136 lines
16 KiB
XML
136 lines
16 KiB
XML
<document id="9150614E76C4F9B311F03CFEC6E7774A" ID-CLB-Dataset="25983" ID-DOI="10.11646/zootaxa.4657.3.1" ID-GBIF-Dataset="3414318d-7570-49ac-9013-be4e1f1e6347" ID-ISSN="1175-5326" ID-Zenodo-Dep="3371886" ID-ZooBank="86DD93B2-E8F4-4174-B105-9436357CB4B6" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1566284110186" checkinUser="plazi" docAuthor="Lambert, Gretchen" docDate="2019" docId="6A2E3761A92AFFDD1390FD1ADFCBFAA9" docLanguage="en" docName="zootaxa.4657.3.1.pdf" docOrigin="Zootaxa 4657 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pyura haustor" docType="treatment" docVersion="11" lastPageNumber="425" masterDocId="96174F19A932FFC51307FFF3DC63FFD6" masterDocTitle="The Ascidiacea collected during the 2017 British Columbia Hakai MarineGEO BioBlitz" masterLastPageNumber="436" masterPageNumber="401" pageNumber="425" updateTime="1698751689998" updateUser="ExternalLinkService">
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<mods:title id="4199A50E28E8A09E9638BE2A233BC8EB">The Ascidiacea collected during the 2017 British Columbia Hakai MarineGEO BioBlitz</mods:title>
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<mods:namePart id="55485FD3296E6F6061105D14BE1D4A59">Lambert, Gretchen</mods:namePart>
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<mods:title id="E3AEF8FB0B339B67459869F8ABAABF33">Zootaxa</mods:title>
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<mods:date id="4F0170A22B1BBFEC01C1C1E926B24AF4">2019</mods:date>
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<mods:number id="3437E342208A772B594AC4F3B3CEE31E">2019-08-20</mods:number>
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<mods:number id="E1EA6750DFCEBD893131293F3DFC4305">4657</mods:number>
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<treatment id="6A2E3761A92AFFDD1390FD1ADFCBFAA9" ID-DOI="http://doi.org/10.5281/zenodo.5941231" ID-GBIF-Taxon="159167942" ID-Zenodo-Dep="5941231" LSID="urn:lsid:plazi:treatment:6A2E3761A92AFFDD1390FD1ADFCBFAA9" httpUri="http://treatment.plazi.org/id/6A2E3761A92AFFDD1390FD1ADFCBFAA9" lastPageNumber="425" pageId="24" pageNumber="425">
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<subSubSection id="AA9DD5FCA92AFFDD1390FD1ADE7DFCD4" box="[151,542,744,771]" pageId="24" pageNumber="425" type="nomenclature">
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<paragraph id="E2388677A92AFFDD1390FD1ADE7DFCD4" blockId="24.[151,542,744,806]" box="[151,542,744,771]" pageId="24" pageNumber="425">
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<heading id="B970311BA92AFFDD1390FD1ADE7DFCD4" bold="true" box="[151,542,744,771]" fontSize="11" level="1" pageId="24" pageNumber="425" reason="1">
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<taxonomicName id="2587FDF4A92AFFDD1390FD1ADE7DFCD4" authority="(Stimpson, 1864)" baseAuthorityName="Stimpson" baseAuthorityYear="1864" box="[151,542,744,771]" class="Ascidiacea" family="Pyuridae" genus="Pyura" kingdom="Animalia" order="Pleurogona" pageId="24" pageNumber="425" phylum="Chordata" rank="species" species="haustor">
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<emphasis id="D0F35A65A92AFFDD1390FD1ADE7DFCD4" bold="true" box="[151,542,744,771]" pageId="24" pageNumber="425">
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<emphasis id="D0F35A65A92AFFDD1390FD1ADD26FCD5" bold="true" box="[151,325,744,771]" italics="true" pageId="24" pageNumber="425">Pyura haustor</emphasis>
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(
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<bibRefCitation id="8616FB86A92AFFDD1253FD1BDE75FCD4" author="Stimpson, W." box="[340,534,744,770]" pageId="24" pageNumber="425" pagination="153 - 161" refId="ref17941" refString="Stimpson, W. (1864) Description of new species of marine Invertebrata from Puget Sound, collected by the naturalists of the North-west Boundary Commission. Proceedings of the Academy of Natural Sciences of Philadelphia, 16, 153 - 161. https: // doi. org / 10.5962 / bhl. title. 5972 0" type="journal article" year="1864">Stimpson, 1864</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="AA9DD5FCA92AFFDD1390FCFFDFCBFAA9" pageId="24" pageNumber="425" type="description">
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<paragraph id="E2388677A92AFFDD1390FCFFDD74FCF0" blockId="24.[151,542,744,806]" box="[151,279,780,806]" pageId="24" pageNumber="425">
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<figureCitation id="7ABC9AF2A92AFFDD1390FCFFDD74FCF0" box="[151,279,780,806]" captionStart="FIGURE 12" captionStartId="25.[151,250,1266,1291]" captionTargetBox="[151,1435,417,1247]" captionTargetId="figure@25.[151,1435,417,1247]" captionTargetPageId="25" captionText="FIGURE 12.A–D: Pyuridae; E, F: Molgulidae.A: Halocynthia aurantium 10 cm in length; B: Halocynthia igaboja; C: Boltenia villosa; D: Pyura haustor; E: Molgula pacifica whole animal left side, tunic removed; F: M. pacifica close-up of siphons. Scale bars: B, 5 mm; C, 5 mm; D, 5 mm; E, 1.6 mm." httpUri="https://zenodo.org/record/3371910/files/figure.png" pageId="24" pageNumber="425">Figure 12D</figureCitation>
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</paragraph>
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<paragraph id="E2388677A92AFFDD1390FCA3D97CFCBF" blockId="24.[151,1311,848,903]" box="[151,1311,848,873]" pageId="24" pageNumber="425">IHAK 12 BHAK 0610, 0611, 0612 UF 2467, 2468, 2469. Common in the low intertidal under rocks by lab, small.</paragraph>
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<paragraph id="E2388677A92AFFDD1390FC9CD805FC50" blockId="24.[151,1311,848,903]" box="[151,1126,878,903]" pageId="24" pageNumber="425">
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<materialsCitation id="52EF8C2AA92AFFDD1390FC9CD803FC51" ID-GBIF-Occurrence="2336841838" box="[151,1120,878,903]" collectionCode="IHAK" county="Scuba" elevation="8" location="Triquet Island Macro" municipality="Triquet Island" pageId="24" pageNumber="425" specimenCode="IHAK 31" specimenCount="8">
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<specimenCode id="B2212E0CA92AFFDD1390FC9CDC95FC51" box="[151,246,879,903]" collectionCode="IHAK" pageId="24" pageNumber="425">IHAK 31</specimenCode>
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BHAK 1696 UF 2520.
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<location id="E758D0ACA92AFFDD12EEFC9DDEDCFC50" LSID="urn:lsid:plazi:treatment:6A2E3761A92AFFDD1390FD1ADFCBFAA9:E758D0ACA92AFFDD12EEFC9DDEDCFC50" box="[489,703,878,903]" county="Scuba" municipality="Triquet Island" name="Triquet Island Macro" pageId="24" pageNumber="425">
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<collectingMunicipality id="025C1C0DA92AFFDD12EEFC9DDE14FC51" box="[489,631,878,903]" pageId="24" pageNumber="425">Triquet Island</collectingMunicipality>
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Macro
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</location>
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site,
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<collectingCounty id="0B59FEFBA92AFFDD11F1FC9DDF55FC51" box="[758,822,878,903]" pageId="24" pageNumber="425">Scuba</collectingCounty>
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,
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<specimenCount id="F4814DFEA92AFFDD1047FC9CDF0DFC50" box="[832,878,879,903]" pageId="24" pageNumber="425" type="generic">
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<quantity id="257F2B92A92AFFDD1047FC9CDF09FC50" box="[832,874,879,903]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="24" pageNumber="425" unit="m" value="8.0">
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<elevation id="69AA6144A92AFFDD1047FC9CDF09FC50" box="[832,874,879,903]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="24" pageNumber="425" unit="m" value="8.0">8 m</elevation>
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</quantity>
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.
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</specimenCount>
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Tiny,
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<collectedFrom id="519BC58AA92AFFDD10AAFC9CD803FC51" box="[941,1120,879,903]" pageId="24" pageNumber="425">
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on
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<taxonomicName id="2587FDF4A92AFFDD10CAFC83D803FC51" authorityName="Dana" authorityYear="1851" box="[973,1120,879,903]" class="Malacostraca" family="Epialtidae" genus="Pugettia" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="425" phylum="Arthropoda" rank="species" species="richii">
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<emphasis id="D0F35A65A92AFFDD10CAFC83D803FC51" box="[973,1120,879,903]" italics="true" pageId="24" pageNumber="425">Pugettia richii</emphasis>
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</taxonomicName>
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</collectedFrom>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="E2388677A92AFFDD1390FC46DDB0FBB6" blockId="24.[151,1437,949,1408]" pageId="24" pageNumber="425">
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This leathery solitary stolidobranch can be very variable in shape, and so confusing to determine if all the variants are the same species that
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<bibRefCitation id="8616FB86A92AFFDD12A8FC2ADE5CFC25" author="Ritter, W. E." box="[431,575,985,1011]" pageId="24" pageNumber="425" pagination="65 - 114" refId="ref17743" refString="Ritter, W. E. (1909) Halocynthia johnsoni n. sp., a comprehensive inquiry as to the extent of law and order that prevails in a single animal species. University of California Publications in Zoology, 6, 65 - 114." type="journal article" year="1909">Ritter (1909)</bibRefCitation>
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undertook a detailed internal and external examination of many specimens in southern
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<collectingRegion id="20434895A92AFFDD13F8FC0EDD12FBC1" box="[255,369,1021,1047]" country="United States of America" name="California" pageId="24" pageNumber="425">California</collectingRegion>
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to determine what he called the “law and order” (!) that delimits a species, long before genetic sequencing was possible. He separated one variant as
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<taxonomicName id="2587FDF4A92AFFDD11FFFBD1D875FBED" authority="(Ritter, 1909)" baseAuthorityName="Ritter" baseAuthorityYear="1909" box="[760,1046,1057,1083]" class="Malacostraca" family="Epialtidae" genus="Pugettia" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="425" phylum="Arthropoda" rank="species" species="johnsoni">
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<emphasis id="D0F35A65A92AFFDD11FFFBD1DF17FBED" box="[760,884,1057,1083]" italics="true" pageId="24" pageNumber="425">P. johnsoni</emphasis>
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(
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<bibRefCitation id="8616FB86A92AFFDD1084FBD2D86DFBED" author="Ritter, W. E." box="[899,1038,1057,1083]" pageId="24" pageNumber="425" pagination="65 - 114" refId="ref17743" refString="Ritter, W. E. (1909) Halocynthia johnsoni n. sp., a comprehensive inquiry as to the extent of law and order that prevails in a single animal species. University of California Publications in Zoology, 6, 65 - 114." type="journal article" year="1909">Ritter, 1909</bibRefCitation>
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)
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</taxonomicName>
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, but this species was later synonymized by
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<bibRefCitation id="8616FB86A92AFFDD1202FBB5DDB3FBB6" author="Van Name, W. G." box="[261,464,1093,1120]" pageId="24" pageNumber="425" pagination="1 - 476" refId="ref18197" refString="Van Name, W. G. (1945) The North and South American ascidians. Bulletin of the American Museum of Natural History, 84, 1 - 476." type="journal article" year="1945">Van Name (1945)</bibRefCitation>
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.
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</paragraph>
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<paragraph id="E2388677A92AFFDD13C0FB9ADFCBFAA9" blockId="24.[151,1437,949,1408]" pageId="24" pageNumber="425">
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Specimens may attain a height of more than
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<quantity id="257F2B92A92AFFDD11C4FB9ADE98FB52" box="[707,763,1129,1156]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="425" unit="cm" value="5.0">5 cm</quantity>
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not including the length of the siphons. The reddish brown tunic is very tough and irregularly rugose, the long tubular siphons at the anterior end are divergent, and there are sharply pointed siphonal spines (
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<bibRefCitation id="8616FB86A92AFFDD110DFB42DEA5FB1A" author="Van Name, W. G." box="[522,710,1201,1228]" pageId="24" pageNumber="425" pagination="1 - 476" refId="ref18197" refString="Van Name, W. G. (1945) The North and South American ascidians. Bulletin of the American Museum of Natural History, 84, 1 - 476." type="journal article" year="1945">Van Name 1945</bibRefCitation>
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;
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<bibRefCitation id="8616FB86A92AFFDD11D4FB42DF14FB1D" author="Lambert, G." box="[723,887,1201,1227]" pageId="24" pageNumber="425" pagination="132 - 149" refId="ref16904" refString="Lambert, G. (2019) Fouling ascidians (Chordata: Ascidiacea) of the Galapagos: Santa Cruz and Baltra Islands. Aquatic Inva- sions, 14, 132 - 149. https: // doi. org / 10.3391 / ai. 2019.14.1.0 5" type="journal article" year="2019">Lambert 2019</bibRefCitation>
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). The pharyngeal sac contains six folds per side. There is great variability in the number of the branched oral tentacles, with southern
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<collectingRegion id="20434895A92AFFDD175DFB26D8AFFB39" box="[1114,1228,1237,1263]" country="United States of America" name="California" pageId="24" pageNumber="425">California</collectingRegion>
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specimens having up to 42 while northern
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<collectingRegion id="20434895A92AFFDD12A9FB0ADE50FAC5" box="[430,563,1273,1299]" country="United States of America" name="Washington" pageId="24" pageNumber="425">Washington</collectingRegion>
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and
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<collectingRegion id="20434895A92AFFDD116AFB0ADF53FAC5" box="[621,816,1273,1299]" country="Canada" name="British Columbia" pageId="24" pageNumber="425">British Columbia</collectingRegion>
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specimens averaging about half that number. See
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<bibRefCitation id="8616FB86A92AFFDD1676FB09DD49FAEE" author="Van Name, W. G." pageId="24" pageNumber="425" pagination="1 - 476" refId="ref18197" refString="Van Name, W. G. (1945) The North and South American ascidians. Bulletin of the American Museum of Natural History, 84, 1 - 476." type="journal article" year="1945">Van Name (1945)</bibRefCitation>
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for a detailed morphological description. Distribution:
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<collectingRegion id="20434895A92AFFDD108AFAEEDFB8FAE1" box="[909,987,1309,1335]" country="United States of America" name="Alaska" pageId="24" pageNumber="425">Alaska</collectingRegion>
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to southern
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<collectingRegion id="20434895A92AFFDD1765FAEED8B7FAE1" box="[1122,1236,1309,1335]" country="United States of America" name="California" pageId="24" pageNumber="425">California</collectingRegion>
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(
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<bibRefCitation id="8616FB86A92AFFDD17E7FAEED9F4FAEE" author="Van Name, W. G." box="[1248,1431,1309,1336]" pageId="24" pageNumber="425" pagination="1 - 476" refId="ref18197" refString="Van Name, W. G. (1945) The North and South American ascidians. Bulletin of the American Museum of Natural History, 84, 1 - 476." type="journal article" year="1945">Van Name 1945</bibRefCitation>
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;
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<bibRefCitation id="8616FB86A92AFFDD1390FAB2DDC4FA8D" author="O'Clair, R. M. & O'Clair, C. E." box="[151,423,1345,1371]" pageId="24" pageNumber="425" refId="ref17413" refString="O'Clair, R. M. & O'Clair, C. E. (1998) Southeast Alaska's Rocky Shores. Plant Press, Auke Bay, Alaska, 564 pp." type="book" year="1998">O’Clair & O’Clair 1998</bibRefCitation>
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;
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<bibRefCitation id="8616FB86A92AFFDD12B4FAB2DEC0FA8A" author="Lamb, A. & Hanby, B. P." box="[435,675,1345,1372]" pageId="24" pageNumber="425" refId="ref16347" refString="Lamb, A. & Hanby, B. P. (2005) Marine Life of the Pacific Northwest - A Photographic Encyclopedia of Invertebrates, Seaweeds and Selected Fishes. Harbour Publishing, Madeira Park, BC., 398 pp." type="book" year="2005">Lamb & Hanby 2005</bibRefCitation>
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) and the Galápagos (
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<bibRefCitation id="8616FB86A92AFFDD1089FAB2D850FA8D" author="Lambert, G." box="[910,1075,1345,1371]" pageId="24" pageNumber="425" pagination="132 - 149" refId="ref16904" refString="Lambert, G. (2019) Fouling ascidians (Chordata: Ascidiacea) of the Galapagos: Santa Cruz and Baltra Islands. Aquatic Inva- sions, 14, 132 - 149. https: // doi. org / 10.3391 / ai. 2019.14.1.0 5" type="journal article" year="2019">Lambert 2019</bibRefCitation>
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). In the present study only small specimens were collected, mostly under rocks near the marine station.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |