436 lines
36 KiB
XML
436 lines
36 KiB
XML
<document id="D28A514A3C5910F6950757EAEDF4AA9A" ID-DOI="10.3897/zse.100.130929" ID-ZooBank="4D6C62E4-224B-4AB5-A251-AA7ABEE66D74" ID-publisher-id="130929" URI-arpha="62A0369D-D7AA-53B4-8EA3-723D47A8742A" URI-zoobank="http://zoobank.org/4D6C62E4-224B-4AB5-A251-AA7ABEE66D74" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1728062842149" checkinUser="pensoft" docAuthor="Jeratthitikul, Ekgachai, Sutcharit, Chirasak & Prasankok, Pongpun" docDate="2024" docId="6A0BC990AA2155A9A787499DACD59067" docLanguage="en" docName="Zoosystematics and Evolution 100(4): 1333-1345" docOrigin="Zoosystematics and Evolution 100 (4)" docSource="https://zse.pensoft.net/article/130929/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Lannanaia kokensis Jeratthitikul, Sutcharit & Prasankok, 2024, sp. nov." docType="treatment" docUuid="873CE4DB-F862-43DF-BFF7-8863B7CBA3C7" docUuidSource="ZooBank" docVersion="3" dtd-version="3.0" lastPageNumber="1345" masterDocId="62A0369DD7AA53B48EA3723D47A8742A" masterDocTitle="Two new genera and three new species of exceptionally rare and endemic freshwater mussels (Bivalvia, Unionidae) from the Mekong Basin" masterLastPageNumber="1345" masterPageNumber="1333" pageNumber="1333" updateTime="1728066362166" updateUser="admin">
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<mods:title id="C84D53CA43F05A6B3CD2BAD74CE7D868">Two new genera and three new species of exceptionally rare and endemic freshwater mussels (Bivalvia, Unionidae) from the Mekong Basin</mods:title>
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<mods:name id="AE6EF9463EAB57A3AF32DD3AB39A148B" type="personal">
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<mods:namePart id="892A6F20816007ACEEB4F146F1156996">Jeratthitikul, Ekgachai</mods:namePart>
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<mods:affiliation id="263F8B9E9CF90CFCD4C6B0C62D12779C">Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10400, Thailand</mods:affiliation>
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<mods:namePart id="C4E2CBE549090F7013BC15069CF1BB13">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="0C63A6A4521FCBD3242058C6A6769992">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="AE44E36EBEB2DC61F51EBB5C7378287B">Prasankok, Pongpun</mods:namePart>
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<mods:affiliation id="BE98529FAB9F048308C8913675694447">School of Biology, Institute of Science, Suranaree University of Technology, Nakhon Ratchasima 30000, Thailand</mods:affiliation>
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<mods:title id="E54CCE06B47F3DEC05D35A906BE75915">Zoosystematics and Evolution</mods:title>
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<mods:date id="D493C42426070F76FC923810302E5AD9">2024</mods:date>
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<mods:number id="9A69613735E098240F54B03A62925605">2024-10-04</mods:number>
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<mods:number id="1438E1E312D8DE789584174FCF8DA0B0">100</mods:number>
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<mods:identifier id="332044DB81AE1C4317807BA53C7955E4" type="ZooBank">4D6C62E4-224B-4AB5-A251-AA7ABEE66D74</mods:identifier>
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<treatment id="6A0BC990AA2155A9A787499DACD59067" ID-DOI="http://doi.org/10.5281/zenodo.13891547" ID-Zenodo-Dep="13891547" ID-arpha="6A0BC990-AA21-55A9-A787-499DACD59067" ID-zoobank="https://zoobank.org/873CE4DB-F862-43DF-BFF7-8863B7CBA3C7" LSID="urn:lsid:zoobank.org:act:873CE4DB-F862-43DF-BFF7-8863B7CBA3C7" LSID-ZBK="urn:lsid:zoobank.org:act:873CE4DB-F862-43DF-BFF7-8863B7CBA3C7" httpUri="http://treatment.plazi.org/id/6A0BC990AA2155A9A787499DACD59067">
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<subSubSection id="B8D9CA501B212253BC9D152C3258C427" type="nomenclature">
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<paragraph id="DA3327AAA4A3A3E73BC9F0A5B7E52E23">
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<taxonomicName id="F8EC63800F8643D8BE4EF606A09C1FDB" authorityName="Jeratthitikul & Sutcharit & Prasankok" authorityYear="2024" class="Bivalvia" family="Unionidae" genus="Lannanaia" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="kokensis" status="sp. nov.">
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<emphasis id="34F94184F6020EA0367171A10993BBDC" italics="true">Lannanaia kokensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="909045C2B90804CE044D52FDB8D5F7FF" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="FA098BE0B2C1D73B37A3554215EDB271" type="description">
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<paragraph id="C0F2D36D21B0E3953B23F8F6C6A56FA0">
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||
<figureCitation id="0BB56E00C52D137C2D76C8E806CCBD5C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Maximum likelihood (ML) tree of freshwater mussels in subfamily Gonideinae based on 1,935 - bp concatenated alignment dataset of COI and 16 S rRNA mitochondrial genes, and 28 S rRNA nuclear gene. The numbers on nodes represent the bootstrap support (BS) from the ML analysis and the bipartition posterior probability (bpp) from the BI analysis, and are shown as ML / BI. Scale bar indicates the branch length." figureDoi="10.3897/zse.100.130929.figure1" httpUri="https://binary.pensoft.net/fig/1148326">Figs 1</figureCitation>
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,
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<figureCitation id="26D5DCC2C98ED27B1566F786D2B155A3" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Shells of Lannanaia kokensis sp. nov., showing the inner side of the right valve, the outer side of the left valve, and the dorsal view of both valves. A. Holotype MUMNH - UNI 0825; B. Paratype MUMNH - UNI 0824. Scale bars: 10 mm." figureDoi="10.3897/zse.100.130929.figure2" httpUri="https://binary.pensoft.net/fig/1148327">2</figureCitation>
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,
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<figureCitation id="E9D351B2C06282E542B28C2202C1C31C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Map showing type localities of the new species: circle Lannanaia kokensis sp. nov., triangle Isannaia fortunata sp. nov., and square Isannaia occultata sp. nov. Map was generated using QGIS v 3.24. 3 with the outline of Mekong basin from the Freshwater Ecoregions of the World (Abell et al. 2008), river and lake topology from the HydroSHEDS database (https://www.hydrosheds.org), and the map raster data from the NASA EARTHDATA (https://www.earthdata.nasa.gov/)." figureDoi="10.3897/zse.100.130929.figure3" httpUri="https://binary.pensoft.net/fig/1148328">3</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="SECID0EVMAG" type="type material">
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<paragraph id="CB044A82F6CE83A4FBB664BF0071F60E">
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<heading id="579E52F95D72C572A74C19209BF713B9" reason="title">Type material.</heading>
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||
</paragraph>
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<paragraph id="84543462399B0595B32FAF924193F859">
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<materialsCitation id="F25E3FD0AAF50BE77D0BD029FE322C29" collectionCode="MUMNH" collectorName="E. Jeratthitikul & K. Wisittikoson" country="Thailand" county="Mueang Chiang Rai District" latitude="19.9131" location="Kok River" longLatPrecision="7" longitude="99.8697" municipality="Rop Wiang Subdistrict" specimenCode="MUMNH-UNI 0825" specimenCount="1" stateProvince="Chiang Rai Province" typeStatus="holotype">
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<emphasis id="B7DB38A15B6ED6D15C91C63A4BD6F563" bold="true">
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||
<emphasis id="B8B86423E9F7B817EEB9D3AE455656AC" bold="true" italics="true">
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<typeStatus id="67EB432A8663EC963883806E570350DA" type="holotype">Holotype</typeStatus>
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</emphasis>
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.
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</emphasis>
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<collectingCountry id="1BF6D8F44952B40F8EC19A7382382132" name="Thailand">Thailand</collectingCountry>
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•
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<collectingRegion id="8A7EEF840F8081C525F078BC7170BB31" country="Thailand" name="Chiang Rai">Chiang Rai Province</collectingRegion>
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,
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<collectingCounty id="295F46DE5832E3D3283760BCCB22CE57">Mueang Chiang Rai District</collectingCounty>
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,
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<collectingMunicipality id="D2282F6086D7D48D90B1469FFE8A26DC">Rop Wiang Subdistrict</collectingMunicipality>
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,
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||
<location id="17D31E957049E5371A58F4662BB65451" LSID="urn:lsid:plazi:treatment:6A0BC990AA2155A9A787499DACD59067:17D31E957049E5371A58F4662BB65451" country="Thailand" county="Mueang Chiang Rai District" latitude="19.9131" longLatPrecision="7" longitude="99.8697" municipality="Rop Wiang Subdistrict" name="Kok River" stateProvince="Chiang Rai Province">Kok River</location>
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;
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<named-content content-type="dwc:verbatimCoordinates" id="NCID0EDNAG" specific-use="{"type":"Point","coordinates":[99.869700,19.913100]}">
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||
<geoCoordinate id="B1D893227D4E285DE5BBF7D9AA969F03" degrees="19.9131" direction="north" orientation="latitude" precision="5" value="19.9131">19.9131 ° N</geoCoordinate>
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,
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||
<geoCoordinate id="924622C7760DCACD82D76181057DFF12" degrees="99.8697" direction="east" orientation="longitude" precision="5" value="99.8697">99.8697 ° E</geoCoordinate>
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</named-content>
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;
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<collectorName id="B8F19D6FC32695C3C39D1CF722D5C7AE">E. Jeratthitikul</collectorName>
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and
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<collectorName id="C370EB994C6001018E740178FE58E046">K. Wisittikoson</collectorName>
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leg.;
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<specimenCode id="E5880BB665A04E61247EA71BBBFE593A">
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||
<named-content id="8AB522022DE5471883EF7045D16E558B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/mahidol-university" xlink_title="Mahidol University Museum of Natural History, Department of Biology, Faculty of Science, Mahidol University, Bangkok, Thailand">MUMNH</named-content>
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-UNI 0825
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</specimenCode>
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; shell length
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<quantity id="6D91A842B340BA4E083DC9D110B7C835" metricMagnitude="-2" metricUnit="m" metricValue="5.72" unit="mm" value="57.2">57.2 mm</quantity>
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, shell height
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<quantity id="B35A01DE3DCB2C46CF55E0F086483120" metricMagnitude="-2" metricUnit="m" metricValue="4.08" unit="mm" value="40.8">40.8 mm</quantity>
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, and shell width
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<quantity id="92D3BDB6BD19A4D7C189581FB0A22151" metricMagnitude="-2" metricUnit="m" metricValue="1.44" unit="mm" value="14.4">14.4 mm</quantity>
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||
</materialsCitation>
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||
.
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<materialsCitation id="7A0ED01A8CF4895A7505439391C4C052" collectionCode="MUMNH" collectorName="E. Jeratthitikul & K. Wisittikoson" country="Thailand" county="Mueang Chiang Rai District" latitude="19.9131" location="Thailand" longLatPrecision="7" longitude="99.8697" municipality="Rop Wiang Subdistrict" specimenCode="MUMNH-UNI 0824" specimenCount="1" stateProvince="Chiang Rai" typeStatus="paratype">
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<emphasis id="3D721ECFC654A27FCAEF1185CEADDDAB" bold="true">
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||
<emphasis id="21DC25062F8C95AC18A205A47D30B2A5" bold="true" italics="true">
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<typeStatus id="B27077C026E39DE47C175E14069836EB" type="paratype">Paratype</typeStatus>
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</emphasis>
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.
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</emphasis>
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<collectingCountry id="6BB24BD2AA97A116B099FC3FA8649361" name="Thailand">Thailand</collectingCountry>
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• 1 shell; same collection data as for holotype;
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<specimenCode id="FF0BB07DBB899E010CE4ABFD1040A036">
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<named-content id="1CA796C311C15647C3BC885106D87A5D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/mahidol-university" xlink_title="Mahidol University Museum of Natural History, Department of Biology, Faculty of Science, Mahidol University, Bangkok, Thailand">MUMNH</named-content>
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-UNI 0824
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</specimenCode>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="SECID0EWNAG" type="description">
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<paragraph id="D939F7B953B2A554B4EE8CC16F3FD640">
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<heading id="BC4CA1284410FDA38E0C7A19E84B82D7" reason="title">Description.</heading>
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||
</paragraph>
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<paragraph id="7DA7D8809BEF2E714BA22404C5CE0C22">
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Shell medium-sized (shell length
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<quantity id="DC31D79015A7396508FA182892A9B701" metricMagnitude="-2" metricUnit="m" metricValue="5.715" metricValueMax="5.72" metricValueMin="5.71" unit="mm" value="57.15" valueMax="57.2" valueMin="57.1">57.1–57.2 mm</quantity>
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, shell height
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<quantity id="5F81B93C9283F962575362DE36CB2E9E" metricMagnitude="-2" metricUnit="m" metricValue="4.2299999999999995" metricValueMax="4.38" metricValueMin="4.08" unit="mm" value="42.3" valueMax="43.8" valueMin="40.8">40.8–43.8 mm</quantity>
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, shell width
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<quantity id="F470C27F73624069D58DC5FCAB508890" metricMagnitude="-2" metricUnit="m" metricValue="1.46" metricValueMax="1.48" metricValueMin="1.44" unit="mm" value="14.6" valueMax="14.8" valueMin="14.4">14.4–14.8 mm</quantity>
|
||
; Table
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<tableCitation id="92864F9E3CFB7F039C02493D268C5828" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Shell measurements and GenBank accession numbers for the type series of the new species described in this study. Measurements in millimetres (mm)." httpUri="http://table.plazi.org/id/2EDC89F0C67A5A15CACAB0A86B456858" tableUuid="2EDC89F0C67A5A15CACAB0A86B456858">2</tableCitation>
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||
), thin, rather short and high (H / L ratio = 0.71–0.77), winged, sub-trigonal outline, inequilateral, rather compressed. Anterior margin rounded; posterior margin angulate, somewhat pointed; ventral margin curved. Dorsal margin straight; anterior low and short, gradually elevated to posterior end; posterior end high and winged. Umbo tiny, not prominent, eroded. Posterior ridge wide and obtuse, not prominent; posterior slope with two fine lines running as curved line from umbo to posterior margin. Periostracum thin, greenish to dark brown, the eroded part white to coppery-brown. Shell surface with very fine growth lines. Ligament long, very narrow, dark brown. Hinge without dentition, or with very rudimentary broad pseudocardinal tooth in each valve; posterior end of hinge structure with rather broad V-shaped fossette. Anterior adductor muscle scar very shallow, ovate, contiguous with anterior pedal retractor muscle scar, but separated from anterior protractor scar; posterior adductor muscle scar round, very shallow, almost invisible. Pallial line very faint, continuous. Umbo cavity shallow. Nacre blue-whitish.
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</paragraph>
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<caption id="2EDC89F0C67A5A15CACAB0A86B456858" ID-Table-UUID="2EDC89F0C67A5A15CACAB0A86B456858" httpUri="http://table.plazi.org/id/2EDC89F0C67A5A15CACAB0A86B456858" startId="T2" targetIsTable="true">
|
||
<paragraph id="3950A18DC4AC117C294F57D3DB15DC09">
|
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<label id="CE107A683B57D0CC2C00F0DA414F7164">Table 2.</label>
|
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</paragraph>
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<paragraph id="9C5CA8EDFD31C20DCC272080CFA88AEF">Shell measurements and GenBank accession numbers for the type series of the new species described in this study. Measurements in millimetres (mm).</paragraph>
|
||
</caption>
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||
<paragraph id="243CA88DC1AB5EEB9925174CD6471EBB">
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||
<table id="TID0EOJBI" rules="all">
|
||
<tbody id="6F911D9DE9CB2D6284B1A6831A289595">
|
||
<tr id="39133741E14545E701773972201BE0EF">
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||
<th id="8B24934AA4BDC5CDDC1F8BD134A91A67" colspan="1" rowspan="2">Taxa</th>
|
||
<th id="5991FD5DCB19860E17520013FCD4D905" colspan="1" rowspan="2">Status of specimen</th>
|
||
<th id="185693CC4C64A5BF7A883AB9A64656C9" colspan="1" rowspan="2">Specimen voucher</th>
|
||
<th id="71F65DF077715037C6A7D422F2E9BC32" colspan="1" rowspan="2">Shell length</th>
|
||
<th id="10947A5A02C071BA7DD99D32E092CF28" colspan="1" rowspan="2">Shell height</th>
|
||
<th id="C14431C6D59DE2A6A95127FBE13AC2F2" colspan="1" rowspan="2">Shell width</th>
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<th id="346802A1DDBB57A9B5CB54C90CBA1667" colspan="3" rowspan="1">Genbank accession</th>
|
||
</tr>
|
||
<tr id="FAE1AB29F87EC6989617F21B14F5BDD6">
|
||
<th id="9AE49482DFEA8A993240846B24D4559A" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EHPAG" xlink_title="mitochondrial cytochrome c oxidase subunit-I gene">COI</abbrev>
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</th>
|
||
<th id="4F327652C9179F7FAD44678B7548D093" colspan="1" rowspan="1">16 S rRNA</th>
|
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<th id="9BA1A8B05BC32BD66D1A10CD9A0556BD" colspan="1" rowspan="1">28 S rRNA</th>
|
||
</tr>
|
||
<tr id="A9E41341F5C96991E461CA790DC20C6F">
|
||
<td id="41421CDC661F099E8B306B4379F7CD4B" colspan="1" rowspan="2">
|
||
<taxonomicName id="F560D50182A321576D045405659BDE0A" authorityName="Jeratthitikul & Sutcharit & Prasankok" authorityYear="2024" class="Bivalvia" family="Unionidae" genus="Lannanaia" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="kokensis" status="sp. nov.">
|
||
<emphasis id="AC6DD3B63D87A4AA95A35CFE9C239AB0" italics="true">Lannanaia kokensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="4F3D265EBE8743285D4E44E9356BC62E" bold="true">
|
||
<taxonomicNameLabel id="46E941E97E20A9C5A7327EEA12C466C0" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</td>
|
||
<td id="5E775D3E7D5D7E96E6C58B59A34EB223" colspan="1" rowspan="1">Holotype</td>
|
||
<td id="E5CF4CBC772EAC89C7607E53BAB6B7B6" colspan="1" rowspan="1">
|
||
<specimenCode id="A16FA952237CC32807FFDB25CDF70669">
|
||
<named-content id="73635AF2947ECE1AC43C372BC1FD7D57" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/mahidol-university" xlink_title="Mahidol University Museum of Natural History, Department of Biology, Faculty of Science, Mahidol University, Bangkok, Thailand">MUMNH</named-content>
|
||
-UNI 0825
|
||
</specimenCode>
|
||
</td>
|
||
<td id="F30A715A35A02C772C377638FC06F655" colspan="1" rowspan="1">57.2</td>
|
||
<td id="C03D929A489D4565C87A558366A1C798" colspan="1" rowspan="1">40.8</td>
|
||
<td id="378EEB8041EBA24129B1226699B018D7" colspan="1" rowspan="1">14.4</td>
|
||
<td id="5BC3A76196BA6BE0C8713C9C1D0719A7" colspan="1" rowspan="1" style="color: #710f10">
|
||
<ext-link id="1D7EA0DF5A10A898AD2415E37228F1B5" ext-link-type="gen" xlink_href="OR987589" xlink_type="simple">OR 987589</ext-link>
|
||
</td>
|
||
<td id="589398E893E8D3FA80A5D4D4A37D0E26" colspan="1" rowspan="1" style="color: #710f10">
|
||
<ext-link id="34D5DC48B5E66A15CF06FDCB744EA784" ext-link-type="gen" xlink_href="OR987515" xlink_type="simple">OR 987515</ext-link>
|
||
</td>
|
||
<td id="9076996B8F1CF48F56D8C2C3ACB39386" colspan="1" rowspan="1" style="color: #710f10">
|
||
<ext-link id="859552662107E656F7F4A76ACAA1933E" ext-link-type="gen" xlink_href="OR987524" xlink_type="simple">OR 987524</ext-link>
|
||
</td>
|
||
</tr>
|
||
<tr id="2F70B20A3B92BAF174A5962DBED535F6">
|
||
<td id="3FE02A91A78A801627BF90B8AF125D1E" colspan="1" rowspan="1">Paratype</td>
|
||
<td id="56CC5D72619CD0781522CD85B5597316" colspan="1" rowspan="1">
|
||
<specimenCode id="0628673BAB1E94A23C2604B4C6585F19">
|
||
<named-content id="1B4AFBF1300ACA47F01AD4F352BF3B5C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/mahidol-university" xlink_title="Mahidol University Museum of Natural History, Department of Biology, Faculty of Science, Mahidol University, Bangkok, Thailand">MUMNH</named-content>
|
||
-UNI 0824
|
||
</specimenCode>
|
||
</td>
|
||
<td id="87C0C88179409516BF3D71402A2AB61C" colspan="1" rowspan="1">57.1</td>
|
||
<td id="1A00F41B1D79CBC03B10677361F98B21" colspan="1" rowspan="1">43.8</td>
|
||
<td id="6AB85EC3B77D54F50719A659160894E9" colspan="1" rowspan="1">14.8</td>
|
||
<td id="E5E36F993F2F682C719A498A9439E4E5" colspan="1" rowspan="1" style="color: #710f10">
|
||
<ext-link id="D461FFC1129B3196265F50CE5A3FCFA4" ext-link-type="gen" xlink_href="OR987588" xlink_type="simple">OR 987588</ext-link>
|
||
</td>
|
||
<td id="4C1D85961CF985A42F83FA9AD597BA50" colspan="1" rowspan="1" style="color: #710f10">
|
||
<ext-link id="5DEC6723BA267824B88EAD6122517514" ext-link-type="gen" xlink_href="OR987514" xlink_type="simple">OR 987514</ext-link>
|
||
</td>
|
||
<td id="24478CF3EFE913C87C0C73158A9540F8" colspan="1" rowspan="1" style="color: #710f10">
|
||
<ext-link id="7462981165DF49A64685535720C6692B" ext-link-type="gen" xlink_href="OR987523" xlink_type="simple">OR 987523</ext-link>
|
||
</td>
|
||
</tr>
|
||
<tr id="1C7106C0C75DF3D3E155D5E9F7BD5FF8">
|
||
<td id="1BEF221A0B29551EFAAF7291C026115D" colspan="1" rowspan="4">
|
||
<taxonomicName id="A9ED12BCAFE8A8C0B603FA9E6B735BFA" authorityName="Jeratthitikul & Sutcharit & Prasankok" authorityYear="2024" class="Bivalvia" family="Unionidae" genus="Isannaia" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="fortunata" status="sp. nov.">
|
||
<emphasis id="2704951FA83C8879592E05AF0E877B46" italics="true">Isannaia fortunata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="912066D0BEF2BD2D4BF4316DC42C8DA1" bold="true">
|
||
<taxonomicNameLabel id="2BAC1D2851DCEB97981C04F130D0E0A6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</td>
|
||
<td id="49C784054CF0AA1120A46129704D1DBC" colspan="1" rowspan="1">Holotype</td>
|
||
<td id="5944C0BFFEC8CEC201DEE8A283CC9BF1" colspan="1" rowspan="1">
|
||
<specimenCode id="67B5EB2B8AB579797B0DDC4FBBD3C194">
|
||
<named-content id="B9D0FAF522B8FACE922E9B3518D4CD32" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/mahidol-university" xlink_title="Mahidol University Museum of Natural History, Department of Biology, Faculty of Science, Mahidol University, Bangkok, Thailand">MUMNH</named-content>
|
||
-UNI 1950
|
||
</specimenCode>
|
||
</td>
|
||
<td id="2E060A01E08E9FA447B74A5260C3191F" colspan="1" rowspan="1">54.1</td>
|
||
<td id="4E10D62C5DA8D8D3C267F0FCE4947146" colspan="1" rowspan="1">29.9</td>
|
||
<td id="AB92EEDF5E46B1C25F8FC70DC1C16854" colspan="1" rowspan="1">18.4</td>
|
||
<td id="B157D68474D59E8343CC71E861214F14" colspan="1" rowspan="1" style="color: #710f10">
|
||
<ext-link id="C3D9D3CBEA82C61E495EFA5F7D7945C2" ext-link-type="gen" xlink_href="OR987590" xlink_type="simple">OR 987590</ext-link>
|
||
</td>
|
||
<td id="8294D452A21FE48416021AD805CDE654" colspan="1" rowspan="1" style="color: #710f10">
|
||
<ext-link id="1428DEE9B1C9C53CFE5CE12F2CE60418" ext-link-type="gen" xlink_href="OR987516" xlink_type="simple">OR 987516</ext-link>
|
||
</td>
|
||
<td id="B5684202EA04A581A6A092C3E88C367A" colspan="1" rowspan="1" style="color: #710f10">
|
||
<ext-link id="BEC4606DC829DDCEB3ED8DD462680E51" ext-link-type="gen" xlink_href="OR987525" xlink_type="simple">OR 987525</ext-link>
|
||
</td>
|
||
</tr>
|
||
<tr id="A769F6DDE1682A59D7AE1B14072F9050">
|
||
<td id="6953FF1A549010411EA87707EB1CFAD5" colspan="1" rowspan="1">Paratype</td>
|
||
<td id="08B06C9DBDF345E737D9F9EC881D97DE" colspan="1" rowspan="1">
|
||
<specimenCode id="505E2B410435FB9C3020F26D9F87E256">
|
||
<named-content id="26B3DBEF69ABFAEC472650C290EFDFA9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/mahidol-university" xlink_title="Mahidol University Museum of Natural History, Department of Biology, Faculty of Science, Mahidol University, Bangkok, Thailand">MUMNH</named-content>
|
||
-UNI 1951
|
||
</specimenCode>
|
||
</td>
|
||
<td id="2F01E69F7495A90A8703D3749BFC3B9F" colspan="1" rowspan="1">45.5</td>
|
||
<td id="2CD2F3DF1B01C7F6BC675C3E6F8F785B" colspan="1" rowspan="1">25.6</td>
|
||
<td id="4DE57EFD1ABF79CF16CB191D09F616BD" colspan="1" rowspan="1">14.4</td>
|
||
<td id="9C6548527EF85325DC82D26E8D17146C" colspan="1" rowspan="1" style="color: #710f10">
|
||
<ext-link id="9760429352C3B6AF938EDDA4C6C695CE" ext-link-type="gen" xlink_href="OR987591" xlink_type="simple">OR 987591</ext-link>
|
||
</td>
|
||
<td id="64BA3C08E5609CBE5FFFE0EE393F8CDE" colspan="1" rowspan="1" style="color: #710f10">
|
||
<ext-link id="B757B1D5418949574EAB15965322987B" ext-link-type="gen" xlink_href="OR987517" xlink_type="simple">OR 987517</ext-link>
|
||
</td>
|
||
<td id="3501ABBC2F31173E54E073E4304D84E8" colspan="1" rowspan="1" style="color: #710f10">
|
||
<ext-link id="4E11A980DF063FBB1423E86550A09216" ext-link-type="gen" xlink_href="OR987526" xlink_type="simple">OR 987526</ext-link>
|
||
</td>
|
||
</tr>
|
||
<tr id="90739CCB79457337A41386DF85F17437">
|
||
<td id="B076091EB85C1D1D652D00F0C2CF04D7" colspan="1" rowspan="1">Paratype</td>
|
||
<td id="4317B1F7B14AB01C1C03517C8AEA73ED" colspan="1" rowspan="1">
|
||
<specimenCode id="56304C64A3B13528261D14371F886AA1">
|
||
<named-content id="5BB7C0A062384C3E366FF4D83FD24EB8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/mahidol-university" xlink_title="Mahidol University Museum of Natural History, Department of Biology, Faculty of Science, Mahidol University, Bangkok, Thailand">MUMNH</named-content>
|
||
-UNI 1952
|
||
</specimenCode>
|
||
</td>
|
||
<td id="7C054C049DBFF0B5F71F7B2797DE6AC7" colspan="1" rowspan="1">61.4</td>
|
||
<td id="6A9F69283EDC725DEF229E5EDE0566D8" colspan="1" rowspan="1">35.1</td>
|
||
<td id="DA92509BD7D2C57C5EC5331A372C9073" colspan="1" rowspan="1">21.6</td>
|
||
<td id="D60887AB4634D13D206E1CE5ED81BF1B" colspan="1" rowspan="1" style="color: #710f10">
|
||
<ext-link id="243093A4C2DBE5925ED8B7755DD9AF4A" ext-link-type="gen" xlink_href="OR987592" xlink_type="simple">OR 987592</ext-link>
|
||
</td>
|
||
<td id="DBF718DD92654B2B55C7EDB479642C2B" colspan="1" rowspan="1" style="color: #710f10">
|
||
<ext-link id="6E41C7D0A29EADC86250D7A575C3093F" ext-link-type="gen" xlink_href="OR987518" xlink_type="simple">OR 987518</ext-link>
|
||
</td>
|
||
<td id="4D70DF399B9A3B6DADC70113DF23B164" colspan="1" rowspan="1" style="color: #710f10">
|
||
<ext-link id="2FE343F389A0E2BF2888FA1208B1371B" ext-link-type="gen" xlink_href="OR987527" xlink_type="simple">OR 987527</ext-link>
|
||
</td>
|
||
</tr>
|
||
<tr id="171ACD1FD1B56702590F034C32E8724B">
|
||
<td id="7CD06440D87A2FE7F5D1569435AD6F73" colspan="1" rowspan="1">Paratype</td>
|
||
<td id="C0FA3B84E7A9EC1F128B956BCC47DFC1" colspan="1" rowspan="1">
|
||
<specimenCode id="0634EA573DAE0BAE713542976D3CC6A9">
|
||
<named-content id="C5837137127A48BF2F7E2B6701A4F44B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/mahidol-university" xlink_title="Mahidol University Museum of Natural History, Department of Biology, Faculty of Science, Mahidol University, Bangkok, Thailand">MUMNH</named-content>
|
||
-UNI 1953
|
||
</specimenCode>
|
||
</td>
|
||
<td id="305E5AA31C5A2396A24953C6854735A8" colspan="1" rowspan="1">43.3</td>
|
||
<td id="B8D980C93A81EE0DBCDDE73B9FC673DA" colspan="1" rowspan="1">24.5</td>
|
||
<td id="C2D195A74DA79AFD742F59E84A3033C3" colspan="1" rowspan="1">13.4</td>
|
||
<td id="1CCB7B44B66E9D5478F9F3E85ABDE861" colspan="1" rowspan="1" style="color: #710f10">
|
||
<ext-link id="4A3F4BBCE9D3615D2C545E8A7B00E32A" ext-link-type="gen" xlink_href="OR987593" xlink_type="simple">OR 987593</ext-link>
|
||
</td>
|
||
<td id="F7ED3E76008765751233A4EFBCC6F93F" colspan="1" rowspan="1" style="color: #710f10">
|
||
<ext-link id="7B47608022D2724827BC29AE81BCD660" ext-link-type="gen" xlink_href="OR987519" xlink_type="simple">OR 987519</ext-link>
|
||
</td>
|
||
<td id="88856C9E2357E3E2D566A624C4670EF1" colspan="1" rowspan="1" style="color: #710f10">
|
||
<ext-link id="FC07935D27DC69166CA70AD6BBF53B86" ext-link-type="gen" xlink_href="OR987528" xlink_type="simple">OR 987528</ext-link>
|
||
</td>
|
||
</tr>
|
||
<tr id="068642CBF54259E52354C3682FE90E6B">
|
||
<td id="3DB613D819E89387CD6C72C9A0AC2BCE" colspan="1" rowspan="3">
|
||
<taxonomicName id="9C49127C31C1E614A7C6FF908E134FD8" authorityName="Jeratthitikul & Sutcharit & Prasankok" authorityYear="2024" class="Bivalvia" family="Unionidae" genus="Isannaia" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="occultata" status="sp. nov.">
|
||
<emphasis id="9A9E8E5EABE6FD7FA50363B05FE55ABF" italics="true">Isannaia occultata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A43EA56D241DC0BCA88D704EBD7C9179" bold="true">
|
||
<taxonomicNameLabel id="EBC16217ECF6823832BC4B677FE21A2F" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</td>
|
||
<td id="6A857C8929EE8AA1CA5BE88C4C98E4CE" colspan="1" rowspan="1">Holotype</td>
|
||
<td id="47473CA048128D6068E6B8CFE0F476D8" colspan="1" rowspan="1">
|
||
<specimenCode id="7B5FD5765094F909A4C71B897C6659A2">
|
||
<named-content id="2825769E718C040FE7A603626925EF94" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/mahidol-university" xlink_title="Mahidol University Museum of Natural History, Department of Biology, Faculty of Science, Mahidol University, Bangkok, Thailand">MUMNH</named-content>
|
||
-UNI 2895
|
||
</specimenCode>
|
||
</td>
|
||
<td id="5145216E69A9405F57D45E88207ACC2F" colspan="1" rowspan="1">53.8</td>
|
||
<td id="1FD0895416ADEBD1F17B764EA0AC9691" colspan="1" rowspan="1">31.7</td>
|
||
<td id="A9682DD4BDC231A89D24C5B583E7FA85" colspan="1" rowspan="1">20.4</td>
|
||
<td id="52201BF4198BC01A1F3AF4CC901CE39A" colspan="1" rowspan="1" style="color: #710f10">
|
||
<ext-link id="D841A5BDF7E1FA68719FB66A44DB77A3" ext-link-type="gen" xlink_href="OR987594" xlink_type="simple">OR 987594</ext-link>
|
||
</td>
|
||
<td id="95D0693A36C59D0A3B08921B89BE062F" colspan="1" rowspan="1" style="color: #710f10">
|
||
<ext-link id="AD8F1B89D3C5660B1CF46A2FFAA73672" ext-link-type="gen" xlink_href="OR987520" xlink_type="simple">OR 987520</ext-link>
|
||
</td>
|
||
<td id="AD0F7CC13CFF4C2DAA28CCA39CACB8E5" colspan="1" rowspan="1" style="color: #710f10">
|
||
<ext-link id="9E39E72B8679B183368B8CDB1F359CA3" ext-link-type="gen" xlink_href="OR987529" xlink_type="simple">OR 987529</ext-link>
|
||
</td>
|
||
</tr>
|
||
<tr id="A3D67E5F1FB78B4B46890D5270025F70">
|
||
<td id="07181E109DE371EF81BF7B28DE9A0A62" colspan="1" rowspan="1">Paratype</td>
|
||
<td id="2D72E9DC6F0E1247C72853B192047672" colspan="1" rowspan="1">
|
||
<specimenCode id="5CDFA6B70AFB02D9BD4BB86B0461333F">
|
||
<named-content id="7ABABE4AA66618FB7CD4D56FCDED4DDB" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/mahidol-university" xlink_title="Mahidol University Museum of Natural History, Department of Biology, Faculty of Science, Mahidol University, Bangkok, Thailand">MUMNH</named-content>
|
||
-UNI 2896
|
||
</specimenCode>
|
||
</td>
|
||
<td id="1F845D187339ABDBAB4B6C953FC51B7D" colspan="1" rowspan="1">43.8</td>
|
||
<td id="BEF8EC51229B52492AED4D00008A24D7" colspan="1" rowspan="1">26.7</td>
|
||
<td id="CEB7D63B2F7C5BECFD025D9BF8816563" colspan="1" rowspan="1">13.8</td>
|
||
<td id="AF13C19B7926F72C9B82CEA84BFE9DDD" colspan="1" rowspan="1" style="color: #710f10">
|
||
<ext-link id="079AA81D7D27613C6934648410870C0B" ext-link-type="gen" xlink_href="OR987595" xlink_type="simple">OR 987595</ext-link>
|
||
</td>
|
||
<td id="D75EA6EC4E7BB330E902C478A8A60BCA" colspan="1" rowspan="1" style="color: #710f10">
|
||
<ext-link id="29A95968BC0910E746479F773F0B473A" ext-link-type="gen" xlink_href="OR987521" xlink_type="simple">OR 987521</ext-link>
|
||
</td>
|
||
<td id="C724E2A730BB186FB4048EBA4D366D45" colspan="1" rowspan="1" style="color: #710f10">
|
||
<ext-link id="39DB3C59DC991D8769B36A8F41ACB88C" ext-link-type="gen" xlink_href="OR987530" xlink_type="simple">OR 987530</ext-link>
|
||
</td>
|
||
</tr>
|
||
<tr id="5067D02F3F0EB3342AB199F35E92C134">
|
||
<td id="3741CD5E99D132371155DE361F3ED4DB" colspan="1" rowspan="1">Paratype</td>
|
||
<td id="27F9D7862F85D7059127E98D8C923CF8" colspan="1" rowspan="1">
|
||
<specimenCode id="2EB710DFBABD4125B0181CB9CE883736">
|
||
<named-content id="C8759A5D97B8406A643CBB82B615C957" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/mahidol-university" xlink_title="Mahidol University Museum of Natural History, Department of Biology, Faculty of Science, Mahidol University, Bangkok, Thailand">MUMNH</named-content>
|
||
-UNI 2897
|
||
</specimenCode>
|
||
</td>
|
||
<td id="9A2FFE0488A4352DE30646998DA1AF32" colspan="1" rowspan="1">47.9</td>
|
||
<td id="E80BE647904253145F13E4B7192A0A20" colspan="1" rowspan="1">27.9</td>
|
||
<td id="7E809F28798FDEFAA8C9CB0D3945E527" colspan="1" rowspan="1">14.8</td>
|
||
<td id="E47C717042EAC0FD7BED213BE75825D1" colspan="1" rowspan="1" style="color: #710f10">
|
||
<ext-link id="55017CF11BE8B39F8DC7E8BFE7012E28" ext-link-type="gen" xlink_href="OR987596" xlink_type="simple">OR 987596</ext-link>
|
||
</td>
|
||
<td id="9407F0645B9295E982A99DD15AF6A3EE" colspan="1" rowspan="1" style="color: #710f10">
|
||
<ext-link id="055EBAE8736E41504831407DBCA66744" ext-link-type="gen" xlink_href="OR987522" xlink_type="simple">OR 987522</ext-link>
|
||
</td>
|
||
<td id="3E1FE59B46C6027EED4473E7C678EA56" colspan="1" rowspan="1" style="color: #710f10">
|
||
<ext-link id="FE942080A8E43733FC64DF17EE6637B3" ext-link-type="gen" xlink_href="OR987531" xlink_type="simple">OR 987531</ext-link>
|
||
</td>
|
||
</tr>
|
||
</tbody>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="3106FF8954F9D24A01DE5128FF14E004">Exhalant aperture smooth, shorter than inhalant. Inhalant aperture with a row of conical papillae, varying in length. Small epithelial fold divides exhalant and inhalant apertures. Gills elongated and slightly ribbed; outer gills much narrower (about half) than inner gills; anterior margin of inner gills slightly longer than that of outer gills. Labial palps elongate, round at tip. Glochidia unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EW6AG" type="etymology">
|
||
<paragraph id="177ACF31433AF2643DDEFCAC0B94DDF8">
|
||
<heading id="6AA45AB6236DCF5102EE7AD08E483DF5" reason="title">Etymology.</heading>
|
||
</paragraph>
|
||
<paragraph id="2F7A846E92FAC64063A9E42B07C92FA9">
|
||
The species name “
|
||
<emphasis id="BDAA4749CF1A2FBE1515600B25714DA4" italics="true">kokensis</emphasis>
|
||
” refers to the type locality, the Kok River in
|
||
<collectingRegion id="55F81C540EA346175FFCF01DD133B1CB" country="Thailand" name="Chiang Rai">Chiang Rai Province</collectingRegion>
|
||
, northern
|
||
<collectingCountry id="BA626D59AE16CCA2D50134F691605133" name="Thailand">Thailand</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EGABG" type="distribution">
|
||
<paragraph id="CC419C773160DAC812163E1CF64981DA">
|
||
<heading id="CC168FA1C35161DAC522B1ABF0165C3C" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="37AC31059E697497ED4FD8C6BE780266">This new species is known only from the type locality.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ELABG" type="remarks">
|
||
<paragraph id="8C3AE560811C13B77CB1CF7E4CB0D65F">
|
||
<heading id="CE0AC38D5BDDF630EDA10381845CF325" reason="title">Remarks.</heading>
|
||
</paragraph>
|
||
<paragraph id="96B0DA0A574818D18A518BF99802DD05">
|
||
Two living specimens were collected from a small stream that is a tributary to the Kok River, with a muddy bottom. We searched multiple times and locations for this species but failed to find additional specimens. In contrast,
|
||
<taxonomicName id="1DA1F642D01EA47A40BD856519AB3C9F" authority="(Lea, 1838)" baseAuthorityName="Lea" baseAuthorityYear="1838" class="Bivalvia" family="Unionidae" genus="Lens" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="contradens">
|
||
<emphasis id="1F0C4B627C6EA9DFD2BE2F1782EA75ED" italics="true">Lens contradens</emphasis>
|
||
(
|
||
<bibRefCitation id="6190391D35A7D7583A97BE24B12A9FD8" DOI="10.2307/1005319" author="Lea I" journalOrPublisher="Transactions of the American Philosophical Society" pagination="1-154" refId="B30" refString="Lea I (1838) Description of new freshwater and land shells. Transactions of the American Philosophical Society 6: 1 - 154. https: // doi. org / 10.2307 / 1005319" title="Description of new freshwater and land shells." volume="6" year="1838">Lea, 1838</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="25B3DA720626709533420FCAFA2EA867" authority="(Lea, 1838)" baseAuthorityName="Lea" baseAuthorityYear="1838" class="Bivalvia" family="Unionidae" genus="Pilsbryoconcha" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="exilis">
|
||
<emphasis id="7625206259AE7EB9263A4C3BA6F37802" italics="true">Pilsbryoconcha exilis</emphasis>
|
||
(
|
||
<bibRefCitation id="CB8693E3D4FBCFFFC977C15502CC42FE" DOI="10.2307/1005319" author="Lea I" journalOrPublisher="Transactions of the American Philosophical Society" pagination="1-154" refId="B30" refString="Lea I (1838) Description of new freshwater and land shells. Transactions of the American Philosophical Society 6: 1 - 154. https: // doi. org / 10.2307 / 1005319" title="Description of new freshwater and land shells." volume="6" year="1838">Lea, 1838</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
were found nearby in high abundance, both living and as empty shells. We assume this new species has a low population density and is possibly endemic to the Kok River Basin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |