treatments-xml/data/03/D9/87/03D9879BE84E17797EFDF94E68B5A77F.xml
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<document id="BB044438F4C8CC58EB12B65E1EDFA66D" ID-CLB-Dataset="99517" ID-DOI="10.11646/zootaxa.5165.4.1" ID-GBIF-Dataset="cd84b880-d77f-401a-bf4c-a98021a5e166" ID-ISSN="1175-5326" ID-Zenodo-Dep="6853942" ID-ZooBank="DBE812FF-7B25-4522-97FE-45A86ECAEFD7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658127043518" checkinUser="plazi" docAuthor="Yang, Qiao-Zhi &amp; Pathomwattananurak, Wuttipon" docDate="2022" docId="03D9879BE84E17797EFDF94E68B5A77F" docLanguage="en" docName="zootaxa.5165.4.1.pdf" docOrigin="Zootaxa 5165 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eophileurus prelli Yang &amp; Pathomwattananurak 2022, new species" docType="treatment" docVersion="6" lastPageNumber="472" masterDocId="FFE0FFE3E85D176C7E6AFFEE6B79A22A" masterDocTitle="Four new species of the genus Eophileurus Arrow, 1908 (Coleoptera: Scarabaeidae Dynastinae) from Thailand and Vietnam, with notes on some Indochinese species" masterLastPageNumber="485" masterPageNumber="451" pageNumber="470" updateTime="1699360166553" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="1477FC354D15218682E339D0D335DFA3">Four new species of the genus Eophileurus Arrow, 1908 (Coleoptera: Scarabaeidae Dynastinae) from Thailand and Vietnam, with notes on some Indochinese species</mods:title>
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<mods:namePart id="AA294AA8D46EB7444243F5C1DF9355F2">Yang, Qiao-Zhi</mods:namePart>
<mods:affiliation id="534966973631E8C432ACF9BFA6EB42E8">3 B, Pingguoshequ, Baiziwan Road, Chaoyang Dist., Beijing, 100022, P. R. China.</mods:affiliation>
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<mods:namePart id="8569E1131D12EA75C464952480C7BFC0">Pathomwattananurak, Wuttipon</mods:namePart>
<mods:affiliation id="55A7E8213F7403C08551BED3DE5FF1C7">1051 / 38 Wiang, Mueang Dist., Chiang Rai Province, 57000, Thailand.</mods:affiliation>
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Yang &amp; Pathomwattananurak
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,
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,
<figureCitation id="134B2A08E84E177F7F59F92B6AEDA4F5" box="[307,404,1733,1759]" captionStart="FIGURES109116" captionStartId="26.[151,264,1724,1748]" captionTargetBox="[158,1431,400,1698]" captionTargetId="figure-122@26.[151,1435,381,1701]" captionTargetPageId="26" captionText="FIGURES109116.HabitusofEophileurusArrow,1908species(obliqueview).109110,E.minorYang&amp;Pathomwattananurak, new species, 109, holotype, 110, female paratype; 111112, E. prelli Yang &amp; Pathomwattananurak, new species, 111, male paratype; 112, female paratype; 113114, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 113, holotype, 114, paratype (Mu Si), 115116, E. sidereus Yang &amp; Pathomwattananurak, new species, 115, holotype, 116, female paratype." figureDoi="http://doi.org/10.5281/zenodo.6853978" httpUri="https://zenodo.org/record/6853978/files/figure.png" pageId="19" pageNumber="470">111112</figureCitation>
,
<figureCitation id="134B2A08E84E177F7FCAF92B697BA4F5" box="[416,514,1733,1759]" captionStart="FIGURES 165184" captionStartId="30.[151,264,1476,1500]" captionTargetBox="[157,1431,189,1441]" captionTargetId="figure-18@30.[151,1435,181,1453]" captionTargetPageId="30" captionText="FIGURES 165184. Parameres of Eophileurus Arrow, 1908 species. 165166, E. confinis Prell, 1913 (Bao Loc); 167168, E. heyrovskyi Kral &amp; Strnad, 1992 (Longli); 169172, E. minor Yang &amp; Pathomwattananurak, new species, 169170, holotype, 171172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173174, E. prelli Yang &amp; Pathomwattananurak, new species, holotype; 174180, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 175176, holotype, 177178, paratype (Mu Si), 179180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181184, E. sidereus Yang &amp; Pathomwattananurak, new species, 181182, holotype, 183184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853988" httpUri="https://zenodo.org/record/6853988/files/figure.png" pageId="19" pageNumber="470">173174</figureCitation>
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, local collector leg.”
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, local collector leg.”
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,
<date id="FFCE104DE84E177F7ED1F8736A5DA59D" box="[187,292,1949,1975]" pageId="19" pageNumber="470" value="2019-07">
<collectingDate id="EF8AE9A5E84E177F7ED1F8736A5DA59D" box="[187,292,1949,1975]" pageId="19" pageNumber="470" value="2019-07">VII.2019</collectingDate>
</date>
, local collector leg.”
</materialsCitation>
;
<materialsCitation id="3B183CD0E84E177F7C74F8736825A5F0" ID-GBIF-Occurrence="3863916309" collectingDate="2020-06" collectionCode="CYQZ, VI" country="Vietnam" elevation="1300" latitude="15.964556" location="Mt. Axan" longLatPrecision="1" longitude="107.43906" municipality="Tay Giang Dist." pageId="19" pageNumber="470" specimenCount="1" specimenCount-female="1" stateProvince="Quang Nam" typeStatus="paratype">
<specimenCount id="9D76FD04E84E177F7C74F8736939A59D" box="[542,576,1949,1975]" pageId="19" pageNumber="470" type="female">1♀</specimenCount>
(
<collectionCode id="ED61AE48E84E177F7C3AF87369D9A59D" box="[592,672,1949,1975]" pageId="19" pageNumber="470">CYQZ</collectionCode>
) “
<geoCoordinate id="EE44504AE84E177F7CD0F872682EA59D" box="[698,855,1948,1975]" degrees="15" direction="north" minutes="57" orientation="latitude" pageId="19" pageNumber="470" precision="1" seconds="52.4" value="15.964556">15°57'52.4&quot;N</geoCoordinate>
,
<geoCoordinate id="EE44504AE84E177F7D09F8726F73A59C" box="[867,1034,1948,1974]" degrees="107" direction="east" minutes="26" orientation="longitude" pageId="19" pageNumber="470" precision="1" seconds="20.6" value="107.43906">107°26'20.6&quot;E</geoCoordinate>
,
<location id="8EAF6056E84E177F7A7CF8736FFAA59D" LSID="urn:lsid:plazi:treatment:03D9879BE84E17797EFDF94E68B5A77F:8EAF6056E84E177F7A7CF8736FFAA59D" box="[1046,1155,1949,1975]" country="Vietnam" latitude="15.964556" longLatPrecision="1" longitude="107.43906" municipality="Tay Giang Dist." name="Mt. Axan" pageId="19" pageNumber="470" stateProvince="Quang Nam">Mt. Axan</location>
,
<collectingMunicipality id="6BABACF7E84E177F7AE5F8736E3FA59C" box="[1167,1350,1948,1975]" pageId="19" pageNumber="470">Tay Giang Dist.</collectingMunicipality>
,
<collectingRegion id="49B4F86FE84E177F7B38F8736A6BA5F1" country="Vietnam" name="Quang Nam" pageId="19" pageNumber="470">Quang Nam Prov.</collectingRegion>
,
<collectingCountry id="F367761DE84E177F7F76F82E6A07A5F0" box="[284,382,1984,2010]" name="Vietnam" pageId="19" pageNumber="470">Vietnam</collectingCountry>
, alt.
<quantity id="4C889B68E84E177F7FD9F82F6973A5F1" box="[435,522,1985,2011]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="19" pageNumber="470" unit="m" value="1300.0">
<elevation id="005DD1BEE84E177F7FD9F82F6973A5F1" box="[435,522,1985,2011]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="19" pageNumber="470" unit="m" value="1300.0">1300 m</elevation>
</quantity>
,
<date id="FFCE104DE84E177F7C7FF82F690CA5F1" box="[533,629,1985,2011]" pageId="19" pageNumber="470" value="2020-06">
<collectingDate id="EF8AE9A5E84E177F7C7FF82F690CA5F1" box="[533,629,1985,2011]" pageId="19" pageNumber="470" value="2020-06">
<collectionCode id="ED61AE48E84E177F7C7FF82F694DA5F1" box="[533,564,1985,2011]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="19" pageNumber="470">VI</collectionCode>
.2020
</collectingDate>
</date>
, local collector leg.”
</materialsCitation>
</paragraph>
<caption id="DF0F6605E84917787EFDFA7068F9A7F0" ID-DOI="http://doi.org/10.5281/zenodo.6853972" ID-Zenodo-Dep="6853972" httpUri="https://zenodo.org/record/6853972/files/figure.png" pageId="20" pageNumber="471" startId="20.[151,264,1438,1462]" targetBox="[159,1425,193,1408]" targetPageId="20">
<paragraph id="8BCF368DE84917787EFDFA7068F9A7F0" blockId="20.[151,1437,1438,1499]" pageId="20" pageNumber="471">
<emphasis id="B904EA9FE84917787EFDFA706A2BA79C" bold="true" box="[151,338,1438,1462]" pageId="20" pageNumber="471">FIGURES 9398.</emphasis>
Habitus of
<taxonomicName id="4C704D0EE84917787FAEFA7068D1A79C" authority="Yang &amp; Pathomwattananurak" authorityName="Yang &amp; Pathomwattananurak" authorityYear="2022" box="[452,936,1438,1463]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="prelli" status="sp. nov.">
<emphasis id="B904EA9FE84917787FAEFA706903A79C" box="[452,634,1438,1462]" italics="true" pageId="20" pageNumber="471">Eophileurus prelli</emphasis>
Yang &amp; Pathomwattananurak
</taxonomicName>
,
<emphasis id="B904EA9FE84917787DDAFA706F49A79C" bold="true" box="[944,1072,1438,1462]" pageId="20" pageNumber="471">
<taxonomicNameLabel id="A23757E4E84917787DDAFA706F55A79C" box="[944,1068,1438,1462]" pageId="20" pageNumber="471" rank="species">new species</taxonomicNameLabel>
.
</emphasis>
9394, holotype; 95, male paratype; 9698, female paratypes; 93, 95, 96, 98 dorsal view; 94, 97, ventral view.
</paragraph>
</caption>
<paragraph id="8BCF368DE84917797EADF9E76F4DA3DF" blockId="20.[151,1437,1544,2039]" lastBlockId="21.[151,1437,150,1365]" lastPageId="21" lastPageNumber="472" pageId="20" pageNumber="471">
<emphasis id="B904EA9FE84917787EADF9E76A4DA409" bold="true" box="[199,308,1545,1571]" pageId="20" pageNumber="471">
<typeStatus id="54CB882FE84917787EADF9E76A4DA409" box="[199,308,1545,1571]" pageId="20" pageNumber="471" type="holotype">Holotype</typeStatus>
</emphasis>
(male).
<emphasis id="B904EA9FE84917787FFEF9E66A8DA408" bold="true" box="[404,500,1544,1570]" pageId="20" pageNumber="471">General</emphasis>
(
<figureCitation id="134B2A08E84917787C6FF9E669F4A409" box="[517,653,1544,1571]" captionStart="FIGURES 9398" captionStartId="20.[151,264,1438,1462]" captionTargetBox="[159,1425,193,1408]" captionTargetId="figure-18@20.[151,1435,181,1414]" captionTargetPageId="20" captionText="FIGURES 9398. Habitus of Eophileurus prelli Yang &amp; Pathomwattananurak, new species. 9394, holotype; 95, male paratype; 9698, female paratypes; 93, 95, 96, 98 dorsal view; 94, 97, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6853972" httpUri="https://zenodo.org/record/6853972/files/figure.png" pageId="20" pageNumber="471">Figs. 9394</figureCitation>
): Body black, rather oblong, flattened, surface slightly shiny; ventral surface clads with dense, long reddish setae.
<emphasis id="B904EA9FE84917787CEFF9C369B7A46D" bold="true" box="[645,718,1581,1607]" pageId="20" pageNumber="471">Head:</emphasis>
Clypeus sparsely punctate, punctures large, with a short, slightly backward horn at middle and a distinct ridge from horn to apex; clypeal apex slightly acute. Frons moderately depressed, disk with sparse micropunctures. Mandible sinuate at outer margin.
<emphasis id="B904EA9FE84917787A6DF99B6FF0A4A5" bold="true" box="[1031,1161,1653,1679]" pageId="20" pageNumber="471">Pronotum:</emphasis>
Surface with dense, fine punctures, becoming much denser and larger near all margins. Anterior region with a large, rather deep, round fovea; punctures in fovea very dense and large, round or in sub-U-shape, sometimes coalescent mostly at anterior region; with two very weak protuberances at each side. All margins beaded except before fovea, more strongly beaded at sides. Anterior angle acute, posterior angle rounded.
<emphasis id="B904EA9FE84917787C84F8EA6809A534" bold="true" box="[750,880,1796,1822]" pageId="20" pageNumber="471">Scutellum:</emphasis>
Surface impunctate.
<emphasis id="B904EA9FE84917787A0BF8EB6FCFA535" bold="true" box="[1121,1206,1797,1823]" pageId="20" pageNumber="471">Elytra:</emphasis>
Surface with sparse micropunctures, and dense and large, round punctures. Interstice 1 with punctures in two irregular rows combined at middle to one irregular row, interstice 2 with punctures in one irregular row. Primary costa A distinct, primary costa B indistinct. Subapical umbones moderately prominent. Margin of elytron slightly curved outward, dilated horizontally from base to about posterior three-seventh.
<emphasis id="B904EA9FE84917787D42F87A68DCA584" bold="true" box="[808,933,1940,1966]" pageId="20" pageNumber="471">Pygidium:</emphasis>
convex, almost impunctate, except regions near all sides sparsely punctate.
<emphasis id="B904EA9FE84917787C6FF85769C9A5F9" bold="true" box="[517,688,1977,2003]" pageId="20" pageNumber="471">Metasternum:</emphasis>
Surface mainly black, with slight reddish color at medial portion; with a lance-shaped depression at center; region near all sides with dense and small punctures; central portion with sparse and small punctures; setae reddish, long, mainly at anterior region.
<emphasis id="B904EA9FE84817797D9FFF786F8FA29A" bold="true" box="[1013,1270,150,176]" pageId="21" pageNumber="472">Abdominal ventrites:</emphasis>
Surfaces with sparse micropunctures throughout and larger punctures at each side; ventrite 6 with a shallow, horizontally oval depression at center.
<emphasis id="B904EA9FE84817797FE2FF316AB0A2D3" bold="true" box="[392,457,223,249]" pageId="21" pageNumber="472">Legs:</emphasis>
Protibia tridentate, protarsi strongly thickened, inner protibial claw strongly enlarged and moderately elongated. Inner metatibial spur long, moderately curved outward at middle; outer metatibial spur shorter, strongly curved outwards.
<emphasis id="B904EA9FE84817797C5BFEC969CBA36B" bold="true" box="[561,690,295,321]" pageId="21" pageNumber="472">Parameres</emphasis>
(
<figureCitation id="134B2A08E84817797CACFEC96814A36B" box="[710,877,295,321]" captionStart="FIGURES 165184" captionStartId="30.[151,264,1476,1500]" captionTargetBox="[157,1431,189,1441]" captionTargetId="figure-18@30.[151,1435,181,1453]" captionTargetPageId="30" captionText="FIGURES 165184. Parameres of Eophileurus Arrow, 1908 species. 165166, E. confinis Prell, 1913 (Bao Loc); 167168, E. heyrovskyi Kral &amp; Strnad, 1992 (Longli); 169172, E. minor Yang &amp; Pathomwattananurak, new species, 169170, holotype, 171172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173174, E. prelli Yang &amp; Pathomwattananurak, new species, holotype; 174180, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 175176, holotype, 177178, paratype (Mu Si), 179180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181184, E. sidereus Yang &amp; Pathomwattananurak, new species, 181182, holotype, 183184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853988" httpUri="https://zenodo.org/record/6853988/files/figure.png" pageId="21" pageNumber="472">Figs. 173174</figureCitation>
): Slender and symmetric; in frontal view (
<figureCitation id="134B2A08E84817797B05FEC96BBDA34F" captionStart="FIGURES 165184" captionStartId="30.[151,264,1476,1500]" captionTargetBox="[157,1431,189,1441]" captionTargetId="figure-18@30.[151,1435,181,1453]" captionTargetPageId="30" captionText="FIGURES 165184. Parameres of Eophileurus Arrow, 1908 species. 165166, E. confinis Prell, 1913 (Bao Loc); 167168, E. heyrovskyi Kral &amp; Strnad, 1992 (Longli); 169172, E. minor Yang &amp; Pathomwattananurak, new species, 169170, holotype, 171172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173174, E. prelli Yang &amp; Pathomwattananurak, new species, holotype; 174180, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 175176, holotype, 177178, paratype (Mu Si), 179180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181184, E. sidereus Yang &amp; Pathomwattananurak, new species, 181182, holotype, 183184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853988" httpUri="https://zenodo.org/record/6853988/files/figure.png" pageId="21" pageNumber="472">Fig. 173</figureCitation>
), outer sides prominent at anterior one-third; inner margins sinuate, slightly curved outward between middle and apex; with a pair of processes emerging from inner margins near apex, apices curved forward, and a pair of processes at middle of outer margins, curved inward at apices; apex enlarged, outer margins extending outward; a pair of triangular processes from lower base of parameres visible; in lateral view (
<figureCitation id="134B2A08E84817797A5EFE596FEFA3FB" box="[1076,1174,439,465]" captionStart="FIGURES 165184" captionStartId="30.[151,264,1476,1500]" captionTargetBox="[157,1431,189,1441]" captionTargetId="figure-18@30.[151,1435,181,1453]" captionTargetPageId="30" captionText="FIGURES 165184. Parameres of Eophileurus Arrow, 1908 species. 165166, E. confinis Prell, 1913 (Bao Loc); 167168, E. heyrovskyi Kral &amp; Strnad, 1992 (Longli); 169172, E. minor Yang &amp; Pathomwattananurak, new species, 169170, holotype, 171172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173174, E. prelli Yang &amp; Pathomwattananurak, new species, holotype; 174180, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 175176, holotype, 177178, paratype (Mu Si), 179180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181184, E. sidereus Yang &amp; Pathomwattananurak, new species, 181182, holotype, 183184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853988" httpUri="https://zenodo.org/record/6853988/files/figure.png" pageId="21" pageNumber="472">Fig. 174</figureCitation>
), upper margin slightly depressed near base; outer processes bent backward; apex strongly bent backward.
</paragraph>
<paragraph id="8BCF368DE84817797EADFE1168D9A04B" blockId="21.[151,1437,150,1365]" pageId="21" pageNumber="472">
<emphasis id="B904EA9FE84817797EADFE116A4DA033" bold="true" box="[199,308,511,537]" pageId="21" pageNumber="472">
<typeStatus id="54CB882FE84817797EADFE116A4DA033" box="[199,308,511,537]" pageId="21" pageNumber="472" type="paratype">Paratype</typeStatus>
</emphasis>
(male). Slightly underdeveloped than the
<typeStatus id="54CB882FE84817797D42FE1168F7A033" box="[808,910,511,537]" pageId="21" pageNumber="472" type="holotype">holotype</typeStatus>
; characters stable, with clypeal horn slightly shorter; fovea slightly smaller and shallower, in longitudinal oval shape, widest at anterior one-third; scutellum sparsely punctate, with only a few punctures near base (
<figureCitation id="134B2A08E84817797D63FDA96822A04B" box="[777,859,582,609]" captionStart="FIGURES 9398" captionStartId="20.[151,264,1438,1462]" captionTargetBox="[159,1425,193,1408]" captionTargetId="figure-18@20.[151,1435,181,1414]" captionTargetPageId="20" captionText="FIGURES 9398. Habitus of Eophileurus prelli Yang &amp; Pathomwattananurak, new species. 9394, holotype; 95, male paratype; 9698, female paratypes; 93, 95, 96, 98 dorsal view; 94, 97, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6853972" httpUri="https://zenodo.org/record/6853972/files/figure.png" pageId="21" pageNumber="472">Fig. 95</figureCitation>
,
<figureCitation id="134B2A08E84817797D0DFDA968EAA04B" box="[871,915,583,609]" captionStart="FIGURES109116" captionStartId="26.[151,264,1724,1748]" captionTargetBox="[158,1431,400,1698]" captionTargetId="figure-122@26.[151,1435,381,1701]" captionTargetPageId="26" captionText="FIGURES109116.HabitusofEophileurusArrow,1908species(obliqueview).109110,E.minorYang&amp;Pathomwattananurak, new species, 109, holotype, 110, female paratype; 111112, E. prelli Yang &amp; Pathomwattananurak, new species, 111, male paratype; 112, female paratype; 113114, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 113, holotype, 114, paratype (Mu Si), 115116, E. sidereus Yang &amp; Pathomwattananurak, new species, 115, holotype, 116, female paratype." figureDoi="http://doi.org/10.5281/zenodo.6853978" httpUri="https://zenodo.org/record/6853978/files/figure.png" pageId="21" pageNumber="472">111</figureCitation>
).
</paragraph>
<paragraph id="8BCF368DE84817797EADFD856F6CA13F" blockId="21.[151,1437,150,1365]" pageId="21" pageNumber="472">
<emphasis id="B904EA9FE84817797EADFD856A46A0AF" bold="true" box="[199,319,619,645]" pageId="21" pageNumber="472">
<typeStatus id="54CB882FE84817797EADFD856A46A0AF" box="[199,319,619,645]" pageId="21" pageNumber="472" type="paratype">Paratypes</typeStatus>
</emphasis>
(female). Similar to male, except clypeus and frons much more densely rugopunctate throughout; clypeal horn much shorter; fovea absent, with a very shallow and densely punctate furrow on anterior region of pronotum; scutellum sparsely punctate near base; pygidium flattened at middle; depression on metasternum shallower and slightly narrower; ventrite 6 without depression, with dense rugopunctures near anterior margin; protarsi not thickened, inner claw not enlarged and elongated (
<figureCitation id="134B2A08E84817797D39FD1568A9A13F" box="[851,976,763,789]" captionStart="FIGURES 9398" captionStartId="20.[151,264,1438,1462]" captionTargetBox="[159,1425,193,1408]" captionTargetId="figure-18@20.[151,1435,181,1414]" captionTargetPageId="20" captionText="FIGURES 9398. Habitus of Eophileurus prelli Yang &amp; Pathomwattananurak, new species. 9394, holotype; 95, male paratype; 9698, female paratypes; 93, 95, 96, 98 dorsal view; 94, 97, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6853972" httpUri="https://zenodo.org/record/6853972/files/figure.png" pageId="21" pageNumber="472">Fig. 9698</figureCitation>
,
<figureCitation id="134B2A08E84817797DB1FD156F71A13F" box="[987,1032,763,789]" captionStart="FIGURES109116" captionStartId="26.[151,264,1724,1748]" captionTargetBox="[158,1431,400,1698]" captionTargetId="figure-122@26.[151,1435,381,1701]" captionTargetPageId="26" captionText="FIGURES109116.HabitusofEophileurusArrow,1908species(obliqueview).109110,E.minorYang&amp;Pathomwattananurak, new species, 109, holotype, 110, female paratype; 111112, E. prelli Yang &amp; Pathomwattananurak, new species, 111, male paratype; 112, female paratype; 113114, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 113, holotype, 114, paratype (Mu Si), 115116, E. sidereus Yang &amp; Pathomwattananurak, new species, 115, holotype, 116, female paratype." figureDoi="http://doi.org/10.5281/zenodo.6853978" httpUri="https://zenodo.org/record/6853978/files/figure.png" pageId="21" pageNumber="472">112</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36A6506E84817797EADFCF1681FA177" pageId="21" pageNumber="472" type="description">
<paragraph id="8BCF368DE84817797EADFCF1681FA177" blockId="21.[151,1437,150,1365]" pageId="21" pageNumber="472">
<emphasis id="B904EA9FE84817797EADFCF16A07A113" bold="true" box="[199,382,799,825]" pageId="21" pageNumber="472">Measurements.</emphasis>
Body length: male
<quantity id="4C889B68E84817797C0EFCF06873A113" box="[612,778,798,825]" metricMagnitude="-2" metricUnit="m" metricValue="2.32" metricValueMax="2.35" metricValueMin="2.29" pageId="21" pageNumber="472" unit="mm" value="23.2" valueMax="23.5" valueMin="22.9">22.923.5 mm</quantity>
(
<typeStatus id="54CB882FE84817797D70FCF16807A113" box="[794,894,799,825]" pageId="21" pageNumber="472" type="holotype">holotype</typeStatus>
<quantity id="4C889B68E84817797DEDFCF1688BA113" box="[903,1010,799,825]" metricMagnitude="-2" metricUnit="m" metricValue="2.29" pageId="21" pageNumber="472" unit="mm" value="22.9">22.9 mm</quantity>
), female
<quantity id="4C889B68E84817797A37FCF06E7CA113" box="[1117,1285,798,825]" metricMagnitude="-2" metricUnit="m" metricValue="2.445" metricValueMax="2.54" metricValueMin="2.35" pageId="21" pageNumber="472" unit="mm" value="24.45" valueMax="25.4" valueMin="23.5">23.525.4 mm</quantity>
; body width: male
<quantity id="4C889B68E84817797EBFFCAC6A0FA177" box="[213,374,834,861]" metricMagnitude="-2" metricUnit="m" metricValue="1.135" metricValueMax="1.15" metricValueMin="1.1199999999999999" pageId="21" pageNumber="472" unit="mm" value="11.35" valueMax="11.5" valueMin="11.2">11.211.5 mm</quantity>
(
<typeStatus id="54CB882FE84817797FEEFCAD6A91A177" box="[388,488,835,861]" pageId="21" pageNumber="472" type="holotype">holotype</typeStatus>
<quantity id="4C889B68E84817797F84FCAD692CA177" box="[494,597,835,861]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="21" pageNumber="472" unit="mm" value="11.2">11.2 mm</quantity>
), female
<quantity id="4C889B68E84817797CD1FCAD6818A177" box="[699,865,835,861]" metricMagnitude="-2" metricUnit="m" metricValue="1.145" metricValueMax="1.26" metricValueMin="1.03" pageId="21" pageNumber="472" unit="mm" value="11.45" valueMax="12.6" valueMin="10.3">10.312.6 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C36A6506E84817797EADFC886952A61F" pageId="21" pageNumber="472" type="diagnosis">
<paragraph id="8BCF368DE84817797EADFC886952A61F" blockId="21.[151,1437,150,1365]" pageId="21" pageNumber="472">
<emphasis id="B904EA9FE84817797EADFC886A3BA1AA" bold="true" box="[199,322,870,896]" pageId="21" pageNumber="472">Diagnosis.</emphasis>
This species resembles
<taxonomicName id="4C704D0EE84817797C3AFC8968ABA1AB" authority="Kral &amp; Strnad, 1992" authorityName="Kral &amp; Strnad" authorityYear="1992" box="[592,978,871,897]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="heyrovskyi">
<emphasis id="B904EA9FE84817797C3AFC89699EA1AB" box="[592,743,871,897]" italics="true" pageId="21" pageNumber="472">E. heyrovskyi</emphasis>
<bibRefCitation id="EFE14B7CE84817797C87FC8968ABA1AB" author="Kral, D. &amp; Strnad, J." box="[749,978,871,897]" pageId="21" pageNumber="472" pagination="3 - 6" refId="ref17217" refString="Kral, D. &amp; Strnad, J. (1992) New Eophileurus species from Vietnam (Scarabaeidae, Dynastinae). Folia Heyrovskyana, 1, 3 - 6." type="journal article" year="1992">Král &amp; Strnad, 1992</bibRefCitation>
</taxonomicName>
(
<figureCitation id="134B2A08E84817797D88FC896F10A1AB" box="[994,1129,870,897]" captionStart="FIGURES 6570" captionStartId="14.[151,264,1486,1510]" captionTargetBox="[164,1420,247,1458]" captionTargetId="figure-46@14.[151,1435,237,1462]" captionTargetPageId="14" captionText="FIGURES 6570. Habitus of Eophileurus heyrovskyi Král &amp; Strnad, 1992. 6566, male (Mt. Laoshan); 67, male (Longli); 6869, female (Luoyingou); 70, male (Luoyingou); 65, 67, 68, 70, dorsal view; 66, 69, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6853964" httpUri="https://zenodo.org/record/6853964/files/figure.png" pageId="21" pageNumber="472">Figs. 6570</figureCitation>
) due to the similar external characters and the backward processes at the outer sides of the parameres. It can be distinguished from the latter by the following points: elytral punctures relatively smaller; elytral margin more weakly dilated (
<figureCitation id="134B2A08E84817797A8EFC416BCEA1C7" captionStart="FIGURES 9398" captionStartId="20.[151,264,1438,1462]" captionTargetBox="[159,1425,193,1408]" captionTargetId="figure-18@20.[151,1435,181,1414]" captionTargetPageId="20" captionText="FIGURES 9398. Habitus of Eophileurus prelli Yang &amp; Pathomwattananurak, new species. 9394, holotype; 95, male paratype; 9698, female paratypes; 93, 95, 96, 98 dorsal view; 94, 97, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6853972" httpUri="https://zenodo.org/record/6853972/files/figure.png" pageId="21" pageNumber="472">Figs. 93, 9596, 98</figureCitation>
); metasternum with a lance-shaped depression at center (
<figureCitation id="134B2A08E84817797D5AFC3D68CEA1C7" box="[816,951,979,1005]" captionStart="FIGURES 9398" captionStartId="20.[151,264,1438,1462]" captionTargetBox="[159,1425,193,1408]" captionTargetId="figure-18@20.[151,1435,181,1414]" captionTargetPageId="20" captionText="FIGURES 9398. Habitus of Eophileurus prelli Yang &amp; Pathomwattananurak, new species. 9394, holotype; 95, male paratype; 9698, female paratypes; 93, 95, 96, 98 dorsal view; 94, 97, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6853972" httpUri="https://zenodo.org/record/6853972/files/figure.png" pageId="21" pageNumber="472">Figs. 94, 97</figureCitation>
), whereas absent in
<taxonomicName id="4C704D0EE84817797AFDFC3D6E55A1C7" authorityName="Kral &amp; Strnad" authorityYear="1992" box="[1175,1324,979,1005]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="heyrovskyi">
<emphasis id="B904EA9FE84817797AFDFC3D6E55A1C7" box="[1175,1324,979,1005]" italics="true" pageId="21" pageNumber="472">E. heyrovskyi</emphasis>
</taxonomicName>
(
<figureCitation id="134B2A08E84817797B51FC3D6BCEA63B" captionStart="FIGURES 6570" captionStartId="14.[151,264,1486,1510]" captionTargetBox="[164,1420,247,1458]" captionTargetId="figure-46@14.[151,1435,237,1462]" captionTargetPageId="14" captionText="FIGURES 6570. Habitus of Eophileurus heyrovskyi Král &amp; Strnad, 1992. 6566, male (Mt. Laoshan); 67, male (Longli); 6869, female (Luoyingou); 70, male (Luoyingou); 65, 67, 68, 70, dorsal view; 66, 69, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6853964" httpUri="https://zenodo.org/record/6853964/files/figure.png" pageId="21" pageNumber="472">Figs. 66, 69</figureCitation>
); protarsi more strongly thickened (
<figureCitation id="134B2A08E84817797C29FC1969B2A63B" box="[579,715,1014,1041]" captionStart="FIGURES 9398" captionStartId="20.[151,264,1438,1462]" captionTargetBox="[159,1425,193,1408]" captionTargetId="figure-18@20.[151,1435,181,1414]" captionTargetPageId="20" captionText="FIGURES 9398. Habitus of Eophileurus prelli Yang &amp; Pathomwattananurak, new species. 9394, holotype; 95, male paratype; 9698, female paratypes; 93, 95, 96, 98 dorsal view; 94, 97, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6853972" httpUri="https://zenodo.org/record/6853972/files/figure.png" pageId="21" pageNumber="472">Figs. 9395</figureCitation>
); shapes of parameres different (
<figureCitation id="134B2A08E84817797A59FC196FAAA63B" box="[1075,1235,1015,1041]" captionStart="FIGURES 165184" captionStartId="30.[151,264,1476,1500]" captionTargetBox="[157,1431,189,1441]" captionTargetId="figure-18@30.[151,1435,181,1453]" captionTargetPageId="30" captionText="FIGURES 165184. Parameres of Eophileurus Arrow, 1908 species. 165166, E. confinis Prell, 1913 (Bao Loc); 167168, E. heyrovskyi Kral &amp; Strnad, 1992 (Longli); 169172, E. minor Yang &amp; Pathomwattananurak, new species, 169170, holotype, 171172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173174, E. prelli Yang &amp; Pathomwattananurak, new species, holotype; 174180, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 175176, holotype, 177178, paratype (Mu Si), 179180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181184, E. sidereus Yang &amp; Pathomwattananurak, new species, 181182, holotype, 183184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853988" httpUri="https://zenodo.org/record/6853988/files/figure.png" pageId="21" pageNumber="472">Figs. 167168</figureCitation>
for
<taxonomicName id="4C704D0EE84817797B6AFC196EEFA63B" authorityName="Kral &amp; Strnad" authorityYear="1992" box="[1280,1430,1015,1041]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="heyrovskyi">
<emphasis id="B904EA9FE84817797B6AFC196EEFA63B" box="[1280,1430,1015,1041]" italics="true" pageId="21" pageNumber="472">E. heyrovskyi</emphasis>
</taxonomicName>
, 173174 for
<taxonomicName id="4C704D0EE84817797F42FBF56AFAA61F" authority="Yang &amp; Pathomwattananurak, 2022" authorityName="Yang &amp; Pathomwattananurak" authorityYear="2022" box="[296,387,1051,1077]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="prelli" status="sp. nov.">
<emphasis id="B904EA9FE84817797F42FBF56AFAA61F" box="[296,387,1051,1077]" italics="true" pageId="21" pageNumber="472">E. prelli</emphasis>
</taxonomicName>
,
<emphasis id="B904EA9FE84817797FFBFBF56963A61E" bold="true" box="[401,538,1050,1077]" pageId="21" pageNumber="472">
<taxonomicNameLabel id="A23757E4E84817797FFBFBF56963A61E" box="[401,538,1050,1077]" pageId="21" pageNumber="472" rank="species">new species</taxonomicNameLabel>
</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C36A6506E84817797EADFBD1680BA657" pageId="21" pageNumber="472" type="etymology">
<paragraph id="8BCF368DE84817797EADFBD1680BA657" blockId="21.[151,1437,150,1365]" pageId="21" pageNumber="472">
<emphasis id="B904EA9FE84817797EADFBD16A37A673" bold="true" box="[199,334,1087,1113]" pageId="21" pageNumber="472">Etymology.</emphasis>
This species is named in honor of the German zoologist Prof. Heinrich Prell (18881962) for his significant contribution to the study of the subfamily
<taxonomicName id="4C704D0EE84817797C87FB8D6816A657" authorityName="MacLeay" authorityYear="1819" box="[749,879,1123,1149]" class="Magnoliopsida" family="Malvaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="21" pageNumber="472" phylum="Tracheophyta" rank="subFamily" subFamily="Dynastinae">Dynastinae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36A6506E84817797EADFB686AB7A68B" box="[199,462,1158,1185]" pageId="21" pageNumber="472" type="distribution">
<paragraph id="8BCF368DE84817797EADFB686AB7A68B" blockId="21.[151,1437,150,1365]" box="[199,462,1158,1185]" pageId="21" pageNumber="472">
<emphasis id="B904EA9FE84817797EADFB686A18A68A" bold="true" box="[199,353,1158,1184]" pageId="21" pageNumber="472">Distribution.</emphasis>
<collectingCountry id="F367761DE84817797F0DFB696AB3A68B" box="[359,458,1159,1185]" name="Vietnam" pageId="21" pageNumber="472">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C36A6506E84817797EADFB4568B5A77F" pageId="21" pageNumber="472" type="discussion">
<paragraph id="8BCF368DE84817797EADFB4568B5A77F" blockId="21.[151,1437,150,1365]" pageId="21" pageNumber="472">
<emphasis id="B904EA9FE84817797EADFB456A42A6EF" bold="true" box="[199,315,1195,1221]" pageId="21" pageNumber="472">Remarks.</emphasis>
Amongst other
<taxonomicName id="4C704D0EE84817797F9DFB4569F9A6EF" authorityName=", Endrodi" authorityYear="1977" box="[503,640,1195,1221]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="genus">
<emphasis id="B904EA9FE84817797F9DFB4569F9A6EF" box="[503,640,1195,1221]" italics="true" pageId="21" pageNumber="472">Eophileurus</emphasis>
</taxonomicName>
from
<collectingCountry id="F367761DE84817797CA2FB456853A6EF" box="[712,810,1195,1221]" name="Vietnam" pageId="21" pageNumber="472">Vietnam</collectingCountry>
,
<taxonomicName id="4C704D0EE84817797D5DFB4568EDA6EF" authority="Yang &amp; Pathomwattananurak, 2022" authorityName="Yang &amp; Pathomwattananurak" authorityYear="2022" box="[823,916,1195,1221]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="prelli" status="sp. nov.">
<emphasis id="B904EA9FE84817797D5DFB4568EDA6EF" box="[823,916,1195,1221]" italics="true" pageId="21" pageNumber="472">E. prelli</emphasis>
</taxonomicName>
,
<emphasis id="B904EA9FE84817797DCEFB456F56A6EE" bold="true" box="[932,1071,1194,1221]" pageId="21" pageNumber="472">
<taxonomicNameLabel id="A23757E4E84817797DCEFB456F56A6EE" box="[932,1071,1194,1221]" pageId="21" pageNumber="472" rank="species">new species</taxonomicNameLabel>
</emphasis>
is related to both
<taxonomicName id="4C704D0EE84817797B6EFB456A02A6C3" authority="Kral &amp; Strnad, 1992" authorityName="Kral &amp; Strnad" authorityYear="1992" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="heyrovskyi">
<emphasis id="B904EA9FE84817797B6EFB456EE4A6EF" box="[1284,1437,1195,1221]" italics="true" pageId="21" pageNumber="472">E. heyrovskyi</emphasis>
<bibRefCitation id="EFE14B7CE84817797EFDFB216A02A6C3" author="Kral, D. &amp; Strnad, J." box="[151,379,1231,1257]" pageId="21" pageNumber="472" pagination="3 - 6" refId="ref17217" refString="Kral, D. &amp; Strnad, J. (1992) New Eophileurus species from Vietnam (Scarabaeidae, Dynastinae). Folia Heyrovskyana, 1, 3 - 6." type="journal article" year="1992">Král &amp; Strnad, 1992</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C704D0EE84817797FDBFB21686EA6C3" authority="Muramoto, 2004" authorityName="Muramoto" authorityYear="2004" box="[433,791,1231,1257]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="baolocensis">
<emphasis id="B904EA9FE84817797FDBFB21692DA6C3" box="[433,596,1231,1257]" italics="true" pageId="21" pageNumber="472">E. baolocensis</emphasis>
<bibRefCitation id="EFE14B7CE84817797C31FB21686EA6C3" author="Muramoto, R." box="[603,791,1231,1257]" pageId="21" pageNumber="472" pagination="21 - 24" refId="ref17354" refString="Muramoto, R. (2004) Notes on the genus Eophileurus Arrow (Coleoptera, Scarabaeidae, Dynastinae) from Southeast Asia, with description of a new species from southern Vietnam. Kogane, 5, 21 - 24." type="journal article" year="2004">Muramoto, 2004</bibRefCitation>
</taxonomicName>
by the rounded posterior angle of pronotum. Although its
<typeStatus id="54CB882FE84817797EFDFB1D6BBEA727" box="[151,199,1267,1293]" pageId="21" pageNumber="472">type</typeStatus>
locality is closer to the distributing area of
<taxonomicName id="4C704D0EE84817797CC9FB1D683CA727" authorityName="Muramoto" authorityYear="2004" box="[675,837,1267,1293]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="baolocensis">
<emphasis id="B904EA9FE84817797CC9FB1D683CA727" box="[675,837,1267,1293]" italics="true" pageId="21" pageNumber="472">E. baolocensis</emphasis>
</taxonomicName>
, it is more closely related to
<taxonomicName id="4C704D0EE84817797AEBFB1D6E6CA727" authorityName="Kral &amp; Strnad" authorityYear="1992" box="[1153,1301,1267,1293]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="heyrovskyi">
<emphasis id="B904EA9FE84817797AEBFB1D6E6CA727" box="[1153,1301,1267,1293]" italics="true" pageId="21" pageNumber="472">E. heyrovskyi</emphasis>
</taxonomicName>
due to them both having backward processes on the outer sides of the parameres.Additionally, since its
<typeStatus id="54CB882FE84817797AEFFAF96FCCA71B" box="[1157,1205,1303,1329]" pageId="21" pageNumber="472">type</typeStatus>
locality is extremely close to
<collectingCountry id="F367761DE84817797E9FFAD56A57A77F" box="[245,302,1339,1365]" name="Laos" pageId="21" pageNumber="472">Laos</collectingCountry>
, occurrence of this species in
<collectingCountry id="F367761DE84817797C17FAD569CAA77F" box="[637,691,1339,1365]" name="Laos" pageId="21" pageNumber="472">Laos</collectingCountry>
would not be surprising.
</paragraph>
</subSubSection>
</treatment>
</document>