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<document id="8CB077357290662B33567478BA2E2228" ID-CLB-Dataset="5601" ID-DOI="10.11646/zootaxa.3796.2.8" ID-GBIF-Dataset="72f6fdb3-a9cb-4823-ada5-233a4510b0db" ID-ISSN="1175-5326" ID-Zenodo-Dep="4915243" ID-ZooBank="9F9DA323-128B-44A7-9414-798B50DAAC44" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623199009682" checkinUser="felipe" docAuthor="Maruyama, Munetoshi, Komatsu, Takashi, Katayama, Yuji, Song, Xiao-Bin &amp; Sakchoowong, Watana" docDate="2014" docId="470287817148FFB67E91FAFBA864FD5D" docLanguage="en" docName="zt03796p373.pdf" docOrigin="Zootaxa 3796 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aenictosymbia cornuta Maruyama 2014, sp. n." docType="treatment" docVersion="6" lastPageNumber="369" masterDocId="BB3BFFF9714CFFBE7E06FFECAA48FFF5" masterDocTitle="Myrmecophilous rove beetles (Coleoptera: Staphylinidae) associated with Aenictus hodgsoni (Hymenoptera: Formicidae) from Thailand, with description of two new genera and three new species" masterLastPageNumber="373" masterPageNumber="361" pageNumber="365" updateTime="1699006430599" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="B9DBB0DBB8DDF7F3D51715A06CCC4193" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E628D8D4AA98CD73E99BCD83C211A1D0">
<mods:title id="3A0372A0FD09A2685EE93072D1C0FD4D">Myrmecophilous rove beetles (Coleoptera: Staphylinidae) associated with Aenictus hodgsoni (Hymenoptera: Formicidae) from Thailand, with description of two new genera and three new species</mods:title>
</mods:titleInfo>
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<mods:roleTerm id="CC1CF1E25EA9F3FB37207CE9C72E52B9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B63B0448A431D8E04D713355B7F0E950">Maruyama, Munetoshi</mods:namePart>
<mods:affiliation id="17F83FEB32467D79FE269DC875D043AB">The Kyushu University Museum, Hakozaki 6 - 10 - 1, Higashi-ku, Fukuoka-shi, Fukuoka, 812 - 8581 Japan</mods:affiliation>
<mods:nameIdentifier id="A75C3A4388711125C3B770D3199C470F" type="email">dendrolasius@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="B5B2B9276D1F4E69E30604E40643775D" type="personal">
<mods:role id="06138BE055804CFDF08AB40870670B41">
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</mods:role>
<mods:namePart id="495CF5B612F0544188E6B977F2F42975">Komatsu, Takashi</mods:namePart>
<mods:affiliation id="CD3FBFCC84798F6F16A2E1DFB82A05D1">Department of Biology, Faculty of Science, Shinshu University, 3 - 1 - 1 Asahi, Matsumoto, Nagano 390 - 8621, Japan</mods:affiliation>
</mods:name>
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<mods:roleTerm id="61EE81118726641D59324ADEA31BC1C7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1B60B740D8AAF620C0AE92A675E143D1">Katayama, Yuji</mods:namePart>
<mods:affiliation id="EF7B303CF8E14D8015A4F6318B57FB7C">Entomological Laboratory, Faculty of Agriculture, Ehime University, Tarumi, Matsuyama 790 - 8566, Japan</mods:affiliation>
</mods:name>
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<mods:role id="6FB432F34123387B64E446DCEE8BA32E">
<mods:roleTerm id="428846FF72B68E8A3F71C84AA7BE3E78">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E69F1C5F5DF00DC0FB354EF87888D539">Song, Xiao-Bin</mods:namePart>
<mods:affiliation id="2EF66FAAF6145DB4941B2DFF76715B53">Department of Biology, College of Life and Environmental Sciences, Shanghai Normal University, 100 Guilin Road, Shanghai, 200234, P. R. China</mods:affiliation>
</mods:name>
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<mods:role id="C783EB4E5546233F67ED2F0219AE02EA">
<mods:roleTerm id="5F22DE36F9CD45FB64482178CB67D4AF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A04E0F5C36B8D88FD7A4B0C72905FD7C">Sakchoowong, Watana</mods:namePart>
<mods:affiliation id="F3B330A036C764D6169EC7828586FF46">Forest Entomology and Microbiology Group, Department of National Parks, Wildlife and Plant Conservation, 61 Paholyothin Road, Jatujak, Bangkok 10900, Thailand</mods:affiliation>
</mods:name>
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<mods:title id="A5644B0438AA535B53EFB947573CAAFE">Zootaxa</mods:title>
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<mods:part id="F83802408D81A4F611A00C985BC1B931">
<mods:date id="FE68AA99844B39C9FB306CD5982DDB9C">2014</mods:date>
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<mods:number id="36C957243C813B19940D7A22C78B5FC0">2014-05-19</mods:number>
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<mods:number id="E34C91B8F36B926C463BD64939FC3F21">3796</mods:number>
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<mods:classification id="206DA379DC71ED2C6A633D331C0DB084">journal article</mods:classification>
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<mods:identifier id="185DCAAA549CC195CA9D1992DEEFC4B4" type="DOI">10.11646/zootaxa.3796.2.8</mods:identifier>
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<treatment id="470287817148FFB67E91FAFBA864FD5D" ID-DOI="http://doi.org/10.5281/zenodo.4915261" ID-GBIF-Taxon="183403954" ID-Zenodo-Dep="4915261" LSID="urn:lsid:plazi:treatment:470287817148FFB67E91FAFBA864FD5D" httpUri="http://treatment.plazi.org/id/470287817148FFB67E91FAFBA864FD5D" lastPageId="8" lastPageNumber="369" pageId="4" pageNumber="365">
<subSubSection id="87B1651C7148FFBA7E91FAFBA833FAC5" box="[151,635,1303,1328]" pageId="4" pageNumber="365" type="nomenclature">
<paragraph id="CF1436977148FFBA7E91FAFBA833FAC5" blockId="4.[151,635,1303,1365]" box="[151,635,1303,1328]" pageId="4" pageNumber="365">
<heading id="945C81FB7148FFBA7E91FAFBA833FAC5" bold="true" box="[151,635,1303,1328]" fontSize="10" level="2" pageId="4" pageNumber="365" reason="2">
<emphasis id="FDDFEA857148FFBA7E91FAFBA833FAC5" bold="true" box="[151,635,1303,1328]" pageId="4" pageNumber="365">
<taxonomicName id="08AB4D147148FFBA7E91FAFBA867FAC5" ID-CoL="659TB" authority="Maruyama" authorityName="Maruyama" authorityYear="2014" box="[151,559,1303,1328]" class="Insecta" family="Staphylinidae" genus="Aenictosymbia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="365" phylum="Arthropoda" rank="species" species="cornuta" status="sp. nov.">
<emphasis id="FDDFEA857148FFBA7E91FAFBABE9FADA" bold="true" box="[151,417,1303,1328]" italics="true" pageId="4" pageNumber="365">Aenictosymbia cornuta</emphasis>
Maruyama
</taxonomicName>
,
<taxonomicNameLabel id="E6EC57FE7148FFBA7C3AFAFBA833FAC5" box="[572,635,1303,1328]" pageId="4" pageNumber="365" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="87B1651C7148FFBA7E91FAD0AB66FAA0" box="[151,302,1340,1365]" pageId="4" pageNumber="365" type="description">
<paragraph id="CF1436977148FFBA7E91FAD0AB66FAA0" blockId="4.[151,635,1303,1365]" box="[151,302,1340,1365]" pageId="4" pageNumber="365">
(
<figureCitation id="57902A127148FFBA7E99FAD0AB6EFAA0" box="[159,294,1340,1365]" captionStart-0="FIGURES 10–12" captionStart-1="FIGURES 13–23" captionStart-2="FIGURES 24–32" captionStartId-0="5.[151,263,1891,1914]" captionStartId-1="6.[151,263,1837,1860]" captionStartId-2="7.[151,263,1952,1975]" captionTargetBox-0="[189,1408,208,1848]" captionTargetBox-1="[151,1422,204,1800]" captionTargetBox-2="[161,1430,212,1917]" captionTargetId-0="figure-39@5.[151,1436,193,1870]" captionTargetId-1="figure-66@6.[151,1436,193,1817]" captionTargetId-2="figure-65@7.[151,1436,193,1932]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURES 10–12. Habitus of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 10―dorsal view; 11―ventral view; 12―lateral view." captionText-1="FIGURES 13–23. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 13―head, dorsal view; 14―labrum (right, epipharynx); 15―left mandible; 16―memtum; 17―labium; 18, right maxilla; 19―pronotum, dorsal view; 20―ditto, ventral view; 21―meso- and meta traces; 22―right elytra; 23―left fore leg." captionText-2="FIGURES 24–32. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 24―left mid leg; 25―left hind leg; 26―male tergite VIII; 27―male sternite VIII; 28―male abdominal tergites IX and X; 29―median lobe of aedeagus, lateral view; 30―ditto, apical part, ventra view; 31―paramere; 32―spermatheca." figureDoi-0="http://doi.org/10.5281/zenodo.4915251" figureDoi-1="http://doi.org/10.5281/zenodo.4915255" figureDoi-2="http://doi.org/10.5281/zenodo.4915259" httpUri-0="https://zenodo.org/record/4915251/files/figure.png" httpUri-1="https://zenodo.org/record/4915255/files/figure.png" httpUri-2="https://zenodo.org/record/4915259/files/figure.png" pageId="4" pageNumber="365">Figs. 10–32</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="87B1651C7148FFBA7E91FA68A949F9D8" pageId="4" pageNumber="365" type="materials_examined">
<paragraph id="CF1436977148FFBA7E91FA68A949F9D8" blockId="4.[151,1437,1412,1903]" pageId="4" pageNumber="365">
<emphasis id="FDDFEA857148FFBA7E91FA68AB6BFA68" bold="true" box="[151,291,1412,1437]" inLineHeading="true" pageId="4" pageNumber="365" reason="1">Type series.</emphasis>
<materialsCitation id="7FC33CCA7148FFBA7F2CFA68AB7EF9FD" ID-GBIF-Occurrence="3319798302" collectingDate="2007-09-30" collectionCode="THAI, T, KUM" determinerName="Maruyama" elevation="750" location="Heaw Suwat" municipality="Khao Yai National Park" pageId="4" pageNumber="365" specimenCount="1" specimenCount-male="1" stateProvince="Nakhon Ratchasima" typeStatus="holotype">
<typeStatus id="101088357148FFBA7F2CFA68ABDDFA68" box="[298,405,1412,1437]" pageId="4" pageNumber="365" type="holotype">Holotype</typeStatus>
,
<specimenCount id="D9ADFD1E7148FFBA7FA7FA68ABF0FA6E" box="[417,440,1412,1435]" pageId="4" pageNumber="365" type="male">♂</specimenCount>
, &quot;
<collectionCode id="A9BAAE527148FFBA7FCBFA68A85FFA68" box="[461,535,1412,1437]" pageId="4" pageNumber="365">THAI</collectionCode>
:
<collectingRegion id="0D6FF8757148FFBA7C24FA68A940FA68" box="[546,776,1412,1437]" country="Thailand" name="Nakhon Ratchasima" pageId="4" pageNumber="365">Nakhon Ratchasima</collectingRegion>
, /
<collectingMunicipality id="2F70ACED7148FFBA7D24FA68AE7AFA68" box="[802,1074,1412,1437]" pageId="4" pageNumber="365">Khao Yai National Park</collectingMunicipality>
, /
<location id="CA74604C7148FFBA7A4BFA68AE94FA68" LSID="urn:lsid:plazi:treatment:470287817148FFB67E91FAFBA864FD5D:CA74604C7148FFBA7A4BFA68AE94FA68" box="[1101,1244,1412,1437]" municipality="Khao Yai National Park" name="Heaw Suwat" pageId="4" pageNumber="365" stateProvince="Nakhon Ratchasima">Heaw Suwat</location>
(
<quantity id="08539B727148FFBA7AEDFA68AF7FFA68" box="[1259,1335,1412,1437]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="365" unit="m" value="750.0">
<elevation id="4486D1A47148FFBA7AEDFA68AF7FFA68" box="[1259,1335,1412,1437]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="365" unit="m" value="750.0">750 m</elevation>
</quantity>
), /
<date id="BB1510577148FFBA7B5DFA68AA9AFA35" pageId="4" pageNumber="365" value="2007-09-30">
<collectingDate id="AB51E9BF7148FFBA7B5DFA68AA9AFA35" pageId="4" pageNumber="365" value="2007-09-30">30 IX 2007</collectingDate>
</date>
, Maruyama M., / Komatsu
<collectionCode id="A9BAAE527148FFBA7C00FA44A851FA4A" box="[518,537,1448,1471]" name="Tavera, Department of Geology and Geophysics" pageId="4" pageNumber="365">T</collectionCode>
. &amp; Katayama Y. / MM-AE029 //
<typeStatus id="101088357148FFBA7D89FA4BAE6BFA35" box="[911,1059,1447,1472]" pageId="4" pageNumber="365" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="08AB4D147148FFBA7A3EFA44AEA9FA35" authorityName="Maruyama" authorityYear="2014" box="[1080,1249,1448,1472]" class="Insecta" family="Staphylinidae" genus="Aenictoxenides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="FDDFEA857148FFBA7A3EFA44AEA9FA35" box="[1080,1249,1448,1472]" italics="true" pageId="4" pageNumber="365">Aenictoxenides</emphasis>
</taxonomicName>
/
<taxonomicName id="08AB4D147148FFBA7AFEFA44AF15FA35" authorityName="Maruyama" authorityYear="2014" box="[1272,1373,1448,1472]" class="Insecta" family="Staphylinidae" genus="Aenictoxenides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="365" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="FDDFEA857148FFBA7AFEFA44AF15FA35" box="[1272,1373,1448,1472]" italics="true" pageId="4" pageNumber="365">mirabilis</emphasis>
</taxonomicName>
/ det.
<determinerName id="3AC8DA537148FFBA7E91FA20AB5BFA11" box="[151,275,1484,1508]" pageId="4" pageNumber="365">Maruyama</determinerName>
, 2014&quot; (male 8th abdominal segment and aedeagus dissected and glued on paper card together with body) (
<collectionCode id="A9BAAE527148FFBA7EE1FA03AB66F9FD" box="[231,302,1519,1544]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="4" pageNumber="365" type="Herbarium">KUM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="7FC33CCA7148FFBA7F47FA1CA8B5F9D8" ID-GBIF-Occurrence="3319798307" collectingDate="2007-10-01" collectionCode="THAI, KUM, DNP" elevation="700" location="Kong Kaew" municipality="Khao Yai National Park" pageId="4" pageNumber="365" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Nakhon Ratchasima" typeStatus="paratype">
<typeStatus id="101088357148FFBA7F47FA1CABF9F9FD" box="[321,433,1520,1544]" pageId="4" pageNumber="365" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="D9ADFD1E7148FFBA7FBAFA03ABA8F9FD" box="[444,480,1519,1544]" pageId="4" pageNumber="365" type="male">1♂</specimenCount>
,
<specimenCount id="D9ADFD1E7148FFBA7FEAFA03A858F9FD" box="[492,528,1519,1544]" pageId="4" pageNumber="365" type="female">3♀</specimenCount>
, &quot;
<collectionCode id="A9BAAE527148FFBA7C20FA03A838F9FD" box="[550,624,1519,1544]" pageId="4" pageNumber="365">THAI</collectionCode>
:
<collectingRegion id="0D6FF8757148FFBA7C7CFA03A928F9FD" box="[634,864,1519,1544]" country="Thailand" name="Nakhon Ratchasima" pageId="4" pageNumber="365">Nakhon Ratchasima</collectingRegion>
, /
<collectingMunicipality id="2F70ACED7148FFBA7D7FFA03AECEF9FD" box="[889,1158,1519,1544]" pageId="4" pageNumber="365">Khao Yai National Park</collectingMunicipality>
, /
<location id="CA74604C7148FFBA7AA6FA1CAF6EF9F2" LSID="urn:lsid:plazi:treatment:470287817148FFB67E91FAFBA864FD5D:CA74604C7148FFBA7AA6FA1CAF6EF9F2" box="[1184,1318,1520,1544]" municipality="Khao Yai National Park" name="Kong Kaew" pageId="4" pageNumber="365" stateProvince="Nakhon Ratchasima">Kong Kaew</location>
(
<quantity id="08539B727148FFBA7B33FA03AFC8F9FD" box="[1333,1408,1519,1544]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="365" unit="m" value="700.0">
<elevation id="4486D1A47148FFBA7B33FA03AFC8F9FD" box="[1333,1408,1519,1544]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="365" unit="m" value="700.0">700 m</elevation>
</quantity>
), /
<date id="BB1510577148FFBA7E91F9F8AB4AF9D8" box="[151,258,1556,1581]" pageId="4" pageNumber="365" value="2007-10-01">
<collectingDate id="AB51E9BF7148FFBA7E91F9F8AB4AF9D8" box="[151,258,1556,1581]" pageId="4" pageNumber="365" value="2007-10-01">1 X 2007</collectingDate>
</date>
, Maruyama M. / MM-AE032&quot; (
<collectionCode id="A9BAAE527148FFBA7C61F9F8A8E5F9D8" box="[615,685,1556,1581]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="4" pageNumber="365" type="Herbarium">KUM</collectionCode>
,
<collectionCode id="A9BAAE527148FFBA7CBEF9F8A8BDF9D8" box="[696,757,1556,1581]" pageId="4" pageNumber="365">DNP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="87B1651C7148FFB67EC1F9DBA855FE25" lastPageId="8" lastPageNumber="369" pageId="4" pageNumber="365" type="description">
<paragraph id="CF1436977148FFBA7EC1F9DBAAA1F980" blockId="4.[151,1437,1412,1903]" pageId="4" pageNumber="365">
<emphasis id="FDDFEA857148FFBA7EC1F9DBAB11F9A5" bold="true" box="[199,345,1591,1616]" inLineHeading="true" pageId="4" pageNumber="365" reason="1">Description.</emphasis>
Body (
<figureCitation id="57902A127148FFBA7FB1F9DBA80BF9A5" box="[439,579,1591,1616]" captionStart="FIGURES 10–12" captionStartId="5.[151,263,1891,1914]" captionTargetBox="[189,1408,208,1848]" captionTargetId="figure-39@5.[151,1436,193,1870]" captionTargetPageId="5" captionText="FIGURES 10–12. Habitus of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 10―dorsal view; 11―ventral view; 12―lateral view." figureDoi="http://doi.org/10.5281/zenodo.4915251" httpUri="https://zenodo.org/record/4915251/files/figure.png" pageId="4" pageNumber="365">Figs. 10–12</figureCitation>
) small. Reddish brown, but head and elytra darker; mouthparts reddish yellow.
</paragraph>
<paragraph id="CF1436977148FFBA7EC1F993AB60F8F0" blockId="4.[151,1437,1412,1903]" pageId="4" pageNumber="365">
Head (
<figureCitation id="57902A127148FFBA7F14F993ABD1F96D" box="[274,409,1663,1688]" captionStart-0="FIGURES 10–12" captionStart-1="FIGURES 13–23" captionStartId-0="5.[151,263,1891,1914]" captionStartId-1="6.[151,263,1837,1860]" captionTargetBox-0="[189,1408,208,1848]" captionTargetBox-1="[151,1422,204,1800]" captionTargetId-0="figure-39@5.[151,1436,193,1870]" captionTargetId-1="figure-66@6.[151,1436,193,1817]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 10–12. Habitus of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 10―dorsal view; 11―ventral view; 12―lateral view." captionText-1="FIGURES 13–23. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 13―head, dorsal view; 14―labrum (right, epipharynx); 15―left mandible; 16―memtum; 17―labium; 18, right maxilla; 19―pronotum, dorsal view; 20―ditto, ventral view; 21―meso- and meta traces; 22―right elytra; 23―left fore leg." figureDoi-0="http://doi.org/10.5281/zenodo.4915251" figureDoi-1="http://doi.org/10.5281/zenodo.4915255" httpUri-0="https://zenodo.org/record/4915251/files/figure.png" httpUri-1="https://zenodo.org/record/4915255/files/figure.png" pageId="4" pageNumber="365">Figs. 10–14</figureCitation>
) with horn truncate at apex, with 12–15 macrosetae on dorsal disc. Antennae (
<figureCitation id="57902A127148FFBA7B0BF993AFDCF96D" box="[1293,1428,1663,1688]" captionStart-0="FIGURES 10–12" captionStart-1="FIGURES 13–23" captionStartId-0="5.[151,263,1891,1914]" captionStartId-1="6.[151,263,1837,1860]" captionTargetBox-0="[189,1408,208,1848]" captionTargetBox-1="[151,1422,204,1800]" captionTargetId-0="figure-39@5.[151,1436,193,1870]" captionTargetId-1="figure-66@6.[151,1436,193,1817]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 10–12. Habitus of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 10―dorsal view; 11―ventral view; 12―lateral view." captionText-1="FIGURES 13–23. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 13―head, dorsal view; 14―labrum (right, epipharynx); 15―left mandible; 16―memtum; 17―labium; 18, right maxilla; 19―pronotum, dorsal view; 20―ditto, ventral view; 21―meso- and meta traces; 22―right elytra; 23―left fore leg." figureDoi-0="http://doi.org/10.5281/zenodo.4915251" figureDoi-1="http://doi.org/10.5281/zenodo.4915255" httpUri-0="https://zenodo.org/record/4915251/files/figure.png" httpUri-1="https://zenodo.org/record/4915255/files/figure.png" pageId="4" pageNumber="365">Figs. 10–13</figureCitation>
) slender, with all segments longer than wide; segment I shorter than II and III combined; segment II slightly shorter than III; segment IV–X equally in length but becoming wider toward apex; segment XI slightly shorter than IX and X combined.
</paragraph>
<paragraph id="CF1436977148FFBA7EC1F8FCAA9EF89A" blockId="4.[151,1437,1412,1903]" pageId="4" pageNumber="365">
Pronotum (
<figureCitation id="57902A127148FFBA7F43F8E3A86AF8DD" box="[325,546,1807,1832]" captionStart-0="FIGURES 10–12" captionStart-1="FIGURES 13–23" captionStartId-0="5.[151,263,1891,1914]" captionStartId-1="6.[151,263,1837,1860]" captionTargetBox-0="[189,1408,208,1848]" captionTargetBox-1="[151,1422,204,1800]" captionTargetId-0="figure-39@5.[151,1436,193,1870]" captionTargetId-1="figure-66@6.[151,1436,193,1817]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 10–12. Habitus of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 10―dorsal view; 11―ventral view; 12―lateral view." captionText-1="FIGURES 13–23. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 13―head, dorsal view; 14―labrum (right, epipharynx); 15―left mandible; 16―memtum; 17―labium; 18, right maxilla; 19―pronotum, dorsal view; 20―ditto, ventral view; 21―meso- and meta traces; 22―right elytra; 23―left fore leg." figureDoi-0="http://doi.org/10.5281/zenodo.4915251" figureDoi-1="http://doi.org/10.5281/zenodo.4915255" httpUri-0="https://zenodo.org/record/4915251/files/figure.png" httpUri-1="https://zenodo.org/record/4915255/files/figure.png" pageId="4" pageNumber="365">Figs. 10–13, 19, 20</figureCitation>
) constricted at basal 1/3; PL/PW= 1.53, with 12–14 macrosetae on dorsal disc and lateral margin. Elytra (
<figureCitation id="57902A127148FFBA7FC9F8D8A8CDF8B8" box="[463,645,1844,1869]" captionStart-0="FIGURES 10–12" captionStart-1="FIGURES 13–23" captionStartId-0="5.[151,263,1891,1914]" captionStartId-1="6.[151,263,1837,1860]" captionTargetBox-0="[189,1408,208,1848]" captionTargetBox-1="[151,1422,204,1800]" captionTargetId-0="figure-39@5.[151,1436,193,1870]" captionTargetId-1="figure-66@6.[151,1436,193,1817]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 10–12. Habitus of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 10―dorsal view; 11―ventral view; 12―lateral view." captionText-1="FIGURES 13–23. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 13―head, dorsal view; 14―labrum (right, epipharynx); 15―left mandible; 16―memtum; 17―labium; 18, right maxilla; 19―pronotum, dorsal view; 20―ditto, ventral view; 21―meso- and meta traces; 22―right elytra; 23―left fore leg." figureDoi-0="http://doi.org/10.5281/zenodo.4915251" figureDoi-1="http://doi.org/10.5281/zenodo.4915255" httpUri-0="https://zenodo.org/record/4915251/files/figure.png" httpUri-1="https://zenodo.org/record/4915255/files/figure.png" pageId="4" pageNumber="365">Figs. 10–13, 22</figureCitation>
) almost as long as pronotum, with 3 macrosetae and several minute setae.
</paragraph>
<caption id="9BD4661F7149FFBB7E91F88FAB0EF86C" ID-DOI="http://doi.org/10.5281/zenodo.4915251" ID-Zenodo-Dep="4915251" httpUri="https://zenodo.org/record/4915251/files/figure.png" pageId="5" pageNumber="366" startId="5.[151,263,1891,1914]" targetBox="[189,1408,208,1848]" targetPageId="5">
<paragraph id="CF1436977149FFBB7E91F88FAB0EF86C" blockId="5.[151,1436,1891,1945]" pageId="5" pageNumber="366">
<emphasis id="FDDFEA857149FFBB7E91F88FAB13F88C" bold="true" box="[151,347,1891,1914]" pageId="5" pageNumber="366">FIGURES 10–12.</emphasis>
Habitus of
<taxonomicName id="08AB4D147149FFBB7FEFF888A911F88F" authority="Maruyama" authorityName="Maruyama" authorityYear="2014" box="[489,857,1892,1914]" class="Insecta" family="Staphylinidae" genus="Aenictosymbia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="FDDFEA857149FFBB7FEFF888A893F88F" box="[489,731,1892,1914]" italics="true" pageId="5" pageNumber="366">Aenictosymbia cornuta</emphasis>
Maruyama
</taxonomicName>
,
<emphasis id="FDDFEA857149FFBB7D6AF888AE42F88C" bold="true" box="[876,1034,1892,1914]" pageId="5" pageNumber="366">gen. &amp; sp. n.</emphasis>
10―dorsal view; 11―ventral view; 12―lateral view.
</paragraph>
</caption>
<caption id="9BD4661F714AFFB87E91F8C1A91EF874" ID-DOI="http://doi.org/10.5281/zenodo.4915255" ID-Zenodo-Dep="4915255" httpUri="https://zenodo.org/record/4915255/files/figure.png" pageId="6" pageNumber="367" startId="6.[151,263,1837,1860]" targetBox="[151,1422,204,1800]" targetPageId="6">
<paragraph id="CF143697714AFFB87E91F8C1A91EF874" blockId="6.[151,1436,1837,1921]" pageId="6" pageNumber="367">
<emphasis id="FDDFEA85714AFFB87E91F8C1AB1CF8B6" bold="true" box="[151,340,1837,1860]" pageId="6" pageNumber="367">FIGURES 13–23.</emphasis>
Body parts of
<taxonomicName id="08AB4D14714AFFB87FEAF8C2A906F8B1" authority="Maruyama" authorityName="Maruyama" authorityYear="2014" box="[492,846,1838,1860]" class="Insecta" family="Staphylinidae" genus="Aenictosymbia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="367" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="FDDFEA85714AFFB87FEAF8C2A89FF8B1" box="[492,727,1838,1860]" italics="true" pageId="6" pageNumber="367">Aenictosymbia cornuta</emphasis>
Maruyama
</taxonomicName>
,
<emphasis id="FDDFEA85714AFFB87D5FF8C3A9AAF8B1" bold="true" box="[857,994,1838,1860]" pageId="6" pageNumber="367">gen. &amp; sp. n.</emphasis>
13―head, dorsal view; 14―labrum (right, epipharynx); 15―left mandible; 16―memtum; 17―labium; 18, right maxilla; 19―pronotum, dorsal view; 20―ditto, ventral view; 21―meso- and meta traces; 22―right elytra; 23―left fore leg.
</paragraph>
</caption>
<caption id="9BD4661F714BFFB97E91F84CA9DCF801" ID-DOI="http://doi.org/10.5281/zenodo.4915259" ID-Zenodo-Dep="4915259" httpUri="https://zenodo.org/record/4915259/files/figure.png" pageId="7" pageNumber="368" startId="7.[151,263,1952,1975]" targetBox="[161,1430,212,1917]" targetPageId="7">
<paragraph id="CF143697714BFFB97E91F84CA9DCF801" blockId="7.[151,1436,1952,2036]" pageId="7" pageNumber="368">
<emphasis id="FDDFEA85714BFFB97E91F84CAB10F843" bold="true" box="[151,344,1952,1975]" pageId="7" pageNumber="368">FIGURES 24–32.</emphasis>
Body parts of
<taxonomicName id="08AB4D14714BFFB97C07F84DA923F842" authority="Maruyama" authorityName="Maruyama" authorityYear="2014" box="[513,875,1953,1975]" class="Insecta" family="Staphylinidae" genus="Aenictosymbia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="368" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="FDDFEA85714BFFB97C07F84DA8B9F842" box="[513,753,1953,1975]" italics="true" pageId="7" pageNumber="368">Aenictosymbia cornuta</emphasis>
Maruyama
</taxonomicName>
,
<emphasis id="FDDFEA85714BFFB97D7DF84DAE58F843" bold="true" box="[891,1040,1953,1975]" pageId="7" pageNumber="368">gen. &amp; sp. n.</emphasis>
24―left mid leg; 25―left hind leg; 26―male tergite VIII; 27―male sternite VIII; 28―male abdominal tergites IX and X; 29―median lobe of aedeagus, lateral view; 30―ditto, apical part, ventra view; 31―paramere; 32―spermatheca.
</paragraph>
</caption>
<paragraph id="CF1436977144FFB67EC1FF7BAED2FF0D" blockId="8.[151,1436,151,680]" pageId="8" pageNumber="369">
Abdomen with convexity of sternite III (
<figureCitation id="57902A127144FFB67C96FF7BA8A8FF45" box="[656,736,151,176]" captionStart="FIGURES 10–12" captionStartId="5.[151,263,1891,1914]" captionTargetBox="[189,1408,208,1848]" captionTargetId="figure-39@5.[151,1436,193,1870]" captionTargetPageId="5" captionText="FIGURES 10–12. Habitus of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 10―dorsal view; 11―ventral view; 12―lateral view." figureDoi="http://doi.org/10.5281/zenodo.4915251" httpUri="https://zenodo.org/record/4915251/files/figure.png" pageId="8" pageNumber="369">Fig. 11</figureCitation>
) large, flattened ventrally, slightly notched on anterior margin in ventral view, shallowly depressed in midline; tergite VIII (
<figureCitation id="57902A127144FFB67D4DFF50A9D6FF20" box="[843,926,188,213]" captionStart="FIGURES 24–32" captionStartId="7.[151,263,1952,1975]" captionTargetBox="[161,1430,212,1917]" captionTargetId="figure-65@7.[151,1436,193,1932]" captionTargetPageId="7" captionText="FIGURES 24–32. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 24―left mid leg; 25―left hind leg; 26―male tergite VIII; 27―male sternite VIII; 28―male abdominal tergites IX and X; 29―median lobe of aedeagus, lateral view; 30―ditto, apical part, ventra view; 31―paramere; 32―spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4915259" httpUri="https://zenodo.org/record/4915259/files/figure.png" pageId="8" pageNumber="369">Fig. 26</figureCitation>
) shallowly emarginated at apex, with 4 short macrosetae; sternite VIII (
<figureCitation id="57902A127144FFB67FBBFF33A846FF0D" box="[445,526,223,248]" captionStart="FIGURES 24–32" captionStartId="7.[151,263,1952,1975]" captionTargetBox="[161,1430,212,1917]" captionTargetId="figure-65@7.[151,1436,193,1932]" captionTargetPageId="7" captionText="FIGURES 24–32. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 24―left mid leg; 25―left hind leg; 26―male tergite VIII; 27―male sternite VIII; 28―male abdominal tergites IX and X; 29―median lobe of aedeagus, lateral view; 30―ditto, apical part, ventra view; 31―paramere; 32―spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4915259" httpUri="https://zenodo.org/record/4915259/files/figure.png" pageId="8" pageNumber="369">Fig. 27</figureCitation>
) with 5 macrosetae and 2 or 3 thin setae postero medially.
</paragraph>
<paragraph id="CF1436977144FFB67EC1FEE8A98DFECA" blockId="8.[151,1436,151,680]" pageId="8" pageNumber="369">
Male: Median lobe of aedeagus (
<figureCitation id="57902A127144FFB67C39FEE8A88EFEE8" box="[575,710,260,285]" captionStart="FIGURES 24–32" captionStartId="7.[151,263,1952,1975]" captionTargetBox="[161,1430,212,1917]" captionTargetId="figure-65@7.[151,1436,193,1932]" captionTargetPageId="7" captionText="FIGURES 24–32. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 24―left mid leg; 25―left hind leg; 26―male tergite VIII; 27―male sternite VIII; 28―male abdominal tergites IX and X; 29―median lobe of aedeagus, lateral view; 30―ditto, apical part, ventra view; 31―paramere; 32―spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4915259" httpUri="https://zenodo.org/record/4915259/files/figure.png" pageId="8" pageNumber="369">Figs. 29–30</figureCitation>
) with basal ridge small, serrate at apex; sclerites of internal sac dentate; apical lobe of paramere (
<figureCitation id="57902A127144FFB67C0BFECBA817FEB5" box="[525,607,295,320]" captionStart="FIGURES 24–32" captionStartId="7.[151,263,1952,1975]" captionTargetBox="[161,1430,212,1917]" captionTargetId="figure-65@7.[151,1436,193,1932]" captionTargetPageId="7" captionText="FIGURES 24–32. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 24―left mid leg; 25―left hind leg; 26―male tergite VIII; 27―male sternite VIII; 28―male abdominal tergites IX and X; 29―median lobe of aedeagus, lateral view; 30―ditto, apical part, ventra view; 31―paramere; 32―spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4915259" httpUri="https://zenodo.org/record/4915259/files/figure.png" pageId="8" pageNumber="369">Fig. 31</figureCitation>
) long, thick, with 4 small setae.
</paragraph>
<paragraph id="CF1436977144FFB67EC1FEA0AA9BFE72" blockId="8.[151,1436,151,680]" pageId="8" pageNumber="369">
Female: Spermatheca (
<figureCitation id="57902A127144FFB67FCEFEA0A851FE90" box="[456,537,332,357]" captionStart="FIGURES 24–32" captionStartId="7.[151,263,1952,1975]" captionTargetBox="[161,1430,212,1917]" captionTargetId="figure-65@7.[151,1436,193,1932]" captionTargetPageId="7" captionText="FIGURES 24–32. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 24―left mid leg; 25―left hind leg; 26―male tergite VIII; 27―male sternite VIII; 28―male abdominal tergites IX and X; 29―median lobe of aedeagus, lateral view; 30―ditto, apical part, ventra view; 31―paramere; 32―spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4915259" httpUri="https://zenodo.org/record/4915259/files/figure.png" pageId="8" pageNumber="369">Fig. 32</figureCitation>
) with basal part about 3 times as long as apical part, which is slightly swollen at apex.
</paragraph>
<paragraph id="CF1436977144FFB67EC1FE78A855FE25" blockId="8.[151,1436,151,680]" pageId="8" pageNumber="369">
<emphasis id="FDDFEA857144FFB67EC1FE78AB35FE58" bold="true" box="[199,381,404,429]" inLineHeading="true" pageId="8" pageNumber="369" reason="1">Measurements.</emphasis>
BL, ca. 2.6
<emphasis id="FDDFEA857144FFB67C17FE78A857FE58" bold="true" box="[529,543,404,429]" pageId="8" pageNumber="369">–</emphasis>
2.7; FBL, ca. 1.6
<emphasis id="FDDFEA857144FFB67CEBFE78A8B3FE58" bold="true" box="[749,763,404,429]" pageId="8" pageNumber="369">–</emphasis>
1.7; HW, 0.47
<emphasis id="FDDFEA857144FFB67DA2FE78A9FAFE58" bold="true" box="[932,946,404,429]" pageId="8" pageNumber="369">–</emphasis>
0.53; AL, 1.55
<emphasis id="FDDFEA857144FFB67A67FE78AE27FE58" bold="true" box="[1121,1135,404,429]" pageId="8" pageNumber="369">–</emphasis>
1.63; PL, 0.65
<emphasis id="FDDFEA857144FFB67B1CFE78AF60FE58" bold="true" box="[1306,1320,404,429]" pageId="8" pageNumber="369">–</emphasis>
0.67; PW, 0.40
<emphasis id="FDDFEA857144FFB67ECEFE5BAA9EFE25" bold="true" box="[200,214,439,464]" pageId="8" pageNumber="369">–</emphasis>
0.41; HTL, 0.99
<emphasis id="FDDFEA857144FFB67F8DFE5BABD1FE25" bold="true" box="[395,409,439,464]" pageId="8" pageNumber="369">–</emphasis>
1.02 (N=3).
</paragraph>
</subSubSection>
<subSubSection id="87B1651C7144FFB67EC1FE30AF68FDED" pageId="8" pageNumber="369" type="diagnosis">
<paragraph id="CF1436977144FFB67EC1FE30AF68FDED" blockId="8.[151,1436,151,680]" pageId="8" pageNumber="369">
<emphasis id="FDDFEA857144FFB67EC1FE30AB90FE00" bold="true" box="[199,472,476,501]" inLineHeading="true" pageId="8" pageNumber="369" reason="1">Differential diagnosis.</emphasis>
This species is similar to
<taxonomicName id="08AB4D147144FFB67D3EFE30AF59FE00" authority="Seevers, 1965" authorityName="Seevers" authorityYear="1965" box="[824,1297,476,501]" class="Insecta" family="Staphylinidae" genus="Aenictophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="369" phylum="Arthropoda" rank="species" species="thailandica">
<emphasis id="FDDFEA857144FFB67D3EFE30AE11FE01" box="[824,1113,476,500]" italics="true" pageId="8" pageNumber="369">Aenictophila thailandica</emphasis>
<bibRefCitation id="AB3A4B667144FFB67A6CFE30AF59FE00" author="Seevers, C. H." box="[1130,1297,476,501]" pageId="8" pageNumber="369" pagination="138 - 351" refId="ref5738" refString="Seevers, C. H. (1965) The systematics, evolution and zoogeography of staphylinid beetles associated with army ants (Coleoptera: Staphylinidae). Fieldiana: Zoology, 47, 138 - 351." type="journal article" year="1965">
Seevers,
<quantity id="08539B727144FFB67ADFFE30AF59FE00" box="[1241,1297,476,501]" metricMagnitude="1" metricUnit="m" metricValue="4.9911" pageId="8" pageNumber="369" unit="in" value="1965.0">1965</quantity>
</bibRefCitation>
</taxonomicName>
in general appearance, but easily distinguished from it by the much smaller body. See also Diagnosis of the genus.
</paragraph>
</subSubSection>
<subSubSection id="87B1651C7144FFB67EC1FDC8A812FD76" pageId="8" pageNumber="369" type="etymology">
<paragraph id="CF1436977144FFB67EC1FDC8AA9BFD95" blockId="8.[151,1436,151,680]" pageId="8" pageNumber="369">
<emphasis id="FDDFEA857144FFB67EC1FDC8AB06FDC8" bold="true" box="[199,334,548,573]" inLineHeading="true" pageId="8" pageNumber="369" reason="1">Etymology.</emphasis>
The Latin adjective
<taxonomicName id="08AB4D147144FFB67C39FDC9A8DEFDC9" authorityName="Maruyama" authorityYear="2014" box="[575,662,549,572]" class="Insecta" family="Staphylinidae" genus="Aenictosymbia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="369" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="FDDFEA857144FFB67C39FDC9A8DEFDC9" box="[575,662,549,572]" italics="true" pageId="8" pageNumber="369">cornuta</emphasis>
</taxonomicName>
meaning &quot;horned&quot;, bringing attention to a horn on the frons of the head.
</paragraph>
<paragraph id="CF1436977144FFB67EC1FD80A812FD76" blockId="8.[151,1436,151,680]" box="[199,602,620,645]" pageId="8" pageNumber="369">
<emphasis id="FDDFEA857144FFB67EC1FD80AB36FD70" bold="true" box="[199,382,620,645]" inLineHeading="true" pageId="8" pageNumber="369" reason="1">Symbiotic host.</emphasis>
<taxonomicName id="08AB4D147144FFB67F83FD80A81AFD71" authorityName="Forel" authorityYear="1901" box="[389,594,620,644]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="369" phylum="Arthropoda" rank="species" species="hodgsoni">
<emphasis id="FDDFEA857144FFB67F83FD80A81AFD71" box="[389,594,620,644]" italics="true" pageId="8" pageNumber="369">Aenictus hodgsoni</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="87B1651C7144FFB67EC1FD63A864FD5D" box="[199,556,655,680]" pageId="8" pageNumber="369" type="distribution">
<paragraph id="CF1436977144FFB67EC1FD63A864FD5D" blockId="8.[151,1436,151,680]" box="[199,556,655,680]" pageId="8" pageNumber="369">
<emphasis id="FDDFEA857144FFB67EC1FD63AB28FD5D" bold="true" box="[199,352,655,680]" inLineHeading="true" pageId="8" pageNumber="369" reason="1">Distribution.</emphasis>
Central
<collectingCountry id="B7BC76077144FFB67FC6FD63A86FFD5D" box="[448,551,655,680]" name="Thailand" pageId="8" pageNumber="369">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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