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<document id="9ED7D0D305D6690C535DAAA715F590C1" ID-DOI="10.5281/zenodo.191235" ID-GBIF-Dataset="8b2da9f6-b81d-4b48-8d0d-105e01ff3b0e" ID-ISSN="1175-5326" ID-Zenodo-Dep="191235" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461125932140" checkinUser="plazi" docAuthor="Micó, Estefanía &amp; Galante, Eduardo" docDate="2009" docId="03BE87E3FFCAF53F108AFA1285D24867" docLanguage="en" docName="zt02284p047.pdf" docOrigin="Zootaxa 2284" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pachydema" docType="key" docVersion="9" lastPageNumber="46" masterDocId="FF87FF9BFFCEF53A101DFFA8804E4D08" masterDocTitle="A new species of Pachydema Laporte (Coleoptera: Scarabaeoidea: Scarabaeidae: Melolonthinae: Pachydemini) from the Canary Islands" masterLastPageNumber="47" masterPageNumber="41" pageNumber="45" updateTime="1698596751661" updateUser="plazi">
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<mods:title id="7E41009ACA8DBEC8A4B83AF8670C622B">A new species of Pachydema Laporte (Coleoptera: Scarabaeoidea: Scarabaeidae: Melolonthinae: Pachydemini) from the Canary Islands</mods:title>
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<mods:namePart id="6F32D3E5B215BA9906CA86DDE19C4133">Micó, Estefanía</mods:namePart>
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<mods:namePart id="456485E1B0E8C1DF426F4B0972AB5D96">Galante, Eduardo</mods:namePart>
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<mods:typeOfResource id="C1FB3B73C0EC8996FAE594735C12D00B">text</mods:typeOfResource>
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<mods:date id="EC3FE5C0B1E2610C50FF80EA39E90FC1">2009</mods:date>
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<treatment id="03BE87E3FFCAF53F108AFA1285D24867" ID-DOI="http://doi.org/10.5281/zenodo.6220113" ID-GBIF-Taxon="119626366" ID-Zenodo-Dep="6220113" LSID="urn:lsid:plazi:treatment:03BE87E3FFCAF53F108AFA1285D24867" httpUri="http://treatment.plazi.org/id/03BE87E3FFCAF53F108AFA1285D24867" lastPageId="5" lastPageNumber="46" pageId="4" pageNumber="45">
<subSubSection id="C30D657EFFCAF53E108AFA1283374B63" pageId="4" pageNumber="45" type="nomenclature">
<paragraph id="8BA836F5FFCAF53E108AFA1283C248DC" blockId="4.[151,908,1466,1492]" box="[151,908,1466,1492]" pageId="4" pageNumber="45">
<heading id="D0E08199FFCAF53E108AFA1283C248DC" bold="true" box="[151,908,1466,1492]" fontSize="11" level="1" pageId="4" pageNumber="45" reason="1">
<emphasis id="B963EAE7FFCAF53E108AFA1283C248DC" bold="true" box="[151,908,1466,1492]" pageId="4" pageNumber="45">
Key to the males of
<taxonomicName id="4C174D76FFCAF53E1190FA12825748DC" box="[397,537,1466,1492]" class="Insecta" family="Melolonthidae" genus="Pachydema" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B963EAE7FFCAF53E1190FA12825748DC" bold="true" box="[397,537,1466,1492]" italics="true" pageId="4" pageNumber="45">Pachydema</emphasis>
</taxonomicName>
species of the Canary Islands
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA836F5FFCAF53E108AF9AC83374B63" blockId="4.[151,1437,1540,1643]" pageId="4" pageNumber="45">
All species of Canarian
<taxonomicName id="4C174D76FFCAF53E11DDF9AC82034B15" box="[448,589,1540,1565]" class="Insecta" family="Melolonthidae" genus="Pachydema" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B963EAE7FFCAF53E11DDF9AC82034B15" box="[448,589,1540,1565]" italics="true" pageId="4" pageNumber="45">Pachydema</emphasis>
</taxonomicName>
are included in the key with the exception of
<taxonomicName id="4C174D76FFCAF53E1498F9AC85D64B16" authority="Wollaston" authorityName="Wollaston" box="[1157,1432,1540,1566]" class="Insecta" family="Melolonthidae" genus="Pachydema" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="B963EAE7FFCAF53E1498F9AC84B24B15" box="[1157,1276,1540,1565]" italics="true" pageId="4" pageNumber="45">P. integra</emphasis>
(Wollaston)
</taxonomicName>
, which was described using a single female. The species distribution (modified from
<bibRefCitation id="EF864B04FFCAF53E1476F98285444B4C" author="Lacroix" box="[1131,1290,1578,1604]" pageId="4" pageNumber="47" refString="Lacroix, M. (2000) Les Pachydema des Iles Canaries: histoire, biogeographie historique et biologie larvaire d'une espece inedite (Coleoptera, Scarabaeoidea, Melolonthidae). Revue francaise d'Entomologie, 22 (1), 1 - 16." type="journal article" year="2000">Lacroix 2000</bibRefCitation>
) is shown in brackets. Locality records in bold have not been verified by us.
</paragraph>
</subSubSection>
<subSubSection id="C30D657EFFCAF53F108AF93685D24867" lastPageId="5" lastPageNumber="46" pageId="4" pageNumber="45" type="key">
<keyStep id="30E32E50FFCAF53E108AF93685D54BFD" pageId="4" pageNumber="45">
<paragraph id="8BA836F5FFCAF53E108AF93685D54BBD" blockId="4.[151,1436,1694,2037]" box="[151,1435,1694,1717]" pageId="4" pageNumber="45">
<keyLead id="30E695C0FFCAF53E108AF93685D54BBD" box="[151,1435,1694,1717]" pageId="4" pageNumber="45">
1. Prementum convex (
<figureCitation id="132C2A70FFCAF53E11B8F93681BF4BBD" box="[421,497,1694,1717]" captionStart="FIGURES 11 15. 11" captionStartId="2.[151,272,1893,1917]" captionTargetBox="[157,1419,206,1826]" captionTargetId="figure@2.[151,1436,134,1869]" captionTargetPageId="2" captionText="FIGURES 11 15. 11 — Mentum: a) P. ob s cu ra, b) P. ob s cu rell a, c) P. keithi. 12 — Galea and lacinia of maxilla: a) P. obscura, b) P. obscurella. 13 — Antenna: a) P. o b s c u r a, b) P. oromi, c) P. l e s n e i. 14 — Head: a) P. o b s c u r a, b) P. oromi, c) P. castanea. 15 — Apical maxillary palpomere: a) P. fuscipennis, b) P. o b s c u re l l a, c) P. l e s n e i, d) P. keithi, e) P. m e n i e r i, f) P. wollastoni, g) P. bipartita, h) P. navatteae. Scale bar in mm." httpUri="https://zenodo.org/record/191237/files/figure.png" pageId="4" pageNumber="45">Fig. 11</figureCitation>
a). Maxillary teeth reduced, with setae or spine-like setae at apex (
<figureCitation id="132C2A70FFCAF53E14A6F93685444BBD" box="[1211,1290,1694,1717]" captionStart="FIGURES 11 15. 11" captionStartId="2.[151,272,1893,1917]" captionTargetBox="[157,1419,206,1826]" captionTargetId="figure@2.[151,1436,134,1869]" captionTargetPageId="2" captionText="FIGURES 11 15. 11 — Mentum: a) P. ob s cu ra, b) P. ob s cu rell a, c) P. keithi. 12 — Galea and lacinia of maxilla: a) P. obscura, b) P. obscurella. 13 — Antenna: a) P. o b s c u r a, b) P. oromi, c) P. l e s n e i. 14 — Head: a) P. o b s c u r a, b) P. oromi, c) P. castanea. 15 — Apical maxillary palpomere: a) P. fuscipennis, b) P. o b s c u re l l a, c) P. l e s n e i, d) P. keithi, e) P. m e n i e r i, f) P. wollastoni, g) P. bipartita, h) P. navatteae. Scale bar in mm." httpUri="https://zenodo.org/record/191237/files/figure.png" pageId="4" pageNumber="45">Fig. 12</figureCitation>
