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<document id="CEDEF56996BD5BEBD0C822FC11F18370" ID-CLB-Dataset="31787" ID-DOI="10.11646/zootaxa.4337.4.3" ID-GBIF-Dataset="c9973f7f-1bbb-4f66-95b0-77913ab30c09" ID-ISSN="1175-5326" ID-Zenodo-Dep="1034252" ID-ZooBank="2D4807A5-EAF1-4B93-8833-B3C71BC53A9D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1508523566360" checkinUser="plazi" docAuthor="Gupta, Devanshu &amp; Chandra, Kailash" docDate="2017" docId="03D987F3ED4FFFB2ADB7F967FBF4FDAB" docLanguage="en" docName="zootaxa.4337.4.3.pdf" docOrigin="Zootaxa 4337 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tropiorhynchus orientis" docType="treatment" docVersion="8" lastPageNumber="513" masterDocId="FFE0FF8BED4CFFB6AD20FFC2FF8BFFB3" masterDocTitle="Tropiorhynchus annandalei (Coleoptera: Scarabaeidae: Rutelinae), a new species from Maharashtra, India" masterLastPageNumber="522" masterPageNumber="509" pageNumber="512" updateTime="1698469998896" updateUser="ExternalLinkService">
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<mods:title id="1767B9C0BE510F1AAE6A832D90D3ECBE">Tropiorhynchus annandalei (Coleoptera: Scarabaeidae: Rutelinae), a new species from Maharashtra, India</mods:title>
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<mods:namePart id="1A9D705D463CA04DFD78179510503586">Gupta, Devanshu</mods:namePart>
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<mods:namePart id="44AC5553B4CFBF423A07267B3C528B59">Chandra, Kailash</mods:namePart>
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<mods:title id="27AF25186A2AF3E17115DF066BE483D1">Zootaxa</mods:title>
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<mods:date id="BDE67FF324157FBF27473CA942EABB99">2017</mods:date>
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<mods:number id="50B881A3C9B0A58E0E2953D071FDE9FD">2017-10-20</mods:number>
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<treatment id="03D987F3ED4FFFB2ADB7F967FBF4FDAB" ID-DOI="http://doi.org/10.5281/zenodo.6042688" ID-GBIF-Taxon="135452493" ID-Zenodo-Dep="6042688" LSID="urn:lsid:plazi:treatment:03D987F3ED4FFFB2ADB7F967FBF4FDAB" httpUri="http://treatment.plazi.org/id/03D987F3ED4FFFB2ADB7F967FBF4FDAB" lastPageId="4" lastPageNumber="513" pageId="3" pageNumber="512">
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<paragraph id="8BCF36E5ED4FFFB5ADB7F967FD05F90C" blockId="3.[151,722,1701,1761]" box="[151,654,1701,1727]" pageId="3" pageNumber="512">
<heading id="D0878189ED4FFFB5ADB7F967FD05F90C" bold="true" box="[151,654,1701,1727]" fontSize="11" level="1" pageId="3" pageNumber="512" reason="1">
<taxonomicName id="4C704D66ED4FFFB5ADB7F967FD05F90C" authority="(Newman, 1838)" baseAuthorityName="Newman" baseAuthorityYear="1838" box="[151,654,1701,1727]" class="Insecta" family="Rutelidae" genus="Tropiorhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="512" phylum="Arthropoda" rank="species" species="orientis">
<emphasis id="B904EAF7ED4FFFB5ADB7F967FD05F90C" bold="true" box="[151,654,1701,1727]" pageId="3" pageNumber="512">
<emphasis id="B904EAF7ED4FFFB5ADB7F967FE30F90C" bold="true" box="[151,443,1701,1727]" italics="true" pageId="3" pageNumber="512">Tropiorhynchus orientis</emphasis>
(
<bibRefCitation id="EFE14B14ED4FFFB5ACEBF967FD0DF90C" author="Newman" box="[459,646,1701,1727]" pageId="3" pageNumber="512" refString="Newman, E. (1838) Entomological notes. Entomological Magazine, 5, 372 - 402." type="journal article" year="1838">Newman, 1838</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BCF36E5ED4FFFB5ADB7F90AFD59F953" blockId="3.[151,722,1701,1761]" box="[151,722,1736,1761]" pageId="3" pageNumber="512">
(
<figureCitation id="134B2A60ED4FFFB5ADBFF90AFF67F953" box="[159,236,1736,1761]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1909,1932]" captionTargetBox="[163,1436,460,1878]" captionTargetId="figure@2.[151,1436,445,1889]" captionTargetPageId="2" captionText="FIGURES 1 4. Habitus, male dorsal view. 1, Tropiorhynchus orientis; 2, T. podagricus; 3, T. umbrinus; 4, T. annandalei (holotype)." httpUri="https://zenodo.org/record/1034254/files/figure.png" pageId="3" pageNumber="512">Figs. 1</figureCitation>
,
<figureCitation id="134B2A60ED4FFFB5ADDAF90AFEADF952" box="[250,294,1736,1761]" captionStart-0="FIGURE 5" captionStart-1="FIGURES 6 9" captionStartId-0="3.[151,250,1316,1338]" captionStartId-1="4.[151,264,1573,1596]" captionTargetBox-0="[157,1436,155,1294]" captionTargetBox-1="[166,1424,611,1552]" captionTargetId-0="figure@3.[151,1436,147,1294]" captionTargetId-1="figure@4.[151,1436,591,1552]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 5. Map of India and adjacent countries showing distribution of the species belonging to the genus Tropiorhynchus." captionText-1="FIGURES 6 9. Head. 6, Tropiorhynchus orientis; 7, T. podagricus; 8, T. umbrinus; 9, T. annandalei (holotype)." httpUri-0="https://zenodo.org/record/1034256/files/figure.png" httpUri-1="https://zenodo.org/record/1034258/files/figure.png" pageId="3" pageNumber="512">56</figureCitation>
,
<figureCitation id="134B2A60ED4FFFB5AC12F90AFEDBF953" box="[306,336,1736,1760]" captionStart="FIGURES 10 13" captionStartId="5.[151,264,1474,1497]" captionTargetBox="[152,1430,375,1452]" captionTargetId="figure@5.[151,1436,375,1452]" captionTargetPageId="5" captionText="FIGURES 10 13. Pronotum. 10, Tropiorhynchus orientis; 11, T. podagricus; 12, T. umbrinus; 13, T. annandalei (holotype)." httpUri="https://zenodo.org/record/1034260/files/figure.png" pageId="3" pageNumber="512">10</figureCitation>
,
<figureCitation id="134B2A60ED4FFFB5AC7DF90AFEF0F953" box="[349,379,1736,1760]" captionStart="FIGURES 14 17" captionStartId="6.[151,264,1832,1855]" captionTargetBox="[152,1433,227,1811]" captionTargetId="figure@6.[151,1436,227,1811]" captionTargetPageId="6" captionText="FIGURES 14 17. Elytra. 14, Tropiorhynchus orientis; 15, T. podagricus; 16, T. umbrinus; 17, T. annandalei (holotype)." httpUri="https://zenodo.org/record/1034262/files/figure.png" pageId="3" pageNumber="512">14</figureCitation>
,
<figureCitation id="134B2A60ED4FFFB5ACA6F90AFE72F953" box="[390,505,1736,1760]" captionStart-0="FIGURES 18 21" captionStart-1="FIGURES 22 24" captionStartId-0="7.[151,264,1156,1179]" captionStartId-1="10.[151,264,1755,1778]" captionTargetBox-0="[151,1432,193,1136]" captionTargetId-0="figure@7.[151,1436,193,1136]" captionTargetPageId-0="7" captionText-0="FIGURES 18 21. Pygidium. 18, Tropiorhynchus orientis; 19, T. podagricus; 20, T. umbrinus; 21, T. annandalei (holotype)." captionText-1="FIGURES 22 24. Tropiorhynchus orientis. 22, profemur; 23, mesofemur; 24, metafemur." httpUri-0="https://zenodo.org/record/1034264/files/figure.png" pageId="3" pageNumber="512">18, 2124</figureCitation>
, 34, 37, 41, 45, 49)
</paragraph>
</subSubSection>
<subSubSection id="C36A656EED4FFFB5ADB7F8D2FDD3F831" pageId="3" pageNumber="512" type="reference_group">
<paragraph id="8BCF36E5ED4FFFB5ADB7F8D2FC9AF895" blockId="3.[151,1436,1808,1922]" box="[151,785,1808,1830]" pageId="3" pageNumber="512">
<treatmentCitationGroup id="AB6011CBED4FFFB5ADB7F8D2FC9AF895" box="[151,785,1808,1830]" pageId="3" pageNumber="512">
<taxonomicName id="4C704D66ED4FFFB5ADB7F8D2FDA8F895" authority="Newman, 1838: 384" authorityName="Newman" authorityPageNumber="384" authorityYear="1838" box="[151,547,1808,1830]" class="Insecta" family="Rutelidae" genus="Anisoplia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="512" phylum="Arthropoda" rank="species" species="orientis">
<emphasis id="B904EAF7ED4FFFB5ADB7F8D2FEC0F895" box="[151,331,1808,1830]" italics="true" pageId="3" pageNumber="512">Anisoplia orientis</emphasis>
<treatmentCitation id="0AD110F4ED4FFFB5AC72F8D3FDA8F895" author="Newman" box="[338,547,1808,1830]" page="384" pageId="3" pageNumber="512" year="1838">
<bibRefCitation id="EFE14B14ED4FFFB5AC72F8D3FE7BF895" author="Newman" box="[338,496,1808,1830]" pageId="3" pageNumber="512" refString="Newman, E. (1838) Entomological notes. Entomological Magazine, 5, 372 - 402." type="journal article" year="1838">Newman, 1838</bibRefCitation>
: 384
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0AD110F4ED4FFFB5AF0DF8D2FC86F895" author="Burmeister" box="[557,781,1808,1830]" page="226" pageId="3" pageNumber="512" year="1844">
<bibRefCitation id="EFE14B14ED4FFFB5AF0DF8D2FD50F895" author="Burmeister" box="[557,731,1808,1830]" pageId="3" pageNumber="512" refString="Burmeister, H. C. C. (1844) Coleoptera Lamellicornia, Anthob Phyllophaga systellochela. Handbuch der Entomologie, 4 (1), 1 - 588." type="journal article" year="1844">Burmeister 1844</bibRefCitation>
: 226
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCF36E5ED4FFFB5ADB7F8ECFDD3F831" blockId="3.[151,1436,1808,1922]" pageId="3" pageNumber="512">
<treatmentCitationGroup id="AB6011CBED4FFFB5ADB7F8ECFDD3F831" pageId="3" pageNumber="512">
<taxonomicName id="4C704D66ED4FFFB5ADB7F8ECFDF9F8F7" authority=": Blanchard 1851: 176" authorityName=": Blanchard" authorityPageNumber="176" authorityYear="1851" box="[151,626,1838,1861]" class="Insecta" family="Rutelidae" genus="Tropiorhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="512" phylum="Arthropoda" rank="species" species="orientis">
<emphasis id="B904EAF7ED4FFFB5ADB7F8ECFE01F8F6" box="[151,394,1838,1861]" italics="true" pageId="3" pageNumber="512">Tropiorhynchus orientis</emphasis>
:
<treatmentCitation id="0AD110F4ED4FFFB5ACB8F8ECFDF9F8F7" author="Blanchard" box="[408,626,1838,1860]" page="176" pageId="3" pageNumber="512" year="1851">
<bibRefCitation id="EFE14B14ED4FFFB5ACB8F8ECFDB5F8F7" author="Blanchard" box="[408,574,1838,1860]" pageId="3" pageNumber="512" refString="Blanchard, C. E. (1851) IerFamille - Scarabaeidae. In: Milne-Edwards H., Blanchard C. E. &amp; Lucas H., Catalogue de la Collection Entomologique du Museum d'Histoire Naturelle de Paris. Classe des Insectes. Ordre des Coleopteres. Tome 1. Deuxieme livraison. Gide et Baudry, Paris, pp. 129 - 240." type="journal article" year="1851">Blanchard 1851</bibRefCitation>
: 176
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0AD110F4ED4FFFB5AF5DF8EDFCA8F8F7" author="Arrow" box="[637,803,1838,1860]" page="54" pageId="3" pageNumber="512" year="1917">
<bibRefCitation id="EFE14B14ED4FFFB5AF5DF8EDFD77F8F7" author="Arrow" box="[637,764,1838,1860]" pageId="3" pageNumber="512" refString="Arrow, G. J. (1917) The Fauna of British India, Including Ceylon and Burma. Coleoptera Lamellicornia part II (Rutelinae, Desmonycinae, and Euchirinae). Taylor &amp; Francis. London, 387 pp." type="book" year="1917">Arrow 1917</bibRefCitation>
: 54
</treatmentCitation>
, plate 1, fig. 2 (habitus, male);
<treatmentCitation id="0AD110F4ED4FFFB5A944F8ECFAC6F8F7" author="Machatschke" box="[1124,1357,1838,1860]" page="64" pageId="3" pageNumber="512" year="1954">
<bibRefCitation id="EFE14B14ED4FFFB5A944F8ECFAACF8F7" author="Machatschke" box="[1124,1319,1838,1860]" pageId="3" pageNumber="512" refString="Machatschke, J. W. (1954) Ein neuer Tropiorhynchus, gleichzeitig Revision der Gattung. Beitragezur Entomologie, 4, 57 - 70." type="journal article" year="1954">Machatschke 1954</bibRefCitation>
: 64
</treatmentCitation>
, fig. 1b (lateral side of pronotum), fig. 2b (metathoracic leg); 6ab (frontal view paramere, lateral view aedeagus), fig. 7 (endophallus);
<treatmentCitation id="0AD110F4ED4FFFB5AC7FF8AEFDDFF831" author="Machatschke" box="[351,596,1900,1922]" page="254" pageId="3" pageNumber="512" year="1972">
<bibRefCitation id="EFE14B14ED4FFFB5AC7FF8AEFDAAF831" author="Machatschke" box="[351,545,1900,1922]" pageId="3" pageNumber="512" refString="Machatschke, J. W. (1972) Superfamilie Scarabaeoidae, Familie Melolonthidae, Subfamilie Rutelinae. Coleopterorum catalogus, 66 (1), 1 - 361." type="journal article" year="1972">Machatschke 1972</bibRefCitation>
: 254
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36A656EED4FFFB5ADB7F86CFC25F85F" pageId="3" pageNumber="512" type="materials_examined">
<paragraph id="8BCF36E5ED4FFFB5ADB7F86CFC25F85F" blockId="3.[151,1436,1966,2028]" pageId="3" pageNumber="512">
<materialsCitation id="3B183CB8ED4FFFB5ADB7F86CFC25F85F" ID-GBIF-Occurrence="1675496974" collectingDate="1916-10-11" collectingDateMax="1916-10-26" collectingDateMin="1916-10-11" collectionCode="ZSCI" collectorName="S. Kemp" country="India" county="Material" location="Castle Rock" municipality="North Kanara district" pageId="3" pageNumber="512" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Himachal Pradesh">
<emphasis id="B904EAF7ED4FFFB5ADB7F86CFEF4F874" bold="true" box="[151,383,1966,1991]" pageId="3" pageNumber="512">
<collectingCounty id="62AE4E69ED4FFFB5ADB7F86CFF74F874" box="[151,255,1966,1991]" pageId="3" pageNumber="512">Material</collectingCounty>
examined
</emphasis>
.
<collectingCountry id="F3677675ED4FFFB5ACB4F86CFE5BF874" box="[404,464,1966,1991]" name="India" pageId="3" pageNumber="512">India</collectingCountry>
,
<collectingRegion id="49B4F807ED4FFFB5ACC2F86CFD33F874" box="[482,696,1966,1991]" country="India" name="Himachal Pradesh" pageId="3" pageNumber="512">Himachal Pradesh</collectingRegion>
,
<collectingMunicipality id="6BABAC9FED4FFFB5AFE9F86CFC4DF874" box="[713,966,1966,1991]" pageId="3" pageNumber="512">North Kanara district</collectingMunicipality>
,
<location id="8EAF603EED4FFFB5AEF8F86CFBE1F874" LSID="urn:lsid:plazi:treatment:03D987F3ED4FFFB2ADB7F967FBF4FDAB:8EAF603EED4FFFB5AEF8F86CFBE1F874" box="[984,1130,1966,1991]" country="India" county="Material" municipality="North Kanara district" name="Castle Rock" pageId="3" pageNumber="512" stateProvince="Himachal Pradesh">Castle Rock</location>
,
<specimenCount id="9D76FD6CED4FFFB5A95BF86DFB50F874" box="[1147,1243,1966,1991]" pageId="3" pageNumber="512" type="male">2 males</specimenCount>
,
<specimenCount id="9D76FD6CED4FFFB5A9CCF86DFAE9F874" box="[1260,1378,1966,1991]" pageId="3" pageNumber="512" type="female">2 females</specimenCount>
,
<collectingDate id="EF8AE9CDED4FFFB5A853F86DFE82F85F" pageId="3" pageNumber="512" value="1916-10-11" valueMax="1916-10-26" valueMin="1916-10-11">
<date id="FFCE1025ED4FFFB5A853F86DFE82F85F" pageId="3" pageNumber="512" value="1916-10-11" valueMax="1916-10-26" valueMin="1916-10-11">11 26.x.1916</date>
</collectingDate>
,
<collectorName id="26855333ED4FFFB5AC35F811FEFCF858" box="[277,375,2003,2027]" pageId="3" pageNumber="512">S. Kemp</collectorName>
(
<collectionCode id="ED61AE20ED4FFFB5ACA5F811FE48F858" box="[389,451,2003,2027]" pageId="3" pageNumber="512">ZSCI</collectionCode>
registration number: 5461/H1 to 5464/H1).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36A656EED48FFB2ADE7FF55FB77FF66" pageId="4" pageNumber="513" type="distribution">
<paragraph id="8BCF36E5ED48FFB2ADE7FF55FB77FF66" blockId="4.[151,1436,151,536]" pageId="4" pageNumber="513">
<emphasis id="B904EAF7ED48FFB2ADE7FF55FED2FF03" bold="true" box="[199,345,151,176]" pageId="4" pageNumber="513">Distribution</emphasis>
(
<figureCitation id="134B2A60ED48FFB2AC4AFF55FE3AFF03" box="[362,433,151,176]" captionStart="FIGURE 5" captionStartId="3.[151,250,1316,1338]" captionTargetBox="[157,1436,155,1294]" captionTargetId="figure@3.[151,1436,147,1294]" captionTargetPageId="3" captionText="FIGURE 5. Map of India and adjacent countries showing distribution of the species belonging to the genus Tropiorhynchus." httpUri="https://zenodo.org/record/1034256/files/figure.png" pageId="4" pageNumber="513">Fig. 5</figureCitation>
).
<collectingCountry id="F3677675ED48FFB2ACE8FF55FD8DFF03" box="[456,518,151,176]" name="India" pageId="4" pageNumber="513">India</collectingCountry>
:
<collectingRegion id="49B4F807ED48FFB2AF32FF55FDE8FF03" box="[530,611,151,176]" country="India" name="Sikkim" pageId="4" pageNumber="513">Sikkim</collectingRegion>
(Darjeeling and Kurseong),
<collectingRegion id="49B4F807ED48FFB2AE8CFF55FB94FF03" box="[940,1055,151,176]" country="India" name="Karnataka" pageId="4" pageNumber="513">Karnataka</collectingRegion>
(Belgaum, Kanara),
<collectingRegion id="49B4F807ED48FFB2A830FF55FA17FF03" box="[1296,1436,151,176]" country="India" name="Maharashtra" pageId="4" pageNumber="513">Maharashtra</collectingRegion>
(
<collectingCountry id="F3677675ED48FFB2ADBFFF7EFE8FFF66" box="[159,260,188,213]" name="India" pageId="4" pageNumber="513">Bombay</collectingCountry>
), and
<collectingRegion id="49B4F807ED48FFB2AC66FF7EFD98FF66" box="[326,531,188,213]" country="India" name="Himachal Pradesh" pageId="4" pageNumber="513">Himachal Pradesh</collectingRegion>
(Shimla). Elsewhere: Baluchistan (Chaman) (
<bibRefCitation id="EFE14B14ED48FFB2A936FF7EFB64FF66" author="Machatschke" box="[1046,1263,188,213]" pageId="4" pageNumber="513" refString="Machatschke, J. W. (1954) Ein neuer Tropiorhynchus, gleichzeitig Revision der Gattung. Beitragezur Entomologie, 4, 57 - 70." type="journal article" year="1954">Machatschke 1954</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36A656EED48FFB2ADE7FF1DFBF4FDAB" pageId="4" pageNumber="513" type="discussion">
<paragraph id="8BCF36E5ED48FFB2ADE7FF1DFBF4FDAB" blockId="4.[151,1436,151,536]" pageId="4" pageNumber="513">
<emphasis id="B904EAF7ED48FFB2ADE7FF1DFEB0FF4B" bold="true" box="[199,315,223,248]" pageId="4" pageNumber="513">Remarks.</emphasis>
The original description of this species (
<bibRefCitation id="EFE14B14ED48FFB2AE18FF22FC6CFF4B" author="Newman" box="[824,999,224,248]" pageId="4" pageNumber="513" refString="Newman, E. (1838) Entomological notes. Entomological Magazine, 5, 372 - 402." type="journal article" year="1838">Newman 1838</bibRefCitation>
) is short though it includes several diagnostic morphological characters revealing its identity such as head, pronotum, and scutellum rugopunctate and setose, with green and bronze colouration; and elytra deeply striate, striae rugopunctate.
<bibRefCitation id="EFE14B14ED48FFB2A9D7FEEAFA17FEF3" author="Arrow" box="[1271,1436,296,320]" pageId="4" pageNumber="513" refString="Arrow, G. J. (1917) The Fauna of British India, Including Ceylon and Burma. Coleoptera Lamellicornia part II (Rutelinae, Desmonycinae, and Euchirinae). Taylor &amp; Francis. London, 387 pp." type="book" year="1917">Arrow (1917)</bibRefCitation>
differentiated this species from
<taxonomicName id="4C704D66ED48FFB2ACD9FE8FFD19FED6" baseAuthorityName="Burmeister" baseAuthorityYear="1844" box="[505,658,332,357]" class="Insecta" family="Rutelidae" genus="Tropiorhynchus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="513" phylum="Arthropoda" rank="species" species="podagricus">
<emphasis id="B904EAF7ED48FFB2ACD9FE8FFD8DFED7" box="[505,518,333,356]" italics="true" pageId="4" pageNumber="513">T</emphasis>
.
<emphasis id="B904EAF7ED48FFB2AF34FE8EFD19FED6" box="[532,658,332,357]" italics="true" pageId="4" pageNumber="513">podagricus</emphasis>
</taxonomicName>
by the presence of produced mesosternum, densely setose pronotum, and apically split outer protarsal claw. Thorough examination of all the congeners revealed that produced mesosternum and apically split outer claw of protarsus are species specific characters, which are not found in any other species of the genus. Moreover, the aedeagus in this particular species is unique with the ventral plate having a raised lamina medially (see
<bibRefCitation id="EFE14B14ED48FFB2ACC7FE1EFD35FE46" author="Machatschke" box="[487,702,476,501]" pageId="4" pageNumber="513" refString="Machatschke, J. W. (1954) Ein neuer Tropiorhynchus, gleichzeitig Revision der Gattung. Beitragezur Entomologie, 4, 57 - 70." type="journal article" year="1954">Machatschke 1954</bibRefCitation>
: figs. 6ab). The aedeagal structure in the specimen we observed (
<figureCitation id="134B2A60ED48FFB2ADBFFE3DFF77FDAB" box="[159,252,511,536]" captionStart="FIGURES 45 48" captionStartId="12.[151,264,1546,1569]" captionTargetBox="[155,1434,193,1508]" captionTargetId="figure@12.[151,1436,193,1525]" captionTargetPageId="12" captionText="FIGURES 45 48. Parameres (in dorsal view). 45, T. orientis; 46, T. podagricus; 47, T. umbrinus; 48, T. annandalei (holotype). FIGURES 49 52. Parameres (in lateral view). 49, T. orientis; 50, T. podagricus; 51, T. umbrinus; 52, T. annandalei (holotype)." httpUri="https://zenodo.org/record/1034268/files/figure.png" pageId="4" pageNumber="513">Figs. 45</figureCitation>
, 49) is identical to the illustrations provided by
<bibRefCitation id="EFE14B14ED48FFB2AE35FE3DFC7EFDAB" author="Machatschke" box="[789,1013,511,536]" pageId="4" pageNumber="513" refString="Machatschke, J. W. (1954) Ein neuer Tropiorhynchus, gleichzeitig Revision der Gattung. Beitragezur Entomologie, 4, 57 - 70." type="journal article" year="1954">Machatschke (1954</bibRefCitation>
: figs. 6ab).
</paragraph>
</subSubSection>
</treatment>
</document>