treatments-xml/data/A2/1F/87/A21F8799FFCAFF9BFF6EDFCBFDC3F113.xml
2024-06-21 12:46:14 +02:00

374 lines
36 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document ID-DOI="10.11646/phytotaxa.560.1.6" ID-GBIF-Dataset="59400094-97b1-4022-8d25-8b29ef41866e" ID-ISSN="1179-3163" ID-Zenodo-Dep="7031279" checkinTime="1661766202837" checkinUser="plazi" docAuthor="Xu, Rong-Ju, Li, Lu &amp; Zhao, Qi" docDate="2022" docId="A21F8799FFCAFF9BFF6EDFCBFDC3F113" docLanguage="en" docName="phytotaxa.560.1.6.pdf" docOrigin="Phytotaxa 560 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.560.1.6" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.4:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="4" docTitle="Helvella lacunosa Afzelius 1783" docType="treatment" docVersion="7" lastPageNumber="89" masterDocId="5E26FFE1FFCCFF9CFFE6DF56FFC6F50A" masterDocTitle="Helvella cystidiata sp. nov. (Helvellaceae, Ascomycota) from Tibetan Plateau, China" masterLastPageNumber="92" masterPageNumber="82" pageNumber="88" updateTime="1661927200450" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Helvella cystidiata sp. nov. (Helvellaceae, Ascomycota) from Tibetan Plateau, China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Xu, Rong-Ju</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-3968-8442</mods:nameIdentifier>
<mods:affiliation>Yunnan Key Laboratory of Fungal Diversity and Green Development, Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650201, China &amp; Center of Excellence in Fungal Research, Mae Fah Luang University, Chiang Rai 57100, Thailand &amp; School of Science, Mae Fah Luang University, Chiang Rai 57100, Thailand &amp; xurongju 1005 @ outlook. com; https: // orcid. org / 0000 - 0002 - 3968 - 8442</mods:affiliation>
<mods:nameIdentifier type="email">xurongju1005@outlook.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Lu</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-4695-2528</mods:nameIdentifier>
<mods:affiliation>Yunnan Key Laboratory of Fungal Diversity and Green Development, Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650201, China &amp; lilu @ mail. kib. ac. cn; https: // orcid. org / 0000 - 0003 - 4695 - 2528</mods:affiliation>
<mods:nameIdentifier type="email">lilu@mail.kib.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhao, Qi</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8169-0573</mods:nameIdentifier>
<mods:affiliation>Yunnan Key Laboratory of Fungal Diversity and Green Development, Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650201, China &amp; zhaoqi @ mail. kib. ac. cn; https: // orcid. org / 0000 - 0001 - 8169 - 0573</mods:affiliation>
<mods:nameIdentifier type="email">zhaoqi@mail.kib.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-08-29</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>560</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>82</mods:start>
<mods:end>92</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/phytotaxa.560.1.6</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.560.1.6</mods:identifier>
<mods:identifier type="GBIF-Dataset">59400094-97b1-4022-8d25-8b29ef41866e</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7031279</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7036831" ID-GBIF-Taxon="199898919" ID-Zenodo-Dep="7036831" LSID="urn:lsid:plazi:treatment:A21F8799FFCAFF9BFF6EDFCBFDC3F113" httpUri="http://treatment.plazi.org/id/A21F8799FFCAFF9BFF6EDFCBFDC3F113" lastPageId="7" lastPageNumber="89" pageId="6" pageNumber="88">
<subSubSection box="[136,1130,157,184]" pageId="6" pageNumber="88" type="nomenclature">
<paragraph blockId="6.[136,1130,157,184]" box="[136,1130,157,184]" pageId="6" pageNumber="88">
<heading box="[136,1130,157,184]" fontSize="11" level="2" pageId="6" pageNumber="88" reason="3">
<taxonomicName authority="Afzelius, 1783" authorityName="Afzelius" authorityYear="1783" box="[136,517,157,184]" class="Leotiomycetes" family="Chlorociboriaceae" genus="Helvella" kingdom="Fungi" order="Helotiales" pageId="6" pageNumber="88" phylum="Ascomycota" rank="species" species="lacunosa">
<emphasis bold="true" box="[136,343,157,184]" italics="true" pageId="6" pageNumber="88">Helvella lacunosa</emphasis>
<bibRefCitation author="Afzelius, A. &amp; Akrel, F." box="[349,517,158,184]" pageId="6" pageNumber="88" pagination="299 - 313" refId="ref4609" refString="Afzelius, A. &amp; Akrel, F. (1783) Svamp-Slagtet Helvella. Kungliga Svenska Vetenskapsakademiens Handlingar 4: 299 - 313." type="journal article" year="1783">Afzelius, 1783</bibRefCitation>
</taxonomicName>
, Kongl. Vetensk. Akad. Nya Hand. 4: 303.
<figureCitation box="[1000,1130,157,184]" captionStart="FIGURE 4" captionStartId="6.[136,229,1993,2015]" captionTargetBox="[151,1435,530,1964]" captionTargetId="figure-149@6.[151,1436,530,1965]" captionTargetPageId="6" captionText="FIGURE 4. Helvella lacunosa. a, b Mature specimens (a HKAS 87877; b HKAS 87878); c, d Receptacle surface of pileus; e, f Stipitipellis; g, h Asci and paraphyses; ik Asci; l Ascospores. Scale bars: a, b = 1 cm; cl = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.7031294" httpUri="https://zenodo.org/record/7031294/files/figure.png" pageId="6" pageNumber="88">
<emphasis bold="true" box="[1000,1130,157,184]" pageId="6" pageNumber="88">FIGURE 4</emphasis>
</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[136,822,197,219]" pageId="6" pageNumber="88" type="multiple">
<paragraph blockId="6.[136,1451,197,291]" box="[136,822,197,219]" pageId="6" pageNumber="88">
<emphasis box="[136,376,197,219]" italics="true" pageId="6" pageNumber="88">Index Fungorum number</emphasis>
:
<materialsCitation ID-GBIF-Occurrence="3897150302" accessionNumber="IF 495899" box="[389,822,197,219]" pageId="6" pageNumber="88" specimenCount="1">
<accessionNumber box="[389,490,197,219]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/IF495899" pageId="6" pageNumber="88">IF 495899</accessionNumber>
,
<emphasis box="[500,704,197,218]" italics="true" pageId="6" pageNumber="88">Facesoffungi number</emphasis>
: FoF 02201
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="88" type="materials_examined">
<paragraph blockId="6.[136,1451,197,291]" pageId="6" pageNumber="88">
<emphasis box="[136,215,234,255]" italics="true" pageId="6" pageNumber="88">
<typeStatus box="[136,215,234,255]" pageId="6" pageNumber="88">Neotype</typeStatus>
</emphasis>
(here designated):
<collectingCountry box="[398,501,233,254]" name="Sweden" pageId="6" pageNumber="88">SWEDEN</collectingCountry>
, Lapland, Vindeljallets naturreservat, Gieravard, on the ground, under
<taxonomicName authority=", Salix" authorityName="Salix" box="[1175,1349,233,255]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="88" phylum="Tracheophyta" rank="species" species="nana">Betula nana, Salix</taxonomicName>
sp.,
<taxonomicName class="Magnoliopsida" family="Rosaceae" genus="Dryas" kingdom="Plantae" order="Rosales" pageId="6" pageNumber="88" phylum="Tracheophyta" rank="species" species="undetermined">Dryas sp.</taxonomicName>
forests,
<date box="[299,431,269,291]" pageId="6" pageNumber="88" value="2010-08-20">20 Aug. 2010</date>
, K. Hansen, K. Gillen &amp; I. Olariaga (KH.10.97).
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="88" type="diagnosis">
<paragraph blockId="6.[136,1451,338,508]" pageId="6" pageNumber="88">
<emphasis box="[136,206,338,364]" italics="true" pageId="6" pageNumber="88">Pileus</emphasis>
irregularly lobed to slightly saddle-shaped,
<quantity box="[704,788,338,364]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="6" pageNumber="88" unit="cm" value="3.0" valueMax="4.0" valueMin="2.0">24 cm</quantity>
high,
<quantity box="[860,944,338,365]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="6" pageNumber="88" unit="cm" value="3.5" valueMax="5.0" valueMin="2.0">25 cm</quantity>
broad, margin reflexed and fused with stipe;
<emphasis box="[136,251,374,400]" italics="true" pageId="6" pageNumber="88">hymenium</emphasis>
wrinkled to folded towards the center, taupe or nearly black when fresh, blackish when dry, receptacle surface glabrous, grey to pale greyish when fresh, becoming light blackish when dry.
<emphasis box="[1110,1166,410,436]" italics="true" pageId="6" pageNumber="88">Stipe</emphasis>
<quantity box="[1174,1272,410,436]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" metricValueMax="10.0" metricValueMin="3.0" pageId="6" pageNumber="88" unit="cm" value="6.5" valueMax="10.0" valueMin="3.0">310 cm</quantity>
long,
<quantity box="[1346,1451,410,437]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="4.0" metricValueMin="1.5" pageId="6" pageNumber="88" unit="cm" value="2.75" valueMax="4.0" valueMin="1.5">1.54 cm</quantity>
broad, lacunose, with sharp to rounded ribs, internally chambered, pubescent, taupe, yellowish brown to brownish when fresh, becoming dark brown when dry.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7031294" ID-Zenodo-Dep="7031294" httpUri="https://zenodo.org/record/7031294/files/figure.png" pageId="6" pageNumber="88" startId="6.[136,229,1993,2015]" targetBox="[151,1435,530,1964]" targetPageId="6">
<paragraph blockId="6.[136,1452,1993,2051]" pageId="6" pageNumber="88">
<emphasis bold="true" box="[136,252,1993,2015]" pageId="6" pageNumber="88">FIGURE 4.</emphasis>
<emphasis box="[257,436,1993,2014]" italics="true" pageId="6" pageNumber="88">
<taxonomicName authorityName="Afzelius" authorityYear="1783" box="[257,432,1993,2014]" class="Leotiomycetes" family="Chlorociboriaceae" genus="Helvella" kingdom="Fungi" order="Helotiales" pageId="6" pageNumber="88" phylum="Ascomycota" rank="species" species="lacunosa">Helvella lacunosa</taxonomicName>
.
</emphasis>
a, b Mature specimens (a HKAS 87877; b HKAS 87878); c, d Receptacle surface of pileus; e, f Stipitipellis; g, h Asci and paraphyses; ik Asci; l Ascospores. Scale bars: a, b
<emphasis box="[765,781,2030,2051]" italics="true" pageId="6" pageNumber="88">=</emphasis>
1 cm; cl
<emphasis box="[883,899,2030,2051]" italics="true" pageId="6" pageNumber="88">=</emphasis>
50 μm.
</paragraph>
</caption>
<subSubSection pageId="7" pageNumber="89" type="description">
<paragraph blockId="7.[136,1452,158,1049]" pageId="7" pageNumber="89">
<emphasis box="[189,429,158,184]" italics="true" pageId="7" pageNumber="89">Medullary excipulum</emphasis>
270330 μm broad, of textura intricata, hyaline, composed of 34 μm broad hyphae, blue in cotton blue, J- in Melzers reagent.
<emphasis box="[551,732,194,220]" italics="true" pageId="7" pageNumber="89">Ectal excipulum</emphasis>
50110 μm broad, of
<emphasis box="[977,1170,194,220]" italics="true" pageId="7" pageNumber="89">textura angularis</emphasis>
, outermost cells 1537 × 712 μm, catenuliform in long fasciculate tufts, hyaline, evenly blue in cotton blue, J- in Melzers reagent.
<emphasis box="[1324,1452,230,256]" italics="true" pageId="7" pageNumber="89">Stipitipellis</emphasis>
6090 μm, hyaline, composed of cells of
<emphasis box="[610,805,266,292]" italics="true" pageId="7" pageNumber="89">textura angularis</emphasis>
, terminal cells 1330 × 714 μm, clavate, with a yellow refractive content in Melzers reagent, blue in cotton blue.
<emphasis box="[827,875,303,329]" italics="true" pageId="7" pageNumber="89">Asci</emphasis>
arising from croziers, 8-spored, subcylindrical to clavate, with apex rounded, 230275 × 1417 μm, J- in Melzers reagent, blue in cotton blue.
<emphasis box="[1172,1303,338,364]" italics="true" pageId="7" pageNumber="89">Paraphyses</emphasis>
filiform, 35 μm broad, slightly exceeding the asci, with a slightly yellow refractive content in Melzers reagent, blue in cotton blue, apex slightly enlarged, 57 μm broad.
<emphasis box="[561,688,411,437]" italics="true" pageId="7" pageNumber="89">Ascospores</emphasis>
[40/2/2, in H
<subScript attach="both" box="[836,844,429,443]" fontSize="6" pageId="7" pageNumber="89">2</subScript>
O] (13) 1416 (17) × 811 μm [Q = (1.361.81), Qm = (1.52 ± 0.09)], ellipsoid, smooth-walled under the light microscope, uniguttulate, with large central globose to broadly ellipsoidal oil droplet.
<emphasis bold="true" box="[390,572,482,508]" pageId="7" pageNumber="89">Asexual morph</emphasis>
: Undetermined.
</paragraph>
</subSubSection>
<subSubSection box="[189,1236,518,545]" pageId="7" pageNumber="89" type="biology_ecology">
<paragraph blockId="7.[136,1452,158,1049]" box="[189,1236,518,545]" pageId="7" pageNumber="89">
<emphasis bold="true" box="[189,291,518,544]" pageId="7" pageNumber="89">Habitat:</emphasis>
Scattered or gregarious on the ground, under
<taxonomicName box="[804,918,518,545]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="89" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis box="[804,877,518,544]" italics="true" pageId="7" pageNumber="89">Betula</emphasis>
sp.
</taxonomicName>
,
<taxonomicName box="[930,1027,518,545]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="89" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis box="[930,986,518,544]" italics="true" pageId="7" pageNumber="89">Salix</emphasis>
sp.
</taxonomicName>
,
<taxonomicName box="[1039,1147,519,545]" class="Magnoliopsida" family="Rosaceae" genus="Dryas" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="89" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis box="[1039,1107,519,544]" italics="true" pageId="7" pageNumber="89">Dryas</emphasis>
sp.
</taxonomicName>
forests.
</paragraph>
</subSubSection>
<subSubSection box="[189,1392,554,581]" pageId="7" pageNumber="89" type="distribution">
<paragraph blockId="7.[136,1452,158,1049]" box="[189,1392,554,581]" pageId="7" pageNumber="89">
<emphasis bold="true" box="[189,345,554,580]" pageId="7" pageNumber="89">Distribution:</emphasis>
Known in Europe (Dissing, 1966;
<bibRefCitation author="Haffner, J." box="[738,894,554,580]" pageId="7" pageNumber="89" pagination="1 - 165" refId="ref5505" refString="Haffner, J. (1987) Die Gattung Helvella - Morphologie und Taxonomie. Beihefte zur Zeitschrift fur My-kologie 7: 1 - 165." type="journal article" year="1987">Häffner, 1987</bibRefCitation>
) and
<collectingCountry box="[957,1024,554,580]" name="China" pageId="7" pageNumber="89">China</collectingCountry>
(
<bibRefCitation author="Liu, B. &amp; Du, F. &amp; Cao, J. Z." box="[1038,1208,554,581]" pageId="7" pageNumber="89" pagination="208 - 217" refId="ref6582" refString="Liu, B., Du, F. &amp; Cao, J. Z. (1985) New species and new combinations of the genus Helvella. Acta Mycologica Sinica 4: 208 - 217." type="journal article" year="1985">
Liu
<emphasis box="[1086,1135,554,580]" italics="true" pageId="7" pageNumber="89">et al</emphasis>
, 1985
</bibRefCitation>
;
<bibRefCitation author="Zhuang, W. Y." box="[1220,1380,554,580]" pageId="7" pageNumber="89" pagination="35 - 42" refId="ref8023" refString="Zhuang, W. Y. (2004) Preliminary survey of the Helvellaceae from Xinjiang, China. Mycotaxon 90: 35 - 42." type="journal article" year="2004">Zhuang, 2004</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="89" type="materials_examined">
<paragraph blockId="7.[136,1452,158,1049]" pageId="7" pageNumber="89">
<emphasis bold="true" box="[189,429,590,616]" pageId="7" pageNumber="89">Material examined:</emphasis>
<materialsCitation ID-GBIF-Occurrence="3897150301" collectingDate="2014-07-28" collectionCode="HKAS" country="China" county="Linzhi County" elevation="3300" location="Lulang Town" pageId="7" pageNumber="89" specimenCode="HKAS 87877, HKAS 87878" specimenCount="1" stateProvince="Xizang">
<collectingCountry box="[440,532,590,616]" name="China" pageId="7" pageNumber="89">CHINA</collectingCountry>
,
<collectingRegion box="[546,607,590,616]" country="China" name="Xizang" pageId="7" pageNumber="89">Tibet</collectingRegion>
,
<collectingCounty box="[622,790,590,616]" pageId="7" pageNumber="89">Linzhi County</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:A21F8799FFCAFF9BFF6EDFCBFDC3F113:2F696054FFCBFF9BFCC2DD18FC06F763" box="[804,960,590,617]" country="China" county="Linzhi County" name="Lulang Town" pageId="7" pageNumber="89" stateProvince="Xizang">Lulang Town</location>
, alt.
<quantity box="[1021,1112,590,617]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="7" pageNumber="89" unit="m" value="3300.0">
<elevation box="[1021,1112,590,617]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="7" pageNumber="89" unit="m" value="3300.0">3300 m</elevation>
</quantity>
,
<date box="[1127,1276,590,616]" pageId="7" pageNumber="89" value="2014-07-28">
<collectingDate box="[1127,1276,590,616]" pageId="7" pageNumber="89" value="2014-07-28">28 Jul. 2014</collectingDate>
</date>
, Qi Zhao 2111 (
<specimenCode box="[144,303,626,652]" collectionCode="HKAS" country="China" httpUri="http://grbio.org/cool/di2a-40jb" name="Cryptogamic Herbarium of Kunming Institute of Botany" pageId="7" pageNumber="89">HKAS 87877</specimenCode>
), same location,
<date box="[491,631,626,652]" pageId="7" pageNumber="89" value="2014-07-28">
<collectingDate box="[491,631,626,652]" pageId="7" pageNumber="89" value="2014-07-28">28 Jul. 2014</collectingDate>
</date>
, Qi Zhao 2112 (
<specimenCode box="[811,971,626,652]" collectionCode="HKAS" country="China" httpUri="http://grbio.org/cool/di2a-40jb" name="Cryptogamic Herbarium of Kunming Institute of Botany" pageId="7" pageNumber="89">HKAS 87878</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="89" type="discussion">
<paragraph blockId="7.[136,1452,158,1049]" pageId="7" pageNumber="89">
<emphasis bold="true" box="[189,265,662,688]" pageId="7" pageNumber="89">Notes:</emphasis>
<taxonomicName authorityName="Afzelius" authorityYear="1783" box="[272,474,662,688]" class="Leotiomycetes" family="Chlorociboriaceae" genus="Helvella" kingdom="Fungi" order="Helotiales" pageId="7" pageNumber="89" phylum="Ascomycota" rank="species" species="lacunosa">
<emphasis box="[272,474,662,688]" italics="true" pageId="7" pageNumber="89">Helvella lacunosa</emphasis>
</taxonomicName>
was described by
<bibRefCitation author="Afzelius, A. &amp; Akrel, F." box="[675,853,662,688]" pageId="7" pageNumber="89" pagination="299 - 313" refId="ref4609" refString="Afzelius, A. &amp; Akrel, F. (1783) Svamp-Slagtet Helvella. Kungliga Svenska Vetenskapsakademiens Handlingar 4: 299 - 313." type="journal article" year="1783">Afzelius (1783)</bibRefCitation>
, and the
<typeStatus box="[951,999,663,689]" pageId="7" pageNumber="89">type</typeStatus>
was studied by
<bibRefCitation author="Abbott, S. P. &amp; Currah, R. S." box="[1174,1447,662,688]" pageId="7" pageNumber="89" pagination="1 - 125" refId="ref4634" refString="Abbott, S. P. &amp; Currah, R. S. (1997) The Hevellaceae: systematic revision and occurrence in northern and northwestern North America. Mycotaxon 62: 1 - 125." type="journal article" year="1997">Abbott &amp; Currah (1997)</bibRefCitation>
. The distinctive characteristics of our collections, such as irregularly-lobed pileus with a dark brown to blackish hymenium, wrinkled to folded towards the center, a reflexed margin and fusing with stipe in several places, a glabrous, yellowish brown to brownish receptacle surface, and a lacunose, pubescent, yellowish brown, slightly greyish brown to brownish stipe, matches the character variation reported for
<taxonomicName authorityName="Afzelius" authorityYear="1783" box="[839,1042,806,832]" class="Leotiomycetes" family="Chlorociboriaceae" genus="Helvella" kingdom="Fungi" order="Helotiales" pageId="7" pageNumber="89" phylum="Ascomycota" rank="species" species="lacunosa">
<emphasis box="[839,1042,806,832]" italics="true" pageId="7" pageNumber="89">Helvella lacunosa</emphasis>
</taxonomicName>
(Dissing 1966,
<bibRefCitation author="Haffner, J." box="[1221,1374,806,832]" pageId="7" pageNumber="89" pagination="1 - 165" refId="ref5505" refString="Haffner, J. (1987) Die Gattung Helvella - Morphologie und Taxonomie. Beihefte zur Zeitschrift fur My-kologie 7: 1 - 165." type="journal article" year="1987">Häffner 1987</bibRefCitation>
).
</paragraph>
<paragraph blockId="7.[136,1452,158,1049]" pageId="7" pageNumber="89">
<taxonomicName authorityName="Afzelius" authorityYear="1783" box="[189,394,842,868]" class="Leotiomycetes" family="Chlorociboriaceae" genus="Helvella" kingdom="Fungi" order="Helotiales" pageId="7" pageNumber="89" phylum="Ascomycota" rank="species" species="lacunosa">
<emphasis box="[189,394,842,868]" italics="true" pageId="7" pageNumber="89">Helvella lacunosa</emphasis>
</taxonomicName>
has been described and illustrated repeatedly by mycologists worldwide (
<emphasis box="[1238,1271,843,868]" italics="true" pageId="7" pageNumber="89">e.g</emphasis>
., Dissing 1966,
<bibRefCitation author="Weber, N. S." box="[136,274,878,904]" pageId="7" pageNumber="89" pagination="147 - 201" refId="ref7449" refString="Weber, N. S. (1972) The genus Helvella in Michigan. The Michigan Botanist 11: 147 - 201." type="journal article" year="1972">Weber 1972</bibRefCitation>
,
<bibRefCitation author="Haffner, J." box="[286,438,878,904]" pageId="7" pageNumber="89" pagination="1 - 165" refId="ref5505" refString="Haffner, J. (1987) Die Gattung Helvella - Morphologie und Taxonomie. Beihefte zur Zeitschrift fur My-kologie 7: 1 - 165." type="journal article" year="1987">Häffner 1987</bibRefCitation>
,
<bibRefCitation author="Abbott, S. P. &amp; Currah, R. S." box="[448,705,878,904]" pageId="7" pageNumber="89" pagination="1 - 125" refId="ref4634" refString="Abbott, S. P. &amp; Currah, R. S. (1997) The Hevellaceae: systematic revision and occurrence in northern and northwestern North America. Mycotaxon 62: 1 - 125." type="journal article" year="1997">Abbott &amp; Currah 1997</bibRefCitation>
,
<bibRefCitation author="Zhuang, W. Y." box="[716,870,878,904]" pageId="7" pageNumber="89" pagination="35 - 42" refId="ref8023" refString="Zhuang, W. Y. (2004) Preliminary survey of the Helvellaceae from Xinjiang, China. Mycotaxon 90: 35 - 42." type="journal article" year="2004">Zhuang 2004</bibRefCitation>
). The relationship and identity of
<taxonomicName authorityName="Afzelius" authorityYear="1783" box="[1248,1451,878,904]" class="Leotiomycetes" family="Chlorociboriaceae" genus="Helvella" kingdom="Fungi" order="Helotiales" pageId="7" pageNumber="89" phylum="Ascomycota" rank="species" species="lacunosa">
<emphasis box="[1248,1451,878,904]" italics="true" pageId="7" pageNumber="89">Helvella lacunosa</emphasis>
</taxonomicName>
with other species of
<taxonomicName box="[390,485,914,940]" class="Leotiomycetes" family="Chlorociboriaceae" genus="Helvella" kingdom="Fungi" order="Helotiales" pageId="7" pageNumber="89" phylum="Ascomycota" rank="genus">
<emphasis box="[390,485,914,940]" italics="true" pageId="7" pageNumber="89">Helvella</emphasis>
</taxonomicName>
, e.g.,
<taxonomicName box="[561,669,915,941]" class="Leotiomycetes" family="Chlorociboriaceae" genus="Helvella" kingdom="Fungi" order="Helotiales" pageId="7" pageNumber="89" phylum="Ascomycota" rank="species" species="crispa">
<emphasis box="[561,581,915,940]" italics="true" pageId="7" pageNumber="89">H</emphasis>
.
<emphasis box="[599,669,915,940]" italics="true" pageId="7" pageNumber="89">crispa</emphasis>
</taxonomicName>
,
<taxonomicName box="[687,834,914,941]" class="Leotiomycetes" family="Chlorociboriaceae" genus="Helvella" kingdom="Fungi" order="Helotiales" pageId="7" pageNumber="89" phylum="Ascomycota" rank="species" species="dryophila">
<emphasis box="[687,707,915,940]" italics="true" pageId="7" pageNumber="89">H</emphasis>
.
<emphasis box="[725,834,914,940]" italics="true" pageId="7" pageNumber="89">dryophila</emphasis>
</taxonomicName>
,
<taxonomicName box="[851,948,914,941]" class="Leotiomycetes" family="Chlorociboriaceae" genus="Helvella" kingdom="Fungi" order="Helotiales" pageId="7" pageNumber="89" phylum="Ascomycota" rank="species" species="fusca">
<emphasis box="[851,871,915,940]" italics="true" pageId="7" pageNumber="89">H</emphasis>
.
<emphasis box="[889,948,914,940]" italics="true" pageId="7" pageNumber="89">fusca</emphasis>
</taxonomicName>
,
<taxonomicName box="[966,1093,915,941]" class="Leotiomycetes" family="Chlorociboriaceae" genus="Helvella" kingdom="Fungi" order="Helotiales" pageId="7" pageNumber="89" phylum="Ascomycota" rank="species" species="juniperi">
<emphasis box="[966,986,915,940]" italics="true" pageId="7" pageNumber="89">H</emphasis>
.
<emphasis box="[1004,1093,915,940]" italics="true" pageId="7" pageNumber="89">juniperi</emphasis>
</taxonomicName>
,
<taxonomicName box="[1111,1252,914,941]" class="Leotiomycetes" family="Chlorociboriaceae" genus="Helvella" kingdom="Fungi" order="Helotiales" pageId="7" pageNumber="89" phylum="Ascomycota" rank="species" species="maculata">
<emphasis box="[1111,1131,915,940]" italics="true" pageId="7" pageNumber="89">H</emphasis>
.
<emphasis box="[1148,1252,914,940]" italics="true" pageId="7" pageNumber="89">maculata</emphasis>
</taxonomicName>
,
<taxonomicName box="[1270,1406,914,941]" class="Leotiomycetes" family="Chlorociboriaceae" genus="Helvella" kingdom="Fungi" order="Helotiales" pageId="7" pageNumber="89" phylum="Ascomycota" rank="species" species="palustris">
<emphasis box="[1270,1290,915,940]" italics="true" pageId="7" pageNumber="89">H</emphasis>
.
<emphasis box="[1308,1406,914,940]" italics="true" pageId="7" pageNumber="89">palustris</emphasis>
</taxonomicName>
,
<taxonomicName class="Leotiomycetes" family="Chlorociboriaceae" genus="Helvella" kingdom="Fungi" order="Helotiales" pageId="7" pageNumber="89" phylum="Ascomycota" rank="species" species="phlebophora">
<emphasis box="[1424,1444,915,940]" italics="true" pageId="7" pageNumber="89">H</emphasis>
.
<emphasis box="[136,279,950,976]" italics="true" pageId="7" pageNumber="89">phlebophora</emphasis>
</taxonomicName>
,
<taxonomicName box="[294,509,950,977]" class="Leotiomycetes" family="Chlorociboriaceae" genus="Helvella" kingdom="Fungi" order="Helotiales" pageId="7" pageNumber="89" phylum="Ascomycota" rank="species" species="pseudolacunosa">
<emphasis box="[294,314,951,976]" italics="true" pageId="7" pageNumber="89">H</emphasis>
.
<emphasis box="[329,509,950,976]" italics="true" pageId="7" pageNumber="89">pseudolacunosa</emphasis>
</taxonomicName>
and
<taxonomicName box="[566,687,950,977]" class="Leotiomycetes" family="Chlorociboriaceae" genus="Helvella" kingdom="Fungi" order="Helotiales" pageId="7" pageNumber="89" phylum="Ascomycota" rank="species" species="robusta">
<emphasis box="[566,586,951,976]" italics="true" pageId="7" pageNumber="89">H</emphasis>
.
<emphasis box="[602,687,950,976]" italics="true" pageId="7" pageNumber="89">robusta</emphasis>
</taxonomicName>
,
<taxonomicName box="[701,815,951,977]" class="Leotiomycetes" family="Chlorociboriaceae" genus="Helvella" kingdom="Fungi" order="Helotiales" pageId="7" pageNumber="89" phylum="Ascomycota" rank="species" species="rugosa">
<emphasis box="[701,721,951,976]" italics="true" pageId="7" pageNumber="89">H</emphasis>
.
<emphasis box="[737,815,951,976]" italics="true" pageId="7" pageNumber="89">rugosa</emphasis>
</taxonomicName>
,
<taxonomicName box="[829,946,950,977]" class="Leotiomycetes" family="Chlorociboriaceae" genus="Helvella" kingdom="Fungi" order="Helotiales" pageId="7" pageNumber="89" phylum="Ascomycota" rank="species" species="sulcata">
<emphasis box="[829,849,951,976]" italics="true" pageId="7" pageNumber="89">H</emphasis>
.
<emphasis box="[865,946,950,976]" italics="true" pageId="7" pageNumber="89">sulcata</emphasis>
</taxonomicName>
,
<taxonomicName authority="N. H. Nguyen &amp; Vellinga" authorityName="N. H. Nguyen &amp; Vellinga" box="[961,1403,950,977]" class="Leotiomycetes" family="Chlorociboriaceae" genus="Helvella" kingdom="Fungi" order="Helotiales" pageId="7" pageNumber="89" phylum="Ascomycota" rank="species" species="vespertina">
<emphasis box="[961,981,951,976]" italics="true" pageId="7" pageNumber="89">H</emphasis>
.
<emphasis box="[996,1113,951,976]" italics="true" pageId="7" pageNumber="89">vespertina</emphasis>
N.H. Nguyen &amp; Vellinga
</taxonomicName>
and
<taxonomicName box="[136,336,986,1013]" class="Leotiomycetes" family="Chlorociboriaceae" genus="Helvella" kingdom="Fungi" order="Helotiales" pageId="7" pageNumber="89" phylum="Ascomycota" rank="species" species="zhongtiaoensis">
<emphasis box="[136,156,987,1012]" italics="true" pageId="7" pageNumber="89">H</emphasis>
.
<emphasis box="[170,336,986,1012]" italics="true" pageId="7" pageNumber="89">zhongtiaoensis</emphasis>
</taxonomicName>
, have been addressed several times (Dissing 1966,
<bibRefCitation author="Haffner, J." box="[909,1061,986,1012]" pageId="7" pageNumber="89" pagination="1 - 165" refId="ref5505" refString="Haffner, J. (1987) Die Gattung Helvella - Morphologie und Taxonomie. Beihefte zur Zeitschrift fur My-kologie 7: 1 - 165." type="journal article" year="1987">Häffner 1987</bibRefCitation>
,
<bibRefCitation author="Abbott, S. P. &amp; Currah, R. S." box="[1070,1326,986,1012]" pageId="7" pageNumber="89" pagination="1 - 125" refId="ref4634" refString="Abbott, S. P. &amp; Currah, R. S. (1997) The Hevellaceae: systematic revision and occurrence in northern and northwestern North America. Mycotaxon 62: 1 - 125." type="journal article" year="1997">Abbott &amp; Currah 1997</bibRefCitation>
,
<bibRefCitation author="Nguyen, N. H. &amp; Landeros, F. &amp; Garibay-Orijel, R. &amp; Hansen, K. &amp; Vellinga, E. C." pageId="7" pageNumber="89" pagination="1275 - 1286" refId="ref6989" refString="Nguyen, N. H., Landeros, F., Garibay-Orijel, R., Hansen, K. &amp; Vellinga, E. C. (2013) The Helvella lacunosa species complex in western north america: cryptic species, misapplied names and parasites. Mycologia 105: 1275 - 1286. https: // doi. org / 10.3852 / 12 - 391" type="journal article" year="2013">
Nguyen
<emphasis italics="true" pageId="7" pageNumber="89">et al</emphasis>
. 2013
</bibRefCitation>
,
<bibRefCitation author="Ariyawansa, H. &amp; Hyde, K. D. &amp; Jayasiri, S. &amp; Buyck, B. &amp; Chethana, K. W. T. &amp; Dai, D. &amp; Dai, Y. C. &amp; Daranagama, D. &amp; Jayawardena, R. &amp; Lucking, R. &amp; Ghobad-Nejhad, M. &amp; Niskanen, T. &amp; Thambugala, K. &amp; Voigt, K. &amp; Zhao, R. &amp; Li, G. J. &amp; Doilom, M. &amp; Boonmee, S. &amp; Yang, Z. &amp; Cai, Q. &amp; Cui, Y. Y. &amp; Bahkali, A. &amp; Chen, J. &amp; Cui, B. &amp; Chen, J. &amp; Dayarathne, M. &amp; Dissanayake, A. &amp; Ekanayaka, A. &amp; Hashimoto, A. &amp; Hongsanan, S. &amp; Jones, E. B. G. &amp; Larsson, E. &amp; Li, W. &amp; Li, Q. R. &amp; Liu, J. &amp; Luo, Z. &amp; Maharachchikumbura, S. N. &amp; Mapook, A. &amp; McKenzie, E. C. &amp; Norphanphoun, C. &amp; Konta, S. &amp; Pang, K. &amp; Perera, R. &amp; Phookamsak, R. &amp; Phukhamsakda, C. &amp; Pinruan, U. &amp; Randrianjohany, E. &amp; Singtripop, C. &amp; Tanaka, K. &amp; Tian, C. &amp; Tibpromma, S. &amp; Abdel-Wahab, M. &amp; Wanasinghe, D. &amp; Wijayawardene, N. &amp; Zhang, J. F. &amp; Zhang, H. &amp; Abdel-Aziz, F. &amp; Wedin, M. &amp; Westberg, M. &amp; Ammirati, J. &amp; Bulgakov, T. &amp; Lima, D. &amp; Callaghan, T. &amp; Callac, P. &amp; Chang, C. H. &amp; Coca, L. &amp; Dal-Forno, M. &amp; Dollhofer, V. &amp; Fliegerova, K. &amp; Greiner, K. &amp; Griffith, G. &amp; Ho, H. M. &amp; Hofstetter, V. &amp; Jeewon, R. &amp; Kang, J. &amp; Wen, T. C. &amp; Kirk, P. &amp; Kytovuori, I. &amp; Lawrey, J. &amp; Xing, J. &amp; Li, H. &amp; Liu, Z. &amp; Liu, X. &amp; Liimatainen, K. &amp; Lumbsch, H. T. &amp; Matsumura, M. &amp; Moncada, B. &amp; Nuankaew, S. &amp; Parnmen, S. &amp; de Azevedo Santiago, A. L. C. M. &amp; Sommai, S. &amp; Song, Y. &amp; de Souza, C. F. &amp; de Souza-Motta, C. &amp; Su, H. &amp; Suetrong, S. &amp; Wang, Y. &amp; Wei, S. F. &amp; Wen, T. &amp; Yuan, H. &amp; Zhou, L. &amp; Reblova, M. &amp; Fournier, J. &amp; Camporesi, E. &amp; Tasanathai, K. &amp; Khonsanit, A. &amp; Thanakitpipattana, D. &amp; Somrithipol, S. &amp; Diederich, P. &amp; Millanes, A. &amp; Common, R. &amp; Stadler, M. &amp; Yan, J. &amp; Li, X. &amp; Lee, H. &amp; Nguyen, T. T. &amp; Lee, H. &amp; Battistin, E. &amp; Marsico, O. &amp; Vizzini, A. &amp; Vila, J. &amp; Ercole, E. &amp; Eberhardt, U. &amp; Simonini, G. &amp; Wen, H. A. &amp; Chen, X. H." box="[240,505,1022,1049]" pageId="7" pageNumber="89" pagination="1 - 248" refId="ref4671" refString="Ariyawansa, H., Hyde, K. D., Jayasiri, S., Buyck, B., Chethana, K. W. T., Dai, D., Dai, Y. C., Daranagama, D., Jayawardena, R., Lucking, R., Ghobad-Nejhad, M., Niskanen, T., Thambugala, K., Voigt, K., Zhao, R., Li, G. J., Doilom, M., Boonmee, S., Yang, Z., Cai, Q., Cui, Y. Y., Bahkali, A., Chen, J., Cui, B., Chen, J., Dayarathne, M., Dissanayake, A., Ekanayaka, A., Hashimoto, A., Hongsanan, S., Jones, E. B. G., Larsson, E., Li, W., Li, Q. R., Liu, J., Luo, Z., Maharachchikumbura, S. N., Mapook, A., McKenzie, E. C., Norphanphoun, C., Konta, S., Pang, K., Perera, R., Phookamsak, R., Phukhamsakda, C., Pinruan, U., Randrianjohany, E., Singtripop, C., Tanaka, K., Tian, C., Tibpromma, S., Abdel-Wahab, M., Wanasinghe, D., Wijayawardene, N., Zhang, J. F., Zhang, H., Abdel-Aziz, F., Wedin, M., Westberg, M., Ammirati, J., Bulgakov, T., Lima, D., Callaghan, T., Callac, P., Chang, C. H., Coca, L., Dal-Forno, M., Dollhofer, V., Fliegerova, K., Greiner, K., Griffith, G., Ho, H. M., Hofstetter, V., Jeewon, R., Kang, J., Wen, T. C., Kirk, P., Kytovuori, I., Lawrey, J., Xing, J., Li, H., Liu, Z., Liu, X., Liimatainen, K., Lumbsch, H. T., Matsumura, M., Moncada, B., Nuankaew, S., Parnmen, S., de Azevedo Santiago, A. L. C. M., Sommai, S., Song, Y., de Souza, C. F., de Souza-Motta, C., Su, H., Suetrong, S., Wang, Y., Wei, S. F., Wen, T., Yuan, H., Zhou, L., Reblova, M., Fournier, J., Camporesi, E., Luangsa-ard, J. J., Tasanathai, K., Khonsanit, A., Thanakitpipattana, D., Somrithipol, S., Diederich, P., Millanes, A., Common, R., Stadler, M., Yan, J., Li, X., Lee, H., Nguyen, T. T., Lee, H., Battistin, E., Marsico, O., Vizzini, A., Vila, J., Ercole, E., Eberhardt, U., Simonini, G., Wen, H. A. &amp; Chen, X. H. (2015) Fungal diversity notes 111 - 252 - taxonomic and phylogenetic contributions to fungal taxa. Fungal Diversity 75: 1 - 248. https: // doi. org / 10.1007 / s 13225 - 015 - 0346 - 5" type="journal article" year="2015">
Ariyawansa
<emphasis box="[381,430,1022,1048]" italics="true" pageId="7" pageNumber="89">et al</emphasis>
. 2015
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>