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<document id="407F6903BB610801DA811DF2506ABBB7" ID-CLB-Dataset="99517" ID-DOI="10.11646/zootaxa.5165.4.1" ID-GBIF-Dataset="cd84b880-d77f-401a-bf4c-a98021a5e166" ID-ISSN="1175-5326" ID-Zenodo-Dep="6853942" ID-ZooBank="DBE812FF-7B25-4522-97FE-45A86ECAEFD7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658127043518" checkinUser="plazi" docAuthor="Yang, Qiao-Zhi &amp; Pathomwattananurak, Wuttipon" docDate="2022" docId="03D9879BE85417677EFDFB686898A3A3" docLanguage="en" docName="zootaxa.5165.4.1.pdf" docOrigin="Zootaxa 5165 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eophileurus felschei Prell 1913" docType="treatment" docVersion="6" lastPageNumber="460" masterDocId="FFE0FFE3E85D176C7E6AFFEE6B79A22A" masterDocTitle="Four new species of the genus Eophileurus Arrow, 1908 (Coleoptera: Scarabaeidae Dynastinae) from Thailand and Vietnam, with notes on some Indochinese species" masterLastPageNumber="485" masterPageNumber="451" pageNumber="460" updateTime="1699360166553" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9D3BB69068C578F1B0E5BE80A31CC59D">Four new species of the genus Eophileurus Arrow, 1908 (Coleoptera: Scarabaeidae Dynastinae) from Thailand and Vietnam, with notes on some Indochinese species</mods:title>
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<mods:namePart id="9DEF70358881161FADB4AACAF537B00D">Pathomwattananurak, Wuttipon</mods:namePart>
<mods:affiliation id="0AECC52518E2D5A9A8A6DE0696E9E24B">1051 / 38 Wiang, Mueang Dist., Chiang Rai Province, 57000, Thailand.</mods:affiliation>
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<heading id="D08781E1E85417657EFDFB686967A68A" bold="true" box="[151,542,1158,1185]" fontSize="11" level="1" pageId="9" pageNumber="460" reason="1">
<taxonomicName id="4C704D0EE85417657EFDFB686967A68A" authority="Prell, 1913" authorityName="Prell" authorityYear="1913" box="[151,542,1158,1185]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="felschei">
<emphasis id="B904EA9FE85417657EFDFB686967A68A" bold="true" box="[151,542,1158,1185]" pageId="9" pageNumber="460">
<emphasis id="B904EA9FE85417657EFDFB686AE8A68B" bold="true" box="[151,401,1158,1185]" italics="true" pageId="9" pageNumber="460">Eophileurus felschei</emphasis>
<bibRefCitation id="EFE14B7CE85417657FF3FB696967A68A" author="Prell, H." box="[409,542,1158,1185]" pageId="9" pageNumber="460" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell, 1913</bibRefCitation>
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<paragraph id="8BCF368DE85417657EFDFB456A2EA6EF" blockId="9.[151,542,1158,1221]" box="[151,343,1194,1221]" pageId="9" pageNumber="460">
(
<figureCitation id="134B2A08E85417657EF5FB456B83A6EF" box="[159,250,1195,1221]" captionStart="FIGURE 1" captionStartId="2.[151,250,1829,1853]" captionTargetBox="[151,1435,1065,1805]" captionTargetId="figure-436@2.[151,1435,1065,1805]" captionTargetPageId="2" captionText="FIGURE 1. Elytra of Eophileurus Arrow, 1908 species. a, E. quadrigeminatus Arrow, 1914; b, E. quadratifovealis Yang &amp; Pathomwattananurak, new species; c, E. felschei Prell, 1913. Primary costae marked in yellow and labeled in capital letters, interstices labeled in numbers without additional coloring." figureDoi="http://doi.org/10.5281/zenodo.6853944" httpUri="https://zenodo.org/record/6853944/files/figure.png" pageId="9" pageNumber="460">Figs. 1c</figureCitation>
,
<figureCitation id="134B2A08E85417657F6DFB446A36A6EF" box="[263,335,1194,1221]" captionStart-0="FIGURES 38–48" captionStart-1="FIGURES 49–53" captionStartId-0="10.[151,264,1944,1968]" captionStartId-1="11.[151,264,1333,1357]" captionTargetBox-0="[253,1339,186,1912]" captionTargetBox-1="[156,1433,418,1309]" captionTargetId-0="figure-18@10.[241,1346,181,1920]" captionTargetId-1="figure-149@11.[151,1435,417,1309]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURES 38–48. Type specimens of Eophileurus felschei Prell, 1913. 38–31, lectotype, female, 38, habitus, dorsal; 39, habitus, lateral; 40, habitus, ventral; 41, labels; 42–48, lectoparatype, male (images ©MTD/Olaf Jäger), 42, habitus, dorsal; 43, habitus, lateral; 44, habitus, ventral, 45–46, parameres, apical; 47, parameres, lateral; 48, labels." captionText-1="FIGURES 49–53. Holotype of Eophileurus grossepunctatus Dupuis, 2014 (images ©MNHN/A. Mantilleri). 49, habitus, dorsal; 50, habitus, lateral; 51, parameres, apical; 52, parameres, lateral; 53, labels." figureDoi-0="http://doi.org/10.5281/zenodo.6853956" figureDoi-1="http://doi.org/10.5281/zenodo.6853958" httpUri-0="https://zenodo.org/record/6853956/files/figure.png" httpUri-1="https://zenodo.org/record/6853958/files/figure.png" pageId="9" pageNumber="460">38–53</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36A6506E85417657EFDFB0168CBA7A8" pageId="9" pageNumber="460" type="reference_group">
<paragraph id="8BCF368DE85417657EFDFB016E1BA76F" blockId="9.[151,1437,1263,1410]" pageId="9" pageNumber="460">
<treatmentCitationGroup id="AB6011A3E85417657EFDFB016E1BA76F" pageId="9" pageNumber="460">
<treatmentCitation id="0AD1109CE85417657EFDFB01696BA72D" author="Prell, H." box="[151,530,1263,1287]" page="111" pageId="9" pageNumber="460" year="1913">
<taxonomicName id="4C704D0EE85417657EFDFB01696BA72D" authority="Prell, 1913: 111" authorityName="Prell" authorityPageNumber="111" authorityYear="1913" box="[151,530,1263,1287]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="felschei">
<emphasis id="B904EA9FE85417657EFDFB016A1EA72D" box="[151,359,1263,1287]" italics="true" pageId="9" pageNumber="460">Eophileurus felschei</emphasis>
<bibRefCitation id="EFE14B7CE85417657F05FB01696BA72D" author="Prell, H." box="[367,530,1263,1287]" pageId="9" pageNumber="460" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell, 1913: 111</bibRefCitation>
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(
<typeStatus id="54CB882FE85417657C4AFB016934A72D" box="[544,589,1263,1287]" pageId="9" pageNumber="460">type</typeStatus>
locality: “
<collectingRegion id="49B4F86FE85417657CD2FB01698AA72D" box="[696,755,1263,1287]" country="India" name="Delhi" pageId="9" pageNumber="460">Delhi</collectingRegion>
Genji, Coromandel”), plates 1–2, fig. 4 (parameres);
<treatmentCitation id="0AD1109CE85417657B73FB016B9AA70C" author="Arrow, G. J." page="84" pageId="9" pageNumber="460" year="1937">
<bibRefCitation id="EFE14B7CE85417657B73FB016B9AA70C" author="Arrow, G. J." pageId="9" pageNumber="460" pagination="1 - 124" refId="ref16830" refString="Arrow, G. J. (1937) Scarabaeidae: Dynastinae. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Pars 156. W. Junk, Gravenhage, pp. 1 - 124." type="book chapter" year="1937">Arrow 1937: 84</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AD1109CE85417657E85FAE06AD6A70C" author="Endrodi, S." box="[239,431,1294,1318]" page="101" pageId="9" pageNumber="460" year="1977">
<bibRefCitation id="EFE14B7CE85417657E85FAE06AD6A70C" author="Endrodi, S." box="[239,431,1294,1318]" pageId="9" pageNumber="460" pagination="93 - 134" refId="ref16934" refString="Endrodi, S. (1977) Monographie de Dynastinae VIII Tribus Phileurini. Entomologische Abhandlungen Staatlichen Museum fur Tierkunde Dresden, 41 (4), 93 - 134. https: // doi. org / 10.1515 / 9783112546161" type="journal article" year="1977">Endrődi 1977: 101</bibRefCitation>
</treatmentCitation>
(Coromandel;
<collectingRegion id="49B4F86FE85417657C20FAE069FCA70C" box="[586,645,1294,1318]" country="India" name="Delhi" pageId="9" pageNumber="460">Delhi</collectingRegion>
; Genji; Madras; Taniore; Bhadravati), fig. 4 (parameres);
<treatmentCitation id="0AD1109CE85417657ABCFAE06EE1A70C" author="Endrodi, S." box="[1238,1432,1294,1318]" page="664" pageId="9" pageNumber="460" year="1985">
<bibRefCitation id="EFE14B7CE85417657ABCFAE06EE1A70C" author="Endrodi, S." box="[1238,1432,1294,1318]" pageId="9" pageNumber="460" refId="ref17001" refString="Endrodi, S. (1985) The Dynastinae of the World. W. Junk, Dordrecht, 800 pp." type="book" year="1985">Endrődi 1985: 664</bibRefCitation>
</treatmentCitation>
, fig. 2005 (parameres);
<treatmentCitation id="0AD1109CE85417657FC5FAC26927A76E" author="Krajcik, M." box="[431,606,1324,1348]" page="64" pageId="9" pageNumber="460" year="2005">
<bibRefCitation id="EFE14B7CE85417657FC5FAC26927A76E" author="Krajcik, M." box="[431,606,1324,1348]" pageId="9" pageNumber="460" pagination="1 - 122" refId="ref17162" refString="Krajcik, M. (2005) Dynastinae of the World Checklist (Coleoptera, Scarabaeidae, Dynastinae). Animma. x, Supplement 2, 1 - 122." type="journal article" year="2005">Krajčik 2005: 64</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AD1109CE85417657C0DFAC368DEA76E" author="Yamaya, S. &amp; Muramoto, R." box="[615,935,1324,1349]" page="26" pageId="9" pageNumber="460" year="2008">
<bibRefCitation id="EFE14B7CE85417657C0DFAC368DEA76E" author="Yamaya, S. &amp; Muramoto, R." box="[615,935,1324,1349]" pageId="9" pageNumber="460" pagination="1 - 34" refId="ref17677" refString="Yamaya, S. &amp; Muramoto, R. (2008) On the genus Eophileurus Arrow, 1908 (Scarabaeidae, Dynastinae), with descriptions of five new species. Kogane, Supplement 2, 1 - 34." type="journal article" year="2008">Yamaya &amp; Muramoto 2008: 26</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AD1109CE85417657DD8FAC26F14A76F" author="Krajcik, M." box="[946,1133,1324,1349]" page="101" pageId="9" pageNumber="460" year="2012">
<bibRefCitation id="EFE14B7CE85417657DD8FAC26F14A76F" author="Krajcik, M." box="[946,1133,1324,1349]" pageId="9" pageNumber="460" pagination="1 - 278" refId="ref17193" refString="Krajcik, M. (2012) Checklist of the World Scarabaeoidea. Animma. x, Supplement 5, 1 - 278." type="journal article" year="2012">Krajčik 2012: 101</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AD1109CE85417657A1DFAC26E27A76F" author="Jakl, S. &amp; Zidek, J." box="[1143,1374,1324,1349]" page="19" pageId="9" pageNumber="460" year="2015">
<bibRefCitation id="EFE14B7CE85417657A1DFAC26E27A76F" author="Jakl, S. &amp; Zidek, J." box="[1143,1374,1324,1349]" pageId="9" pageNumber="460" pagination="14 - 26" refId="ref17117" refString="Jakl, S. &amp; Zidek, J. (2015) Contribution to the knowledge of Oriental and Austro-Papuan Eophileurus Arrow, with descriptions of two new species. Folia Heyrovskyana, Series A, 23 (2), 14 - 26." type="journal article" year="2015">Jákl &amp; Zídek 2015: 19</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BCF368DE85417657EFDFAA568CBA7A8" blockId="9.[151,1437,1263,1410]" pageId="9" pageNumber="460">
<treatmentCitationGroup id="AB6011A3E85417657EFDFAA568CBA7A8" pageId="9" pageNumber="460">
<treatmentCitation id="0AD1109CE85417657EFDFAA56903A749" author="Dupuis, F." box="[151,634,1355,1379]" page="13" pageId="9" pageNumber="460" year="2014">
<taxonomicName id="4C704D0EE85417657EFDFAA56903A749" authority="Dupuis, 2014: 13" authorityName="Dupuis" authorityPageNumber="13" authorityYear="2014" box="[151,634,1355,1379]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="grossepunctatus">
<emphasis id="B904EA9FE85417657EFDFAA56AC6A749" box="[151,447,1355,1379]" italics="true" pageId="9" pageNumber="460">Eophileurus grossepunctatus</emphasis>
<bibRefCitation id="EFE14B7CE85417657FACFAA56903A749" author="Dupuis, F." box="[454,634,1355,1379]" pageId="9" pageNumber="460" pagination="13 - 18" refId="ref16872" refString="Dupuis, F. (2014) Deux nouvelles especes d' Eophileurus Arrow, 1908. Coleopteres, 20 (3), 13 - 18." type="journal article" year="2014">Dupuis, 2014: 13</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, (
<typeStatus id="54CB882FE85417657CE7FAA569C3A749" box="[653,698,1355,1379]" pageId="9" pageNumber="460">type</typeStatus>
locality: “
<collectingCountry id="F367761DE85417657D4CFAA5682CA749" box="[806,853,1355,1379]" name="India" pageId="9" pageNumber="460">Inde</collectingCountry>
Anglaise, Shimoga”), plate 13, fig. 7 (parameres); fig. 1 (male habitus), fig. 3 (parameres);
<treatmentCitation id="0AD1109CE85417657C42FA846877A7A8" author="Jakl, S. &amp; Zidek, J." box="[552,782,1386,1410]" page="19" pageId="9" pageNumber="460" year="2015">
<bibRefCitation id="EFE14B7CE85417657C42FA846877A7A8" author="Jakl, S. &amp; Zidek, J." box="[552,782,1386,1410]" pageId="9" pageNumber="460" pagination="14 - 26" refId="ref17117" refString="Jakl, S. &amp; Zidek, J. (2015) Contribution to the knowledge of Oriental and Austro-Papuan Eophileurus Arrow, with descriptions of two new species. Folia Heyrovskyana, Series A, 23 (2), 14 - 26." type="journal article" year="2015">Jákl &amp; Zídek 2015: 19</bibRefCitation>
</treatmentCitation>
;
<emphasis id="B904EA9FE85417657D73FA8468CBA7A8" bold="true" box="[793,946,1386,1410]" pageId="9" pageNumber="460">new synonym.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36A6506E85417657EFDFA5E6988A524" pageId="9" pageNumber="460" type="materials_examined">
<paragraph id="8BCF368DE85417657EFDFA5E6A09A4EC" blockId="9.[151,1437,1455,2023]" pageId="9" pageNumber="460">
<emphasis id="B904EA9FE85417657EFDFA5E68BDA7E0" bold="true" box="[151,964,1455,1482]" pageId="9" pageNumber="460">
Type material examined.
<typeStatus id="54CB882FE85417657FD5FA5E694CA7E0" box="[447,565,1456,1482]" pageId="9" pageNumber="460" type="lectotype">Lectotype</typeStatus>
of
<taxonomicName id="4C704D0EE85417657C3CFA4168BDA7E0" authority="Prell, 1913" authorityName="Prell" authorityYear="1913" box="[598,964,1455,1482]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="felschei">
<emphasis id="B904EA9FE85417657C3CFA41683AA7E0" bold="true" box="[598,835,1455,1482]" italics="true" pageId="9" pageNumber="460">Eophileurus felschei</emphasis>
<bibRefCitation id="EFE14B7CE85417657D2DFA5E68BDA7E0" author="Prell, H." box="[839,964,1456,1482]" pageId="9" pageNumber="460" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell, 1913</bibRefCitation>
</taxonomicName>
</emphasis>
(
<specimenCount id="9D76FD04E85417657DBEFA5E689EA7E0" box="[980,999,1456,1482]" pageId="9" pageNumber="460" type="female">♀</specimenCount>
,
<collectionCode id="ED61AE48E85417657D99FA5F6F3CA7E1" box="[1011,1093,1457,1483]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="460">MFNB</collectionCode>
,
<figureCitation id="134B2A08E85417657A24FA5E6FAAA7E0" box="[1102,1235,1456,1482]" captionStart="FIGURES 38–48" captionStartId="10.[151,264,1944,1968]" captionTargetBox="[253,1339,186,1912]" captionTargetId="figure-18@10.[241,1346,181,1920]" captionTargetPageId="10" captionText="FIGURES 38–48. Type specimens of Eophileurus felschei Prell, 1913. 38–31, lectotype, female, 38, habitus, dorsal; 39, habitus, lateral; 40, habitus, ventral; 41, labels; 42–48, lectoparatype, male (images ©MTD/Olaf Jäger), 42, habitus, dorsal; 43, habitus, lateral; 44, habitus, ventral, 45–46, parameres, apical; 47, parameres, lateral; 48, labels." figureDoi="http://doi.org/10.5281/zenodo.6853956" httpUri="https://zenodo.org/record/6853956/files/figure.png" pageId="9" pageNumber="460">Figs. 38–41</figureCitation>
), labeled: “
<collectionCode id="ED61AE48E85417657B26FA5E6EE5A7E0" box="[1356,1436,1456,1482]" pageId="9" pageNumber="460">GENJI</collectionCode>
<date id="FFCE104DE85417657EFDFA3A6AE6A7C4" box="[151,415,1492,1519]" pageId="9" pageNumber="460" value="1901-08-25" valueMax="1901-09-15" valueMin="1901-08-25">25 août – 15 sept 1901</date>
// COROMANDEL M. MAINDRON //
<taxonomicName id="4C704D0EE85417657D18FA3A6FEEA7C4" authority="Prell" authorityName="Prell" authorityYear="1913" box="[882,1175,1492,1518]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="felschei">
<emphasis id="B904EA9FE85417657D18FA3A6F23A7C4" box="[882,1114,1492,1518]" italics="true" pageId="9" pageNumber="460">Eophileurus felschei</emphasis>
Prell
</taxonomicName>
<specimenCount id="9D76FD04E85417657ACAFA3B6FCAA7C5" box="[1184,1203,1493,1519]" pageId="9" pageNumber="460" type="female">♀</specimenCount>
- Type //
<typeStatus id="54CB882FE85417657B4AFA3B6EE5A7C5" box="[1312,1436,1493,1519]" pageId="9" pageNumber="460" type="lectotype">Lectotypus</typeStatus>
<taxonomicName id="4C704D0EE85417657EFDFA166AC5A438" authority="Prell" authorityName="Prell" authorityYear="1913" box="[151,444,1528,1554]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="felschei">
<emphasis id="B904EA9FE85417657EFDFA166A06A438" box="[151,383,1528,1554]" italics="true" pageId="9" pageNumber="460">Eophileurus felschei</emphasis>
Prell
</taxonomicName>
SEndrődi”.
<emphasis id="B904EA9FE85417657C21FA166F66A438" bold="true" box="[587,1055,1527,1554]" pageId="9" pageNumber="460">
<typeStatus id="54CB882FE85417657C21FA166989A438" box="[587,752,1528,1554]" pageId="9" pageNumber="460" type="paralectotype">Paralectotype</typeStatus>
of
<taxonomicName id="4C704D0EE85417657D70FA166F66A438" authority="Prell, 1913" authorityName="Prell" authorityYear="1913" box="[794,1055,1527,1554]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="felschei">
<emphasis id="B904EA9FE85417657D70FA1668ECA438" bold="true" box="[794,917,1527,1554]" italics="true" pageId="9" pageNumber="460">E. felschei</emphasis>
<bibRefCitation id="EFE14B7CE85417657DF4FA166F66A438" author="Prell, H." box="[926,1055,1528,1554]" pageId="9" pageNumber="460" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell, 1913</bibRefCitation>
</taxonomicName>
</emphasis>
(
<specimenCount id="9D76FD04E85417657A5BFA166F31A438" box="[1073,1096,1528,1554]" pageId="9" pageNumber="460" type="male">♂</specimenCount>
,
<collectionCode id="ED61AE48E85417657A3CFA176FEEA439" box="[1110,1175,1529,1555]" country="Germany" httpUri="http://grbio.org/cool/4j3t-w40g" name="Museum of Zoology Senckenberg Dresden" pageId="9" pageNumber="460">MTD</collectionCode>
,
<figureCitation id="134B2A08E85417657ACEFA166E56A438" box="[1188,1327,1528,1554]" captionStart="FIGURES 38–48" captionStartId="10.[151,264,1944,1968]" captionTargetBox="[253,1339,186,1912]" captionTargetId="figure-18@10.[241,1346,181,1920]" captionTargetPageId="10" captionText="FIGURES 38–48. Type specimens of Eophileurus felschei Prell, 1913. 38–31, lectotype, female, 38, habitus, dorsal; 39, habitus, lateral; 40, habitus, ventral; 41, labels; 42–48, lectoparatype, male (images ©MTD/Olaf Jäger), 42, habitus, dorsal; 43, habitus, lateral; 44, habitus, ventral, 45–46, parameres, apical; 47, parameres, lateral; 48, labels." figureDoi="http://doi.org/10.5281/zenodo.6853956" httpUri="https://zenodo.org/record/6853956/files/figure.png" pageId="9" pageNumber="460">Figs. 42–48</figureCitation>
), labeled: “Sum.
<collectingRegion id="49B4F86FE85417657E8EF9F26A6DA41C" box="[228,276,1564,1590]" country="India" name="Delhi" pageId="9" pageNumber="460">Deli</collectingRegion>
// Typus // Coll. C. Felsche Kauf 20, 1918 // Staatl. Museum für Tierkunde Dresden”, examined through photographs provided by Olaf Jäger.
<emphasis id="B904EA9FE85417657C2FF9AE6F1CA470" bold="true" box="[581,1125,1600,1626]" pageId="9" pageNumber="460">
<typeStatus id="54CB882FE85417657C2FF9AE69CBA470" box="[581,690,1600,1626]" pageId="9" pageNumber="460" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C704D0EE85417657CB4F9AE6F1CA470" authority="Dupuis, 2014" authorityName="Dupuis" authorityYear="2014" box="[734,1125,1600,1626]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="grossepunctatus">
<emphasis id="B904EA9FE85417657CB4F9AE68C5A470" bold="true" box="[734,956,1600,1626]" italics="true" pageId="9" pageNumber="460">E. grossepunctatus</emphasis>
<bibRefCitation id="EFE14B7CE85417657DACF9AE6F1CA470" author="Dupuis, F." box="[966,1125,1600,1626]" pageId="9" pageNumber="460" pagination="13 - 18" refId="ref16872" refString="Dupuis, F. (2014) Deux nouvelles especes d' Eophileurus Arrow, 1908. Coleopteres, 20 (3), 13 - 18." type="journal article" year="2014">Dupuis, 2014</bibRefCitation>
</taxonomicName>
</emphasis>
(
<specimenCount id="9D76FD04E85417657A12F9AE6FF6A470" box="[1144,1167,1600,1626]" pageId="9" pageNumber="460" type="male">♂</specimenCount>
,
<collectionCode id="ED61AE48E85417657AF4F9AF6F8FA471" box="[1182,1270,1601,1627]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="460">MNHN</collectionCode>
,
<figureCitation id="134B2A08E85417657B6FF9AE6EF6A471" box="[1285,1423,1600,1627]" captionStart="FIGURES 49–53" captionStartId="11.[151,264,1333,1357]" captionTargetBox="[156,1433,418,1309]" captionTargetId="figure-149@11.[151,1435,417,1309]" captionTargetPageId="11" captionText="FIGURES 49–53. Holotype of Eophileurus grossepunctatus Dupuis, 2014 (images ©MNHN/A. Mantilleri). 49, habitus, dorsal; 50, habitus, lateral; 51, parameres, apical; 52, parameres, lateral; 53, labels." figureDoi="http://doi.org/10.5281/zenodo.6853958" httpUri="https://zenodo.org/record/6853958/files/figure.png" pageId="9" pageNumber="460">Figs. 49–53</figureCitation>
), labeled: “
<collectingCountry id="F367761DE85417657F6EF98A6A30A454" box="[260,329,1636,1662]" name="India" pageId="9" pageNumber="460">INDE</collectingCountry>
ANGLAISE Shimoga //
<taxonomicName id="4C704D0EE85417657C1BF98A685DA455" box="[625,804,1636,1663]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B904EA9FE85417657C1BF98A6983A454" box="[625,762,1636,1662]" italics="true" pageId="9" pageNumber="460">Eophileurus</emphasis>
sp.
</taxonomicName>
<collectionCode id="ED61AE48E85417657D44F98B683AA455" box="[814,835,1637,1663]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="460">R</collectionCode>
.-P. Dechambre //
<taxonomicName id="4C704D0EE85417657A7FF98A6E24A454" authority="Yang &amp; Pathomwattananurak, 2022" authorityName="Yang &amp; Pathomwattananurak" authorityYear="2022" box="[1045,1373,1636,1662]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="grossepunctatus" status="sp. nov.">
<emphasis id="B904EA9FE85417657A7FF98A6E24A454" box="[1045,1373,1636,1662]" italics="true" pageId="9" pageNumber="460">Eophileurus grossepunctatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23757E4E85417657B0DF98B6EE5A455" box="[1383,1436,1637,1663]" pageId="9" pageNumber="460" rank="species">n.sp.</taxonomicNameLabel>
<typeStatus id="54CB882FE85417657EFDF9666B86A488" box="[151,255,1672,1698]" pageId="9" pageNumber="460" type="holotype">Holotype</typeStatus>
F.
<bibRefCitation id="EFE14B7CE85417657F48F9666ACBA488" author="Dupuis, F." box="[290,434,1672,1698]" pageId="9" pageNumber="460" pagination="13 - 18" refId="ref16872" refString="Dupuis, F. (2014) Deux nouvelles especes d' Eophileurus Arrow, 1908. Coleopteres, 20 (3), 13 - 18." type="journal article" year="2014">Dupuis 2014</bibRefCitation>
//
<typeStatus id="54CB882FE85417657FBAF966691DA488" box="[464,612,1672,1698]" pageId="9" pageNumber="460" type="holotype">HOLOTYPE</typeStatus>
//
<collectionCode id="ED61AE48E85417657CE8F96769A1A489" box="[642,728,1673,1699]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="460">MNHN</collectionCode>
EC4413”, examined through website photographs provided by Antoine Mantilleri.
</paragraph>
<paragraph id="8BCF368DE85417657EADF93E6988A524" blockId="9.[151,1437,1455,2023]" pageId="9" pageNumber="460">
<emphasis id="B904EA9FE85417657EADF93E6957A4C0" bold="true" box="[199,558,1744,1770]" pageId="9" pageNumber="460">Additional material examined</emphasis>
(
<specimenCount id="9D76FD04E85417657C2AF93E691DA4C0" box="[576,612,1744,1770]" pageId="9" pageNumber="460" type="male">1♂</specimenCount>
).
<emphasis id="B904EA9FE85417657C10F93E69BAA4C0" bold="true" box="[634,707,1744,1770]" pageId="9" pageNumber="460">
<collectingCountry id="F367761DE85417657C10F93E69C7A4C0" box="[634,702,1744,1770]" name="India" pageId="9" pageNumber="460">India</collectingCountry>
:
</emphasis>
<materialsCitation id="3B183CD0E85417657CA1F93E6988A524" ID-GBIF-Occurrence="3863916337" collectingDate="2010-10-06" collectionCode="CQZY" collectorName="Local Collector" country="India" county="Dwarakapuram Village" location="Nellore Dist." municipality="Naidupet Mandal" pageId="9" pageNumber="460" specimenCount="1" specimenCount-male="1" stateProvince="Andhra Pradesh">
<specimenCount id="9D76FD04E85417657CA1F93E6997A4C0" box="[715,750,1744,1770]" pageId="9" pageNumber="460" type="male">1♂</specimenCount>
(
<collectionCode id="ED61AE48E85417657C97F93E6836A4C0" box="[765,847,1744,1770]" pageId="9" pageNumber="460">CQZY</collectionCode>
),
<collectingCounty id="62AE4E01E85417657D09F93E6F12A4C0" box="[867,1131,1744,1770]" pageId="9" pageNumber="460">Dwarakapuram Village</collectingCounty>
,
<collectingMunicipality id="6BABACF7E85417657A1CF93E6E45A4C0" box="[1142,1340,1744,1770]" pageId="9" pageNumber="460">Naidupet Mandal</collectingMunicipality>
,
<location id="8EAF6056E85417657B22F93E6BA9A524" LSID="urn:lsid:plazi:treatment:03D9879BE85417677EFDFB686898A3A3:8EAF6056E85417657B22F93E6BA9A524" country="India" county="Dwarakapuram Village" municipality="Naidupet Mandal" name="Nellore Dist." pageId="9" pageNumber="460" stateProvince="Andhra Pradesh">Nellore Dist.</location>
,
<collectingRegion id="49B4F86FE85417657EB3F91A6AE9A524" box="[217,400,1780,1806]" country="India" name="Andhra Pradesh" pageId="9" pageNumber="460">Andhra Pradesh</collectingRegion>
,
<date id="FFCE104DE85417657FF1F91A697FA524" box="[411,518,1780,1806]" pageId="9" pageNumber="460" value="2010-10-06">
<collectingDate id="EF8AE9A5E85417657FF1F91A697FA524" box="[411,518,1780,1806]" pageId="9" pageNumber="460" value="2010-10-06">6.X.2010</collectingDate>
</date>
,
<collectorName id="2685535BE85417657C7BF91A69B9A524" box="[529,704,1780,1806]" pageId="9" pageNumber="460">Local Collector</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36A6506E85417657EADF8F66AD0A518" box="[199,425,1816,1842]" pageId="9" pageNumber="460" type="distribution">
<paragraph id="8BCF368DE85417657EADF8F66AD0A518" blockId="9.[151,1437,1455,2023]" box="[199,425,1816,1842]" pageId="9" pageNumber="460">
<emphasis id="B904EA9FE85417657EADF8F66A18A518" bold="true" box="[199,353,1816,1842]" pageId="9" pageNumber="460">Distribution.</emphasis>
<collectingCountry id="F367761DE85417657F0DF8F66ADDA518" box="[359,420,1816,1842]" name="India" pageId="9" pageNumber="460">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C36A6506E85417677EADF8D26898A3A3" lastPageId="11" lastPageNumber="462" pageId="9" pageNumber="460" type="discussion">
<paragraph id="8BCF368DE85417677EADF8D26898A3A3" blockId="9.[151,1437,1455,2023]" lastBlockId="11.[151,1437,151,393]" lastPageId="11" lastPageNumber="462" pageId="9" pageNumber="460">
<emphasis id="B904EA9FE85417657EADF8D26A42A57C" bold="true" box="[199,315,1852,1878]" pageId="9" pageNumber="460">Remarks.</emphasis>
<taxonomicName id="4C704D0EE85417657F28F8D269D0A57C" authority="Prell, 1913" authorityName="Prell" authorityYear="1913" box="[322,681,1852,1878]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="felschei">
<emphasis id="B904EA9FE85417657F28F8D26951A57C" box="[322,552,1852,1878]" italics="true" pageId="9" pageNumber="460">Eophileurus felschei</emphasis>
<bibRefCitation id="EFE14B7CE85417657C44F8D269D0A57C" author="Prell, H." box="[558,681,1852,1878]" pageId="9" pageNumber="460" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell, 1913</bibRefCitation>
</taxonomicName>
was described based on one pair of specimens from “Genji,
<collectingCountry id="F367761DE85417657B25F8D26EF7A57C" box="[1359,1422,1852,1878]" name="India" pageId="9" pageNumber="460">India</collectingCountry>
”. The
<typeStatus id="54CB882FE85417657EA6F88E6A4AA550" box="[204,307,1888,1914]" pageId="9" pageNumber="460" type="lectotype">lectotype</typeStatus>
(
<figureCitation id="134B2A08E85417657F29F88E6AB5A550" box="[323,460,1888,1914]" captionStart="FIGURES 38–48" captionStartId="10.[151,264,1944,1968]" captionTargetBox="[253,1339,186,1912]" captionTargetId="figure-18@10.[241,1346,181,1920]" captionTargetPageId="10" captionText="FIGURES 38–48. Type specimens of Eophileurus felschei Prell, 1913. 38–31, lectotype, female, 38, habitus, dorsal; 39, habitus, lateral; 40, habitus, ventral; 41, labels; 42–48, lectoparatype, male (images ©MTD/Olaf Jäger), 42, habitus, dorsal; 43, habitus, lateral; 44, habitus, ventral, 45–46, parameres, apical; 47, parameres, lateral; 48, labels." figureDoi="http://doi.org/10.5281/zenodo.6853956" httpUri="https://zenodo.org/record/6853956/files/figure.png" pageId="9" pageNumber="460">Figs. 38–41</figureCitation>
) was designated by
<bibRefCitation id="EFE14B7CE85417657CDCF88E681FA550" author="Endrodi, S." box="[694,870,1888,1914]" pageId="9" pageNumber="460" pagination="93 - 134" refId="ref16934" refString="Endrodi, S. (1977) Monographie de Dynastinae VIII Tribus Phileurini. Entomologische Abhandlungen Staatlichen Museum fur Tierkunde Dresden, 41 (4), 93 - 134. https: // doi. org / 10.1515 / 9783112546161" type="journal article" year="1977">Endrődi (1977)</bibRefCitation>
, which is the female specimen housed in MFNB. The
<typeStatus id="54CB882FE85417657EA3F86A6A19A5B4" box="[201,352,1924,1950]" pageId="9" pageNumber="460" type="paralectotype">paralectotype</typeStatus>
(
<figureCitation id="134B2A08E85417657F04F86A6A8AA5B4" box="[366,499,1924,1950]" captionStart="FIGURES 38–48" captionStartId="10.[151,264,1944,1968]" captionTargetBox="[253,1339,186,1912]" captionTargetId="figure-18@10.[241,1346,181,1920]" captionTargetPageId="10" captionText="FIGURES 38–48. Type specimens of Eophileurus felschei Prell, 1913. 38–31, lectotype, female, 38, habitus, dorsal; 39, habitus, lateral; 40, habitus, ventral; 41, labels; 42–48, lectoparatype, male (images ©MTD/Olaf Jäger), 42, habitus, dorsal; 43, habitus, lateral; 44, habitus, ventral, 45–46, parameres, apical; 47, parameres, lateral; 48, labels." figureDoi="http://doi.org/10.5281/zenodo.6853956" httpUri="https://zenodo.org/record/6853956/files/figure.png" pageId="9" pageNumber="460">Figs. 42–48</figureCitation>
) of this species is the male specimen housed in MTD. In recent years,
<bibRefCitation id="EFE14B7CE85417657A91F86A6EE5A5B4" author="Dupuis, F." box="[1275,1436,1924,1950]" pageId="9" pageNumber="460" pagination="13 - 18" refId="ref16872" refString="Dupuis, F. (2014) Deux nouvelles especes d' Eophileurus Arrow, 1908. Coleopteres, 20 (3), 13 - 18." type="journal article" year="2014">Dupuis (2014)</bibRefCitation>
described
<taxonomicName id="4C704D0EE85417657F60F8476904A5E8" authority="Dupuis, 2014" authorityName="Dupuis" authorityYear="2014" box="[266,637,1960,1986]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="grossepunctatus">
<emphasis id="B904EA9FE85417657F60F8476AA7A5E8" box="[266,478,1961,1986]" italics="true" pageId="9" pageNumber="460">E. grossepunctatus</emphasis>
<bibRefCitation id="EFE14B7CE85417657F8FF8466904A5E8" author="Dupuis, F." box="[485,637,1960,1986]" pageId="9" pageNumber="460" pagination="13 - 18" refId="ref16872" refString="Dupuis, F. (2014) Deux nouvelles especes d' Eophileurus Arrow, 1908. Coleopteres, 20 (3), 13 - 18." type="journal article" year="2014">Dupuis, 2014</bibRefCitation>
</taxonomicName>
from “Shimoga,
<collectingCountry id="F367761DE85417657D2AF8466805A5E8" box="[832,892,1960,1986]" name="India" pageId="9" pageNumber="460">India</collectingCountry>
” (
<typeStatus id="54CB882FE85417657DFFF8466F7AA5E8" box="[917,1027,1960,1986]" pageId="9" pageNumber="460" type="holotype">Holotype</typeStatus>
:
<figureCitation id="134B2A08E85417657A64F8466FEAA5E9" box="[1038,1171,1960,1987]" captionStart="FIGURES 49–53" captionStartId="11.[151,264,1333,1357]" captionTargetBox="[156,1433,418,1309]" captionTargetId="figure-149@11.[151,1435,417,1309]" captionTargetPageId="11" captionText="FIGURES 49–53. Holotype of Eophileurus grossepunctatus Dupuis, 2014 (images ©MNHN/A. Mantilleri). 49, habitus, dorsal; 50, habitus, lateral; 51, parameres, apical; 52, parameres, lateral; 53, labels." figureDoi="http://doi.org/10.5281/zenodo.6853958" httpUri="https://zenodo.org/record/6853958/files/figure.png" pageId="9" pageNumber="460">Figs. 49–53</figureCitation>
) without providing any comparison to
<taxonomicName id="4C704D0EE85417657F51F8236AD6A5CC" authorityName="Prell" authorityYear="1913" box="[315,431,1996,2022]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="felschei">
<emphasis id="B904EA9FE85417657F51F8236AD6A5CC" box="[315,431,1996,2022]" italics="true" pageId="9" pageNumber="460">E. felschei</emphasis>
</taxonomicName>
. After the examination of the type material of both taxa, it is clear that no major differences are present in both the appearance and the parameres other than slight differences on the lateral processes of the parameres of the
<specimenCount id="9D76FD04E85617677F3DFF556AC4A2FF" box="[343,445,187,213]" pageId="11" pageNumber="462" type="male">two male</specimenCount>
types (the lateral processes are slightly more enlarged and expanded at their apices in the type of
<taxonomicName id="4C704D0EE85617677E84FF316A1AA2D3" authorityName="Prell" authorityYear="1913" box="[238,355,223,249]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="felschei">
<emphasis id="B904EA9FE85617677E84FF316A1AA2D3" box="[238,355,223,249]" italics="true" pageId="11" pageNumber="462">E. felschei</emphasis>
</taxonomicName>
and less so in
<emphasis id="B904EA9FE85617677C63FF31699CA2D2" box="[521,741,223,248]" italics="true" pageId="11" pageNumber="462">
<taxonomicName id="4C704D0EE85617677C63FF316998A2D2" authorityName="Dupuis" authorityYear="2014" box="[521,737,223,248]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="grossepunctatus">E. grossepunctatus</taxonomicName>
.
</emphasis>
This mild difference can be seen in
<figureCitation id="134B2A08E85617677A14FF316FB7A2D3" box="[1150,1230,223,249]" captionStart="FIGURES 38–48" captionStartId="10.[151,264,1944,1968]" captionTargetBox="[253,1339,186,1912]" captionTargetId="figure-18@10.[241,1346,181,1920]" captionTargetPageId="10" captionText="FIGURES 38–48. Type specimens of Eophileurus felschei Prell, 1913. 38–31, lectotype, female, 38, habitus, dorsal; 39, habitus, lateral; 40, habitus, ventral; 41, labels; 42–48, lectoparatype, male (images ©MTD/Olaf Jäger), 42, habitus, dorsal; 43, habitus, lateral; 44, habitus, ventral, 45–46, parameres, apical; 47, parameres, lateral; 48, labels." figureDoi="http://doi.org/10.5281/zenodo.6853956" httpUri="https://zenodo.org/record/6853956/files/figure.png" pageId="11" pageNumber="462">Fig. 46</figureCitation>
versus
<figureCitation id="134B2A08E85617677B4EFF316E0DA2D3" box="[1316,1396,222,249]" captionStart="FIGURES 49–53" captionStartId="11.[151,264,1333,1357]" captionTargetBox="[156,1433,418,1309]" captionTargetId="figure-149@11.[151,1435,417,1309]" captionTargetPageId="11" captionText="FIGURES 49–53. Holotype of Eophileurus grossepunctatus Dupuis, 2014 (images ©MNHN/A. Mantilleri). 49, habitus, dorsal; 50, habitus, lateral; 51, parameres, apical; 52, parameres, lateral; 53, labels." figureDoi="http://doi.org/10.5281/zenodo.6853958" httpUri="https://zenodo.org/record/6853958/files/figure.png" pageId="11" pageNumber="462">Fig. 52</figureCitation>
for the front view, and
<figureCitation id="134B2A08E85617677F04FEED6AC5A337" box="[366,444,259,285]" captionStart="FIGURES 38–48" captionStartId="10.[151,264,1944,1968]" captionTargetBox="[253,1339,186,1912]" captionTargetId="figure-18@10.[241,1346,181,1920]" captionTargetPageId="10" captionText="FIGURES 38–48. Type specimens of Eophileurus felschei Prell, 1913. 38–31, lectotype, female, 38, habitus, dorsal; 39, habitus, lateral; 40, habitus, ventral; 41, labels; 42–48, lectoparatype, male (images ©MTD/Olaf Jäger), 42, habitus, dorsal; 43, habitus, lateral; 44, habitus, ventral, 45–46, parameres, apical; 47, parameres, lateral; 48, labels." figureDoi="http://doi.org/10.5281/zenodo.6853956" httpUri="https://zenodo.org/record/6853956/files/figure.png" pageId="11" pageNumber="462">Fig. 47</figureCitation>
versus
<figureCitation id="134B2A08E85617677C7BFEED6926A337" box="[529,607,258,285]" captionStart="FIGURES 49–53" captionStartId="11.[151,264,1333,1357]" captionTargetBox="[156,1433,418,1309]" captionTargetId="figure-149@11.[151,1435,417,1309]" captionTargetPageId="11" captionText="FIGURES 49–53. Holotype of Eophileurus grossepunctatus Dupuis, 2014 (images ©MNHN/A. Mantilleri). 49, habitus, dorsal; 50, habitus, lateral; 51, parameres, apical; 52, parameres, lateral; 53, labels." figureDoi="http://doi.org/10.5281/zenodo.6853958" httpUri="https://zenodo.org/record/6853958/files/figure.png" pageId="11" pageNumber="462">Fig. 51</figureCitation>
for the lateral view). Following the conclusion of the discussion section at the end of this paper, it is observed that the lateral processes bare the largest individual and geographical variations, and with differences this mild, the differences are too insignificant to separate them from a single species. Hence,
<taxonomicName id="4C704D0EE85617677BEEFEA56A35A3A2" authorityName="Dupuis" authorityYear="2014" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="grossepunctatus">
<emphasis id="B904EA9FE85617677BEEFEA56A35A3A2" italics="true" pageId="11" pageNumber="462">E. grossepunctatus</emphasis>
</taxonomicName>
is conspecific with
<taxonomicName id="4C704D0EE85617677C46FE8169D8A3A3" authorityName="Prell" authorityYear="1913" box="[556,673,367,393]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="felschei">
<emphasis id="B904EA9FE85617677C46FE8169D8A3A3" box="[556,673,367,393]" italics="true" pageId="11" pageNumber="462">E. felschei</emphasis>
</taxonomicName>
and thus synonymized here.
</paragraph>
</subSubSection>
</treatment>
</document>
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