259 lines
38 KiB
XML
259 lines
38 KiB
XML
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<mods:title id="C4A21B6665A3FFDDD635C0EC851B9EE3">Two new genera of pinnotherid crabs (Crustacea: Brachyura: Pinnotheroidea) from the Americas and the Western Pacific</mods:title>
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<mods:namePart id="D6014FE4609C6A97EFEE141E50980D7C">Ahyong, Shane T.</mods:namePart>
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<mods:namePart id="88D6EE1B579164413AD5745DFDABC2D6">Campos, Ernesto</mods:namePart>
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<emphasis id="B904EAE9FFB38B70114E4E6723B4FC16" bold="true" box="[834,1430,895,920]" pageId="9" pageNumber="346">
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<taxonomicName id="4C704D78FFB38B70114E4E6722EAFC16" ID-CoL="84GBP" authority="(Burger, 1895)" authorityName="Ng & Ahyong & Campos" authorityYear="2019" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[834,1224,895,919]" class="Malacostraca" family="Pinnotheridae" genus="Tacitotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="346" phylum="Arthropoda" rank="species" species="glaber" status="comb. nov.">
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<emphasis id="B904EAE9FFB38B70114E4E67223BFC16" bold="true" box="[834,1049,895,919]" italics="true" pageId="9" pageNumber="346">Tacitotheres glaber</emphasis>
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(
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<bibRefCitation id="EFE14B0AFFB38B7016244E6722E0FC16" author="Burger O" box="[1064,1218,895,919]" pageId="9" pageNumber="346" pagination="361 - 390" refId="ref7836" refString="Burger O (1895) Ein Beitrag zur Kenntnis der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8 (3): 361 - 390, pls. 9, 10." type="book chapter" year="1895">Bürger, 1895</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="A2375792FFB38B7016DD4E9823B4FC16" box="[1233,1430,895,920]" pageId="9" pageNumber="346" rank="species">new combination</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C36A6570FFB38B7016224EB82288FC39" box="[1070,1194,928,952]" pageId="9" pageNumber="346" type="description">
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<paragraph id="8BCF36FBFFB38B7016224EB82288FC39" blockId="9.[834,1430,895,952]" box="[1070,1194,928,952]" pageId="9" pageNumber="346">
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(
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<figureCitation id="134B2A7EFFB38B70163A4EB8225BFC39" box="[1078,1145,928,952]" captionStart="Fig" captionStartId="10.[135,170,2064,2086]" captionTargetBox="[156,1454,193,2048]" captionTargetId="figure-8@10.[156,1454,193,2048]" captionTargetPageId="10" captionText="Fig. 9. Tacitotheres glaber (Bürger, 1895) new combination, A, paralectotype male (4.6 × 4.2 mm) (SMF-ZMG 173); B, C, lectotype ovigerous female (7.8 × 6.5 mm) (USNM 32431), Palau. A, B, overall dorsal habitus; C, ventral surface of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.4576195" httpUri="https://zenodo.org/record/4576195/files/figure.png" pageId="9" pageNumber="346">Fig. 9</figureCitation>
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,
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<figureCitation id="134B2A7EFFB38B70168A4EB82283FC39" box="[1158,1185,928,952]" captionStart="Fig" captionStartId="11.[113,149,1951,1973]" captionTargetBox="[116,1444,255,1930]" captionTargetId="figure-13@11.[113,1451,250,1935]" captionTargetPageId="11" captionText="Fig. 10. Tacitotheres glaber (Bürger, 1895) new combination, A–E, lectotype female (7.8 × 6.5 mm) (USNM 32431), Palau; F–I, paralectotype male (4.0 × 3.6 mm) (SMF-ZMG 173); J–M, paralectotype male (4.6 × 4.2 mm) (SMF-ZMG 173). A, F, dorsal overall habitus; B, sternum, anterior margin; C, anterior cephalothorax; E, H, right MXP3; D, G, right chela, outer view; I, J, male pleon; K, left G1, pleonal view; L, left G2, pleonal view, with long exopod; M, left MXP3. A, D–I, modified after Ahyong & Ng (2007a: fig. 24). Scale: A = 2 mm; B–D, F–G, I, J = 1.0 mm; E = 0.5; H, K, L, M = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4576198" httpUri="https://zenodo.org/record/4576198/files/figure.png" pageId="9" pageNumber="346">10</figureCitation>
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)
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</paragraph>
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<paragraph id="8BCF36FBFFB38B7011214EC6238AFBAC" blockId="9.[813,1452,990,1214]" pageId="9" pageNumber="346">
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<treatmentCitationGroup id="AB6011D5FFB38B7011214EC6238AFBAC" pageId="9" pageNumber="346">
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<treatmentCitation id="0AD110EAFFB38B7011214EC622E2FC72" author="Burger O" box="[813,1216,990,1012]" page="379" pageId="9" pageNumber="346" year="1895">
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<taxonomicName id="4C704D78FFB38B7011214EC622E2FC72" ID-CoL="7Y7Y2" authority="Burger, 1895: 379" authorityName="Burger" authorityPageNumber="379" authorityYear="1895" box="[813,1216,990,1012]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="346" phylum="Arthropoda" rank="species" species="glaber">
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<emphasis id="B904EAE9FFB38B7011214EC625D1FC72" box="[813,1011,990,1011]" italics="true" pageId="9" pageNumber="346">Pinnotheres glaber</emphasis>
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<bibRefCitation id="EFE14B0AFFB38B7011F24EC722E2FC72" author="Burger O" box="[1022,1216,990,1012]" pageId="9" pageNumber="346" pagination="361 - 390" refId="ref7836" refString="Burger O (1895) Ein Beitrag zur Kenntnis der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8 (3): 361 - 390, pls. 9, 10." type="book chapter" year="1895">Bürger, 1895: 379</bibRefCitation>
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, pl. 9: fig. 23, pl. 10: fig. 21;
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<treatmentCitation id="0AD110EAFFB38B7011AF4EE32271FB91" author="Tesch JJ" box="[931,1107,1019,1041]" page="248" pageId="9" pageNumber="346" year="1918">
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<bibRefCitation id="EFE14B0AFFB38B7011AF4EE32271FB91" author="Tesch JJ" box="[931,1107,1019,1041]" pageId="9" pageNumber="346" pagination="149 - 295" refId="ref10190" refString="Tesch JJ (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1: 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch, 1918: 248</bibRefCitation>
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;
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<treatmentCitation id="0AD110EAFFB38B70166C4EE3233DFB91" author="Miyake S" box="[1120,1311,1019,1041]" page="221" pageId="9" pageNumber="346" year="1939">
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<bibRefCitation id="EFE14B0AFFB38B70166C4EE3233DFB91" author="Miyake S" box="[1120,1311,1019,1041]" pageId="9" pageNumber="346" pagination="168 - 247" refId="ref9373" refString="Miyake S (1939) Notes on crustacea Brachyura collected by Prof. Teiso Esaki's Micronesia expeditions 1937 - 1938 together with check list of Micronesian Brachyura. Records of Oceanographic Works in Japan, 10 (2): 168 - 247, figs. 1 - 13, pls. 12 - 17." type="journal article" year="1939">Miyake, 1939: 221</bibRefCitation>
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, 241;
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<treatmentCitation id="0AD110EAFFB38B70176C4EE32273FBAC" author="Silas EG & Alagarswami K" page="1199" pageId="9" pageNumber="346" year="1967">
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<bibRefCitation id="EFE14B0AFFB38B70176C4EE32273FBAC" author="Silas EG & Alagarswami K" pageId="9" pageNumber="346" pagination="1161 - 1227" refId="ref10101" refString="Silas EG & Alagarswami K (1967) On an instance of parasitisation by the pea-crab (Pinnotheres sp.) on the backwater clam [Meretrix casta (Chemnitz)] from India, with a review of the work on systematics, ecology, biology and ethology of pea crabs of the genus Pinnotheres, Latreille, 1802. Proceedings of the Symposium on Crustacea held at Ernakulam, 1965, Symposium Series 2. Marine Biological Association of India, 3: 1161 - 1227." type="journal article" year="1967">Silas & Alagarswami, 1967: 1199</bibRefCitation>
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</treatmentCitation>
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, 1216;
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<treatmentCitation id="0AD110EAFFB38B70169749002386FBAC" author="Ng PKL & Manning RB" box="[1179,1444,1048,1070]" page="902" pageId="9" pageNumber="346" year="2003">
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<bibRefCitation id="EFE14B0AFFB38B70169749002386FBAC" author="Ng PKL & Manning RB" box="[1179,1444,1048,1070]" pageId="9" pageNumber="346" pagination="901 - 919" refId="ref9519" refString="Ng PKL & Manning RB (2003) On two new genera of pea crabs parasitic in holothurians (Crustacea: Decapoda: Brachyura: Pinnotheridae) from the IndoWest Pacific, with notes on allied genera. Proceedings of the Biological Society of Washington, 116 (4): 901 - 919." type="journal article" year="2003">Ng & Manning, 2003: 902</bibRefCitation>
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</treatmentCitation>
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.
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</paragraph>
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<paragraph id="8BCF36FBFFB38B701121492D25E5FB05" blockId="9.[813,1452,990,1214]" pageId="9" pageNumber="346">
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<treatmentCitationGroup id="AB6011D5FFB38B701121492D25E5FB05" pageId="9" pageNumber="346">
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<treatmentCitation id="0AD110EAFFB38B701121492D22EDFBCB" author="Burger O" box="[813,1231,1077,1099]" page="380" pageId="9" pageNumber="346" year="1895">
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<taxonomicName id="4C704D78FFB38B701121492D22EDFBCB" ID-CoL="7Y7Y5" authority="Burger, 1895: 380" authorityName="Burger" authorityPageNumber="380" authorityYear="1895" box="[813,1231,1077,1099]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="346" phylum="Arthropoda" rank="species" species="impressus">
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<emphasis id="B904EAE9FFB38B701121492D222EFBCB" box="[813,1036,1077,1098]" italics="true" pageId="9" pageNumber="346">Pinnotheres impressus</emphasis>
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<bibRefCitation id="EFE14B0AFFB38B701619492D22EDFBCB" author="Burger O" box="[1045,1231,1077,1099]" pageId="9" pageNumber="346" pagination="361 - 390" refId="ref7836" refString="Burger O (1895) Ein Beitrag zur Kenntnis der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8 (3): 361 - 390, pls. 9, 10." type="book chapter" year="1895">Bürger, 1895: 380</bibRefCitation>
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, pl. 9: fig. 24, pl. 10: fig. 23;
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<treatmentCitation id="0AD110EAFFB38B7011AF49492273FBE6" author="Tesch JJ" box="[931,1105,1105,1127]" page="249" pageId="9" pageNumber="346" year="1918">
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<bibRefCitation id="EFE14B0AFFB38B7011AF49492273FBE6" author="Tesch JJ" box="[931,1105,1105,1127]" pageId="9" pageNumber="346" pagination="149 - 295" refId="ref10190" refString="Tesch JJ (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1: 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch, 1918: 249</bibRefCitation>
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</treatmentCitation>
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, 255;
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<treatmentCitation id="0AD110EAFFB38B70169E494925A6FB02" author="Silas EG & Alagarswami K" page="1200" pageId="9" pageNumber="346" year="1967">
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<bibRefCitation id="EFE14B0AFFB38B70169E494925A6FB02" author="Silas EG & Alagarswami K" pageId="9" pageNumber="346" pagination="1161 - 1227" refId="ref10101" refString="Silas EG & Alagarswami K (1967) On an instance of parasitisation by the pea-crab (Pinnotheres sp.) on the backwater clam [Meretrix casta (Chemnitz)] from India, with a review of the work on systematics, ecology, biology and ethology of pea crabs of the genus Pinnotheres, Latreille, 1802. Proceedings of the Symposium on Crustacea held at Ernakulam, 1965, Symposium Series 2. Marine Biological Association of India, 3: 1161 - 1227." type="journal article" year="1967">Silas & Alagarswami, 1967: 1200</bibRefCitation>
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</treatmentCitation>
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, 1216.
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</paragraph>
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<paragraph id="8BCF36FBFFB38B701121499325C3FB3F" blockId="9.[813,1452,990,1214]" pageId="9" pageNumber="346">
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<treatmentCitationGroup id="AB6011D5FFB38B701121499325C3FB3F" pageId="9" pageNumber="346">
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<taxonomicName id="4C704D78FFB38B701121499325CBFB21" ID-CoL="7XSLV" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[813,1001,1163,1184]" class="Malacostraca" family="Pinnotheridae" genus="Orthotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="346" phylum="Arthropoda" rank="species" species="glaber">
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<emphasis id="B904EAE9FFB38B701121499325CBFB21" box="[813,1001,1163,1184]" italics="true" pageId="9" pageNumber="346">Orthotheres glaber</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="0AD110EAFFB38B70160B49932334FB21" author="Ahyong ST & Ng PKL" box="[1031,1302,1163,1185]" page="218" pageId="9" pageNumber="346" year="2007">
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<bibRefCitation id="EFE14B0AFFB38B70160B49932334FB21" author="Ahyong ST & Ng PKL" box="[1031,1302,1163,1185]" pageId="9" pageNumber="346" pagination="191 - 226" refId="ref7584" refString="Ahyong ST & Ng PKL (2007 a) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura. Raffles Bulletin of Zoology, Supplement 16: 191 - 226." type="journal article" year="2007">Ahyong & Ng, 2007a: 218</bibRefCitation>
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|
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, fig. 24;
|
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<treatmentCitation id="0AD110EAFFB38B70177F499425FFFB3F" author="Ng PKL & Guinot D & Davie PJF" page="250" pageId="9" pageNumber="346" year="2008">
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<bibRefCitation id="EFE14B0AFFB38B70177F499425FFFB3F" author="Ng PKL & Guinot D & Davie PJF" pageId="9" pageNumber="346" pagination="1 - 286" refId="ref9429" refString="Ng PKL, Guinot D & Davie PJF (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286." type="journal article" year="2008">Ng et al., 2008: 250</bibRefCitation>
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</treatmentCitation>
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.
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<subSubSection id="C36A6570FFB38B70112149FE2299FAFF" pageId="9" pageNumber="346" type="materials_examined">
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<paragraph id="8BCF36FBFFB38B70112149FE2299FAFF" blockId="9.[813,1454,1254,1406]" pageId="9" pageNumber="346">
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<emphasis id="B904EAE9FFB38B70112149FE2232FB7F" bold="true" box="[813,1040,1254,1278]" pageId="9" pageNumber="346">Material examined.</emphasis>
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<materialsCitation id="3B183CA6FFB38B70161549FF25A5FABF" collectionCode="USNM" collectorName="C. Semper." country="Palau" location="Palau" pageId="9" pageNumber="346" specimenCode="USNM 32431" specimenCount="1" typeStatus="lectotype">
|
||
<typeStatus id="54CB8859FFB38B70161549FF22ABFB7E" box="[1049,1161,1255,1279]" pageId="9" pageNumber="346" type="lectotype">Lectotype</typeStatus>
|
||
:
|
||
<specimenCode id="DBD69E80FFB38B70169A49FE2316FB7F" box="[1174,1332,1254,1278]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="346" type="Museum">USNM 32431</specimenCode>
|
||
, ovigerous female (7.8 × 6.5 mm),
|
||
<collectingCountry id="F367766BFFB38B70163D481E224DFA9F" box="[1073,1135,1286,1310]" name="Palau" pageId="9" pageNumber="346">Palau</collectingCountry>
|
||
, from
|
||
<taxonomicName id="4C704D78FFB38B7016B4481F236FFA9F" baseAuthorityName="Lamarck" baseAuthorityYear="1818" box="[1208,1357,1286,1310]" class="Bivalvia" family="Veneridae" genus="Tapes" kingdom="Animalia" order="Veneroida" pageId="9" pageNumber="346" phylum="Mollusca" rank="species" species="turgida">
|
||
<emphasis id="B904EAE9FFB38B7016B4481F236FFA9F" box="[1208,1357,1286,1310]" italics="true" pageId="9" pageNumber="346">Tapes turgida</emphasis>
|
||
</taxonomicName>
|
||
, coll.
|
||
<collectorName id="2685532DFFB38B70179F481E25A5FABF" pageId="9" pageNumber="346">C. Semper.</collectorName>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B183CA6FFB38B701183483E2294FAFF" collectionCode="USNM" collectorName="C. Semper." country="Palau" location="Palau" pageId="9" pageNumber="346" specimenCount="10" specimenCount-female="4" specimenCount-male="6" typeStatus="paralectotype">
|
||
<typeStatus id="54CB8859FFB38B701183483E220DFABF" box="[911,1071,1318,1342]" pageId="9" pageNumber="346" type="paralectotype">Paralectotypes</typeStatus>
|
||
: SMF-ZMG 173, Go304a,
|
||
<specimenCount id="9D76FD72FFB38B701754483E238FFABF" box="[1368,1453,1318,1342]" pageId="9" pageNumber="346" type="male">6 males</specimenCount>
|
||
(3.4 × 3.1 to 4.8 × 4.3 mm),
|
||
<specimenCount id="9D76FD72FFB38B70167C485E22FBFADF" box="[1136,1241,1350,1374]" pageId="9" pageNumber="346" type="female">4 females</specimenCount>
|
||
(3.5 × 3.5 to 8.3 × 7.4 mm), same locality as lectotype
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A6570FFB38B75112148BE279FFD36" lastPageId="12" lastPageNumber="349" pageId="9" pageNumber="346" type="diagnosis">
|
||
<paragraph id="8BCF36FBFFB38B75112148BE279FFD36" blockId="9.[811,1454,1446,2079]" lastBlockId="12.[135,777,191,696]" lastPageId="12" lastPageNumber="349" pageId="9" pageNumber="346">
|
||
<emphasis id="B904EAE9FFB38B70112148BE2582FA3F" bold="true" box="[813,928,1446,1470]" pageId="9" pageNumber="346">Diagnosis.</emphasis>
|
||
<emphasis id="B904EAE9FFB38B7011A948BE25DEFA3F" box="[933,1020,1446,1470]" italics="true" pageId="9" pageNumber="346">Female:</emphasis>
|
||
Carapace subcircular, slightly wider than long, front faintly projecting, convex. Eyes small, not visible in dorsal view. MXP3 with propodus more than twice as long as high, apex rounded; dactylus inserting subdistally, outreaching propodal apex; ischiomerus exceeding twice length, outer margin convex, inner margin with proximal ¾ concave, angular at distal quarter; exopod outer margin convex, inner margin relatively straight, flagellum 2-segmented. Cheliped with dactylus about 2/3 palm length; palm 2½ times as long as high, mesioventral margin setose; dactylus occlusal margin finely setose, with large proximal tooth; pollex occlusal margin sinuous, sparsely setose. Ambulatory legs asymmetrical in length from right to left; propodi broadening distally; relative lengths P3>P4>P2>P5. P2–P5 dactyli sparsely setose, shorter than half-length of respective propodi; relative lengths P3>P4>P2>P5. P4 and P5 propodus with sparse ventral setae. Pleon extending to buccal region and covering bases of ambulatory legs.
|
||
<emphasis id="B904EAE9FFB38B7011214AFE254EF87F" box="[813,876,2022,2046]" italics="true" pageId="9" pageNumber="346">Male:</emphasis>
|
||
Carapace rounded, wider than long, front projecting, medially emarginate. Eyes visible in dorsal view. MXP3 as in female, but sometimes with small distinct tooth at inner distal angle of ischiomerus. Cheliped robust; dactylus almost as long as palm; palm almost as high as long, outer surface with shallow, irregular pits; dactylus occlusal margin with prominent, blunt proximal tooth, distal margin crenulate, sparsely setose; pollex occlusal margin finely dentate, sparsely setose; mesial margin of carpus setose. Ambulatory legs dorsally and ventrally unarmed, length asymmetrical from left to right; relative lengths P3>P2>P4>P5. Dactyli shorter than half-length of respective propodi; P2–P4 dactyli subequal in length and slightly longer than P5 dactylus. P3 and P4 propodus with dorsal and ventral setae; carpus with diagonal setal row. Pleon widest at somites 3 and 4, tapering distally to evenly rounded telson. G1 slender, arcuate. G2 with exopod distinctly longer then endopod. (Modified from
|
||
<bibRefCitation id="EFE14B0AFFB68B7512844F872793FD36" author="Ahyong ST & Ng PKL" box="[136,433,671,696]" pageId="12" pageNumber="349" pagination="191 - 226" refId="ref7584" refString="Ahyong ST & Ng PKL (2007 a) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura. Raffles Bulletin of Zoology, Supplement 16: 191 - 226." type="journal article" year="2007">Ahyong & Ng, 2007a: 218</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF0F6673FFB08B73128B4508231FF7C2" ID-DOI="http://doi.org/10.5281/zenodo.4576195" ID-Zenodo-Dep="4576195" httpUri="https://zenodo.org/record/4576195/files/figure.png" pageId="10" pageNumber="347" startId="10.[135,170,2064,2086]" targetBox="[156,1454,193,2048]" targetPageId="10">
|
||
<paragraph id="8BCF36FBFFB08B73128B4508231FF7C2" blockId="10.[135,1474,2064,2115]" pageId="10" pageNumber="347">
|
||
Fig. 9.
|
||
<taxonomicName id="4C704D78FFB08B7312DD4508240EF7A7" authority="(Burger, 1895)" authorityName="Ng & Ahyong & Campos" authorityYear="2019" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[209,556,2064,2086]" class="Malacostraca" family="Pinnotheridae" genus="Tacitotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="347" phylum="Arthropoda" rank="species" species="glaber" status="comb. nov.">
|
||
<emphasis id="B904EAE9FFB08B7312DD450827B3F7A4" box="[209,401,2064,2085]" italics="true" pageId="10" pageNumber="347">Tacitotheres glaber</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE14B0AFFB08B7313AD45082407F7A7" author="Burger O" box="[417,549,2064,2086]" pageId="10" pageNumber="347" pagination="361 - 390" refId="ref7836" refString="Burger O (1895) Ein Beitrag zur Kenntnis der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8 (3): 361 - 390, pls. 9, 10." type="book chapter" year="1895">Bürger, 1895</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2375792FFB08B731039450824C2F7A7" box="[565,736,2064,2086]" pageId="10" pageNumber="347" rank="species">new combination</taxonomicNameLabel>
|
||
, A, paralectotype male (4.6 × 4.2 mm) (SMF-ZMG 173); B, C, lectotype ovigerous female (7.8 × 6.5 mm) (USNM 32431), Palau. A, B, overall dorsal habitus; C, ventral surface of cephalothorax.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0F6673FFB18B72127D4A872555F7A6" ID-DOI="http://doi.org/10.5281/zenodo.4576198" ID-Zenodo-Dep="4576198" httpUri="https://zenodo.org/record/4576198/files/figure.png" pageId="11" pageNumber="348" startId="11.[113,149,1951,1973]" targetBox="[116,1444,255,1930]" targetPageId="11">
|
||
<paragraph id="8BCF36FBFFB18B72127D4A872555F7A6" blockId="11.[113,1452,1950,2088]" pageId="11" pageNumber="348">
|
||
Fig. 10.
|
||
<taxonomicName id="4C704D78FFB18B7212C74A872412F835" authority="(Burger, 1895)" authorityName="Ng & Ahyong & Campos" authorityYear="2019" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[203,560,1950,1972]" class="Malacostraca" family="Pinnotheridae" genus="Tacitotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="348" phylum="Arthropoda" rank="species" species="glaber" status="comb. nov.">
|
||
<emphasis id="B904EAE9FFB18B7212C74A8727B2F835" box="[203,400,1951,1972]" italics="true" pageId="11" pageNumber="348">Tacitotheres glaber</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE14B0AFFB18B7213AD4A87240BF835" author="Burger O" box="[417,553,1950,1972]" pageId="11" pageNumber="348" pagination="361 - 390" refId="ref7836" refString="Burger O (1895) Ein Beitrag zur Kenntnis der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8 (3): 361 - 390, pls. 9, 10." type="book chapter" year="1895">Bürger, 1895</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2375792FFB18B7210364A8724C8F834" box="[570,746,1951,1973]" pageId="11" pageNumber="348" rank="species">new combination</taxonomicNameLabel>
|
||
, A–E, lectotype female (7.8 × 6.5 mm) (USNM 32431), Palau; F–I, paralectotype male (4.0 × 3.6 mm) (SMF-ZMG 173); J–M, paralectotype male (4.6 × 4.2 mm) (SMF-ZMG 173). A, F, dorsal overall habitus; B, sternum, anterior margin; C, anterior cephalothorax; E, H, right MXP3; D, G, right chela, outer view; I, J, male pleon; K, left G1, pleonal view; L, left G2, pleonal view, with long exopod; M, left MXP3. A, D–I, modified after
|
||
<bibRefCitation id="EFE14B0AFFB18B7216764AED2376F78B" author="Ahyong ST & Ng PKL" box="[1146,1364,2037,2059]" pageId="11" pageNumber="348" pagination="191 - 226" refId="ref7584" refString="Ahyong ST & Ng PKL (2007 a) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura. Raffles Bulletin of Zoology, Supplement 16: 191 - 226." type="journal article" year="2007">Ahyong & Ng (2007a</bibRefCitation>
|
||
: fig. 24). Scale: A = 2 mm; B–D, F–G, I, J = 1.0 mm; E = 0.5; H, K, L, M = 0.25 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C36A6570FFB68B7512844FC72524FAB6" pageId="12" pageNumber="349" type="discussion">
|
||
<paragraph id="8BCF36FBFFB68B7512844FC72524FAB6" blockId="12.[134,775,735,1335]" pageId="12" pageNumber="349">
|
||
<emphasis id="B904EAE9FFB68B7512844FC726D5FD76" bold="true" box="[136,247,735,759]" pageId="12" pageNumber="349">Remarks.</emphasis>
|
||
It is somewhat surprising that
|
||
<taxonomicName id="4C704D78FFB68B7510324FF82482FD76" authorityName="Ng & Ahyong & Campos" authorityYear="2019" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[574,672,735,759]" class="Malacostraca" family="Pinnotheridae" genus="Tacitotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="349" phylum="Arthropoda" rank="species" species="glaber">
|
||
<emphasis id="B904EAE9FFB68B7510324FF82482FD76" box="[574,672,735,759]" italics="true" pageId="12" pageNumber="349">T. glaber</emphasis>
|
||
</taxonomicName>
|
||
is known only from the
|
||
<typeStatus id="54CB8859FFB68B75133C4E18277DFC99" box="[304,351,768,792]" pageId="12" pageNumber="349">type</typeStatus>
|
||
material given that it was originally reported from venerid clams, which are widely collected for human consumption. The original host,
|
||
<taxonomicName id="4C704D78FFB68B7510624E58276CFCF6" authority="(Lamarck, 1818)" baseAuthorityName="Lamarck" baseAuthorityYear="1818" class="Bivalvia" family="Veneridae" genus="Tapes" kingdom="Animalia" order="Veneroida" pageId="12" pageNumber="349" phylum="Mollusca" rank="species" species="turgida">
|
||
<emphasis id="B904EAE9FFB68B7510624E582525FCD6" box="[622,775,831,855]" italics="true" pageId="12" pageNumber="349">Tapes turgida</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE14B0AFFB68B75129C4E472767FCF6" author="Lamarck JBPA de" box="[144,325,863,887]" pageId="12" pageNumber="349" pagination="1 - 612" refId="ref8845" refString="Lamarck JBPA de (1818) Histoire Naturelle des Animaux sans Vert ebres, presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent; pr ecedee d'une introduction offrant la determination des caracteres essentiels de l'Animal, sa distinction du vegetal et des autres corps naturels, enfin, l'Exposition des principes fondamentaux de la zoologie. Deterville, Paris, 5: 1 - 612." type="journal article" year="1818">Lamarck, 1818</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
is currently a junior synonym of
|
||
<taxonomicName id="4C704D78FFB68B7510FC4E782791FC16" authority="(Gmelin, 1791)" baseAuthorityName="Gmelin" baseAuthorityYear="1791" class="Bivalvia" family="Veneridae" genus="Tapes" kingdom="Animalia" order="Veneroida" pageId="12" pageNumber="349" phylum="Mollusca" rank="species" species="conspersus">
|
||
<emphasis id="B904EAE9FFB68B7510FC4E7826DDFC16" italics="true" pageId="12" pageNumber="349">T. conspersus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE14B0AFFB68B75131C4E672789FC16" author="Gmelin JF" box="[272,427,895,919]" pageId="12" pageNumber="349" pagination="3021 - 3910" refId="ref8540" refString="Gmelin JF (1791) Vermes. In: Gmelin JF (ed.) Caroli a Linnaei Systema Naturae per Regna Tria Naturae, Ed. 13. Tome 1 (6). GE Beer, Lipsiae [Leipzig]. Pp. 3021 - 3910." type="journal article" year="1791">Gmelin, 1791</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. An unusual aspect of male
|
||
<taxonomicName id="4C704D78FFB68B7510FD4E9826ECFC36" authorityName="Ng & Ahyong & Campos" authorityYear="2019" baseAuthorityName="Burger" baseAuthorityYear="1895" class="Malacostraca" family="Pinnotheridae" genus="Tacitotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="349" phylum="Arthropoda" rank="species" species="glaber">
|
||
<emphasis id="B904EAE9FFB68B7510FD4E9826ECFC36" italics="true" pageId="12" pageNumber="349">T. glaber</emphasis>
|
||
</taxonomicName>
|
||
is the size of the G2 exopod and pleonal form. The G2 exopod of
|
||
<taxonomicName id="4C704D78FFB68B7513214ED827B1FC56" authorityName="Ng & Ahyong & Campos" authorityYear="2019" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[301,403,959,983]" class="Malacostraca" family="Pinnotheridae" genus="Tacitotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="349" phylum="Arthropoda" rank="species" species="glaber">
|
||
<emphasis id="B904EAE9FFB68B7513214ED827B1FC56" box="[301,403,959,983]" italics="true" pageId="12" pageNumber="349">T. glaber</emphasis>
|
||
</taxonomicName>
|
||
is remarkable as it is the largest known of any pinnotherid (relatively), being about quadruple the length of the endopod (
|
||
<figureCitation id="134B2A7EFFB68B7513BA4EE72438FB96" box="[438,538,1023,1047]" captionStart="Fig" captionStartId="11.[113,149,1951,1973]" captionTargetBox="[116,1444,255,1930]" captionTargetId="figure-13@11.[113,1451,250,1935]" captionTargetPageId="11" captionText="Fig. 10. Tacitotheres glaber (Bürger, 1895) new combination, A–E, lectotype female (7.8 × 6.5 mm) (USNM 32431), Palau; F–I, paralectotype male (4.0 × 3.6 mm) (SMF-ZMG 173); J–M, paralectotype male (4.6 × 4.2 mm) (SMF-ZMG 173). A, F, dorsal overall habitus; B, sternum, anterior margin; C, anterior cephalothorax; E, H, right MXP3; D, G, right chela, outer view; I, J, male pleon; K, left G1, pleonal view; L, left G2, pleonal view, with long exopod; M, left MXP3. A, D–I, modified after Ahyong & Ng (2007a: fig. 24). Scale: A = 2 mm; B–D, F–G, I, J = 1.0 mm; E = 0.5; H, K, L, M = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4576198" httpUri="https://zenodo.org/record/4576198/files/figure.png" pageId="12" pageNumber="349">Fig. 10L</figureCitation>
|
||
). A large G2 exopod is also present in
|
||
<taxonomicName id="4C704D78FFB68B75134349072710FBD6" authority="Ahyong, Komai & Watanabe, 2012" authorityName="Ahyong, Komai & Watanabe" authorityYear="2012" class="Malacostraca" family="Pinnotheridae" genus="Viridotheres" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="349" phylum="Arthropoda" rank="species" species="takedai">
|
||
<emphasis id="B904EAE9FFB68B7513434907240CFBB6" box="[335,558,1055,1079]" italics="true" pageId="12" pageNumber="349">Viridotheres takedai</emphasis>
|
||
<bibRefCitation id="EFE14B0AFFB68B75103449072710FBD6" author="Ahyong ST & Komai T & Watanabe T" pageId="12" pageNumber="349" pagination="35 - 47" refId="ref7471" refString="Ahyong ST, Komai T & Watanabe T (2012) First Viridotheres Manning, 1996, from Japan, with a key to the species (Decapoda, Brachyura, Pinnotheridae). In: Komatsu H, Okuno J & Fukuoka K (eds.). Studies on Eumalacostraca: a homage to Takeda Masatsune. Crustaceana Monographs, Volume 17. Pp. 35 - 47." type="journal article" year="2012">Ahyong, Komai & Watanabe, 2012</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C704D78FFB68B75136F49582445FBD6" authority="Ahyong, 2019" authorityName="Ahyong" authorityYear="2019" box="[355,615,1087,1111]" class="Malacostraca" family="Pinnotheridae" genus="Viridotheres" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="349" phylum="Arthropoda" rank="species" species="kupang">
|
||
<emphasis id="B904EAE9FFB68B75136F495827E9FBD6" box="[355,459,1087,1111]" italics="true" pageId="12" pageNumber="349">V. kupang</emphasis>
|
||
<bibRefCitation id="EFE14B0AFFB68B7513DD49272445FBD6" author="Ahyong ST" box="[465,615,1087,1111]" pageId="12" pageNumber="349" pagination="107 - 118" refId="ref7429" refString="Ahyong ST (2019) First Indonesian Viridotheres Manning, 1996, and redescription of male Afropinnotheres dofleini Manning, 1993, from South Africa (Decapoda, Pinnotheridae). Crustaceana, 92 (1): 107 - 118." type="journal article" year="2019">Ahyong, 2019</bibRefCitation>
|
||
</taxonomicName>
|
||
, but the exopod is shorter than three times the length of the endopod. Of note is also the form of the male pleon. Unusually, the male pleon in
|
||
<taxonomicName id="4C704D78FFB68B7512AB49B8272CFB36" authorityName="Ng & Ahyong & Campos" authorityYear="2019" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[167,270,1183,1207]" class="Malacostraca" family="Pinnotheridae" genus="Tacitotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="349" phylum="Arthropoda" rank="species" species="glaber">
|
||
<emphasis id="B904EAE9FFB68B7512AB49B8272CFB36" box="[167,270,1183,1207]" italics="true" pageId="12" pageNumber="349">T. glaber</emphasis>
|
||
</taxonomicName>
|
||
is narrowest in the smallest specimens (
|
||
<figureCitation id="134B2A7EFFB68B7510D74987268EFB56" captionStart="Fig" captionStartId="11.[113,149,1951,1973]" captionTargetBox="[116,1444,255,1930]" captionTargetId="figure-13@11.[113,1451,250,1935]" captionTargetPageId="11" captionText="Fig. 10. Tacitotheres glaber (Bürger, 1895) new combination, A–E, lectotype female (7.8 × 6.5 mm) (USNM 32431), Palau; F–I, paralectotype male (4.0 × 3.6 mm) (SMF-ZMG 173); J–M, paralectotype male (4.6 × 4.2 mm) (SMF-ZMG 173). A, F, dorsal overall habitus; B, sternum, anterior margin; C, anterior cephalothorax; E, H, right MXP3; D, G, right chela, outer view; I, J, male pleon; K, left G1, pleonal view; L, left G2, pleonal view, with long exopod; M, left MXP3. A, D–I, modified after Ahyong & Ng (2007a: fig. 24). Scale: A = 2 mm; B–D, F–G, I, J = 1.0 mm; E = 0.5; H, K, L, M = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4576198" httpUri="https://zenodo.org/record/4576198/files/figure.png" pageId="12" pageNumber="349">Fig. 10I</figureCitation>
|
||
) becoming wider with increasing body size (
|
||
<figureCitation id="134B2A7EFFB68B75109249A724D8FB56" box="[670,762,1215,1239]" captionStart="Fig" captionStartId="11.[113,149,1951,1973]" captionTargetBox="[116,1444,255,1930]" captionTargetId="figure-13@11.[113,1451,250,1935]" captionTargetPageId="11" captionText="Fig. 10. Tacitotheres glaber (Bürger, 1895) new combination, A–E, lectotype female (7.8 × 6.5 mm) (USNM 32431), Palau; F–I, paralectotype male (4.0 × 3.6 mm) (SMF-ZMG 173); J–M, paralectotype male (4.6 × 4.2 mm) (SMF-ZMG 173). A, F, dorsal overall habitus; B, sternum, anterior margin; C, anterior cephalothorax; E, H, right MXP3; D, G, right chela, outer view; I, J, male pleon; K, left G1, pleonal view; L, left G2, pleonal view, with long exopod; M, left MXP3. A, D–I, modified after Ahyong & Ng (2007a: fig. 24). Scale: A = 2 mm; B–D, F–G, I, J = 1.0 mm; E = 0.5; H, K, L, M = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4576198" httpUri="https://zenodo.org/record/4576198/files/figure.png" pageId="12" pageNumber="349">Fig. 10J</figureCitation>
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), potentially becoming more female-like. The significance of this phenomenon is presently unknown and must await further collections allowing assessment of developmental changes.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |