74D56FFF0E6084B9FFB9A751FBD22" authorityName="Tirmizi &amp; Javed" authorityYear="1980" box="[1172,1334,1059,1085]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="bengalensis">P. bengalensis</taxonomicName>
,
</emphasis>
whereas the sternite has the anterior margin convex, with feeble median excavation in
<taxonomicName id="64D74D56FFF0E6084CF6FBF67475BD7E" baseAuthorityName="Miyake &amp; Baba" baseAuthorityYear="1965" box="[1021,1116,1095,1121]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ikedai">
<emphasis id="91A3EAC7FFF0E6084CF6FBF67475BD7E" box="[1021,1116,1095,1121]" italics="true" pageId="26" pageNumber="27">P. ikedai</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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