a). .............. 2
</keyLead>
</paragraph>
<paragraph id="8BA836F5FFCAF53E108AF91685D54BFD" blockId="4.[151,1436,1694,2037]" pageId="4" pageNumber="45">
<keyLead id="30E695C0FFCAF53E108AF91685D54BFD" pageId="4" pageNumber="45">
- Prementum showing a deep concave area at apical third (
<figureCitation id="132C2A70FFCAF53E1334F91683304BDD" box="[809,894,1726,1749]" captionStart="FIGURES 11 15. 11" captionStartId="2.[151,272,1893,1917]" captionTargetBox="[157,1419,206,1826]" captionTargetId="figure@2.[151,1436,134,1869]" captionTargetPageId="2" captionText="FIGURES 11 15. 11 — Mentum: a) P. ob s cu ra, b) P. ob s cu rell a, c) P. keithi. 12 — Galea and lacinia of maxilla: a) P. obscura, b) P. obscurella. 13 — Antenna: a) P. o b s c u r a, b) P. oromi, c) P. l e s n e i. 14 — Head: a) P. o b s c u r a, b) P. oromi, c) P. castanea. 15 — Apical maxillary palpomere: a) P. fuscipennis, b) P. o b s c u re l l a, c) P. l e s n e i, d) P. keithi, e) P. m e n i e r i, f) P. wollastoni, g) P. bipartita, h) P. navatteae. Scale bar in mm." httpUri="https://zenodo.org/record/191237/files/figure.png" pageId="4" pageNumber="45">Figs. 11</figureCitation>
bc). Maxillary teeth well developed (
<figureCitation id="132C2A70FFCAF53E150BF916852D4BDD" box="[1302,1379,1726,1749]" captionStart="FIGURES 11 15. 11" captionStartId="2.[151,272,1893,1917]" captionTargetBox="[157,1419,206,1826]" captionTargetId="figure@2.[151,1436,134,1869]" captionTargetPageId="2" captionText="FIGURES 11 15. 11 — Mentum: a) P. ob s cu ra, b) P. ob s cu rell a, c) P. keithi. 12 — Galea and lacinia of maxilla: a) P. obscura, b) P. obscurella. 13 — Antenna: a) P. o b s c u r a, b) P. oromi, c) P. l e s n e i. 14 — Head: a) P. o b s c u r a, b) P. oromi, c) P. castanea. 15 — Apical maxillary palpomere: a) P. fuscipennis, b) P. o b s c u re l l a, c) P. l e s n e i, d) P. keithi, e) P. m e n i e r i, f) P. wollastoni, g) P. bipartita, h) P. navatteae. Scale bar in mm." httpUri="https://zenodo.org/record/191237/files/figure.png" pageId="4" pageNumber="45">Fig. 12</figureCitation>
b).. ..................................................................................................................................................................................... 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E32E50FFCAF53E108AF95685D54A7D" pageId="4" pageNumber="45">
<paragraph id="8BA836F5FFCAF53E108AF95685D54A3D" blockId="4.[151,1436,1694,2037]" pageId="4" pageNumber="45">
<keyLead id="30E695C0FFCAF53E108AF95685D54A3D" pageId="4" pageNumber="45">
2. Antennal club more than twice as long the preceding 4 antennomeres combined (
<figureCitation id="132C2A70FFCAF53E1447F95684E24A1D" box="[1114,1196,1790,1813]" captionStart="FIGURES 11 15. 11" captionStartId="2.[151,272,1893,1917]" captionTargetBox="[157,1419,206,1826]" captionTargetId="figure@2.[151,1436,134,1869]" captionTargetPageId="2" captionText="FIGURES 11 15. 11 — Mentum: a) P. ob s cu ra, b) P. ob s cu rell a, c) P. keithi. 12 — Galea and lacinia of maxilla: a) P. obscura, b) P. obscurella. 13 — Antenna: a) P. o b s c u r a, b) P. oromi, c) P. l e s n e i. 14 — Head: a) P. o b s c u r a, b) P. oromi, c) P. castanea. 15 — Apical maxillary palpomere: a) P. fuscipennis, b) P. o b s c u re l l a, c) P. l e s n e i, d) P. keithi, e) P. m e n i e r i, f) P. wollastoni, g) P. bipartita, h) P. navatteae. Scale bar in mm." httpUri="https://zenodo.org/record/191237/files/figure.png" pageId="4" pageNumber="45">Fig. 13</figureCitation>
a). Genae protruding beyond canthus (
<figureCitation id="132C2A70FFCAF53E1161F8B681864A3D" box="[380,456,1822,1845]" captionStart="FIGURES 11 15. 11" captionStartId="2.[151,272,1893,1917]" captionTargetBox="[157,1419,206,1826]" captionTargetId="figure@2.[151,1436,134,1869]" captionTargetPageId="2" captionText="FIGURES 11 15. 11 — Mentum: a) P. ob s cu ra, b) P. ob s cu rell a, c) P. keithi. 12 — Galea and lacinia of maxilla: a) P. obscura, b) P. obscurella. 13 — Antenna: a) P. o b s c u r a, b) P. oromi, c) P. l e s n e i. 14 — Head: a) P. o b s c u r a, b) P. oromi, c) P. castanea. 15 — Apical maxillary palpomere: a) P. fuscipennis, b) P. o b s c u re l l a, c) P. l e s n e i, d) P. keithi, e) P. m e n i e r i, f) P. wollastoni, g) P. bipartita, h) P. navatteae. Scale bar in mm." httpUri="https://zenodo.org/record/191237/files/figure.png" pageId="4" pageNumber="45">Fig. 14</figureCitation>
a). ......................................................................................................................................... 3
</keyLead>
</paragraph>
<paragraph id="8BA836F5FFCAF53E108AF89685D54A7D" blockId="4.[151,1436,1694,2037]" pageId="4" pageNumber="45">
<keyLead id="30E695C0FFCAF53E108AF89685D54A7D" pageId="4" pageNumber="45">
- Antennal club less than 1.5 times the preceding 4 antennomeres combined (
<figureCitation id="132C2A70FFCAF53E13ECF89684714A5D" box="[1009,1087,1854,1877]" captionStart="FIGURES 11 15. 11" captionStartId="2.[151,272,1893,1917]" captionTargetBox="[157,1419,206,1826]" captionTargetId="figure@2.[151,1436,134,1869]" captionTargetPageId="2" captionText="FIGURES 11 15. 11 — Mentum: a) P. ob s cu ra, b) P. ob s cu rell a, c) P. keithi. 12 — Galea and lacinia of maxilla: a) P. obscura, b) P. obscurella. 13 — Antenna: a) P. o b s c u r a, b) P. oromi, c) P. l e s n e i. 14 — Head: a) P. o b s c u r a, b) P. oromi, c) P. castanea. 15 — Apical maxillary palpomere: a) P. fuscipennis, b) P. o b s c u re l l a, c) P. l e s n e i, d) P. keithi, e) P. m e n i e r i, f) P. wollastoni, g) P. bipartita, h) P. navatteae. Scale bar in mm." httpUri="https://zenodo.org/record/191237/files/figure.png" pageId="4" pageNumber="45">Fig. 13</figureCitation>
b). Genae not protruding beyond canthus (14b). [La Gomera].
<emphasis id="B963EAE7FFCAF53E121CF8F785474A7E" box="[513,1289,1887,1910]" italics="true" pageId="4" pageNumber="45">
.....................................................................................................
<taxonomicName id="4C174D76FFCAF53E14B5F8F785474A7E" authority="Lopez-Colon" authorityName="Lopez-Colon" box="[1192,1289,1887,1910]" class="Insecta" family="Melolonthidae" genus="Pachydema" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="oromii">P. oromii</taxonomicName>
</emphasis>
López-Colón
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E32E50FFCAF53E108AF8D685D54ADD" pageId="4" pageNumber="45">
<paragraph id="8BA836F5FFCAF53E108AF8D685D54ABD" blockId="4.[151,1436,1694,2037]" pageId="4" pageNumber="45">
<keyLead id="30E695C0FFCAF53E108AF8D685D54ABD" pageId="4" pageNumber="45">
3. Genae sharply angled and protruding beyond canthus (
<figureCitation id="132C2A70FFCAF53E1309F8D6832C4A9D" box="[788,866,1918,1941]" captionStart="FIGURES 11 15. 11" captionStartId="2.[151,272,1893,1917]" captionTargetBox="[157,1419,206,1826]" captionTargetId="figure@2.[151,1436,134,1869]" captionTargetPageId="2" captionText="FIGURES 11 15. 11 — Mentum: a) P. ob s cu ra, b) P. ob s cu rell a, c) P. keithi. 12 — Galea and lacinia of maxilla: a) P. obscura, b) P. obscurella. 13 — Antenna: a) P. o b s c u r a, b) P. oromi, c) P. l e s n e i. 14 — Head: a) P. o b s c u r a, b) P. oromi, c) P. castanea. 15 — Apical maxillary palpomere: a) P. fuscipennis, b) P. o b s c u re l l a, c) P. l e s n e i, d) P. keithi, e) P. m e n i e r i, f) P. wollastoni, g) P. bipartita, h) P. navatteae. Scale bar in mm." httpUri="https://zenodo.org/record/191237/files/figure.png" pageId="4" pageNumber="45">Fig. 14</figureCitation>
a). Parameres as in
<figureCitation id="132C2A70FFCAF53E1421F8D684C74A9D" box="[1084,1161,1918,1941]" captionStart="FIGURES 16 17. 16" captionStartId="3.[151,271,1654,1678]" captionTargetBox="[245,1345,200,1541]" captionTargetId="figure@3.[226,1360,194,1630]" captionTargetPageId="3" captionText="FIGURES 16 17. 16 — Parameres, dorsal view: a) P. obscura, b) P. tinerfensis, c) P. fortunatorum, d) P. jeannei, e) P. castanea, f) P. fuscipennis, g) P. obscurella, h) P. g o m er a e, i) P. l e s n e i, j) P. keithi, k) P. m e n i e r i, l) P. bipartita, m) P. wollastoni, n) P. navatteae. 17 — Metatibia. a) P. obscurella, b) P. ke i t h i. Scale bar in mm." httpUri="https://zenodo.org/record/191238/files/figure.png" pageId="4" pageNumber="45">Fig. 16</figureCitation>
a. [Tenerife,
<emphasis id="B963EAE7FFCAF53E150CF8D685DB4A9E" bold="true" box="[1297,1429,1918,1942]" pageId="4" pageNumber="45">La Gomera</emphasis>
,
<emphasis id="B963EAE7FFCAF53E10DAF83681254ABE" bold="true" box="[199,363,1950,1974]" pageId="4" pageNumber="45">Fuerteventura</emphasis>
].
<emphasis id="B963EAE7FFCAF53E119FF837849E4ABE" box="[386,1232,1951,1974]" italics="true" pageId="4" pageNumber="45">...............................................................................................................................</emphasis>
<taxonomicName id="4C174D76FFCAF53E14CCF83785D54ABD" authority="(Brulle)" authorityName="Brulle" baseAuthorityName="Brulle" box="[1233,1435,1950,1974]" class="Insecta" family="Melolonthidae" genus="Pachydema" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="B963EAE7FFCAF53E14CCF837850F4ABE" box="[1233,1345,1951,1974]" italics="true" pageId="4" pageNumber="45">P. obscura</emphasis>
(Brullé)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA836F5FFCAF53E108AF81685D54ADD" blockId="4.[151,1436,1694,2037]" box="[151,1435,1982,2005]" pageId="4" pageNumber="45">
<keyLead id="30E695C0FFCAF53E108AF81685D54ADD" box="[151,1435,1982,2005]" pageId="4" pageNumber="45">- Genae rounded and slightly protruding beyond canthus. ............................................................................................ 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E32E50FFCAF53F108AF87685D54DC7" lastPageId="5" lastPageNumber="46" pageId="4" pageNumber="45">
<paragraph id="8BA836F5FFCAF53E108AF87685D64AFD" blockId="4.[151,1436,1694,2037]" box="[151,1432,2014,2037]" pageId="4" pageNumber="45">
<keyLead id="30E695C0FFCAF53E108AF87685D64AFD" box="[151,1432,2014,2037]" pageId="4" pageNumber="45">
4. Second protarsomere not widened (1.8 times longer than wide) Parameres as in
<figureCitation id="132C2A70FFCAF53E1401F87684274AFD" box="[1052,1129,2014,2037]" captionStart="FIGURES 16 17. 16" captionStartId="3.[151,271,1654,1678]" captionTargetBox="[245,1345,200,1541]" captionTargetId="figure@3.[226,1360,194,1630]" captionTargetPageId="3" captionText="FIGURES 16 17. 16 — Parameres, dorsal view: a) P. obscura, b) P. tinerfensis, c) P. fortunatorum, d) P. jeannei, e) P. castanea, f) P. fuscipennis, g) P. obscurella, h) P. g o m er a e, i) P. l e s n e i, j) P. keithi, k) P. m e n i e r i, l) P. bipartita, m) P. wollastoni, n) P. navatteae. 17 — Metatibia. a) P. obscurella, b) P. ke i t h i. Scale bar in mm." httpUri="https://zenodo.org/record/191238/files/figure.png" pageId="4" pageNumber="45">Fig. 16</figureCitation>
b. [Tenerife]. .......................
</keyLead>
</paragraph>
<paragraph id="8BA836F5FFCBF53F10D3FF3085D54DC7" blockId="5.[151,1436,152,1391]" pageId="5" pageNumber="46">
<emphasis id="B963EAE7FFCBF53F10D3FF3084F64DA7" box="[206,1208,152,175]" italics="true" pageId="5" pageNumber="46">
.................................................................................................................................
<taxonomicName id="4C174D76FFCBF53F1433FF3084F64DA7" authority="Galante &amp; Stebnicka" authorityName="Galante &amp; Stebnicka" box="[1070,1208,152,175]" class="Insecta" family="Melolonthidae" genus="Pachydema" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="tinerfensis">P. tinerfensis</taxonomicName>
</emphasis>
Galante &amp; Stebnicka
<keyLead id="30E695C0FFCBF53F108AFF1085D54DC7" box="[151,1435,184,207]" pageId="5" pageNumber="46">- Second protarsomere slightly widened (1.3 times longer than wide). ......................................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E32E50FFCBF53F108AFF7085D24C47" pageId="5" pageNumber="46">
<paragraph id="8BA836F5FFCBF53F108AFF7085D24C07" blockId="5.[151,1436,152,1391]" pageId="5" pageNumber="46">
<keyLead id="30E695C0FFCBF53F108AFF7085D24C07" pageId="5" pageNumber="46">
5. Elytra conspicuously convex and slightly punctate. Anterior border of clypeus emarginate. Parameres as in
<figureCitation id="132C2A70FFCBF53F1523FF7085C24DE7" box="[1342,1420,216,239]" captionStart="FIGURES 16 17. 16" captionStartId="3.[151,271,1654,1678]" captionTargetBox="[245,1345,200,1541]" captionTargetId="figure@3.[226,1360,194,1630]" captionTargetPageId="3" captionText="FIGURES 16 17. 16 — Parameres, dorsal view: a) P. obscura, b) P. tinerfensis, c) P. fortunatorum, d) P. jeannei, e) P. castanea, f) P. fuscipennis, g) P. obscurella, h) P. g o m er a e, i) P. l e s n e i, j) P. keithi, k) P. m e n i e r i, l) P. bipartita, m) P. wollastoni, n) P. navatteae. 17 — Metatibia. a) P. obscurella, b) P. ke i t h i. Scale bar in mm." httpUri="https://zenodo.org/record/191238/files/figure.png" pageId="5" pageNumber="46">Fig. 16</figureCitation>
c. [Tenerife].
<emphasis id="B963EAE7FFCBF53F1122FF5085074C07" box="[319,1353,248,271]" italics="true" pageId="5" pageNumber="46">
.................................................................................................................................
<taxonomicName id="4C174D76FFCBF53F14BDFF5085074C07" authority="Baraud" authorityName="Baraud" box="[1184,1353,248,271]" class="Insecta" family="Melolonthidae" genus="Pachydema" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="fortunatorum">P. fortunatorum</taxonomicName>
</emphasis>
Baraud
</keyLead>
</paragraph>
<paragraph id="8BA836F5FFCBF53F108AFEB085D24C47" blockId="5.[151,1436,152,1391]" pageId="5" pageNumber="46">
<keyLead id="30E695C0FFCBF53F108AFEB085D24C47" pageId="5" pageNumber="46">
- Elytra less convex. Anterior border of clypeus very slightly emarginate. Parameres as in
<figureCitation id="132C2A70FFCBF53F1464FEB084884C27" box="[1145,1222,280,303]" captionStart="FIGURES 16 17. 16" captionStartId="3.[151,271,1654,1678]" captionTargetBox="[245,1345,200,1541]" captionTargetId="figure@3.[226,1360,194,1630]" captionTargetPageId="3" captionText="FIGURES 16 17. 16 — Parameres, dorsal view: a) P. obscura, b) P. tinerfensis, c) P. fortunatorum, d) P. jeannei, e) P. castanea, f) P. fuscipennis, g) P. obscurella, h) P. g o m er a e, i) P. l e s n e i, j) P. keithi, k) P. m e n i e r i, l) P. bipartita, m) P. wollastoni, n) P. navatteae. 17 — Metatibia. a) P. obscurella, b) P. ke i t h i. Scale bar in mm." httpUri="https://zenodo.org/record/191238/files/figure.png" pageId="5" pageNumber="46">Fig. 16</figureCitation>
d. [Tenerife]. .........
<emphasis id="B963EAE7FFCBF53F10D3FE9085074C47" box="[206,1353,312,335]" italics="true" pageId="5" pageNumber="46">
............................................................................................................................................................
<taxonomicName id="4C174D76FFCBF53F14FDFE9085074C47" authority="Baraud" authorityName="Baraud" box="[1248,1353,312,335]" class="Insecta" family="Melolonthidae" genus="Pachydema" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="jeannei">P. jeannei</taxonomicName>
</emphasis>
Baraud
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E32E50FFCBF53F108AFEF085D54CC7" pageId="5" pageNumber="46">
<paragraph id="8BA836F5FFCBF53F108AFEF085D54C87" blockId="5.[151,1436,152,1391]" pageId="5" pageNumber="46">
<keyLead id="30E695C0FFCBF53F108AFEF085D54C87" pageId="5" pageNumber="46">
6. Scutellum with punctures at disc. Metatibia internal border not dilated at middle (
<figureCitation id="132C2A70FFCBF53F1438FEF0843D4C67" box="[1061,1139,344,367]" captionStart="FIGURES 16 17. 16" captionStartId="3.[151,271,1654,1678]" captionTargetBox="[245,1345,200,1541]" captionTargetId="figure@3.[226,1360,194,1630]" captionTargetPageId="3" captionText="FIGURES 16 17. 16 — Parameres, dorsal view: a) P. obscura, b) P. tinerfensis, c) P. fortunatorum, d) P. jeannei, e) P. castanea, f) P. fuscipennis, g) P. obscurella, h) P. g o m er a e, i) P. l e s n e i, j) P. keithi, k) P. m e n i e r i, l) P. bipartita, m) P. wollastoni, n) P. navatteae. 17 — Metatibia. a) P. obscurella, b) P. ke i t h i. Scale bar in mm." httpUri="https://zenodo.org/record/191238/files/figure.png" pageId="5" pageNumber="46">Fig. 17</figureCitation>
a). Parameres forcep shaped (narrow and sinuate, as in
<figureCitation id="132C2A70FFCBF53F11FDFED082784C87" box="[480,566,376,399]" captionStart="FIGURES 16 17. 16" captionStartId="3.[151,271,1654,1678]" captionTargetBox="[245,1345,200,1541]" captionTargetId="figure@3.[226,1360,194,1630]" captionTargetPageId="3" captionText="FIGURES 16 17. 16 — Parameres, dorsal view: a) P. obscura, b) P. tinerfensis, c) P. fortunatorum, d) P. jeannei, e) P. castanea, f) P. fuscipennis, g) P. obscurella, h) P. g o m er a e, i) P. l e s n e i, j) P. keithi, k) P. m e n i e r i, l) P. bipartita, m) P. wollastoni, n) P. navatteae. 17 — Metatibia. a) P. obscurella, b) P. ke i t h i. Scale bar in mm." httpUri="https://zenodo.org/record/191238/files/figure.png" pageId="5" pageNumber="46">Figs. 16</figureCitation>
eh). ...................................................................................................................... 7
</keyLead>
</paragraph>
<paragraph id="8BA836F5FFCBF53F108AFE3085D54CC7" blockId="5.[151,1436,152,1391]" pageId="5" pageNumber="46">
<keyLead id="30E695C0FFCBF53F108AFE3085D54CC7" pageId="5" pageNumber="46">
- Scutellum without punctures at disc. Metatibia internal border dilated at middle. Parameres not sinuate or arrow shaped (as in
<figureCitation id="132C2A70FFCBF53F1144FE1081E14CC7" box="[345,431,440,463]" captionStart="FIGURES 16 17. 16" captionStartId="3.[151,271,1654,1678]" captionTargetBox="[245,1345,200,1541]" captionTargetId="figure@3.[226,1360,194,1630]" captionTargetPageId="3" captionText="FIGURES 16 17. 16 — Parameres, dorsal view: a) P. obscura, b) P. tinerfensis, c) P. fortunatorum, d) P. jeannei, e) P. castanea, f) P. fuscipennis, g) P. obscurella, h) P. g o m er a e, i) P. l e s n e i, j) P. keithi, k) P. m e n i e r i, l) P. bipartita, m) P. wollastoni, n) P. navatteae. 17 — Metatibia. a) P. obscurella, b) P. ke i t h i. Scale bar in mm." httpUri="https://zenodo.org/record/191238/files/figure.png" pageId="5" pageNumber="46">
Figs.
<date id="FFA91035FFCBF53F1189FE1081E14CC7" box="[404,431,440,463]" pageId="5" pageNumber="46" value="1916-01">16</date>
</figureCitation>
in). ......................................................................................................................................... 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E32E50FFCBF53F108AFE7085D54F27" pageId="5" pageNumber="46">
<paragraph id="8BA836F5FFCBF53F108AFE7085D54F07" blockId="5.[151,1436,152,1391]" pageId="5" pageNumber="46">
<keyLead id="30E695C0FFCBF53F108AFE7085D54F07" pageId="5" pageNumber="46">
7. Eyes large and laterally exposed. Front between eyes nearly twice as wide as eye diameter (
<figureCitation id="132C2A70FFCBF53F14BCFE7084BF4CE7" box="[1185,1265,472,495]" captionStart="FIGURES 11 15. 11" captionStartId="2.[151,272,1893,1917]" captionTargetBox="[157,1419,206,1826]" captionTargetId="figure@2.[151,1436,134,1869]" captionTargetPageId="2" captionText="FIGURES 11 15. 11 — Mentum: a) P. ob s cu ra, b) P. ob s cu rell a, c) P. keithi. 12 — Galea and lacinia of maxilla: a) P. obscura, b) P. obscurella. 13 — Antenna: a) P. o b s c u r a, b) P. oromi, c) P. l e s n e i. 14 — Head: a) P. o b s c u r a, b) P. oromi, c) P. castanea. 15 — Apical maxillary palpomere: a) P. fuscipennis, b) P. o b s c u re l l a, c) P. l e s n e i, d) P. keithi, e) P. m e n i e r i, f) P. wollastoni, g) P. bipartita, h) P. navatteae. Scale bar in mm." httpUri="https://zenodo.org/record/191237/files/figure.png" pageId="5" pageNumber="46">Fig. 14</figureCitation>
c). [Tenerife, La Gomera].
<emphasis id="B963EAE7FFCBF53F1124FE5084F34F07" box="[313,1213,504,527]" italics="true" pageId="5" pageNumber="46">.......................................................................................................................................</emphasis>
<taxonomicName id="4C174D76FFCBF53F14D5FE5085D54F07" authority="(Brulle)" authorityName="Brulle" baseAuthorityName="Brulle" box="[1224,1435,504,527]" class="Insecta" family="Melolonthidae" genus="Pachydema" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="B963EAE7FFCBF53F14D5FE50850F4F07" box="[1224,1345,504,527]" italics="true" pageId="5" pageNumber="46">P. castanea</emphasis>
(Brullé)
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</keyLead>
</paragraph>
<paragraph id="8BA836F5FFCBF53F108AFDB085D54F27" blockId="5.[151,1436,152,1391]" box="[151,1435,536,559]" pageId="5" pageNumber="46">
<keyLead id="30E695C0FFCBF53F108AFDB085D54F27" box="[151,1435,536,559]" pageId="5" pageNumber="46">- Eyes small and not laterally exposed. Front between eyes more than three times as wide as eye diameter. .............. 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E32E50FFCBF53F108AFD9085D54F87" pageId="5" pageNumber="46">
<paragraph id="8BA836F5FFCBF53F108AFD9085D54F67" blockId="5.[151,1436,152,1391]" pageId="5" pageNumber="46">
<keyLead id="30E695C0FFCBF53F108AFD9085D54F67" pageId="5" pageNumber="46">
8. Apical maxillary palpomere narrowly oval (
<figureCitation id="132C2A70FFCBF53F1286FD9082A64F47" box="[667,744,568,591]" captionStart="FIGURES 11 15. 11" captionStartId="2.[151,272,1893,1917]" captionTargetBox="[157,1419,206,1826]" captionTargetId="figure@2.[151,1436,134,1869]" captionTargetPageId="2" captionText="FIGURES 11 15. 11 — Mentum: a) P. ob s cu ra, b) P. ob s cu rell a, c) P. keithi. 12 — Galea and lacinia of maxilla: a) P. obscura, b) P. obscurella. 13 — Antenna: a) P. o b s c u r a, b) P. oromi, c) P. l e s n e i. 14 — Head: a) P. o b s c u r a, b) P. oromi, c) P. castanea. 15 — Apical maxillary palpomere: a) P. fuscipennis, b) P. o b s c u re l l a, c) P. l e s n e i, d) P. keithi, e) P. m e n i e r i, f) P. wollastoni, g) P. bipartita, h) P. navatteae. Scale bar in mm." httpUri="https://zenodo.org/record/191237/files/figure.png" pageId="5" pageNumber="46">Fig. 15</figureCitation>
a). Parameres as in
<figureCitation id="132C2A70FFCBF53F13A8FD90844E4F47" box="[949,1024,568,591]" captionStart="FIGURES 16 17. 16" captionStartId="3.[151,271,1654,1678]" captionTargetBox="[245,1345,200,1541]" captionTargetId="figure@3.[226,1360,194,1630]" captionTargetPageId="3" captionText="FIGURES 16 17. 16 — Parameres, dorsal view: a) P. obscura, b) P. tinerfensis, c) P. fortunatorum, d) P. jeannei, e) P. castanea, f) P. fuscipennis, g) P. obscurella, h) P. g o m er a e, i) P. l e s n e i, j) P. keithi, k) P. m e n i e r i, l) P. bipartita, m) P. wollastoni, n) P. navatteae. 17 — Metatibia. a) P. obscurella, b) P. ke i t h i. Scale bar in mm." httpUri="https://zenodo.org/record/191238/files/figure.png" pageId="5" pageNumber="46">
Fig.
<specimenCount id="9D11FD7CFFCBF53F13F8FD90844E4F47" box="[997,1024,568,591]" pageId="5" pageNumber="46" type="generic">16</specimenCount>
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f. [
<emphasis id="B963EAE7FFCBF53F1403FD9F84F04F47" bold="true" box="[1054,1214,567,591]" pageId="5" pageNumber="46">Gran Canaria</emphasis>
, Tenerife, La Palma,
<emphasis id="B963EAE7FFCBF53F10DAFDFF81254F67" bold="true" box="[199,363,599,623]" pageId="5" pageNumber="46">Fuerteventura</emphasis>
,
<emphasis id="B963EAE7FFCBF53F1165FDFF81B54F67" bold="true" box="[376,507,599,623]" pageId="5" pageNumber="46">La Gomera</emphasis>
]. ..................................................................................................
<taxonomicName id="4C174D76FFCBF53F14B2FDF085D54F67" authority="Brulle" authorityName="Brulle" box="[1199,1435,599,623]" class="Insecta" family="Melolonthidae" genus="Pachydema" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="fuscipennis">
<emphasis id="B963EAE7FFCBF53F14B2FDF0850F4F67" box="[1199,1345,600,623]" italics="true" pageId="5" pageNumber="46">P. fuscipennis</emphasis>
<emphasis id="B963EAE7FFCBF53F155AFDFF851E4F67" bold="true" box="[1351,1360,599,623]" pageId="5" pageNumber="46">(</emphasis>
Brullé)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA836F5FFCBF53F108AFDD085D54F87" blockId="5.[151,1436,152,1391]" box="[151,1435,632,655]" pageId="5" pageNumber="46">
<keyLead id="30E695C0FFCBF53F108AFDD085D54F87" box="[151,1435,632,655]" pageId="5" pageNumber="46">
- Apical maxillary palpomere largely oval as in
<figureCitation id="132C2A70FFCBF53F12AAFDD0834D4F87" box="[695,771,632,655]" captionStart="FIGURES 11 15. 11" captionStartId="2.[151,272,1893,1917]" captionTargetBox="[157,1419,206,1826]" captionTargetId="figure@2.[151,1436,134,1869]" captionTargetPageId="2" captionText="FIGURES 11 15. 11 — Mentum: a) P. ob s cu ra, b) P. ob s cu rell a, c) P. keithi. 12 — Galea and lacinia of maxilla: a) P. obscura, b) P. obscurella. 13 — Antenna: a) P. o b s c u r a, b) P. oromi, c) P. l e s n e i. 14 — Head: a) P. o b s c u r a, b) P. oromi, c) P. castanea. 15 — Apical maxillary palpomere: a) P. fuscipennis, b) P. o b s c u re l l a, c) P. l e s n e i, d) P. keithi, e) P. m e n i e r i, f) P. wollastoni, g) P. bipartita, h) P. navatteae. Scale bar in mm." httpUri="https://zenodo.org/record/191237/files/figure.png" pageId="5" pageNumber="46">Fig. 15</figureCitation>
b. ............................................................................................. 9
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</paragraph>
</keyStep>
<keyStep id="30E32E50FFCBF53F108AFD3085D54FC7" pageId="5" pageNumber="46">
<paragraph id="8BA836F5FFCBF53F108AFD3085D24FA7" blockId="5.[151,1436,152,1391]" box="[151,1436,663,687]" pageId="5" pageNumber="46">
<keyLead id="30E695C0FFCBF53F108AFD3085D24FA7" box="[151,1436,663,687]" pageId="5" pageNumber="46">
9. Frons and pronotum densely punctate. Parameres as in
<figureCitation id="132C2A70FFCBF53F1308FD30832C4FA7" box="[789,866,664,687]" captionStart="FIGURES 16 17. 16" captionStartId="3.[151,271,1654,1678]" captionTargetBox="[245,1345,200,1541]" captionTargetId="figure@3.[226,1360,194,1630]" captionTargetPageId="3" captionText="FIGURES 16 17. 16 — Parameres, dorsal view: a) P. obscura, b) P. tinerfensis, c) P. fortunatorum, d) P. jeannei, e) P. castanea, f) P. fuscipennis, g) P. obscurella, h) P. g o m er a e, i) P. l e s n e i, j) P. keithi, k) P. m e n i e r i, l) P. bipartita, m) P. wollastoni, n) P. navatteae. 17 — Metatibia. a) P. obscurella, b) P. ke i t h i. Scale bar in mm." httpUri="https://zenodo.org/record/191238/files/figure.png" pageId="5" pageNumber="46">
Fig.
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g. [El Hierro,
<emphasis id="B963EAE7FFCBF53F13E8FD3F842D4FA7" bold="true" box="[1013,1123,663,687]" pageId="5" pageNumber="46">La Palma</emphasis>
].
<emphasis id="B963EAE7FFCBF53F1467FD3084C34FA7" box="[1146,1165,664,687]" italics="true" pageId="5" pageNumber="46">...</emphasis>
<taxonomicName id="4C174D76FFCBF53F1491FD3085D24FA7" authority="(Wollaston)" authorityName="Wollaston" baseAuthorityName="Wollaston" box="[1164,1436,664,687]" class="Insecta" family="Melolonthidae" genus="Pachydema" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="obscurella">
<emphasis id="B963EAE7FFCBF53F1491FD3085564FA7" box="[1164,1304,664,687]" italics="true" pageId="5" pageNumber="46">P. obscurella</emphasis>
(Wollaston)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA836F5FFCBF53F108AFD1085D54FC7" blockId="5.[151,1436,152,1391]" box="[151,1435,696,719]" pageId="5" pageNumber="46">
<keyLead id="30E695C0FFCBF53F108AFD1085D54FC7" box="[151,1435,696,719]" pageId="5" pageNumber="46">
- Frons and pronotum less densely punctate. Parameres as in
<figureCitation id="132C2A70FFCBF53F1359FD1083DE4FC7" box="[836,912,696,719]" captionStart="FIGURES 16 17. 16" captionStartId="3.[151,271,1654,1678]" captionTargetBox="[245,1345,200,1541]" captionTargetId="figure@3.[226,1360,194,1630]" captionTargetPageId="3" captionText="FIGURES 16 17. 16 — Parameres, dorsal view: a) P. obscura, b) P. tinerfensis, c) P. fortunatorum, d) P. jeannei, e) P. castanea, f) P. fuscipennis, g) P. obscurella, h) P. g o m er a e, i) P. l e s n e i, j) P. keithi, k) P. m e n i e r i, l) P. bipartita, m) P. wollastoni, n) P. navatteae. 17 — Metatibia. a) P. obscurella, b) P. ke i t h i. Scale bar in mm." httpUri="https://zenodo.org/record/191238/files/figure.png" pageId="5" pageNumber="46">Fig. 16</figureCitation>
h. [La Gomera]. ...........
<taxonomicName id="4C174D76FFCBF53F148CFD1085D54FC7" authority="Lopez-Colon" authorityName="Lopez-Colon" box="[1169,1435,696,719]" class="Insecta" family="Melolonthidae" genus="Pachydema" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="gomerae">
<emphasis id="B963EAE7FFCBF53F148CFD1085464FC7" box="[1169,1288,696,719]" italics="true" pageId="5" pageNumber="46">P. gomerae</emphasis>
López-Colón
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</paragraph>
</keyStep>
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<paragraph id="8BA836F5FFCBF53F108AFD7085D24FE7" blockId="5.[151,1436,152,1391]" box="[151,1436,728,751]" pageId="5" pageNumber="46">
<keyLead id="30E695C0FFCBF53F108AFD7085D24FE7" box="[151,1436,728,751]" pageId="5" pageNumber="46">10. Eyes large and laterally exposed. Front between eyes nearly twice as wide as eye diameter. .................................. 11</keyLead>
</paragraph>
<paragraph id="8BA836F5FFCBF53F108AFD5085D54E07" blockId="5.[151,1436,152,1391]" box="[151,1435,760,783]" pageId="5" pageNumber="46">
<keyLead id="30E695C0FFCBF53F108AFD5085D54E07" box="[151,1435,760,783]" pageId="5" pageNumber="46">- Eyes small and not laterally exposed. Front between eyes more than three times as wide as eye diameter. .......... 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E32E50FFCBF53F108AFCB085D24EC7" pageId="5" pageNumber="46">
<paragraph id="8BA836F5FFCBF53F108AFCB085D24E67" blockId="5.[151,1436,152,1391]" pageId="5" pageNumber="46">
<keyLead id="30E695C0FFCBF53F108AFCB085D24E67" pageId="5" pageNumber="46">
11. Apical maxillary palpomere large (ratio palpomere length/habitus length: 0.03) and widened (length 2.2 times the width). Third and fourth antennomeres short and wide (1.7 times longer than wide) (
<figureCitation id="132C2A70FFCBF53F146AFC9084F24E47" box="[1143,1212,824,847]" captionStart="FIGURES 1 10" captionStartId="1.[151,273,1601,1625]" captionTargetBox="[152,1435,551,1600]" captionTargetId="figure@1.[151,1436,545,1622]" captionTargetPageId="1" captionText="FIGURES 1 10. Pachydema megalops Micó sp. nov. 1 — Head. 2 — Antenna. 3 — Galea and lacinia of maxilla. 4 — Apical maxillary palpomere. 5 — Mentum. 6 — Pronotum. 7 — Scutellum. 8 — Metatibia. 9 — Protarsus. 10 — Parameres, dorsal view. Scale bar in mm." httpUri="https://zenodo.org/record/191236/files/figure.png" pageId="5" pageNumber="46">Fig. 2</figureCitation>
). Parameres slightly arrow shaped (
<figureCitation id="132C2A70FFCBF53F1179FCF081FA4E67" box="[356,436,856,879]" captionStart="FIGURES 1 10" captionStartId="1.[151,273,1601,1625]" captionTargetBox="[152,1435,551,1600]" captionTargetId="figure@1.[151,1436,545,1622]" captionTargetPageId="1" captionText="FIGURES 1 10. Pachydema megalops Micó sp. nov. 1 — Head. 2 — Antenna. 3 — Galea and lacinia of maxilla. 4 — Apical maxillary palpomere. 5 — Mentum. 6 — Pronotum. 7 — Scutellum. 8 — Metatibia. 9 — Protarsus. 10 — Parameres, dorsal view. Scale bar in mm." httpUri="https://zenodo.org/record/191236/files/figure.png" pageId="5" pageNumber="46">Fig. 10</figureCitation>
). [Gran Canaria].
<emphasis id="B963EAE7FFCBF53F1264FCF0854F4E67" box="[633,1281,856,879]" italics="true" pageId="5" pageNumber="46">
..............................................................................
<taxonomicName id="4C174D76FFCBF53F149CFCF0854F4E67" authority="Mico" authorityName="Mico" box="[1153,1281,856,879]" class="Insecta" family="Melolonthidae" genus="Pachydema" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="megalops" status="sp. nov.">P. megalops</taxonomicName>
</emphasis>
Micó
<emphasis id="B963EAE7FFCBF53F155AFCFF85D24E67" bold="true" box="[1351,1436,855,879]" pageId="5" pageNumber="46">
<taxonomicNameLabel id="A250579CFFCBF53F155AFCFF85D24E67" box="[1351,1436,855,879]" pageId="5" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BA836F5FFCBF53F108AFCD085D24EC7" blockId="5.[151,1436,152,1391]" pageId="5" pageNumber="46">
<keyLead id="30E695C0FFCBF53F108AFCD085D24EC7" pageId="5" pageNumber="46">
- Apical maxillary palpomere small (ratio palpomere length/habitus length: 0.01) and narrow (length 3.5 times the width). Third and fourth antennomeres long and narrow (3 times longer than wide) (
<figureCitation id="132C2A70FFCBF53F1441FC3084E54EA7" box="[1116,1195,920,943]" captionStart="FIGURES 11 15. 11" captionStartId="2.[151,272,1893,1917]" captionTargetBox="[157,1419,206,1826]" captionTargetId="figure@2.[151,1436,134,1869]" captionTargetPageId="2" captionText="FIGURES 11 15. 11 — Mentum: a) P. ob s cu ra, b) P. ob s cu rell a, c) P. keithi. 12 — Galea and lacinia of maxilla: a) P. obscura, b) P. obscurella. 13 — Antenna: a) P. o b s c u r a, b) P. oromi, c) P. l e s n e i. 14 — Head: a) P. o b s c u r a, b) P. oromi, c) P. castanea. 15 — Apical maxillary palpomere: a) P. fuscipennis, b) P. o b s c u re l l a, c) P. l e s n e i, d) P. keithi, e) P. m e n i e r i, f) P. wollastoni, g) P. bipartita, h) P. navatteae. Scale bar in mm." httpUri="https://zenodo.org/record/191237/files/figure.png" pageId="5" pageNumber="46">Fig. 13</figureCitation>
c). Parameres strongly arrow shaped (
<figureCitation id="132C2A70FFCBF53F1179FC1081FD4EC7" box="[356,435,952,975]" captionStart="FIGURES 16 17. 16" captionStartId="3.[151,271,1654,1678]" captionTargetBox="[245,1345,200,1541]" captionTargetId="figure@3.[226,1360,194,1630]" captionTargetPageId="3" captionText="FIGURES 16 17. 16 — Parameres, dorsal view: a) P. obscura, b) P. tinerfensis, c) P. fortunatorum, d) P. jeannei, e) P. castanea, f) P. fuscipennis, g) P. obscurella, h) P. g o m er a e, i) P. l e s n e i, j) P. keithi, k) P. m e n i e r i, l) P. bipartita, m) P. wollastoni, n) P. navatteae. 17 — Metatibia. a) P. obscurella, b) P. ke i t h i. Scale bar in mm." httpUri="https://zenodo.org/record/191238/files/figure.png" pageId="5" pageNumber="46">
Fig.
<date id="FFA91035FFCBF53F118BFC1081FD4EC7" box="[406,435,952,975]" pageId="5" pageNumber="46" value="1916-01">16</date>
</figureCitation>
i). [Gran Canaria].
<emphasis id="B963EAE7FFCBF53F1262FC10855E4EC7" box="[639,1296,952,975]" italics="true" pageId="5" pageNumber="46">
.....................................................................................
<taxonomicName id="4C174D76FFCBF53F14A5FC10855E4EC7" authority="Peyerimhoff" authorityName="Peyerimhoff" box="[1208,1296,952,975]" class="Insecta" family="Melolonthidae" genus="Pachydema" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="lesnei">P. lesnei</taxonomicName>
</emphasis>
Peyerimhoff
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E32E50FFCBF53F108AFC7085D54947" pageId="5" pageNumber="46">
<paragraph id="8BA836F5FFCBF53F108AFC7085D24907" blockId="5.[151,1436,152,1391]" pageId="5" pageNumber="46">
<keyLead id="30E695C0FFCBF53F108AFC7085D24907" pageId="5" pageNumber="46">
12. Second protarsomere strongly widened (almost as long as wide). Prementum widely notched and with the concave area very reduced (
<figureCitation id="132C2A70FFCBF53F118EFC5081914907" box="[403,479,1016,1039]" captionStart="FIGURES 11 15. 11" captionStartId="2.[151,272,1893,1917]" captionTargetBox="[157,1419,206,1826]" captionTargetId="figure@2.[151,1436,134,1869]" captionTargetPageId="2" captionText="FIGURES 11 15. 11 — Mentum: a) P. ob s cu ra, b) P. ob s cu rell a, c) P. keithi. 12 — Galea and lacinia of maxilla: a) P. obscura, b) P. obscurella. 13 — Antenna: a) P. o b s c u r a, b) P. oromi, c) P. l e s n e i. 14 — Head: a) P. o b s c u r a, b) P. oromi, c) P. castanea. 15 — Apical maxillary palpomere: a) P. fuscipennis, b) P. o b s c u re l l a, c) P. l e s n e i, d) P. keithi, e) P. m e n i e r i, f) P. wollastoni, g) P. bipartita, h) P. navatteae. Scale bar in mm." httpUri="https://zenodo.org/record/191237/files/figure.png" pageId="5" pageNumber="46">Fig. 11</figureCitation>
c). Parameres as in
<figureCitation id="132C2A70FFCBF53F12B3FC5082B54907" box="[686,763,1016,1039]" captionStart="FIGURES 16 17. 16" captionStartId="3.[151,271,1654,1678]" captionTargetBox="[245,1345,200,1541]" captionTargetId="figure@3.[226,1360,194,1630]" captionTargetPageId="3" captionText="FIGURES 16 17. 16 — Parameres, dorsal view: a) P. obscura, b) P. tinerfensis, c) P. fortunatorum, d) P. jeannei, e) P. castanea, f) P. fuscipennis, g) P. obscurella, h) P. g o m er a e, i) P. l e s n e i, j) P. keithi, k) P. m e n i e r i, l) P. bipartita, m) P. wollastoni, n) P. navatteae. 17 — Metatibia. a) P. obscurella, b) P. ke i t h i. Scale bar in mm." httpUri="https://zenodo.org/record/191238/files/figure.png" pageId="5" pageNumber="46">Fig. 16</figureCitation>
j. [Gran Canaria]. .............................................
<taxonomicName id="4C174D76FFCBF53F14F0FC5085D24907" authority="Lacroix" authorityName="Lacroix" box="[1261,1436,1016,1039]" class="Insecta" family="Melolonthidae" genus="Pachydema" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="keithi">
<emphasis id="B963EAE7FFCBF53F14F0FC50850C4907" box="[1261,1346,1016,1039]" italics="true" pageId="5" pageNumber="46">P. keithi</emphasis>
Lacroix
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA836F5FFCBF53F108AFBB085D54947" blockId="5.[151,1436,152,1391]" pageId="5" pageNumber="46">
<keyLead id="30E695C0FFCBF53F108AFBB085D54947" pageId="5" pageNumber="46">
- Second protarsomere moderately widened (always longer than wide). Prementum shortly notched, with a wide concave area in the apical third as in
<figureCitation id="132C2A70FFCBF53F1231FB9082374947" box="[556,633,1080,1103]" captionStart="FIGURES 11 15. 11" captionStartId="2.[151,272,1893,1917]" captionTargetBox="[157,1419,206,1826]" captionTargetId="figure@2.[151,1436,134,1869]" captionTargetPageId="2" captionText="FIGURES 11 15. 11 — Mentum: a) P. ob s cu ra, b) P. ob s cu rell a, c) P. keithi. 12 — Galea and lacinia of maxilla: a) P. obscura, b) P. obscurella. 13 — Antenna: a) P. o b s c u r a, b) P. oromi, c) P. l e s n e i. 14 — Head: a) P. o b s c u r a, b) P. oromi, c) P. castanea. 15 — Apical maxillary palpomere: a) P. fuscipennis, b) P. o b s c u re l l a, c) P. l e s n e i, d) P. keithi, e) P. m e n i e r i, f) P. wollastoni, g) P. bipartita, h) P. navatteae. Scale bar in mm." httpUri="https://zenodo.org/record/191237/files/figure.png" pageId="5" pageNumber="46">Fig. 12</figureCitation>
b. ................................................................................................................ 13
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E32E50FFCBF53F108AFBF085D549A7" pageId="5" pageNumber="46">
<paragraph id="8BA836F5FFCBF53F108AFBF085D24987" blockId="5.[151,1436,152,1391]" pageId="5" pageNumber="46">
<keyLead id="30E695C0FFCBF53F108AFBF085D24987" pageId="5" pageNumber="46">
13. Apical maxillary palpomere narrowly oval (
<figureCitation id="132C2A70FFCBF53F1280FBF082A54967" box="[669,747,1112,1135]" captionStart="FIGURES 11 15. 11" captionStartId="2.[151,272,1893,1917]" captionTargetBox="[157,1419,206,1826]" captionTargetId="figure@2.[151,1436,134,1869]" captionTargetPageId="2" captionText="FIGURES 11 15. 11 — Mentum: a) P. ob s cu ra, b) P. ob s cu rell a, c) P. keithi. 12 — Galea and lacinia of maxilla: a) P. obscura, b) P. obscurella. 13 — Antenna: a) P. o b s c u r a, b) P. oromi, c) P. l e s n e i. 14 — Head: a) P. o b s c u r a, b) P. oromi, c) P. castanea. 15 — Apical maxillary palpomere: a) P. fuscipennis, b) P. o b s c u re l l a, c) P. l e s n e i, d) P. keithi, e) P. m e n i e r i, f) P. wollastoni, g) P. bipartita, h) P. navatteae. Scale bar in mm." httpUri="https://zenodo.org/record/191237/files/figure.png" pageId="5" pageNumber="46">Fig. 15</figureCitation>
e). Parameres as in
<figureCitation id="132C2A70FFCBF53F13A7FBF084484967" box="[954,1030,1112,1135]" captionStart="FIGURES 16 17. 16" captionStartId="3.[151,271,1654,1678]" captionTargetBox="[245,1345,200,1541]" captionTargetId="figure@3.[226,1360,194,1630]" captionTargetPageId="3" captionText="FIGURES 16 17. 16 — Parameres, dorsal view: a) P. obscura, b) P. tinerfensis, c) P. fortunatorum, d) P. jeannei, e) P. castanea, f) P. fuscipennis, g) P. obscurella, h) P. g o m er a e, i) P. l e s n e i, j) P. keithi, k) P. m e n i e r i, l) P. bipartita, m) P. wollastoni, n) P. navatteae. 17 — Metatibia. a) P. obscurella, b) P. ke i t h i. Scale bar in mm." httpUri="https://zenodo.org/record/191238/files/figure.png" pageId="5" pageNumber="46">Fig. 16</figureCitation>
k. [Lanzarote, Fuerteventura]. ...........
<emphasis id="B963EAE7FFCBF53F10D3FBD0850B4987" box="[206,1349,1144,1167]" italics="true" pageId="5" pageNumber="46">........................................................................................................................................................... P. m e n i e r i</emphasis>
Baraud
</keyLead>
</paragraph>
<paragraph id="8BA836F5FFCBF53F108AFB3085D549A7" blockId="5.[151,1436,152,1391]" box="[151,1435,1176,1199]" pageId="5" pageNumber="46">
<keyLead id="30E695C0FFCBF53F108AFB3085D549A7" box="[151,1435,1176,1199]" pageId="5" pageNumber="46">- Apical maxillary palpomere largely oval (Figs. fh). ................................................................................................ 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E32E50FFCBF53F108AFB1085D54807" pageId="5" pageNumber="46">
<paragraph id="8BA836F5FFCBF53F108AFB1085D549E7" blockId="5.[151,1436,152,1391]" pageId="5" pageNumber="46">
<keyLead id="30E695C0FFCBF53F108AFB1085D549E7" pageId="5" pageNumber="46">
14. Parameres slightly arrow shaped (
<figureCitation id="132C2A70FFCBF53F1255FB1082DF49C7" box="[584,657,1208,1231]" captionStart="FIGURES 16 17. 16" captionStartId="3.[151,271,1654,1678]" captionTargetBox="[245,1345,200,1541]" captionTargetId="figure@3.[226,1360,194,1630]" captionTargetPageId="3" captionText="FIGURES 16 17. 16 — Parameres, dorsal view: a) P. obscura, b) P. tinerfensis, c) P. fortunatorum, d) P. jeannei, e) P. castanea, f) P. fuscipennis, g) P. obscurella, h) P. g o m er a e, i) P. l e s n e i, j) P. keithi, k) P. m e n i e r i, l) P. bipartita, m) P. wollastoni, n) P. navatteae. 17 — Metatibia. a) P. obscurella, b) P. ke i t h i. Scale bar in mm." httpUri="https://zenodo.org/record/191238/files/figure.png" pageId="5" pageNumber="46">Fig.16</figureCitation>
l). Scutellum bordered by more or less deeply impressed groove. [
<emphasis id="B963EAE7FFCBF53F159EFB1F815D49E7" bold="true" pageId="5" pageNumber="46">El Hierro</emphasis>
,
<emphasis id="B963EAE7FFCBF53F113CFB7F818F49E7" bold="true" box="[289,449,1239,1263]" pageId="5" pageNumber="46">Gran Canaria</emphasis>
, Tenerife, La Gomera]. ...........................................................................
<taxonomicName id="4C174D76FFCBF53F14D4FB7085D549E7" authority="Brulle" authorityName="Brulle" box="[1225,1435,1240,1263]" class="Insecta" family="Melolonthidae" genus="Pachydema" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="bipartita">
<emphasis id="B963EAE7FFCBF53F14D4FB70850F49E7" box="[1225,1345,1240,1263]" italics="true" pageId="5" pageNumber="46">P. bipartita</emphasis>
(Brullé)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA836F5FFCBF53F108AFB5085D54807" blockId="5.[151,1436,152,1391]" box="[151,1435,1272,1295]" pageId="5" pageNumber="46">
<keyLead id="30E695C0FFCBF53F108AFB5085D54807" box="[151,1435,1272,1295]" pageId="5" pageNumber="46">- Parameres elongated. Scutellum without marginal groove or only present minimally at the base. .......................... 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E32E50FFCBF53F108AFAB085D24867" pageId="5" pageNumber="46">
<paragraph id="8BA836F5FFCBF53F108AFAB085D24827" blockId="5.[151,1436,152,1391]" box="[151,1436,1304,1327]" pageId="5" pageNumber="46">
<keyLead id="30E695C0FFCBF53F108AFAB085D24827" box="[151,1436,1304,1327]" pageId="5" pageNumber="46">
15. Pronotum with fine, sparse punctures. Parameres as in
<figureCitation id="132C2A70FFCBF53F130CFAB083104827" box="[785,862,1304,1327]" captionStart="FIGURES 16 17. 16" captionStartId="3.[151,271,1654,1678]" captionTargetBox="[245,1345,200,1541]" captionTargetId="figure@3.[226,1360,194,1630]" captionTargetPageId="3" captionText="FIGURES 16 17. 16 — Parameres, dorsal view: a) P. obscura, b) P. tinerfensis, c) P. fortunatorum, d) P. jeannei, e) P. castanea, f) P. fuscipennis, g) P. obscurella, h) P. g o m er a e, i) P. l e s n e i, j) P. keithi, k) P. m e n i e r i, l) P. bipartita, m) P. wollastoni, n) P. navatteae. 17 — Metatibia. a) P. obscurella, b) P. ke i t h i. Scale bar in mm." httpUri="https://zenodo.org/record/191238/files/figure.png" pageId="5" pageNumber="46">
Fig.
<specimenCount id="9D11FD7CFFCBF53F135FFAB083104827" box="[834,862,1304,1327]" pageId="5" pageNumber="46" type="generic">16</specimenCount>
</figureCitation>
n. [Gran Canaria, Lanzarote].
<emphasis id="B963EAE7FFCBF53F14BFFAB085074827" box="[1186,1353,1304,1327]" italics="true" pageId="5" pageNumber="46">
.....
<taxonomicName id="4C174D76FFCBF53F14DBFAB085074827" authority="Baraud" authorityName="Baraud" box="[1222,1353,1304,1327]" class="Insecta" family="Melolonthidae" genus="Pachydema" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="navatteae">P. navatteae</taxonomicName>
</emphasis>
Baraud
</keyLead>
</paragraph>
<paragraph id="8BA836F5FFCBF53F108AFA9085D24867" blockId="5.[151,1436,152,1391]" pageId="5" pageNumber="46">
<keyLead id="30E695C0FFCBF53F108AFA9085D24867" pageId="5" pageNumber="46">
- Pronotal punctures moderate to coarse, densely distributed. Parameres as in
<figureCitation id="132C2A70FFCBF53F13E9FA90840E4847" box="[1012,1088,1336,1359]" captionStart="FIGURES 16 17. 16" captionStartId="3.[151,271,1654,1678]" captionTargetBox="[245,1345,200,1541]" captionTargetId="figure@3.[226,1360,194,1630]" captionTargetPageId="3" captionText="FIGURES 16 17. 16 — Parameres, dorsal view: a) P. obscura, b) P. tinerfensis, c) P. fortunatorum, d) P. jeannei, e) P. castanea, f) P. fuscipennis, g) P. obscurella, h) P. g o m er a e, i) P. l e s n e i, j) P. keithi, k) P. m e n i e r i, l) P. bipartita, m) P. wollastoni, n) P. navatteae. 17 — Metatibia. a) P. obscurella, b) P. ke i t h i. Scale bar in mm." httpUri="https://zenodo.org/record/191238/files/figure.png" pageId="5" pageNumber="46">
Fig.
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m. [Fuerteventura]. ...................
<emphasis id="B963EAE7FFCBF53F10D3FAF0855E4867" box="[206,1296,1368,1391]" italics="true" pageId="5" pageNumber="46">
...............................................................................................................................................
<taxonomicName id="4C174D76FFCBF53F149AFAF0855E4867" authority="Peyerimhoff" authorityName="Peyerimhoff" box="[1159,1296,1368,1391]" class="Insecta" family="Melolonthidae" genus="Pachydema" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="wollastoni">P. wollastoni</taxonomicName>
</emphasis>
Peyerimhoff
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>