320 lines
51 KiB
XML
320 lines
51 KiB
XML
<document id="48ED4FEE95B5D6B8EDA4C87B090AF6B1" ID-CLB-Dataset="5601" ID-DOI="10.11646/zootaxa.3796.2.8" ID-GBIF-Dataset="72f6fdb3-a9cb-4823-ada5-233a4510b0db" ID-ISSN="1175-5326" ID-Zenodo-Dep="4915243" ID-ZooBank="9F9DA323-128B-44A7-9414-798B50DAAC44" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623199009682" checkinUser="felipe" docAuthor="Maruyama, Munetoshi, Komatsu, Takashi, Katayama, Yuji, Song, Xiao-Bin & Sakchoowong, Watana" docDate="2014" docId="470287817144FFB57E91FD10A8F0FA68" docLanguage="en" docName="zt03796p373.pdf" docOrigin="Zootaxa 3796 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aenictoxenides Maruyama 2014, gen. n." docType="treatment" docVersion="7" lastPageNumber="372" masterDocId="BB3BFFF9714CFFBE7E06FFECAA48FFF5" masterDocTitle="Myrmecophilous rove beetles (Coleoptera: Staphylinidae) associated with Aenictus hodgsoni (Hymenoptera: Formicidae) from Thailand, with description of two new genera and three new species" masterLastPageNumber="373" masterPageNumber="361" pageNumber="369" updateTime="1699006430599" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="7542B3CB8EF44EB3FF0A80CC059B3CD2">
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<mods:title id="0B150E400F8569578CF64F2A829C0A7A">Myrmecophilous rove beetles (Coleoptera: Staphylinidae) associated with Aenictus hodgsoni (Hymenoptera: Formicidae) from Thailand, with description of two new genera and three new species</mods:title>
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<mods:name id="333CD61435A8612BCECB1E66090A55C2" type="personal">
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<mods:roleTerm id="E15ADC830424504D4AB75264B66141E1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0ED3C04CB32432E945DCA75DB17B83E2">Maruyama, Munetoshi</mods:namePart>
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<mods:affiliation id="FC71E11A34D9C9A1E6C4746773296B69">The Kyushu University Museum, Hakozaki 6 - 10 - 1, Higashi-ku, Fukuoka-shi, Fukuoka, 812 - 8581 Japan</mods:affiliation>
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<mods:nameIdentifier id="6AD4D9D931FC0F18546FBD0178D7FBB7" type="email">dendrolasius@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="36EF2CFF7B928E18CD7D17018961B4EE" type="personal">
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<mods:roleTerm id="0EBF1DFABE807F654661CD05F2887D76">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3678D074590439968C58B9F5CE5337D3">Komatsu, Takashi</mods:namePart>
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<mods:affiliation id="F6EDA7976C18EB91B73C20BD38BD0302">Department of Biology, Faculty of Science, Shinshu University, 3 - 1 - 1 Asahi, Matsumoto, Nagano 390 - 8621, Japan</mods:affiliation>
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</mods:name>
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<mods:name id="4A757E47F19B434ABA024E56A6069701" type="personal">
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<mods:role id="69FCA0EC4ACE2FDAE56CFDFB1AEBCCAA">
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<mods:roleTerm id="A05535F2EC51A6C79DB6B53C54615179">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="79EA5004DF960BD35C221017E1150533">Katayama, Yuji</mods:namePart>
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<mods:affiliation id="8AA0C12AB1A4BF21C8D0423C2820B070">Entomological Laboratory, Faculty of Agriculture, Ehime University, Tarumi, Matsuyama 790 - 8566, Japan</mods:affiliation>
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</mods:name>
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<mods:name id="B2F0E3E9C84A3DC81C90D4D18DADEF2C" type="personal">
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</mods:role>
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<mods:namePart id="3C116EAB7ED2C076D79E446709F29C9C">Song, Xiao-Bin</mods:namePart>
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<mods:affiliation id="AF982E287A2D49B35C46348623C24BF8">Department of Biology, College of Life and Environmental Sciences, Shanghai Normal University, 100 Guilin Road, Shanghai, 200234, P. R. China</mods:affiliation>
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<mods:name id="5B7B82C4842FC495DAFBFFDC125249F7" type="personal">
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</mods:role>
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<mods:namePart id="B075E4A50BB64B2BA0B9AC2D9CAB3752">Sakchoowong, Watana</mods:namePart>
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<mods:affiliation id="827B73CCE87610131644BF91C924C96F">Forest Entomology and Microbiology Group, Department of National Parks, Wildlife and Plant Conservation, 61 Paholyothin Road, Jatujak, Bangkok 10900, Thailand</mods:affiliation>
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<mods:typeOfResource id="922F141DC018F9E673B4132D23B230CD">text</mods:typeOfResource>
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<mods:title id="113BAD9F96A2682DB5198910FBD89A15">Zootaxa</mods:title>
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<mods:part id="3BD75AA9CEC26B7C4C7902139EC75341">
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<mods:date id="91EE65D42EAB6749B4E97344940ECBBD">2014</mods:date>
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<mods:number id="DC5DD29CA0C31A9FBBB7644D27C9B9AE">2014-05-19</mods:number>
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<mods:number id="5D8D470562F01CB50AA2131C09596531">3796</mods:number>
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<mods:identifier id="E1272D663D19C789FA0BB14317A87843" type="DOI">10.11646/zootaxa.3796.2.8</mods:identifier>
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<treatment id="470287817144FFB57E91FD10A8F0FA68" ID-DOI="http://doi.org/10.5281/zenodo.5083092" ID-GBIF-Taxon="183403955" ID-Zenodo-Dep="5083092" LSID="urn:lsid:plazi:treatment:470287817144FFB57E91FD10A8F0FA68" httpUri="http://treatment.plazi.org/id/470287817144FFB57E91FD10A8F0FA68" lastPageId="11" lastPageNumber="372" pageId="8" pageNumber="369">
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<subSubSection id="87B1651C7144FFB67E91FD10A80BFCE3" box="[151,579,764,790]" pageId="8" pageNumber="369" type="nomenclature">
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<paragraph id="CF1436977144FFB67E91FD10A80BFCE3" blockId="8.[151,579,764,823]" box="[151,579,764,790]" pageId="8" pageNumber="369">
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<heading id="945C81FB7144FFB67E91FD10A80BFCE3" bold="true" box="[151,579,764,790]" fontSize="11" level="1" pageId="8" pageNumber="369" reason="1">
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<emphasis id="FDDFEA857144FFB67E91FD10A80BFCE3" bold="true" box="[151,579,764,790]" pageId="8" pageNumber="369">
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<taxonomicName id="08AB4D147144FFB67E91FD10ABACFCE3" ID-CoL="8XJC5" authority="Maruyama" authorityName="Maruyama" authorityYear="2014" box="[151,484,764,790]" class="Insecta" family="Staphylinidae" genus="Aenictoxenides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="369" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="FDDFEA857144FFB67E91FD10AB07FCE3" bold="true" box="[151,335,764,790]" italics="true" pageId="8" pageNumber="369">Aenictoxenides</emphasis>
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Maruyama
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</taxonomicName>
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,
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<taxonomicNameLabel id="E6EC57FE7144FFB67FF7FD10A80BFCE3" box="[497,579,764,790]" pageId="8" pageNumber="369" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="87B1651C7144FFB67E91FCF2AB66FCC2" box="[151,302,798,823]" pageId="8" pageNumber="369" type="description">
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<paragraph id="CF1436977144FFB67E91FCF2AB66FCC2" blockId="8.[151,579,764,823]" box="[151,302,798,823]" pageId="8" pageNumber="369">
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(
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<figureCitation id="57902A127144FFB67E99FCF2AB6EFCC2" box="[159,294,798,823]" captionStart-0="FIGURES 33–35" captionStart-1="FIGURES 36–46" captionStart-2="FIGURES 47–54" captionStartId-0="8.[151,263,1970,1993]" captionStartId-1="9.[151,263,1952,1975]" captionStartId-2="10.[151,263,1518,1541]" captionTargetBox-0="[202,1381,1000,1927]" captionTargetBox-1="[151,1431,198,1926]" captionTargetBox-2="[172,1425,195,1494]" captionTargetId-0="figure-253@8.[151,1436,967,1948]" captionTargetId-1="figure-59@9.[151,1436,193,1932]" captionTargetId-2="figure-224@10.[159,1426,193,1503]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIGURES 33–35. Habitus of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 33―dorsal view; 34―ventral view; 35―lateral view." captionText-1="FIGURES 36–46. Body parts of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 36―head, dorsal view; 37―left antenna; 38―labrum; 39―left mandible; 40―memtum; 41―labium; 42, right maxilla; 43―pronotum, ventral view; 44―meso- and meta traces; 45―right elytra; 46―left fore leg." captionText-2="FIGURES 47–54. Body parts of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 47―left mid leg; 48―left hind leg; 49―male tergite VIII; 50―male sternite VIII; 51―male abdominal tergites IX and X; 52―median lobe of aedeagus, lateral view; 53―paramere; 54―spermatheca." figureDoi-0="http://doi.org/10.5281/zenodo.4915263" figureDoi-1="http://doi.org/10.5281/zenodo.4915267" figureDoi-2="http://doi.org/10.5281/zenodo.4915269" httpUri-0="https://zenodo.org/record/4915263/files/figure.png" httpUri-1="https://zenodo.org/record/4915267/files/figure.png" httpUri-2="https://zenodo.org/record/4915269/files/figure.png" pageId="8" pageNumber="369">Figs. 33–52</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="87B1651C7144FFB57E91FC8AAE6EFCCD" lastPageId="11" lastPageNumber="372" pageId="8" pageNumber="369" type="type_taxon">
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||
<paragraph id="CF1436977144FFB67E91FC8AA937FC56" blockId="8.[151,895,870,931]" pageId="8" pageNumber="369">
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||
<emphasis id="FDDFEA857144FFB67E91FC8AAB7CFC8A" bold="true" box="[151,308,870,895]" pageId="8" pageNumber="369">
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||
<typeStatus id="101088357144FFB67E91FC8AAA9AFC8A" box="[151,210,870,895]" pageId="8" pageNumber="369">Type</typeStatus>
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species:
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</emphasis>
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||
<taxonomicName id="08AB4D147144FFB67F3DFC8AA89CFC8A" ID-CoL="64XTG" authority="Maruyama" authorityName="Maruyama" authorityYear="2014" box="[315,724,870,895]" class="Insecta" family="Staphylinidae" genus="Aenictoxenides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="369" phylum="Arthropoda" rank="species" species="mirabilis" status="sp. nov.">
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<emphasis id="FDDFEA857144FFB67F3DFC8AA819FC8B" box="[315,593,870,894]" italics="true" pageId="8" pageNumber="369">Aenictoxenides mirabilis</emphasis>
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Maruyama
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</taxonomicName>
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,
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<emphasis id="FDDFEA857144FFB67CD9FC8AAB10FC56" bold="true" pageId="8" pageNumber="369">
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<taxonomicNameLabel id="E6EC57FE7144FFB67CD9FC8AA956FC8A" box="[735,798,870,895]" pageId="8" pageNumber="369" rank="species">sp. n.</taxonomicNameLabel>
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<emphasis id="FDDFEA857144FFB67EC1FC66AB10FC56" box="[199,344,906,931]" inLineHeading="true" pageId="8" pageNumber="369" reason="1">Description.</emphasis>
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</emphasis>
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Body (
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<figureCitation id="57902A127144FFB67FADFC66A87AFC56" box="[427,562,906,931]" captionStart="FIGURES 33–35" captionStartId="8.[151,263,1970,1993]" captionTargetBox="[202,1381,1000,1927]" captionTargetId="figure-253@8.[151,1436,967,1948]" captionTargetPageId="8" captionText="FIGURES 33–35. Habitus of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 33―dorsal view; 34―ventral view; 35―lateral view." figureDoi="http://doi.org/10.5281/zenodo.4915263" httpUri="https://zenodo.org/record/4915263/files/figure.png" pageId="8" pageNumber="369">Figs. 33–35</figureCitation>
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) foliaceous, flattened, glossy.
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</paragraph>
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<caption id="9BD4661F7144FFB67E91F85EAB0EF812" ID-DOI="http://doi.org/10.5281/zenodo.4915263" ID-Zenodo-Dep="4915263" httpUri="https://zenodo.org/record/4915263/files/figure.png" pageId="8" pageNumber="369" startId="8.[151,263,1970,1993]" targetBox="[202,1381,1000,1927]" targetPageId="8">
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<paragraph id="CF1436977144FFB67E91F85EAB0EF812" blockId="8.[151,1436,1970,2023]" pageId="8" pageNumber="369">
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||
<emphasis id="FDDFEA857144FFB67E91F85EAB12F832" bold="true" box="[151,346,1970,1993]" pageId="8" pageNumber="369">FIGURES 33–35.</emphasis>
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Habitus of
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<taxonomicName id="08AB4D147144FFB67FE2F85EA92BF83D" ID-CoL="64XTG" authority="Maruyama" authorityName="Maruyama" authorityYear="2014" box="[484,867,1970,1992]" class="Insecta" family="Staphylinidae" genus="Aenictoxenides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="369" phylum="Arthropoda" rank="species" species="mirabilis">
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<emphasis id="FDDFEA857144FFB67FE2F85EA8ADF83D" box="[484,741,1970,1992]" italics="true" pageId="8" pageNumber="369">Aenictoxenides mirabilis</emphasis>
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Maruyama
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||
</taxonomicName>
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,
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<emphasis id="FDDFEA857144FFB67D72F85FAE46F83D" bold="true" box="[884,1038,1970,1992]" pageId="8" pageNumber="369">gen. & sp. n.</emphasis>
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33―dorsal view; 34―ventral view; 35―lateral view.
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</paragraph>
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||
</caption>
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||
<caption id="9BD4661F7145FFB77E91F84CA83EF801" ID-DOI="http://doi.org/10.5281/zenodo.4915267" ID-Zenodo-Dep="4915267" httpUri="https://zenodo.org/record/4915267/files/figure.png" pageId="9" pageNumber="370" startId="9.[151,263,1952,1975]" targetBox="[151,1431,198,1926]" targetPageId="9">
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||
<paragraph id="CF1436977145FFB77E91F84CA83EF801" blockId="9.[151,1436,1952,2036]" pageId="9" pageNumber="370">
|
||
<emphasis id="FDDFEA857145FFB77E91F84CAB1CF843" bold="true" box="[151,340,1952,1975]" pageId="9" pageNumber="370">FIGURES 36–46.</emphasis>
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||
Body parts of
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||
<taxonomicName id="08AB4D147145FFB77FEAF84DA916F842" ID-CoL="64XTG" authority="Maruyama" authorityName="Maruyama" authorityYear="2014" box="[492,862,1953,1975]" class="Insecta" family="Staphylinidae" genus="Aenictoxenides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="370" phylum="Arthropoda" rank="species" species="mirabilis">
|
||
<emphasis id="FDDFEA857145FFB77FEAF84DA8A0F842" box="[492,744,1953,1975]" italics="true" pageId="9" pageNumber="370">Aenictoxenides mirabilis</emphasis>
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||
Maruyama
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||
</taxonomicName>
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||
,
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||
<emphasis id="FDDFEA857145FFB77D6FF84DA9B9F843" bold="true" box="[873,1009,1953,1975]" pageId="9" pageNumber="370">gen. & sp. n.</emphasis>
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36―head, dorsal view; 37―left antenna; 38―labrum; 39―left mandible; 40―memtum; 41―labium; 42, right maxilla; 43―pronotum, ventral view; 44―meso- and meta traces; 45―right elytra; 46―left fore leg.
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||
</paragraph>
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||
</caption>
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||
<caption id="9BD4661F7146FFB47E91FA02A864F9B7" ID-DOI="http://doi.org/10.5281/zenodo.4915269" ID-Zenodo-Dep="4915269" httpUri="https://zenodo.org/record/4915269/files/figure.png" pageId="10" pageNumber="371" startId="10.[151,263,1518,1541]" targetBox="[172,1425,195,1494]" targetPageId="10">
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||
<paragraph id="CF1436977146FFB47E91FA02A864F9B7" blockId="10.[151,1436,1518,1602]" pageId="10" pageNumber="371">
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||
<emphasis id="FDDFEA857146FFB47E91FA02AB1FF9F1" bold="true" box="[151,343,1518,1541]" pageId="10" pageNumber="371">FIGURES 47–54.</emphasis>
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||
Body parts of
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||
<taxonomicName id="08AB4D147146FFB47FFAFA02A93DF9F1" ID-CoL="64XTG" authority="Maruyama" authorityName="Maruyama" authorityYear="2014" box="[508,885,1518,1540]" class="Insecta" family="Staphylinidae" genus="Aenictoxenides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="371" phylum="Arthropoda" rank="species" species="mirabilis">
|
||
<emphasis id="FDDFEA857146FFB47FFAFA02A8B3F9F1" box="[508,763,1518,1540]" italics="true" pageId="10" pageNumber="371">Aenictoxenides mirabilis</emphasis>
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Maruyama
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||
</taxonomicName>
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,
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||
<emphasis id="FDDFEA857146FFB47D82FA03AE5EF9F1" bold="true" box="[900,1046,1518,1540]" pageId="10" pageNumber="371">gen. & sp. n.</emphasis>
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47―left mid leg; 48―left hind leg; 49―male tergite VIII; 50―male sternite VIII; 51―male abdominal tergites IX and X; 52―median lobe of aedeagus, lateral view; 53―paramere; 54―spermatheca.
|
||
</paragraph>
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||
</caption>
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||
<paragraph id="CF1436977146FFB47EC1F982AAA4F8E2" blockId="10.[151,1436,1646,2031]" pageId="10" pageNumber="371">
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Head (
|
||
<figureCitation id="57902A127146FFB47F1EF982ABECF972" box="[280,420,1646,1671]" captionStart-0="FIGURES 33–35" captionStart-1="FIGURES 36–46" captionStartId-0="8.[151,263,1970,1993]" captionStartId-1="9.[151,263,1952,1975]" captionTargetBox-0="[202,1381,1000,1927]" captionTargetBox-1="[151,1431,198,1926]" captionTargetId-0="figure-253@8.[151,1436,967,1948]" captionTargetId-1="figure-59@9.[151,1436,193,1932]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURES 33–35. Habitus of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 33―dorsal view; 34―ventral view; 35―lateral view." captionText-1="FIGURES 36–46. Body parts of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 36―head, dorsal view; 37―left antenna; 38―labrum; 39―left mandible; 40―memtum; 41―labium; 42, right maxilla; 43―pronotum, ventral view; 44―meso- and meta traces; 45―right elytra; 46―left fore leg." figureDoi-0="http://doi.org/10.5281/zenodo.4915263" figureDoi-1="http://doi.org/10.5281/zenodo.4915267" httpUri-0="https://zenodo.org/record/4915263/files/figure.png" httpUri-1="https://zenodo.org/record/4915267/files/figure.png" pageId="10" pageNumber="371">Figs. 33–36</figureCitation>
|
||
) semicircular, strongly flattened above; frons protruded anteriad and laterad to form “arcade” which conceals eyes, mouthparts, bases of antennae underside of head; temples projected laterad, pointed at apex; eyes rather large, located laterally, underside of head; antennae (
|
||
<figureCitation id="57902A127146FFB47DD8F95AAE2FF93A" box="[990,1127,1718,1743]" captionStart="FIGURES 33–35" captionStartId="8.[151,263,1970,1993]" captionTargetBox="[202,1381,1000,1927]" captionTargetId="figure-253@8.[151,1436,967,1948]" captionTargetPageId="8" captionText="FIGURES 33–35. Habitus of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 33―dorsal view; 34―ventral view; 35―lateral view." figureDoi="http://doi.org/10.5281/zenodo.4915263" httpUri="https://zenodo.org/record/4915263/files/figure.png" pageId="10" pageNumber="371">Figs. 33–35</figureCitation>
|
||
,
|
||
<figureCitation id="57902A127146FFB47A70F95AAEDBF93A" box="[1142,1171,1718,1743]" captionStart="FIGURES 36–46" captionStartId="9.[151,263,1952,1975]" captionTargetBox="[151,1431,198,1926]" captionTargetId="figure-59@9.[151,1436,193,1932]" captionTargetPageId="9" captionText="FIGURES 36–46. Body parts of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 36―head, dorsal view; 37―left antenna; 38―labrum; 39―left mandible; 40―memtum; 41―labium; 42, right maxilla; 43―pronotum, ventral view; 44―meso- and meta traces; 45―right elytra; 46―left fore leg." figureDoi="http://doi.org/10.5281/zenodo.4915267" httpUri="https://zenodo.org/record/4915267/files/figure.png" pageId="10" pageNumber="371">37</figureCitation>
|
||
) 7-segmented, inserted near mouthparts; segment I large, thin, concealed in antennal cavity; segments II–VII connected closely, fusiform overall.
|
||
</paragraph>
|
||
<paragraph id="CF1436977146FFB57EC1F8CEA9B7FF20" blockId="10.[151,1436,1646,2031]" lastBlockId="11.[151,1437,151,1437]" lastPageId="11" lastPageNumber="372" pageId="10" pageNumber="371">
|
||
Mouthparts: Labrum (
|
||
<figureCitation id="57902A127146FFB47FBBF8CEA846F8CE" box="[445,526,1826,1851]" captionStart="FIGURES 36–46" captionStartId="9.[151,263,1952,1975]" captionTargetBox="[151,1431,198,1926]" captionTargetId="figure-59@9.[151,1436,193,1932]" captionTargetPageId="9" captionText="FIGURES 36–46. Body parts of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 36―head, dorsal view; 37―left antenna; 38―labrum; 39―left mandible; 40―memtum; 41―labium; 42, right maxilla; 43―pronotum, ventral view; 44―meso- and meta traces; 45―right elytra; 46―left fore leg." figureDoi="http://doi.org/10.5281/zenodo.4915267" httpUri="https://zenodo.org/record/4915267/files/figure.png" pageId="10" pageNumber="371">Fig. 38</figureCitation>
|
||
) rather generalized, with some pseudopores, and a row of 3 campaniform sensilla around antero-lateral corner (Fig, 38: arrow). Mandibles (
|
||
<figureCitation id="57902A127146FFB47D1CF8AAA923F8AA" box="[794,875,1862,1887]" captionStart="FIGURES 36–46" captionStartId="9.[151,263,1952,1975]" captionTargetBox="[151,1431,198,1926]" captionTargetId="figure-59@9.[151,1436,193,1932]" captionTargetPageId="9" captionText="FIGURES 36–46. Body parts of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 36―head, dorsal view; 37―left antenna; 38―labrum; 39―left mandible; 40―memtum; 41―labium; 42, right maxilla; 43―pronotum, ventral view; 44―meso- and meta traces; 45―right elytra; 46―left fore leg." figureDoi="http://doi.org/10.5281/zenodo.4915267" httpUri="https://zenodo.org/record/4915267/files/figure.png" pageId="10" pageNumber="371">Fig. 39</figureCitation>
|
||
) almost symmetric, with 2 setae around middle of outer margin. Mentum (
|
||
<figureCitation id="57902A127146FFB47FA3F886ABBFF876" box="[421,503,1898,1923]" captionStart="FIGURES 36–46" captionStartId="9.[151,263,1952,1975]" captionTargetBox="[151,1431,198,1926]" captionTargetId="figure-59@9.[151,1436,193,1932]" captionTargetPageId="9" captionText="FIGURES 36–46. Body parts of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 36―head, dorsal view; 37―left antenna; 38―labrum; 39―left mandible; 40―memtum; 41―labium; 42, right maxilla; 43―pronotum, ventral view; 44―meso- and meta traces; 45―right elytra; 46―left fore leg." figureDoi="http://doi.org/10.5281/zenodo.4915267" httpUri="https://zenodo.org/record/4915267/files/figure.png" pageId="10" pageNumber="371">Fig. 40</figureCitation>
|
||
) sub-trapezoidal, but lateral margin rounded, truncate on anterior margin, sparsely with pseudopores, some setae around antero-lateral area. Labium (
|
||
<figureCitation id="57902A127146FFB47DADF862AE4DF852" box="[939,1029,1934,1959]" captionStart="FIGURES 36–46" captionStartId="9.[151,263,1952,1975]" captionTargetBox="[151,1431,198,1926]" captionTargetId="figure-59@9.[151,1436,193,1932]" captionTargetPageId="9" captionText="FIGURES 36–46. Body parts of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 36―head, dorsal view; 37―left antenna; 38―labrum; 39―left mandible; 40―memtum; 41―labium; 42, right maxilla; 43―pronotum, ventral view; 44―meso- and meta traces; 45―right elytra; 46―left fore leg." figureDoi="http://doi.org/10.5281/zenodo.4915267" httpUri="https://zenodo.org/record/4915267/files/figure.png" pageId="10" pageNumber="371">Fig. 41</figureCitation>
|
||
): apodeme with lateral lobe short, narrowed toward pointed apex, with apically truncate medial projection; prementum with 2 real pores and 1 setal pore antero-laterally; palpus with segment I long, 3 times as long as II; segment III narrow, elongate; maxilla (
|
||
<figureCitation id="57902A127146FFB57B69F83AAAFDFF45" captionStart="FIGURES 36–46" captionStartId="9.[151,263,1952,1975]" captionTargetBox="[151,1431,198,1926]" captionTargetId="figure-59@9.[151,1436,193,1932]" captionTargetPageId="9" captionText="FIGURES 36–46. Body parts of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 36―head, dorsal view; 37―left antenna; 38―labrum; 39―left mandible; 40―memtum; 41―labium; 42, right maxilla; 43―pronotum, ventral view; 44―meso- and meta traces; 45―right elytra; 46―left fore leg." figureDoi="http://doi.org/10.5281/zenodo.4915267" httpUri="https://zenodo.org/record/4915267/files/figure.png" lastPageId="11" lastPageNumber="372" pageId="10" pageNumber="371">Fig. 42</figureCitation>
|
||
) with cardo elongate; lacinia with 4 spines near apex, densely with pubescence; galea short, with tuft of long pubescence at apex; palpus sparsely with pseudopores; segment IV narrowed.
|
||
</paragraph>
|
||
<paragraph id="CF1436977147FFB57EC1FF0CAE1EFEE8" blockId="11.[151,1437,151,1437]" pageId="11" pageNumber="372">
|
||
Pronotum (
|
||
<figureCitation id="57902A127147FFB57F4EFF33AB9AFF0D" box="[328,466,223,248]" captionStart="FIGURES 33–35" captionStartId="8.[151,263,1970,1993]" captionTargetBox="[202,1381,1000,1927]" captionTargetId="figure-253@8.[151,1436,967,1948]" captionTargetPageId="8" captionText="FIGURES 33–35. Habitus of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 33―dorsal view; 34―ventral view; 35―lateral view." figureDoi="http://doi.org/10.5281/zenodo.4915263" httpUri="https://zenodo.org/record/4915263/files/figure.png" pageId="11" pageNumber="372">Figs. 33–35</figureCitation>
|
||
,
|
||
<figureCitation id="57902A127147FFB57FE7FF33ABB6FF0D" box="[481,510,223,248]" captionStart="FIGURES 36–46" captionStartId="9.[151,263,1952,1975]" captionTargetBox="[151,1431,198,1926]" captionTargetId="figure-59@9.[151,1436,193,1932]" captionTargetPageId="9" captionText="FIGURES 36–46. Body parts of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 36―head, dorsal view; 37―left antenna; 38―labrum; 39―left mandible; 40―memtum; 41―labium; 42, right maxilla; 43―pronotum, ventral view; 44―meso- and meta traces; 45―right elytra; 46―left fore leg." figureDoi="http://doi.org/10.5281/zenodo.4915267" httpUri="https://zenodo.org/record/4915267/files/figure.png" pageId="11" pageNumber="372">43</figureCitation>
|
||
) transverse, with a row of thick setae on lateral margin; hypomera laterally not visible, with several macrosetae; prosternite (
|
||
<figureCitation id="57902A127147FFB57C94FEE8A8ACFEE8" box="[658,740,260,285]" captionStart="FIGURES 13–23" captionStartId="6.[151,263,1837,1860]" captionTargetBox="[151,1422,204,1800]" captionTargetId="figure-66@6.[151,1436,193,1817]" captionTargetPageId="6" captionText="FIGURES 13–23. Body parts of Aenictosymbia cornuta Maruyama, gen. & sp. n. 13―head, dorsal view; 14―labrum (right, epipharynx); 15―left mandible; 16―memtum; 17―labium; 18, right maxilla; 19―pronotum, dorsal view; 20―ditto, ventral view; 21―meso- and meta traces; 22―right elytra; 23―left fore leg." figureDoi="http://doi.org/10.5281/zenodo.4915255" httpUri="https://zenodo.org/record/4915255/files/figure.png" pageId="11" pageNumber="372">Fig. 20</figureCitation>
|
||
) trapezoidal, carinate at midline.
|
||
</paragraph>
|
||
<paragraph id="CF1436977147FFB57EC1FECBAF16FE7D" blockId="11.[151,1437,151,1437]" pageId="11" pageNumber="372">
|
||
Mesoventrite (
|
||
<figureCitation id="57902A127147FFB57F76FECBAB80FEB5" box="[368,456,295,320]" captionStart="FIGURES 36–46" captionStartId="9.[151,263,1952,1975]" captionTargetBox="[151,1431,198,1926]" captionTargetId="figure-59@9.[151,1436,193,1932]" captionTargetPageId="9" captionText="FIGURES 36–46. Body parts of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 36―head, dorsal view; 37―left antenna; 38―labrum; 39―left mandible; 40―memtum; 41―labium; 42, right maxilla; 43―pronotum, ventral view; 44―meso- and meta traces; 45―right elytra; 46―left fore leg." figureDoi="http://doi.org/10.5281/zenodo.4915267" httpUri="https://zenodo.org/record/4915267/files/figure.png" pageId="11" pageNumber="372">Fig. 44</figureCitation>
|
||
) with process not differentiated from that of metaventrite; mesocoxal cavity well margined, clearly separated. Elytra (
|
||
<figureCitation id="57902A127147FFB57C28FEA0A8C8FE90" box="[558,640,332,357]" captionStart="FIGURES 36–46" captionStartId="9.[151,263,1952,1975]" captionTargetBox="[151,1431,198,1926]" captionTargetId="figure-59@9.[151,1436,193,1932]" captionTargetPageId="9" captionText="FIGURES 36–46. Body parts of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 36―head, dorsal view; 37―left antenna; 38―labrum; 39―left mandible; 40―memtum; 41―labium; 42, right maxilla; 43―pronotum, ventral view; 44―meso- and meta traces; 45―right elytra; 46―left fore leg." figureDoi="http://doi.org/10.5281/zenodo.4915267" httpUri="https://zenodo.org/record/4915267/files/figure.png" pageId="11" pageNumber="372">Fig. 45</figureCitation>
|
||
) transverse, slightly convex above, with a row of thick setae on lateral margin; hypomera laterally not visible, with several macrosetae along lateral margins; hind wings developed.
|
||
</paragraph>
|
||
<paragraph id="CF1436977147FFB57EC1FE78A817FDC8" blockId="11.[151,1437,151,1437]" pageId="11" pageNumber="372">
|
||
Legs (
|
||
<figureCitation id="57902A127147FFB57F0AFE78ABDBFE58" box="[268,403,404,429]" captionStart="FIGURES 33–35" captionStartId="8.[151,263,1970,1993]" captionTargetBox="[202,1381,1000,1927]" captionTargetId="figure-253@8.[151,1436,967,1948]" captionTargetPageId="8" captionText="FIGURES 33–35. Habitus of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 33―dorsal view; 34―ventral view; 35―lateral view." figureDoi="http://doi.org/10.5281/zenodo.4915263" httpUri="https://zenodo.org/record/4915263/files/figure.png" pageId="11" pageNumber="372">Figs. 34–35</figureCitation>
|
||
,
|
||
<figureCitation id="57902A127147FFB57F98FE78ABAEFE58" box="[414,486,404,429]" captionStart-0="FIGURES 36–46" captionStart-1="FIGURES 47–54" captionStartId-0="9.[151,263,1952,1975]" captionStartId-1="10.[151,263,1518,1541]" captionTargetBox-0="[151,1431,198,1926]" captionTargetBox-1="[172,1425,195,1494]" captionTargetId-0="figure-59@9.[151,1436,193,1932]" captionTargetId-1="figure-224@10.[159,1426,193,1503]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURES 36–46. Body parts of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 36―head, dorsal view; 37―left antenna; 38―labrum; 39―left mandible; 40―memtum; 41―labium; 42, right maxilla; 43―pronotum, ventral view; 44―meso- and meta traces; 45―right elytra; 46―left fore leg." captionText-1="FIGURES 47–54. Body parts of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 47―left mid leg; 48―left hind leg; 49―male tergite VIII; 50―male sternite VIII; 51―male abdominal tergites IX and X; 52―median lobe of aedeagus, lateral view; 53―paramere; 54―spermatheca." figureDoi-0="http://doi.org/10.5281/zenodo.4915267" figureDoi-1="http://doi.org/10.5281/zenodo.4915269" httpUri-0="https://zenodo.org/record/4915267/files/figure.png" httpUri-1="https://zenodo.org/record/4915269/files/figure.png" pageId="11" pageNumber="372">46–48</figureCitation>
|
||
) short, compressed, with inner margins of femora sulcate to hold tibia; tarsal formula 4–4–5; fore leg (
|
||
<figureCitation id="57902A127147FFB57F57FE5BABEAFE25" box="[337,418,439,464]" captionStart="FIGURES 36–46" captionStartId="9.[151,263,1952,1975]" captionTargetBox="[151,1431,198,1926]" captionTargetId="figure-59@9.[151,1436,193,1932]" captionTargetPageId="9" captionText="FIGURES 36–46. Body parts of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 36―head, dorsal view; 37―left antenna; 38―labrum; 39―left mandible; 40―memtum; 41―labium; 42, right maxilla; 43―pronotum, ventral view; 44―meso- and meta traces; 45―right elytra; 46―left fore leg." figureDoi="http://doi.org/10.5281/zenodo.4915267" httpUri="https://zenodo.org/record/4915267/files/figure.png" pageId="11" pageNumber="372">Fig. 46</figureCitation>
|
||
) with tibia slender widest at middle; tarsal segments I–III subequal in length; mid leg (
|
||
<figureCitation id="57902A127147FFB57B69FE5BAAFCFE00" captionStart="FIGURES 47–54" captionStartId="10.[151,263,1518,1541]" captionTargetBox="[172,1425,195,1494]" captionTargetId="figure-224@10.[159,1426,193,1503]" captionTargetPageId="10" captionText="FIGURES 47–54. Body parts of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 47―left mid leg; 48―left hind leg; 49―male tergite VIII; 50―male sternite VIII; 51―male abdominal tergites IX and X; 52―median lobe of aedeagus, lateral view; 53―paramere; 54―spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4915269" httpUri="https://zenodo.org/record/4915269/files/figure.png" pageId="11" pageNumber="372">Fig. 47</figureCitation>
|
||
) with coxa oval; tibia thick, widest around basal 1/3; tarsal segments I–III subequal in length; hind leg (
|
||
<figureCitation id="57902A127147FFB57B45FE30AFDCFE00" box="[1347,1428,476,501]" captionStart="FIGURES 47–54" captionStartId="10.[151,263,1518,1541]" captionTargetBox="[172,1425,195,1494]" captionTargetId="figure-224@10.[159,1426,193,1503]" captionTargetPageId="10" captionText="FIGURES 47–54. Body parts of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 47―left mid leg; 48―left hind leg; 49―male tergite VIII; 50―male sternite VIII; 51―male abdominal tergites IX and X; 52―median lobe of aedeagus, lateral view; 53―paramere; 54―spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4915269" httpUri="https://zenodo.org/record/4915269/files/figure.png" pageId="11" pageNumber="372">Fig. 48</figureCitation>
|
||
) with coxa transverse, with thick setae around lateral area; tibia thick, widest around basal 1/3; tarsal segments I–III subequal in length; segment IV smallest.
|
||
</paragraph>
|
||
<paragraph id="CF1436977147FFB57EC1FDABAEF1FD38" blockId="11.[151,1437,151,1437]" pageId="11" pageNumber="372">
|
||
Abdomen (
|
||
<figureCitation id="57902A127147FFB57F40FDABAB87FD95" box="[326,463,583,608]" captionStart="FIGURES 33–35" captionStartId="8.[151,263,1970,1993]" captionTargetBox="[202,1381,1000,1927]" captionTargetId="figure-253@8.[151,1436,967,1948]" captionTargetPageId="8" captionText="FIGURES 33–35. Habitus of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 33―dorsal view; 34―ventral view; 35―lateral view." figureDoi="http://doi.org/10.5281/zenodo.4915263" httpUri="https://zenodo.org/record/4915263/files/figure.png" pageId="11" pageNumber="372">Figs. 33–35</figureCitation>
|
||
) widest at segment IV, gently narrowed toward apex; tergite VIII (
|
||
<figureCitation id="57902A127147FFB57AC8FDABAF69FD95" box="[1230,1313,583,608]" captionStart="FIGURES 47–54" captionStartId="10.[151,263,1518,1541]" captionTargetBox="[172,1425,195,1494]" captionTargetId="figure-224@10.[159,1426,193,1503]" captionTargetPageId="10" captionText="FIGURES 47–54. Body parts of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 47―left mid leg; 48―left hind leg; 49―male tergite VIII; 50―male sternite VIII; 51―male abdominal tergites IX and X; 52―median lobe of aedeagus, lateral view; 53―paramere; 54―spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4915269" httpUri="https://zenodo.org/record/4915269/files/figure.png" pageId="11" pageNumber="372">Fig. 49</figureCitation>
|
||
) narrowed apicad, sub-parallel near apex, rounded on apical margin; sternite (
|
||
<figureCitation id="57902A127147FFB57D98FD80A9BAFD70" box="[926,1010,620,645]" captionStart="FIGURES 47–54" captionStartId="10.[151,263,1518,1541]" captionTargetBox="[172,1425,195,1494]" captionTargetId="figure-224@10.[159,1426,193,1503]" captionTargetPageId="10" captionText="FIGURES 47–54. Body parts of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 47―left mid leg; 48―left hind leg; 49―male tergite VIII; 50―male sternite VIII; 51―male abdominal tergites IX and X; 52―median lobe of aedeagus, lateral view; 53―paramere; 54―spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4915269" httpUri="https://zenodo.org/record/4915269/files/figure.png" pageId="11" pageNumber="372">Fig. 50</figureCitation>
|
||
) gently narrowed apicad, rounded at apex; tergites IX and X (
|
||
<figureCitation id="57902A127147FFB57FB3FD63A840FD5D" box="[437,520,655,680]" captionStart="FIGURES 47–54" captionStartId="10.[151,263,1518,1541]" captionTargetBox="[172,1425,195,1494]" captionTargetId="figure-224@10.[159,1426,193,1503]" captionTargetPageId="10" captionText="FIGURES 47–54. Body parts of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 47―left mid leg; 48―left hind leg; 49―male tergite VIII; 50―male sternite VIII; 51―male abdominal tergites IX and X; 52―median lobe of aedeagus, lateral view; 53―paramere; 54―spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4915269" httpUri="https://zenodo.org/record/4915269/files/figure.png" pageId="11" pageNumber="372">Fig. 51</figureCitation>
|
||
) cylindrical; tergite IX narrowed around base; sclerite at base of tergite IX well developed, transverse (
|
||
<figureCitation id="57902A127147FFB57F9FFD58ABA4FD38" box="[409,492,692,717]" captionStart="FIGURES 47–54" captionStartId="10.[151,263,1518,1541]" captionTargetBox="[172,1425,195,1494]" captionTargetId="figure-224@10.[159,1426,193,1503]" captionTargetPageId="10" captionText="FIGURES 47–54. Body parts of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 47―left mid leg; 48―left hind leg; 49―male tergite VIII; 50―male sternite VIII; 51―male abdominal tergites IX and X; 52―median lobe of aedeagus, lateral view; 53―paramere; 54―spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4915269" httpUri="https://zenodo.org/record/4915269/files/figure.png" pageId="11" pageNumber="372">Fig. 51</figureCitation>
|
||
: arrow); tergite X emarginate on anterior margin in dorsal view.
|
||
</paragraph>
|
||
<paragraph id="CF1436977147FFB57EC1FD3BABACFCE0" blockId="11.[151,1437,151,1437]" pageId="11" pageNumber="372">
|
||
Male: median lobe of aedeagus (
|
||
<figureCitation id="57902A127147FFB57C32FD3BA8CDFD05" box="[564,645,727,752]" captionStart="FIGURES 47–54" captionStartId="10.[151,263,1518,1541]" captionTargetBox="[172,1425,195,1494]" captionTargetId="figure-224@10.[159,1426,193,1503]" captionTargetPageId="10" captionText="FIGURES 47–54. Body parts of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 47―left mid leg; 48―left hind leg; 49―male tergite VIII; 50―male sternite VIII; 51―male abdominal tergites IX and X; 52―median lobe of aedeagus, lateral view; 53―paramere; 54―spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4915269" httpUri="https://zenodo.org/record/4915269/files/figure.png" pageId="11" pageNumber="372">Fig. 52</figureCitation>
|
||
) bulbous at base; apical lobe elongate, pointed at apex; paramere (
|
||
<figureCitation id="57902A127147FFB57B69FD3BAAFCFCE0" captionStart="FIGURES 47–54" captionStartId="10.[151,263,1518,1541]" captionTargetBox="[172,1425,195,1494]" captionTargetId="figure-224@10.[159,1426,193,1503]" captionTargetPageId="10" captionText="FIGURES 47–54. Body parts of Aenictoxenides mirabilis Maruyama, gen. & sp. n. 47―left mid leg; 48―left hind leg; 49―male tergite VIII; 50―male sternite VIII; 51―male abdominal tergites IX and X; 52―median lobe of aedeagus, lateral view; 53―paramere; 54―spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4915269" httpUri="https://zenodo.org/record/4915269/files/figure.png" pageId="11" pageNumber="372">Fig. 53</figureCitation>
|
||
) with apical lobe elongate.
|
||
</paragraph>
|
||
<paragraph id="CF1436977147FFB57EC1FCF3AE6EFCCD" blockId="11.[151,1437,151,1437]" box="[199,1062,799,824]" pageId="11" pageNumber="372">
|
||
Female: spermatheca (
|
||
<figureCitation id="57902A127147FFB57FC5FCF3A85DFCCD" box="[451,533,799,824]" captionStart="FIGURES 24–32" captionStartId="7.[151,263,1952,1975]" captionTargetBox="[161,1430,212,1917]" captionTargetId="figure-65@7.[151,1436,193,1932]" captionTargetPageId="7" captionText="FIGURES 24–32. Body parts of Aenictosymbia cornuta Maruyama, gen. & sp. n. 24―left mid leg; 25―left hind leg; 26―male tergite VIII; 27―male sternite VIII; 28―male abdominal tergites IX and X; 29―median lobe of aedeagus, lateral view; 30―ditto, apical part, ventra view; 31―paramere; 32―spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4915259" httpUri="https://zenodo.org/record/4915259/files/figure.png" pageId="11" pageNumber="372">Fig. 32</figureCitation>
|
||
) coiled around base, with apical part spherical.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="87B1651C7147FFB57EC1FCA8AE86FBC0" pageId="11" pageNumber="372" type="diagnosis">
|
||
<paragraph id="CF1436977147FFB57EC1FCA8AE86FBC0" blockId="11.[151,1437,151,1437]" pageId="11" pageNumber="372">
|
||
<emphasis id="FDDFEA857147FFB57EC1FCA8AB99FCA8" bold="true" box="[199,465,836,861]" inLineHeading="true" pageId="11" pageNumber="372" reason="1">Differential diagnosis.</emphasis>
|
||
This genus is closely related to
|
||
<taxonomicName id="08AB4D147147FFB57D57FCA8AEDAFCA8" authority="Seevers, 1953" authorityName="Seevers" authorityYear="1953" box="[849,1170,836,861]" class="Insecta" family="Staphylinidae" genus="Aenictoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FDDFEA857147FFB57D57FCA8A9AEFCAE" box="[849,998,836,859]" italics="true" pageId="11" pageNumber="372">Aenictoxenus</emphasis>
|
||
<bibRefCitation id="AB3A4B667147FFB57DF7FCA8AEDAFCA8" author="Seevers, C. H." box="[1009,1170,836,861]" pageId="11" pageNumber="372" pagination="125 - 131" refId="ref5706" refString="Seevers, C. H. (1953) Two genera of myrmecophilous Staphylinidae from the Philippines. Philippine Journal of Science, 81, 125 - 131 + 1 pl." type="journal article" year="1953">
|
||
Seevers,
|
||
<quantity id="08539B727147FFB57A5CFCA8AEDAFCA8" box="[1114,1170,836,861]" metricMagnitude="1" metricUnit="m" metricValue="4.96062" pageId="11" pageNumber="372" unit="in" value="1953.0">1953</quantity>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
in numerous character states, especially those of the mouthparts and thorax, but distinguished from it by the temples which widen behind the eyes, forming acute posterior head angles (although
|
||
<taxonomicName id="08AB4D147147FFB57D1CFC60A9E7FC56" authorityName="Seevers" authorityYear="1953" box="[794,943,908,931]" class="Insecta" family="Staphylinidae" genus="Aenictoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FDDFEA857147FFB57D1CFC60A9E7FC56" box="[794,943,908,931]" italics="true" pageId="11" pageNumber="372">Aenictoxenus</emphasis>
|
||
</taxonomicName>
|
||
has an “arcade” formed by a fronto-lateral expansion of the head, the temples are not expanded, and instead narrow smoothly and roundly to the head base), the antennae which are 7-segmented (in
|
||
<taxonomicName id="08AB4D147147FFB57CA8FC38A90BFC1E" authorityName="Seevers" authorityYear="1953" box="[686,835,980,1003]" class="Insecta" family="Staphylinidae" genus="Aenictoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FDDFEA857147FFB57CA8FC38A90BFC1E" box="[686,835,980,1003]" italics="true" pageId="11" pageNumber="372">Aenictoxenus</emphasis>
|
||
</taxonomicName>
|
||
, 8-segmented), the sub-trapezoidal mentum (in
|
||
<taxonomicName id="08AB4D147147FFB57E91FC14AB64FBFA" authorityName="Seevers" authorityYear="1953" box="[151,300,1016,1039]" class="Insecta" family="Staphylinidae" genus="Aenictoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FDDFEA857147FFB57E91FC14AB64FBFA" box="[151,300,1016,1039]" italics="true" pageId="11" pageNumber="372">Aenictoxenus</emphasis>
|
||
</taxonomicName>
|
||
, the anterior margin is rounded), and tergite X more shallowly emarginate on the anterior margin in dorsal view (in
|
||
<taxonomicName id="08AB4D147147FFB57F40FBF0AB93FBC6" authorityName="Seevers" authorityYear="1953" box="[326,475,1052,1075]" class="Insecta" family="Staphylinidae" genus="Aenictoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FDDFEA857147FFB57F40FBF0AB93FBC6" box="[326,475,1052,1075]" italics="true" pageId="11" pageNumber="372">Aenictoxenus</emphasis>
|
||
</taxonomicName>
|
||
, it is deeply emarginate, deeper than 1/2 of the length of tergite X).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="87B1651C7147FFB57EC1FBD3A987FB88" pageId="11" pageNumber="372" type="etymology">
|
||
<paragraph id="CF1436977147FFB57EC1FBD3A987FB88" blockId="11.[151,1437,151,1437]" pageId="11" pageNumber="372">
|
||
<emphasis id="FDDFEA857147FFB57EC1FBD3AB06FBAD" bold="true" box="[199,334,1087,1112]" inLineHeading="true" pageId="11" pageNumber="372" reason="1">Etymology.</emphasis>
|
||
A combination of the related genus name
|
||
<taxonomicName id="08AB4D147147FFB57D48FBACA9ABFBA2" authorityName="Seevers" authorityYear="1953" box="[846,995,1088,1111]" class="Insecta" family="Staphylinidae" genus="Aenictoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FDDFEA857147FFB57D48FBACA9ABFBA2" box="[846,995,1088,1111]" italics="true" pageId="11" pageNumber="372">Aenictoxenus</emphasis>
|
||
</taxonomicName>
|
||
, and the Latin suffix
|
||
<emphasis id="FDDFEA857147FFB57AEDFBACAF6EFBAD" box="[1259,1318,1088,1112]" italics="true" pageId="11" pageNumber="372">oides</emphasis>
|
||
(meaning "similar"), indicating a similar genus of
|
||
<taxonomicName id="08AB4D147147FFB57C5FFB88A8A6FB8E" authorityName="Seevers" authorityYear="1953" box="[601,750,1124,1147]" class="Insecta" family="Staphylinidae" genus="Aenictoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FDDFEA857147FFB57C5FFB88A8A6FB8E" box="[601,750,1124,1147]" italics="true" pageId="11" pageNumber="372">Aenictoxenus</emphasis>
|
||
</taxonomicName>
|
||
. Gender masculine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="87B1651C7147FFB57EC1FB6BA8F0FA68" pageId="11" pageNumber="372" type="distribution">
|
||
<paragraph id="CF1436977147FFB57EC1FB6BA864FB55" blockId="11.[151,1437,151,1437]" box="[199,556,1159,1184]" pageId="11" pageNumber="372">
|
||
<emphasis id="FDDFEA857147FFB57EC1FB6BAB28FB55" bold="true" box="[199,352,1159,1184]" inLineHeading="true" pageId="11" pageNumber="372" reason="1">Distribution.</emphasis>
|
||
Central
|
||
<collectingCountry id="B7BC76077147FFB57FC6FB6BA86FFB55" box="[448,551,1159,1184]" name="Thailand" pageId="11" pageNumber="372">Thailand</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="CF1436977147FFB57EC1FB40A8F0FA68" blockId="11.[151,1437,151,1437]" pageId="11" pageNumber="372">
|
||
<emphasis id="FDDFEA857147FFB57EC1FB40ABDDFB30" bold="true" box="[199,405,1196,1221]" inLineHeading="true" pageId="11" pageNumber="372" reason="1">Systematic notes.</emphasis>
|
||
Tribal affiliations of
|
||
<taxonomicName id="08AB4D147147FFB57C8DFB40A968FB36" authorityName="Seevers" authorityYear="1953" box="[651,800,1196,1219]" class="Insecta" family="Staphylinidae" genus="Aenictoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FDDFEA857147FFB57C8DFB40A968FB36" box="[651,800,1196,1219]" italics="true" pageId="11" pageNumber="372">Aenictoxenus</emphasis>
|
||
</taxonomicName>
|
||
are controversial.
|
||
<bibRefCitation id="AB3A4B667147FFB57DF3FB40AF6FFB30" author="Jacobson, H. R. & Kistner, D. H." box="[1013,1319,1196,1221]" pageId="11" pageNumber="372" pagination="163 - 200" refId="ref5319" refString="Jacobson, H. R. & Kistner, D. H. (1975) The natural history of Pygostenini. Section 1. New species and new records of Pygostenini. Sociobiology, 1, 163 - 200." type="journal article" year="1975">Jacobson & Kistner (1975)</bibRefCitation>
|
||
stated the genus does not belong to
|
||
<taxonomicName id="08AB4D147147FFB57FB4FB23A872FB1D" authorityName="Fauvel" authorityYear="1899" box="[434,570,1231,1256]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="372" phylum="Arthropoda" rank="tribe" tribe="Pygostenini">Pygostenini</taxonomicName>
|
||
. However, the related genus
|
||
<taxonomicName id="08AB4D147147FFB57D7CFB3CAE6BFB1D" authorityName="Maruyama" authorityYear="2014" box="[890,1059,1232,1256]" class="Insecta" family="Staphylinidae" genus="Aenictoxenides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FDDFEA857147FFB57D7CFB3CAE6BFB1D" box="[890,1059,1232,1256]" italics="true" pageId="11" pageNumber="372">Aenictoxenides</emphasis>
|
||
</taxonomicName>
|
||
shares a sclerite at the base of the tergite IX which is one of autapomorphies of
|
||
<taxonomicName id="08AB4D147147FFB57CADFB18A967FAF8" authorityName="Fauvel" authorityYear="1899" box="[683,815,1268,1293]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="372" phylum="Arthropoda" rank="tribe" tribe="Pygostenini">Pygostenini</taxonomicName>
|
||
(Maruyama
|
||
<emphasis id="FDDFEA857147FFB57DC3FB19AE49FAF9" box="[965,1025,1268,1292]" italics="true" pageId="11" pageNumber="372">et al.</emphasis>
|
||
, in prep.), and the well-sclerotized, cylindrical abdominal segments IX + X are also characteristic in
|
||
<taxonomicName id="08AB4D147147FFB57D97FAFBAE51FAC5" authorityName="Fauvel" authorityYear="1899" box="[913,1049,1303,1328]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="372" phylum="Arthropoda" rank="tribe" tribe="Pygostenini">Pygostenini</taxonomicName>
|
||
. Although
|
||
<taxonomicName id="08AB4D147147FFB57A9DFAF4AF78FADA" authorityName="Seevers" authorityYear="1953" box="[1179,1328,1304,1327]" class="Insecta" family="Staphylinidae" genus="Aenictoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FDDFEA857147FFB57A9DFAF4AF78FADA" box="[1179,1328,1304,1327]" italics="true" pageId="11" pageNumber="372">Aenictoxenus</emphasis>
|
||
</taxonomicName>
|
||
does not share the sclerite at base of tergite X, it could be a secondary loss of the state due to extreme miniaturization of the body. This sclerite is ill-defined also in some other minute-sized pygostenines. We propose that both
|
||
<taxonomicName id="08AB4D147147FFB57B00FA8CAFD3FA82" authorityName="Seevers" authorityYear="1953" box="[1286,1435,1376,1399]" class="Insecta" family="Staphylinidae" genus="Aenictoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FDDFEA857147FFB57B00FA8CAFD3FA82" box="[1286,1435,1376,1399]" italics="true" pageId="11" pageNumber="372">Aenictoxenus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="08AB4D147147FFB57EC0FA68AB38FA69" authorityName="Maruyama" authorityYear="2014" box="[198,368,1412,1436]" class="Insecta" family="Staphylinidae" genus="Aenictoxenides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FDDFEA857147FFB57EC0FA68AB38FA69" box="[198,368,1412,1436]" italics="true" pageId="11" pageNumber="372">Aenictoxenides</emphasis>
|
||
</taxonomicName>
|
||
are members of
|
||
<taxonomicName id="08AB4D147147FFB57C2AFA68A8FCFA68" authorityName="Fauvel" authorityYear="1899" box="[556,692,1412,1437]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="372" phylum="Arthropoda" rank="tribe" tribe="Pygostenini">Pygostenini</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |