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<document id="1402DED43252FBF87EB252BB9B9DBC1E" ID-DOI="10.5281/zenodo.14681675" ID-ISSN="0253-116X" ID-Zenodo-Dep="14681675" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737164562754" checkinUser="felipe" docAuthor="Fery, Hans" docDate="2024" docId="A10856059348FFA2FF24A5F5FC2EF778" docLanguage="en" docName="LinzerbiolBeitr.56.1.365-377.pdf" docOrigin="Linzer biologische Beiträge 56 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.12:LinzerbiolBeitr.2006-.journal_article.type1" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article.type1" docStyleVersion="12" docTitle="Hygrotus (Leptolambus) natrun Fery, 2024, nov.sp." docType="treatment" docVersion="2" lastPageNumber="369" masterDocId="5D312E7D934AFFA6FF87A474FFF8F17B" masterDocTitle="Hygrotus (Leptolambus) natrun nov. sp. from northern Egypt (Coleoptera, Dytiscidae, Hydroporinae)" masterLastPageNumber="377" masterPageNumber="365" pageNumber="367" updateTime="1737164934474" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="203647F67AEB94AFEA57BCB0782AFC32" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="65A84A10A786CBA9EE7AE494DA8D5BCC">Hygrotus (Leptolambus) natrun nov. sp. from northern Egypt (Coleoptera, Dytiscidae, Hydroporinae)</mods:title>
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<mods:namePart id="969858555E0F7170A5E5E97276B198B7">Fery, Hans</mods:namePart>
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<mods:title id="DD30AF452FCB5BD4A4799023668BF4C5">Linzer biologische Beiträge</mods:title>
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<mods:date id="E27531149D16029CBD85898302600227">2024</mods:date>
<mods:detail id="8AAEA801FAA868E8114333E441DB3AC0" type="volume">
<mods:number id="185D59D07A70C57FAF7AFDA8B41E0E1A">56</mods:number>
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<mods:classification id="9F45E7DB4F06BBFBF779E3C3443603EA">journal article</mods:classification>
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<mods:identifier id="B417373309E4D4173C0651A6703A2C91" type="ISSN">0253-116X</mods:identifier>
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<treatment id="A10856059348FFA2FF24A5F5FC2EF778" LSID="urn:lsid:plazi:treatment:A10856059348FFA2FF24A5F5FC2EF778" httpUri="http://treatment.plazi.org/id/A10856059348FFA2FF24A5F5FC2EF778" lastPageId="4" lastPageNumber="369" pageId="2" pageNumber="367">
<subSubSection id="61BBB4989348FFA4FF24A5F5FDBBF0EE" box="[163,579,384,406]" pageId="2" pageNumber="367" type="nomenclature">
<paragraph id="291EE7139348FFA4FF24A5F5FDBBF0EE" blockId="2.[163,579,384,406]" box="[163,579,384,406]" pageId="2" pageNumber="367">
<heading id="7256507F9348FFA4FF24A5F5FDBBF0EE" box="[163,579,384,406]" fontSize="8" level="2" pageId="2" pageNumber="367" reason="2">
<emphasis id="1BD53B019348FFA4FF24A5F5FDBBF0EE" bold="true" box="[163,579,384,406]" pageId="2" pageNumber="367">
<taxonomicName id="EEA19C909348FFA4FF24A5F5FE08F0ED" authorityName="Fery" authorityYear="2024" box="[163,496,384,406]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="367" phylum="Arthropoda" rank="species" species="natrun" status="sp. nov." subGenus="Leptolambus">
<emphasis id="1BD53B019348FFA4FF24A5F5FEFCF0ED" bold="true" box="[163,260,385,406]" italics="true" pageId="2" pageNumber="367">Hygrotus</emphasis>
(
<emphasis id="1BD53B019348FFA4FE94A5F4FE62F0ED" bold="true" box="[275,410,384,406]" italics="true" pageId="2" pageNumber="367">Leptolambus</emphasis>
)
<emphasis id="1BD53B019348FFA4FE2FA5F5FE08F0ED" bold="true" box="[424,496,385,406]" italics="true" pageId="2" pageNumber="367">natrun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00E6867A9348FFA4FE70A5F4FDBBF0EE" box="[503,579,384,405]" pageId="2" pageNumber="367" rank="species">nov.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61BBB4989348FFA4FF24A5D2FD9AF39D" pageId="2" pageNumber="367" type="reference_group">
<paragraph id="291EE7139348FFA4FF24A5D2FCFEF0C7" blockId="2.[163,1071,422,1038]" box="[163,774,422,444]" pageId="2" pageNumber="367">
<treatmentCitationGroup id="09B1C03D9348FFA4FF24A5D2FCFEF0C7" box="[163,774,422,444]" pageId="2" pageNumber="367">
?
<taxonomicName id="EEA19C909348FFA4FF34A5D2FDFFF0C7" authority="SCHAUM, 1857" authorityName="SCHAUM" authorityYear="1857" box="[179,519,422,444]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="367" phylum="Arthropoda" rank="species" species="lernaeus">
<emphasis id="1BD53B019348FFA4FF34A5D2FE8CF0C0" box="[179,372,422,443]" italics="true" pageId="2" pageNumber="367">Hydroporus lernaeus</emphasis>
<bibRefCitation id="4D309AE29348FFA4FEFDA5D2FDFFF0C7" author="SCHAUM H." box="[378,519,422,444]" pageId="2" pageNumber="367" pagination="116 - 158" refId="ref6309" refString="SCHAUM H. (1857): Beitrag zur Kaferfauna Griechenlands. Erstes Stuck: Cicindelidae, Carabici, Dytiscidae, Gyrinidae. - Berliner Entomologische Zeitschrift 1: 116 - 158." type="journal article" year="1857">
<smallCapsWord id="2FF871CF9348FFA4FEFDA5D2FE33F0C0" baselines="438,439" box="[378,459,422,443]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Schaum" pageId="2" pageNumber="367">SCHAUM</smallCapsWord>
, 1857
</bibRefCitation>
</taxonomicName>
;
<treatmentCitation id="A800C1029348FFA4FD97A5D2FD57F0C7" author="SHARP D." box="[528,687,422,444]" page="405" pageId="2" pageNumber="367" year="1882">
<bibRefCitation id="4D309AE29348FFA4FD97A5D2FD57F0C7" author="SHARP D." box="[528,687,422,444]" pageId="2" pageNumber="367" pagination="179 - 1003" refId="ref6342" refString="SHARP D. (1882): On aquatic carnivorous Coleoptera or Dytiscidae. - Scientific Transactions of the Royal Dublin Society (2) 2: 179 - 1003 + pls. 7 - 18." type="journal article" year="1882">
<smallCapsWord id="2FF871CF9348FFA4FD97A5D2FDB7F0C0" baselines="438,439" box="[528,591,422,443]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Sharp" pageId="2" pageNumber="367">SHARP</smallCapsWord>
1882: 405
</bibRefCitation>
</treatmentCitation>
(partim).
</treatmentCitationGroup>
</paragraph>
<paragraph id="291EE7139348FFA4FF24A5B4FED1F095" blockId="2.[163,1071,422,1038]" pageId="2" pageNumber="367">
?
<emphasis id="1BD53B019348FFA4FF3EA5B4FE20F0AE" box="[185,472,448,469]" italics="true" pageId="2" pageNumber="367">Coelambus parallellogrammus</emphasis>
<taxonomicName id="EEA19C909348FFA4FE63A5B4FD19F0AD" authority="(SCHAUM, 1857)" baseAuthorityName="SCHAUM" baseAuthorityYear="1857" box="[484,737,448,470]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="367" phylum="Arthropoda" rank="species" species="lernaeus">
<emphasis id="1BD53B019348FFA4FE63A5B4FDCAF0AE" box="[484,562,448,469]" italics="true" pageId="2" pageNumber="367">lernaeus</emphasis>
(
<bibRefCitation id="4D309AE29348FFA4FDC2A5B4FD22F0AD" author="SCHAUM H." box="[581,730,448,470]" pageId="2" pageNumber="367" pagination="116 - 158" refId="ref6309" refString="SCHAUM H. (1857): Beitrag zur Kaferfauna Griechenlands. Erstes Stuck: Cicindelidae, Carabici, Dytiscidae, Gyrinidae. - Berliner Entomologische Zeitschrift 1: 116 - 158." type="journal article" year="1857">
<smallCapsWord id="2FF871CF9348FFA4FDC2A5B4FD6FF0AE" baselines="464,465" box="[581,663,448,469]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Schaum" pageId="2" pageNumber="367">SCHAUM</smallCapsWord>
, 1857
</bibRefCitation>
)
</taxonomicName>
;
<smallCapsWord id="2FF871CF9348FFA4FD77A5B5FCD6F0AF" baselines="465,464" box="[752,814,449,470]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Bedel" pageId="2" pageNumber="367">BEDEL</smallCapsWord>
1895-1925: 339, footnote (partim).
</paragraph>
<paragraph id="291EE7139348FFA4FF24A586FF0DF34C" blockId="2.[163,1071,422,1038]" pageId="2" pageNumber="367">
<treatmentCitationGroup id="09B1C03D9348FFA4FF24A586FF0DF34C" pageId="2" pageNumber="367">
<emphasis id="1BD53B019348FFA4FF24A586FEF1F37C" box="[163,265,498,519]" italics="true" pageId="2" pageNumber="367">Coelambus</emphasis>
<taxonomicName id="EEA19C909348FFA4FE96A586FDFFF373" authority="(SCHAUM, 1857)" baseAuthorityName="SCHAUM" baseAuthorityYear="1857" box="[273,519,498,520]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="367" phylum="Arthropoda" rank="species" species="lernaeus">
<emphasis id="1BD53B019348FFA4FE96A586FEA7F37C" box="[273,351,498,519]" italics="true" pageId="2" pageNumber="367">lernaeus</emphasis>
(
<bibRefCitation id="4D309AE29348FFA4FEE8A586FDF9F373" author="SCHAUM H." box="[367,513,498,520]" pageId="2" pageNumber="367" pagination="116 - 158" refId="ref6309" refString="SCHAUM H. (1857): Beitrag zur Kaferfauna Griechenlands. Erstes Stuck: Cicindelidae, Carabici, Dytiscidae, Gyrinidae. - Berliner Entomologische Zeitschrift 1: 116 - 158." type="journal article" year="1857">
<smallCapsWord id="2FF871CF9348FFA4FEE8A586FE39F37C" baselines="514,515" box="[367,449,498,519]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Schaum" pageId="2" pageNumber="367">SCHAUM</smallCapsWord>
, 1857
</bibRefCitation>
)
</taxonomicName>
;
<treatmentCitation id="A800C1029348FFA4FD94A587FD18F373" author="REGIMBART M." box="[531,736,498,520]" page="35" pageId="2" pageNumber="367" year="1895">
<bibRefCitation id="4D309AE29348FFA4FD94A587FD18F373" author="REGIMBART M." box="[531,736,498,520]" pageId="2" pageNumber="367" pagination="1 - 244" refId="ref6276" refString="REGIMBART M. (1895): Revision des Dytiscidae et Gyrinidae d'Afrique et Madagascar et iles voisines. - Memoires de la Societe Entomologiques de Belgique 4: 1 - 244." type="journal article" year="1895">
<smallCapsWord id="2FF871CF9348FFA4FD94A587FD7CF37C" baselines="515,514" box="[531,644,498,520]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Regimbart" pageId="2" pageNumber="367">RÉGIMBART</smallCapsWord>
1895: 35
</bibRefCitation>
</treatmentCitation>
(partim);
<treatmentCitation id="A800C1029348FFA4FCC6A587FBD6F373" author="ZIMMERMANN A." box="[833,1070,498,520]" page="151" pageId="2" pageNumber="367" year="1919">
<bibRefCitation id="4D309AE29348FFA4FCC6A587FBD6F373" author="ZIMMERMANN A." box="[833,1070,498,520]" pageId="2" pageNumber="367" pagination="68 - 249" refId="ref6528" refString="ZIMMERMANN A. (1919): Die Schwimmkafer des Deutschen Entomologischen Museums in Berlin-Dahlem. - Archiv fur Naturgeschichte 83 (1917) (A 12): 68 - 249." type="journal article" year="1919">
<smallCapsWord id="2FF871CF9348FFA4FCC6A587FC3FF37C" baselines="515,515" box="[833,967,499,520]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Zimmermann" pageId="2" pageNumber="367">ZIMMERMANN</smallCapsWord>
1919: 151
</bibRefCitation>
</treatmentCitation>
(partim); 1920: 68 (partim); 1930: 101 (partim);
<treatmentCitation id="A800C1029348FFA4FD14A67EFCB5F364" author="GUIGNOT F." box="[659,845,522,544]" page="332" pageId="2" pageNumber="367" year="1959">
<bibRefCitation id="4D309AE29348FFA4FD14A67EFCB5F364" author="GUIGNOT F." box="[659,845,522,544]" pageId="2" pageNumber="367" pagination="323 - 648" refId="ref5985" refString="GUIGNOT F. (1959): Revision des hydrocanthares d'Afrique (Coleoptera Dytiscoidea). 2. - Annales du Musee Royal du Congo Belge Serie 8 vo (Sciences Zoologiques) 78: 323 - 648." type="journal article" year="1959">
<smallCapsWord id="2FF871CF9348FFA4FD14A67EFD13F364" baselines="538,539" box="[659,747,522,543]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Guignot" pageId="2" pageNumber="367">GUIGNOT</smallCapsWord>
1959: 332
</bibRefCitation>
</treatmentCitation>
(partim);
<treatmentCitation id="A800C1029348FFA4FC2DA67EFF09F34C" author="ALFIERI A." page="33" pageId="2" pageNumber="367" year="1976">
<bibRefCitation id="4D309AE29348FFA4FC2DA67EFF09F34C" author="ALFIERI A." pageId="2" pageNumber="367" pagination="1 - 361" refId="ref5595" refString="ALFIERI A. (1976): The Coleoptera of Egypt. - Memoires de la Societe Entomologique d'Egypte 5: i-xvi, 1 - 361." type="journal article" year="1976">
<smallCapsWord id="2FF871CF9348FFA4FC2DA67EFC0CF365" baselines="538,538" box="[938,1012,522,543]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Alfieri" pageId="2" pageNumber="367">ALFIERI</smallCapsWord>
1976: 33
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="291EE7139348FFA4FF24A649FE2DF311" blockId="2.[163,1071,422,1038]" pageId="2" pageNumber="367">
<treatmentCitationGroup id="09B1C03D9348FFA4FF24A649FE2DF311" pageId="2" pageNumber="367">
<taxonomicName id="EEA19C909348FFA4FF24A649FD82F329" ID-CoL="8LZGZ" authority="(SCHAUM, 1857)" baseAuthorityName="SCHAUM" baseAuthorityYear="1857" box="[163,634,572,594]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="367" phylum="Arthropoda" rank="species" species="lernaeus" subGenus="Coelambus">
<emphasis id="1BD53B019348FFA4FF24A649FF00F329" box="[163,248,573,594]" italics="true" pageId="2" pageNumber="367">Hygrotus</emphasis>
(
<emphasis id="1BD53B019348FFA4FE8EA648FE97F32A" box="[265,367,572,593]" italics="true" pageId="2" pageNumber="367">Coelambus</emphasis>
)
<emphasis id="1BD53B019348FFA4FE06A648FE37F32A" box="[385,463,572,593]" italics="true" pageId="2" pageNumber="367">lernaeus</emphasis>
(
<bibRefCitation id="4D309AE29348FFA4FE67A648FD8BF329" author="SCHAUM H." box="[480,627,572,594]" pageId="2" pageNumber="367" pagination="116 - 158" refId="ref6309" refString="SCHAUM H. (1857): Beitrag zur Kaferfauna Griechenlands. Erstes Stuck: Cicindelidae, Carabici, Dytiscidae, Gyrinidae. - Berliner Entomologische Zeitschrift 1: 116 - 158." type="journal article" year="1857">
<smallCapsWord id="2FF871CF9348FFA4FE67A648FDCAF32A" baselines="588,589" box="[480,562,572,593]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Schaum" pageId="2" pageNumber="367">SCHAUM</smallCapsWord>
, 1857
</bibRefCitation>
)
</taxonomicName>
;
<treatmentCitation id="A800C1029348FFA4FD0FA649FC9CF329" author="ZALAT S. &amp; SALEH AHMED R. &amp; ANGUS R. B." box="[648,868,572,594]" page="26" pageId="2" pageNumber="367" year="2000">
<bibRefCitation id="4D309AE29348FFA4FD0FA649FC9CF329" author="ZALAT S. &amp; SALEH AHMED R. &amp; ANGUS R. B." box="[648,868,572,594]" pageId="2" pageNumber="367" pagination="1 - 107" refId="ref6482" refString="ZALAT S., SALEH AHMED R., ANGUS R. B. &amp; A. KASCHEF (2000): Diving beetles (Coleoptera: Dytiscidae and Noteridae) of Egypt. - Egyptian Journal of Natural History 2: 1 - 107." type="journal article" year="2000">
<smallCapsWord id="2FF871CF9348FFA4FD0FA649FD30F32A" baselines="589,589" box="[648,712,573,594]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Zalat" pageId="2" pageNumber="367">ZALAT</smallCapsWord>
et al. 2000: 26
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="A800C1029348FFA4FCF5A649FBD5F32A" author="NILSSON A. N." box="[882,1069,572,594]" page="207" pageId="2" pageNumber="367" year="2001">
<bibRefCitation id="4D309AE29348FFA4FCF5A649FBD5F32A" author="NILSSON A. N." box="[882,1069,572,594]" pageId="2" pageNumber="367" pagination="1 - 395" refId="ref6087" refString="NILSSON A. N. (2001): Dytiscidae (Coleoptera). - In: World Catalogue of insects 3: 1 - 395." type="journal article" year="2001">
<smallCapsWord id="2FF871CF9348FFA4FCF5A649FC3BF32A" baselines="589,589" box="[882,963,573,594]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Nilsson" pageId="2" pageNumber="367">NILSSON</smallCapsWord>
2001: 207
</bibRefCitation>
</treatmentCitation>
(partim); 2003: 72 (partim).
</treatmentCitationGroup>
</paragraph>
<paragraph id="291EE7139348FFA4FF24A61BFEA8F3E7" blockId="2.[163,1071,422,1038]" pageId="2" pageNumber="367">
<taxonomicName id="EEA19C909348FFA4FF24A61BFD81F3FF" ID-CoL="8LZGZ" authority="(SCHAUM, 1857)" baseAuthorityName="SCHAUM" baseAuthorityYear="1857" box="[163,633,622,644]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="367" phylum="Arthropoda" rank="species" species="lernaeus" subGenus="Leptolambus">
<emphasis id="1BD53B019348FFA4FF24A61BFF00F3FF" box="[163,248,623,644]" italics="true" pageId="2" pageNumber="367">Hygrotus</emphasis>
(
<emphasis id="1BD53B019348FFA4FE82A61AFE82F3F8" box="[261,378,622,643]" italics="true" pageId="2" pageNumber="367">Leptolambus</emphasis>
)
<emphasis id="1BD53B019348FFA4FE0FA61AFE2EF3F8" box="[392,470,622,643]" italics="true" pageId="2" pageNumber="367">lernaeus</emphasis>
(
<bibRefCitation id="4D309AE29348FFA4FE64A61BFD8BF3FF" author="SCHAUM H." box="[483,627,622,644]" pageId="2" pageNumber="367" pagination="116 - 158" refId="ref6309" refString="SCHAUM H. (1857): Beitrag zur Kaferfauna Griechenlands. Erstes Stuck: Cicindelidae, Carabici, Dytiscidae, Gyrinidae. - Berliner Entomologische Zeitschrift 1: 116 - 158." type="journal article" year="1857">
<smallCapsWord id="2FF871CF9348FFA4FE64A61BFDCDF3F8" baselines="639,639" box="[483,565,623,644]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Schaum" pageId="2" pageNumber="367">SCHAUM</smallCapsWord>
, 1857
</bibRefCitation>
)
</taxonomicName>
;
<smallCapsWord id="2FF871CF9348FFA4FD04A61BFD2CF3F8" baselines="639,639" box="[643,724,623,644]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Nilsson" pageId="2" pageNumber="367">NILSSON</smallCapsWord>
&amp;
<smallCapsWord id="2FF871CF9348FFA4FD74A61BFCC9F3F8" baselines="639,638" box="[755,817,622,644]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Hajek" pageId="2" pageNumber="367">HÁJEK</smallCapsWord>
2024a: 46 (partim); 2024b: 197 (partim).
</paragraph>
<paragraph id="291EE7139348FFA4FF24A6D5FD9AF39D" blockId="2.[163,1071,422,1038]" pageId="2" pageNumber="367">
T y p e l o c a l i t y: Wadi El Natrun (also Wadi Natrun, Wadi an-Natrun, Wadi Natroun), ca.
<quantity id="EE594AF69348FFA4FF5FA6CDFEE7F3B5" box="[216,287,696,718]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="367" unit="km" value="100.0">100 km</quantity>
NW
<collectingRegion id="EB6529F19348FFA4FED3A6CCFE71F3B5" box="[340,393,696,718]" country="Egypt" name="Al Qahirah" pageId="2" pageNumber="367">Cairo</collectingRegion>
, ca.
<geoCoordinate id="4C9581D49348FFA4FE32A6CDFE01F3B5" box="[437,505,697,718]" degrees="30.41" direction="north" orientation="latitude" pageId="2" pageNumber="367" precision="555" value="30.41">30.41N</geoCoordinate>
<geoCoordinate id="4C9581D49348FFA4FD87A6CDFDBCF3B5" box="[512,580,697,718]" degrees="30.30" direction="east" orientation="longitude" pageId="2" pageNumber="367" precision="555" value="30.3">30.30E</geoCoordinate>
. N o t e s: This wadi is about
<quantity id="EE594AF69348FFA4FCE0A6CCFC6FF3B5" box="[871,919,696,718]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="367" unit="m" value="25.0">25 m</quantity>
below sea level and about
<quantity id="EE594AF69348FFA4FEB1A6A5FE9DF39D" box="[310,357,721,742]" metricMagnitude="1" metricUnit="m" metricValue="3.8" pageId="2" pageNumber="367" unit="m" value="38.0">38 m</quantity>
below the level of the Nile.
</paragraph>
</subSubSection>
<subSubSection id="61BBB4989348FFA2FF24A685FC2EF778" lastPageId="4" lastPageNumber="369" pageId="2" pageNumber="367" type="description">
<paragraph id="291EE7139348FFA4FF24A685FDE5F575" blockId="2.[163,1071,422,1038]" pageId="2" pageNumber="367">
<materialsCitation id="99C9ED4E9348FFA4FF24A685FC39F265" collectorName="Coll. Alfieri &amp; Coll. F. Guignot" country="Egypt" location="Wadi Natroun" pageId="2" pageNumber="367" specimenCount="2" specimenCount-female="2" typeStatus="holotype">
M a t e r i a l e x a m i n e d:
<typeStatus id="F61A59B19348FFA4FE6FA684FDB8F27D" box="[488,576,752,774]" pageId="2" pageNumber="367" type="holotype">Holotype</typeStatus>
:
<specimenCount id="3FA72C9A9348FFA4FDCBA685FDA5F27D" box="[588,605,753,774]" count="1" pageId="2" pageNumber="367" type="female">♀</specimenCount>
, &quot;
<location id="2C7EB1C89348FFA4FDF6A684FD0FF27D" LSID="urn:lsid:plazi:treatment:A10856059348FFA2FF24A5F5FC2EF778:2C7EB1C89348FFA4FDF6A684FD0FF27D" box="[625,759,752,774]" country="Egypt" name="Wadi Natroun" pageId="2" pageNumber="367">Wadi Natroun</location>
, 2.5.[19]27&quot; [hw Guignot], &quot;
<collectorName id="845482C59348FFA4FB87A684FEF4F265" pageId="2" pageNumber="367">Coll. Alfieri</collectorName>
,
<collectingCountry id="51B6A7839348FFA4FE9FA77DFEA1F265" box="[280,345,777,798]" name="Egypt" pageId="2" pageNumber="367">Egypte</collectingCountry>
&quot; [printed], &quot;
<specimenCount id="3FA72C9A9348FFA4FE4AA77DFE25F265" box="[461,477,777,798]" count="1" pageId="2" pageNumber="367" type="female">♀</specimenCount>
&quot;, &quot;Muséum Paris, 1960,
<collectorName id="845482C59348FFA4FD4DA77CFC9CF265" box="[714,868,776,798]" pageId="2" pageNumber="367">Coll. F. Guignot</collectorName>
&quot; [printed]
</materialsCitation>
;
<materialsCitation id="99C9ED4E9348FFA4FC49A77CFBD2F235" collectionCode="MNHN" collectorName="Guignot &amp; Coelambus" country="Egypt" determinerName="H. Fery" pageId="2" pageNumber="367" specimenCount="1" typeStatus="holotype">
&quot;
<typeStatus id="F61A59B19348FFA4FC53A77CFBD2F265" box="[980,1066,776,798]" pageId="2" pageNumber="367" type="holotype">Holotype</typeStatus>
,
<taxonomicName id="EEA19C909348FFA4FF57A755FE1EF24D" authorityName="Fery" authorityYear="2024" box="[208,486,800,822]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="367" phylum="Arthropoda" rank="species" species="natrun" status="sp. nov." subGenus="Leptolambus">
<emphasis id="1BD53B019348FFA4FF57A755FED9F24D" box="[208,289,801,822]" italics="true" pageId="2" pageNumber="367">Hygrotus</emphasis>
(
<emphasis id="1BD53B019348FFA4FEA9A754FE67F24E" box="[302,415,800,821]" italics="true" pageId="2" pageNumber="367">Leptolambus</emphasis>
)
<emphasis id="1BD53B019348FFA4FE2BA755FE1EF24D" box="[428,486,801,822]" italics="true" pageId="2" pageNumber="367">natrun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00E6867A9348FFA4FE6AA755FDD6F24D" box="[493,558,801,822]" pageId="2" pageNumber="367" rank="species">nov.sp.</taxonomicNameLabel>
,
<determinerName id="DCC20BD79348FFA4FDB0A755FD82F24D" box="[567,634,801,822]" pageId="2" pageNumber="367">H. Fery</determinerName>
det. 2024&quot; [red, printed] (
<collectionCode id="4FB07FD69348FFA4FCE7A755FC5EF24D" box="[864,934,801,822]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="367">MNHN</collectionCode>
) [for labels see
<figureCitation id="B19AFB969348FFA4FF57A74CFEFDF235" box="[208,261,824,846]" captionStart="Fig" captionStartId="9.[163,195,1508,1530]" captionTargetBox="[164,1067,201,1493]" captionTargetId="figure-23@9.[164,1067,201,1493]" captionTargetPageId="9" captionText="Fig. 1: Habitus of Hygrotus (Leptolambus) natrun nov.sp. and labels (holotype from Wadi El Natrun, Egypt)." figureDoi="http://doi.org/10.5281/zenodo.14681677" httpUri="https://zenodo.org/record/14681677/files/figure.png" pageId="2" pageNumber="367">Fig. 1</figureCitation>
; specimen until now kept in coll.
<collectorName id="845482C59348FFA4FDB1A74CFD86F235" box="[566,638,824,846]" pageId="2" pageNumber="367">Guignot</collectorName>
under
<emphasis id="1BD53B019348FFA4FD3AA74CFCD8F236" box="[701,800,824,845]" italics="true" pageId="2" pageNumber="367">
<collectorName id="845482C59348FFA4FD3AA74CFCD8F236" box="[701,800,824,845]" pageId="2" pageNumber="367">Coelambus</collectorName>
</emphasis>
<taxonomicName id="EEA19C909348FFA4FCA0A74CFBD9F236" box="[807,1057,824,846]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="367" phylum="Arthropoda" rank="variety" species="lernaeus" variety="orthogrammus">
<emphasis id="1BD53B019348FFA4FCA0A74CFC8AF236" box="[807,882,824,845]" italics="true" pageId="2" pageNumber="367">lernaeus</emphasis>
var.
<emphasis id="1BD53B019348FFA4FC27A74CFBD9F236" box="[928,1057,824,845]" italics="true" pageId="2" pageNumber="367">orthogrammus</emphasis>
</taxonomicName>
]
</materialsCitation>
.
<materialsCitation id="99C9ED4E9348FFA4FF57A725FDE1F575" collectingDate="1982-09-27" collectingDateMax="1996-04-30" collectingDateMin="1982-09-27" collectionCode="NHMW, CHF, OLML, CLH, ZSM" collectorName="Coll. A. Alfieri &amp; Coll. Gschwendtner &amp; W Desert Oasis Bahariva &amp; W. G. Ullrich &amp; Zimmermann &amp; Samml. A. Zimmermann &amp; Coelambus &amp; Fery &amp; All" country="Egypt" determinerName="Gschwendtner &amp; Fery" latitude="28.33" location="Desert Oasis" longLatPrecision="740" longitude="29.0" municipality="Bahariva Oases" pageId="2" pageNumber="367" specimenCount="2" specimenCount-female="2" stateProvince="Al Qahirah" typeStatus="paratype">
<typeStatus id="F61A59B19348FFA4FF57A725FED4F21D" box="[208,300,849,870]" pageId="2" pageNumber="367" type="paratype">Paratypes</typeStatus>
: 2♁♁, &quot;Ägypten
<date id="5D1FC1D39348FFA4FE42A725FDE6F21D" box="[453,542,849,870]" pageId="2" pageNumber="367" value="1982-09-27">
<collectingDate id="4D5B383B9348FFA4FE42A725FDE6F21D" box="[453,542,849,870]" pageId="2" pageNumber="367" value="1982-09-27">27.9.1982</collectingDate>
</date>
, Wadi Natrun, Kafr Daoud, Sattm.&quot; [hw?] (
<collectionCode id="4FB07FD69348FFA4FC26A725FC15F21D" box="[929,1005,849,870]" country="Austria" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="367" type="Museum">NHMW</collectionCode>
,
<collectionCode id="4FB07FD69348FFA4FC71A725FBDDF21D" box="[1014,1061,849,870]" pageId="2" pageNumber="367">CHF</collectionCode>
). 2♁♁, &quot;
<collectingMunicipality id="C97A7D699348FFA4FE89A71CFE64F205" box="[270,412,872,894]" pageId="2" pageNumber="367">Bahariva Oases</collectingMunicipality>
, 20.–24.3 [19]25&quot; [printed], &quot;
<collectingCountry id="51B6A7839348FFA4FD27A71DFD21F205" box="[672,729,873,894]" name="Egypt" pageId="2" pageNumber="367">Egypt</collectingCountry>
, Min. Agric. (
<collectingCountry id="51B6A7839348FFA4FCD1A71DFC76F205" box="[854,910,873,894]" name="Egypt" pageId="2" pageNumber="367">Egypt</collectingCountry>
),
<collectorName id="845482C59348FFA4FC19A71CFBD0F205" box="[926,1064,872,894]" pageId="2" pageNumber="367">Coll. A. Alfieri</collectorName>
&quot; [printed], &quot;
<collectorName id="845482C59348FFA4FEA9A7F4FE1BF2ED" box="[302,483,896,918]" pageId="2" pageNumber="367">Coll. Gschwendtner</collectorName>
&quot; [printed],
<taxonomicName id="EEA19C909348FFA4FDC0A7F5FD0EF2ED" authority="SCHAUM." authorityName="SCHAUM." box="[583,758,896,918]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="367" phylum="Arthropoda" rank="species" species="lernaeus">
&quot;
<emphasis id="1BD53B019348FFA4FDD7A7F4FD63F2EE" box="[592,667,896,917]" italics="true" pageId="2" pageNumber="367">lernaeus</emphasis>
<smallCapsWord id="2FF871CF9348FFA4FD26A7F5FD09F2EE" baselines="913,913" box="[673,753,897,918]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Schaum" pageId="2" pageNumber="367">SCHAUM</smallCapsWord>
.
</taxonomicName>
, det.
<determinerName id="DCC20BD79348FFA4FCAFA7F4FC51F2ED" box="[808,937,896,918]" pageId="2" pageNumber="367">Gschwendtner</determinerName>
&quot; [first line hw, second line printed] (
<collectionCode id="4FB07FD69348FFA4FE13A7EDFE2FF2D5" box="[404,471,921,942]" country="Austria" httpUri="http://grbio.org/cool/pp0t-48s3" name="Oberösterreichisches Landesmuseum" pageId="2" pageNumber="367" type="Museum">OLML</collectionCode>
).
<specimenCount id="3FA72C9A9348FFA4FE6EA7EDFDFDF2D5" box="[489,517,921,942]" count="1" pageId="2" pageNumber="367" type="female">1♀</specimenCount>
, &quot;
<collectingCountry id="51B6A7839348FFA4FD9EA7EDFDAAF2D5" box="[537,594,921,942]" name="Egypt" pageId="2" pageNumber="367">Egypt</collectingCountry>
,
<collectorName id="845482C59348FFA4FDD8A7EDFCB0F2D5" box="[607,840,920,942]" pageId="2" pageNumber="367">
W
<location id="2C7EB1C89348FFA4FDFAA7EDFD08F2D5" LSID="urn:lsid:plazi:treatment:A10856059348FFA2FF24A5F5FC2EF778:2C7EB1C89348FFA4FDFAA7EDFD08F2D5" box="[637,752,920,942]" country="Egypt" latitude="28.33" longLatPrecision="740" longitude="29.0" municipality="Bahariva Oases" name="Desert Oasis" pageId="2" pageNumber="367" stateProvince="Al Qahirah">Desert Oasis</location>
Bahariva
</collectorName>
[= Bahariya, ca.
<geoCoordinate id="4C9581D49348FFA4FC6BA7EDFBD6F2D5" box="[1004,1070,921,942]" degrees="28.33" direction="north" orientation="latitude" pageId="2" pageNumber="367" precision="555" value="28.33">28.33N</geoCoordinate>
<geoCoordinate id="4C9581D49348FFA4FF57A7C5FEE9F2BD" box="[208,273,945,966]" degrees="29.00" direction="east" orientation="longitude" pageId="2" pageNumber="367" precision="555" value="29.0">29.00E</geoCoordinate>
, ca.
<quantity id="EE594AF69348FFA4FEB8A7C5FE7FF2BD" box="[319,391,944,966]" metricMagnitude="5" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="367" unit="km" value="300.0">300 km</quantity>
SW
<collectingRegion id="EB6529F19348FFA4FE3DA7C4FE17F2BD" box="[442,495,944,966]" country="Egypt" name="Al Qahirah" pageId="2" pageNumber="367">Cairo</collectingRegion>
],
<date id="5D1FC1D39348FFA4FD87A7C4FD9CF2BD" box="[512,612,944,966]" pageId="2" pageNumber="367" value="1996-04-30">
<collectingDate id="4D5B383B9348FFA4FD87A7C4FD9CF2BD" box="[512,612,944,966]" pageId="2" pageNumber="367" value="1996-04-30">30.IV.1996</collectingDate>
</date>
<collectorName id="845482C59348FFA4FDEAA7C5FD1FF2BD" box="[621,743,944,966]" pageId="2" pageNumber="367">W.G. Ullrich</collectorName>
&quot; [printed] (
<collectionCode id="4FB07FD69348FFA4FCD5A7C5FC7BF2BD" box="[850,899,945,966]" pageId="2" pageNumber="367">CLH</collectionCode>
).
<specimenCount id="3FA72C9A9348FFA4FC13A7C5FC48F2BD" box="[916,944,945,966]" count="1" pageId="2" pageNumber="367" type="female">1♀</specimenCount>
, &quot;
<collectingRegion id="EB6529F19348FFA4FC42A7C4FC04F2BD" box="[965,1020,944,966]" country="Egypt" name="Al Qahirah" pageId="2" pageNumber="367">Kairo</collectingRegion>
&quot; [hw
<collectorName id="845482C59348FFA4FF57A7BCFEB4F2A5" box="[208,332,968,990]" pageId="2" pageNumber="367">Zimmermann</collectorName>
], &quot;
<collectorName id="845482C59348FFA4FEE7A7BCFDBCF2A5" box="[352,580,968,990]" pageId="2" pageNumber="367">Samml. A. Zimmermann</collectorName>
&quot; [printed], &quot;
<emphasis id="1BD53B019348FFA4FD37A7BCFCEAF2A6" box="[688,786,968,989]" italics="true" pageId="2" pageNumber="367">
<collectorName id="845482C59348FFA4FD37A7BCFCEAF2A6" box="[688,786,968,989]" pageId="2" pageNumber="367">Coelambus</collectorName>
</emphasis>
,
<taxonomicName id="EEA19C909348FFA4FC98A7BCFC2CF2A5" authority=", (SCHAUM)" baseAuthorityName="SCHAUM" box="[799,980,968,990]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="367" phylum="Arthropoda" rank="species" species="lernaeus">
<emphasis id="1BD53B019348FFA4FC98A7BCFC92F2A6" box="[799,874,968,989]" italics="true" pageId="2" pageNumber="367">lernaeus</emphasis>
, (
<smallCapsWord id="2FF871CF9348FFA4FCFBA7BDFC35F2A6" baselines="985,985" box="[892,973,969,990]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Schaum" pageId="2" pageNumber="367">SCHAUM</smallCapsWord>
)
</taxonomicName>
,
<determinerName id="DCC20BD79348FFA4FC58A7BDFBFFF2A5" box="[991,1031,969,990]" pageId="2" pageNumber="367">Fery</determinerName>
det. 19[91] [hw
<collectorName id="845482C59348FFA4FEC5A795FE96F28D" box="[322,366,993,1014]" pageId="2" pageNumber="367">Fery</collectorName>
], &quot;SNSB-ZSM
<taxonomicName id="EEA19C909348FFA4FD87A794FD4BF28D" authority=", ID No." authorityName="ID No." box="[512,691,992,1014]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="367" phylum="Arthropoda" rank="order">Coleoptera, ID No.</taxonomicName>
000124&quot; [printed] (
<collectionCode id="4FB07FD69348FFA4FCE9A795FC58F28D" box="[878,928,993,1014]" country="Germany" name="Bavarian State Collection of Zoology" pageId="2" pageNumber="367" type="Museum">ZSM</collectionCode>
).
<collectorName id="845482C59348FFA4FC33A794FC28F28D" box="[948,976,992,1014]" pageId="2" pageNumber="367">All</collectorName>
<typeStatus id="F61A59B19348FFA4FC5DA795FBD6F28D" box="[986,1070,993,1014]" pageId="2" pageNumber="367" type="paratype">paratypes</typeStatus>
provided with the respective red label
</materialsCitation>
.
</paragraph>
<paragraph id="291EE7139348FFA4FF25A069FD71F549" blockId="2.[162,649,1053,1075]" box="[162,649,1053,1075]" pageId="2" pageNumber="367">D e s c r i p t i o n o f t h e h o l o t y p e:</paragraph>
<paragraph id="291EE7139348FFA4FF24A037FEA5F5B6" blockId="2.[163,1070,1090,1230]" pageId="2" pageNumber="367">
H a b i t u s elongate oval, sides evenly rounded (
<figureCitation id="B19AFB969348FFA4FD3FA036FD0DF52C" box="[696,757,1090,1112]" captionStart="Fig" captionStartId="9.[163,195,1508,1530]" captionTargetBox="[164,1067,201,1493]" captionTargetId="figure-23@9.[164,1067,201,1493]" captionTargetPageId="9" captionText="Fig. 1: Habitus of Hygrotus (Leptolambus) natrun nov.sp. and labels (holotype from Wadi El Natrun, Egypt)." figureDoi="http://doi.org/10.5281/zenodo.14681677" httpUri="https://zenodo.org/record/14681677/files/figure.png" pageId="2" pageNumber="367">Fig. 1</figureCitation>
), body outline in dorsal view without distinct discontinuity at shoulders. Maximum width near midlength of body. Elytra brownish yellow, with blackish longitudinal vittae. Head and pronotum more reddish yellow, with brownish spots. Entire dorsal surface shiny although weakly reticulated in part.
</paragraph>
<paragraph id="291EE7139348FFA5FF24A0A9FE8DF078" blockId="2.[163,1071,1245,1560]" lastBlockId="3.[163,1069,208,259]" lastPageId="3" lastPageNumber="368" pageId="2" pageNumber="367">H e a d: Anterior margin medially almost straight, but appearing semicircular if observed in perpendicular view on entire body of specimens because anterior margin not visible; anteriorly between eyes with two clypeal grooves. Whole surface finely reticulated, but not matt. Third and fourth antennomeres equal in length, thinner and shorter than first and second as well as rest of them. Punctation on clypeus fine and sparse, diameter of punctures that of one mesh, distance between punctures that of about 5 meshes; on frons punctures larger, diameter that of about three meshes, distance that of about three meshes; on vertex punctures again smaller and sparser; in clypeal groves punctures very dense; line of coarse punctures alongside inner margin of eyes. Clypeus with two ill-defined, brownish interocular spots; above insertions of antennae (between clypeal grooves and anterior margin) shining through brownish; vertex dark brownish. Antennae reddish-yellowish with antennomers progressively darkened distally, beginning with fifth.</paragraph>
<paragraph id="291EE7139349FFA5FF24A567FDEDF368" blockId="3.[163,1093,274,531]" pageId="3" pageNumber="368">P r o n o t u m: Sides slightly rounded, with distinct brownish rim; posterior angles more or less right-angled. Punctation on disc rather fine (as on clypeus), becoming coarser laterally; centre of disc with one small longitudinal puncture; directly behind anterior margin punctation very fine and sparse, but then before disc very coarse; before posterior margin again very coarse; near lateral margins punctation much denser. Few coarser punctures near margin with extremely short seta. Surface of pronotum smooth and shiny, but directly behind anterior margin distinctly reticulated and matt. Disc with one large ill-defined, brownish spot; anterior margin appearing dark brownish because dark vertex of head shining through.</paragraph>
<paragraph id="291EE7139349FFA5FF24A657FD66F550" blockId="3.[162,1070,546,1067]" pageId="3" pageNumber="368">E l y t r a: In dorsal view lateral margin of elytron visible only near elytral base and before apex because else sides of elytra turned inwards. Epipleural rim in lateral view weakly ascending towards humeral angle. Surface smooth in anterior two thirds, behind reticulated, but not distinctly matt. Punctation double; coarse punctures interspersed with small punctures; coarse punctures progressively becoming more numerous posteriad, but near apex punctures smaller. Each elytron in anterior half with two indistinct puncture lines, which coincide more or less with second and fourth vittae; in anterior quarter between fifth vitta and elytral margin some larger punctures indicating a third line; no sutural line perceptible. Most coarser punctures with very short seta, almost imperceptible on disc, otherwise slightly longer; in posterior half a very few punctures with a longer seta; however, generally setation only well visible if adequately illuminated. Each elytron with suture, four discal vittae, a lateral and a submarginal vitta blackish; elytral base medially with blackish stripe. Second vitta reaching base, fourth vitta ending just before base, first and third vittae ending far before base; lateral vitta short, only present in anterior half, and ending far before base; submarginal vitta present only in posterior half, more or less parallel to elytral margin, reaching until apex, and here fused with sutural line. First, second and third vittae fused just before apex, third and fourth vittae short behind midlength of elytra.</paragraph>
<paragraph id="291EE7139349FFA2FF24A04FFCE0F370" blockId="3.[163,1071,1082,1574]" lastBlockId="4.[163,1070,208,523]" lastPageId="4" lastPageNumber="369" pageId="3" pageNumber="368">
V e n t r a l s u r f a c e: Head, prosternum, epipleura and legs light brownish, metaventrite and abdomen predominantly black; metacoxal processes apically, neighbouring small parts of metacoxal plates, first abdominal ventrite, hind margins of third to fifth abdominal ventrites and sixth abdominal ventrite apically shining through brownish. Almost entire venter reticulated, only slightly shining, but not distinctly matt; metaventrite medially without reticulation and here more shiny; small brownish parts of metacoxal plates neighbouring apex of metacoxal processes smooth and shiny. Punctation on metacoxal plates and metaventrite distinct; on metaventrite punctures somewhat larger; punctures absent in small posterior part of metacoxal plates and on areas right and left of medial part of metaventrite; medial part of metaventrite with irregular lines of small punctures; metacoxal processes with sparse small punctures. Abdominal ventrites generally punctured; punctures progressively becoming smaller to last ventrite; second and third ventrites medially fused and here with only very few small punctures. Third to fifth ventrites medially with few densely arranged punctures, these with long and backwardly directed setae. Prosternal process (
<figureCitation id="B19AFB969349FFA5FCE6A1A1FC50F491" box="[865,936,1493,1515]" captionStart="Figs 2-3" captionStartId="10.[163,203,1041,1063]" captionTargetBox="[161,1066,197,1027]" captionTargetId="figure-29@10.[161,1066,197,1027]" captionTargetPageId="10" captionText="Figs 2-3: Prosternal process. (2) Hygrotus (Leptolambus) natrun nov.sp.; (3) Hygrotus (L.) orthogrammus (SHARP, 1882)." figureDoi="http://doi.org/10.5281/zenodo.14681679" httpUri="https://zenodo.org/record/14681679/files/figure.png" pageId="3" pageNumber="368">Fig. 2</figureCitation>
) lanceolate, longitudinally carinate, with tectiform cross section; apex rounded, sides slightly flattened and here with some punctures, lateral margin rimmed (for comparison see flat process of
<emphasis id="1BD53B01934EFFA2FE92A4A4FEDFF19E" box="[277,295,208,229]" italics="true" pageId="4" pageNumber="369">H</emphasis>
.
<taxonomicName id="EEA19C90934EFFA2FEB1A4A4FE01F19E" baseAuthorityName="SHARP" baseAuthorityYear="1882" box="[310,505,208,229]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="369" phylum="Arthropoda" rank="species" species="orthogrammus" subGenus="Leptolambus">
(
<emphasis id="1BD53B01934EFFA2FEB8A4A4FEB5F19E" box="[319,333,208,229]" italics="true" pageId="4" pageNumber="369">L</emphasis>
.)
<emphasis id="1BD53B01934EFFA2FEE3A4A4FE01F19E" box="[356,505,208,229]" italics="true" pageId="4" pageNumber="369">orthogrammus</emphasis>
</taxonomicName>
in
<figureCitation id="B19AFB96934EFFA2FD99A4A4FDA5F19E" box="[542,605,208,230]" captionStart="Figs 2-3" captionStartId="10.[163,203,1041,1063]" captionTargetBox="[161,1066,197,1027]" captionTargetId="figure-29@10.[161,1066,197,1027]" captionTargetPageId="10" captionText="Figs 2-3: Prosternal process. (2) Hygrotus (Leptolambus) natrun nov.sp.; (3) Hygrotus (L.) orthogrammus (SHARP, 1882)." figureDoi="http://doi.org/10.5281/zenodo.14681679" httpUri="https://zenodo.org/record/14681679/files/figure.png" pageId="4" pageNumber="369">Fig. 3</figureCitation>
); prosternal column coarsely punctured with small transverse elevation between anterior coxae; medial part of prosternum (before column) vaulted and very coarsely sculptured. Epipleura in anterior third more or less as broad as apex of mesotibia, with some punctures and extremely weak reticulation; evenly narrowed behind; humeral part of epipleura with oblique carina delimiting genicular fossa; carina forming with epipleural rim an angle of about 25° (see details in
<bibRefCitation id="4D309AE2934EFFA2FF24A5F4FE36F0EE" author="VILLASTRIGO A. &amp; RIBERA I. &amp; MANUEL M. &amp; MILLAN A." box="[163,462,384,406]" pageId="4" pageNumber="369" pagination="499 - 529" refId="ref6430" refString="VILLASTRIGO A., RIBERA I., MANUEL M., MILLAN A. &amp; H. FERY (2017): A new classification of the tribe Hygrotini PORTEVIN, 1929 (Coleoptera: Dytiscidae: Hydroporinae). - Zootaxa 4317 (3): 499 - 529." type="journal article" year="2017">VILLASTRIGO et al. (2017))</bibRefCitation>
. Hind margins of metacoxal processes slightly slanted, interlaminary bridge well exposed, forming a more or less triangular (or wing-like shaped) elevation on first and second abdominal ventrites (see details in FERY &amp; RIBERA (2018)). Metacoxal lines diverging forwards, ending before hind margin of metaventrite. Protarsal claws thin, evenly curved and more or less identical.
</paragraph>
<paragraph id="291EE713934EFFA2FF24A66EFD82F3B9" blockId="4.[163,1070,538,707]" pageId="4" pageNumber="369">
G e n i t a l i a: Gonocoxae in ventral view as in
<figureCitation id="B19AFB96934EFFA2FD1BA66EFD21F354" box="[668,729,538,560]" captionStart="Figs 4-12" captionStartId="11.[163,203,1416,1438]" captionTargetBox="[166,1068,194,1403]" captionTargetId="figure-63@11.[164,1068,194,1403]" captionTargetPageId="11" captionText="Figs 4-12: Gonocoxae in ventral and lateral view and gonocoxosternum. (4-6) Hygrotus (Leptolambus) natrun nov.sp.; (7-9) Hygrotus (L.) lernaeus (SCHAUM, 1857); (10-12) Hygrotus (L.) orthogrammus (SHARP, 1882)." figureDoi="http://doi.org/10.5281/zenodo.14681681" httpUri="https://zenodo.org/record/14681681/files/figure.png" pageId="4" pageNumber="369">Fig. 4</figureCitation>
; in lateral view (
<figureCitation id="B19AFB96934EFFA2FC03A66EFC39F34B" box="[900,961,538,560]" captionStart="Figs 4-12" captionStartId="11.[163,203,1416,1438]" captionTargetBox="[166,1068,194,1403]" captionTargetId="figure-63@11.[164,1068,194,1403]" captionTargetPageId="11" captionText="Figs 4-12: Gonocoxae in ventral and lateral view and gonocoxosternum. (4-6) Hygrotus (Leptolambus) natrun nov.sp.; (7-9) Hygrotus (L.) lernaeus (SCHAUM, 1857); (10-12) Hygrotus (L.) orthogrammus (SHARP, 1882)." figureDoi="http://doi.org/10.5281/zenodo.14681681" httpUri="https://zenodo.org/record/14681681/files/figure.png" pageId="4" pageNumber="369">Fig. 5</figureCitation>
) distinctly flatter than in
<emphasis id="1BD53B01934EFFA2FEB2A64CFEB6F336" box="[309,334,568,589]" italics="true" pageId="4" pageNumber="369">H.</emphasis>
<taxonomicName id="EEA19C90934EFFA2FED2A64CFE21F336" box="[341,473,568,589]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="369" phylum="Arthropoda" rank="species" species="lernaeus" subGenus="Leptolambus">
(
<emphasis id="1BD53B01934EFFA2FED9A64CFE94F336" box="[350,364,568,589]" italics="true" pageId="4" pageNumber="369">L</emphasis>
.)
<emphasis id="1BD53B01934EFFA2FE05A64CFE21F336" box="[386,473,568,589]" italics="true" pageId="4" pageNumber="369">lernaeus</emphasis>
</taxonomicName>
(
<figureCitation id="B19AFB96934EFFA2FE6FA64CFDDFF336" box="[488,551,568,590]" captionStart="Figs 4-12" captionStartId="11.[163,203,1416,1438]" captionTargetBox="[166,1068,194,1403]" captionTargetId="figure-63@11.[164,1068,194,1403]" captionTargetPageId="11" captionText="Figs 4-12: Gonocoxae in ventral and lateral view and gonocoxosternum. (4-6) Hygrotus (Leptolambus) natrun nov.sp.; (7-9) Hygrotus (L.) lernaeus (SCHAUM, 1857); (10-12) Hygrotus (L.) orthogrammus (SHARP, 1882)." figureDoi="http://doi.org/10.5281/zenodo.14681681" httpUri="https://zenodo.org/record/14681681/files/figure.png" pageId="4" pageNumber="369">Fig. 8</figureCitation>
); proximal portion less curved upwards than in
<emphasis id="1BD53B01934EFFA2FB92A64CFBDFF336" box="[1045,1063,568,589]" italics="true" pageId="4" pageNumber="369">H</emphasis>
.
<taxonomicName id="EEA19C90934EFFA2FF24A621FED1F311" box="[163,297,597,619]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="369" phylum="Arthropoda" rank="species" species="lernaeus" subGenus="Leptolambus">
(
<emphasis id="1BD53B01934EFFA2FF2CA622FF41F310" box="[171,185,598,619]" italics="true" pageId="4" pageNumber="369">L</emphasis>
.)
<emphasis id="1BD53B01934EFFA2FF55A621FED1F311" box="[210,297,597,618]" italics="true" pageId="4" pageNumber="369">lernaeus</emphasis>
</taxonomicName>
(
<figureCitation id="B19AFB96934EFFA2FEBDA621FE63F311" box="[314,411,597,619]" captionStart="Figs 4-12" captionStartId="11.[163,203,1416,1438]" captionTargetBox="[166,1068,194,1403]" captionTargetId="figure-63@11.[164,1068,194,1403]" captionTargetPageId="11" captionText="Figs 4-12: Gonocoxae in ventral and lateral view and gonocoxosternum. (4-6) Hygrotus (Leptolambus) natrun nov.sp.; (7-9) Hygrotus (L.) lernaeus (SCHAUM, 1857); (10-12) Hygrotus (L.) orthogrammus (SHARP, 1882)." figureDoi="http://doi.org/10.5281/zenodo.14681681" httpUri="https://zenodo.org/record/14681681/files/figure.png" pageId="4" pageNumber="369">Figs 7, 8</figureCitation>
), but more than in
<emphasis id="1BD53B01934EFFA2FDEAA622FD87F310" box="[621,639,598,619]" italics="true" pageId="4" pageNumber="369">H</emphasis>
.
<taxonomicName id="EEA19C90934EFFA2FD08A621FCABF311" baseAuthorityName="SHARP" baseAuthorityYear="1882" box="[655,851,597,619]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="369" phylum="Arthropoda" rank="species" species="orthogrammus" subGenus="Leptolambus">
(
<emphasis id="1BD53B01934EFFA2FD1FA622FD5EF310" box="[664,678,598,619]" italics="true" pageId="4" pageNumber="369">L</emphasis>
.)
<emphasis id="1BD53B01934EFFA2FD39A621FCABF311" box="[702,851,597,618]" italics="true" pageId="4" pageNumber="369">orthogrammus</emphasis>
</taxonomicName>
(
<figureCitation id="B19AFB96934EFFA2FCE3A621FC27F311" box="[868,991,597,619]" captionStart="Figs 4-12" captionStartId="11.[163,203,1416,1438]" captionTargetBox="[166,1068,194,1403]" captionTargetId="figure-63@11.[164,1068,194,1403]" captionTargetPageId="11" captionText="Figs 4-12: Gonocoxae in ventral and lateral view and gonocoxosternum. (4-6) Hygrotus (Leptolambus) natrun nov.sp.; (7-9) Hygrotus (L.) lernaeus (SCHAUM, 1857); (10-12) Hygrotus (L.) orthogrammus (SHARP, 1882)." figureDoi="http://doi.org/10.5281/zenodo.14681681" httpUri="https://zenodo.org/record/14681681/files/figure.png" pageId="4" pageNumber="369">Figs 10, 11</figureCitation>
); distal margins slightly concavely emarginate (see arrow in
<figureCitation id="B19AFB96934EFFA2FD3AA606FD03F3FC" box="[701,763,626,648]" captionStart="Figs 4-12" captionStartId="11.[163,203,1416,1438]" captionTargetBox="[166,1068,194,1403]" captionTargetId="figure-63@11.[164,1068,194,1403]" captionTargetPageId="11" captionText="Figs 4-12: Gonocoxae in ventral and lateral view and gonocoxosternum. (4-6) Hygrotus (Leptolambus) natrun nov.sp.; (7-9) Hygrotus (L.) lernaeus (SCHAUM, 1857); (10-12) Hygrotus (L.) orthogrammus (SHARP, 1882)." figureDoi="http://doi.org/10.5281/zenodo.14681681" httpUri="https://zenodo.org/record/14681681/files/figure.png" pageId="4" pageNumber="369">Fig. 4</figureCitation>
). Gonocoxosternum as in
<figureCitation id="B19AFB96934EFFA2FB82A606FF4AF3DE" captionStart="Figs 4-12" captionStartId="11.[163,203,1416,1438]" captionTargetBox="[166,1068,194,1403]" captionTargetId="figure-63@11.[164,1068,194,1403]" captionTargetPageId="11" captionText="Figs 4-12: Gonocoxae in ventral and lateral view and gonocoxosternum. (4-6) Hygrotus (Leptolambus) natrun nov.sp.; (7-9) Hygrotus (L.) lernaeus (SCHAUM, 1857); (10-12) Hygrotus (L.) orthogrammus (SHARP, 1882)." figureDoi="http://doi.org/10.5281/zenodo.14681681" httpUri="https://zenodo.org/record/14681681/files/figure.png" pageId="4" pageNumber="369">Fig. 6</figureCitation>
; differences to those of
<emphasis id="1BD53B01934EFFA2FE36A6E4FE32F3DE" box="[433,458,656,677]" italics="true" pageId="4" pageNumber="369">H.</emphasis>
<taxonomicName id="EEA19C90934EFFA2FE55A6E4FDA0F3DE" box="[466,600,656,677]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="369" phylum="Arthropoda" rank="species" species="lernaeus" subGenus="Leptolambus">
(
<emphasis id="1BD53B01934EFFA2FE5CA6E4FE11F3DE" box="[475,489,656,677]" italics="true" pageId="4" pageNumber="369">L</emphasis>
.)
<emphasis id="1BD53B01934EFFA2FD86A6E4FDA0F3DE" box="[513,600,656,677]" italics="true" pageId="4" pageNumber="369">lernaeus</emphasis>
</taxonomicName>
(
<figureCitation id="B19AFB96934EFFA2FDEFA6E4FD5FF3DE" box="[616,679,656,678]" captionStart="Figs 4-12" captionStartId="11.[163,203,1416,1438]" captionTargetBox="[166,1068,194,1403]" captionTargetId="figure-63@11.[164,1068,194,1403]" captionTargetPageId="11" captionText="Figs 4-12: Gonocoxae in ventral and lateral view and gonocoxosternum. (4-6) Hygrotus (Leptolambus) natrun nov.sp.; (7-9) Hygrotus (L.) lernaeus (SCHAUM, 1857); (10-12) Hygrotus (L.) orthogrammus (SHARP, 1882)." figureDoi="http://doi.org/10.5281/zenodo.14681681" httpUri="https://zenodo.org/record/14681681/files/figure.png" pageId="4" pageNumber="369">Fig. 9</figureCitation>
) and
<emphasis id="1BD53B01934EFFA2FD62A6E4FD0FF3DE" box="[741,759,656,677]" italics="true" pageId="4" pageNumber="369">H</emphasis>
.
<taxonomicName id="EEA19C90934EFFA2FC80A6E4FC32F3DE" baseAuthorityName="SHARP" baseAuthorityYear="1882" box="[775,970,656,677]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="369" phylum="Arthropoda" rank="species" species="orthogrammus" subGenus="Leptolambus">
(
<emphasis id="1BD53B01934EFFA2FC88A6E4FCE5F3DE" box="[783,797,656,677]" italics="true" pageId="4" pageNumber="369">L</emphasis>
.)
<emphasis id="1BD53B01934EFFA2FCB2A6E4FC32F3DE" box="[821,970,656,677]" italics="true" pageId="4" pageNumber="369">orthogrammus</emphasis>
</taxonomicName>
(
<figureCitation id="B19AFB96934EFFA2FC5DA6E4FBDEF3DE" box="[986,1062,656,678]" captionStart="Figs 4-12" captionStartId="11.[163,203,1416,1438]" captionTargetBox="[166,1068,194,1403]" captionTargetId="figure-63@11.[164,1068,194,1403]" captionTargetPageId="11" captionText="Figs 4-12: Gonocoxae in ventral and lateral view and gonocoxosternum. (4-6) Hygrotus (Leptolambus) natrun nov.sp.; (7-9) Hygrotus (L.) lernaeus (SCHAUM, 1857); (10-12) Hygrotus (L.) orthogrammus (SHARP, 1882)." figureDoi="http://doi.org/10.5281/zenodo.14681681" httpUri="https://zenodo.org/record/14681681/files/figure.png" pageId="4" pageNumber="369">Fig. 12</figureCitation>
) present, but irrelevant for diagnostic purposes.
</paragraph>
<paragraph id="291EE713934EFFA2FF24A6A7FE9EF2CE" blockId="4.[163,1070,722,950]" pageId="4" pageNumber="369">
M a l e s: Externally similar to females, but pro- and mesotarsi broader and anterior (inner) claw of protarsi strongly enlarged and curved (
<figureCitation id="B19AFB96934EFFA2FD49A684FCE1F27E" box="[718,793,752,774]" captionStart="Figs 13-22" captionStartId="12.[163,203,1387,1409]" captionTargetBox="[167,1073,199,1377]" captionTargetId="figure-86@12.[167,1074,198,1377]" captionTargetPageId="12" captionText="Figs 13-22: Median lobe in ventral and lateral view (13-18) and left paramere (19-21). (13, 14, 19) Hygrotus (Leptolambus) natrun nov.sp.; (15, 16, 20) Hygrotus (L.) lernaeus (SCHAUM, 1857); (17, 18, 21) Hygrotus (L.) orthogrammus (SHARP, 1882) (18a: detail of apex). Fig. 22: Claws of left protarsus of male H. (L.) natrun nov.sp. (anterior claw in front)." figureDoi="http://doi.org/10.5281/zenodo.14681683" httpUri="https://zenodo.org/record/14681683/files/figure.png" pageId="4" pageNumber="369">Fig. 22</figureCitation>
). Median lobe of aedeagus in ventral and lateral view as in
<figureCitation id="B19AFB96934EFFA2FE7DA779FD63F259" box="[506,667,781,803]" captionStart="Figs 13-22" captionStartId="12.[163,203,1387,1409]" captionTargetBox="[167,1073,199,1377]" captionTargetId="figure-86@12.[167,1074,198,1377]" captionTargetPageId="12" captionText="Figs 13-22: Median lobe in ventral and lateral view (13-18) and left paramere (19-21). (13, 14, 19) Hygrotus (Leptolambus) natrun nov.sp.; (15, 16, 20) Hygrotus (L.) lernaeus (SCHAUM, 1857); (17, 18, 21) Hygrotus (L.) orthogrammus (SHARP, 1882) (18a: detail of apex). Fig. 22: Claws of left protarsus of male H. (L.) natrun nov.sp. (anterior claw in front)." figureDoi="http://doi.org/10.5281/zenodo.14681683" httpUri="https://zenodo.org/record/14681683/files/figure.png" pageId="4" pageNumber="369">Figs 13 and 14</figureCitation>
; apex simple, not bulged as in
<emphasis id="1BD53B01934EFFA2FC61A77AFBF8F258" box="[998,1024,782,803]" italics="true" pageId="4" pageNumber="369">H.</emphasis>
<taxonomicName id="EEA19C90934EFFA2FB8FA779FEC0F244" baseAuthorityName="SHARP" baseAuthorityYear="1882" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="369" phylum="Arthropoda" rank="species" species="orthogrammus" subGenus="Leptolambus">
(
<emphasis id="1BD53B01934EFFA2FB96A77AFBE7F258" box="[1041,1055,782,803]" italics="true" pageId="4" pageNumber="369">L</emphasis>
.)
<emphasis id="1BD53B01934EFFA2FF24A75EFEC0F244" box="[163,312,810,831]" italics="true" pageId="4" pageNumber="369">orthogrammus</emphasis>
</taxonomicName>
(
<figureCitation id="B19AFB96934EFFA2FEC1A75EFE14F244" box="[326,492,810,832]" captionStart="Figs 13-22" captionStartId="12.[163,203,1387,1409]" captionTargetBox="[167,1073,199,1377]" captionTargetId="figure-86@12.[167,1074,198,1377]" captionTargetPageId="12" captionText="Figs 13-22: Median lobe in ventral and lateral view (13-18) and left paramere (19-21). (13, 14, 19) Hygrotus (Leptolambus) natrun nov.sp.; (15, 16, 20) Hygrotus (L.) lernaeus (SCHAUM, 1857); (17, 18, 21) Hygrotus (L.) orthogrammus (SHARP, 1882) (18a: detail of apex). Fig. 22: Claws of left protarsus of male H. (L.) natrun nov.sp. (anterior claw in front)." figureDoi="http://doi.org/10.5281/zenodo.14681683" httpUri="https://zenodo.org/record/14681683/files/figure.png" pageId="4" pageNumber="369">Figs 18 and 18a</figureCitation>
); left paramere as in
<figureCitation id="B19AFB96934EFFA2FD44A75EFCF6F244" box="[707,782,810,832]" captionStart="Figs 13-22" captionStartId="12.[163,203,1387,1409]" captionTargetBox="[167,1073,199,1377]" captionTargetId="figure-86@12.[167,1074,198,1377]" captionTargetPageId="12" captionText="Figs 13-22: Median lobe in ventral and lateral view (13-18) and left paramere (19-21). (13, 14, 19) Hygrotus (Leptolambus) natrun nov.sp.; (15, 16, 20) Hygrotus (L.) lernaeus (SCHAUM, 1857); (17, 18, 21) Hygrotus (L.) orthogrammus (SHARP, 1882) (18a: detail of apex). Fig. 22: Claws of left protarsus of male H. (L.) natrun nov.sp. (anterior claw in front)." figureDoi="http://doi.org/10.5281/zenodo.14681683" httpUri="https://zenodo.org/record/14681683/files/figure.png" pageId="4" pageNumber="369">Fig. 19</figureCitation>
. Medial part of median lobe in lateral view significantly thinner (see arrow in
<figureCitation id="B19AFB96934EFFA2FD39A73CFCF3F226" box="[702,779,840,862]" captionStart="Figs 13-22" captionStartId="12.[163,203,1387,1409]" captionTargetBox="[167,1073,199,1377]" captionTargetId="figure-86@12.[167,1074,198,1377]" captionTargetPageId="12" captionText="Figs 13-22: Median lobe in ventral and lateral view (13-18) and left paramere (19-21). (13, 14, 19) Hygrotus (Leptolambus) natrun nov.sp.; (15, 16, 20) Hygrotus (L.) lernaeus (SCHAUM, 1857); (17, 18, 21) Hygrotus (L.) orthogrammus (SHARP, 1882) (18a: detail of apex). Fig. 22: Claws of left protarsus of male H. (L.) natrun nov.sp. (anterior claw in front)." figureDoi="http://doi.org/10.5281/zenodo.14681683" httpUri="https://zenodo.org/record/14681683/files/figure.png" pageId="4" pageNumber="369">Fig. 14</figureCitation>
) than in other two species (compare
<figureCitation id="B19AFB96934EFFA2FE89A711FE40F201" box="[270,440,869,891]" captionStart="Figs 13-22" captionStartId="12.[163,203,1387,1409]" captionTargetBox="[167,1073,199,1377]" captionTargetId="figure-86@12.[167,1074,198,1377]" captionTargetPageId="12" captionText="Figs 13-22: Median lobe in ventral and lateral view (13-18) and left paramere (19-21). (13, 14, 19) Hygrotus (Leptolambus) natrun nov.sp.; (15, 16, 20) Hygrotus (L.) lernaeus (SCHAUM, 1857); (17, 18, 21) Hygrotus (L.) orthogrammus (SHARP, 1882) (18a: detail of apex). Fig. 22: Claws of left protarsus of male H. (L.) natrun nov.sp. (anterior claw in front)." figureDoi="http://doi.org/10.5281/zenodo.14681683" httpUri="https://zenodo.org/record/14681683/files/figure.png" pageId="4" pageNumber="369">Figs 16 and 18</figureCitation>
); corrugated membrane (see FOSTER &amp; ANGUS (1985: 8)) extending less far distally (compare
<figureCitation id="B19AFB96934EFFA2FDA6A7F6FD13F2EC" box="[545,747,898,920]" captionStart="Figs 13-22" captionStartId="12.[163,203,1387,1409]" captionTargetBox="[167,1073,199,1377]" captionTargetId="figure-86@12.[167,1074,198,1377]" captionTargetPageId="12" captionText="Figs 13-22: Median lobe in ventral and lateral view (13-18) and left paramere (19-21). (13, 14, 19) Hygrotus (Leptolambus) natrun nov.sp.; (15, 16, 20) Hygrotus (L.) lernaeus (SCHAUM, 1857); (17, 18, 21) Hygrotus (L.) orthogrammus (SHARP, 1882) (18a: detail of apex). Fig. 22: Claws of left protarsus of male H. (L.) natrun nov.sp. (anterior claw in front)." figureDoi="http://doi.org/10.5281/zenodo.14681683" httpUri="https://zenodo.org/record/14681683/files/figure.png" pageId="4" pageNumber="369">Figs 13, 15 and 17</figureCitation>
); for comparison of parameres see
<figureCitation id="B19AFB96934EFFA2FF4EA7D4FE9EF2CE" box="[201,358,928,950]" captionStart="Figs 13-22" captionStartId="12.[163,203,1387,1409]" captionTargetBox="[167,1073,199,1377]" captionTargetId="figure-86@12.[167,1074,198,1377]" captionTargetPageId="12" captionText="Figs 13-22: Median lobe in ventral and lateral view (13-18) and left paramere (19-21). (13, 14, 19) Hygrotus (Leptolambus) natrun nov.sp.; (15, 16, 20) Hygrotus (L.) lernaeus (SCHAUM, 1857); (17, 18, 21) Hygrotus (L.) orthogrammus (SHARP, 1882) (18a: detail of apex). Fig. 22: Claws of left protarsus of male H. (L.) natrun nov.sp. (anterior claw in front)." figureDoi="http://doi.org/10.5281/zenodo.14681683" httpUri="https://zenodo.org/record/14681683/files/figure.png" pageId="4" pageNumber="369">Figs 20 and 21.</figureCitation>
</paragraph>
<paragraph id="291EE713934EFFA2FF24A7B1FD20F28C" blockId="4.[163,1069,965,1016]" pageId="4" pageNumber="369">
M e a s u r e m e n t s: TL:
<quantity id="EE594AF6934EFFA2FE5DA7B1FD95F2A1" box="[474,621,965,987]" metricMagnitude="-3" metricUnit="m" metricValue="3.975" metricValueMax="4.25" metricValueMin="3.7" pageId="4" pageNumber="369" unit="mm" value="3.975" valueMax="4.25" valueMin="3.7">3.7–4.25 mm</quantity>
, MW:
<quantity id="EE594AF6934EFFA2FD42A7B1FCB2F2A1" box="[709,842,965,986]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="4" pageNumber="369" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.8–2.2 mm</quantity>
, TL/MW: 1.93–2.01 (
<typeStatus id="F61A59B1934EFFA2FF2DA796FEF0F283" box="[170,264,994,1016]" pageId="4" pageNumber="369" type="holotype">holotype</typeStatus>
: TL:
<quantity id="EE594AF6934EFFA2FEBAA796FE76F283" box="[317,398,994,1016]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="4" pageNumber="369" unit="mm" value="3.9">3.9 mm</quantity>
, MW:
<quantity id="EE594AF6934EFFA2FE53A796FDD7F283" box="[468,559,994,1016]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" pageId="4" pageNumber="369" unit="mm" value="1.95">1.95 mm</quantity>
, TL/MW: 2.00).
</paragraph>
<paragraph id="291EE713934EFFA2FF24A07CFC62F438" blockId="4.[163,1070,1031,1347]" pageId="4" pageNumber="369">V a r i a b i l i t y: The variation of external morphological characters of the specimens studied is scanty. Worth mentioning are only: the spots on head and pronotum and the elytral vittae can be more blackish; the degree of fusion of the elytral vittae can vary; the small brownish parts on metathorax and abdomen can be darker or even black; the areas with coarser punctures on pronotum can be extended more mediad. The variation in the shape of the median lobe is small, but in some specimens the corrugated membrane extends a little more distally (ventral view). The variation in the shape of the parameres is somewhat greater, particularly because the distal part may be slightly curved inward. This also applies to the other two species and therefore the shape of the parameres is more or less unsuitable for diagnostic purposes. N o t e s: The females studied have a shiny upper surface, but the existence of a matt morph cannot be excluded.</paragraph>
<paragraph id="291EE713934EFFA2FF24A127FC2DF4A5" blockId="4.[163,1070,1362,1502]" pageId="4" pageNumber="369">
D i s t r i b u t i o n: Endemic to
<collectingCountry id="51B6A783934EFFA2FD8CA127FDB3F413" box="[523,587,1363,1384]" name="Egypt" pageId="4" pageNumber="369">Egypt</collectingCountry>
. In
<bibRefCitation id="4D309AE2934EFFA2FDFEA126FCBCF41C" author="ALFIERI A." box="[633,836,1362,1384]" pageId="4" pageNumber="369" pagination="1 - 361" refId="ref5595" refString="ALFIERI A. (1976): The Coleoptera of Egypt. - Memoires de la Societe Entomologique d'Egypte 5: i-xvi, 1 - 361." type="journal article" year="1976">ALFIERI (1976: 33)</bibRefCitation>
is given: &quot;
<emphasis id="1BD53B01934EFFA2FC38A127FEC9F4FE" italics="true" pageId="4" pageNumber="369">
in swamps around
<collectingRegion id="EB6529F1934EFFA2FF71A104FEC9F4FE" box="[246,305,1392,1413]" country="Egypt" name="Al Qahirah" pageId="4" pageNumber="369">Cairo</collectingRegion>
</emphasis>
&quot;. In
<bibRefCitation id="4D309AE2934EFFA2FEEBA104FD91F4FE" author="ZALAT S. &amp; SALEH AHMED R. &amp; ANGUS R. B." box="[364,617,1391,1414]" pageId="4" pageNumber="369" pagination="1 - 107" refId="ref6482" refString="ZALAT S., SALEH AHMED R., ANGUS R. B. &amp; A. KASCHEF (2000): Diving beetles (Coleoptera: Dytiscidae and Noteridae) of Egypt. - Egyptian Journal of Natural History 2: 1 - 107." type="journal article" year="2000">ZALAT et al. (2000: 26)</bibRefCitation>
is given: &quot;
<emphasis id="1BD53B01934EFFA2FD65A104FC1CF4BB" italics="true" pageId="4" pageNumber="369">Nile valley &amp; Delta, eastern &amp; western desert. Months of abundance: April, July-August. Habitat: collected from margins of swamps and marshes of rich vegetation; fresh and brackish water.</emphasis>
&quot; (both under
<taxonomicName id="EEA19C90934EFFA2FF64A1BCFEBCF4A6" box="[227,324,1480,1501]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="369" phylum="Arthropoda" rank="species" species="lernaeus">
&quot;
<emphasis id="1BD53B01934EFFA2FF6AA1BCFEBCF4A6" box="[237,324,1480,1501]" italics="true" pageId="4" pageNumber="369">lernaeus</emphasis>
</taxonomicName>
&quot;). Unfortunately, I was not enabled to study respective material.
</paragraph>
<paragraph id="291EE713934EFFA2FF24A199FC2EF778" blockId="4.[163,982,1517,1539]" box="[163,982,1517,1539]" pageId="4" pageNumber="369">
D e r i v a t i o n o m i n i s: The new species is named after the
<typeStatus id="F61A59B1934EFFA2FCD6A199FC85F779" box="[849,893,1517,1538]" pageId="4" pageNumber="369">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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